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A Neoceratopsian Dinosaur from the Early Cretaceous of Mongolia And
ARTICLE https://doi.org/10.1038/s42003-020-01222-7 OPEN A neoceratopsian dinosaur from the early Cretaceous of Mongolia and the early evolution of ceratopsia ✉ Congyu Yu 1 , Albert Prieto-Marquez2, Tsogtbaatar Chinzorig 3,4, Zorigt Badamkhatan4,5 & Mark Norell1 1234567890():,; Ceratopsia is a diverse dinosaur clade from the Middle Jurassic to Late Cretaceous with early diversification in East Asia. However, the phylogeny of basal ceratopsians remains unclear. Here we report a new basal neoceratopsian dinosaur Beg tse based on a partial skull from Baruunbayan, Ömnögovi aimag, Mongolia. Beg is diagnosed by a unique combination of primitive and derived characters including a primitively deep premaxilla with four pre- maxillary teeth, a trapezoidal antorbital fossa with a poorly delineated anterior margin, very short dentary with an expanded and shallow groove on lateral surface, the derived presence of a robust jugal having a foramen on its anteromedial surface, and five equally spaced tubercles on the lateral ridge of the surangular. This is to our knowledge the earliest known occurrence of basal neoceratopsian in Mongolia, where this group was previously only known from Late Cretaceous strata. Phylogenetic analysis indicates that it is sister to all other neoceratopsian dinosaurs. 1 Division of Vertebrate Paleontology, American Museum of Natural History, New York 10024, USA. 2 Institut Català de Paleontologia Miquel Crusafont, ICTA-ICP, Edifici Z, c/de les Columnes s/n Campus de la Universitat Autònoma de Barcelona, 08193 Cerdanyola del Vallès Sabadell, Barcelona, Spain. 3 Department of Biological Sciences, North Carolina State University, Raleigh, NC 27695, USA. 4 Institute of Paleontology, Mongolian Academy of Sciences, ✉ Ulaanbaatar 15160, Mongolia. -
A Phylogenetic Analysis of the Basal Ornithischia (Reptilia, Dinosauria)
A PHYLOGENETIC ANALYSIS OF THE BASAL ORNITHISCHIA (REPTILIA, DINOSAURIA) Marc Richard Spencer A Thesis Submitted to the Graduate College of Bowling Green State University in partial fulfillment of the requirements of the degree of MASTER OF SCIENCE December 2007 Committee: Margaret M. Yacobucci, Advisor Don C. Steinker Daniel M. Pavuk © 2007 Marc Richard Spencer All Rights Reserved iii ABSTRACT Margaret M. Yacobucci, Advisor The placement of Lesothosaurus diagnosticus and the Heterodontosauridae within the Ornithischia has been problematic. Historically, Lesothosaurus has been regarded as a basal ornithischian dinosaur, the sister taxon to the Genasauria. Recent phylogenetic analyses, however, have placed Lesothosaurus as a more derived ornithischian within the Genasauria. The Fabrosauridae, of which Lesothosaurus was considered a member, has never been phylogenetically corroborated and has been considered a paraphyletic assemblage. Prior to recent phylogenetic analyses, the problematic Heterodontosauridae was placed within the Ornithopoda as the sister taxon to the Euornithopoda. The heterodontosaurids have also been considered as the basal member of the Cerapoda (Ornithopoda + Marginocephalia), the sister taxon to the Marginocephalia, and as the sister taxon to the Genasauria. To reevaluate the placement of these taxa, along with other basal ornithischians and more derived subclades, a phylogenetic analysis of 19 taxonomic units, including two outgroup taxa, was performed. Analysis of 97 characters and their associated character states culled, modified, and/or rescored from published literature based on published descriptions, produced four most parsimonious trees. Consistency and retention indices were calculated and a bootstrap analysis was performed to determine the relative support for the resultant phylogeny. The Ornithischia was recovered with Pisanosaurus as its basalmost member. -
71St Annual Meeting Society of Vertebrate Paleontology Paris Las Vegas Las Vegas, Nevada, USA November 2 – 5, 2011 SESSION CONCURRENT SESSION CONCURRENT
