Dinosaur Warriors
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A Neoceratopsian Dinosaur from the Early Cretaceous of Mongolia And
ARTICLE https://doi.org/10.1038/s42003-020-01222-7 OPEN A neoceratopsian dinosaur from the early Cretaceous of Mongolia and the early evolution of ceratopsia ✉ Congyu Yu 1 , Albert Prieto-Marquez2, Tsogtbaatar Chinzorig 3,4, Zorigt Badamkhatan4,5 & Mark Norell1 1234567890():,; Ceratopsia is a diverse dinosaur clade from the Middle Jurassic to Late Cretaceous with early diversification in East Asia. However, the phylogeny of basal ceratopsians remains unclear. Here we report a new basal neoceratopsian dinosaur Beg tse based on a partial skull from Baruunbayan, Ömnögovi aimag, Mongolia. Beg is diagnosed by a unique combination of primitive and derived characters including a primitively deep premaxilla with four pre- maxillary teeth, a trapezoidal antorbital fossa with a poorly delineated anterior margin, very short dentary with an expanded and shallow groove on lateral surface, the derived presence of a robust jugal having a foramen on its anteromedial surface, and five equally spaced tubercles on the lateral ridge of the surangular. This is to our knowledge the earliest known occurrence of basal neoceratopsian in Mongolia, where this group was previously only known from Late Cretaceous strata. Phylogenetic analysis indicates that it is sister to all other neoceratopsian dinosaurs. 1 Division of Vertebrate Paleontology, American Museum of Natural History, New York 10024, USA. 2 Institut Català de Paleontologia Miquel Crusafont, ICTA-ICP, Edifici Z, c/de les Columnes s/n Campus de la Universitat Autònoma de Barcelona, 08193 Cerdanyola del Vallès Sabadell, Barcelona, Spain. 3 Department of Biological Sciences, North Carolina State University, Raleigh, NC 27695, USA. 4 Institute of Paleontology, Mongolian Academy of Sciences, ✉ Ulaanbaatar 15160, Mongolia. -
MEET the DINOSAURS! WHAT’S in a NAME? a LOT If You Are a Dinosaur!
MEET THE DINOSAURS! WHAT’S IN A NAME? A LOT if you are a dinosaur! I will call you Dyoplosaurus! Most dinosaurs get their names from the ancient Greek and Latin languages. And I will call you Mojoceratops! And sometimes they are named after a defining feature on their body. Their names are made up of word parts that describe the dinosaur. The name must be sent to a special group of people called the International Commission on Zoological Nomenclature to be approved! Did You The word DINOSAUR comes from the Greek word meaning terrible lizard and was first said by Know? Sir Richard Owen in 1841. © 2013 Omaha’s Henry Doorly Zoo & Aquarium® | 3 SEE IF YOU CAN FIND OUT THE MEANING OF SOME OF OUR DINOSAUR’S NAMES. Dinosaur names are not just tough to pronounce, they often have meaning. Dinosaur Name MEANING of Dinosaur Name Carnotaurus (KAR-no-TORE-us) Means “flesh-eating bull” Spinosaurus (SPY-nuh-SORE-us) Dyoplosaurus (die-o-pluh-SOR-us) Amargasaurus (ah-MAR-guh-SORE-us) Omeisaurus (Oh-MY-ee-SORE-us) Pachycephalosaurus (pak-ee-SEF-uh-low-SORE-us) Tuojiangosaurus (toh-HWANG-uh-SORE-us) Yangchuanosaurus (Yang-chew-ON-uh-SORE-us) Quetzalcoatlus (KWET-zal-coe-AT-lus) Ouranosaurus (ooh-RAN-uh-SORE-us) Parasaurolophus (PAIR-uh-so-ROL-uh-PHUS) Kosmoceratops (KOZ-mo-SARA-tops) Mojoceratops (moe-joe-SEH-rah-tops) Triceratops (try-SER-uh-TOPS) Tyrannosaurus rex (tuh-RAN-uh-SORE-us) Find the answers by visiting the Resource Library at Carnotaurus A: www.OmahaZoo.com/Education. Search word: Dinosaurs 4 | © 2013 Omaha’s Henry Doorly Zoo & Aquarium® DINO DEFENSE All animals in the wild have to protect themselves. -
Offer Your Guests a Visually Stunning and Interactive Experience They Won’T Find Anywhere Else World of Animals
