Downloaded from Brill.Com09/30/2021 07:16:35PM Via Free Access 278 H.-P
Total Page:16
File Type:pdf, Size:1020Kb
Load more
Recommended publications
-
Early Tetrapod Relationships Revisited
Biol. Rev. (2003), 78, pp. 251–345. f Cambridge Philosophical Society 251 DOI: 10.1017/S1464793102006103 Printed in the United Kingdom Early tetrapod relationships revisited MARCELLO RUTA1*, MICHAEL I. COATES1 and DONALD L. J. QUICKE2 1 The Department of Organismal Biology and Anatomy, The University of Chicago, 1027 East 57th Street, Chicago, IL 60637-1508, USA ([email protected]; [email protected]) 2 Department of Biology, Imperial College at Silwood Park, Ascot, Berkshire SL57PY, UK and Department of Entomology, The Natural History Museum, Cromwell Road, London SW75BD, UK ([email protected]) (Received 29 November 2001; revised 28 August 2002; accepted 2 September 2002) ABSTRACT In an attempt to investigate differences between the most widely discussed hypotheses of early tetrapod relation- ships, we assembled a new data matrix including 90 taxa coded for 319 cranial and postcranial characters. We have incorporated, where possible, original observations of numerous taxa spread throughout the major tetrapod clades. A stem-based (total-group) definition of Tetrapoda is preferred over apomorphy- and node-based (crown-group) definitions. This definition is operational, since it is based on a formal character analysis. A PAUP* search using a recently implemented version of the parsimony ratchet method yields 64 shortest trees. Differ- ences between these trees concern: (1) the internal relationships of aı¨stopods, the three selected species of which form a trichotomy; (2) the internal relationships of embolomeres, with Archeria -
Physical and Environmental Drivers of Paleozoic Tetrapod Dispersal Across Pangaea
ARTICLE https://doi.org/10.1038/s41467-018-07623-x OPEN Physical and environmental drivers of Paleozoic tetrapod dispersal across Pangaea Neil Brocklehurst1,2, Emma M. Dunne3, Daniel D. Cashmore3 &Jӧrg Frӧbisch2,4 The Carboniferous and Permian were crucial intervals in the establishment of terrestrial ecosystems, which occurred alongside substantial environmental and climate changes throughout the globe, as well as the final assembly of the supercontinent of Pangaea. The fl 1234567890():,; in uence of these changes on tetrapod biogeography is highly contentious, with some authors suggesting a cosmopolitan fauna resulting from a lack of barriers, and some iden- tifying provincialism. Here we carry out a detailed historical biogeographic analysis of late Paleozoic tetrapods to study the patterns of dispersal and vicariance. A likelihood-based approach to infer ancestral areas is combined with stochastic mapping to assess rates of vicariance and dispersal. Both the late Carboniferous and the end-Guadalupian are char- acterised by a decrease in dispersal and a vicariance peak in amniotes and amphibians. The first of these shifts is attributed to orogenic activity, the second to increasing climate heterogeneity. 1 Department of Earth Sciences, University of Oxford, South Parks Road, Oxford OX1 3AN, UK. 2 Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstraße 43, 10115 Berlin, Germany. 3 School of Geography, Earth and Environmental Sciences, University of Birmingham, Birmingham B15 2TT, UK. 4 Institut -
1 1 Appendix S1: Complete List of Characters And
1 1 Appendix S1: Complete list of characters and modifications to the data matrix of RC07, with 2 reports of new observations of specimens. 3 The names, the abbreviations and the order of all characters and their states are 4 unchanged from RC07 unless a change is explained. We renumbered the characters we did 5 not delete from 1 to 277, so the character numbers do not match those of RC07. However, 6 merged characters retain the abbreviations of all their components: PREMAX 1-2-3 (our 7 character 1) consists of the characters PREMAX 1, PREMAX 2 and PREMAX 3 of RC07, 8 while MAX 5/PAL 5 (our ch. 22) is assembled from MAX 5 and PAL 5 of RC07. We did not 9 add any characters, except for splitting state 1 of INT FEN 1 into the new state 1 of INT FEN 10 1 (ch. 84) and states 1 and 2 of the new character MED ROS 1 (ch. 85), undoing the merger 11 of PIN FOR 1 and PIN FOR 2 (ch. 91 and 92) and splitting state 0 of TEETH 3 into the new 12 state 0 of TEETH 3 (ch. 183) and the entire new character TEETH 10 (ch. 190). A few 13 characters have additional states or are recoded in other ways. Deleted characters are retained 14 here, together with the reasons why we deleted them and the changes we made to their scores. 15 All multistate characters mention in their names whether they are ordered, unordered, 16 or treated according to a stepmatrix. -
