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Revista Mexicana de Biodiversidad ISSN: 1870-3453 [email protected] Universidad Nacional Autónoma de México México García Aldrete, Alfonso N.; Mockford, Edward L. A list of Psocoptera (Insecta: Psocodea) from Brazil Revista Mexicana de Biodiversidad, vol. 80, núm. 3, 2009, pp. 665-673 Universidad Nacional Autónoma de México Distrito Federal, México Available in: http://www.redalyc.org/articulo.oa?id=42515996009 How to cite Complete issue Scientific Information System More information about this article Network of Scientific Journals from Latin America, the Caribbean, Spain and Portugal Journal's homepage in redalyc.org Non-profit academic project, developed under the open access initiative Revista Mexicana de Biodiversidad 80: 665- 673, 2009 A list of Psocoptera (Insecta: Psocodea) from Brazil Listado de Psocoptera (Insecta: Psocodea) de Brasil Alfonso N. García Aldrete1* and Edward L. Mockford 2 1Instituto de Biología, Universidad Nacional Autónoma de México, Apartado postal 70-153, 04510 México, D. F., Mexico. 2Department of Biological Sciences, Illinois State University, Campus Box 4120, Normal, Illinois 61790-4120, USA. *Correspondent: [email protected] Abstract. The species of Psocoptera currently known for Brazil are listed, with state distribution and biogeographic status. The categories of geographic distribution are discussed, as well as some of the evidence indicating that the present size of the Brazilian fauna is underestimated. Key words: Psocodea, Psocoptera, Brazil, geographic distribution. Resumen. Se listan las especies de Psocoptera actualmente registradas en Brasil, incluyendo la distribución por estado y su categoría biogeográfi ca. Se presenta alguna de la evidencia que hace suponer que el tamaño actual de la fauna de Psocoptera de Brasil está subestimada. Palabras clave: Psocodea, Psocoptera, Brasil, distribución geográfi ca. Introduction de Feira de Santana, Bahia, Brasil. The distributions in Table 1 were taken from the literature and from specimens The Catalogue of the Psocoptera (Lienhard and in collections (mostly INPA and Colección Nacional Smithers, 2002) lists for Brazil 374 species in 87 genera de Insectos, CNIN, Instituto de Biología, Universidad and 29 families. We have updated this information, and Nacional Autónoma de México, México City). the list presented here includes 425 species in 94 genera and 28 families (Table 1). The ‘Psocoptera’, a paraphyletic group, are the non-parasitic members of the order Psocodea Results (sensu Yoshizawa and Johnson, 2006: 358), which comprises also the ectoparasitic chewing and sucking Table 1 presents the list of species of Psocoptera lice (‘Phthiraptera’) (see Johnson et al., 2004). The list known in Brazil; 425 species are listed, in 94 genera and 28 below is to be considered as a work in progress subject to families. The largest and most diverse family is Psocidae, modifi cation as more information on the Brazilian psocid with 89 species in 23 genera, followed by Ptiloneuridae fauna becomes available. (50 species in 5 genera), Epipsocidae (48 species in 6 genera), and Lachesillidae (41 species in 6 genera). The most speciose genera (Lachesilla, Triplocania, Epipsocus, Material and methods Archipsocus, Mesepipsocus, and Valenzuela) account for 135 species, and on the other extreme, 32 genera are The updating of the information on the Brazilian fauna represented by 1 species and 11 genera have 2 species. in Lienhard and Smithers (2002) is derived mostly from It is likely that Notolepium brasiliense New will have the ongoing study of the vast Psocoptera collection in the to be transferred to Thylacella (cf. Mockford, 2005). Instituto Nacional de Pesquisas da Amazônia (INPA), in Twelve additional genera (Thylacella, Cyptophania, Manaus, Amazonas, and from the study of the Psocoptera Neolepolepis, Lepinotus, Nanopsocus, Peritroctes, collected through the program PPBio, housed in the Epitroctes, Compsocus, Spurostigma, Pararchipsocus, Laboratory of Insect Systematics, Universidade Estadual Tricholachesilla and Palmicola) probably occur in Brazil (García Aldrete and Mockford, unpublished data), but they Recibido: 14 octubre 2008; aceptado: 14 enero 2009 have not yet been detected. 