Creative Arts & Attractions Offer Your Guests a Visually Stunning and Interactive Experience They Won’t Find Anywhere Else World of Animals Entertain guests with giant illuminated, eye-catching displays of animals from around the world. Animal displays are made in the ancient Eastern tradition of lantern-making with 3-D metal frames, fiberglass, and acrylic materials. Unique and captivating displays will provide families and friends with a lifetime of memories. Animatronic Dinosaurs Fascinate patrons with an impressive visual spectacle in our exhibitions. Featured with lifelike appearances, vivid movements and roaring sounds. From the very small to the gigantic dinosaurs, we have them all. Everything needed for a realistic and immersive experience for your patrons. Providing interactive options for your event including riding dinosaurs and dinosaur inflatable slides. Enhancing your merchandise stores with dinosaur balloons and toys. Animatronic Dinosaur Options Abelisaurus Maiasaura Acrocanthosaurus Megalosaurus Agilisaurus Olorotitan arharensis Albertosaurus Ornithomimus Allosaurus Ouranosaurus nigeriensis Ankylosaurus Oviraptor philoceratops Apatosaurus Pachycephalosaurus wyomingensis Archaeopteryx Parasaurolophus Baryonyx Plateosaurus Brachiosaurus Protoceratops andrewsi Carcharodontosaurus Pterosauria Carnotaurus Pteranodon longiceps Ceratosaurus Raptorex Coelophysis Rugops Compsognathus Spinosaurus Deinonychus Staurikosaurus pricei Dilophosaurus Stegoceras Diplodocus Stegosaurus Edmontosaurus Styracosaurus Eoraptor Lunensis Suchomimus -
At Carowinds
at Carowinds EDUCATOR’S GUIDE CLASSROOM LESSON PLANS & FIELD TRIP ACTIVITIES Table of Contents at Carowinds Introduction The Field Trip ................................... 2 The Educator’s Guide ....................... 3 Field Trip Activity .................................. 4 Lesson Plans Lesson 1: Form and Function ........... 6 Lesson 2: Dinosaur Detectives ....... 10 Lesson 3: Mesozoic Math .............. 14 Lesson 4: Fossil Stories.................. 22 Games & Puzzles Crossword Puzzles ......................... 29 Logic Puzzles ................................. 32 Word Searches ............................... 37 Answer Keys ...................................... 39 Additional Resources © 2012 Dinosaurs Unearthed Recommended Reading ................. 44 All rights reserved. Except for educational fair use, no portion of this guide may be reproduced, stored in a retrieval system, or transmitted in any form or by any Dinosaur Data ................................ 45 means—electronic, mechanical, photocopy, recording, or any other without Discovering Dinosaurs .................... 52 explicit prior permission from Dinosaurs Unearthed. Multiple copies may only be made by or for the teacher for class use. Glossary .............................................. 54 Content co-created by TurnKey Education, Inc. and Dinosaurs Unearthed, 2012 Standards www.turnkeyeducation.net www.dinosaursunearthed.com Curriculum Standards .................... 59 Introduction The Field Trip From the time of the first exhibition unveiled in 1854 at the Crystal -
Investigating Sexual Dimorphism in Ceratopsid Horncores
University of Calgary PRISM: University of Calgary's Digital Repository Graduate Studies The Vault: Electronic Theses and Dissertations 2013-01-25 Investigating Sexual Dimorphism in Ceratopsid Horncores Borkovic, Benjamin Borkovic, B. (2013). Investigating Sexual Dimorphism in Ceratopsid Horncores (Unpublished master's thesis). University of Calgary, Calgary, AB. doi:10.11575/PRISM/26635 http://hdl.handle.net/11023/498 master thesis University of Calgary graduate students retain copyright ownership and moral rights for their thesis. You may use this material in any way that is permitted by the Copyright Act or through licensing that has been assigned to the document. For uses that are not allowable under copyright legislation or licensing, you are required to seek permission. Downloaded from PRISM: https://prism.ucalgary.ca UNIVERSITY OF CALGARY Investigating Sexual Dimorphism in Ceratopsid Horncores by Benjamin Borkovic A THESIS SUBMITTED TO THE FACULTY OF GRADUATE STUDIES IN PARTIAL FULFILMENT OF THE REQUIREMENTS FOR THE DEGREE OF MASTER OF SCIENCE DEPARTMENT OF BIOLOGICAL SCIENCES CALGARY, ALBERTA JANUARY, 2013 © Benjamin Borkovic 2013 Abstract Evidence for sexual dimorphism was investigated in the horncores of two ceratopsid dinosaurs, Triceratops and Centrosaurus apertus. A review of studies of sexual dimorphism in the vertebrate fossil record revealed methods that were selected for use in ceratopsids. Mountain goats, bison, and pronghorn were selected as exemplar taxa for a proof of principle study that tested the selected methods, and informed and guided the investigation of sexual dimorphism in dinosaurs. Skulls of these exemplar taxa were measured in museum collections, and methods of analysing morphological variation were tested for their ability to demonstrate sexual dimorphism in their horns and horncores. -
