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Texto Completo (Pdf) Acta Botanica Malacitana 44 (2019). 93-97 Nota corta C. DOI: Romero-Zarco http://dx.doi.org/10.24310/abm.v44i0.5331 et al. Notas nomenclaturales gramíneas mediterráneas 93 Notas nomenclaturales sobre algunas gramíneas mediterráneas Carlos Romero-Zarco1, Juan Luis García-Castaño1 & Regina Berjano Pérez1,2 1Departamento de Biología Vegetal y Ecología, Universidad de Sevilla 2Departamento de Botánica, Ecología y Fisiología Vegetal, Universidad de Córdoba Correspondencia Nomenclatural notes on some Mediterranean grasses C. Romero Zarco e-mail: [email protected] Palabras clave: Nomenclatura, tipificación, Gramineae, Poaceae, región Recibido: 5 noviembre 2018 mediterránea Aceptado: 2 septiembre 2019 Publicado on-line: 25 septiembre 2019 Key words: Nomenclature, typification, Gramineae, Poaceae, Mediterranean Editado por: B. Cabezudo region Durante la elaboración de los manuscritos Basiónimo: Arrhenatherum fernandesii Rivas destinados al volumen XIX de Flora iberica Mart. in Itinera Geobot. 15: 698 (2002) (Castroviejo, 1986 en adelante) se han resuelto Arrhenatherum erianthum var. montanum R. algunas cuestiones de nomenclatura o tipificación Fernandes in Mem. Soc. Brot. 6: 10 (1950) [syn. que, siguiendo la tradición del proyecto, es subst.] preferible publicar previamente en formato de Arrhenatherum album var. montanum (R. notas breves. Fernandes) P. Silva in Agron. Lusit. 40: 6 (1980) Indicación locotípica: “prope Caldeirão, circa Airopsis tenella (Cav.) Asch. & Graebn., Syn. Guarda” [actualmente embalse de Caldeirão en la Mitteleur. Fl. 2(1): 298 (1899) Beira Alta] Millium tenellum Cav., Icon. 3: 37, t. 274 (1796) Lectotipo: COI 9526 (photographiam vidit), [basión.] designado aquí Airopsis globosa (Thore) Desv., Journ. Bot. Isolectotipi: LISI 3202-A!, LISU P 17030!, (Desvaux) 1: 200 (1809) designados aquí Indicación locotípica: “Habitat in arenosis prope Las plantas de esta subespecie se caracterizan oppidum vulgo Pobla tornesa” [La Pobla Tornesa, por la ausencia –o la reducción extrema– de los Castellón]. tubérculos basales que se desarrollan siempre Lectotipo: MA 475983!, designado aquí en Arrhenatherum album subsp. album. Son Según el catálogo de los materiales de A. frecuentes en las crestas cuarcíticas de la parte J. Cavanilles (cf. Garilleti, 1993) el ejemplar occidental del Sistema Central y en otras zonas designado aquí como lectotipo es el único material occidentales de los valles del Duero, del Tajo y del original encontrado en el Herbario del Jardín Guadiana [Esp.: Ba Cc Sa Za. Port.: AAl BA TM]; Botánico de Madrid (MA). Paunero (1957: 189) al parecer se trata de un taxón diploide (n = 7, cf. citó material de esa localidad en su monografía Devesa & al., 1990 sub A. elatius subsp. sardoum, sin indicar su número de registro en el herbario UNEX 8431!). Las características morfológicas de ni tipificación alguna. Hay otro material original, estas plantas y su distribución geográfica hacen presumiblemente enviado por el mismo Cavanilles más recomendable un tratamiento subespecífico a Londres (LINN-Smith 121.9, “Spain-Cavanilles- dentro de A. album (Vahl) W.D. Clayton, en lugar del 1793.-N.º 8.”, cf. The Linnean Collections-Smith tratamiento específico o varietal de las anteriores Herbarium, photographiam vidit) pero parece más propuestas. La existencia de un Arrhenatherum recomendable elegir como lectotipo el material de montanum (Dumort.) Potztal desaconseja el uso la colección principal de Cavanilles, donde figura del epíteto montanum en este género. En su lugar expresamente la localidad mencionada en el se prefiere el epíteto fernandesii, anteriormente protólogo. usado por Rivas Martínez en honor de la botánica que describió el taxón. Arrhenatherum album subsp. fernandesii (Rivas Mart.) Romero Zarco, comb. et. stat. nov. Arrhenatherum elatius subsp. cypricola (H. 94 Acta Botanica Malacitana, 44, 2019 Scholz) Romero Zarco, comb. nov. Presl”. El primer nombre corresponde a la especie Basiónimo: Arrhenatherum album subsp. que hoy llamamos Helictotrichon sempervirens cypricola H. Scholz in Willdenowia 37: 211 (2007) (Vill.) Pilg. (cf. Index Synonymique de la Flore de Indicación locotípica: “Cyprus, Kaminaria, France) y el segundo a H. convolutum (C. Presl) Platys valley, at the picnic site Komititzi, W of the Henrard, ninguna de ellas presentes en España. A summit Agios Ilias, dry open slope, 700 m.” continuación separa dos variedades. La primera la Holotipo: Hand 4809, 5.5.2005 (B) denomina var. glabra, donde se podría interpretar Taxón endémico de la isla de Chipre, que se que incluye los sinónimos citados para la especie, caracteriza principalmente por tener el artejo de y la segunda la describe como var. velutina “in la raquilla muy corto (c. 0,4 mm), carácter que alpinâ Sierra Tejeda”, actualmente