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The Blowflies of California
BULLETIN OF THE CALIFORNIA INSECT SURVEY VOLUME 4,NO. 1 THE BLOWFLIES OF CALIFORNIA (Diptera: Calliphoridae) BY MAURICE T. JAMES (Department of Zo'dlogy, State College of Washington, Pullman) UNIVERSITY OF CALIFORNIA PRESS BERKELEY AND LOS ANGELES 1955 BULLETIN OF THE CALIFORNIA INSECT SURVEY Editors: E. G. Linsley, S. B. Freeborn, R. L. Usinger Volume 4, No. 1, pp. 1-34, plates 1-2, 1 figure in text Submitted by Editors, January 13, 1955 Issued October 28, 1955 Price, 50 cents UNIVERSITY OF CALIFORNIA PRESS BERKELEY AND LOS ANGELES CALIFORNIA CAMBRIDGE UNIVERSITY PRESS LONDON. ENGLAND PRINTED BY OFFSET IN THE UNITED STATES OF AMERICA THE BLOWFLIES OF CALIFORNIA (Diptera: Calliphoridae) by Maurice T. James Identification of the blowflies of North America Blowflies are important from a medical and has been made much easier and more secure in veterinary standpoint. Some are obligatory or recent years by the publication of the monograph facultative parasites on man or on domestic or of that family by Hall (1948). However, there other useful animals. In our area, the primary exists ti0 regional treatment that covers any screwworm, Callitroga hominivorax (Coquerel), definite part of the United States. Hall's mono- 'is the only obligatory parasite that invades graph gives only general information about the living tissue, although larvae of Pmtocalliphora, geographical distribution of most of the species. represented in the Califdrnia fauna by seven These considerations, together with the fact that known species, feed on the blood of nesting Hall had obviously examined an insufficient birds, often with fatal results. Of the facultative amount of material from the western states, parasites, Callitroga macellaria (Fabricius), makes a review of the California species partic- Phaenicia sericata (Meigen), and Phormia regina ularly desirable. -
Die Steppe Lebt
Buchrücken 1200 Stück:Layout 1 04.04.2008 14:39 Seite 1 Die Steppe lebt Felssteppen und Trockenrasen in Niederösterreich Heinz Wiesbauer (Hrsg.) Die Steppe lebt ISBN 3-901542-28-0 Die Steppe lebt Felssteppen und Trockenrasen in Niederösterreich Heinz Wiesbauer (Hrsg.) Mit Beiträgen von Roland Albert, Horst Aspöck, Ulrike Aspöck, Hans-Martin Berg, Peter Buchner, Erhard Christian, Margret Bunzel-Drüke, Manuel Denner, Joachim Drüke, Michael Duda, Rudolf Eis, Karin Enzinger, Ursula Göhlich, Mathias Harzhauser, Johannes Hill, Werner Holzinger, Franz Humer, Rudolf Klepsch, Brigitte Komposch, Christian Komposch, Ernst Lauermann, Erwin Neumeister, Mathias Pacher, Wolfgang Rabitsch, Birgit C. Schlick-Steiner, Luise Schratt-Ehrendorfer, Florian M. Steiner, Otto H. Urban, Henning Vierhaus, Wolfgang Waitzbauer, Heinz Wiesbauer und Herbert Zettel St. Pölten 2008 Die Steppe lebt – Felssteppen und Trockenrasen in Niederösterreich Begleitband zur gleichnamigen Ausstellung in Hainburg an der Donau Bibliografische Information der Deutschen Bibliothek Die Deutsche Bibliothek verzeichnet diese Publikation in der Deutschen Nationalbibliografie; detaillierte bibliografische Daten sind im Internet über http://dnb.ddb.de abrufbar. ISBN 3-901542-28-0 Die Erstellung des Buches wurde aus Mitteln von LIFE-Natur gefördert. LIFE-Natur-Projekt „Pannonische Steppen und Trockenrasen“ Gestaltung: Manuel Denner und Heinz Wiesbauer Lektorat: caout:chouc Umschlagbilder: Heinz Wiesbauer Druck: Gugler Druck, Melk Medieninhaber: Amt der NÖ Landesregierung, Abteilung Naturschutz Landhausplatz 1 A-3109 St. Pölten Bestellung: Tel.: +43/(0)2742/9005-15238 oder [email protected] © 2008 Autoren der jeweiligen Beiträge, Bilder: Bildautoren Sämtliche Rechte vorbehalten Inhalt 1. Einleitung 5 2. Eiszeitliche Steppen und Großsäuger 9 2.1 Was ist Eiszeit? 11 2.2 Die Tierwelt der Eiszeit 14 2.3 Der Einfluss von Großherbivoren auf die Naturlandschaft Mitteleuropas 17 3. -
From Chewing to Sucking Via Phylogeny—From Sucking to Chewing Via Ontogeny: Mouthparts of Neuroptera