New Book Catalog
New Book Catalog www.doverpublications.com Winter 2020 Connect with us! New Books for a OVER 50 NEW TITLES! New Year! Save 25% on 50 Favorites! Take an Additional $10 Off & Free Shipping See pages 46–47 See page 2 New Book Q1 2020 01-09.indd 1 12/5/19 11:03 AM 2020: New Year, New You $10 off your order of $40 or more TO REDEEM: When ordering by mail or fax, please IXIA PRESS NEW! enter the coupon code in the title column on the last Named after the South African flower that represents happiness, line of the order form. When ordering online, enter the Ixia Press presents inspiring books on leadership, business, spirituality, NEW! coupon code during checkout. and wellness that foster a spirit of personal and professional growth PLEASE NOTE: You must provide the Coupon Code Use to receive your discount. Orders must be received by and exploration. The imprint combines completely original works with seminal February 22, 2020. Shipping and handling, taxes, Coupon and gift certificates do not apply toward the classics given a fresh new lease on life, all written by some of the most respected Code purchase requirement. We reserve the right to change or discontinue this offer at any time. Your authors in their fields. merchandise total after your coupon deduction must CNWC be $50 or more to be eligible for Free Shipping. Can’t be combined with any other coupon. Reader-friendly handbook is structured around the ways in which crystals can Offer Ends 2/22/20 be used to improve specifi c life areas HIGHLIGHTS Valentine’s Day . -
A Revision of the Ceratopsia Or Horned Dinosaurs
MEMOIRS OF THE PEABODY MUSEUM OF NATURAL HISTORY VOLUME III, 1 A.R1 A REVISION orf tneth< CERATOPSIA OR HORNED DINOSAURS BY RICHARD SWANN LULL STERLING PROFESSOR OF PALEONTOLOGY AND DIRECTOR OF PEABODY MUSEUM, YALE UNIVERSITY LVXET NEW HAVEN, CONN. *933 MEMOIRS OF THE PEABODY MUSEUM OF NATURAL HISTORY YALE UNIVERSITY Volume I. Odontornithes: A Monograph on the Extinct Toothed Birds of North America. By Othniel Charles Marsh. Pp. i-ix, 1-201, pis. 1-34, text figs. 1-40. 1880. To be obtained from the Peabody Museum. Price $3. Volume II. Part 1. Brachiospongidae : A Memoir on a Group of Silurian Sponges. By Charles Emerson Beecher. Pp. 1-28, pis. 1-6, text figs. 1-4. 1889. To be obtained from the Peabody Museum. Price $1. Volume III. Part 1. American Mesozoic Mammalia. By George Gaylord Simp- son. Pp. i-xvi, 1-171, pis. 1-32, text figs. 1-62. 1929. To be obtained from the Yale University Press, New Haven, Conn. Price $5. Part 2. A Remarkable Ground Sloth. By Richard Swann Lull. Pp. i-x, 1-20, pis. 1-9, text figs. 1-3. 1929. To be obtained from the Yale University Press, New Haven, Conn. Price $1. Part 3. A Revision of the Ceratopsia or Horned Dinosaurs. By Richard Swann Lull. Pp. i-xii, 1-175, pis. I-XVII, text figs. 1-42. 1933. To be obtained from the Peabody Museum. Price $5 (bound in cloth), $4 (bound in paper). Part 4. The Merycoidodontidae, an Extinct Group of Ruminant Mammals. By Malcolm Rutherford Thorpe. In preparation. -
Histology and Ontogeny of Pachyrhinosaurus Nasal Bosses By
Histology and Ontogeny of Pachyrhinosaurus Nasal Bosses by Elizabeth Kruk A thesis submitted in partial fulfillment of the requirements for the degree of Master of Science in Systematics and Evolution Department of Biological Sciences University of Alberta © Elizabeth Kruk, 2015 Abstract Pachyrhinosaurus is a peculiar ceratopsian known only from Upper Cretaceous strata of Alberta and the North Slope of Alaska. The genus consists of three described species Pachyrhinosaurus canadensis, Pachyrhinosaurus lakustai, and Pachyrhinosaurus perotorum that are distinguishable by cranial characteristics, including parietal horn shape and orientation, absence/presence of a rostral comb, median parietal bar horns, and profile of the nasal boss. A fourth species of Pachyrhinosaurus is described herein and placed into its phylogenetic context within Centrosaurinae. This new species forms a polytomy at the crown with Pachyrhinosaurus canadensis and Pachyrhinosaurus perotorum, with Pachyrhinosaurus lakustai falling basal to that polytomy. The diagnostic features of this new species are an apomorphic, laterally curved Process 3 horns and a thick longitudinal ridge separating the supraorbital bosses. Another focus is investigating