植物研究雑誌 J. J. Jpn. Bo t. Originals Originals 75: 1-4 1 (2000)

A Systematic Enumeration of J apanese Salix ()

Hiroyoshi Hiroyoshi OHASHI

Biological Biological Institute ，Graduate School of Science ，Tohoku University ，Sendai ，980-8578 JAPAN (Received (Received on August 2， 1999)

A systematic enumeration of the genus Salix of Japan is provided. Chosenia and Toisusu Toisusu are included in Salix. About 40 have been recognized as distinct in recent recent floras of Japan ，bu ，( they were reduced to 31 species in this work. Of these ， 28 28 are recognized as nativetOJapan and three are supposedly imported from China or or Korea. These are classified into four subgenera and 17 sections. Subgenus Urbanianae Urbanianae (Seemen) H.Ohashi is newly proposed. Thirty-seven natural hybrids are recognized. recognized. Four subspecies ， one variety ， ten forms and one nothosubspecies are newly newly proposed. They are indicated in a systematic list in this pape 工 Each taxon is enumerated enumerated with its original reference ，selected bibliography and principal synonyms. Types Types of specific and infraspecific names are quoted after the original reference. Areas Areas of distribution are shown for species and infraspecific taxa.

Key words: Bibliography ，enumeration ，J apan ，Salix ，

Introduction Introduction 1979; Kimura 1989 and Ohwi and Kitagawa Salix Salix comprises some 400 (Newsholme 1992) ， but they were reduced to 31 species 1992 ，Ohashi 1995) or 450 species (Argus with 37 hybrids in this work. These are 1997). 1997). Within the genus ，Chosenia and/or newly classified into four subgenera and 17 Toisusu Toisusu are often treated as distinc t. In this sections. A new subgenus ，Urbanianae paper ， the broadest concept of Salix is (Seemen) H.Ohashi ，is proposed for accom- adopted adopted on the basis of mo ゅhological stud- modating Chosenia ， Toisusu and Salix sub- ies ies of the three genera (Kimura 1988) and genus Protitea in the system of Kimura m01ecular m01ecular analysis of Salicaceae (Leskinen (1928 ，1938 ，1989) and Kitamura and and Alstrom-Rapaport 1999 ，Azuma et al. in 乱1urata (1 979); Chosenia and Salix sections press). press). Glandulosae and Urbanianae in Nakai Salix Salix is distributed mostly in cold and (1930); Chosenia and Salix sections temperate temperate regions in Northern Hemisphere Tetraspermae and Wilsonianae in Chou and a few species extending to South (1 984); Chosenia and Salix sections Flori- America and Africa. The species are rich in danae and Humboldianae in Argus (1 997); East East Asia ，especially in China. According to or subgenera Pleuradenia and Protitea in Argus (1997) ，there 訂 e about 270 species in Kimura (1 988). The subgenus contains all China ，120 in the former Soviet Union ， 103 species having the bud scale with a free and in in North America and 65 in Europe. imbricate margin and short or absent necta- About 40 species with 30 hybrids are rec- ries. Subgen. Urbanianae is the most primi- ognized in Japan in recent works (Ohwi tive group of Salix. 1953 ，1965a & b; Kitamura and Murata Salix does not occur naturally in the 2 植物研究雑誌第75 巻第1号 平成12 年2月

Ryukyu Islands (Hatusima 1971 ，Shimabuku hardly comparable with those of Skvortsov 1997) ， although Walker (1976) recorded (1968a ， 1968b) or Argus (1 986 ， 1997). Salix Salix integra from Okinawa Island as a na- Many hybrids have been recorded mainly tive. tive. Kagoshima Prefecture is generally re- by Ki mura in Japanese Salix ， but these have garded garded to be the southernmost boundary of not been treated systematically. Only names Salix Salix in Japan for its natural distribution. were cited in Floras of Japan (Ohwi 1953 ， Among seven species and two hybrids re- 1965a ，1965b ， Kitamura and Murata 1979 ， corded corded in this Prefecture ， Salix eriocarpa Ohwi and Kitagawa 1992). Any citations of known in Yakushima ，Tanegashima and references for hybrids of the J apanese Salix Amami-oshima (culti vated?) and S. in the Floras have not been made. sieboldiana sieboldiana in Tanegashima (Hatusima The importance of citation of taxonomic 1978) 1978) grow in the southernmost places in references was emphasized in my previous Japan. Japan. paper on genera of J apanese Leguminosae Since Since the comprehensive taxonomic work (Ohashi 1999). Generally ，1 believe this is by Seemen in 1903 ，a number of studies true for every taxon in biology. Citation of have have been made on the Japanese Salix. Es- selected taxonomic references for a given pecially ，Koidzumi (1 913) ，Kimura (1 926 ， taxon is important for co 町 ect understanding ， 1934 ，1981 ，1988 ，etc.) ， Nakai (1 930) ，and because present-day knowledge of a taxon is Ki tagawa (1939 ， 1979) made great contribu- constructed mostly by the accumulation of tions tions to the genus of Japan ，Korea ，Manchu- knowledge since it was first described. Usu- ria ， Kuril Islands and Sakhalin. Recent ally ，taxa 訂 e not fully recognized when they works on Japanese Salix have treated the are initially circumscribed; charac- various taxa taxa occurring in present-day Japan (Ohwi teristics are added by later students of the 1953 ，1965a ，1965b ， Kitamura and Murata group. Additions or revisions are made not 1979 ，Kimura 1989 ，Ohwi and Kitagawa only in gross morphological characters but 1992); 1992); Chosenia and Toisusu are recognized also in various cryptic characters such as as as distinct. These works ，however ， lack any chromosomes ，pollen ， chemical components ， citation citation of taxonomic references or bibliog- DNA sequences ，etc. Naturally ，moreover ， raphy raphy for the taxa ，detailed synonymy and discovery of future new characteristics are citation citation of the types of the names. to be expected for a given taxon. Accumu- Infrageneric Infrageneric system for the J apanese Salix lation of characteristics is necessary for full is is incomplete. Seemen (1903) and Schneider recognition of a taxon ， and citation of taxo- (1916) (1916) founded system for J apanese Sα lix. nomic references contributes important in- Recent Recent systematic works and nomenclature formation to its true understanding. of of Sα lix are greatly advanced by Skvortsov 1 believe that citation of synonyms for a (1 968a ， 1968b) and Argus (1986 ， 1997). correct name has a role similar in impor- Kimura (1 928 ，1938 ， 1988) proposed a sys- tance to the citation of selected taxonomic tem of Salicaceae ， but did only subgeneric references. In this paper each synonym is system system for Salix without any references to listed with its references and the type under works of Skvortsov (1968a ， 1968b) and the correct name. Argus Argus (1986). Kitamura and Murata (1979) This paper is intended to provide a basic merely merely quoted sectional names in a key to systematic information for J apanese Salix the the species of J apanese Salix in their Flora accumulated to date and my opinions on the of of Japanese Woody . It lacks any ci- taxa examined. tation tation of synonymy and references for the sections sections and their infrageneric taxa are February February 2000 Journal of Japanese Botany Vo l. 75 No. 1 3

Systematic Ii st of Japanese Salix with 6-3. subsp. yezoalpina (Koidz.) H.Ohashi エ the the Japanese name ゾタカネヤナギ (New names in boldface) 6-3- 1. f. yezoalpina (Koidz.) H.Ohashi *Cultivated *Cultivated plants 6-3-2. f. neoreticulata (Nakai) H.Ohashi イ Fam. Salicaceae Mirb. ヤナギ科 ヌマルパヤナギ Subfam. Salicoideae Kimura ヤナギ亜科 11 1. Subgen. Salix ヤナギ亜属 Gen. Salix L. ヤナギ属 III-i. Sec t. Triandrae Dumortier 1. 1. Subgen. Urbanianae (Seemen) タチヤナギ節 H.Ohashi トカチヤナギ亜属 7. Salix triandra L. タチヤナギ I-i. I-i. Sec t. Chosenia (Nakai) Kimura III-ii. Sec t. Subalbae Koidz. ケショウヤナギ節 シダレヤナギ節 1. 1. Salix arbutifolia Pal l.ケショウヤナギ 8. *Sα lix babylonica L. i. I-i i. Sect. Urbanianae (Seemen) 8 ..， 1. *f. babylonic αシダレヤナギ C.K.Schneid. トカチヤナギ節 8-2. *f. rokkaku Kimura ロッカクヤナギ 2. 2. Salix cardiophylla Trautv. & Mey. トカ 8-3. *f. seiko Kimura セイコヤナギ チヤナギ 9. *Salix matsudana Koidz. va r. tortuosa 2-1. 2-1. var. cardiophylla Vilm. ウンリュウヤナギ、 2-2. 2-2. var. urbaniana (Seemen) Kudo オオパ 10. Salix eriocarpa Franch. & Sav. ジャヤ ヤナギ ナギ I-iii. I-iii. Sec t. Glandulosae Kimura 11. Salix pierotii Miq. オオタチヤナギ マルバヤナギ節 11-1. f. pierotii 3. 3. Salix chaenomeloides Kimura マルパヤ 11-2. *f. auricoma Kimura コガネヤナギ ナギ 12. Salix yoshinoi Koidz. ヨシノヤナギ 11. 11. Subgen. Chamaetia (Dumortier) Nasarov 13. Salix jessoensis Seemen シロヤナギ

エゾマメヤナギ亜属 13 同 1. subsp. jessoensis II-i. II-i. Sec t. Herbella Seringe 13-2. subsp. serissaefolia (Kimura) エゾマメヤナギ節 H.Ohashi コゴメヤナギ 4. 4. Salix nummularia Andersson エゾマメヤ IV. Subgen. Vetrix バッコヤナギ亜属 ナギ IV-i. Sec t. Hastatae (Fries) A.Kerner 4-1. 4-1. f. nummulari α シパヤナギ節 4-2. 4-2. f. hebecarp α(Kimura) H.Ohashi キヌ 14. Salix japonic αThunb. シパヤナギ ゲエゾマメヤナギ 15. Salix shiraii Seemen シライヤナギ II-ii. II-ii. Sec t. Myrtilloides (Borrer) Andersson 16. Salix rupifraga Koidz. コマイワヤナギ ミヤマヤチヤナギ節 16- 1. f. rupifrag α 5. 5. Salix fuscescens Andersson ミヤマヤチ 16-2. f. eriocarpa (Kimura) Kimura ex ヤナギ H.Ohashi ケコマイワヤナギ II-iii. II-iii. Sec t. Glaucae (Fries) Andersson IV -ii. Sec t. Sieboldianae C.K.Schneid. タカネイワヤナギ百行 ヤマヤナギ節 6. 6. Salix nakamurana Koidz. タカネイワヤ 17. Salix sieboldiana Blume ヤマヤナギ ナギ 18. Salix reinii Seemen ミネヤナギ 6・1. subsp. nakamurana 18-1. f. reinii 6-1-1. 6-1-1. f. nakamurana 18-2. f. pendula Kimura シダレミネヤナギ 6-1-2. 6-1-2. f. eriocarp α(Kimura) T .Shimizu ケ IV -iii. Sec t. Helix Dumortier タカネイワヤナギ コリヤナギ節 ふ2. subsp. kurilensis (Koidz.) H.Ohashi ヒ 19. Salix miy αbean αSeemen エゾノカワヤ ダカミネヤナギ ナギ 4 植物研究雑誌第75 巻第l号 平成12 年2月

19- 1. subsp. miy αbean α IV -ix. Sec t. Cinerella Seringe 19-2. 19-2. subsp. gilgiana (Seemen) H.Ohashi カ バッコヤナギ節 ワヤナギ 28. Salix taraikensis Kimura タライカヤナ 19-2- 1. f. gilgiana (Seemen) H.Ohashi ギ 19-2-2. 19-2-2. f. pendula (Kimura) H.Ohashi シダ 29. Sα lix cα :prea L. パッコヤナギ レカワヤナギ 30. Salix ルtura Seemen オオキツネヤナギ 20. 20. Salix integra Thunb. イヌコリヤナギ 30- 1. f. ルtura 20-1. 20-1. f. integra 30-2. f. rufa (Kimura) H.Ohashi アカゲオ 20-2. 20-2. f. pendula Kimura シダレイヌコリヤ オキツネヤナギ ナギ 31. Salix vulpina Andersson キツネヤナギ 1. 21. *Salix koriy α nagi Kimura ex Goerz コ リヤナギ Putative natural hybrids IV-iv. IV-iv. Sec t. Incubaceae A.Kerner (in alphabetical order) ノヤナギ節 H 1. Salix xα 19ista C.K.Schneid. トウゲヤ 22. 22. Salix subopposita Miq. ノヤナギ ナギ IV-v. IV-v. Sec t. Subviminales C.K.Schneid. H2. Salix xα mpherista C.K.Schneid. ハコ ネコヤナギ節 ダテヤナギ 23. 23. Salix gracilistyla Miq. ネコヤナギ H3. Salix x arakian αKoidz. ナガパノネコ 23-1. 23-1. var. gracilistyl α ヤナギ 23-1-1. 23-1-1. f. gracilistyla H4. Salix caprea L. x S. futura Seemen 23-1-2. 23-1-2. f. adscendens (Kimura) H.Ohashi H5. Salix x cremnophylla Kimura ネコシノて タチネコヤナギ ヤナギ

23 聞 1-3. f. pendula (Kimura) H.Ohashi シダ H6. Salix x eriocataphylla Kimura シグレ レネコヤナギ ヤナギ 23-1-4. 23-1-4. f. variegata (Kimura) Kimura フイ H7. SaUx x eriocataphylloides Kimura ナス リネコヤナギ ノシグレヤナギ 23-2. 23-2. va r. graciliglans (Nakai) H.Ohashi H8. Salix x euerata Kimura カワオノエヤ チョウセンネコヤナギ ナギ 23-3. 23-3. *var. melanostachys (Makino) H9. Salix futura Seemen x S. udensis C.K.Schneid. クロヤナギ Trautv. & Mey. IV -v i. Sec t. Hukaoana Kimura HI0. Salix x gracilis り110ides Kimura ロッコ ユピソヤナギ節 ウヤナギ 24. 24. Salix hukaoana Kimura ユピソヤナギ Hll. Salix x hα pala Kimura カワイヌコリ IV -vii. Sec t. Daphnella Seringe ヤナギ エゾヤナギ節 H12. Salix x hatusimai Kimura チクゼンヤ 25. 25. Salix rorida Lacksch. エゾヤナギ ナギ 25 -1. f. ro rida H 13. Salix x hayatana Kimura リュウゾウ 25-2. 25-2. f. pendula Kimura シダレエゾヤナギ ジヤナギ

25-3. 25-3. f. roridaeformis (Nakai) Kimura ex H14. Salix x hir αoα， na Kimura ヒラオヤナ H.Ohashi コエゾヤナギ ギ. -viii. IV -viii. Sec t. Viminella Seringe H 15. Salix x hisauchiana Koidz. フジヤナ キヌヤナギ節 ギ 26. 26. Salix schwerinii E. Wolf キヌヤナギ H16. Salix x ikenoana Kimura イケノヤナ 27. 27. Salix udensis Trautv. & Mey. オノエヤ ギ

ナギ H 17. Salix integr αThunb.， x S. miy αbeana Seemen February February 2000 Journal of Japanese Botany Vo l. 75 No. 1 5

H 18. Salix x iwahisana Kimura イワヒサヤ Systematic enumeration of ナギ Japanese 8α lix H19. Salix x kamikotica Kimura カミコウ チヤナギ Fam. Salicaceae Mirb. ，Elem. Physio l. H20. Salix x kawamurae Kimura ミョウジ Veg. Bo t. 2: 905 (1 815). Ty pe. Salix L. ンヤナギ Subfam. Salicoideae Kimura in Bo t. H2 1. Salix x koidzumii Kimura トヨハラヤ Mag. Tokyo 42: 290 (1 928); Kimura in Sci. ナギ Rep. Tohoku Imp. Univ. 4 ser. (Bio l.) 13: H22. Salix x koiei Kimura コイエヤナギ 391 (1 938). Ty pe. Salix L. H23. Salix x lasiogyne Seemen シロシダレ ヤナギ Gen. Salix L. H.23-1. H.23-1. Sα lix x lasiogyne nothosubsp. Salix L. ， Sp. Pl.: 1015 (1 753) & Gen. Pl. lasiogyne lasiogyne ed. 5，447 (1 754); Thunb. ， F l. Japon.: 24 H.23-2. H.23-2. Sa !i x x lasiogyne nothosubsp. yuhkii (1 784); Andersson in DC. ，Prodr. 16 (2): 191 (Kimura) (Kimura) H.Ohashi ユウキシダレ (1 868); Franch. & Sav. ，Enum. Pl. Jap. 1: H24. Salix x leucopithecia Kimura フリソ 458 (1 875) & 2: 502 (1 878); Pax in Engler デヤナギ & Prantl ，Na t. Pflanzenfam. 3， 1: 36 (1 887); H25. Salix x matsumurae Seemen オクヤマ Seemen in Bo t. Jahrb. 21: Beib l. 52: 6 サルコ (1896) & Salic. Jap.:1 (1903); C. K. H26. Sa !i x x microstemon Kimura ワケノカ Schneid. ，Illus t. Handb. Laubh. 1: 2 (1 904) ワヤナギ & in Sarg. ， P l. Wils. 3: 40 (1 916); Koidz. in H27. Salix x nasuensis Kimura ナスノイワ Bo t. Mag. Tokyo 27: 87 & 264 (1 913); ヤナギ Kimura in Bo t. Mag. Tokyo 40: 7 (1 926) & H28. Salix x pedionoma Kimura パッコキ 42: 290 (1 928); Nakai in Bull. Soc. Dendr. ヌヤナギ Fr. no. 66: 6 (1 928) & Fl. Sylv. Korea. 18: H29. Salix x pseudopaludicola Kimura オオ 65 (1 930); Makino & Nemoto ，Nippon- ミヤマヤチヤナギ Shokubutsu-Soran 2nd ed.: 161 (1 931); H30. Salix x send α ica Kimura センダイヤ Nemoto ，Nippon-Shokubutsu-Soran-Hoi: ナギ 105 (1 936); Kimura in Sci. Rep. Tohoku H31. Salix x sigemitui Kimura フカオヤナ Imp. Univ. 4 ser. (Bio l.) 13: 393 (1 938); ギ Rehder ，Man. Cul t. Tr. Shr. ed. 2: 82 (1 940); H32. Salix x sirakawensis Kimura コセキヤ Ohwi ， F l. Jap.: 398 (1 953) ， F l. Jap. ed. ナギ Eng l.: 362 (1 965) & F l. Jap. ed. rev.: 465 H33. Salix x sugayana Kimura スガヤヤナ (1 965); Hutch. ，Gen. Fl. Pl. 2: 118 (1967); ギ Skvortsov ， USSR: 92 (1 968); Dorn H34. Salix x sumiyosensis Kimura スミヨシ in Can. J. Bot. 54: 2774 (1976); Kitam. & ヤナギ Murata ，Co l. Il lus t. Wood. P l. J ap. 2: 304 H35. Salix x thaymasta Kimura ミヤコヤナ (1 979); Kitag. ，Neo-Lineam. F l. Mansh.: ギ 204 (1 979); Chou in Fl. Reipub. Pop. Sin. H36. Salix x turumatii Kimura ヒタチヤナ 20 (2): 81 (1 984); Kimura in Sci. Rep. ギ Tohoku Univ. 4 ser. (Bio l.) 39: 143 & 147 H37. *Salix xjapopina Kimura シノミキツネ (1 988); Ohwi & Kitag. ，New F l. Jap.: 530 ヤナギ (1 992); Newsholme ， Willows: 22 (1 992); Sohma in Sci. Rep. Tohoku Univ. 4 ser. (Bio l.) 40: 77 (1993) [pollen]; Nedoluzhko in in Charkevicz ， P l. Vascu l. Ori. Extr. Sovie t. 6 植物研究雑誌第75 巻第1号 平成12 年2 月