666 García Aldrete and Mockford.- Psocoptera from Brazil Table 1. Psocoptera recorded from Brazil, distribution and biogeographic status. The species not listed by Lienhard and Smithers (2002) are marked with an asterisk (*). Most of them are new to Brazil, otherwise the references to published records are given Trogiomorpha 27. S. fasciatus Mockford. Roraima (Endemic). Lepidopsocidae 28. S. rafaeli Mockford. Roraima (Endemic). 1. Nepticulomima brasiliensis (Enderlein). Pará 29. S. rotundatus Roesler. Santa Catarina (Endemic). (Endemic). Liposcelididae 2. N. pulvillata New. Mato Grosso (Endemic). 30. Belaphopsocus badonneli New. Mato Grosso (Brazil, 3. Notolepium brasiliense New. Mato Grosso (Endemic). Colombia, Mexico, Paraguay). 4. Proentomum personatum Badonnel. Roraima (Tropical 31. Belaphotroctes brunneus Badonnel. Sao Paulo waif). (Endemic). 5. Soa fl aviterminata Enderlein. Amazonas, Pará, 32. B. ghesquierei Badonnel. Mato Grosso, Roraima, Sao Roraima (Widely distributed). Paulo (Widespread in tropical Africa and America). 6. S. violacea New. Amazonas, Mato Grosso (Endemic). 33. B. major Badonnel. Mato Grosso (Endemic). 7. Echmepteryx angusta New. Mato Grosso (Endemic). 34. B. mimulus Badonnel. Sao Paulo (Endemic). 8. E. bishopi New. Mato Grosso (Endemic). 35. B. ocularis Badonnel. Sao Paulo (Endemic). 9. E. falco (Badonnel). Amazonas, Bahia (Tropical 36. B. striatus Badonnel. Sao Paulo (Endemic). waif). 37. B. vaginatus Badonnel. Sao Paulo (Endemic). 10. E. fuscata New. Mato Grosso (Endemic). 38. Belapha sp. Pará (Endemic). (*). New for Brazil. 11. E. lealae New. Pernambuco (Endemic). 39. Chaetotroctes lenkoi Badonnel. Mato Grosso 12. E. lutosa Mockford. Roraima (Endemic). (Endemic). 13. E. madagascariensis (Kolbe). Rio de Janeiro, Bahia 40. Embidopsocopsis newi Badonnel. Mato Grosso (Tropical waif). (*). New for Brazil. (Endemic). 14. E. uniformis Mockford. Roraima (Endemic). 41. Embidopsocus brasiliensis Badonnel. Sao Paulo 15. Pteroxanium sp. “Brazil” (Endemic). (Endemic). Trogiidae 42. E. fl exuosus parvulus Badonnel. Mato Grosso 16. Cerobasis guestfalica (Kolbe). Pernambuco, Rio de (Endemic). Janeiro (Widely distributed). 43. E. frater Badonnel. Sao Paulo (Endemic). Psoquillidae 44. E. leucomelas (Enderlein). Mato Grosso (Brazil, 17. Psoquilla marginepunctata Hagen. Mato Grosso, Santa Paraguay). Catarina (Widely distributed). 45. E. luteus Hagen. Mato Grosso (Brazil, Cuba, Mexico). 18. Rhyopsocus rafaeli García Aldrete. Amazonas 46. E. mendax Badonnel. Mato Grosso, Sao Paulo (Brazil, (Endemic). (*). See García Aldrete (2004a). Argentina). Psyllipsocidae 47. E. pilosus Badonnel. Mato Grosso, Sao Paulo 19. Psocathropos lachlani Ribaga. Amazonas, Sao Paulo (Endemic). (Widely distributed). 48. E. similis Badonnel. Mato Grosso (Endemic). 20. Psyllipsocus yucatan Gurney. Pará (Brazil, Mexico). 49. E. virgatus (Enderlein). Mato Grosso, Sao Paulo (*). New for Brazil. (Brazil, Argentina, Paraguay). Troctomorpha 50. Liposcelis bostrychophila Badonnel. Rio de Janeiro Troctopsocidae (Cosmopolitan). 21. Troctopsoculus brasiliensis New. Mato Grosso (Brazil, 51. L. entomophila (Enderlein). Amazonas, Rio de Janeiro Mexico). (Cosmopolitan). (*). New for Brazil. 22. Troctopsocus separatus (Roesler). Santa Catarina 52. L. fuscipes Badonnel. Sao Paulo (Brazil, Mexico). (Brazil, Trinidad). 