(Temnospondyli), and the Evolution of Modern
THE LOWER PERMIAN DISSOROPHOID DOLESERPETON (TEMNOSPONDYLI), AND THE EVOLUTION OF MODERN AMPHIBIANS Trond Sigurdsen Department of Biology McGill University, Montreal November 2009 A thesis submitted to McGill University in partial fulfillment of the requirements of the degree of Doctor of Philosophy © Trond Sigurdsen 2009 1 ACKNOWLEDGMENTS I am deeply grateful to my supervisors Robert L. Carroll and David M. Green for their support, and for revising and correcting the drafts of the individual chapters. Without their guidance, encouragement, and enthusiasm this project would not have been possible. Hans Larsson has also provided invaluable help, comments, and suggestions. Special thanks go to John R. Bolt, who provided specimens and contributed to Chapters 1 and 3. I thank Farish Jenkins, Jason Anderson, and Eric Lombard for making additional specimens available. Robert Holmes, Jean-Claude Rage, and Zbyněk Roček have all provided helpful comments and observations. Finally, I would like to thank present and past members of the Paleolab at the Redpath Museum, Montreal, for helping out in various ways. Specifically, Thomas Alexander Dececchi, Nadia Fröbisch, Luke Harrison, Audrey Heppleston and Erin Maxwell have contributed helpful comments and technical insight. Funding was provided by NSERC, the Max Stern Recruitment Fellowship (McGill), the Delise Allison and Alma Mater student travel grants (McGill), and the Society of Vertebrate Paleontology Student Travel Grant. 2 CONTRIBUTIONS OF AUTHORS Chapters 1 and 3 were written in collaboration with Dr. John R. Bolt from the Field Museum of Chicago. The present author decided the general direction of these chapters, studied specimens, conducted the analyses, and wrote the final drafts. -
Of Modern Amphibians: a Commentary
The origin(s) of modern amphibians: a commentary. D. Marjanovic, Michel Laurin To cite this version: D. Marjanovic, Michel Laurin. The origin(s) of modern amphibians: a commentary.. Journal of Evolutionary Biology, Wiley, 2009, 36, pp.336-338. 10.1007/s11692-009-9065-8. hal-00549002 HAL Id: hal-00549002 https://hal.archives-ouvertes.fr/hal-00549002 Submitted on 7 May 2020 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. The origin(s) of modern amphibians: a commentary By David Marjanović1 and Michel Laurin1* 1Address: UMR CNRS 7207 “Centre de Recherches sur la Paléobiodiversité et les Paléoenvironnements”, Muséum National d’Histoire Naturelle, Département Histoire de la Terre, Bâtiment de Géologie, case postale 48, 57 rue Cuvier, F-75231 Paris cedex 05, France *Corresponding author tel/fax. (+33 1) 44 27 36 92 E-mail: [email protected] Number of words: 1884 Number of words in text section only: 1378 2 Anderson (2008) recently reviewed the controversial topic of extant amphibian origins, on which three (groups of) hypotheses exist at the moment. Anderson favors the “polyphyly hypothesis” (PH), which considers the extant amphibians to be polyphyletic with respect to many Paleozoic limbed vertebrates and was most recently supported by the analysis of Anderson et al. -
Angela Milner Keraterpeton.Pdf