Chapter 11 From Chewing to Sucking via Phylogeny—From Sucking to Chewing via Ontogeny: Mouthparts of Neuroptera Dominique Zimmermann, Susanne Randolf, and Ulrike Aspöck Abstract The Neuroptera are highly heterogeneous endopterygote insects. While their relatives Megaloptera and Raphidioptera have biting mouthparts also in their larval stage, the larvae of Neuroptera are characterized by conspicuous sucking jaws that are used to imbibe fluids, mostly the haemolymph of prey. They comprise a mandibular and a maxillary part and can be curved or straight, long or short. In the pupal stages, a transformation from the larval sucking to adult biting and chewing mouthparts takes place. The development during metamorphosis indicates that the larval maxillary stylet contains the Anlagen of different parts of the adult maxilla and that the larval mandibular stylet is a lateral outgrowth of the mandible. The mouth- parts of extant adult Neuroptera are of the biting and chewing functional type, whereas from the Mesozoic era forms with siphonate mouthparts are also known. Various food sources are used in larvae and in particular in adult Neuroptera. Morphological adaptations of the mouthparts of adult Neuroptera to the feeding on honeydew, pollen and arthropods are described in several examples. New hypoth- eses on the diet of adult Nevrorthidae and Dilaridae are presented. 11.1 Introduction The order Neuroptera, comprising about 5820 species (Oswald and Machado 2018), constitutes together with its sister group, the order Megaloptera (about 370 species), and their joint sister group Raphidioptera (about 250 species) the superorder Neuropterida. Neuroptera, formerly called Planipennia, are distributed worldwide and comprise 16 families of extremely heterogeneous insects. -
Terry Whitworth 3707 96Th ST E, Tacoma, WA 98446
Terry Whitworth 3707 96th ST E, Tacoma, WA 98446 Washington State University E-mail: [email protected] or [email protected] Published in Proceedings of the Entomological Society of Washington Vol. 108 (3), 2006, pp 689–725 Websites blowflies.net and birdblowfly.com KEYS TO THE GENERA AND SPECIES OF BLOW FLIES (DIPTERA: CALLIPHORIDAE) OF AMERICA, NORTH OF MEXICO UPDATES AND EDITS AS OF SPRING 2017 Table of Contents Abstract .......................................................................................................................... 3 Introduction .................................................................................................................... 3 Materials and Methods ................................................................................................... 5 Separating families ....................................................................................................... 10 Key to subfamilies and genera of Calliphoridae ........................................................... 13 See Table 1 for page number for each species Table 1. Species in order they are discussed and comparison of names used in the current paper with names used by Hall (1948). Whitworth (2006) Hall (1948) Page Number Calliphorinae (18 species) .......................................................................................... 16 Bellardia bayeri Onesia townsendi ................................................... 18 Bellardia vulgaris Onesia bisetosa ..................................................... -
Dating Pupae of the Blow Fly Calliphora Vicina Robineau
G C A T T A C G G C A T genes Article Dating Pupae of the Blow Fly Calliphora vicina Robineau–Desvoidy 1830 (Diptera: Calliphoridae) for Post Mortem Interval—Estimation: Validation of Molecular Age Markers Barbara K. Zajac *, Jens Amendt, Marcel A. Verhoff and Richard Zehner Institute of Legal Medicine, Goethe University, 60596 Frankfurt, Germany; [email protected] (J.A.); [email protected] (M.A.V.); [email protected] (R.Z.) * Correspondence: [email protected]; Tel.: +49-163-865-3366 Received: 31 December 2017; Accepted: 5 March 2018; Published: 9 March 2018 Abstract: Determining the age of juvenile blow flies is one of the key tasks of forensic entomology when providing evidence for the minimum post mortem interval. While the age determination of blow fly larvae is well established using morphological parameters, the current study focuses on molecular methods for estimating the age of blow flies during the metamorphosis in the pupal stage, which lasts about half the total juvenile development. It has already been demonstrated in several studies that the intraspecific variance in expression of so far used genes in blow flies is often too high to assign a certain expression level to a distinct age, leading to an inaccurate prediction. To overcome this problem, we previously identified new markers, which show a very sharp age dependent expression course during pupal development of the forensically-important blow fly Calliphora vicina Robineau–Desvoidy 1830 (Diptera: Calliphoridae) by analyzing massive parallel sequencing (MPS) generated transcriptome data. We initially designed and validated two quantitative polymerase chain reaction (qPCR) assays for each of 15 defined pupal ages representing a daily progress during the total pupal development if grown at 17 ◦C. -