the ontogeny of Pachyrhinosaurus nasal bosses in a histological context. Previously, little work has been done on cranial histology in ceratopsians, focusing instead on potential integumentary structures, the parietals of Triceratops, and how surface texture relates to underlying histological structures. An ontogenetic series is established for the nasal bosses of Pachyrhinosaurus at both relative (subadult versus adult) and fine scale (Stages 1-5). It was demonstrated that histology alone can indicate relative ontogenetic level, but not stages of a finer scale. Through Pachyrhinosaurus ontogeny the nasal boss undergoes increased vascularity and secondary remodeling with a reduction in osteocyte lacunar density. -
Inferring Body Mass in Extinct Terrestrial Vertebrates and the Evolution of Body Size in a Model-Clade of Dinosaurs (Ornithopoda)
Inferring Body Mass in Extinct Terrestrial Vertebrates and the Evolution of Body Size in a Model-Clade of Dinosaurs (Ornithopoda) by Nicolás Ernesto José Campione Ruben A thesis submitted in conformity with the requirements for the degree of Doctor of Philosophy Ecology and Evolutionary Biology University of Toronto © Copyright by Nicolás Ernesto José Campione Ruben 2013 Inferring body mass in extinct terrestrial vertebrates and the evolution of body size in a model-clade of dinosaurs (Ornithopoda) Nicolás E. J. Campione Ruben Doctor of Philosophy Ecology and Evolutionary Biology University of Toronto 2013 Abstract Organismal body size correlates with almost all aspects of ecology and physiology. As a result, the ability to infer body size in the fossil record offers an opportunity to interpret extinct species within a biological, rather than simply a systematic, context. Various methods have been proposed by which to estimate body mass (the standard measure of body size) that center on two main approaches: volumetric reconstructions and extant scaling. The latter are particularly contentious when applied to extinct terrestrial vertebrates, particularly stem-based taxa for which living relatives are difficult to constrain, such as non-avian dinosaurs and non-therapsid synapsids, resulting in the use of volumetric models that are highly influenced by researcher bias. However, criticisms of scaling models have not been tested within a comprehensive extant dataset. Based on limb measurements of 200 mammals and 47 reptiles, linear models were generated between limb measurements (length and circumference) and body mass to test the hypotheses that phylogenetic history, limb posture, and gait drive the relationship between stylopodial circumference and body mass as critics suggest. -
Ankylosaurus Magniventris
FOR OUR ENGLISH-SPEAKING GUESTS: “Dinosauria” is the scientific name for dinosaurs, and derives from Ancient Greek: “Deinos”, meaning “terrible, potent or fearfully great”, and “sauros”, meaning “lizard or reptile”. Dinosaurs are among the most successful animals in the history of life on Earth. They dominated the planet for nearly 160 million years during the entire Mesozoic era, from the Triassic period 225 million years ago. It was followed by the Jurassic period and then the Cretaceous period, which ended 65 million years ago with the extinction of dinosaurs. Here follows a detailed overview of the dinosaur exhibition that is on display at Dinosauria, which is produced by Dinosauriosmexico. Please have a look at the name printed on top of the Norwegian sign right by each model to identify the correct dinosaur. (Note that the shorter Norwegian text is similar, but not identical to the English text, which is more in-depth) If you have any questions, please ask our staff wearing colorful shirts with flower decorations. We hope you enjoy your visit at INSPIRIA science center! Ankylosaurus magniventris Period: Late Cretaceous (65 million years ago) Known locations: United States and Canada Diet: Herbivore Size: 9 m long Its name means "fused lizard". This dinosaur lived in North America 65 million years ago during the Late Cretaceous. This is the most widely known armoured dinosaur, with a club on its tail. The final vertebrae of the tail were immobilized by overlapping the connections, turning it into a solid handle. The tail club (or tail knob) is composed of several osteoderms fused into one unit, with two larger plates on the sides. -
A Battering Ram?