ISSN 1937-2809 online Journal of Supplement to the November 2011 Vertebrate Paleontology Vertebrate Society of Vertebrate Paleontology Society of Vertebrate 71st Annual Meeting Paleontology Society of Vertebrate Las Vegas Paris Nevada, USA Las Vegas, November 2 – 5, 2011 Program and Abstracts Society of Vertebrate Paleontology 71st Annual Meeting Program and Abstracts COMMITTEE MEETING ROOM POSTER SESSION/ CONCURRENT CONCURRENT SESSION EXHIBITS SESSION COMMITTEE MEETING ROOMS AUCTION EVENT REGISTRATION, CONCURRENT MERCHANDISE SESSION LOUNGE, EDUCATION & OUTREACH SPEAKER READY COMMITTEE MEETING POSTER SESSION ROOM ROOM SOCIETY OF VERTEBRATE PALEONTOLOGY ABSTRACTS OF PAPERS SEVENTY-FIRST ANNUAL MEETING PARIS LAS VEGAS HOTEL LAS VEGAS, NV, USA NOVEMBER 2–5, 2011 HOST COMMITTEE Stephen Rowland, Co-Chair; Aubrey Bonde, Co-Chair; Joshua Bonde; David Elliott; Lee Hall; Jerry Harris; Andrew Milner; Eric Roberts EXECUTIVE COMMITTEE Philip Currie, President; Blaire Van Valkenburgh, Past President; Catherine Forster, Vice President; Christopher Bell, Secretary; Ted Vlamis, Treasurer; Julia Clarke, Member at Large; Kristina Curry Rogers, Member at Large; Lars Werdelin, Member at Large SYMPOSIUM CONVENORS Roger B.J. Benson, Richard J. Butler, Nadia B. Fröbisch, Hans C.E. Larsson, Mark A. Loewen, Philip D. Mannion, Jim I. Mead, Eric M. Roberts, Scott D. Sampson, Eric D. Scott, Kathleen Springer PROGRAM COMMITTEE Jonathan Bloch, Co-Chair; Anjali Goswami, Co-Chair; Jason Anderson; Paul Barrett; Brian Beatty; Kerin Claeson; Kristina Curry Rogers; Ted Daeschler; David Evans; David Fox; Nadia B. Fröbisch; Christian Kammerer; Johannes Müller; Emily Rayfield; William Sanders; Bruce Shockey; Mary Silcox; Michelle Stocker; Rebecca Terry November 2011—PROGRAM AND ABSTRACTS 1 Members and Friends of the Society of Vertebrate Paleontology, The Host Committee cordially welcomes you to the 71st Annual Meeting of the Society of Vertebrate Paleontology in Las Vegas. -
Investigating Sexual Dimorphism in Ceratopsid Horncores
University of Calgary PRISM: University of Calgary's Digital Repository Graduate Studies The Vault: Electronic Theses and Dissertations 2013-01-25 Investigating Sexual Dimorphism in Ceratopsid Horncores Borkovic, Benjamin Borkovic, B. (2013). Investigating Sexual Dimorphism in Ceratopsid Horncores (Unpublished master's thesis). University of Calgary, Calgary, AB. doi:10.11575/PRISM/26635 http://hdl.handle.net/11023/498 master thesis University of Calgary graduate students retain copyright ownership and moral rights for their thesis. You may use this material in any way that is permitted by the Copyright Act or through licensing that has been assigned to the document. For uses that are not allowable under copyright legislation or licensing, you are required to seek permission. Downloaded from PRISM: https://prism.ucalgary.ca UNIVERSITY OF CALGARY Investigating Sexual Dimorphism in Ceratopsid Horncores by Benjamin Borkovic A THESIS SUBMITTED TO THE FACULTY OF GRADUATE STUDIES IN PARTIAL FULFILMENT OF THE REQUIREMENTS FOR THE DEGREE OF MASTER OF SCIENCE DEPARTMENT OF BIOLOGICAL SCIENCES CALGARY, ALBERTA JANUARY, 2013 © Benjamin Borkovic 2013 Abstract Evidence for sexual dimorphism was investigated in the horncores of two ceratopsid dinosaurs, Triceratops and Centrosaurus apertus. A review of studies of sexual dimorphism in the vertebrate fossil record revealed methods that were selected for use in ceratopsids. Mountain goats, bison, and pronghorn were selected as exemplar taxa for a proof of principle study that tested the selected methods, and informed and guided the investigation of sexual dimorphism in dinosaurs. Skulls of these exemplar taxa were measured in museum collections, and methods of analysing morphological variation were tested for their ability to demonstrate sexual dimorphism in their horns and horncores. -
New Oviraptorid Dinosaur (Dinosauria: Oviraptorosauria) from the Nemegt Formation of Southwestern Mongolia
Bull. Natn. Sci. Mus., Tokyo, Ser. C, 30, pp. 95–130, December 22, 2004 New Oviraptorid Dinosaur (Dinosauria: Oviraptorosauria) from the Nemegt Formation of Southwestern Mongolia Junchang Lü1, Yukimitsu Tomida2, Yoichi Azuma3, Zhiming Dong4 and Yuong-Nam Lee5 1 Institute of Geology, Chinese Academy of Geological Sciences, Beijing 100037, China 2 National Science Museum, 3–23–1 Hyakunincho, Shinjukuku, Tokyo 169–0073, Japan 3 Fukui Prefectural Dinosaur Museum, 51–11 Terao, Muroko, Katsuyama 911–8601, Japan 4 Institute of Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China 5 Korea Institute of Geoscience and Mineral Resources, Geology & Geoinformation Division, 30 Gajeong-dong, Yuseong-gu, Daejeon 305–350, South Korea Abstract Nemegtia barsboldi gen. et sp. nov. here described is a new oviraptorid dinosaur from the Late Cretaceous (mid-Maastrichtian) Nemegt Formation of southwestern