Evaluating the Effects of Temperature on Larval Calliphora Vomitoria (Diptera: Calliphoridae) Consumption
Evaluating the effects of temperature on larval Calliphora vomitoria (Diptera: Calliphoridae) consumption Kadeja Evans and Kaleigh Aaron Edited by Steven J. Richardson Abstract: Calliphora vomitoria (Diptera: Calliphoridae) are responsible for more cases of myiasis than any other arthropods. Several species of blowfly, including Cochliomyia hominivorax and Cocholiomya macellaria, parasitize living organisms by feeding on healthy tissues. Medical professionals have taken advantage of myiatic flies, Lucillia sericata, through debridement or maggot therapy in patients with necrotic tissue. This experiment analyzes how temperature influences blue bottle fly, Calliphora vomitoria. consumption of beef liver. After rearing an egg mass into first larval instars, ten maggots were placed into four containers making a total of forty maggots. One container was exposed to a range of temperatures between 18°C and 25°C at varying intervals. The remaining three containers were placed into homemade incubators at constant temperatures of 21°C, 27°C and 33°C respectively. Beef liver was placed into each container and weighed after each group pupation. The mass of liver consumed and the time until pupation was recorded. Three trials revealed that as temperature increased, the average rate of consumption per larva also increased. The larval group maintained at 33°C had the highest consumption with the shortest feeding duration, while the group at 21°C had lower liver consumption in the longest feeding period. Research in this experiment was conducted to understand the optimal temperature at which larval consumption is maximized whether in clinical instances for debridement or in myiasis cases. Keywords: Calliphora vomitoria, Calliphoridae, myiasis, consumption, debridement As an organism begins to decompose, target open wounds or necrotic tissue. -
Non-Commercial Use Only
Journal Journal of Entomological of Entomological and andAcarological Acarological Research Research 2017; 2012; volume volume 49:6823 44:e ENTOMOLOGY First case of traumatic myiasis caused by Calliphora vicina in a crested porcupine Hystrix cristata L. in Italy D. Scaravelli,1 C. Senini,1 T. Bonacci2 1Department of Veterinary Medical Sciences (DiMEVET), University of Bologna; 2Department of Biology, Ecology and Earth Sciences (DiBEST), University of Calabria, Rende, Italy Sarcophagidae and Muscidae are the main responsible of traumat- Abstract ic myiasis in humans and animals (Zumpt, 1965; Hall & Wall, 1995; Hall, 1997; Hall & Farkas, 2000; Sinha, 2012). Coversely, Calliphora vicina (Diptera: Calliphoridae) it is a facultative some Muscidae are active predators of economically important ectoparasite responsible for traumatic myiasis in humans and warm- pests (Bonsignore, 2016). blooded vertebrates in the world. In this work one case of traumatic Calliphora vicina Robineau-Desvoidy, 1830 has an almost myasis caused by C. vicina (Diptera Calliphoirdae) is reported for worldwide distribution; it mainly favours shady situations and the first time in a vulnerable crested porcupine (Hystrix cristata urban habitats, where itonly is often the dominant species on human Linnaeus, 1758). A total of 30 larvae located in the posterior-dorsal corpses (Smith, 1986; Bonacci, et al., 2009). Calliphora vicina it area of the animal were removed from inside the lesion and either is known like facultative ectoparasites of many species of wild preserved in ethanol or reared to the adult stage. This report shows animals and humans (Dehlaes et al., 2001; Sales et al., 2003; the great ability of C. vicina to use many organic matter for the food Knotek et aluse., 2004; Salvetti et al., 2012; Araghi et al., 2015). -
Functional Morphology of the Raptorial Forelegs in Mantispa Styriaca (Insecta: Neuroptera)