A Battering Ram? All evidence suggests that Pachycephalosaur skulls were built to withstand extreme forces 9 inches of solid bone Bone organized in a radial arrangement- structural support Articulation btw back of skull and vertebrae oriented to transfer forces linearly Articulation btw back of skull and vertebral column built to withstand sideways forces Vertebral column has tongue and groove articulations Spinal column is an S-shaped shock absorber BUT There is no ‘locking’ mechanism on skull to keep battering heads aligned Some Pachycephalosaurs have imprinted blood vessels on dome These factors suggests that head- butting may not be likely Intraspecies Competition (typically male-male) Females are typically choosey Why? Because they have more to loose Common rule in biology: Females are expensive to lose, males are cheap (e.g. deer hunting) Females choose the male most likely to provide the most successful offspring Males compete with each other for access to female vs. female chooses the strongest male Choosey females // Strong males have more offspring => SEXUAL selection Many ways to do this... But: In general, maximize competition and minimize accidental deaths (= no fitness) http://www.youtube.com/watch?v=PontCXFgs0M http://www.metacafe.com/watch/1941236/giraffe_fight/ http://www.youtube.com/watch? http://www.youtube.com/watch?v=DYDx1y38vGw http://www.youtube.com/watch?v=ULRtdk-3Yh4 Air-filled horn cores vs. solid bone skull caps... Gotta have a cheezy animated slide. Homalocephale Pachycephalosaurus Prenocephale Tylocephale Stegoceras Head butting Pachycephalosaurs Bone structure was probably strong enough to withstand collision Convex nature would favor glancing blows Instead, dome and spines seem better suited for “flank butting” So.. -
A New Maastrichtian Species of the Centrosaurine Ceratopsid Pachyrhinosaurus from the North Slope of Alaska
A new Maastrichtian species of the centrosaurine ceratopsid Pachyrhinosaurus from the North Slope of Alaska ANTHONY R. FIORILLO and RONALD S. TYKOSKI Fiorillo, A.R. and Tykoski, R.S. 2012. A new Maastrichtian species of the centrosaurine ceratopsid Pachyrhinosaurus from the North Slope of Alaska. Acta Palaeontologica Polonica 57 (3): 561–573. The Cretaceous rocks of the Prince Creek Formation contain the richest record of polar dinosaurs found anywhere in the world. Here we describe a new species of horned dinosaur, Pachyrhinosaurus perotorum that exhibits an apomorphic character in the frill, as well as a unique combination of other characters. Phylogenetic analysis of 16 taxa of ceratopsians failed to resolve relationships between P. perotorum and other Pachyrhinosaurus species (P. canadensis and P. lakustai). P. perotorum shares characters with each of the previously known species that are not present in the other, including very large nasal and supraorbital bosses that are nearly in contact and separated only by a narrow groove as in P. canadensis, and a rostral comb formed by the nasals and premaxillae as in P. lakustai. P. perotorum is the youngest centrosaurine known (70–69 Ma), and the locality that produced the taxon, the Kikak−Tegoseak Quarry, is close to the highest latitude for recovery of ceratopsid remains. Key words: Dinosauria, Centrosaurinae, Cretaceous, Prince Creek Formation, Kikak−Tegoseak Quarry, Arctic. Anthony R. Fiorillo [[email protected]] and Ronald S. Tykoski [[email protected]], Perot Museum of Nature and Science, 2201 N. Field Street, Dallas, TX 75202, USA. Received 4 April 2011, accepted 23 July 2011, available online 26 August 2011.