Raptors in Action 1 Suggested Pre-Visit Activities
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PALEONTOLOGY and BIOSTRATIGRAPHY of MONGOLIA the JOINT SOVIET-MONGOLIAN PALEONTOLOGICAL EXPEDITION (Transactions, Vol
PALEONTOLOGY AND BIOSTRATIGRAPHY OF MONGOLIA THE JOINT SOVIET-MONGOLIAN PALEONTOLOGICAL EXPEDITION (Transactions, vol. 3) EDITORIAL BOARD: N. N. Kramarenko (editor-in-chief) B. Luvsandansan, Yu. I. Voronin, R. Barsbold, A. K. Rozhdestvensky, B. A. Trofimov, V. Yu. Reshetov 1976 S. M. Kurzanov A NEW CARNOSAUR FROM THE LATE CRETACEOUS OF NOGON-TSAV, MONGOLIA* The Upper Cretaceous locality of Nogon-Tsav, located in the Zaltaika Gobi, 20 km NW of Mt. Ongon-Ulan-Ula, was discovered by geologists of the MNR. Then, in a series of years (1969–1971, 1973) the joint Soviet-Mongolian Geological Expedition investigated the site. According to the oral report of B. Yu. Reshetova, the deposits in this region are composed basically of sandstone and clay, clearly subdivided into two deposits. The lower great thickness contains many complete skulls, separate bones of tarbosaurs, ornithomimids, and claws of therizinosaurs. The upper red-colored beds are characterized only by scattered fragments of skulls. The fragments described below are of a new carnivorous dinosaur, Alioramus remotus (coll. South Goby region), from the family Tyrannosauridae, which occurs in the lower gray beds. Genus Alioramus Kurzanov, gen. nov. G e n u s n a m e from alius (Lat.) – other, ramus (Lat.) – branch; masc. G e n o t y p e — A. remotus Kurzanov, sp. nov., Upper Cretaceous, Nemegetinskaya suite, southern Mongolia. D i a g n o s i s . Tyrannosaurid of average size. Skull low, with highly elongated jaws. Nasals with well-developed separate crests of 1–l.5 cm height. Second antorbital fenestrae displaced forward, reaching the anterior end of the 5th tooth. -
Theropod Teeth from the Upper Maastrichtian Hell Creek Formation “Sue” Quarry: New Morphotypes and Faunal Comparisons
Theropod teeth from the upper Maastrichtian Hell Creek Formation “Sue” Quarry: New morphotypes and faunal comparisons TERRY A. GATES, LINDSAY E. ZANNO, and PETER J. MAKOVICKY Gates, T.A., Zanno, L.E., and Makovicky, P.J. 2015. Theropod teeth from the upper Maastrichtian Hell Creek Formation “Sue” Quarry: New morphotypes and faunal comparisons. Acta Palaeontologica Polonica 60 (1): 131–139. Isolated teeth from vertebrate microfossil localities often provide unique information on the biodiversity of ancient ecosystems that might otherwise remain unrecognized. Microfossil sampling is a particularly valuable tool for doc- umenting taxa that are poorly represented in macrofossil surveys due to small body size, fragile skeletal structure, or relatively low ecosystem abundance. Because biodiversity patterns in the late Maastrichtian of North American are the primary data for a broad array of studies regarding non-avian dinosaur extinction in the terminal Cretaceous, intensive sampling on multiple scales is critical to understanding the nature of this event. We address theropod biodiversity in the Maastrichtian by examining teeth collected from the Hell Creek Formation locality that yielded FMNH PR 2081 (the Tyrannosaurus rex specimen “Sue”). Eight morphotypes (three previously undocumented) are identified in the sample, representing Tyrannosauridae, Dromaeosauridae, Troodontidae, and Avialae. Noticeably absent are teeth attributed to the morphotypes Richardoestesia and Paronychodon. Morphometric comparison to dromaeosaurid teeth from multiple Hell Creek and Lance formations microsites reveals two unique dromaeosaurid morphotypes bearing finer distal denticles than present on teeth of similar size, and also differences in crown shape in at least one of these. These findings suggest more dromaeosaurid taxa, and a higher Maastrichtian biodiversity, than previously appreciated. -