Zoomorphology https://doi.org/10.1007/s00435-021-00524-6 ORIGINAL PAPER Functional morphology of the raptorial forelegs in Mantispa styriaca (Insecta: Neuroptera) Sebastian Büsse1 · Fabian Bäumler1 · Stanislav N. Gorb1 Received: 14 September 2020 / Revised: 26 March 2021 / Accepted: 30 March 2021 © The Author(s) 2021 Abstract The insect leg is a multifunctional device, varying tremendously in form and function within Insecta: from a common walking leg, to burrowing, swimming or jumping devices, up to spinning apparatuses or tools for prey capturing. Raptorial forelegs, as predatory striking and grasping devices, represent a prominent example for convergent evolution within insects showing strong morphological and behavioural adaptations for a lifestyle as an ambush predator. However, apart from praying mantises (Mantodea)—the most prominent example of this lifestyle—the knowledge on morphology, anatomy, and the functionality of insect raptorial forelegs, in general, is scarce. Here, we show a detailed morphological description of raptorial forelegs of Mantispa styriaca (Neuroptera), including musculature and the material composition in their cuticle; further, we will discuss the mechanism of the predatory strike. We could confrm all 15 muscles previously described for mantis lacewings, regarding extrinsic and intrinsic musculature, expanding it for one important new muscle—M24c. Combining the information from all of our results, we were able to identify a possible catapult mechanism (latch-mediated spring actuation system) as a driving force of the predatory strike, never proposed for mantis lacewings before. Our results lead to a better understand- ing of the biomechanical aspects of the predatory strike in Mantispidae. This study further represents a starting point for a comprehensive biomechanical investigation of the convergently evolved raptorial forelegs in insects. -
Post-Feeding Larval Behaviour in the Blowfly
Available online at www.sciencedirect.com Forensic Science International 177 (2008) 162–167 www.elsevier.com/locate/forsciint Post-feeding larval behaviour in the blowfly, Calliphora vicina: Effects on post-mortem interval estimates Sophie Arnott, Bryan Turner * Department of Forensic Science and Drug Monitoring, King’s College London, Franklin-Wilkins Building, 150 Stamford Street, London SE1 9NH, United Kingdom Received 9 July 2007; received in revised form 26 September 2007; accepted 5 December 2007 Available online 19 February 2008 Abstract Using the rate of development of blowflies colonising a corpse, accumulated degree hours (ADH), or days (ADD), is an established method used by forensic entomologists to estimate the post-mortem interval (PMI). Derived from laboratory experiments, their application to field situations needs care. This study examines the effect of the post-feeding larval dispersal time on the ADH and therefore the PMI estimate. Post-feeding dispersal in blowfly larvae is typically very short in the laboratory but may extend for hours or days in the field, whilst the larvae try to find a suitable pupariation site. Increases in total ADH (to adult eclosion), due to time spent dispersing, are not simply equal to the dispersal time. The pupal period is increased by approximately 2 times the length of the dispersal period. In practice, this can introduce over-estimation errors in the PMI estimate of between 1 and 2 days if the total ADH calculations do not consider the possibility of an extended larval dispersal period. # 2007 Elsevier Ireland Ltd. All rights reserved. Keywords: Dispersal; Accumulated degree hours; ADH; Accumulated degree days; ADD; Forensic entomology; PMI; Blowflies; Calliphora 1. -
Ultraviolet Vision in European Owlflies (Neuroptera: Ascalaphidae): a Critical Review
REVIEW Eur. J. Entomol. 99: 1-4, 2002 ISSN 1210-5759 Ultraviolet vision in European owlflies (Neuroptera: Ascalaphidae): a critical review Ka r l KRAL Institut fur Zoologie, Karl-Franzens-Universitat Graz, A-8010 Graz, Austria; e-mail: [email protected] Key words.Owlfly, Ascalaphus, Neuroptera, insect vision, ultraviolet sensitivity, visual acuity, visual behaviour, visual pigment Abstract. This review critically examines the ecological costs and benefits of ultraviolet vision in European owlflies. On the one hand it permits the accurate pursuit of flying prey, but on the other, it limits hunting to sunny periods. First the physics of detecting short wave radiation are presented. Then the advantages and disadvantages of the