Darren H. Tanke Darren H. Tanke Langston, Jr. Wann Philip J. Currie, Philip J. Currie is a professor and Canada In the first monographic treatment of Research Chair at The University of Alberta Philip J. Currie, Wann Langston, Jr., & Darren H. Tanke a horned (ceratopsid) dinosaur in almost a (Department of Biological Sciences), is an Adjunct century, this monumental volume presents Professor at the University of Calgary, and was for- merly the Curator of Dinosaurs at the Royal Tyrrell one of the closest looks at the anatomy, re- Museum of Palaeontology. He took his B.Sc. at the lationships, growth and variation, behavior, University of Toronto in 1972, and his M.Sc. and ecology and other biological aspects of a sin- Ph.D. at McGill in 1975 and 1981. He is a Fellow of gle dinosaur species. The research, which was the Royal Society of Canada (1999) and a member A New Horned conducted over two decades, was possible of the Explorers Club (2001). He has published more because of the discovery of a densely packed than 100 scientific articles, 95 popular articles and bonebed near Grande Prairie, Alberta. The fourteen books, focussing on the growth and varia- tion of extinct reptiles, the anatomy and relationships Dinosaur From an locality has produced abundant remains of a of carnivorous dinosaurs, and the origin of birds. new species of horned dinosaur (ceratopsian), Fieldwork connected with his research has been con- and parts of at least 27 individual animals centrated in Alberta, Argentina, British Columbia, were recovered. China, Mongolia, the Arctic and Antarctica. His awards include the Sir Frederick Haultain Award for Upper Cretaceous Upper Cretaceous Bone Bed in Alberta A New Horned Dinosaur From an This new species of Pachyrhinosaurus significant contributions to science in Alberta (1988), the American Association of Petroleum Geologists is closely related to Pachyrhinosaurus Michel T. Halbouty Human Needs Award (1999), the canadensis, which is known from younger Michael Smith Award (2004) and the ASTech (Alberta Bone Bed in Alberta rocks near Drumheller and Lethbridge in Science and Technology Leadership) Award for out- southern Alberta, but is a smaller animal standing leadership in Alberta Science (2006). He has with many differences in the ornamental given hundreds of popular and scientific lectures on spikes and bumps on the skull. The adults of dinosaurs all over the world, and is often interviewed by the press. both species have massive bosses of bone in the positions where other horned dinosaurs Wann Langston is a professor emeritus in the (like Centrosaurus and Triceratops) have Jackson School of Geosciences at the University of horns. However, juveniles of the new species Texas at Austin. His interest in dinosaurs was kindled resemble juveniles of Centrosaurus in hav- early by visits to museums and by the books of Roy ing horns rather than bosses. Skull anatomy Chapman Andrews. Born in Oklahoma, he was edu- cated at the University of Oklahoma and the University undergoes remarkable changes during growth of California at Berkeley, receiving his Ph.D. there in and the horns over the nose and eyes of the 1952. In 1954, he succeeded The Grand Old Man of Pachyrhinosaurus juveniles transform into Canadian Dinosaurology, Charles M. Sternberg, as bosses; spikes and horns develop on the top curator at Ottawa’s Canadian Museum of Nature. In of and at the back of the frill that extends Canada, Langston’s interests focused on the dinosaurs back over the neck. No cause has been de- of Alberta and Saskatchewan, and Permian vertebrates termined for the apparent catastrophic death of Prince Edward Island. Among his discoveries was a Pachyrhinosaurus bone bed in southern Alberta, which of the herd of Pachyrhinosaurus from the yielded several technical studies over ensuing years. Grande Prairie area, but it has been suggested Retired since 1986, he remains active in re- that such herds may have been migratory search at the Texas Memorial Museum’s Vertebrate animals. Paleontology Laboratory in Austin. In addition to the main descriptive paper, Darren H. Tanke has been a senior fossil prepa- ration technician at the Royal Tyrrell Museum of the volume includes information on the dis- Palaeontology in Drumheller, Alberta, since 2005. tribution of bones within the bonebed itself, From 1979 to 2005, Tanke worked with Philip J. and a cutting-edge digital treatment of CT- Currie at the Provincial Museum of Alberta and the scan data of the fossils to reveal the anatomy Royal Tyrrell Museum of Palaeontology. Tanke has of the animal’s brain! authored or co-authored papers on various aspects of dinosaurs, such as ontogeny; dinosaur paleopathology; relocation of “lost” dinosaur quarries and identifica- tion of “mystery quarries”; and especially various human history aspects of Alberta’s paleontological legacy. He was senior editor of Mesozoic Vertebrate Life: New Research Inspired by the Paleontology of Philip J. Currie (co-published by Indiana University Press, Bloomington, and NRC Research Press, Ottawa, 