7: 7: 158 (1995); Lee ， Lineam. Fl. Korea.: 139 mediate between Populus and Salix and has (1996); (1996); Argus in Sys t. Bo t. Monog. 52: 48 been placed at the root of Salix (N akai 1920 ， (1 997). Ty pe: Salix alba L. Kimura 1928 ，1989 ，Chou 1984 ，Argus Choseni αNakai* in Bo t. Mag. Tokyo 34: 1997). However ， occurrence of a form with 67 (1 920) & Fl. Sylv. Kor. 18: 58 (1 930); rudimentary nectaries emphasized by Kimura in Bo t. Mag. Tokyo 42: 290 (1 928) ， Kimura (1 988) may suggest that Chosenia sub sub Salicoideae tribe Chosenieae; Rehder ， was derived from.a species of Salix by loss Man. Cul t. Tr. Shr. ed. 2: 82 (1 940); Ohwi ， of nectaries. The results of chloroplast DNA l. F l. Jap.: 397 (1 953); Ohwi ， F l. Jap. ed. studies of Sα lix ，Chosenia and Toisusu by Eng l.: 362 (1965) & Fl. Jap. ed. rev.: 464 Azuma et al. (1999 in press) did not support (1 965); Kitam. & Murata ，Co l. Illus t. Woody the distinct status of Chosenia and Toisusu. l. Pl. Jap. 2: 336 (1 979); Kitag. ，Neo-Lineam. 1 treat Chosenia ，therefore ， as congeneric l. F l. Mansh.: 201 (1979); Chou in F l. with Salix. Reipub l. Popu l. Sin. 20 (2): 79 (1 984); Kimura in Wild Flow. Jap. Wood. 1: 34 1. Subgen. Urbanianae (Seemen) (1 989); Ohwi & Kitag. ，New Fl. Jap.: 529 H.Ohashi (1 992); Sohma in Sci. Rep. Tohoku Univ. 4 Salix subgen. Urbanianae (Seemen) ser. ser. (Bio l.) 40: 172 (1 993); Nedoluzhko in H.Ohashi ，sta t. nov. Type: Salix cardio- Charkevicz ， P l. Vascu l. Ori. Extr. Sovie t. 7: phyll αTrautv. &乱 1ey. (= S. urb αnian α 155 (1 995); Lee ，Lineam. Fl. Korea.: 135 Seemen). (1 996); Argus in Sys t. Bo t. Monog. 52: 13 Salix subgen. Pleuradenia Kimura in Bo t. (1 997). Ty pe: Chosenia arbut ザ'o lia (Pall.) Mag. Tokyo 41: 498 (1 927) ，as “Pleura- Skvortsov Skvortsov (= Salix arbutifolia Pall. ， Salix deniae" & in Sci. Rep. Tohoku Univ. 4 ser. splendida N akai ，Chosenia splendida (Bio l.) 39: 143 (1 988). Ty pe: Salix urbani- (Nakai) (Nakai) Nakai). ana Seemen. Toisusu Toisusu Kimura in Bot. Mag. Tokyo 42: Salix subgen. Protitea Kimura in Bo t. 288 (1928) ， sub Salicoideae tribe Saliceae; Mag. Tokyo 42: 290 (1 928 May) & in Sci. Ohwi ， F l. Jap.: 397 (1 953); Ohwi ， F l. Jap. Rep. Tohoku Imp. Univ. 4 ser. (Bio l.) 13: ed. ed. Eng l.: 362 (1 965) & Fl. Jap. ed. rev.: 393 (1 938); Dorn in Can. J. Bo t. 54: 2774 464 (1 965); Kitam. in Kitam. & Murata ， (1 976); Kimura in Sci. Rep. Tohoku. Univ. Co l. Illus t. Woody Pl. Jap. 2: 335 (1 979); 4 ser. (Bio l.) 39: 147 (1 988). Ty pe: Salix Ohwi & Kitag. ，New Fl. Jap.: 529 (1 992); glandulosa Seemen (= S. chaenomeloides Kimura in Wild Flow. Jap. Wood. 1: 34 Kimura). (1 989); Zhang in Bul l. Bot. Research 14: Salix subgen. Pleiolepis Nakai in Bull. 304 (1 994); Nedoluzhko in Charkevicz ， P l. Soc. Dendro l. Fr. 66: 42 (1 928 Aug.). Ty pe: Vascu l. Ori. Extr. Sovie t. 7: 155 (1 995). Salix glandulosa Seemen (Lectotype desig- Ty pe: Toisusu cardiophylla (Trautv. & Mey.) nated by Argus 1997). Kimura (= Salix cardiophyll αTrautv. & Salix subgen. Salix ，p.p.: Skvortsov in Mey.). Mey.). Acad. Sci. URSS Nov. Sys t. Pl. 1968: 63 x Toisochosenia Kimura in Sci. Rep. (1 968) & Willows USSR: 97 (1 968) ， p.p. Tohoku Imp. Univ. 4 ser. (Bio l.) 11: 247 inc l. sec t. Humboldianae & Urbanianae; (1 936); Kitam. & Murata ，Co l. Illus t. Wood. Dorn in Canad. J. Bo t. 54: 2774 (1976) ，p.p. Jap. Jap. 2: 337 (1 979). Ty pe: x Toisochosenia inc l. subgen. Pleiolepis & Protitea; Chou in kamikotic α(Kimura) Kimura (= Salix Fl. Reipub. Pop. Sin. 20 (2): 99 (1984) ，p.p. ， kamikotica kamikotica Kimura). inc l. sec t. Tetraspermae & Wilsonianae; *Note: *Note: Chosenia was considered as inter- Argus in Sys t. Bo t. Monogr. 52: 50 (1 997) ， Pebruary Pebruary 2000 Journal of Japanese Botany Vo l. 75 No. 1 7 p.p. ，inc l. sec t. Floridanae & Humboldianae. Wild Flow. Jap. Wood. 1: 34 ， photo 42 ，1- 6 (1 989); Makino ，New Illus t. Fl. Jap. rev. I-i. I-i. Sec t. Chosenia (Nakai) Kimura ed.: fig. 108 (1 989); Sohma in Sci. Rep. Salix Salix sec t. Chosenia (Nakai) [Flod. in Tohoku Univ. 4 ser. (Bio l.) 40: 88 (1 993); Arkiv Bo t. 25A: 2 (1 933) ，comb. nud.] Nedoluzhko in Charkevicz ， P l. Vascu l. Ori. Kimura in Sc i. Rep. Tohoku Univ. 4 ser. Extr. Sovie t. 7: 155 (1 995). (Bio l.) 39: 145 (1 988). Chosenia arbutifolia mst r. adenantha Chosenia Chosenia Nakai in Bo t. Mag. Tokyo 34: (Kimura) Kimura in Sci. Rep. Tohoku Univ. 67 67 (1 920). 4 ser. (Bio l.) 27: 146 (1 961). Sα lix sec t. Pentandrae Dum. ex Chosenia arbutifoli αf. adenantha C.K.Schneid. in Sarg. ， P l. Wils. 3: 98 (Kimura) Kimura ex Kitag. ，Neo-Lineam. (1 916) ，p.p. Fl. Mansh.: 202 (1 979); Kimura in Sci. Rep. Tohoku Univ. 4 ser. (Bio l.) 39: 145 (1988) ， 1. 1. Salix arbutifolia Pall. ， F l. Ross. 1 (2): sub Fig. 2. 79 (1788). Dist r.: Japan ，Korea ，NE. China ， Sakhalin Salix Salix eucalyptoides Mey. ex C.K.Schneid. and East Siberia. in in Sarg. ， P l. Wils. 3: 99 (1 916). *Note: Kimura (1 938) described a form Salix Salix bracteosa Turcz. ex Trautv. in with rudimentary nectaries in fruit in this Middend. Middend. Reise Sibir. 1 (2): 77 (1 856). species. He considered this form as abnor- Salix Salix splendida Nakai in Bo t. Mag. Tokyo mal and named it Chosenia macrolepis 32: 32: 215 (1 918). (Turcz.) Komarov mstr. adenantha Kimura Chosenia Chosenia splendida (Nakai) Nakai in Bo t. or C. arbutifolia mstr. adenantha (Kimura) Mag. Tokyo 34: 67 (1 920). Kimura. The rank of these names are ，how- Chosenia euc αlyptoides (C.K.Schneid.) ever ， not clear whether “mst r." was corre- Nakai Nakai in J. Arn. Arb. 5: 72 (1 924). sponding to a known rank or was designated Chosenia bracteosa (Turcz. ex Trautv.) as a new additional rank between the known Nakai ， F l. Sylv. Kor. 18: 59 ，t. 3-5 (1 930) ， ranks below species. Later ， Kitagawa (1 979) as as C. bracteosa Nakai; Kimura in J. Fac. and Kimura (1988) regarded mst r. Agric. Agric. Hokkaido Imp. Univ. Sapporo 26: adenantha Kimura as a form. Kimura (1 988) 394 (1 934); Makino ，Il lus t. Fl. Nippon: 670 ， considered this abnormal form as an evi- fig. fig. 2008 (1 940); Ohwi ， F l. Jap.: 397 (1 953) ， dence intermediate Chosenia and Salix. as as C. bracteosa (Turcz.) Nakai. Chosenia Chosenia macrolepis (Turcz.) Komarov I-i i. Sect. Urbanianae (Seemen) mst r. adenantha Kimura* in Sci. Rep. C.K.Schneid. Tohoku Univ. 4 ser. (Bio l.) 13: 389 (1 938). Salix sec t. Urbanianae (Seemen) Voucher Voucher specimen: Manchuria. Jalatun. P. H. C.K.Schneid. in Sarg. ， P l. Wilson. 3: 103 & J. H. Dorsett 3481 (TUS-K). (1 916); Nakai ， F l. Sylv. Korea. 18: 71 Chosenia Chosenia arbutifoli α(Pal l.) Skvortsov in (1 930); Skvortsov in Acad. Sci. URSS Nov. No t. Sys t. Herb. Ins t. Bo t. Kom. Acad. Sci. Sys t. Pl. 1968: 64 (1 968) & Willows USSR: URSS 18: 43 (1 957); Kimura in Sci. Rep. 103 (1 968) ， sub subgen. Salix; Chou in Fl. Tohoku Univ. 4 ser. (Bio l.) 27: 146 (1 961); Reipub. Pop. Sin. 20 (2): 97 (1 984); Kimura Ohwi ， F l. Jap. ed. Eng l.: 362 (1 965); Sugaya in Sci. Rep. Tohoku Univ. 4 ser. (Bio l.) 39: in in Terasak i. Il lus t.: 98 ，fig. 380 (1 977); 144 (1988) ，sub subgen. Pleuradenia Kitam. Kitam. & Murata ，Co l. Il lus t. Woody Pl. Jap. Kimura; Newsholme ， Willows: 23 (1 992) ， 2: 2: 336 ，f. 793 (1 979); Y. L.Chou Fl. Reipub l. sub subgen. Salix ， p.p. exc l. Choseni α. Popu l. Sin. 20 (2): 79 (1984); Kimura in Type: Salix urbaniana Seemen (= S. 8 植物研究雑誌第75 巻第I号 平成12 年2月 cardiophylla cardiophylla Trautv. & Mey.). (1 989). Salix Salix [unranked] Urbanianae Seemen in 2-1. Salix cardiophyll αvar. cardiophylla Salic. Salic. Jap.: 15 (1903) ，sub Salix A. Dist r.: Japan: Hokkaido and Honshu ， Didymadeniae a. Pleonandrae s. Dolicho- Kuriles ，Korea ， East China ， SE. Siberia and stylae. stylae. Ty pe: Salix urbaniana Seemen (= S. Far East Russia: Primorye ，Amur ， cardiophylla cardiophylla Trautv. & Mey.). Khabarovsk and Sakhalin. Toisusu Toisusu Kimura in Bo t. Mag. Tokyo 42: 2-2. Salix cardiophyll αvar. urbaniana 288 (1 928). (Seemen) Kudo ，Nippon Yuyo-jumoku Bunruigaku ed. 1: 77 (1 922); Makino & 2. 2. Salix cardiophylla Trautv. & Mey. ， F l. Nemoto ， F l. Jap. ed. 1: 1121 (1 925). Ocho t.: 77 ，tab. 19-20 (1 856); Tokubuchi in Salix urbaniana Seemen in Bo t. J ahrb. Bo t. Mag. Tokyo 10: page (1 22) (1 896); Sys t. 21: Beib l. 52: 9 (1895) [Ty pe: Japan. Koidz. Koidz. in Bo t. Mag. Tokyo 27: 97 (1 913); Hakodate und Nipon. Prov. Nambu. Miyabe & Miyake ， F l. Saghalin: 424 (1 915); Maximowicz ，1861 ， n. 2 (K ，L isosyn.) & C.K.Schneid. C.K.Schneid. in Sarg. ， P l. Wilson. 3 (1): 103 Maximowicz ，1865 ， n. 4 (K ，L isosyn.)] & (1 916); Skvortsov ，Wi l1 0ws USSR: 103 Salic. Jap.: 24 ， tab. 1 (1903); Tokubuchi in (1 968). Bo t. Mag. Tokyo ， 10: page (122) (1896); Salix Salix maximowiczii Kom. in Ac t. Hor t. Shirasawa ，Ic on. Ess. For. Trees Jap. 2: tab. Petrop. Petrop. 18: 442 (1 90 1); C.K.Schneid. in 10 ，1-12 (1 908); Miyabe & Miyake ， F l. Sarg. ， P l. Wilson. 3 (1): 100 (1 916); Nakai ， Saghalin: 424 (1 915); C.K.Schneid. in Sarg. ， l. F l. Sylv. Korea. 18: 72 ，t. 6-7 (1930); P l. Wilson. 3 (1): 103 (1 916); Makino ， Kitag. ，Neo-Lineam. F l. Mansh.: 206 Il lus t. Fl. Nippon: 670 ， fig. 2009 (1940); (1 979); Chou in F l. Reipub. Pop. Sin. 20 Hatusima ，Nihon-no-jyumoku: 690 (1 976). (2): (2): 99 ，t. 26 (1 984). Toisusu cardiophylla va r. urbani αna Salix Salix urbani αna var. schneideri Miyabe & (Seemen) Kimura in Bo t. Mag. Tokyo 42: Kudo ，Ic on. Ess. For. Trees Hokkaido 1: 45 ， 289 (1 928). tab. tab. 13 (1921). Toisusu urbaniana (Seemen) Kimura in J. [Sα lix cardiphylla var. schneideri (Miyabe Fac. Agric. Hokkaido Imp. Univ. Sapporo & Kudo) Kudo ，Nippon Yuyo-jumoku 26: 397 (1 934); Ohwi ， F l. Jap. ed. Eng l.: Bunruigaku ， ed. 1: 78 (1 922) ，comb. nud.] 362 (1 965); Sugaya in Terasaki. Illus t.: 98 ， Toisusu Toisusu cardiophyll α(Trautv. & Mey.) fig. 381 (1 977); Kitam. in Kitam. & Murata ， Kimura in Bo t. Mag. Tokyo 42: 288 (1928) ， Co l. Illus t. Woody Pl. Jap. 2: 335 ，fig. 792 in in J. Jap. Bo t. 24: 546 (1 949) & in Sci. Rep. (1 979); Ohwi & Kitag. ，New F l. Jap.: 530 Tohoku Univ. 4 ser. (Bio l.) 18: 546 ，fig. 2 (1 992); Nova Fl. Nasu. Supp l.: 118 ， figs. (1 950); Nedoluzhko in Charkevicz ， P l. 61-65 (1 985); Kimura in Wild Flow. Jap. Vascu l. Ori. Extr. Sovie t. 7: 155 (1 995). Wood. 1: 34 ， photo 41 ， 1-5 (1 989); Makino ， Toisusu cardiphylla va r. schneideri New Il lus t. Fl. Jap. rev. ed.: fig. 87 (1 989); (Miyabe & Kudo) Kimura in Bo t. Mag. To- Nedoluzhko in Charkevicz ， P l. Vascu l. Or i. kyo 42: 289 (1 928). Extr. Sovie t. 7: 158 (1 995). Toisusu Toisusu cardiophylla var. maximowiczii Salix cα rdiophylla subsp. urbaniana (Kom.) Kimura in Bo t. Mag. Tokyo 42: 289 (Seemen) Skvortsov ，Willows USSR: 104 (1928). (1928). (1 968); Sohma in Sci. Rep. Tohoku Univ. 4 Toisusu urbaniana va r. schneideri ser. (Bio l.) 40: 79 ， 121 (1993). (Miyabe & Kudo) Kimura in J. Fac. Agric. Toisusu urbaniana mstr. nematadenia Hokkaido Imp. Univ. Sapporo 26: 397 Kimura in Sci. Rep. Tohoku Univ. 4 ser. (1 934) & in Wild Flow. Jap. Wood. 1: 34 (Bio l.) 20: 206 (1 954) [Ty pe: Japan. Iwate February February 2000 Journal of Japanese Botany Vo l. 75 No. 1 9