Title A morphological revision of Keraterpeton, the earliest horned nectridean from the Pennsylvanian of England and Ireland. Authors Milner, Angela Date Submitted 2020-02-07 Earth and Environmental Science Transactions of the Royal Society of Edinburgh A morphological revision of the earliest horned nectridean from the Pennsylvanian of England and Ireland Earth and Environmental Science Transactions of the Royal Society of Journal: Edinburgh Manuscript ID Draft Manuscript Type: Early Vertebrate Evolution Date Submitted by the Author: n/a Complete List of Authors: Milner, Angela; The Natural History Museum, Earth Sceinces For<i>Keraterpeton</i>, Peer Review anatomy, evolution, functional morphology, Keywords: systematics Cambridge University Press Page 1 of 43 Earth and Environmental Science Transactions of the Royal Society of Edinburgh 1 A morphological revision of the earliest horned nectridean from the Pennsylvanian of England and Ireland Angela C. Milner Department of Earth Sciences The Natural History Museum Cromwell Road London SW7 5BD [email protected] Running head For Peer Review Keraterpeton from the Pennsylvanian of England and Ireland Cambridge University Press Earth and Environmental Science Transactions of the Royal Society of Edinburgh Page 2 of 43 2 ABSTRACT The aquatic diplocaulid nectridean Keraterpeton galvani, is the commonest taxon represented in the Jarrow Coal assemblage from Kilkenny, Ireland. The Jarrow locality has yielded the earliest known Carboniferous coal-swamp fauna in the fossil record and is therefore of importance in understanding the history and diversity of the diplocaulid clade. The morphology of Keraterpeton is described in detail with emphasis on newly observed anatomical features. A reconstruction of the palate includes the presence of interpterygoid vacuities andnew morphological details of the pterygoid, parasphenoid and basicranial region. -
A Systematic and Ecomorphological Investigation of the Early Amniotes from Mazon Creek, Francis Creek Shale, Illinois, USA
A systematic and ecomorphological investigation of the early amniotes from Mazon Creek, Francis Creek Shale, Illinois, USA. by Arjan Mann A thesis submitted to the Faculty of Graduate and Postdoctoral Affairs in partial fulfillment of the requirements for the degree of Doctor of Philosophy in Earth Sciences Carleton University Ottawa, Ontario ©2020, Arjan Mann Abstract The late Carboniferous-aged (309-307 Ma) Mazon Creek lagerstätte produces some of the earliest tetrapod fossils, including those of major Paleozoic lineages such as the second oldest reptile. Despite this, the Mazon Creek lagerstätte has remained a difficult and unproductive vertebrate locality for researchers to utilize in tetrapod diversity studies due to the scarcity of fossils of this kind. Over the past decades, several new terrestrial tetrapod fossils collected from Mazon Creek have come to light. These include several new virtually-complete fossils of the earliest fossorially adapted recumbirostrans. Here I provide a revised systematic study of the Mazon Creek pan- amniote fauna, in an attempt to reassess the terrestrial ecosystem diversity present at the late Carboniferous lagerstätte. The results accumulate to systematic descriptions of four new and unique recumbirostran taxa (Diabloroter bolti, Infernovenator steenae, FMNH 1309, and MPM VP359229.2) and a re-description of the basal eureptile Cephalerpeton ventriarmatum leading to the anointment of the oldest parareptile Carbonodraco lundi (formerly Cephalerpeton aff. C. ventriarmatum from Linton, Ohio). Descriptions are aided by modern imaging techniques and updated phylogenetic analyses using Maximum parsimony and Bayesian methods where applicable. Across the newly described terrestrial fauna there is an unexpected ecomorphological diversity of bauplans present. These range from the short-bodied Diabloroter to the serpentine, long-bodied, and limb-reduced MPM VP359229.2. -
On the Origin of and Phylogenetic Relationships Among Living Amphibians
On the origin of and phylogenetic relationships among living amphibians Rafael Zardoya*† and Axel Meyer‡ *Museo Nacional de Ciencias Naturales, Jose´Gutierrez Abascal, 2, 28006 Madrid, Spain; and ‡Department of Biology, University of Konstanz, 78457 Konstanz, Germany Edited by David B. Wake, University of California, Berkeley, CA, and approved April 4, 2001 (received for review September 22, 2000) The phylogenetic relationships among the three orders of modern (9) integrated previous morphological and paleontological evi- amphibians (Caudata, Gymnophiona, and Anura) have been esti- dence, and concluded that the Lissamphibia were a natural mated based on both morphological and molecular evidence. Most group. The monophyly of Lissamphibia