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ZOOLOGICAL MUSEUM OF MOSCOW STATE UNIVERSITY K.G. Mikhailov The spiders (Arachnida: Aranei) of Russia and adjacent countries: a non-annotated checklist Arthropoda Selecta. Supplement No. 3. KMK Scientific Press Ltd. Moscow v 2013 Mikhailov K.G.1 The spiders (Arachnida: Aranei) of Russia and adjacent countries: a non- annotated checklist. Arthropoda Selecta. Supplement No.3. Moscow: KMK Scientific Press Ltd. 262 p. 1 Zoological Museum MGU, Bolshaya Nikitskaya Str. 6, Moscow 125009 Russia. E-mail: [email protected] A checklist of 3,340 spider species belonging to 629 genera and 50 families is provided for Russia and the other post-Soviet republics, namely, Armenia, Azerbaijan, Byelorussia (Belarus), Estonia, Georgia, Kazakhstan, Latvia, Lithuania, Moldavia (Moldova), Kirghizia (Kyrgyzstan), Tajikistan, Turkmenia (Turkmenistan), Ukraine, and Uzbekistan, based on comprehensive literature data covering more than 2,600 sources published between 1770 and June 2013. Calculations of the number of spider species in the post-Soviet republics and different physiographical areas are provided in the introductory part. Each species included in the checklist is supplied with an attribution both to physiographical area(s) and republic(s). The necessary synonymies and valid subspecies are also enlisted, same as nomina dubia and nomina nuda. An alphabetic index of all genera and species names is provided as well. Destined for zoologists, mainly arachnologists and entomologists, as well as for local biologists and naturalists. 1 map, 3 tables, 199 references. Key words: Aranei, spiders, checklist, Russia, Armenia, Azerbaijan, Byelorussia, Belarus, Estonia, Georgia, Kazakhstan, Latvia, Lithuania, Moldavia, Moldova, Kirghizia, Kyrgyzstan, Tajikistan, Turkmenia, Turkmenistan, Ukraine, Uzbekistan, physiographical areas. Ìèõàéëîâ Ê.Ã.1 Ïàóêè (Arachnida: Aranei) Ðîññèè è ïðèëåæàùèõ ñòðàí: íåàííîòè- ðîâàííûé ðååñòð. Arthropoda Selecta. Ïðèëîæåíèå ¹ 3. Ìîñêâà: Òîâàðèùåñòâî íàó÷íûõ èçäàíèé ÊÌÊ. 262 ñ. 1 Çîîëîãè÷åñêèé ìóçåé ÌÃÓ, óë. Áîëüøàÿ Íèêèòñêàÿ, 6, Ìîñêâà 125009 Ðîññèÿ. E-mail: [email protected] Íà îñíîâå èñ÷åðïûâàþùèõ ëèòåðàòóðíûõ äàííâõ, ñîáðàííûõ â áîëåå ÷åì 2600 èñòî÷íèêàõ çà ïåðèîä ñ 1770 ã. ïî èþíü 2013 ã. äàí ðååñòð 3340 âèäîâ ïàóêîâ, ïðèíàäëåæàùèõ ê 629 ðîäàì è 50 ñåìåéñòâàì ñ òåððèòîðèè Ðîññèè è äðóãèõ ïîñòñîâåòñêèõ ðåñïóáëèê: Àðìåíèè, Àçåðáàéäæàíà, Áåëîðóññèè (Áåëàðóñè), Ýñòîíèè, Ãðóçèè, Êàçàõñòàíà, Ëàòâèè, Ëèòâû, Ìîë- äàâèè (Ìîëäîâû), Êèðãèçèè (Êûðãûçñòàíà), Òàäæèêèñòàíà, Òóðêìåíèè (Òóðêìåíèñòàíà), Óêðàèíû è Óçáåêèñòàíà. Âî ââîäíîé ÷àñòè ïðèâåäåíû ïîäñ÷åòû ÷èñëà âèäîâ ïàóêîâ íà òåððèòîðèè ïîñòñîâåòñêèõ ðåñïóáëèê è ôèçèêî-ãåîãðàôè÷åñêèõ ðåãèîíîâ. Êàæäûé âèä â ðååñòðå ñíàáæåí óêàçàíèÿìè íà åãî ðàñïðîñòðàíåíèå â ñîîòâåòñòâóþùèõ ôèçèêî-ãåîãðàôè- ÷åñêèõ ðåãèîíàõ è ðåñïóáëèêàõ. Ïåðå÷èñëåíû íåîáõîäèìûå ñèíîíèìû è âàëèäíûå ïîäâè- äû, à òàêæå nomina dubia è nomina nuda. Äàí àëôàâèòíûé óêàçàòåëü âñåõ ðîäîâûõ è âèäîâûõ íàçâàíèé. Äëÿ çîîëîãîâ, ïðåèìóùåñòâåííî àðàõíîëîãîâ è ýíòîìîëîãîâ, à òàêæå äëÿ êðàåâåäîâ è íàòóðàëèñòîâ. 