By Howard Zimmerman
by Howard Zimmerman DINO_COVERS.indd 4 4/24/08 11:58:35 AM [Intentionally Left Blank] by Howard Zimmerman Consultant: Luis M. Chiappe, Ph.D. Director of the Dinosaur Institute Natural History Museum of Los Angeles County 1629_ArmoredandDangerous_FNL.ind1 1 4/11/08 11:11:17 AM Credits Title Page, © Luis Rey; TOC, © De Agostini Picture Library/Getty Images; 4-5, © John Bindon; 6, © De Agostini Picture Library/The Natural History Museum, London; 7, © Luis Rey; 8, © Luis Rey; 9, © Adam Stuart Smith; 10T, © Luis Rey; 10B, © Colin Keates/Dorling Kindersly; 11, © Phil Wilson; 12L, Courtesy of the Royal Tyrrell Museum, Drumheller, Alberta; 12R, © De Agostini Picture Library/Getty Images; 13, © Phil Wilson; 14-15, © Phil Wilson; 16-17, © De Agostini Picture Library/The Natural History Museum, London; 18T, © 2007 by Karen Carr and Karen Carr Studio; 18B, © photomandan/istockphoto; 19, © Luis Rey; 20, © De Agostini Picture Library/The Natural History Museum, London; 21, © John Bindon; 23TL, © Phil Wilson; 23TR, © Luis Rey; 23BL, © Vladimir Sazonov/Shutterstock; 23BR, © Luis Rey. Publisher: Kenn Goin Editorial Director: Adam Siegel Creative Director: Spencer Brinker Design: Dawn Beard Creative Cover Illustration: Luis Rey Photo Researcher: Omni-Photo Communications, Inc. Library of Congress Cataloging-in-Publication Data Zimmerman, Howard. Armored and dangerous / by Howard Zimmerman. p. cm. — (Dino times trivia) Includes bibliographical references and index. ISBN-13: 978-1-59716-712-3 (library binding) ISBN-10: 1-59716-712-6 (library binding) 1. Ornithischia—Juvenile literature. 2. Dinosaurs—Juvenile literature. I. Title. QE862.O65Z56 2009 567.915—dc22 2008006171 Copyright © 2009 Bearport Publishing Company, Inc. All rights reserved. -
The Princeton Field Guide to Dinosaurs, Second Edition
MASS ESTIMATES - DINOSAURS ETC (largely based on models) taxon k model femur length* model volume ml x specific gravity = model mass g specimen (modeled 1st):kilograms:femur(or other long bone length)usually in decameters kg = femur(or other long bone)length(usually in decameters)3 x k k = model volume in ml x specific gravity(usually for whole model) then divided/model femur(or other long bone)length3 (in most models femur in decameters is 0.5253 = 0.145) In sauropods the neck is assigned a distinct specific gravity; in dinosaurs with large feathers their mass is added separately; in dinosaurs with flight ablity the mass of the fight muscles is calculated separately as a range of possiblities SAUROPODS k femur trunk neck tail total neck x 0.6 rest x0.9 & legs & head super titanosaur femur:~55000-60000:~25:00 Argentinosaurus ~4 PVPH-1:~55000:~24.00 Futalognkosaurus ~3.5-4 MUCPv-323:~25000:19.80 (note:downsize correction since 2nd edition) Dreadnoughtus ~3.8 “ ~520 ~75 50 ~645 0.45+.513=.558 MPM-PV 1156:~26000:19.10 Giraffatitan 3.45 .525 480 75 25 580 .045+.455=.500 HMN MB.R.2181:31500(neck 2800):~20.90 “XV2”:~45000:~23.50 Brachiosaurus ~4.15 " ~590 ~75 ~25 ~700 " +.554=~.600 FMNH P25107:~35000:20.30 Europasaurus ~3.2 “ ~465 ~39 ~23 ~527 .023+.440=~.463 composite:~760:~6.20 Camarasaurus 4.0 " 542 51 55 648 .041+.537=.578 CMNH 11393:14200(neck 1000):15.25 AMNH 5761:~23000:18.00 juv 3.5 " 486 40 55 581 .024+.487=.511 CMNH 11338:640:5.67 Chuanjiesaurus ~4.1 “ ~550 ~105 ~38 ~693 .063+.530=.593 Lfch 1001:~10700:13.75 2 M. -