has since gained wide morphological and paleontological studies of living and fossil acceptance among researchers (refs. 1 and 10–16; Fig. 1 a–c). A amphibians support the hypothesis that salamanders and frogs are noteworthy exception is Carroll (17–21), who suggests that sister lineages (the Batrachia hypothesis) and that caecilians are Lissamphibia are nonmonophyletic because he believes that more distantly related. Previous interpretations of molecular data salamanders and caecilians have affinities with different lineages based on nuclear and mitochondrial rRNA sequences suggested of microsauria (an extinct group of Lepospondyl amphibians), that salamanders and caecilians are sister groups to the exclusion whereas frogs are related to another extinct group, the temno- of frogs. In an attempt to resolve this apparent conflict, the spondyl amphibians (Fig. 1d). complete mitochondrial genomes of a salamander (Mertensiella Although monophyly of the Lissamphibia is widely accepted, luschani) and a caecilian (Typhlonectes natans) were determined it is still controversial whether the extinct temnospondyls (Fig. -
(Lepospondyli: Nectridea) Ze Svrchního Karbonu Lokality Nýřany, Česká Republika
PŘÍRODOVĚDECKÁ FAKULTA Redeskripce druhu Sauropleura scalaris (Lepospondyli: Nectridea) ze svrchního karbonu lokality Nýřany, Česká republika Rešerše k diplomové práci Pavel Barták Vedoucí práce: doc. Mgr. Martin Ivanov, Dr. Ústav geologických věd Obsah Úvod ............................................................................................................................... 3 Lokality svrchnokarbonského stáří České republiky s doklady skupin Temnospondyli Zittel, 1888 a Lepospondyli Zittel, 1888 .......................................... 3 Nýřany ................................................................................................................................................. 4 Třemošná ............................................................................................................................................. 5 Kounov ................................................................................................................................................. 6 Systematika svrchnokarbonských tetrapodů: Lepospondyli Zittel, 1888 & Temnospondyli Zittel, 1888 .......................................................................................... 8 Lepospondyli Zittel, 1888 .................................................................................................................... 8 Temnospondyli Zittel, 1888 ................................................................................................................. 9 Chronologický přehled výzkumů bazálních tetrapodů ze svrchního -
Katedra Geologie a Paleontologie Přírodovědecká Fakulta Univerzity Karlovy V Praze
Katedra geologie a paleontologie Přírodovědecká fakulta Univerzity Karlovy v Praze DIPLOMOVÁ PRÁCA Variabilita stavcov permokarbónskych lepospondylných obojživelníkov Pavel Danko Školiteľ: Doc. RNDr. Zbyněk Roček, DrSc. september 2004 2 POĎAKOVANIE Chcem poďakovať predovšetkým svojmu školiteľovi a konzultantovi pánu doc. RNDr. Zbyňkovi Ročkovi DrSc. z katedry zoológie PřF – UK v Prahe za odborné vedenie, cenné rady a pripomienky pri písaní diplomovej práce. Ďakujem RNDr. Martinovi Košťakovi z katedry geologie a paleontologie PřF – UK v Prahe za poskytnutie fosilného materiálu z paleontologických zbierok, Mgr. Borisovi Ekrtovi z paleontologického oddelenia Národního muzea v Prahe a doc. RNDr. Jaroslavovi Kraftovi CSc. zo Západočeského muzea v Plzni za sprístupnenie fosilného materiálu a poskytnutie techniky (digitálny fotoaparát, optický mikroskop, binokulárna lupa) k ďalšiemu spracovaniu získaných informácii počas výskumu. Na záver chcem poďakovať svojim rodičom a kolegom z katedry geológie a paleontológie PřF – UK v Prahe za podporu a uznanie v mojej vedeckej práci. Prehlasujem, že som diplomovú prácu vypracoval samostatne s použitím odbornej literatúry a súhlasím s jej zapožičiavaním. ________________________ 3 Obsah 1. Úvod 5 1.0. Deskriptívna terminológia definitívneho stavca 2.0. Embryonálny pôvod stavca 3.0. Fylogenetická diferenciácia stavca u