1 êàðòà, 3 òàáë., áèáë. 199. Êëþ÷åâûå ñëîâà: Aranei, ïàóêè, ðååñòð, Ðîññèÿ, Àðìåíèÿ, Àçåðáàéäæàí, Áåëîðóññèÿ, Áåëà- ðóñü, Ýñòîíèÿ, Ãðóçèÿ, Êàçàõñòàí, Ëàòâèÿ, Ëèòâà, Ìîëäàâèÿ, Ìîëäîâà, Êèðãèçèÿ, Êûðãûç- ñòàí, Òàäæèêèñòàí, Òóðêìåíèÿ, Òóðêìåíèñòàí, Óêðàèíà, Óçáåêèñòàí, ôèçèêî-ãåîãðàôè÷å- ñêèå ðåãèîíû. © K.G. Mikhailov, 2013 ISBN 978-5-87317-933-6 © KMK Scientific Press, 2013 Preface and Introduction The material presented below is the first 1996b,c, 1997b, 2000b, 2002a, 2011, 2012b, more or less complete result of my project 2013a]. started in 1981. The objective of this project is The following mistakes have been found compiling the literature data on the spiders of since the 1997 checklist [Mikhailov, 1997b]: Russia and other post-Soviet republics, with P.93. Savignya Blackwall, 1833 (a generic some critical observations. name in Linyphiidae) is a subsequent invalid The bibliography list comprising 3,560 pa- emendation. The original correct name is Sav- pers and monographs which appeared between ignia Blackwall, 1833. 1770 until 2011 has been published elsewhere P.117. Cyrtarachne ixodoides (Simon, 1870) [Mikhailov, 2012a]. Regular addenda to this (Araneidae). A lapsus calami. The correct name basic bibliography list are planned. A card cat- is Cyrtarachne ixoides (Simon, 1870). alogue of local/regional data is also complete, P.133138. Tarentula Sundevall, 1832 (a but not digitalized. However, publication of all generic name in Lycosidae). Must be replaced material (comparable in volume to P. Bonnets with Alopecosa Simon, 1885 [Mikhailov, Bibliographia araneorum), as insistently rec- 2013c]. ommended by Heptner [1998] and some col- P.134. Tarentula andreevae Mikhailov, leagues arachnologists, remains impossible, first 1996 (Lycosidae). An incorrect replacement of all due to purely technical reasons. Besides name for Tarentula kronebergi (Andreeva, this, local faunistic data are sometimes very 1976), not homonymous to Tarentula croneber- difficult to verify, since many authors do/did gi Thorell, 1875. not keep their collections properly. Numerous P.144. Tegenaria lapidicinarum Spassky, samples are lost, some others inaccessible. 1934 (Agelenidae). A lapsus calami. The cor- To date, only a checklist of the spiders of the rect name is Tegenaria lapicidinarum Spassky, former USSR together with a bibliographic in- 1934. dex covering all literature sources until July P.169. Drassylus Chamberlin, 1922 (Gna- 2000 has been published [Mikhailov 1997a, b, phosidae). A lapsus calami. The correct name is 1998, 1999, 2000a]. An earlier checklist had Drassyllus Chamberlin, 1922. appeared as well [Mikhailov, 1996a]. Still the P.186. Micromata Latreille, 1804 (Spar- way the bibliographic index was presented in assidae), an attempt to validate a forgotten name. the above series was extremely complicated, The generally accepted valid name (ICZN Arti- changed into a more clear and simple one in the cle 23.9.1) is Micrommata Latreille, 1804 (an latest edition [Mikhailov, 2012a]. The compo- emendation by Latreille [1806]). sition of the basic book [Mikhailov, 1997b] P.197. Thomisis albus (Gmelin, 1789) (Th- which failed to supply each species with proper omisidae). A nomen dubium, after Simon references and clear connection between the 2nd [1918]. Should be replaced by Thomisus