Pref. Pref. (Prov. Rikutyu): secus fl. Sekiguchi- Rep. Tohoku Imp. Univ. 4 ser. (Bio l.) 13: 77 gawa in viculo Sekiya prope oppidulum (1 938); Ohwi ， F l. Jap. ed. Eng l. 364 (1 965) Yamada. S. Sasamura 1188 (fr.) ，10 Ju l. & F l. J ap. ed. rev.: 468 (1 965); Kurata ， 1949 (TUS-K)]. Il lus t. Imp. For. Tr. Jap. 3: 6， p l. 3 (1 971); Toisusu urbaniana mst r. composita Sugaya in Terasaki. Il lus t.: 98 ， fig. 382 Kimura in Sci. Rep. Tohoku Univ. 4 ser. (1977); Hatusima ，Kagoshima-ken (Bio l.) 20: 207 (1954) [Type: Japan. Shokubutsu Mokuroku: 35 (1 978); Kitam. & Hokkaido ， Prov. Ishikari ，ad ripas fl. Murata ，Co l. Il lus t. Woody Pl. Jap. 2: 332 ， Toyohiragawa ，Sapporo ，A. Kimura 642 (♀ fig. 791 (1 979); Chou in Fl. Reipub. Pop. fl.) ，20 May 1927 (TUS-K)]. Sin. 20 (2): 105 (1 984); Kimura in Nova Fl. Distr.: Distr.: Japan: Hokkaido and Honshu ， Nasu. Supp l.: 113 (1 985); Kimura in Wild Kuriles ，Korea ， Eastern China and East Rus- Flow. Jap. Wood. 1: 39 ，photo 43 ，1-3 sia: sia: Primorye ，Amur ，Kh abarowsk ， Sakhalin (1 989); Makino ，New Il lus t. Fl. Jap. ed.: rev. and SE. Siberia. fig. 88 (1 989); Ohwi & Kitag. ，New Fl. Jap.: Miyabe and Miyake (1 915) suggested for 533 (1 992); Sohma in Sci. Rep. Tohoku the the first time Salix urbaniana as a variey of Univ. 4 ser. (Bio l.) 40: 103 (1 993). Sα lix cardiophylla. Salix glandulosa Seemen in Bo t. Jahrb. Sys t. 21: Beib l. 53: 55 (1 896) & Salic. Jap.: I-iii. I-iii. Sec t. Glandulosae Kimura 22 ，tab. 1 (1 903) ，non Raf. (1 838): Koidz. in Salix Salix sect. Glandulosae Kimura in Bo t. Bo t. Mag. Tokyo 27: 87 (1 913); Kimura in Mag. Tokyo 42: 68 (1 928) ，sub subgen. Bo t. Mag. Tokyo 42: 69 (1 928); Nakai ， F l. Protitea Protitea Kimura; Kimura in Sci. Rep. Sy l. Korea. 18: 77 (1 930); Makino ，Il lus t. Tohoku Imp. Univ. 4 ser. (Bio l.) 13: 81 Fl. Nippon: 670 ，fig. 2010 (1 940). (1938); (1938); Kitam. & Murata ，Co l. Il lus t. Woody Distr.: J apan: Honshu (Southern Tohoku l. Pl. Jap. 2: 307 (1 979); Newsholme ，Wil- District and southward) ，Shikoku and lows: lows: 24 (1 992) ，as “Glandulosae Seem." Kyushu ，Korea and China. Type: Salix glandulosa Seemen (= S. chaenomeloides chaenomeloides Kimura). 11. Subgen. Chamaetia (Dumortier) Salix Salix [unranked] Pentandrae Seemen ， Nasarov Salic. Salic. Jap.: 15 (1 903) ， sub Salix A. Didyma- Salix subgen. Chamaetia (Dumortier) deniae deniae a. Pleonandraeα. Brachystylae ，non Nasarov ， F l. URSS 5: 31 (1 936); Rechinger Borrer (1830). Type: Salix glandulosa in Fl. Europ. 1: 46 (1 964); Argus in Sys t. Seemen. Bo t. Monogr. 52: 58 (1 997). Salix Salix sec t. Pentandrae C.K.Schneid. in Salix sect. Chamaetia Dumortier ， Verh. Sarg. ， P l. Wils. 3: 98 (1 916) ， as “ Pentandrae Ges l. Wilgen: 15 (1825) & in Bijd r. Dumortier" ，p.p. ci t. S. glandulosa Seemen. Natuurk. Wetensch. 1: 56 (1 826). Type: Salix Salix sec t. Glandulosae Nakai in Bull. Sα lix reticulata L. Soc. Soc. Dendr. France no. 66: 7 (1 928 Aug.) ， non Kimura (1928 Jan.). Type: Salix II-i. Sec t. Herbella Seringe glandulosa glandulosa Seemen. Salix sec t. Herbella Seringe ， Exemplaires Salix Salix sec t. Wilsonianae Hao in Reper t. desseches de la revision inedit du genre Spec. Spec. Nov. Regni Veg. Beih. 93: 41 (1 936); Salix ， 14th page [no pagination] (1 824); Chou in F l. Reipub. Pop. Sin. 20 (2): 99 Argus in System. Bo t. Monog r. 52: 62 (1 984). Ty pe: Salix wilsonii Seemen. (1 997) ， sub subgen. Chamaetia (Dumortier) Nasarov. Nasarov. Ty pe: Salix herbacea L. 3. 3. Salix chaenomeloides Kimura in Sci. Salix [unranked] Herbaceae Borrer in 10 10 植物研究雑誌第75 巻第1号 平成12 年2月

Hook. ，Bri t. Fl.: 432 (1830). Salix nummularia subsp. pauciflora Salix Salix sec t. Retusae A.Kerner in Verh. (Koidz.) Kimura in Bo t. Mag. Tokyo 59: 87 Zoo l. -Bo t. Ges. Vereins Wien 10: 195 (1 946); Ohwi & Kitag. ，New F l. Jap.: 535 (1 860); Skvortsov in Acad. Sci. URSS Nov. (1 992). Sys t. Pl. 1968: 65 (1968) & Willows USSR: Salix nummularia subsp. pauciflora var. 118 (1968) ，sub subgen. Chamaetia cyclophylla (Kimura) Kimura in Bo t. Mag. (Dumortier) Nasarov; Kitam. & Murata ， Tokyo 59: 87 (1 946). Co l. Il lus t. Woody Pl. Jap. 2: 307 (1 979) ， Salix nummularia subsp. pauciflora var. sub subgen. Salix. Ty pe: Salix retusa L. stenophylla (Kimura) Kimura in Bo t. Mag. Salix Salix sec t. Herbaceae (Seringe) A.Camus Tokyo 59: 88 (1 946). & Camus ，Classif. saules Eur. 1: 106 (1 904); Distr.: J apan: Hokkaido and Russia: C.K.Schneid. in Sarg. ， P l. Wils. 3: 142 Kamchatka ，Magadan ， Siberia to Russia (1 916) ，as “Herbaceae Borrer"; Nakai ， F l. North. Sylv. Sylv. Kor. 18: 145 (1 930) ，as “Herbaceae 4-1. Salix nummularia f. nummularia Borrer". Borrer". Ty pe: Salix herbacea L. 4-2. Salix nummularia f. hebecarpa (Kimura) (Kimura) Kimura in Bo t. Mag. Tokyo 59: 88 4. 4. Salix nummularia Andersson in DC. ， (1 946). Prodr. Prodr. 16 (2): 298 (1 868); Skvortsov ，Wil- Salix pauciflora Koidzumi var. hebec α rpa lows USSR: 122 (1968); Nedoluzhko in Kimura in J. Fac. Agric. Hokkaido Imp. Charkevicz ， P l. Vascu l. Ori. Extr. Sovie t. 7: Univ. 26 (4): 413 (1 934) [Type: Japan. 206 (1 995). Hokkaido. Prov. Ishikari ， in regione alpina Salix Salix pauciflora Koidz. in Bo t. Mag. To- montis Daisetsusan. A. Kimura 1774 (♀)， 7 kyo 32: 61 (1 918) [Ty pe: Japan. Yezo. M t. Ju l. 1928 (TUS-K)]. Nutakkamshpe (= Mt. Daisetsu) ， alpine bel t. Salix pauciflora f. hebecarpa (Kimura) H. Koidzumi s.n. Aug. 1917 (TI holo.)]; Kimura: Ohwi: 365 (1 965); Kimura in Wild Kimura in J. Fac. Agric. Hokkaido Imp. Flow. Jap. Wood. 1: 42 (1 989). Univ. Univ. Sapporo 26: 411 (1 934) & in Bo t. Salix nummularia var. hebecarpa Kimura: Mag. Tokyo 42: 575 (1 928); Ohwi ， F l. Jap. Ohwi & Kitag. ，New F l. Jap.: 535 (1 992). ed. ed. Eng l.: 365 (1 965) & F l. Jap. ed. rev.: Distr.: J apan. Occurs with the typical 469 (1 965); Sugaya in Terasaki. Il lus t.: 100 ， form. fig. fig. 388 (1 977); Kitam. & Murata ，Co l. Il lus t. Woody P l. Jap. 2: 326 ， fig. 783 II-ii. Sec t. Myrtilloides (Borrer) (1 979); Shimizu ，New Alp. Fl. Jap. 2: 110 ， Andersson fig. fig. 318 (1 983); Kimura in Wild Flow. Jap. Salix sec t. Myrtilloides (Borrer) Wood. 1: 41 ，photo 46 ，4-6 (1 989). Andersson in DC. ，Prod r. 16 (2): 229

Salix Salix pauciflor αvar.， stenophylla Kimura (1868) ，as “Salix sec t. Roseae seu in in J. Fac. Agric. Hokkaido Imp. Univ. Myrtilloides"; Koehne ，Deu t. Dendro l.: 89 ， Sapporo 26: 413 (1934) [Type: J apan ， 102 (1 893); C.K.Schneid. in Sarg. ， P l. Wils. Hokkaido. Prov. Ishikari ， in regione alpina 3: 152 (1 916) ， as “ Myrtilloides Borrer"; montis montis Daisetsusan. A. Kimura 1824 (♀)， 26 Nakai ， F l. Sylv. Kor. 18: 153 (1 930) ， as Ju l. 1927 (TUS-K)]. “Myrtilloides Borrer ex Loudon"; Skvortsov Sa !i x pauciflora var. cyclophylla Kimura in Acad. Sci. URSS Nov. Sys t. Pl. 1968: 65 in in Bo t. Mag. Tokyo 42: 575 (1 928) [Ty pe: (1 968) & Willows USSR: 124 (1968) ， ut Japan ，Hokkaido ， Prov. Ishikari ， in regione Myrtilloides Koehne ， sub subgen. Chamaetia alpina alpina montis Daisetsusan ，A. Kimura 1826 (Dumortier) Nasarov; Kitam. & Murata ， (♀)， 28 Ju l. 1927 (TUS-K)]. Co l. Il lus t. Woody Pl. Jap. 2: 305 (1 979) ， ut February February 2000 Journal of Japanese Botany Vo l. 75 No. 1 11

“ Myrtilloides Borrer" ，sub subgen. Salix; [Salix tribe Glaucae Fries in Physiogr. Argus in System. Bo t. Monogr. 52: 67 Sallsk.λrsb. 2: 35 (1 825) ，nom. illeg.]. (1 997) ，sub subgen. Chamaetia (Dumortier) Salix sec t. Ovalifoliae (Rydberg) Nasarov. Nasarov. Ty pe: Salix myrtilloides L. C.K.Schneid. in Sarg. ， P l. Wilson. 3: 140 Salix Salix [group] Myrtilloides Borrer in (1 916); Kitam. & Murata ， Col. Il lus t. Woody Loudon ， Arbor. Frutic. Bri t. 3: 1587 (1 838). Pl. J ap. 2: 305 (1979) ，ut “sec t. Ovalifoliae Rydberg" sub subgen. Salix; Argus in Sys- 5. 5. Salix fuscescens Andersson ，Monogr. tem. Bo t. Monog r. 52: 65 (1997) ，sub Salic.: Salic.: 97 (1 867) & in DC. ， Prodr. 16 (2): subgen. Chamaetia (Dumortier) Nasarov. 230 (1 868); Schneid. in Sarg. ， P l. Wils. 3: 153 (1 916); Kimura in Sci. Rep. Tohoku 6. Salix nakamurana Koidz. in Bo t. Univ. Univ. 4 ser. (Bio l.) 28: 40 (1 962); Skvortsov ， Mag. Tokyo 27: 96 (1 913) [Type: Japan. Willows USSR: 125 (1 968); Argus ， Salix Honshu. N agano Pre f. (Prov. Shinano) ， Alaska Yukon: 113 (1 973); Nedoluzhko in monte Orengezan (sphalm. Dailengezan) ， Charkevicz ， P l. Vascu l. Ori. Extr. Sovie t. 7: [Etigo-guti]. M. Nakamura s.n. (♀)， 4 Aug. 195 (1 995). 1912 (TI lecto. designated by A.Kimura Salix Salix paludicola Koidz. in Bo t. Mag. To- 1981)]; C.K.Schneid. in Sarg. ， P l. Wils. 3: kyo 33: 219 (1 919); Kimura in J. Fac. Agric. 135 (1 916); Kimura in Bo t. Mag. Tokyo 42: Hokkaido Imp. Univ. Sapporo 26: 425 574 (1 928) & in Sci. Rep. Tohoku Imp. (1 934); Ohwi ， F l. Jap. ed. Eng l.: 366 (1 965) Univ. 4 ser. (Bio l.) 2: 198 (1 931); Makino ， & F l. Jap. ed. rev.: 471 (1 965); Kitam. & Illus t. F l. Nippon: 675 ， fig. 2024 (1 940); Murata ，Co l. Il lus t. Woody P l. Jap. 2: 315 ， Ohwi ， F l. Jap. ed. Eng l.: 365 (1 965) & F l. fig. fig. 768 (1 979); T.Shimizu ，New Alp. F l. Jap. ed. rev.: 470 (1 965); Skvortsov ，Wil- Jap. Jap. 2: 110 ，fig. 319 (1 983); Kimura in Wild lows USSR: 136 (1 968); Kitam. & Murata ， Flow. Flow. Jap. Wood. 1: 47 ， photo 54 ，3 (1 989); Co l. Il lus t. Woody Pl. Jap. 2: 313 ，fig. 766 Ohwi & Kitag. ，New F l. Jap.: 536 (1 992). (1 979); T.Shimizu ，New Alp. Fl. Jap. 2: 114 ， Salix Salix arbutifolia auc t. non Pall.: Kimura fig. 322 (1 983); Kimura in Wild Flow. Jap. in in J. Fac. Agr. Hokkaido Imp. Univ. 26: 423 Wood. 1: 47 ， photo 54 ，1-2 (1 989); Makino ， (1 934) & in Sci. Rep. Tohoku Univ. 4 ser. New Il lus t. Fl. Jap. ed.: rev. fig. 106 (1 989); (Bio l.) 19: 196 (1 952). Ohwi & Kitag. ，New Fl. Jap.: 535 (1 992); Dist r.: Japan: Hokkaido ，Russia: Nedoluzhko in Charkevicz ， P l. Vascu l. Or i. Primorye ，Sakhalin ，Kamchatka ，Magadan Extr. Soviet. 7: 199 (1 995). and Siberia and North America: Yukon ， Salix nakamurana va r. stenophylla Alaska and Canada. Kimura in Bo t. Mag. Tokyo 42: 574 (1928) [Ty pe: Japan. Honshu. Nagano Pref. (Prov. 11 11 - ii i. Sect. Glaucae Andersson Shinano) ，Monte Yatsugatake. A. Kimura Salix Salix sec t. Glaucae Andersson in DC. ， 712 (♀ fl.) ， 12 Ju l. 1927 (TUS-K holo.)]. Prodr. Prodr. 16 (2): 273 (1 868); Pax in Eng l. & Salix nak αmurana f. stenophylla (Kimura) Prantl ，Natur l. Pflanzenfam. 3 (1): 37 Kimura in Eco l. Rev. 13: 200 (1 953) & in (1889) ，p.p.; Skvortsov in Acad. Sci. URSS Wild Flow. Jap. Wood. 1: 47 (1 989). Nov. Sys t. Pl. 1968: 65 (1 968) & Willows Salix nakamurana mstr. multic αpsularis USSR: 128 (1968) ，sub subgen. Chamaetia Kimura in Sci. Rep. Tohoku Univ. 4 ser. (Dumortier) (Dumortier) Nasarov; Argus in System. Bo t. (Bio l.) 18: 549 (1950) [Type: J apan. Monog r. 52: 68 (1997) ，sub subgen. Honshu. Nagano Pref. (Prov. Shinano) ， in Chamaetia (Dumortier) Nasarov. Ty pe: Salix regionibus alpinis montis Yatugatake ， inter glauca glauca L. saxa et rupes. A. Kimura 1243 (♀ fl.) ， 10 12 12 植物研究雑誌第75 巻第1号 平成12 年2月

Ju l. 1926 (TUS-K holo.)]. Kitag. ，New Fl. Jap.: 535 (1 992). 6-1. Salix nakamurana subsp. Distr.: J apan: Hokkaido (M t. Rakko in nakamurana 恥1t s. Hidaka) and Kuril Islands. Distr.: Distr.: Japan ，Central Honshu (Yamanashi 6-3. Salix nakamurana subsp. yezoalpina and and Nagano Prefs.). Rare on alpine slopes. (Koidz.) H.Ohashi ，sta t. nov. 6-1- 1. Salix nakamurana subsp. Salix cyclophylla Seemen in Bo t. Jahrb. nakamurana f. nakamurana Sys t. 30: Beib l. 67: 41 (1901) [可 pe: Japan.