Mongolia. It differs from other oviraptorids in the skull having a well-developed crest, the anterior margin of which is nearly vertical, and the dorsal margin of the skull and the anterior margin of the crest form nearly 90°; the nasal process of the premaxilla being less exposed on the dorsal surface of the skull than those in other known oviraptorids; the length of the frontal being approximately one fourth that of the parietal along the midline of the skull. Phylogenetic analysis shows that Nemegtia barsboldi is more closely related to Citipati osmolskae than to any other oviraptorosaurs. Key words : Nemegt Basin, Mongolia, Nemegt Formation, Late Cretaceous, Oviraptorosauria, Nemegtia. dae, and Caudipterygidae (Barsbold, 1976; Stern- Introduction berg, 1940; Currie, 2000; Clark et al., 2001; Ji et Oviraptorosaurs are generally regarded as non- al., 1998; Zhou and Wang, 2000; Zhou et al., avian theropod dinosaurs (Osborn, 1924; Bars- 2000). -
Histology and Ontogeny of Pachyrhinosaurus Nasal Bosses By
Histology and Ontogeny of Pachyrhinosaurus Nasal Bosses by Elizabeth Kruk A thesis submitted in partial fulfillment of the requirements for the degree of Master of Science in Systematics and Evolution Department of Biological Sciences University of Alberta © Elizabeth Kruk, 2015 Abstract Pachyrhinosaurus is a peculiar ceratopsian known only from Upper Cretaceous strata of Alberta and the North Slope of Alaska. The genus consists of three described species Pachyrhinosaurus canadensis, Pachyrhinosaurus lakustai, and Pachyrhinosaurus perotorum that are distinguishable by cranial characteristics, including parietal horn shape and orientation, absence/presence of a rostral comb, median parietal bar horns, and profile of the nasal boss. A fourth species of Pachyrhinosaurus is described herein and placed into its phylogenetic context within Centrosaurinae. This new species forms a polytomy at the crown with Pachyrhinosaurus canadensis and Pachyrhinosaurus perotorum, with Pachyrhinosaurus lakustai falling basal to that polytomy. The diagnostic features of this new species are an apomorphic, laterally curved Process 3 horns and a thick longitudinal ridge separating the supraorbital bosses. Another focus is investigating the ontogeny of Pachyrhinosaurus nasal bosses in a histological context. Previously, little work has been done on cranial histology in ceratopsians, focusing instead on potential integumentary structures, the parietals of Triceratops, and how surface texture relates to underlying histological structures. An ontogenetic series is established for the nasal bosses of Pachyrhinosaurus at both relative (subadult versus adult) and fine scale (Stages 1-5). It was demonstrated that histology alone can indicate relative ontogenetic level, but not stages of a finer scale. Through Pachyrhinosaurus ontogeny the nasal boss undergoes increased vascularity and secondary remodeling with a reduction in osteocyte lacunar density. -
A Subadult Individual of Styracosaurus Albertensis
Vertebrate Anatomy Morphology Palaeontology 8:67–95 67 ISSN 2292-1389 A subadult individual of Styracosaurus albertensis (Ornithischia: Ceratopsidae) with comments on ontogeny and intraspecific variation inStyracosaurus and Centrosaurus Caleb M. Brown1,*, Robert B. Holmes2, Philip J. Currie2 1Royal Tyrrell Museum of Palaeontology, Box 7500, Drumheller, AB, T0J 0Y0, Canada; [email protected] 2Department of Biological Sciences, University of Alberta, Edmonton, Alberta, T6G 2E9, Canada; [email protected]; [email protected] Abstract: Styracosaurus albertensis is an iconic centrosaurine horned dinosaur from the Campanian of Alberta, Canada, known for its large spike-like parietal processes. Although described over 100 years ago, subsequent dis- coveries were rare until the last few decades, during which time several new skulls, skeletons, and bonebeds were found. Here we described an immature individual, the smallest known for the species, represented by a complete skull and fragmentary skeleton. Although ~80% maximum size, it possesses a suite of characters associated with immaturity, and is regarded as a subadult. The ornamentation is characterized by a small, recurved, but fused nasal horncore; short, rounded postorbital horncores; and short, triangular, and flat parietal processes. Using this specimen, and additional skulls and bonebed material, the cranial ontogeny of Styracosaurus is described, and compared to Centrosaurus. In early ontogeny, the nasal horncores of Styracosaurus and Centrosaurus are thin, recurved, and unfused, but in the former the recurved morphology is retained into large adult size and the horncore never develops the procurved morphology common in Centrosaurus. The postorbital horncores of Styracosaurus are shorter and more rounded than those of Centrosaurus throughout ontogeny, and show great- er resorption later in ontogeny. -