53. L. lenkoi Badonnel. Sao Paulo (Endemic). Musapsocidae Pachytroctidae 23. Musapsocus mockfordi New. Amazonas (Endemic). 54. Tapinella gamma Mockford. Roraima (Endemic). 24. M. newi Mockford. Roraima (Endemic). 55. T. maracana Mockford. Roraima (Endemic). Amphientomidae 56. T. ornaticeps Mockford. Roraima (Brazil, Venezuela). 25. Seopsocus acuminatus Roesler. Santa Catarina 57. Antilopsocus nadleri Gurney. Mato Grosso (Brazil, (Endemic). Trinidad). 26. S. albiceps Mockford. Roraima (Endemic). 58. Pachytroctes brasilianus Roesler. Santa Catarina Revista Mexicana de Biodiversidad 80: 665- 673, 2009 667 (Brazil, Venezuela). 101. Neurostigma dispositum Roesler. Amazonas, Mato 59. Psyllotroctes plaumanni Roesler. Santa Catarina Grosso, Santa Catarina, Sao Paulo (Brazil, Mexico, (Endemic). Perú). Psocomorpha 102. N. enderleini New. Amazonas (Endemic). Epipsocidae 103. N. paucivenosum New. Amazonas (Endemic). 60. Epipsocus acanthus New. Amazonas (Endemic). 104. N. radiatum Mockford. Roraima (Endemic). 61. E. argutus New. Amazonas (Endemic). 105. N. roesleri New. Amazonas (Endemic). 62. E. atratus New. Amazonas, Roraima (Endemic). 106. N. xanthopterum New. Amazonas (Endemic). 63. E. badonneli Mockford. Pará, Roraima (Endemic). 107. Papillopsocus oriximinaensis García Aldrete. Pará 64. E. borgmeieri Karny. Rio de Janeiro (Endemic). (Endemic). (*). See García Aldrete 2005. 65. E. foliatus Mockford. Roraima (Endemic). Dolabellopsocidae 66. E. fuscareolatus New. Amazonas (Endemic). 108. Dolabellopsocus carcinus Mockford. Roraima 67. E. latistigma Roesler. Santa Catarina (Endemic). (Endemic). 68. E. maculithorax New. Amazonas (Endemic). 109. D. catenatus Mockford. Roraima (Endemic). 69. E. meruleus New. Amazonas (Endemic). 110. D. ctenatus (New). Amazonas, Bahia, Mato Grosso 70. E. pennyi New. Amazonas (Endemic). (Brazil, Perú). 71. E. pereirai Badonnel. Amazonas, Mato Grosso 111. D. eertmoedi Badonnel. Mato Grosso (Endemic). (Endemic). 112. D. fl avipennis (Roesler). Santa Catarina (Endemic). 72. E. phaeus
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    NINE Phylum ARTHROPODA SUBPHYLUM HEXAPODA Protura, springtails, Diplura, and insects ROD P. MACFARLANE, PETER A. MADDISON, IAN G. ANDREW, JOCELYN A. BERRY, PETER M. JOHNS, ROBERT J. B. HOARE, MARIE-CLAUDE LARIVIÈRE, PENELOPE GREENSLADE, ROSA C. HENDERSON, COURTenaY N. SMITHERS, RicarDO L. PALMA, JOHN B. WARD, ROBERT L. C. PILGRIM, DaVID R. TOWNS, IAN McLELLAN, DAVID A. J. TEULON, TERRY R. HITCHINGS, VICTOR F. EASTOP, NICHOLAS A. MARTIN, MURRAY J. FLETCHER, MARLON A. W. STUFKENS, PAMELA J. DALE, Daniel BURCKHARDT, THOMAS R. BUCKLEY, STEVEN A. TREWICK defining feature of the Hexapoda, as the name suggests, is six legs. Also, the body comprises a head, thorax, and abdomen. The number A of abdominal segments varies, however; there are only six in the Collembola (springtails), 9–12 in the Protura, and 10 in the Diplura, whereas in all other hexapods there are strictly 11. Insects are now regarded as comprising only those hexapods with 11 abdominal segments. Whereas crustaceans are the dominant group of arthropods in the sea, hexapods prevail on land, in numbers and biomass. Altogether, the Hexapoda constitutes the most diverse group of animals – the estimated number of described species worldwide is just over 900,000, with the beetles (order Coleoptera) comprising more than a third of these. Today, the Hexapoda is considered to contain four classes – the Insecta, and the Protura, Collembola, and Diplura. The latter three classes were formerly allied with the insect orders Archaeognatha (jumping bristletails) and Thysanura (silverfish) as the insect subclass Apterygota (‘wingless’). The Apterygota is now regarded as an artificial assemblage (Bitsch & Bitsch 2000).