At Carowinds
at Carowinds EDUCATOR’S GUIDE CLASSROOM LESSON PLANS & FIELD TRIP ACTIVITIES Table of Contents at Carowinds Introduction The Field Trip ................................... 2 The Educator’s Guide ....................... 3 Field Trip Activity .................................. 4 Lesson Plans Lesson 1: Form and Function ........... 6 Lesson 2: Dinosaur Detectives ....... 10 Lesson 3: Mesozoic Math .............. 14 Lesson 4: Fossil Stories.................. 22 Games & Puzzles Crossword Puzzles ......................... 29 Logic Puzzles ................................. 32 Word Searches ............................... 37 Answer Keys ...................................... 39 Additional Resources © 2012 Dinosaurs Unearthed Recommended Reading ................. 44 All rights reserved. Except for educational fair use, no portion of this guide may be reproduced, stored in a retrieval system, or transmitted in any form or by any Dinosaur Data ................................ 45 means—electronic, mechanical, photocopy, recording, or any other without Discovering Dinosaurs .................... 52 explicit prior permission from Dinosaurs Unearthed. Multiple copies may only be made by or for the teacher for class use. Glossary .............................................. 54 Content co-created by TurnKey Education, Inc. and Dinosaurs Unearthed, 2012 Standards www.turnkeyeducation.net www.dinosaursunearthed.com Curriculum Standards .................... 59 Introduction The Field Trip From the time of the first exhibition unveiled in 1854 at the Crystal -
Brains and Intelligence
BRAINS AND INTELLIGENCE The EQ or Encephalization Quotient is a simple way of measuring an animal's intelligence. EQ is the ratio of the brain weight of the animal to the brain weight of a "typical" animal of the same body weight. Assuming that smarter animals have larger brains to body ratios than less intelligent ones, this helps determine the relative intelligence of extinct animals. In general, warm-blooded animals (like mammals) have a higher EQ than cold-blooded ones (like reptiles and fish). Birds and mammals have brains that are about 10 times bigger than those of bony fish, amphibians, and reptiles of the same body size. The Least Intelligent Dinosaurs: The primitive dinosaurs belonging to the group sauropodomorpha (which included Massospondylus, Riojasaurus, and others) were among the least intelligent of the dinosaurs, with an EQ of about 0.05 (Hopson, 1980). Smartest Dinosaurs: The Troodontids (like Troödon) were probably the smartest dinosaurs, followed by the dromaeosaurid dinosaurs (the "raptors," which included Dromeosaurus, Velociraptor, Deinonychus, and others) had the highest EQ among the dinosaurs, about 5.8 (Hopson, 1980). The Encephalization Quotient was developed by the psychologist Harry J. Jerison in the 1970's. J. A. Hopson (a paleontologist from the University of Chicago) did further development of the EQ concept using brain casts of many dinosaurs. Hopson found that theropods (especially Troodontids) had higher EQ's than plant-eating dinosaurs. The lowest EQ's belonged to sauropods, ankylosaurs, and stegosaurids. A SECOND BRAIN? It used to be thought that the large sauropods (like Brachiosaurus and Apatosaurus) and the ornithischian Stegosaurus had a second brain. -
A New Caenagnathid Dinosaur from the Upper Cretaceous Wangshi