Exam 2 Review Sheet
EART 65 Spring 2008 EXAM 2 REVIEW SHEET To prepare for the upcoming exam in EART 65 you should: What to bring? 1. A scantron test form (#F1712: ‘big pink one’) and a Number 2 pencil 2. One 8.5x11” sheet on which you can use one side to write notes. Other Issues 1. Review your lecture notes. The test will cover material from the lecture on 4/24/08 up to and including the material on Ornithopods on 05/13/08. 2. Review the relevant text chapters (including figures and cladograms). 3. Review the section assignments and quizzes for these weeks. The test will include bone identification questions that flow from your work over the last two weeks in section. 4. Make sure you can read and manipulate cladograms. There will be questions on this subject, just like on the last exam. 5. Look over the review sheet and make sure you can answer questions and know the importance of different terms. 6. Don’t forget the geologic time scale!! 7. Do not bring electronics to the exam. No iPods, cellphones, etc. If you are spotted using such a device during the exam, your exam will be confiscated. 8. There will be no makeup exams. Show up and take the exam, or give me the name of your doctor who will vouch for your absence. 9. The test will be one hour long. Class will continue after the exam, though we’ll only be checking out a video on Cretaceous animals. 10. Don’t forget the geologic time scale!! 11. -
A Battering Ram?
A Battering Ram? All evidence suggests that Pachycephalosaur skulls were built to withstand extreme forces 9 inches of solid bone Bone organized in a radial arrangement- structural support Articulation btw back of skull and vertebrae oriented to transfer forces linearly Articulation btw back of skull and vertebral column built to withstand sideways forces Vertebral column has tongue and groove articulations Spinal column is an S-shaped shock absorber BUT There is no ‘locking’ mechanism on skull to keep battering heads aligned Some Pachycephalosaurs have imprinted blood vessels on dome These factors suggests that head- butting may not be likely Intraspecies Competition (typically male-male) Females are typically choosey Why? Because they have more to loose Common rule in biology: Females are expensive to lose, males are cheap (e.g. deer hunting) Females choose the male most likely to provide the most successful offspring Males compete with each other for access to female vs. female chooses the strongest male Choosey females // Strong males have more offspring => SEXUAL selection Many ways to do this... But: In general, maximize competition and minimize accidental deaths (= no fitness) http://www.youtube.com/watch?v=PontCXFgs0M http://www.metacafe.com/watch/1941236/giraffe_fight/ http://www.youtube.com/watch? http://www.youtube.com/watch?v=DYDx1y38vGw http://www.youtube.com/watch?v=ULRtdk-3Yh4 Air-filled horn cores vs. solid bone skull caps... Gotta have a cheezy animated slide. Homalocephale Pachycephalosaurus Prenocephale Tylocephale Stegoceras Head butting Pachycephalosaurs Bone structure was probably strong enough to withstand collision Convex nature would favor glancing blows Instead, dome and spines seem better suited for “flank butting” So.. -
A New Maastrichtian Species of the Centrosaurine Ceratopsid Pachyrhinosaurus from the North Slope of Alaska
A new Maastrichtian species of the centrosaurine ceratopsid Pachyrhinosaurus from the North Slope of Alaska ANTHONY R. FIORILLO and RONALD S. TYKOSKI Fiorillo, A.R. and Tykoski, R.S. 2012. A new Maastrichtian species of the centrosaurine ceratopsid Pachyrhinosaurus from the North Slope of Alaska. Acta Palaeontologica Polonica 57 (3): 561–573. The Cretaceous rocks of the Prince Creek Formation contain the richest record of polar dinosaurs found anywhere in the world. Here we describe a new species of horned dinosaur, Pachyrhinosaurus perotorum that exhibits an apomorphic character in the frill, as well as a unique combination of other characters. Phylogenetic analysis of 16 taxa of ceratopsians failed to