6 司 1-2. Salix nak αmurana subsp. Jesso ，Gipfel des Rishiri. U. Faurie 3713 ， 25 nakamur αna f. eriocarpa (Kimura) Ju l. 1899-n.v.] & Salic. Jap.: 69 ，tab. 16 T. Shimizu ，New Alp. Fl. Jap. 2: 357 (1 983); (1 903) ，non Rydberg (1 899). Kimura in Wild Flow. Jap. Wood. 1: 47 Salix yezoalpina Koidz. in Bo t. Mag. To- (1 989). kyo 30: 332 (1 916); Kimura in 1. Fac. Agric. Salix Salix nakamurana va r. eriocarpa Kimura Hokkaido Imp. Univ. Sapporo 26: 407 in in Sci. Rep. Tohoku Imp. Univ. 4 ser. (Bio l.) (1 934); Ohwi ， F l. Jap. ed. Eng l.: 365 (1 965) 6: 6: 189 (1931) [Type: J apan. Honshu. & Fl. Jap. ed. rev.: 470 (1 965); Kitam. & N agano Pre f. (Prov. Shinano) ，monte Murata ，Co l. Illus t. Woody Pl. Jap. 2: 313 ， Yatsugatake. Yatsugatake. A. Kimura no. 712 (♀ fl.) ， 12 fig. 767 (1 979); T. Shimizu ，New Alp. Fl. Ju l. 1927 (TUS-K holo.)]; Ohwi & Kitag. ， Jap. 2: 113 ，fig. 320 (1 983); Kimura in Wild New Fl. Jap.: 535 (1 992). Flow. Jap. Wood. 1: 48 ，photo 54 ，5-6 Distr.: Distr.: J apan. Occurs with the typical (1 989); Makino ，New Il lus t. Fl. Jap. rev. ed.: form. form. fig. 92 (1 989); Ohwi & Kitag. ，New F l. Jap.: 6-2. 6-2. Salix nakamurana subsp. kurilensis 535 (1 992). (Koidz.) (Koidz.) H.Ohashi ， comb. & sta t. nov. Salix nakamurana va r. yezo α lpina Salix Salix kurilensis Koidz. in Bo t. Mag. To- (Koidz.) Kimura in Bo t. Mag. Tokyo 42: kyo 32: 62 (1 918) [Type: Kuriles. Ins. 574 (1 928). Shumushu. K. Yendo s.n. (♀日.)， 29 Jun. Salix yezoalpina f. grandifolia (N akai) 1903 1903 (TI holo.)]; Kimura in J. Fac. Agric. Kimura in Eco l. Rev. 13: 201 (1 953) & in Hokkaido Imp. Univ. Sapporo 26: 406 Wild Flow. Jap. Wood. 1: 48 (1 989). (1934); (1934); Skvortsov ，Willows USSR: 136 Salix yezoalpina f. stenophylla (Nakai) (1 968); Nedoluzhko in Charkevicz ， P l. Kimura in Eco l. Rev. 13: 201 (1953) & in Vascu l. Ori. Extr. Sovie t. 7: 199 (1 995). Wild Flow. Jap. Wood. 1: 48 (1 989). Salix Salix longepetiolata Flod. in Ark. Bo t. Distr.: Japan: Hokkaido (M t. Taisetsu and 20A ， 6: 14 (1 926). Is l. Rishiri) ， endemic. On rocky slopes. Salix Salix hidakamontana H.Hara in J. Jap. 6-3- 1. Salix nαkamurana subsp. Bo t. 9: 512 ，fig. 1 (1 933) ，as “S. hidaka- yezoalpina f. yezoalpina (Koidz.) H.Ohashi ， montana" [Type: J apan. Hokkaido. Prov. comb. nov. Hidaka ， in summo monte Rakko. H. Hara 6-3-2. Sα lix nakamur αna subsp. 2994 (fr.) ，8 Aug. 1933 (TI holo.)]; Kimura yezoalpina f. neoreticulata (Nakai) in in J. Fac. Agric. Hokkaido Imp. Univ. H.Ohashi ， comb. nov. Sapporo Sapporo 26: 408 (1 934); Ohwi ， F l. Jap.: 402 Salix neoreticulata Nakai (1 953); Ohwi ， F l. Jap. ed. Eng l.: 365 (1 965) [Tennenkinenbutsu Chosahokoku ，Bo t. 12 & F1. Jap. ed. rev.: 470 (1 965); Kitam. & (Rep. Veg. Daisetsusan Mts.): 45 ， 1930] ex Murata ，Co l. Il lust. Woody Pl. Jap. 2: 314 Kimura in J. Fac. Agr. Hokkaido Imp. Univ.

(1 979); T. Shimizu ，New Alp. Fl. Jap. 2: 113 ， 26 (4): 408 (1 934) ，as “neo 四 reticulata" fig. fig. 321 (1 983); Kimura in Wild Flow. Jap. [Type: Hokkaido. Prov. Ishikari ， Wood. 1: 48 ， photo 54 ，4 (1 989); Ohwi & Takanegahara ，monte Daisetsu. T. N akai February February 2000 Journal of Japanese Botany Vo l. 75 No. 1 13

s.n. ，Aug. 1928 (TI holo.)]; Ohwi ， F l. Jap.: Fl. Germ. Helv. 2: 644 (1 837); Skvortsov in 402 (1953). Acad. Sc i. URSS Nov. Sys t. Pl. 1968: 64 Salix Salix yezoalpina va r. neoreticulata (1 968) & Willows USSR: 99 (1 968) ，sub (Nakai) Kimura in Eco l. Rev. 13: 201 subgen. Salix; Kitam. & Murata ，Co l. Illus t. (1 953); Kitam. & Murata ，Co l. I1 1us t. Woody Woody Pl. Jap. 2: 307 (1 979) ， sub subgen. l. Pl. Jap. 2: 314 (1979); Kimura in Wild Flow. Salix; Chou in Fl. Reipub. Pop. Sin. 20 (2): Jap. Jap. Wood. 1: 48 (1 989). 120 (1984); Newsholme ，Willows: 24 Salix Salix yezoalpina f. neoreticulata (Nakai) (1 992). Ty pe. Salix triandra L. T.Shimizu ，New Alp. Fl. Jap. 2: 357 (1 983). Distr.: Distr.: Japan: Hokkaido (M t. Taisetsu). 7. Salix triandra L. ， Sp. P l.: 1016 (1 753); Chou in Fl. Reipub. Pop. Sin. 20 111. 111. Subgen. Salix (2): 121 (1 984); Sohma in Sci. Rep. Tohoku Salix Salix subgen. Salix: Skvortsov in Acad. Univ. 4 ser. (Bio l.) 40: 107 (1 993). Sci. Sci. URSS Nov. Sys t. Pl. 1968: 63 (1 968) ， Salix amygdalina L. ， Sp. Pl.: 1016 (1753). p.p. p.p. exc l. sec t. Humboldtianae & Salix amygdalina va r. discolor Wimmer Urbanianae & Willows USSR: 97 (1968) ， & Grabowski ， F l. Siles. 3: 362 (1 829); p.p. p.p. exc l. subgen. Protitea & genus Toisusu; Trautv. in Maxim. ， Prim. F l. Amur.: 242 Dorn in Can. J. Bo t. 54: 2774 (1 976) ， p.p. (1 859). exc l. subgen. Pleiolepis & Protitea; Argus in Salix subfragilis .f¥_ ndersson in Mem. Sys t. Bo t. Monog. 52: 50 (1 997) ，p.p. exc l. Amer. Acad. 6 (2): 450 (1 858) [Ty pe: Japan. subgen. subgen. Pleiolepis & Protitea; Newsholme ， Hokkaido ，ad Hakodate (ut Hakodadi). Wil- Willows: 22 ，23 (1 992) ， p.p. Type: Salix liams & Morrow (not in GH)-n.v.]: Kimura alba alba L. in Acta Phytotax. Geobot. 13: 188 (1 944); Salix Salix sec t. Amerina Dumortier ， Verh. Ohwi ， F l. Jap. ed. Eng l.: 364 (1 965) & F l. Ges l. Wilgen: 15 (1 825). Jap. ed. rev.: 468 (1 965); Kurata ，Illus t. Imp. Salix Salix subgen. Amerina (Dumortier) For. Tr. Jap. 3: 4， p l. 4 (1 971); Sugaya in Dumortier in Bull. Soc. Roy. Bo t. Belg. 1: Terasaki. Il lus t.: 98 ， fig. 383 (1977); 145 145 (1 862). Ty pe: Salix alb αL. Hatusima ，Kagoshima-ken Shokubutsu Salix Salix subgen. Euitea Kimura in Bo t. Mag. Mokuroku: 35 (1 978); Kitam. & Murata ， Tokyo 42: 290 (1928) & in Sc i. Rep. Co l. Illus t. Woody Pl. Jap. 2: 332 ，fig. 790 Tohoku Imp. Univ. 4th ser. Bio l. 13: 394 (1 979); Lauener & McKean in Notes RBG (1 938). Ty pe: Salix pentandra L. Edinb. 41: 178 (1 983); Kimura in Wild Salix Salix subgen. Calypterolepis Nakai in Flow. Jap. Wood. 1: 39 ，photo 43 ，4-6 Bull. Bull. Soc. Dendro l. Fr. 66: 43 (1 928). Ty pe: (1 989); Makino ，New Illus t. Fl. Jap. rev. ed.: Salix Salix pentandra Seemen. fig. 89 (1 989); Ohwi & Kitag. ，New Fl. Jap.: 533 (1 992); Sohma in Sci. Rep. Tohoku III-i. III-i. Sec t. Tr iandrae Dumortier Univ. 4 ser. (Bio l.) 40: 107 (1 993). Salix Salix sec t. Tr iandrae Dumortier in B りdr. Salix triandra var. discolor (Wimmer & Na t. Wetens. 1: 58 (1 825) & in Bul l. Soc. Grabowski) Andersson in DC. ，Prodr. 12 (2): Roy. Bo t. Belg. 1 ， 1: 146 (1862); 203 (1 868); Nakai ， F l. Sylv. Kor. 18: 87 C.K.Schneid. in Sarg. ， P l. Wils. 3: 106 (1 930); Makino ，Il lus t. Fl. Nippon: 671 ，fig. (1 916); Nakai in Bull. Soc. Dendro l. Fr. 66: 2011 (1 940). 9 (1 928) & F l. Sylv. Kor. 18: 85 (1 930). Salix nipponica Franch. & Sav. ，Enum. Ty pe. Salix triandra L. F l. Pl. Jap. [1: 459 (1 875) ，nom. nud.] 2: Salix Salix sec t. Amygdalinae Koch ， Salic. 502 (1 876) [Type: Nippon media. circa Europ. Europ. Commen t.: 11 & 17 (1828) & Syn. Yokoska. Savatier 1139 (P syn.-n. v.); 14 14 植物研究雑誌第75 巻第1号 平成12 年2 月

Niigata ， Savatier 2717 (P syn.-n. V. ，LE (1944) as identical with S. nipponic αor by isosyn. isosyn. fide Skvortsov)]; Tokubuchi in Bo t. Skvortsov (1 968b) as S. babylonica. Unfor- Mag. Tokyo 10: page (122) (1896); tunately the type specimen is not found in Nedoluzhko in Charkevicz ， P l. Vascu l. Ori. GH ，although so designated in the Extr. Extr. Sovie t. 7: 166 (1995). protologue of S. subf 1i αgilis. Salix Salix triandra L. var. nipponica (Franch. & Sav.) Seemen ， Salic. Jap.: 27 ， tab. 2 III-ii. Sec t. Subalbae Koidz. (1 903); Kimura in J. Fac. Agric. Hokkaido Salix sec t. Subalbae Koidz. in Bo t. Mag. Imp. Imp. Univ. Sapporo 26: 401 (1 934); Chou in Tokyo 27: 88 (1 913) ， in adno t.; Skvortsov in l. Fl. Reipub. Pop. Sin. 20 (2): 121 (1 984). Acad. Sci. URSS Nov. Sys t. Pl. 1968: 64 Salix Salix triandra H.Lev. & Vaniot in Bull. (1 968) & Willows USSR: 113 (1 968) ， sub Soc. Soc. Bo t. Fr. 52: 141 (1 905): 143 (1 906) subgen. Salix; Kitam. & Murata ，Co l. Illus t. [Ty pe: Japan. Honshu. Aomori ，May 1902. Woody Pl. Jap. 2: 307 (1979) ， sub subgen. Kinashi Kinashi 11 (E holo.-n.v.)]. Salix; Argus in System. Bo t. Monogr. 52: 52 Salix Salix hamatidens H.Lev. in Bul l. Soc. (1 997). Type. Salix hondoensis Koidz. (= Bo t. F r. 56: 301 (1909) [Type: J apan. Salix pierotii Miq.). Hokkaido ， secus aquas Sapporo ， 13 June Salix [unranked] Subfragiles Seemen ， 1908. 1908. U. Faurie 263 (E syn.-n.v.) ， U. Faurie Salic. Jap.: 15 (1903) ，sub Salix A. 264 (E syn.-n.v.)]. Didymadeniae b. Diandrae. Salix Salix amygd α lina var. nipponica (Franch. Salix sec t. Subfragiles Nakai ， F l. Sylv. & Sav.) C.K.Schneid. in Sarg. ， P l. Wils. 3: Kor. 18: 163 (1 930); Kimura in Sci. Rep. 106 106 (1 916). Tohoku Univ. 4 ser. (Bio l.) 28: 43 (1 962). Salix Salix triandra L. subsp. nipponica Type. Salix subfragilis L. (= Salix (Franch. (Franch. & Sav.) Skvortsov ， Willows USSR: babylonica L.). 100 100 (1 968). Salix sect. Fragiles Koch: Newsholme ， Distr.: Distr.: Japan: Hokkaido ，Honshu ， Shikoku Willows: 24 (1 992) ，p.p. inc l. S. matsudana. and Kyushu ，Korea ，NE. & N. China and Salix sec t. Albae Borrer: C.K.Schneid. in Russia: Russia: Sakhalin ，Khabarovsk ，Primorye ， Sarg. ， P l. Wils. 3: 109 (1916) ，p.p. Amur and Siberia to Europe. Salix sec t. Salix: Chou in Fl. Reipub. Pop. Note: Note: The Japanese plants have been re- Sin. 20 (2): 122 (1 984) ，p.p.; Newsholme ， garded garded as different from Salix triandra as a Willows: 24 (1992) ， p.p. inc l. sec t. distinct distinct species or an infraspecific taxon. Subalbae. Comparing them with specimens of Salix triandra triandra from Europe no definite differences 8. *Salix babylonica L. ， Sp. Pl.: 1017 are are found ， although tendency in variation is (1 753); Franch. & Sav. ，Enum. Pl. Jap. 1: seen seen in persistency of stipules ， degree of 459 (1 875); Seemen ，Salic. Jap.: 29 ，tab. 3 development of leaves on (1 903); C.K. Schneid. in Sarg. ， P l. Wilson. branchlets branchlets and persistency of hairs on young 3: 42 (1 916); Makino ，Il lus t. Fl. Nippon: leaves leaves and branchlets. Morphological varia- 673 ，fig. 2018 (1 940); Ohwi ， F l. Jap. ed. tion tion seems to be continuous within the ar- Eng l.: 364 (1 965) & Fl. Jap. ed. rev.: 468 eas eas of distribution from Europe to Asia and (1 965); Skvortsov ，Willows USSR: 114 Japan. Japan. Salix triandra L. is used for the Japa- (1 968); Sugaya in Terasaki. Il lus t.: 100 ，fig. nese nese plants. 386 (1 977); Chou in Fl. Reipub. Pop. Sin. Salix Salix subfragilis Andersson was described 20 (2): 138 (1 984); Nova Fl. Nasu. Supp l.: based on a female twig collected in 112 (1 985); Kimura in Wild Flow. J ap. Hokkaido. It was regarded by Kimura Wood. 1: 41 ， photo 46 ， 1-2 (1 989); Makino ， February February 2000 Journal of Japanese Botany Vo l. 75 No. 1 15

New Illus t. F1. Jap. rev. ed.: fig. 99 (1 989); Distr.: Cultivated. Ohwi & Kitag. ，New F1. Jap.: 533 (1 992). Salix Salix pendula 恥10ench ，Meth.: 336 9. *Salix matsudana Koidz. in Bo t. Mag. (1 794). Tokyo 29: 312 (1 915); Chou in F1. Reipub. Sα lix babylonica var. lavallei Dode in Pop. Sin. 20 (2): 132 (1 984). Bul l. Soc. Dendr. Fr. no. 76: 93 (1 930); var. tortuosa Vilm. in J. Soc. Na t. Hor t. Kimura in Sc i. Rep. Tohoku Univ. 4 ser. Fr. ser. 4， 25: 350 (1 924); Ohwi ， F l. Jap.: (Bio 1.) 24: 115 (1958) & in J. Jap. Bo t. 53: 401 (1 953); Sugaya in Terasaki. Illus t.: 99 ， 193 193 (1 978); Kitam. & Murata ，Co 1. Il lus t. fig. 384 (1 977); Kitam. & Murata ，Co l. Woody P1. Jap. 2: 328 ，fig. 786 (1 979). Il lus t. Woody P l. J ap. 2: 327 ， fig. 784 Sα lix ohsidare Kimura in Bo t. Mag. To- (1 979); Kimura in Wild Flow. Jap. Wood. 1: kyo 59: 79 (1946) [Type: Japan. Aomori 41 ， photo 46 ，3 (1989). Pre f. (Prov. Mutsu). Aomori ，cul t. A. Salix matsudan αf. tortuosa Rehder in 1. Kimura 2166 (♀ fl.) ，9 May 1933 (TU ふK Arn. Arb. 6: 206 (1 925); Ohwi ， F 1. Jap. ed. lecto. lecto. desiganted by Kimura 1981)]; Kimura rev.: 468 (1 965); Chou in F1. Reipub. Pop. in in Sci. Rep. Tohoku Univ. 4 ser. (Bio 1.) 24: Sin. 20 (2): 133 (1 984); Ohwi & Kitag. ， 113 ，fig. 1 ，tab. 1-2 (1 958); Ohwi ， F 1. Jap.: New F1. Jap.: 533 (1 992).