www.nature.com/scientificreports OPEN A new caenagnathid dinosaur from the Upper Cretaceous Wangshi Group of Shandong, China, with Received: 12 October 2017 Accepted: 7 March 2018 comments on size variation among Published: xx xx xxxx oviraptorosaurs Yilun Yu1, Kebai Wang2, Shuqing Chen2, Corwin Sullivan3,4, Shuo Wang 5,6, Peiye Wang2 & Xing Xu7 The bone-beds of the Upper Cretaceous Wangshi Group in Zhucheng, Shandong, China are rich in fossil remains of the gigantic hadrosaurid Shantungosaurus. Here we report a new oviraptorosaur, Anomalipes zhaoi gen. et sp. nov., based on a recently collected specimen comprising a partial left hindlimb from the Kugou Locality in Zhucheng. This specimen’s systematic position was assessed by three numerical cladistic analyses based on recently published theropod phylogenetic datasets, with the inclusion of several new characters. Anomalipes zhaoi difers from other known caenagnathids in having a unique combination of features: femoral head anteroposteriorly narrow and with signifcant posterior orientation; accessory trochanter low and confuent with lesser trochanter; lateral ridge present on femoral lateral surface; weak fourth trochanter present; metatarsal III with triangular proximal articular surface, prominent anterior fange near proximal end, highly asymmetrical hemicondyles, and longitudinal groove on distal articular surface; and ungual of pedal digit II with lateral collateral groove deeper and more dorsally located than medial groove. The holotype of Anomalipes zhaoi is smaller than is typical for Caenagnathidae but larger than is typical for the other major oviraptorosaurian subclade, Oviraptoridae. Size comparisons among oviraptorisaurians show that the Caenagnathidae vary much more widely in size than the Oviraptoridae. Oviraptorosauria is a clade of maniraptoran theropod dinosaurs characterized by a short, high skull, long neck and short tail. -
Implications for Predatory Dinosaur Macroecology and Ontogeny in Later Late Cretaceous Asiamerica
Canadian Journal of Earth Sciences Theropod Guild Structure and the Tyrannosaurid Niche Assimilation Hypothesis: Implications for Predatory Dinosaur Macroecology and Ontogeny in later Late Cretaceous Asiamerica Journal: Canadian Journal of Earth Sciences Manuscript ID cjes-2020-0174.R1 Manuscript Type: Article Date Submitted by the 04-Jan-2021 Author: Complete List of Authors: Holtz, Thomas; University of Maryland at College Park, Department of Geology; NationalDraft Museum of Natural History, Department of Geology Keyword: Dinosaur, Ontogeny, Theropod, Paleocology, Mesozoic, Tyrannosauridae Is the invited manuscript for consideration in a Special Tribute to Dale Russell Issue? : © The Author(s) or their Institution(s) Page 1 of 91 Canadian Journal of Earth Sciences 1 Theropod Guild Structure and the Tyrannosaurid Niche Assimilation Hypothesis: 2 Implications for Predatory Dinosaur Macroecology and Ontogeny in later Late Cretaceous 3 Asiamerica 4 5 6 Thomas R. Holtz, Jr. 7 8 Department of Geology, University of Maryland, College Park, MD 20742 USA 9 Department of Paleobiology, National Museum of Natural History, Washington, DC 20013 USA 10 Email address: [email protected] 11 ORCID: 0000-0002-2906-4900 Draft 12 13 Thomas R. Holtz, Jr. 14 Department of Geology 15 8000 Regents Drive 16 University of Maryland 17 College Park, MD 20742 18 USA 19 Phone: 1-301-405-4084 20 Fax: 1-301-314-9661 21 Email address: [email protected] 22 23 1 © The Author(s) or their Institution(s) Canadian Journal of Earth Sciences Page 2 of 91 24 ABSTRACT 25 Well-sampled dinosaur communities from the Jurassic through the early Late Cretaceous show 26 greater taxonomic diversity among larger (>50kg) theropod taxa than communities of the 27 Campano-Maastrichtian, particularly to those of eastern/central Asia and Laramidia. -