  • Kenai National Wildlife Refuge Species List, Version 2018-07-24

    Kenai National Wildlife Refuge Species List, Version 2018-07-24

    Kenai National Wildlife Refuge Species List, version 2018-07-24 Kenai National Wildlife Refuge biology staff July 24, 2018 2 Cover image: map of 16,213 georeferenced occurrence records included in the checklist. Contents Contents 3 Introduction 5 Purpose............................................................ 5 About the list......................................................... 5 Acknowledgments....................................................... 5 Native species 7 Vertebrates .......................................................... 7 Invertebrates ......................................................... 55 Vascular Plants........................................................ 91 Bryophytes ..........................................................164 Other Plants .........................................................171 Chromista...........................................................171 Fungi .............................................................173 Protozoans ..........................................................186 Non-native species 187 Vertebrates ..........................................................187 Invertebrates .........................................................187 Vascular Plants........................................................190 Extirpated species 207 Vertebrates ..........................................................207 Vascular Plants........................................................207 Change log 211 References 213 Index 215 3 Introduction Purpose to avoid implying
  • Morphology of Psocomorpha (Psocodea: 'Psocoptera')

    Morphology of Psocomorpha (Psocodea: 'Psocoptera')

    Title MORPHOLOGY OF PSOCOMORPHA (PSOCODEA: 'PSOCOPTERA') Author(s) Yoshizawa, Kazunori Insecta matsumurana. New series : journal of the Faculty of Agriculture Hokkaido University, series entomology, 62, 1- Citation 44 Issue Date 2005-12 Doc URL http://hdl.handle.net/2115/10524 Type bulletin (article) File Information Yoshizawa-62.pdf Instructions for use Hokkaido University Collection of Scholarly and Academic Papers : HUSCAP INSECTA MATSUMURANA NEW SERIES 62: 1–44 DECEMBER 2005 MORPHOLOGY OF PSOCOMORPHA (PSOCODEA: 'PSOCOPTERA') By KAZUNORI YOSHIZAWA Abstract YOSHIZAWA, K. 2005. Morphology of Psocomorpha (Psocodea: 'Psocoptera'). Ins. matsum. n. s. 62: 1–44, 24 figs. Adult integumental morphology of the suborder Psocomorpha (Psocodea: 'Psocoptera') was examined, and homologies and transformation series of characters throughout the suborder and Psocoptera were discussed. These examinations formed the basis of the recent morphology-based cladistic analysis of the Psocomorpha (Yoshizawa, 2002, Zool. J. Linn. Soc. 136: 371–400). Author's address. Systematic Entomology, Graduate School of Agriculture, Hokkaido University, Sapporo, 060-8589 Japan. E-mail. [email protected]. 1 INTRODUCTION Psocoptera (psocids, booklice or barklice) are a paraphyletic assemblage of non-parasitic members of the order Psocodea (Lyal, 1985; Yoshizawa & Johnson, 2003, 2005; Johnson et al., 2004), containing about 5500 described species (Lienhard, 2003). They are about 1 to 10 mm in length and characterized by well-developed postclypeus, long antennae, pick-like lacinia, reduced prothorax, well-developed pterothorax, etc. Phylogenetically, Psocoptera compose a monophyletic group (the order Psocodea) with parasitic lice ('Phtiraptera': biting lice and sucking lice) (Lyal, 1985; Yoshizawa & Johnson, 2003, in press; Johnson et al., 2004). The order is related to Thysanoptera (thrips) and Hemiptera (bugs, cicadas, etc.) (Yoshizawa & Saigusa, 2001, 2003, but see also Yoshizawa & Johnson, 2005).
  • Along the Salt Fork River at Champaign County Forest Preserve Districtbhomer Lake (Champaign County)

    Along the Salt Fork River at Champaign County Forest Preserve Districtbhomer Lake (Champaign County)

    THE BIOLOGY OF TRICHADENOTECNUM ALEXANDERAE SOMMERMAN (PSOCOPTERA: PSOCIDAE). I11. ANALYSIS OF MATING BEHAVIOR By B. W. BETZ INTRODUCTION Several authors have described mating behavior in species of Pso- coptera (Pearman 1928, Sommerman 1943a, 1943b, 1944, 1956, Badonnel 1951, Thornton and Broadhead 1954, Klier 1956, Mock- ford 1957, 1977, Broadhead 1961, Eertmoed 1966). Only one or at most a few matings in a species were observed. This paper presents a comprehensive analysis of pre- through post-copulatory behavior in Trichadenotecnum alexanderae Sommerman. Evidence is presented for a sex-attractant pheromone, produced only by females that were receptive to mating. Trichadenotecnum alexanderae is a relatively common psocid in eastern United States (Betz 1983a). The species inhabits trees and rock outcroppings providing its principal food source, pleurococ- cine algae. Betz (1983a) found that T. alexanderae is capable of facultative thelytoky. Formerly, the species was confused morpho- logically with three other species, all obligatorily thelytokous, which have been identified and described as T. castum Betz, T. merum Betz, and T. innuptum Betz (Betz 1983a). This paper is part of a series (cf. Betz 1983b, c, d) detailing the life history of T. alexanderae. MATERIALS AND METHODS Cultures of T. alexanderae were obtained from three populations in Illinois: at Moraine View State Park (McLean County), along the Sangamon River at Lake of the Woods (Champaign County), and along the Salt Fork River at Champaign County Forest Preserve DistrictBHomer Lake (Champaign County). Specimens were collected from tree trunks with an aspirator and kept with pieces of bark in cotton-stoppered test tubes. Cultures were transported to the laboratory over ice-water in a cooler.