401 401 (1 953) & F1. Jap. ed. Eng 1.: 365 (1 965); Salix matsud α， nαcv. Tortuosa: Ohwi ， F 1. Ohwi & Kitag. ，New F1. Jap.: 534 (1 992); Jap. ed. Eng 1.: 364 (1 965). Sohma in Sci. Rep. Tohoku Univ. 4 ser. Distr.: Japan ，cultivated in a park or gar- (Bio 1.) 40: 119 (1 993). den for ornamental in Japan. Female plants Distr.: Distr.: Japan: Widely cultivated ， imported are rare; China and Korea. Native in North from Korea or China in old time. Probably China. spontaneous spontaneous in central China. 8-1. 8-1. * Salix babylonica f. babylonica 10. Salix eriocarpa Franch. & Sav. ， 8-2. 8-2. *Salix babylonica f. rokkaku Kimura Enum. P1. Jap. [1: 459 (1 875) ，nom. nud.] 2: in in J. Jap. Bo t. 53: 193 ， figs. 1-4 (1 978) 503 (1 876) [Type: Nippon media. circa [Ty pe: J apan. Miyagi Pre f. (Prov. Rikuzen) ， Hemi prope Yokoska. Savatier 2718 (P- Sendai ，cul t. in Hor t. Bo t. Fac. Sci. Tohoku n.v.)]; Ohwi ， F l. Jap. 401 (1 953) ，1. c. ed. Univ. Univ. A. Kimura 3417 (fo 1.)， 11 Nov. 1968 Eng 1.: 364 (1 965) & F1. Jap. ed. rev.: 468 (TUS-K holo.)]; Sohma in Sc i. Rep. Tohoku (1 965); Chou in Fl. Reipub. Pop. Sin. 20 Univ. Univ. 4 ser. (Bio 1.) 40: 123 (1 993). (2): 141 (1 984); Kimura in Wild Flow. Jap. Sα lix elegantissima auc t. non K.Koch: Wood. 1: 40 (1 989); Ohwi & Kitag. ，New Nakai ， F l. Sylv. Korea. 18: 163 & 171 F1. Jap.: 533 (1 992). (1 930); Ohwi ， F 1. Jap. ed. Eng 1.: 364 (1 965). Salix dolichostyla Seemen in Bo t. Jahrb. Distr.: Distr.: Cultivated. Sys t. 30 ，Beib 1. 67: 39 (1 90 1) & Salic. Jap.: 8-3. 8-3. *Salix babylonica f. seiko Kimura [in 26 ，tab. 2 (1 903). Ishii ，Engei daiziten 4: 2429 (1956) ，nom. Distr.: Japan ， mostly Kanto Districts and nud.] nud.] in Sci. Rep. Tohoku Univ. 4 ser. westward (introduced ?)， Korea and China. (Bio l.) 24: 115 ，fig. 2 (1 958) [Ty pe: Japan. Note: This species is recognized as native Honshu ，Kyoto Pref. (Prov. Yamashiro) ，Uji ， in Japan. Kimura (1 989) supposed ，however ， cul t. A. Kimura 3157 (fo 1.)， 19 Oc t. 1955 this was imported into Japan in an- (TUS-K holo.)]; Seibutsugaku- cient time ， because no male plants found in gokenkyusho: gokenkyusho: F1. Sedis Imperatoris Japon.: Japan and it found usually growing within 186 ，figs. 81-83 (1 989); Sohma in Sci. Rep. living territories of human beings. Tohoku Univ. 4 ser. (Bio 1.) 40: 125 (1 993). 16 16 植物研究雑誌第75 巻第1号 平成12 年2月

11. 11. Salix pierotii Miq. ，Ann. Mus. Lugd.- Japan ， Aichi Pref. (Prov. Mikawa) ，cul t. in Ba t. 3: 27 (1 867) & Prolusio F1. Jap. 4: 215 Kamekubityo urbis Toyota ， K. Kitagawa s.n. (1 867) [Ty pe: Japan. in Monte Iwaja (vel (♀ f1.)， 2 Apr. 1982 (TUS-K holo.)]. Iwajama) ，ubi frequens in ungis et in Distr.: Cultivated. vallibus ，m. Aprili florentem detexit Siebold (L ?-n.v.); in promontorio Nomo-Saki 12. Salix yoshinoi Koidz. in Bo t. Mag. insulase insulase Kiusiu. Pierot 181 (L syn.)*]; To ky 0 29: 3 14 (1 915) [Ty P e: Japan ， C.K.Schneid. in Sarg. ， P l. Wils. 3: 165 Honshu， Okayama Pref. (Prov. Bittyu) ， (1 916); Koidz. in Bo t. Mag. Tokyo 40: 346 Kawakamigori ，Ae (errore Abe) ， Z. Yoshino (1 926); Skvortsov ，Willows USSR: 113 300 (♀ fl.) ，24 Apr. 1914 (TI holo.)] & in (1968) ，p.p. ，exc l. S. jessoensis ， S. Bo t. Mag. Tokyo 30: 332 (1 916); Ohwi ， F 1. hondoensis & S. hirosakensis; Kitam. & Jap. ed. Eng l.: 364 (1 965) & Fl. Jap. ed. Murata ，Co 1. Il lus t. Woody P1. Jap. 2: 328 ， rev.: 469 (1 965); Kitam. & Murata ，Co l. fig. fig. 785 (1979); Kimura in J. Jap. Bo t. 59: Il lus t. Woody P l. Jap. 2: 330 ， fig. 788 1 (1 984); Chou in F1. Reipub. Pop. Sin. 20 (1 979); Kimura in Wild Flow. Jap. Wood. 1: (2): (2): 147 (1 984); Kimura in Wild Flow. Jap. 40 ， photo 50 ，5-7 (1 989); Ohwi & Kitag. ， Wood. 1: 39 ， photo 44 ，1-4 (1 989); Sohma New Fl. Jap.: 534 (1 992). in in Sci. Rep. Tohoku Univ. 4 ser. (Bio 1.) 40: Salix pseudoyoshinoi Koidz. in Acta 123 & 134 (1993); Nedoluzhko in Phytotax. Geobo t. 4: 40 (1 935) [Ty pe: Ja- Charkevicz ， P 1. Vascu 1. Ori. Extr. Sovie t. 7: pan ，Honshu ，Okayama Pref. (Prov. Bittyu) ， 169 (1995). *The type composed of 2 Matsuyamamura ，Awoki ， Z. Yoshino s.n. branchlets; branchlets; one with young (ster.) ， Ju 1. 13 ，1926 (KYO lecto. designated and the other with somewhat young leaves. by A. Kimura 1981)]. The two branchlets belong to the same spe- Salix yoshinoi f. pseudoyoshinoi (Koidz.) cles. cles. Kimura in Wild Flow. Jap. Wood. 1: 40 Salix Salix koreensis Andersson in DC. ， Prodr. (1 989). 16 (2): 271 (1 868); Koidz. in Bo t. Mag. To- Distr.: Endemic to J apan: Honshu (Kinki kyo 27: 89 (1 913) & 30: 332 (1 916); Nakai ， and Chugoku Districts) and Shikoku. 1. F1. Sylv. Kor. 18: 164 (1 930); Ohwi ， F 1. Jap. ed. ed. Eng l.: 365 (1 965) & Fl. J ap. ed. rev.: 13. Salix jessoensis Seemen ，Salic. Jap.: 469 (1 965); Sugaya in Terasaki. Il lus t.: 98 ， 31 ， tab. 3 (1 903) [Ty pe: Japan. Hokkaido ， fig. fig. 385 (1 977); Lauener & McKean in Sapporo ，Tokubuchi ，1891 (SAPT holo.)]; Notes Notes RBG Edinb. 41: 177 (1983); Chou in Miyabe & Kudo ， Icon. Ess. Fo r. Trees l. Fl. Reipub. Pop. Sin. 20 (2): 143 (1 984); Hokkaido 1: 49 ，tab. 14 (1 921); Kimura in Ohwi & Kitag. ，New F1. Jap.: 534 (1 992). J. Fac. Agric. Hokkaido Imp. Univ. Sapporo Salix Salix pseudokoreensis Koidz. in Bo t. 26: 402 (1 934); Ohwi ， F l. Jap. ed. Eng l.: Mag. Tokyo 40: 346 (1926) [Type not des- 365 (1 965) & F1. Jap. ed. rev.: 469 (1 965); ignated]. ignated]. Kitam. & Murata ，Co l. Il lus t. Woody P1. Jap. Distr.: Distr.: Japan: SW. Hokkaido ，Honshu 2: 331 ，fig. 789 (1979); Kimura in Wild (Hokuriku ，Chugoku and Kinki Districts) ， Flow. J ap. Wood. 1: 40 ，photo 45 ，5-7 Shikoku and Kyushu ，Korea and Far East (1 989); Ohwi & Kitag. ，New F1. Jap.: 534 Russia: Russia: Amur ，U ssuri ，Khabarovsk and (1 992); Sohma in Sci. Rep. Tohoku Univ. 4 Primorye. Primorye. ser. (Bio 1.) 40: 117 (1 993). 11- 1. Salix pierotii f. pierotii Salix eriocarpa auc t. non Franch. & Sav.: 11-2. 11-2. *Salix pierotii f. auricomans Tokubuchi in Bo t.島 1ag. Tokyo 10: page Kimura in J. Jap. Bo t. 59: 1 (1 984) [Ty pe: (1 22) (1 896). February February 2000 Journal of Japanese Botany Vo l. 75 No. 1 17

Salix Salix dolichostyl αva r. hirosakensis F l. Europ. 1: 47 (1964) ，as “subgen. H.Lev l. in Reper t. Spec. Nov. Regni Veg. 3: Caprisalix Dumortier"; Newsholme ，Wil- 22 (1 907). lows: 22 & 24 (1 992). Ty pe: Salix caprea L. Salix Salix hondoensis Koidz. in Bo t. Mag. Tokyo 27: 88 (1 913) [Ty pe: Japan. Honshu. IV- i. Sec t. Hastatae A.Kerner Yamagata Pre f. (Prov. Uzen) ， Salix sec t. Hastatae A.Kerner in Verh. Minamiokitama-gun ， Seki-no-sanson. G. Zoo l. -Bo t. Ges. Vereins Wien 10: 241 Koidzumi s.n. (♀ fl.) ，May 1910 (TI-lecto. (1 860); C.K.Schneid. in Sarg. ， P l. Wils. 3: designated designated by A. Kimura 1981)]. 129 (1 916); Skvortsov in Acad. Sc i. URSS Salix Salix hiros α kensis (H.Lev l.) Koidz. in Nov. Sys t. Pl. 1968: 66 (1968) & Willows Bo t. Mag. Tokyo 27: 264 (1 913). USSR: 147 (1 968); Kitam. & Murata ，Co l. 13-1. 13-1. Salix jessoesis subsp. jessoensis Il lust. Woody Pl. Jap. 2: 306 (1979) ，sub Endemic to Japan: Hokkaido and Honshu. subgen. Salix; Argus in System. Bo t. 13-2. 13-2. Salix jessoesis subsp. serissaefolia Monogr. 52: 70 (1997). Ty pe: Salix hastata (Kimura) (Kimura) H. Ohashi ，comb. nov. L. Salix Salix serissaefolia Kimura in Bo t. Mag. Salix sec t. Cordatae Bar r. ex Hook.: Tokyo 40: 639 (1926) [Type: J apan. Newsholme ， Willows: 25 (1 992) ， p.p. inc l. Honshu. Shizuoka Pref. (Prov. Suruga) ， sec t. Hastatae. prope Subashiri ，pede Fujisan. A. Kimura [Salix tribe Hastatae Fries ， Physiogr. 197 197 (♀日.)， 12 May 1924 (TI holo.; TUS- Salls k. Arsb. 2: 34 (1825) ，nom. 日leg.]. K iso.)]; Ohwi ， F l. J ap. ed. Eng l.: 364 (1 965) & Fl. Jap. ed. rev.: 469 (1 965); Nova 14. Salix japonica Thunb. in Murray ， l. Fl. Nasu.: 143 ，figs. 68-70 (1 972); Sugaya Sys t. Veg. ed. 14: 879 (1 784) ， F l. Jap.: 24 in in Terasak i. Illus t.: 100 fig. 387 (1 977); .(784) [Ty pe: Japan. Thunberg (UPS-n.v.)] ， Kitam. Kitam. & Murata ，Co l. Il lus t. Woody Pl. Jap. Icon. Pl. J ap. 4: tab. 1 (1802) ，& Pl. J ap. 2: 2: 330 ， fig. 787 (1 979); Kimura in Wild Nov. Sp.: 4 (1 824); Franch. & Sav. ，Enum. Flow. Jap. Wood. 1: 40 ，photo 45 ，1-4 Pl. Jap. 1: 459 (1875); Koidz. in Bo t.島 1ag. (1 989); Makino ，New Illus t. Fl. Jap. rev. ed.: Tokyo 27: 90 (1 913) ，cum f. typica Koidz.; fig. fig. 100 (1989); Ohwi & Kitag. ，New Fl. Seemen ， Salic. Jap.: 43 ，tab.7 ，figs. A-E Jap.: Jap.: 534 (1992); Sohma in Sci. Rep. (1 903); C.K.Schneid. in Sarg. ， P l. Wils. 3: Tohoku Univ. 4 ser. (Bio l.) 40: 126 (1993). 132 (1 916); Kimura in Sci. Rep. Tohoku Distr.: Distr.: Endemic to Japan: Honshu (Kanto Imp. Univ. 15: 415 (1940); Makino ，Il lus t. to to Kinki Districts). Fl. Nippon: 674 ，fig. 2021 (1 940); Kimura in in Eco l. Rev. 13: 135 (1 953); Ohwi ， F l. Jap. IV. IV. Subgen. Vetrix Dumortier ed. Eng l.: 367 (1 965) & F l. Jap. ed. rev.: Salix Salix subgen. Vetrix Dumortier in Bu l1. 473 (1 965); Kurata ，Il lus t. Imp. For. Tr. Jap. Soc. Soc. Bo t. Belg. 1: 141 (1 862); Skvortsov in 4: 1 ，p l. 1 (1 973); Sugaya in Terasak i. Illus t.: Acad. Sc i. URSS Nov. Sys t. Pl. 1968: 66 102 ，fig. 397 (1 977); Kitam. & Murata ，Co l. (1 968) & Willows USSR: 147 (1 968); Dorn Il lus t. Woody P l. Jap. 2: 325 ， fig. 782 in in Can. J. Bo t. 54: 2776 (1 976); Argus in (1 979); Kimura in Wild Flow. Jap. Wood. 1: System. System. Bo t. Monogr. 52: 70 (1 997). Ty pe: 45 ，photo 52 ，1-4 (1 989); Makino ，New Salix Salix caprea L. Illus t. F l. J ap. rev. ed.: fig. 103 (1 989); Salix Salix subgen. Chamaetia (Dumortier) Ohwi & Kitag. ，New Fl. Jap.: 538 (1 992); Nasarov ， F l. URSS 5: 31 (1936). Sohma in Sci. Rep. Tohoku Univ. 4 ser. Salix Salix subgen. Caprisalix (Dumortier) (Bio l.) 40: 119 (1993). Nasarov ， F l. URSS 5: 57 (1 936); Rech. f. in Salix p αdifolia Andersson in Mem. Am. 18 18 植物研究雑誌第75 巻第l号 平成 12 年2月

Acad. 6: 451 (1 859) ， in Svensk. Vetensk. Salix babylonica auc t. non L.: Thunb. in Akad. Hand 1. 6: 165 (1 867) & in DC. ， Prodr. Nova Acta Reg. Soc. Scien t. Upsa 1. 3: 207 16 (2): 255 (1 868). (1 780). Salix Salix babylonica var. japonica Andersson Distr.: Endemic to Japan: Honshu (south- in in Svensk. Vetensk. Akad. Hand 1. 6: 51 ern Tohoku ，Kanto and Chubu Districts). (1 867) & in DC. ， Prodr. 16 (2): 213 (1 868). Salix Salix oldhami αna Miq. ，Ann. Mus. Ludg.- 15. Salix shiraii Seemen in Bo t. J ahrb. Ba t. 3: 25 (1 867). Sys t. 30: Beib 1. 67: 40 (1901) & Salic. Jap.: Salix Salix japonica va r. oldhamian α(Miq.) 42 ，tab. 6 (1903); C.K.Schneid. in Sarg. ， P 1. Franch. & Sav. ，Enum. P 1. Jap. 1: 459 Wils. 3: 135 (1 916); Ohwi ， F 1. Jap. ed. (1 875). Eng l.: 367 (1 965) & Fl. Jap. ed. rev.: 472 Salix Salix japonica var. pygmaea Franch. & (1965); Sugaya in Terasaki. Il lus t.: 102 ，fig. Sav. ，Enum. P 1. Jap. 1: 459 (1 875) [Ty pe: 396 (1 977); Kitam. & Murata ，Co 1. Illus t. Japan. Japan. circa Yokoska. Savatier (P-n.v.)] & 2: Woody P l. J ap. 2: 324 ， fig. 780 (1 979); 503 (1 913). Kimura in Wild Flow. Jap. Wood. 1: 45 ， Sα lix fauriei Seemen in Bo t. Jahrb. Sys t. photo 50 ，5-7 (1 989); 加lakino ，New Illus t. 30: 30: Beib 1. 67: 40 (190 1) & Salic. Jap.: 48 ， F l. Jap. rev. ed.: fig. 104 (1 989); Ohwi & tab. tab. 8 (1903) [Type: Japan. Honshu ， Kitag. ，New F1. Jap.: 538 (1 992); Sohma in Kanagawa Pre f. (Prov. Sagami) ， Yamakita. Sci. Rep. Tohoku Univ. 4 ser. (Bio l.) 40: 119 U. Faurie 3700 p.p. (♀ fl.) ，8 May 1899 (1993). (KYO iso.)]; C.K.Schneid. in Sarg. ， P 1. Salix kenoensis Koidz. in Bo t. Mag. To- Wils. Wils. 3: 134 (1 916). kyo 27: 265 (1913) [Ty pe: Japan. Honshu ， Salix Salix japonica var. pad 扮li α(Andersson) Saitama Pref. (Prov.. Musashi) ，Chichibu ， Seemen ， Salic. Jap.: 45 ， tab. 8 ，figs. A-B M t. Buko. S. Matsuda 104 (fr.). May 1901 (1 903). (TI lecto. selected by Kimura in Sci. Rep. Salix Salix japonica f. padifoli α(Seemen) Tohoku Univ. 4 ser. (Bio l.) 28: 37. 1962)]; Koidz. Koidz. in Bo t. Mag. Tokyo 27: 91 (1 913). Kitam. & Murata ，Co l. Il lus t. Woody Pl. Jap. Salix Salix japonica f. oldhamian α(Franch. & 2: 310 (1 979). Sav.) Sav.) Koidz. in Bo t. Mag. Tokyo 27: 91 Salix shiraii va r. kenoensis (Koidz.) (1 913) ，as ‘'f. oldhami". Sugimoto ，New Keys Jap. Tr.: 472 (1 961) & Salix Salix japonica f. pygmaea (Franch. & New Keys Woody Pl. Japan: 98 (1 972). Sav.) Sav.) C.K.Schneid. in Sarg. ， P l. Wils. 3: 132 Distr.: Endemic to Japan: Honshu (North- (1 916). ern Kanto and southern Tohoku Districts). Salix Salix japonica var.α ngustifolia Kimura in Bo t. Mag. Tokyo 42: 573 (1 928) [Ty pe: Ja- 16. Salix rupifraga Koidz. in Bo t. Mag. pan. pan. Honshu ，Tokyo Pref. (Prov. Musashi) ， Tokyo 33: 121 (1919) [Type: J apan ， Mitamura-Sawai. K. Hisauchi 2 (fo 1.)， 20 Honshu ，Yamanashi Pref. (Prov. Kai) ， 1. Ju 1. 1919 (TU ふK holo.)]. Komagadake ， [Shinshu Higashi- Salix Salix japonica var.α 時 ustifolia f. fauriei Komagadake]. G. Koidzumi s.n. (fo 1. cum (Seemen) Kimura in Sc i. Rep. Tohoku Imp. 仕.) (TI holo. ，TU ふK iso.)]; Kimura in Sci. Univ. Univ. 15: 416 (1 940). Rep. Tohoku Imp. Univ. 4 ser. (Bio l.) 13: Salix Salix japonica f. angustifolia (Kimura) 74 ，fig. 3 ，tt. 8-9 (1 938); Ohwi ， F 1. Jap. ed. Kimura in Eco 1. Rev. 13: 200 (1 953). Eng 1.: 367 (1 965) & Fl. Jap. ed. rev.: 472 Salix Salix japonica f.α ngustifolia subf. fauriei (1 965); Kitam. & Murata ，Co l. Il lus t. Woody (Seemen) Kimura in Eco 1. Rev. 13: 200 Pl. Jap. 2: 325 ，fig. 781 (1979); Kimura in (1 953). Wild Flow. Jap. Wood. 1: 46 ，photo 50 ，5- February February 2000 Journal of Japanese Botany Vo l. 75 No. 1 19