optical specializations necessary for UV vision are discussed. Finally the question of why several visual pigments are involved in UV vision is addressed. UV vision in predatory European owlflies of R7 means that the former receives only the longer The European owlflies, like Ascalaphus macaronius, wavelengths, since the short wavelengths are absorbed by A. libelluloides, A. longicornis and Libelloides coccajus the latter. However, intracellular electrophysiological are rapidly-flying neuropteran insects, which hunt in open recordings or microspectrophotometry on these tiny pho country for flying insects. These owlflies are only adapted toreceptors have not been done so their spectral sensi for daytime activity. They have large double eyes, which tivity is unknown (P. Stušek, personal communication). structurally correspond to optical refracting superposition Advantages of UV vision eyes (Ast, 1920; Gogala & Michieli, 1965; Schneider et What are the advantages of using UV light for locating al., 1978; forreview, seeNilsson, 1989). -
Test Key to British Blowflies (Calliphoridae) And
Draft key to British Calliphoridae and Rhinophoridae Steven Falk 2016 BRITISH BLOWFLIES (CALLIPHORIDAE) AND WOODLOUSE FLIES (RHINOPHORIDAE) DRAFT KEY March 2016 Steven Falk Feedback to [email protected] 1 Draft key to British Calliphoridae and Rhinophoridae Steven Falk 2016 PREFACE This informal publication attempts to update the resources currently available for identifying the families Calliphoridae and Rhinophoridae. Prior to this, British dipterists have struggled because unless you have a copy of the Fauna Ent. Scand. volume for blowflies (Rognes, 1991), you will have been largely reliant on Van Emden's 1954 RES Handbook, which does not include all the British species (notably the common Pollenia pediculata), has very outdated nomenclature, and very outdated classification - with several calliphorids and tachinids placed within the Rhinophoridae and Eurychaeta palpalis placed in the Sarcophagidae. As well as updating keys, I have also taken the opportunity to produce new species accounts which summarise what I know of each species and act as an invitation and challenge to others to update, correct or clarify what I have written. As a result of my recent experience of producing an attractive and fairly user-friendly new guide to British bees, I have tried to replicate that approach here, incorporating lots of photos and clear, conveniently positioned diagrams. Presentation of identification literature can have a big impact on the popularity of an insect group and the accuracy of the records that result. Calliphorids and rhinophorids are fascinating flies, sometimes of considerable economic and medicinal value and deserve to be well recorded. What is more, many gaps still remain in our knowledge. -
Strasbourg, 19 April 2013
Strasbourg, 25 October 2013 T-PVS (2013) 17 [tpvs17e_2013.doc] CONVENTION ON THE CONSERVATION OF EUROPEAN WILDLIFE AND NATURAL HABITATS Group of Experts on the Conservation of Invertebrates Tirana, Albania 23-24 September 2013 ---ooOoo--- REPORT Document prepared by the Directorate of Democratic Governance This document will not be distributed at the meeting. Please bring this copy. Ce document ne sera plus distribué en réunion. Prière de vous munir de cet exemplaire. T-PVS (2013) 17 - 2 - CONTENTS 1. Meeting report ................................................................................................................................... 3 2. Appendix 1: Agenda .......................................................................................................................... 6 3. Appendix 2: List of participants ........................................................................................................ 9 4. Appendix 3: Compilation of National Reports .................................................................................. 10 5. Appendix 4: Draft Recommendation on threats by neurotoxic insecticides to pollinators ................ 75 * * * The Standing Committee is invited to: 1. Take note of the report of the meeting; 2. Thank the Albanian government for the efficient preparation of the meeting and the excellent hospitality; 3. Continue with Bern Convention engagement with invertebrate conservation issues by further encouraging and monitoring national implementation of European Strategy for the Conservation