2001), and has appeared in the 1998 documentary film Dinosaur Park, and the 1993 educational film on the Pipestone Creek Pachyrhinosaurus bone bed Messages in Stone. © 2008 National Research Council of Canada cover art copyright 2008, ISBN 978–0–660–19820–0 Michael W. Skrepnick NRC No. 49729 http://pubs.nrc-cnrc.gc.ca Downloaded from http://pubs.geoscienceworld.org/books/book/chapter-pdf/3731473/9780660198200_frontmatter.pdf by guest on 02 October 2021 C M Y K 1 A $jobid A Publication of the National Research Council of Canada Monograph Publishing Program Philip J. Currie, Wann Langston, Jr., & Darren H. Tanke A New Horned Dinosaur From an Upper Cretaceous Bone Bed in Alberta with contributions from Patricia E. Ralrick, Ryan C. Ridgely, and Lawrence M. Witmer Downloaded from http://pubs.geoscienceworld.org/books/book/chapter-pdf/3731473/9780660198200_frontmatter.pdf by guest on 02 October 2021 ©2008 National Research Council of Canada All rights reserved. No part of this publication may be reproduced in a retrieval system, or transmitted by any means, electronic, mechanical, photocopying, recording or otherwise, without the prior written permission of the National Research Council of Canada, Ottawa, ON K1A 0R6, Canada. Printed on acid-free paper. ISBN 978-0-660-19819-4 NRC No. 49729 cover art copyright 2008 Michael W. Skrepnick Library and Archives Canada Cataloguing in Publication Currie, Philip J., 1949- A new horned dinosaur from an Upper Cretaceous bone bed in Alberta / Philip John Currie, Wann Langston, Jr., Darren H. Tanke. Issued by: National Research Council Canada. Includes bibliographical references. ISBN 978-0-660-19819-4 1. Pachyrhinosaurus. 2. Paleontology—Alberta. 3. Dinosaurs—Alberta. I. Langston, Wann, 1921- II. Tanke, Darren H III. National Research Council Canada IV. Title. QE862.O65C87 2008 567.915 C2008-980235-7 The Publisher wishes to thank the County of Grande Prairie, AB with a special thanks to Walter Paszkowski for their financial contribution to this book. NRC Monograph Publishing Program Editor: P.B. Cavers (University of Western Ontario) Editorial Board: W.G.E. Caldwell, OC, FRSC (University of Western Ontario); M.E. Cannon, FCAE, FRSC (University of Calgary); K.G. Davey, OC, FRSC (York University); M.M. Ferguson (University of Guelph); S. Gubins (Annual Reviews); B.K. Hall, FRSC (Dalhousie University); W.H. Lewis (Washington University); A.W. May, OC (Memorial University of Newfoundland); B.P. Dancik, Editor-in-Chief, NRC Research Press (University of Alberta) Inquiries: Monograph Publishing Program, NRC Research Press, National Research Council of Canada, Ottawa, Ontario K1A 0R6, Canada. Web site: http://pubs.nrc-cnrc.gc.ca Correct citation for this publication: Currie, P.J., Langston, W., Jr., and Tanke, D.H. 2008. A New Horned Dinosaur from an Upper Cretaceous Bone Bed in Alberta. NRC Research Press, Ottawa, Ontario, Canada. 144p. Downloaded from http://pubs.geoscienceworld.org/books/book/chapter-pdf/3731473/9780660198200_frontmatter.pdf by guest on 02 October 2021 CONTENTS Contributors iv Foreword: SCOTT D. SAMPSON v 1. A new species of Pachyrhinosaurus (Dinosauria, Ceratopsidae) from the Upper Cretaceous of Alberta, Canada • PHILI P J. CU rr I E , WA NN LA NG S T O N , JR., A ND Darre N H. TA NK E 1 2. Comments on the quarry map and preliminary taphonomic observations of the Pachyrhinosaurus (Dinosauria: Ceratopsidae)bone bed at Pipestone Creek, Alberta, Canada • PA T R I C I A E. RA L R I C K A ND Darre N H. TA NK E 109 3. Structure of the brain cavity and inner ear of the centrosaurine ceratopsid dinosaur Pachyrhinosaurus based on CT scanning and 3D visualization Lawre N ce M. WIT mer A ND RY A N C. RIDG E LY 117 Contents • iii Downloaded from http://pubs.geoscienceworld.org/books/book/chapter-pdf/3731473/9780660198200_frontmatter.pdf by guest on 02 October 2021 Contributors Philip J. Currie, University of Alberta, Department of Biological Sci- ences, CW405 Biological Sciences Building, Edmonton, AB T6G 2E9, Canada (e-mail: [email protected]). Wann Langston, Jr., University of Texas at Austin, J.J. Pickle Research Campus, Vertebrate Paleontology Laboratory #6, 10100 Burnet Road, Austin, TX 78758, USA (e-mail: [email protected]). Patricia E. Ralrick, Interdisciplinary Graduate Program, University of Calgary, 2500 University Drive NW, Calgary, AB T2N 1N4, Canada (e-mail: [email protected]). Ryan C. Ridgely, Department of Biomedical Sciences, Ohio University College of Osteopathic Medicine, Athens, OH 45701, USA (e-mail: [email protected]). Darren H. Tanke, Royal Tyrrell Museum of Palaeontology, Box 7500, Drumheller, AB T0J 0Y0, Canada (e-mail: [email protected]). Lawrence M. Witmer, Department of Biomedical Sciences, Ohio University College of Osteopathic Medicine, Athens, OH 45701, USA (email: [email protected]). iv • Contributors Downloaded from http://pubs.geoscienceworld.org/books/book/chapter-pdf/3731473/9780660198200_frontmatter.pdf by guest on 02 October 2021 Foreword I am unlikely to forget my first encounters with Pachyrhinosaurus lakustai.