7 (1 989); Ohwi & Kitag. ，New FL Jap.: 538 no-jyumoku: 691 (1976); Sugaya in (1992); (1992); Sohma in Sci. Rep. Tohoku Univ. 4 Terasaki. Il lus t.: 104 ， fig. 406 (1 977); ser. ser. (Bio l.) 40: 100 (1 993). Kitam. & Murata ，Co l. Illus t. Woody Pl. Jap. Distr.: Distr.: Japan: Central Honshu (Gunma ， 2: 307 ，fig. 760 (1 979); Kimura in Wild Nagano and Yamanashi Prefs.). Flow. Jap. Wood. 1: 51 ，photo 58 ，5-7 16-1. 16-1. Sa !i x rupifraga f. rupi 仕aga (1 989); Makino ，New Il lus t. Fl. Jap. ed.: rev. 16-2. 16-2. Salix rupifraga f. eriocarpa fig. 107 (1 989); Ohwi & Kitag. ，New Fl. (Kimura) (Kimura) Kimura [in Wild Flow. Jap. Wood. Jap.: 541 (1992); Sohma in Sci. Rep. 1: 1: 46 (1 989) ， comb. nud.] ex H.Ohashi ，sta t. Tohoku Univ. 4 ser. (Bio l.) 40: 92 (1 993). nov. nov. * According to Seemen (1 903) ， the speci- Salix Salix rupifraga var. eriocarpa Kimura in mens are “ Japan: ohne Angabe des Standorts Sci. Sci. Rep. Tohoku Imp. Univ. 4 ser. (Bio l.) (Siebold p. par t.， Buerger ， Textor); Insel 13: 13: 75 ，fig. 4 (1 938) [Ty pe: Japan. Honshu ， Kiuschiu; Nagasaki (Eykman)." Yamanashi Pref. (Prov. Kai) ， in rupium Salix buergeriana Miq. in Ann. Mus. Bo t. fissuris ，Komakimura. S. Huno s.n. (fr.) ，7 Lugd.-Ba t. 3: 28 (1 867) [Ty pe: Japan ，locis Jun. Jun. 1936 (TUS-K holo.)]; Kitam. & non indicatis ，ex Siebold ，Buerger & Murata ，Co l. Illus t. Woody Pl. Jap. 2: 325 Mohnike s.n. (?， not in L ， n.v.)]; (1 979); Ohwi & Kitag. ，New Fl. Jap.: 538 C.K.Schneid. in Sarg. ， P l. Wilson. 3: 162 (1 992). (1 916); Ohwi ， F l. Jap.: 407 (1 953); Ohwi ， Distr.: Distr.: Occurs with the typical form. Fl. Jap. ed. Eng l.: 368 (1 965) & Fl. Jap. ed. rev.: rev.: 475 (1 965); Ohwi & Kitag. ，New Fl. IV-ii. IV-ii. Sec t. Sieboldianae (Seemen) Jap.: 541 (1 992). C.K.Schneid. C.K.Schneid. Salix saidaeana Seemen ，Salic. Jap.: 68 ， Salix Salix sec t. Sieboldianae (Seemen) tab. 17 (1 903) [Syntypes: Japan. U. Faurie C.K.Schneid. ，Il lus t. Handb. Laubholz k. 1: 4995 ， U. Faurie 3694 ， U. Faurie 3705 ， U. 69 (1 904) ，as “Sieboldianae Seem." & in Faurie 3706 & U. Faurie 3708; J. Rein s.n. Sarg. ， P l. Wilson. 3: 161 (1 916); Kitam. & Apr. 1875. (B?) ，all n. v.]; C.K.Schneid. in Murata ，Co l. Il lus t. Woody Pl. Jap. 2: 305 Sarg. ， P l. WilSOTI. 3: 163 (1 916). (1 979) ， sub subgen. Salix. Type: Salix Salix daiseniensis Seemen ，Salic. Jap.: 65 ， sieboldiana sieboldiana Blume. tab. 15 (1 903) [Ty pe: Japan. Honshu ，Tottori Salix Salix [unranked] Sieboldianae Seemen in Pre f. (Prov. Hoki) ，Daisen ， U. Faurie 3708 Salic. Salic. Jap.: 21 (1903) ，sub Salix C. [3768] (♂ f1.)， 25 May [25 mai ]， 1899 Monadeniae c. Submonandrae. (KYO-isolecto. fide Kimura 1981)]; C.K.Schneid. C.K.Schneid. in Sarg. ， P l. Wilson. 3: 129 17. 17. Salix sieboldiana Blume ，Bijdr. Fl. (1 916). Ned. Ned. Ind. 10: 517 (1 825) [Ty pe: Cresci t: sub Salix harmsiana Seemen ，Salic. Jap.: 73 ， nomine S. albae a Doc t. v. Siebold ex tab. 18 (1 903) [Ty pe: Japan. Nagasaka i. R. J aponia missa et nunc in horto botanico Oldham 551-anno 1862-n.v.]; C.K.Schneid. Buitenzorgii Buitenzorgii culta. (not in L ，n.v.)]; Miq. ， in Sarg. ， P l. Wilson. 3: 163 (1 916); Ohwi ， Ann. Ann. Mus. Lugd.-Ba t. 3: 28 (1 867); Franch. F l. Jap.: 407 (1 953); Ohwi ， F l. Jap. ed. rev.: & Sav. ，Enum. P l. Jap. 1: 460 (1 875); 476 (1 965); Sugaya in Terasak i. Il lus t.: 104 ， Seemen ，Salic. Jap. 63* ，tab. 14 ，figs. F-K fig. 405 (1 977); Ohwi & Kitag. ，New Fl. (1 903); C.K.Schneid. in Sarg. ， P l. Wilson. 3: Jap.: 541 (1 992). 162 162 (1 916); Ohwi ， F l. Jap.: 408 (1 953); Salix sieboldiana var. sikokiana Koidz. in Ohwi ， F l. Jap. ed. Eng l.: 368 (1 965) & Fl. Bo t. Mag. Tokyo 27: 93 (1913) [Ty pe: Ja- Jap. Jap. ed. rev.: 476 (1 965); Hatusima ，Nihon- pan. Shikoku. Tokushima (Prov. Awa) ， 20 植物研究雑誌第75 巻第1号 平成 12 年2月

Tsurugisan. Tsurugisan. J. Nikai 2072 (fr.) ， 18 Ju 1. 1909 Tokyo 17: 91 (1 913); Kimura in Sci. Rep. (TI (TI lecto. designated by Kimura 1981)]. Tohoku Imp. Univ. 4 ser. (Bio 1.) 6 (2): 189 Salix Salix vulpina var. daiseniensis (Seemen) (1 931) & 15: 412 (1 940) ，Kimura in J. Fac. Koidz. Koidz. in Bo t. Mag. Tokyo 27: 90 (1 913). Agric. Hokkaido Imp. Univ. Sapporo 26: Salix Salix propitia Koidz. in Bo t. Mag. Tokyo 402 (1 934); Makino ，Il lus t. F1. Nippon: 675 ， 27: 27: 266 (1 913); C.K.Schneid. in Sarg. ， P 1. fig. 2023 (1 940); Ohwi ， F 1. Jap. ed. Eng 1.: Wilson. Wilson. 3: 163 (1 916). 366 (1 965) & F1. Jap. ed. rev.: 471 (1 965); Salix Salix tsukushiana Koidzumi in Bo t. Mag. Skvortsov ，Willows USSR: 152 (1 968); To kY 0 33: 220 (1 91 9) [Typ e: Jap an . Sugaya in Terasaki. Illus t.: 101 ， tab. 393 Kyushu. G. Koidzumi s.n. (♀ f1.)， Mar t. (1977); Kitam. & Murata ，Co 1. Il lus t. Woody 1918 (TI holo.)]. P1. Jap. 2: 318 ，fig. 772 (1 979); T. Shimizu ， Salix Salix doi αna Koidz. in F1. Satsum. 2: 103 New Alp. Fl. Jap. 2: 114 ，fig. 323 (1 983); (1 931) & in Acta Phytotax. Geobe t. 1: 21 Kimura in Wild Flow. J ap. Wood. 1: 46 ， (1 932) [Ty pe: Japan. Kyushu ，Kagoshima photo 53 ，1-4 (1 989); Makino ，New Il lus t. Pref. Pref. (Prov. Satsuma) ， Ijuuin. Y. Doi 9 F1. Jap. ed.: rev. fig. 105 (1989); Ohwi & (ster.) ， 12 Oc t. 1928 (KYO-lecto. designated Kitag. ，New F1. Jap.: 537 (1992); Sohma in by Kimura 1981) ，(TUS-K isolecto)]. Sc i. Rep. Tohoku Univ. 4 ser. (Bio 1.) 40: 100 Salix Salix aridaensis Koidz. in Acta Phytotax. (1 993). Geobo t. 8: 111 (1939) [Type: J apan. Salix glabra auc t. non Scopoli: Franch. & Honshu ，Wakayama Pref. (Prov. Kii) ，Arida- Sav. ，Enum. P l. Jap. 2: 503 (1876); gun ，Yawatamura ，Chikai ， in Experim. For. Tokubuchi in Bo t. Mag. Tokyo 10: page Kyoto Univ. S. Okamoto (ster.) ，22 Sep t. (1 23) (1 896). 1938 (KYO lecto. designated by Kimura Sa !i x aequitriens Seemen in Bo t. Jahrb. 1981; 1981; TUS- K isolecto)]. Sys t. 21: Beib 1. 53: 52 (1 896) & Salic. Jap.: Salix Salix daiseniensis va r. rotundifolia 70 ，tab. 15 ，figs. E-G (1 903); Tokubuchi in Kimura in Bo t. Mag. Tokyo 59: 80 (1946) Bo t. Mag. Tokyo 10: page (124) (1 896). [Ty pe: Japan. Shimane Pref. (Prov. Izumo) ， Salix reinii f. aomoriensis Seemen ，Salic. Torikamimura. Torikamimura. 1. Maruyama s.n. (fo 1.)， 12 Jap.: 42 (1903); Kimura in Eco 1. Rev. 13: Aug. 1939 (TUS-K holo.)]. 199 (1 953). Salix Salix daiseniensis f. rotundifolia (Kimura) Salix reinii var. cyclophylloides Koidz. in Kimura in Eco l. Rev. 13: 200 (1 953). Bot. Mag. Tokyo 27: 91 (1 913) [Ty pe: Ja- Distr.: Distr.: Endemic to Japan: Western Honshu pan. Honshu ， Iwate Pref. (Prov. Rikuchu) ， (Kinki (Kinki District and westward) ， Shikoku and Iwatesan. G. Nakahara s.n. (fo 1. &仕.)， 26 Kyushu. Jun. 1907 (TI lecto. ， designated by Kimura 198 1)].

18. 18. Salix reinii Seemen ，Salic. Jap.: 41 ， Sa !i x kakist αC.K.Schneid.， in Sarg. ， P 1. tab. tab. 6 (1 903) ，as “ Salix reinii Franch. & Wils. 3: 128 (1 916) [Type: Japan. Hondo ， Sav." Sav." [Enum. P1. Jap. 1: 459 (1 875) ，nom. Aomori Pref. (Prov. Mutsu) ，Hakkoda-yama ， nud. nud. & 1. c. 2: 503 (1876) ， pro syn.] [Ty pe: common ，lower and middle slopes. E. H. Nippon media. in monte Hak'san. Rein in Wilson 7103 ，5 July 1914 (GH holo.)]. Herb. Herb. Savatier 2923 & 2934 (P syn.-n.v.)]; Salix hidewoi Koidz. in Bo t. Mag. Tokyo Nedoluzhko in Charkevicz ， P 1. Vascu 1. Ori. 33: 220 (1 919) [Type: Japan. Hokkaido ， Extr. Extr. Sovie t. 7: 158 (1 995) ，as “ Salix reinii Prov. Ishikari ，M t. Nutakkamshpe. G. Franch. Franch. & Sav. ex Seem." Koidzumi s.n. (♀ fl.) ， Ju 1. 1916 (TI lecto. Salix Salix reinii Franch. & Sav.: Seemen ， designated by A.Kimura 1981)]; Kimura in Salic. Salic. Jap.: 41 (1 903); Koidz. in Bo t. Mag. J. Fac. Agric. Hokkaido Imp. Univ. Sapporo /

February February 2000 Journal of Japanese Botany Vo l. 75 No. 1 21

26: 26: 403 (1 934); Ohwi ， F l. Jap.: 404 (1 953); sub subgen. Salix; Argus in System. Bo t. Ohwi ， F l. Jap. ed. Eng l.: 366 (1 965) & Fl. Monogr. 52: 90 (1997) ，sub subgen. Vetrix Jap. Jap. ed. rev.: 472 (1 965); Kimura in Wild (Dumortier) Dumortier. Ty pe: Salix helix L. Flow. Jap. Wood. 1: 46 (1 989); Ohwi & (= S. purpurea L.). Kitag. ，New Fl. Jap.: 537 (1 992). Salix subgen. Helice (Dumortier) Salix Salix reinii var. eriocarpa Kimura in Sci. Dumortier in Bull. Soc. Roy. Bo t. Belg. 1: Rep. Rep. Tohoku Imp. Univ. 4 ser. (Bio 1.) 6: 190 145 (1 862). (1 931) [Type: J apan. Honshu ，Shizuoka Salix sec t. Purpurea Koch ， Syn. F1. Germ. Pre f. (Prov. Suruga) ，Subashiri ，pede montis Helv. 2: 646 (1 837); Newsholme ， Willows: F吋isan. A. Kimura 203 (♀ fl.) ，May 14 ， 26 (1 992) ，as “sec t. Purpurea Fries". Ty pe: 1926 (TUS-K holo.)]; Ohwi & Kitag. ，New Salix purpurea L. l. Fl. Jap.: 537 (1 992). Salix sec t. Monandrae Nakai in Bull. Soc. Salix sachalinensis f. αequitriens Dendro 1. Fr. 66: 50 (1 928). Type. Salix (Seemen) Kimura in J. Fac. Agric. Hokkaido purpurea L. Imp. Univ. Sapporo 26: 433 (1 934). Salix reinii f.α omoriensis subf. 19. Salix miyabeana Seemen in Bo t. cyclophylloides cyclophylloides (Koidz.) Kimura in Eco l. Jahrb. Sys t. 21: Beib 1. 53: 50 (1 896) [Ty pe: Rev. Rev. 13: 199 (1 953). Japan. Hokkaido ， Prov. Ishikari ， Sapporo. Sα lix reinii f. erioc αrpa (Kimura) Tokubuchi s.n. (♀ f1.)， 23 Apr. 1890 (as T.Shimizu ，New Alp. F1. Jap. 2: 358 (1 983); 1891) (SAPT isolecto. desiganted by Kimura in Wild Flow. Jap. Wood. 1: 46 Kimura 1981)] & Salic. Jap.: 57 ， tab. 12 (1 989). (1 903); Tokubuchi in Bo t. Mag. Tokyo 10: Dist r.: Japan: Hokkaido and Honshu pages (1 25) & (339); Tokubuchi ，1. c. 10: 69 ， (Central (Central Districts and northward) and Far p1. 6 (1 896); C.K.Schneid. in Sarg. ， P 1. Wils. Eastern Russia: Kuriles ，Primorye ， 3: 166 (1 916); Kimura in J. Fac. Agric. Khabarovsk and Kamchatka. Hokkaido Imp. Univ. Sapporo 26: 435