A Revision of the Ceratopsia Or Horned Dinosaurs
MEMOIRS OF THE PEABODY MUSEUM OF NATURAL HISTORY VOLUME III, 1 A.R1 A REVISION orf tneth< CERATOPSIA OR HORNED DINOSAURS BY RICHARD SWANN LULL STERLING PROFESSOR OF PALEONTOLOGY AND DIRECTOR OF PEABODY MUSEUM, YALE UNIVERSITY LVXET NEW HAVEN, CONN. *933 MEMOIRS OF THE PEABODY MUSEUM OF NATURAL HISTORY YALE UNIVERSITY Volume I. Odontornithes: A Monograph on the Extinct Toothed Birds of North America. By Othniel Charles Marsh. Pp. i-ix, 1-201, pis. 1-34, text figs. 1-40. 1880. To be obtained from the Peabody Museum. Price $3. Volume II. Part 1. Brachiospongidae : A Memoir on a Group of Silurian Sponges. By Charles Emerson Beecher. Pp. 1-28, pis. 1-6, text figs. 1-4. 1889. To be obtained from the Peabody Museum. Price $1. Volume III. Part 1. American Mesozoic Mammalia. By George Gaylord Simp- son. Pp. i-xvi, 1-171, pis. 1-32, text figs. 1-62. 1929. To be obtained from the Yale University Press, New Haven, Conn. Price $5. Part 2. A Remarkable Ground Sloth. By Richard Swann Lull. Pp. i-x, 1-20, pis. 1-9, text figs. 1-3. 1929. To be obtained from the Yale University Press, New Haven, Conn. Price $1. Part 3. A Revision of the Ceratopsia or Horned Dinosaurs. By Richard Swann Lull. Pp. i-xii, 1-175, pis. I-XVII, text figs. 1-42. 1933. To be obtained from the Peabody Museum. Price $5 (bound in cloth), $4 (bound in paper). Part 4. The Merycoidodontidae, an Extinct Group of Ruminant Mammals. By Malcolm Rutherford Thorpe. In preparation. -
The Systematic Position of the Enigmatic Thyreophoran Dinosaur Paranthodon Africanus, and the Use of Basal Exemplifiers in Phyl
1 The systematic position of the enigmatic thyreophoran dinosaur Paranthodon africanus, 2 and the use of basal exemplifiers in phylogenetic analysis 3 4 Thomas J. Raven1,2 ,3 and Susannah C. R. Maidment2 ,3 5 61Department of Earth Science & Engineering, Imperial College London, UK 72School of Environment & Technology, University of Brighton, UK 8 3Department of Earth Sciences, Natural History Museum, London, UK 9 10Corresponding author: Thomas J. Raven 11 12Email address: [email protected] 13 14 15 16 17 18 19 20 21ABSTRACT 22 23The first African dinosaur to be discovered, Paranthodon africanus was found in 1845 in the 24Lower Cretaceous of South Africa. Taxonomically assigned to numerous groups since discovery, 25in 1981 it was described as a stegosaur, a group of armoured ornithischian dinosaurs 26characterised by bizarre plates and spines extending from the neck to the tail. This assignment 27that has been subsequently accepted. The type material consists of a premaxilla, maxilla, a nasal, 28and a vertebra, and contains no synapomorphies of Stegosauria. Several features of the maxilla 29and dentition are reminiscent of Ankylosauria, the sister-taxon to Stegosauria, and the premaxilla 30appears superficially similar to that of some ornithopods. The vertebral material has never been 31described, and since the last description of the specimen, there have been numerous discoveries 32of thyreophoran material potentially pertinent to establishing the taxonomic assignment of the 33specimen. An investigation of the taxonomic and systematic position of Paranthodon is therefore 34warranted. This study provides a detailed re-description, including the first description of the 35vertebra. Numerous phylogenetic analyses demonstrate that the systematic position of 36Paranthodon is highly labile and subject to change depending on which exemplifier for the clade 37Stegosauria is used. -
Ten Little Dinosaurs Free