aceptada como lo aproxima más a A. elatius (L.) J. Presl & C. especie [Helictotrichon sarracenorum (Gand.) Presl que a A. album. El lema de la flor superior Holub]. Sin embargo, en las observaciones al pie es más densamente peloso que en otros taxones de la descripción comenta que las plantas de Italia mediterráneos que se suelen incluir en esta pertenecen probablemente a una tercera variedad. especie, lo que tal vez hizo que H. Scholz se Por tanto, entre estos elementos del protólogo, solo inclinara por clasificar esta subespecie enA. album, las plantas observadas por él mismo en Andalucía cuyas espiguillas sin embargo son muy diferentes y la referencia al tipo de Lagasca –“in Hispanieae debido al artejo de (0,8)1-2 mm que separa ambas regno Granatensi, Murcico (Lag.)”– deben flores (Romero-Zarco, 1985). Esta subespecie tenerse en consideración para tipificar el nombre podría representar el taxón oriental de un conjunto varietal. Se da la circunstancia de que el lectotipo de taxones mediterráneos vicariantes del que designado por Röser (1989) para Avena filifolia formarían parte A. elatius subsp. baeticum Romero Lag. es precisamente un ejemplar del herbario Zarco en la cuenca mediterránea occidental y de Boissier que ya se comentó en la revisión del A. elatius subsp. nebrodense (Brullo, Miniss. & género para la Península Ibérica (Romero-Zarco, Spamp.) Giardina & Raimondo en Sicilia, todas 1984): “Avena filifolia Lag. // dedit Lagasca”. Parece ellas formas bulbosas diploides. lógico designar ese mismo ejemplar como lectotipo de la primera variedad de la especie, aunque eso Arrhenatherum riofrioi Sennen, Diagn. Nouv.: suponga establecer su ilegitimidad. 168 (1936) Nombre aceptado: Arrhenatherum album Corynephorus articulatus (Desf.) P. Beauv., (Vahl) W.D. Clayton var. album Agrost. 90 (1812) Indicación locotípica: “Maroc: Melilla, Gurugu, Aira articulata Desf., Fl. Atlant. 1: 70 (1798) ampelodesmatum, vers 1650 m.” [basión.] Lectotipo: Sennen & Mauricio, F. Sennen Pl. C. divaricatus auct. fl. europ., non Aira Esp. 1932, n° 8557, BC 141849!, designado aquí divaricata Pourr. El exsiccatum de Sennen contiene plantas Indicación locotípica: “Habitat in arvis prope de pequeño porte y espiguillas relativamente Mascar” [Muaskar, Argelia] pequeñas, idénticas a los diploides de la especie Lectotipo: P 00320332 (photographiam vidit), estudiados en los Sistemas Béticos (cf. Romero- designado aquí Zarco, 1985). El género comprende seis especies de origen y distribución principalmente mediterránea, una Avena filifolia var. glabra Boiss., Voy. Bot. perenne –C. canescens (L.) P. Beauv.– de más Espagne 2: 655 (1844) amplia distribución en Europa, y cinco anuales. Nombre aceptado: Helictotricon filifolium (Lag.) Murbeck (1900) hizo un tratamiento –aún válido– Henrard in Blumea 3: 430 (1940) var. filifolium de las especies anuales del género, incluyendo las Avena filifolia Lag., Gen. Sp. Pl.: 4 (1816) tres especies aceptadas presentes en la Península Indicación locotípica: “varietas a in regione Ibérica [C. articulatus, C. macrantherus Boiss. & montanâ[…] Hab. species in Hispaniae regno Reut. y C. divaricatus (Pourr.) Breistr.], junto con Granatensi, Murcico (Lag.), in montibus Calabriae otras dos especies del centro y del este de la región (Ten.), Siciliae (Guss.).” mediterránea, respectivamente: C. oranensis Lectotipo: “Avena filifolia Lag. // dedit Lagasca” Murb. y C. deschampsioides Bornm. (sub C. laxus (G-Boissier 165448!), designado aquí Murb.). Sin embargo, el trabajo de Murbeck no Burdet & al. (1981) no encontraron ningún tuvo la repercusión que merecía, pues los autores material original correspondiente al protólogo. posteriores consideraron una sola especie anual Boissier (l. c.) incluye dos sinónimos para la especie: en el género: C. articulatus s. l. Este fue el caso “A. [Avena] fallax Ten. Guss. Pl. rar. t. 9. A. convoluta de Breistroffer (1950), quien rescató Aira divaricata C. Romero-Zarco et al. Notas nomenclaturales gramíneas mediterráneas 95 Pourr. de la sinonimia de Aira caryophyllea L. y pliego contiene una panícula fértil y una hoja basal. propuso la combinación Corynephorus divaricatus Tuvo que ser incorporado al herbario entre 1800 y (Pourr.) Breist. para sustituir a C. articulatus s. 1830, y posiblemente fue adquirido en los jardines l. De todos los epítetos específicos aplicables botánicos de Kew, París o Madrid (F.J. Salgueiro, a las especies anuales de Corynephorus, el com. pers.). de Pourret resultó ser el más antiguo. Paunero (1956), haciéndose eco del trabajo de Breistroffer, Helictotrichon thorei Röser in Schlechtendalia reconoció dos especies anuales, aplicando 19: 30 (2009) el epíteto “divaricatus” de Pourret a lo que se Avena longifolia Thore, Prom. Golfe Gascogne: había llamado anteriormente “articulatus” de 92
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