18 目 1. Salix reinii f. reinii (1 934); Ohwi ， F l. Jap. ed. Eng l.: 368 (1 965) 18-2. 18-2. Salix reinii f. pendula Kimura in & Fl. Jap. ed. rev.: 474 (l 965)~ Skvortsov ， Sci. Sci. Rep. Tohoku Univ. 4 ser. (Bio l.) 27: 145 Willows USSR: 228 (1 968); Kitam. & (1 96 1) [Type: Japan. Honshu ，Shizuoka Murata ，Co l. Il lus t. Woody Pl. J ap. 2: 312 ， Pref. Pref. (Prov. Suruga) ，Subashiriguchi ，pede fig. 764 (1 979); Lauener & McKean in montis Fujisan ， ca. 970 m s.m. A. Kimura Notes RBG Edinb. 41: 177 (1 983); Kimura 194 (fo 1.)， 5 Oc t. 1924 (TUS-K holo.)]; in Wild Flow. Jap. Wood. 1: 49 ，photo 56 ， Ohwi & Kitag. ，New F l. Jap.: 537 (1 992). 5-7 (1 989); Ohwi & Kitag. ，New F l. Jap.: Distr.: Distr.: Japan. Honshu. Shizuoka Pref. 540 (1 992); Nedoluzhko in Charkevicz ， P l. Vascu l. Or i. Extr. Sovie t. 7: 189 (1 995). IV-iii. IV-iii. Sec t. Helix Dumortier Salix sapporoensis H.Lev l. in Bull. Soc. Salix Salix sec t. Helix Dumortier ， Verh. Ges 1. Bo t. F r. 56: 302 (1909) [Type: Japan. Wilgen: 15 (1825) & in Bijdr. N atuurk. Hokkaido ， Sapporo. U. Faurie 266 ， 12 June Wetensch. 1: 56 (1 826); C.K.Schneid. in 1908 (E lecto. ，n.v. ， designated by Lauener Sarg. ， P l. Wils. 3: 165 (1 916); Nakai ， F l. & McKean 1983)]. Sylv. Sylv. Kor. 18: 110 (1 930); Skvortsov in 19-1. Salix miyabeana subsp. miyabeana Acad. Sci. URSS Nov. Sys t. P1. 1968: 70 Distr.: Japan: Hokkaido ，Korea ，China [as (1 968) & Willows USSR: 220 (1 968) ， sub Sα lix linearistipularis (Franch.) Hao: Chou subgen. subgen. Vetrix Dumortier; Kitam. & Murata ， in F1. Reipub. Pop. Sin. 20 (2): 374. (1 984)] ， Co 1. Il lus t. Woody P1. Jap. 2: 305 (1 979) ， East Russia: Far East Russia and Siberia 22 植物研究雑誌第75 巻第1号 平成12 年2月

(map: (map: Skv. fig. 62-4). 400 (1977); Hatusima ，Kagoshima-ken 19-2. 19-2. Salix miy αbeana subsp. gilgiana Shokubutsu Mokuroku: 35 (1 978); Kitam. & (Seemen) (Seemen) H.Ohashi ，comb. nov. Murata ，Co 1. Il lus t. Woody P1. Jap. 2: 310 ， Salix Salix gilgiana Seemen ，Salic. Jap.: 59 ， fig. 762 (1 979); Yamanaka in 1. Jap. Bo t. 59: tab. tab. 13 (1 903); C.K.Schneid. in Sarg. ， P 1. 185 (1 98 1); Kimura in Wild Flow. Jap. Wils. Wils. 3: 169 (1916); Kimura in J. Fac. Wood. 1: 48 ， photo 55 ，1 -4 (1 989); Makino ， Agric. Agric. Hokkaido Imp. Univ. Sapporo 26: New Illus t. F1. Jap. rev. ed.: fig. 94 (1 989); 437 (1934); Ohwi ， F 1. Jap. ed. Eng 1.: 368 Ohwi & Kitag. ，New F1. Jap.: 540 (1 992); (1 965) & F 1. Jap. ed. rev.: 475 (1 965); Sohma in Sci. Rep. Tohoku Univ. 4 ser. Skvortsov ，Willows USSR: 230 (1968); (Bio l.) 40: 99 (1993); Nedoluzhko in Sugaya in Terasaki. Il lus t.: 104 ， tab. 402 Charkevicz ， P l. Vascu l. Ori. Extr. Sovie t. 7: (1977); (1977); Kitam. & Murata ，Co l. Il lus t. Woody 189 (1 995). l. Pl. Jap. 2: 312 ，fig. 765 (1 979); Kimura in Salix purpure αauc t. non L.: Miq. in Ann. Wild Flow. Jap. Wood. 1: 49 ， photo 56 ，1- Mus. Bo t. Lugd.-Ba t. 3: 26 (1 867) & Pro l. 4 (1 989); Makino ，New Illus t. Fl. Jap. rev. Fl. Jap.: 214 (1 867); Franch. & Sav. ，Enum. ed.: ed.: fig. 91 (1 989); Ohwi & Kitag. ，New Fl. P 1. Jap. 1: 462 (1 875); C.K.Schneid. in Jap.: Jap.: 540 (1992); Sohma in Sci. Rep. Sarg. ， P 1. Wils. 3: 167 (1 916) ，p.p. Tohoku Univ. 4 ser. (Bio 1.) 40: 95 (1 993); Salix multinervis Franch. & Sav. ，Enum. Nedoluzhko in Charkevicz ， P l. Vascu l. Ori. Pl. Jap. 2: 504 (1 876). Extr. Extr. Sovie t. 7: 190 (1 995). Salix purpurea var. multinervis (Franch. Distr.: Distr.: J apan: Southern Hokkaido and & Sav.) Matsum. ，Shokubutsu Meii: 261 Honshu ， northern Korea ，China ，and East (1 895); Seemen ， Salic. Jap.: 54 ， tab. 11 Russia: Russia: U ssuri. (1 903); C.K. Schneid. in Sarg. ， P 1. Wils. 3: 19-2-1. 19-2-1. Salix miyabean αsubsp. gilgiana 168 (1916) ，as “ S. purpurea subsp. f. f. gilgiana (Seemen) H.Ohashi ，comb. nov. amplexicaulis var. multinervis". 19-2-2. 19-2-2. Salix miyabeana subsp. gilgiana Salix purpure αL. subsp. amplexicaulis f. f. pendula (Kimura) H.Ohashi ，comb. nov. auct. non (Bory & Chaub.) Boissier: Koidz. Salix Salix gilgiana var. pendula Kimura in Sci. in Bot. Mag. Tokyo 27: 92 (1 913). Rep. Rep. Tohoku Univ. 4 se r. (Bio 1.) 20: 211 Dist r.: J apan: Hokkaido ，Honshu and (1 96 1) [Ty pe: Japan. Honshu ， Miyagi Pref. Kyushu ，Korea ， E. & NE. China and Far (Prov. (Prov. Rikuzen) ，Sendai ，cul t. A. Kimura East Russia: U ssuri and Kuril: Etorohu. Wet 3096 (♀ fl.) ， 16 Apr. 1953 (TUS-K lecto. ground or riverside. desiganted desiganted by Kimura 1981)]. 20-1. Salix integra f. integra 20-2. 20-2. Salix integra f. pendula Kimura in 20. 20. Salix integra Thunb. in Murray ，Sys t. J. Jap. Bo t. 57: 161 ，figs. 1-2 (1 982) [Ty pe: Veg. Veg. ed. 14: 880 (1 784) ， F l. Jap.: 24 (1 784) J apan. Prov. Shinano ， decli vi in loco [Ty pe: Japan ，Thunberg (UPS holo.)] & P1. graminoso-herboso ， ca. 1200 m s.m. in Jap. Jap. Nov. Sp.: 4 (1 824); Siebold & Zucc. ， N agawamura-Oidaira. A. Kimura 3429 l. Fl. Jap. Fam. Na t. 2: 211 (1 846); Nakai ， F 1. (fo 1.)， 24 Aug. 1978 (TUS-K holo.)]. Sylv. Sylv. Kor. 18: 113 (1 930); Kimura in J. Fac. J apan. N agano Pre f. Agric. Agric. Hokkaido Imp. Univ. Sapporo 26: 434 (1 934); Makino ，Il lus t. Fl. Nippon: 672 ， 21. *Salix koriyanagi Kimura ex G りrz ， fig. fig. 2015 (1 940); Ohwi ， F l. Jap. ed. Eng 1.: Schedae ad fasc. 1. Salicac. Asia t.: 17 (193 1) 367 (1 965) & F1. Jap. ed. rev.: 474 (1 965); [Ty pe: Japan. Honshu ，Miyagi Pref. (Prov. Skvortsov ，Willows USSR: 229 (1 968); Rikuzen) ，Sendai ，cult. A. Kimura 1931 Sugaya in Terasaki. Il lus t. rev. ed.: 103 ，tab. (fo 1.)， 3 Ju l. 1930 (TUS-K lecto. designated February February 2000 Journal of Japanese Botany Vo l. 75 No. 1 23 by Kimura 1981)] & in Fedde ，Reper t. 32: 22. Salix subopposita Miq. in Ann. Mus. 119 ，388 (1 933); H.Hara in Bo t. Mag. Tokyo Bo t. Lugd.-Ba t. 3: 28 (1 867) [Type: legit 48: 48: 799 (1 934); Makino ，Il lus t. F1. Nippon: Siebold ， qui pro S. integra Th. habuit; Pierot 672 ，fig. 2014 (1 940); Ohwi ， F 1. Japan: 406 in montibus Hakone ins. Nippon ad ripas (1 953); Kimura in Sci. Rep. Tohoku Univ. rivulorum utrumque exum et folia legi t.- 4 ser. (Bio 1.) 20: 209 (1 954) & 1. c. 28: 42 Kuro Janagi isp.]; Nakai ， F l. Sylv. Kor. 18: (1 962); Ohwi ， F 1. Jap. ed. Eng 1.: 368 (1 965) 160 ，tab. 37 (1930); Ohwi ， F l. Jap. ed. Eng l.: & F1. Jap. ed. rev.: 474 (1 965); Skvortsov ， 366 (1 965) & F1. Jap. ed. rev.: 471 (1 965); Willows USSR: 226 (1 968); Sugaya in Sugaya in Terasaki. Il lus t.: 101 ， tab. 392 Terasak i. Il lus t.: 103 ， tab. 401 (1 977); (1 977); Kitam. & Murata ，Co l. Il lus t. Woody Kitam. Kitam. & Murata ，Co 1. Illus t. Woody Pl. Jap. Pl. Jap. 2: 315 ，fig. 769 (1 979); Kimura in 2: 2: 311 ， fig. 763 (1 979); Kimura in Wild Wild Flow. J ap. Wood. 1: 44 ， photo 50 ，5- Flow. Jap. Wood. 1: 49 ，photo 55 ，5-7 7 (1989); Ohwi & Kitag. ，New Fl. Jap.: 536 (1989); (1989); Makino ，New Il lus t. Fl. Jap. ed.: rev. (1 992). fig. fig. 93 (1 989); Ohwi & Kitag. ，New Fl. Jap.: Dist r.: Japan: W. Honshu (Chugoku 540 (1 992). Dis t.)， NW. Shikoku ， N. Kyushu and South Salix Salix purpurea L. subsp. amplexicaulis Korea (Quelpaert Is 1.). (Bory & Chaub.) Koidz. var. petiolata subvar.α ngustifolia Koidz. in Bo t. Mag. IV-v. Sec t. Subviminales (Seemen) Tokyo 27: 92 (1 913). C.K.Schneid. Salix Salix purpurea var. japonica Nakai in Salix sec t. Subviminales (Seemen) Bull. Bull. Soc. Dendr. Fr. 66: 50 (1 928) [苛 pe: C.K.Schneid. ，Il lus t. Handb. Laubholzk. 1: Korea. Korea. Heihok ，Teisyu. T. Ishidoya no. 3838 65 (1 904); Skvortsov in Acad. Sci. URSS (fr.) ，2 May 1920 (TI ，lecto. designated by Nov. Sys t. Pl. 1968: 69 (1 968) & Willows Nakai ， 1930)] & Nakai ， F l. Sylv. Kor. 18: USSR: 201 (1968) ，sub subgen. Vetrix 117 117 (1 930). Dumortier; Kitam. & Murata ，Co l. Il lus t. Salix Salix purpurea auc t. Japon. ，non L.: Woody Pl. Jap. 2: 305 (1 979) ， sub subgen. Seemen ，Salic. Jap.: 54 ，tab. 11 (1903) ，exc l. Salix; Newsholme ， Willows: 25 (1 992) ， sub syn.; syn.; Matsum. ， Index Pl. Jap. 2: 12 (1 912) ， subgen. Caprisalix. Ty pe: Salix gracilis η la p.p. p.p. Miq. Distr.: Distr.: Japan (cultivated) and Korea. Salix [unranked] Subviminales Seemen in Salic. Salic. Jap.: 20 (1903) ，sub Salix C. IV-iv. IV-iv. Sec t. Incubaceae A.Kerner Monad 白山e b. Synandrae s. Dolichostylae. Salix Salix sec t. Incubaceae A.Kerner in Verh. Ty pe: Salix gracilis η la Miq. Zoo l.聞 Bo t. Ges. Vereins Wien 10: 264 Salix sec t. Gracilistylae C.K.Schneid. in (1860); (1860); C.K. Schneid. in Sarg. ， P 1. Wils. 3: Sarg. ， P 1. Wils. 3: 163 (1 916); Nakai ， F 1. 153 153 (1 916); Nakai ， F 1. Sylv. Kor. 18: 157 Sylv. Kor. 18: 100 (1 930). Type: Sα lix (1 930); Skvortsov in Acad. Sci. URSS Nov. gracilistyla Miq. Sys t. Pl. 1968: 70 (1968) & Willows USSR: 216 (1 968) ， sub subgen. Vetrix; Newsholme ， 23. Salix gracilistyla Miq. in Ann. Mus. Willows: Willows: 25 (1 992). Ty pe: Salix repens L. Lugd.-Ba t. 3: 26 (1 867) & Pro l. Fl. Jap.: 214 Salix Salix sec t. Repentes Wimmer: Pax in (1 867); Tokubuchi in Bo t. Mag. Tokyo 10:

Engler Engler & Prantl ，Na t. Pflanzenfam. 3， 1: 37 page (1 24) (1 896); Koidz. in Bo t. Mag. To 四 (1 887) ，p.p.; Kitam. & Murata ，Co l. Il lus t. kyo 27: 92 (1913); C.K.Schneid. in Sarg. ， Woody P1. Jap. 2: 305 (1 979) ， sub subgen. Pl. Wils. 3: 164 (1 916); Nakai ， F 1. Sylv. Kor. Salix. Salix. Ty pe: Salix repens L. 18: 104 (1 930); Makino ，Il lus t. F1. Nippon: 24 植物研究雑誌第75 巻第1号 平成 12 年2 月

671 ，fig. 2012 (1940); Ohwi ， F l. Jap. ed. 1254 (♂日.)， 22 Mar. 1927 (TUS-K holo.)]. Eng l.: 368 (1965) & Fl. Jap. ed. rev.: 475 23-1-4. Salix gracilis η la var. gracilistyla (1 965); Skvortsov ，Willows USSR: 201 f. variegata (Kimura) Kimura in Eco l. Rev. (1 968); Sugaya in Terasak i. Il lus t.: 104 ，tab. 13: 200 (1 953). 403 (1 977); Hatusima ，Kagoshima-ken Salix gracilistyla var. variegata Kimura in Shokubutsu Shokubutsu Mokuroku: 35 (1978); Kitam. & Sci. Rep. Tohoku Imp. Univ. 4 ser. (Bio l.) Murata ，Co l. Il lus t. Woody Pl. Jap. 2: 308 ， 12: 107 (1 937). [Type: Japan. Honshu ， fig. fig. 761 (1 979); Kimura in Wild Flow. Jap. Miyagai Pref. (Prov. Rikuzen) ，Sendai ，cul t. Wood. 1: 50 ， photo 57 ， 1-4 (1 989); Makino ， A. Kimura 2409 (TUS-K)]. New Il lus t. Fl. Jap. rev. ed.: fig. 90 (1 989); Dist r.: Cultivated. Ohwi & Kitag. ，New F l. Jap.: 540 (1 992); 23-2. Salix gracilistyla var. graciliglans Sohma in Sci. Rep. Tohoku Univ. 4 ser. (Nakai) H.Ohashi ，comb. nov. (Bio l.) 40: 95 (1993); Nedoluzhko in Salix graciliglans Nakai in Bo t. Mag. Charkevicz ， P l. Vascu l. Ori. Extr. Sovie t. 7: Tokyo 30: 274 (1916) [Type: Korea. 187 187 (1 995). Heihoku ，Kang-gei [Kangkai]. Ralph G. Salix Salix thunbergiana B l. ex Andersson in Mills 312 (♀ fl.) ，28 Apr. 1911 (TIlecto. DC. ， Prodr. 16 (2): 271 (1 868); Seemen ， designated by Nakai 1930)] & Fl. Sylv. Kor. Salic. Salic. Jap.: 61 ，tab. 14 (1 868). 18: 102 (1 930); Ohwi ， F l. Jap. ed. Eng l.: 23-1. 23-1. Salix gracilistyl αvar. gracilistyla 368 (1 965) & Fl. Jap. ed. rev.: 475 (1 965); 23-1-1. 23-1-1. Salix gracilistyla var. gracilistyla Ohwi & Kitag. ，New Fl. Jap.: 541 (1 992). f. f. gracilistyla Salix nakaii Kimura in Bo t. Mag. Tokyo