FREE TEN LITTLE DINOSAURS PDF Mike Brownlow,Simon Rickerty | 32 pages | 01 Oct 2015 | Hachette Children's Group | 9781408334010 | English | London, United Kingdom The First Dinosaurs A period of great transformation, Ten Little Dinosaurs Late Cretaceous Period is when the dinosaurs disappeared from the earth. Learn more about the Late Cretaceous dinosaurs that existed during this era, such as the Ten Little Dinosaurs, Gallimimus, and Brachylophosaurus. Tyrannosaurus rex, one of the fiercest meat-eaters ever, is the animal that probably springs to mind when most of us hear the word "dinosaur. It shared the Cretaceous landscape Ten Little Dinosaurs, and probably was preyed upon by, Tyrannosaurus rex. Constantly compared to the Tyrannosaurus rex, the Giganotosaurus was one of a handful of dinosaurs that rivaled, or possibly exceeded, the creature in size. University of Kansas paleontologists Ten Little Dinosaurs comparing the bones of a new T. Could this exciting find help bridge the gaps Ten Little Dinosaurs Africa's late Cretaceous fossil record and that of other continents? OK, hop in your time machine and go back 67 million years or so to the Cretaceous period. Then find a Tyrannosaurus rex and challenge it to a race. Sounds crazy, huh? Could you really outrun a Tyrannosaurus rex? Learn about Monoclonius, Late Cretaceous dinosaurs and dinosaurs Ten Little Dinosaurs all eras. The recently discovered large theropod Abelisaurus comahuensis, from Patagonia is argentina, looked a little like Albertosaurus from Alberta, Canada, particularly in its size and lifestyle. Find out more about the Late Cretaceous dinosaurs. Albertosaurus was an older "cousin" to the better-known Tyrannosaurus. -
Immigrant Species, Or Native Species?
The Journal of Paleontological Sciences: JPS.C.2017.01 TESTING THE HYPOTHESES OF THE ORIGIN OF TYRANNOSAURUS REX: IMMIGRANT SPECIES, OR NATIVE SPECIES? __________________________________________________________________________________________________________________ Chan-gyu Yun Vertebrate Paleontological Institute of Incheon, Incheon 21974, Republic of Korea & Biological Sciences, Inha University, Incheon 22212, Republic of Korea [email protected] __________________________________________________________________________________________________________________ Abstract: It is an undoubtable fact that Tyrannosaurus rex is the most iconic dinosaur species of all time. However, it is currently debatable whether this species has a North American origin or Asian origin. In this paper, I test these two hypotheses based on current fossil records and former phylogenetic analyses. Phylogenetic and fossil evidence, such as derived tyrannosaurine fossils of Asia, suggests that the hypothesis of an Asian origin of Tyrannosaurus rex is the most plausible one, but this is yet to be certain due to the scarcity of fossil records. INTRODUCTION The most famous and iconic dinosaur of all time, Tyrannosaurus rex, is only known from upper Maastrichtian geological formations in Western North America (e.g. Carr and Williamson, 2004; Larson, 2008). However, older relatives of Tyrannosaurus rex (e.g. Daspletosaurus, Tarbosaurus) are known from both Asia and North America. This leads to an evolutionary question: is the origin of Tyrannosaurus rex from Asia, or North America? About six of the currently valid tyrannosaurine taxa were described in the twenty-first century (based on parsimony analysis of Brusatte and Carr, 2016), with new species which are being described (Sebastian Dalman, Pers. Comm., 2016; Thomas Carr, Pers. Comm., 2016). It can be said that "now" is the "golden age” for studying tyrannosaurine evolution.