resolve relationships between P. perotorum and other Pachyrhinosaurus species (P. canadensis and P. lakustai). P. perotorum shares characters with each of the previously known species that are not present in the other, including very large nasal and supraorbital bosses that are nearly in contact and separated only by a narrow groove as in P. canadensis, and a rostral comb formed by the nasals and premaxillae as in P. lakustai. P. perotorum is the youngest centrosaurine known (70–69 Ma), and the locality that produced the taxon, the Kikak−Tegoseak Quarry, is close to the highest latitude for recovery of ceratopsid remains. Key words: Dinosauria, Centrosaurinae, Cretaceous, Prince Creek Formation, Kikak−Tegoseak Quarry, Arctic. Anthony R. Fiorillo [[email protected]] and Ronald S. Tykoski [[email protected]], Perot Museum of Nature and Science, 2201 N. Field Street, Dallas, TX 75202, USA. Received 4 April 2011, accepted 23 July 2011, available online 26 August 2011. -
Horn Use in Triceratops (Dinosauria: Ceratopsidae): Testing Behavioral Hypotheses Using Scale Models
Palaeontologia Electronica http://palaeo-electronica.org HORN USE IN TRICERATOPS (DINOSAURIA: CERATOPSIDAE): TESTING BEHAVIORAL HYPOTHESES USING SCALE MODELS Andrew A. Farke ABSTRACT Triceratops, a common chasmosaurine ceratopsid dinosaur from the Late Creta- ceous of North America, is known for its cranial ornamentation, including a single nasal horn and large, paired supraorbital horns. It is commonly surmised that Triceratops used its horns in intraspecific combat, but this hypothesis has not been rigorously tested. Scale models of Triceratops skulls were used to determine if it could physically lock horns as has been suggested. Three hypothetical horn locking positions were found, involving varying orientations of the combatants’ skulls. Based on these posi- tions, it was hypothesized that injuries caused by horns were especially likely in certain portions of the frill, jugals, and postorbital horncore tips. This corresponds to some pre- viously reported pathologies in chasmosaurine specimens. Uncertainties in this model- ing exercise center around variations in horn orientation, size, shape, and the possible existence of a keratinous supraorbital horncore sheath. Triceratops differs from modern horned mammals in its horn orientation, which suggests that if it engaged in intraspe- cific combat, its fighting style was quite different from these modern animals. During hypothetical horn locking in Triceratops, most of the force was directed against the medial and lateral surfaces of the horn cores. This has implications for future studies of ceratopsid cranial functional morphology, especially as related to horn architecture and the development of the frontal sinus complex. Andrew A. Farke. Department of Anatomical Sciences, Stony Brook University, Health Sciences Center, Stony Brook, New York, 11794-8081, USA. -
Bayanzag 1-Page
THE FLAMING CLIFFS MONGOLIA’S FAMOUS FOSSIL BEDS The Flaming Cliffs in the heart of the Gobi Desert are known worldwide for containing exceptional dinosaur fossils. The cliffs, the fossils, and the plants and animals who live nearby are protected by local government as part of Bayanzag Park. As you enjoy the park, please leave everything as you find it – for science, for the environment, and for Mongolia. WHO LIVED HERE A DELICATE LIVING ECOSYSTEM The Gobi Desert is home to animals and plants that live 80 MILLION YEARS AGO? nowhere else. Follow these rules to keep Bayanzag’s living The time was the Late Cretaceous: the last era when ecosystem from going extinct like the dinosaurs: dinosaurs ruled the earth. The Flaming Cliffs were sand dunes in a desert oasis, and many animals came here to hunt, Never damage plants or collect wood. forage, and raise their young. Meet three of them: Never feed or harm wildlife. Take your trash with you when you leave. Weighing two tons and covered with FOUND A FOSSIL? spiky armor, Pinacosaurus was a formidable plant-eater first discovered Leave it alone. Fossils lose scientific data at the Flaming Cliffs in 1923. and can easily be destroyed if moved. Plus, removing fossils without a permit is illegal. Protoceratops was the first dinosaur If it’s more than a small discovered in fragment, notify the local Mongolia, at officials of Umnugovi the Flaming Aimag. Cliffs in 1922. It ate plants and Also notify the ISMD at was about the size of MongolianDinosaurs.org. Take a sheep.