raných obojživelníkov 4.0. Problém lepospondylného stavca 1.4.1 Historický prehľad názorov na charakter a vznik lepospondylného stavca 2. Materiál a metodika 26 3. Výsledky 27 Labyrinthodontia Discosauriscus pulcherrimus (Fritsch, 1879) Discosauriscus potamites (Steen, 1938) Discosauriscus sp. Letoverpeton austriacum (Makowsky, 1876) Letoverpeton moravicum (Fritsch, 1879) Lepospondyli Microbrachis pelikani Frič, 1876 Hyloplesion longicostatum (Frič, 1876) Sauropleura scalaris (Frič, 1876) Scincosaurus crassus Frič, 1876 4 Phlegethontia longissima (Frič, 1876) (partim) + Oestocephalus amphiuminum (Cope, 1868) (partim) podľa Andersona, Carrolla & Roweho (2003) Oestocephalus granulosum (Frič, 1879) 4. -
The Palaeozoic Ancestry of Salamanders, Frogs and Caecilians
Blackwell Publishing LtdOxford, UKZOJZoological Journal of the Linnean Society0024-4082The Lin- nean Society of London, 2007? 2007 150s2 1140 Original Articles LISSAMPHIBIAN ANCESTRYR. L. CARROLL Zoological Journal of the Linnean Society, 2007, 150 (Suppl. 1), 1–140. With 78 figures The Palaeozoic Ancestry of Salamanders, Frogs and Caecilians ROBERT L. CARROLL FLS1 1Redpath Museum, McGill University, 859 Sherbrooke St W., Montreal, P.Q. Canada, H3A 2K6 The relationships of frogs, salamanders, and caecilians (Gymnophiona) with one another and with the vast assem- blage of Palaeozoic amphibians remain highly contentious phylogenetic problems. Cladistic analyses support a com- mon ancestry of the three modern orders, but fail to achieve a consensus regarding their affinities with Palaeozoic amphibians. The most exhaustive phylogenetic analyses that have been applied to the ancestry of lissamphibians have recognized few, if any, biologically significant characters differentiating the living orders. These results can be attributed to limiting the database primarily to characters common to Palaeozoic amphibians and including few fea- tures that distinguish the modern orders. Making use of the numerous derived characters that are expressed in either the larvae or adults of extant salamanders, frogs, and caecilians provides the basis for recognizing a nested sequence of synapomorphies that support a common ancestry of salamanders and anurans with temnospondyl lab- yrinthodonts to the exclusion of caecilians. The larvae of Carboniferous and Permian temnospondyl labyrinthodonts provide strong evidence for their being members of the stem group of urodeles. This is based primarily on the great similarity in the sequence of ossification of the bones of the skull and appendicular skeleton, but is also supported by detailed similarities of the hyoid apparatus. -
Paleo Jegyzet.Indd
1 PALEONTOLÓGIA - jegyzet - 2 3 Az állatvilág rendszere Az egysejtű állatokat a telepes növényekkel együtt a Protistákhoz sorolják. Az egysejtűek (Protozoa) és a többsejtűek (Metazoa) között elég nagy az eltérés. Ez mind a test méretében, mind pedig a felépitésében megmutatkozik. Megszokták különböztetni az álszöveteseket a valódi szö- vetes állatoktól is. Az álszövetesek (Parazoa) esetében a testsejtek (szomatikus sejtek) elkülönülnek a szaporodási sejtektől (generatív sejtek), de azonos felépítésű és működésű sejtekből álló szövet azonban még nem alakult ki. Az álszövetesek (Parazoa) közé mindössze egyetlen állatt örzs tartozik, a szivacsok. A többsejtűek többsége valódi szövetes (Eumetazoa). Ezek alacsonyabbrendű csoportjainál a test 2 rétegből épül fel : a külső- és a belső csíralevélből (ektodermából és entodermából). Az en- toderma veszi körül a gasztrális üreget, a cölenteront vagy űrbelet. Ezért ezeket az állatcsoportokat még Coelenteratának is szokták nevezni. Ide tartoznak a csalánozók (Cnidaria) a bordásmedúzák (Ctenophora). A többi Eumetazoának három csíralevele van: ektoderma, entoderma és a köztes me- zoderma. Az űrbelűek sugarasan részarányos testfelépítésűek, míg a fejlett ebb többsejtűek kétoldalian részarányosak (Bilateria). A Bilateriák döntő többségénél az egyedi fejlődés során az ősszáj (pro- tosztóma) megmarad, és a végbélnyílás másodlagosan alakul ki. Ezek az ősszájúak (Protostomia). Az ősszájúaknál a központi idegrendszer a hasi (ventrális) oldalon húzódik, és a váz általában ek- todermális eredetű. A mai állatvilágnak