onus- (checklist) and 3rd (bibliography) chapters was tus Walckenaer, 1805. justly criticized in a review in Zoologicheskiy P.216. Neon valentinulus Falconer, 1912 Zhurnal [Heptner, 1998]. For this reason, only (Salticidae). A lapsus calami. The correct name a checklist is retained in this edition, but this is Neon valentulus Falconer, 1912. time supplied with the necessary, sometimes When preparing this edition, the earlier shortened, references provided in a format stan- checklist was compared with the current World dard for Arthropoda Selecta. In the current Spider Catalog [Platnick, 2013]. Certain differ- checklist, the literature data are mainly present- ences have been found and followed. The se- ed in footnotes, as well as in special cases like quence of families in my checklist basically non-existing species, nomina nuda, etc. As not- follows that in the earlier version [Mikhailov, ed earlier, a general bibliographic index has 1997b], with minor changes according to the been published separately [Mikhailov, 2012a]. modern version of the World Spider Catalog. Calculations of spider families, genera and The genus Cheiracanthium C.L. Koch, 1839 is species amongst the post-USSR republics and left in Clubionidae. The spiders described by physiographical regions have been published in Carl Clerck are dated 1758, not 1757, as de- a number of contributions [Mikhailov, 1992, manded by ICZN Articles 3 and 21; the linyphi- 4 K.G. Mikhailov id taxonomy mostly follows A.V. Tanasevitch; also grateful to K.Yu. Eskov (Moscow), A.A. the name Larinioides folium (Schrenk, 1803) Zyuzin (Alma-Ata, Kazakhstan) and V.I. Ovt- has priority over L. suspicax (O. Pickard-Cam- sharenko (New York, earlier St. Petersburg) for bridge, 1876) [Mikhailov, 2013b]; etc., etc. their remarks on earlier versions of my card I am deeply obliged to my colleagues and catalogue of the 1980s and 1990s. friends Yu.M. Marusik (Magadan), A.V. Ta- Compilation of the tremendous bibliograph- nasevitch (Moscow), D.V. Logunov (Manches- ical index published separately would have been ter, earlier Novosibirsk), M.M. Kovblyuk (Sim- impossible without a flow of papers and copies feropol, Ukraine), S.L. Esyunin (Perm), A.V. from all my Russian/Soviet and a number of Ponomarev (Rostov-na-Donu), G.N. Azarkina foreign colleagues. I would like to thank again (Novosibirsk), A.V. Gromov (Bingen am Rhein, the dozens of persons listed already in Mikhailov Germany, earlier Alma-Ata, Kazakhstan), E.M. [2012a]. Zhukovets (Minsk, Byelorussia) and others for The linguistic help of S.I. Golovatch (Mos- their valuable comments on the checklist. I am cow) is also deeply acknowledged. The spiders of Russia and adjacent countries: a non-annotated check-list 5 Methodology The checklist of spiders is actually an ex- Êèðã Kirghizia (Kyrgyzstan), traction from a complete card catalogue. Òàäæ Tajikistan. Each spider species included is supplied Synonyms are marked with a = sign, only with an attribution both to the main physio- the names cited in the Russian/Soviet arachno- graphical areas delimited after Gvozdetskiy logical literature are reported. Some old combi- [1968] and to post-Soviet republics. The phys- nations not accepted