Distr.: Distr.: Japan: Hokkaido ，Honshu ， Shikoku 40: 637 (1 926) [Ty pe: Japan. Honshu ，To 四 and K yushu ，Korea and N orth & NE. China. kyo ， (Prov. Musashi) ，cul t. in horto Bo t. 23-1-2. 23-1-2. Salix gracilistyla var. gracilistyla Univ. Imp. Tokyo. A. Kimura 371 (♂ fl.) ， f. f. adscendens (Kimura) H.Ohashi ，comb. 25 Feb. 1926 (TI holo.)]. nov. nov. Dist r. Japan: Honshu (Kinki Dis t. and Salix Salix gracilis η la var. adscendens Kimura westward) ，Shikoku ，and N. Kyushu and in in Sci. Rep. Tohoku Imp. Univ. 4 ser. (Bio l.) Korea. 12: 12: 107 (1937) [Type: Japan. Honshu ， 23-3. Salix gracilistyla var. Miyagi Pref. (Prov. Rikuzen) ， Sendai. A. melanostachys (Makino) C.K.Schneid. in Kimura 3042 (♂ fl.) ， 11 Mar. 1949 (TUS-K Sarg. ， P l. Wils. 3: 164 (May 1916); Makino lecto. lecto. designated by A.Kimura 1981)]. ex Matsum. ，Shokubutsu-meii ， ed. 9: 353 Salix Salix gracilistyla va r. αdscendens f. (J une 1916); Ohwi ， F l. Jap.: 407 (1 953) & latifolia latifolia Kimura Sci. Rep. Tohoku Univ. 4 in Bull. Na t. Sci. Mus. no. 33: 70 (1 953) ， as ser. ser. (Bio l.) 26: 4 ，tab. 3 (1 960) [Ty pe: Japan. var. melanostachys (Makino) Ohwi; Ohwi ， Honshu ， Tochigi Pre f. (Prov. Shimotuke) ， Fl. Jap. ed. Eng l.: 368 (1 965) & Fl. Jap. ed. secus secus torrentem Nigatogawa ， in Imperiali rev.: 475 (1 965); Kitam. & Murata ，Co l. Praedio Praedio Rustico N asuensi ，altitudine circa Illus t. Woody Pl. Jap. 2: 309 (1 979); Kimura 660 m s.m. A. Kimura 927 (fo l.)， 18 Aug. in Wild Flow. J ap. Wood. 1: 50 ， photo 57 ， 1956 (TUS-K holo.)]. 5 (1 989). 23-1-3. 23-1-3. Salix gracilistyla var. gracilistyla Salix thunbergiana var. melanostachys f. f. pendula (Kimura) H.Ohashi ，sta t. nov. Makino in Bo t.恥 lag. Tokyo 18: 141 (1 904) Salix Salix gracilistyla var. pendula Kimura in [Type: J apan. Honshu ，Tokyo Pref. (Prov. Bo t. Mag. Tokyo 42: 570 (1 928) [Ty pe: Ja- 恥1usashi) ，Tokyo ，Bo t. Gard. Koishikawa ， pan. Honshu ，Tochigi Pref. (Prov. cul t. S. Okubo 36 (♂ fl.) ，March. anno? (TI Shimotsuke) ，Utsunomiya ，cul t. A. Kimura lecto. designated by A.Kimura 198 1)]. February February 2000 Journal of Japanese Botany Vo l. 75 No. 1 25

Salix Salix gracilistyla subsp. melanostachys Vill. (Makino) Makino in Bo t. Mag. Tokyo 28: Salix sec t. Daphnoideae Dumortier ， F 1. 175 175 (1 914); Makino & Nemoto ， F 1. Japan Belg. Prodr.: 12 (1 827) & in Bul l. Soc. Bo t. ed. ed. 2: 164 (1 931). Belg. 1: 143 (1 862); C.K.Schneid. in Sarg. ， Salix Salix melanostachys Makino ex Gorz ， Pl. Wils. 3: 154 (1 916); Nakai ， F l. Sylv. Kor. Schedae ad fasc. 1. Salicac. Asiatic.: 21 18: 91 (1 930); Newsholme ，Willows: 25 (1 93 1) & in Reper t. Sp. Nov. Reg. Veg. 32: (1 992). 121 121 (1 933); Kimura in Sci. Rep. Tohoku Imp. Univ. 15: 414 (1940); Sugaya in 25. Salix rorida Lacksch. in Sched. Herb. Terasak i. Il lus t.: 104 ，fig. 404 (1 977). Fl. Ross. 7: 131 (1 911); Miyabe & Kudo ， Distr.: Distr.: Cultivated rarely in gardens. Icon. Ess. For. Tr. Hokkaido 5: 55 ， tab. 16 (1 921); Nakai ， F l. Sylv. Kor. 18: 92 (1 930); IV- vi. Sec t. Hukaoana Kimura Ohwi ， F l. Jap. ed. Eng l.: 367 (1 965) & Fl. Salix Salix sec t. Hukaoana Kimura in J. Jap. Jap. ed. rev.: 473 (1 965); Skvortsov ，Wil- Bo t. 49: 46 (1 974); Kitam. & Murata ，Co l. lows USSR: 214 (1 968); Kitam. & Murata ， Illus t. Woody Pl. Jap. 2: 305 (1 979) ， sub Co l. Il lus t. Woody Pl. Jap. 2: 319 ，fig. 773 subgen. Salix. Type: Salix hukaoana (1 979); Kimura in J. Jap. Bo t. 58: 129 Kimura. Kimura. (1 983) & in Wild Flow. Jap. Wood. 1: 43 ， photo photo 49 ，1-6 (1989); Chou in Fl. Reipub. 24. 24. Salix hukaoana Kimura in J. Jap. Pop. Sin. 20 (2): 313 (1984); Ohwi & Bo t. 48: 321 ，fig. 1 (1 973) [Type: Japan. Kitag. ，New F l. Jap.: 538 (1992); Prov. Prov. Kozuke ， secus torrentem Yubisogawa ， Nedoluzhko in Charkevicz ， P l. Vascu l. Ori. Yubiso ， altitudine circa 560 m s.m. A. Extr. Sovie t. 7: 158 (1 995). Kimura & S. Hukao (♀日. sub anth.) ，14 Salix lackschewitziana Toepfer ， Osterr.

Ap r. 1973 (TUS 四 K holo.)]; Kitam. & Bo t. Zeits. 66: 402 (1916); Kimura in Sci. Murata ，Co l. Il lus t. Woody Pl. Jap. 2: 309 Rep. Tohoku Imp. Univ. 6: 185 (1 931) ， as (1 979); Kimura in Wild Flow. Jap. Wood. 1: var. typic αKimura & Kimura in J. Fac. 51 ，photo 58 ，1-4 (1989); Ohwi & Kitag. ， Agric. Hokkaido Imp. Univ. Sapporo 26: New F l. Jap.: 539 (1 992); Sohma in Sci. 428 (1 934). Rep. Tohoku Univ. 4 se r. (Bio l.) 40: 117 Salix daphnoides auc t.， non Vill.: Ledeb. ， (1 993). Fl. Ross. 3 (2): 602 (1850) ， p. p.; Tokubuchi Distr.: Distr.: Endemic to Japan: Honshu (North- in Bo t. Mag. Tokyo 10: page (1 23) (1 896); ern ern Kanto and Tohoku Districts). C.K.Schneid. ，Il lus t. Handb. Laubh. 1: 44 (1 904) ， p. p.; Shirasawa ， Icon. Ess. For. IV-vii. IV-vii. Sec t. Daphnella Seringe Trees Jap. 2: tab. 10 (1 908); Matsum. ， Ind. Salix sec t. Daphnella Seringe ， Pl. Jap. 2 (2): 9 (1 912). Exemplaires Exemplaires desseches de la revision inedit Distr.: Japan: Hokkaido and Honshu ， du genre Salix ， 10th page [no pagination] Korea ，NE. China and East Russia: Amur ， (1824) (1824) [cf. Argus in System. Bo t. Monogr. Primorye ，Khabarovsk ，Magadan ， Sakhalin 52: 52: 5 (1 997)]; Skvortsov in Acad. Sci. and Siberia.

URSS Nov. Sys t. P l. 1968: 69 (1 968) & 25- 1. Sα lix rorid α， f. rorida Willows USSR: 211 (1 968) ， sub subgen. 25-2. Salix rorid αf. pendula Kimura in Vetrix Vetrix Dumortier; Kitam. & Murata ，Co l. J. Jap. Bo t. 58: 132 (1 983) [Ty pe: Japan. Illus t. Woody P l. J ap. 2: 306 (1 979) ， sub Miyagi Pre f. (Prov. Rikuzen). cul t. in Horto subgen. subgen. Salix; Chou in Fl. Reipub. Pop. Sin. Bo t. Fac. Sci. Tohoku Univ. Sendaiensis. A. 20 (2): 311 (1 984). Ty pe: Salix daphnoides Kimura 3442 (fo l.)， 6 Sep t. 1982 (TUS ・K)] 26 26 植物研究雑誌第75 巻第1号 平成 12 年2月

& 59: 4 (1 984). C.K.Schneid. in Sarg. ， P l. Wi1s. 3: 157 Dist r.: Cultivated in Japan ，Honshu ， (1 916); Nakai ， F 1. Sylv. Kor. 18: 174 (1 930); Miyagai Pref. ， Sendai. Newsholme ，Willows: 25 (1 992). Type: 25-3. 25-3. Salix rorid α f. roridaeformis Salix viminalis L. (Nakai) (Nakai) Kimura [in Wild Flow. Jap. Wood. Salix subgen. Vimen (Seringe) Dumortier 1: 1: 43 (1 989) ，comb. nud.; Sohma in Sci. in Bull. Soc. Roy. Bo t. Belg. 1: 143 (1 862). Rep. Tohoku Univ. 4 ser. (Bio l.) 40: 106 (1 993)] ex H.Ohashi ，comb. nov. 26. Salix schwerinii E. Wolf in Mitt. Salix Salix roridaeformis N akai in Bo t. Mag. Deutsch. Dendro l. Ges.: 407 (1929) & in Tokyo 33: 5 (1 919) [Ty pe: Korea. Kannan ， Bull. Jard. Bo t. Princ. URSS 28: 421 (1 929); Taichu-ri. Taichu-ri. T. Ishidoya 2738 (♀ fl.) ，22 May Skvortsov ，Willows USSR: 196 (1 968); 1918 (TI lecto. designated by Nakai 1930)] Nedoluzhko in Charkevicz ， P 1. Vascu l. Ori. & Fl. Sylv. Kor. 18: 96 (1 930). Extr. Sovie t. 7: 183 (1 995). Salix Salix lackschewitzian αvar. roridaeformis Salix viminalis auc t. non L.: Sugiyama ， (Nakai) (Nakai) Kimura in Sc i. Rep. Tohoku Imp. Hokkaido-Zyumokusiryo n. 158 (1 890) ， fide

Univ. Univ. 4 ser. (Bio l.) 6: 187 (1 931). Kimura 1937; Tokubuchi in Bo t. Mag. To 同 Salix Salix rorida var. roridaeformis (Nakai) kyo 10: page (1 24) (1 896): Seemen ，Salic. Ohwi ， F l. Jap.: 405 (1 953) & in Bul l. Na t. Jap.: 50 ， tab. 9，F-K (1 903); Miyabe & Sci. Sci. Mus. no. 33: 70 (1 953); Ohwi ， F l. Jap. Miyake ， F l. Sagha l.: 428 (1 915); Nakai ， F l. ed. ed. Eng l.: 367 (1 965); Kitam. & Murata ， Sylv. Kor. 18: 175 (1 930). Co l. Il lus t. Woody Pl. Jap. 2: 319 (1 979) ， ut Salix viminalis L. va r. yezoensis var. “var. roridaeformis (Nakai) Kimura"; Chou C.K.Schneid. in Sarg. ， P l. Wilson. 3: 158 in in Fl. Reipub. Pop. Sin. 20 (2): 313 (1 984); (1 916); Miyabe & Kudo ， Icon. Ess. For. Tr. Ohwi & Kitag. ，New Fl. Jap.: 539 (1992). Hokkaido 1: 58 ，tab. 17 (1 92 1); Makino & Distr.: Distr.: Japan ， as in the species. Nemoto ， F l. Jap. ed. 2: 173 (1 931). Salix Salix yezoensis (C.K.Schneid.) Kimura in IV-vii i. Sec t. Viminella Seringe Bot. Mag. Tokyo 45: 28 (1 93 1) & in Sci. Salix sec t. Viminella Seringe ， Rep. Tohoku Imp. Univ. 4 ser. (Bio 1.) 6: 190 Exemplaires Exemplaires desseches de la revision inedit (1 931); Kimura in J. Fac. Agric. Hokkaido du genre Salix ， 10th page [no pagination] Imp. Univ. Sapporo 26: 430 (1934); (1824); (1824); Argus in System. Bo t. Monogr. 52: Nemoto ， F l. Jap. Supp l.: 118 (1936); 83 83 (1 997) ， sub subgen. Vetrix (Dumortier) Makino ，Illus t. Fl. Nippon: 674 ，fig. 2020 Dumortier. Dumortier. Ty pe: Salix viminalis L. (1 940). Salix Salix sec t. Vimen (Seringe) Dumortier ， Salix yezoensis var. angust ザ'o li αKimura Verh. Verh. Ges l. Wilgen: 14 (1 825) & in B りdr. in Sci. Rep. Tohoku Imp. Univ. 4 ser. (Bio 1.) Natuurk. Wetensch. 1: 56 (1 826) ， p.p.; 6: 194 (1 931) [Ty pe: Sachalin. Distr. Sikka ， Skvortsov Skvortsov in Acad. Sci. URSS Nov. Sys t. P1. Siska. A. Kimura 1541 (fo 1.)， 21 Aug. 1927

1968: 69 (1 968) & Willows USSR: 191 (TUS 四 K holo.)]. (1968) ，sub subgen. Vetrix Dumortier; Salix yezoensis var. abbre νiata Kimura in Kitam. Kitam. & Murata ，Co 1. Illus t. Woody Pl. Jap. J. Fac. Agr. Hokkaido Imp. Univ. 26: 431 2: 2: 306 (1 979) ，sub subgen. Salix. Type: (1 934) [Type: Japan. Hokkaido ， Prov. Sa !i x viminalis L. Tokati ，Satsunai. M. Tatewaki 10483 (fo 1.) ， Salix Salix [unranked] Viminales Bluff & 3 Aug. 1927 (SAPT holo. TUS-K iso.)]. Fingerhuth ，Cimp. Fl. Germ. 2: 256 (1 825). Salix serotin αPallas va r. yezoensis Salix Salix sec t. Viminales Bluff & Fingerhuth: Kimura ex G り rz in Reper t. Spec. Nov. Regni W. Koch ，Syn. F l. Germ.: 647 (1837); Veg. 36: 26 (1 934). February February 2000 JournaI of Japanese Botany Vo l. 75 No. 1 27

Salix Salix petsusu Kimura in Sci. Rep. Tohoku 27. Salix udensis Trautv. & Mey. ， F l. Imp. Univ. 4 ser. (Bio l.) 12: 317 (1 937) ， as Ocho t.: 81 (1856); Skvortsov ，Willows S. “S. Pet-susu" & in Sci. Rep. Tohoku Univ. USSR: 200 (1968); Nedo1uzhko in 4 ser. (Bio l.) 18: 548 (1 950); Ohwi ， F l. Jap. Charkevicz ， P l. Vascu l. Ori. Extr. Sovie t. 7: ed. ed. Eng l.: 367 (1965); Kitam. & Murata ， 181 (1 995). Co l. Illus t. Woody Pl. Jap. 2: 320 ，fig. 775 Salix sachalinensis F. Schmidt in Mem. (1 979); Kimura in Wi1d F1ow. Jap. Wood. 1: Acad. Imp. Sci. St. Petersb. ser. 7， 12: 173 42 ，photo 48 ，4-5 (1 989); Ohwi & Kitag. ， (1 868); Tokubuchi in Bo t. Mag. Tokyo 10: New Fl. Jap.: 539 (1 992). page (1 24) (1 896); Seemen ，Sa1ic. Jap.: 53 ， Salix Salix petsusu var. abbreviata (Kimura) tab. 10 (1 903); C.K.Schneid. in Sarg. ， P l. Kimura in Sci. Rep. Tohoku Imp. Univ. 4 Wi1s.3: 158 (1 916); Miyabe & Kudo ， Icon. ser. ser. (Bio l.) 12: 318 (1 937). Ess. For. Trees Hokkaido 1: 59 ， tab. 18 Salix Salix petsusu var. angustifolia (Kimura) (1 921); Kimura in J. Fac. Agric. Hokkaido Kimura in Sci. Rep. Tohoku Imp. Univ. 4 Imp. Univ. Sapporo 26: 431 (1 934); Makino ， ser. ser. (Bio l.) 12: 318 (1 937). Il 1us t. Fl. Nippon: 673 ， fig. 2019 (1940); Salix Salix kinuyanagi Kimura in Sci. Rep. Kimura in Sci. Rep. Tohoku Univ. 4 ser. Tohoku Imp. Univ. 4 ser. (Bio l.) 15: 401 (Bio l.) 27: 141 (1 961); Ohwi ， F l. Jap.: 406 (1 940) [Ty pe: Japan. Honshu ，Miyagi Pref. (1 953); Kurata ，Il 1us t. Imp. For. Tr. Jap. 1: (Prov. (Prov. Rikuzen) ，Sendai ，cu1 t. A. Kimura n. 54 ， p l. 27 (1 964); Ohwi ， F l. Jap. ed. Eng l.: 2192 (♂f1.)， 11 Apr. 1933 (TUS-K 1ecto.)]; 367 (1 965) & Fl. Jap. ed. rev.: 474 (1 965); Ohwi ， F l. Jap. ed. Eng l.: 367 (1 965) & F l. Sugaya in Terasak i. Il 1us t.: 103 ， tab. 399 Jap. Jap. ed. rev.: 473 (1965); Sugaya in (1 977); Kitam. & Murata ，Co l. Illus t. Woody Terasaki. Terasaki. Il 1us t.: 103 ， tab. 398 (1 977); Pl. Jap. 2: 321 ，fig. 776 (1 979); Chou in F l. Kitamura & Murata ，Co l. Illus t. Woody Pl. Reipub. Pop. Sin. 20 (2): 326 (1 984); Jap. Jap. 2: 320 ，fig. 774 (1 979); Kimura in Wi1d Mizuno & al. in Bo t. Mag. Tokyo 102: 403 F1ow. Jap. Wood. 1: 43 ，photo 48 ，6-7 (1989) [chemotaxonomy*]; Kimura in Wi1d (1 989); Makino ，New Il 1us t. Fl. Jap. ed.: rev. F1ow. Jap. Wood. 1: 42 ，photo 48 ，1-3 fig. fig. 102 (1 989); Ohwi & Kitag. ，New F l. (1 989); Makino ，New Illus t. Fl. Jap. rev. ed.: Jap.: Jap.: 539 (1992); Sohma in Sci. Rep. fig. 101 (1 989); Ohwi & Kitag. ，New F l. Tohoku Univ. 4 ser. (Bio l.) 40: 99 (1 993). Jap.: 539 (1992); Sohma in Sci. Rep. Salix Salix petsusu f.α ngust ザ'o lia (Kimura) Tohoku Univ. 4 ser. (Bio l.) 40: 106 (1993). Kimura in Eco l. Rev. 13: 200 (1 953). [Sα lix sikokiana Makino in Bo t. Mag. Salix Salix kinuyanagi mstr. ligul α ta Kimura in Tokyo 6: 49 (1 892) ，nom. nud. Specimen Sci. Sci. Rep. Tohoku Univ. 4 ser. (Bio l.) 27: cited: Japan. Tosa ，Nanokawa. K. Watanabe 143 ， tab. 9 (1 961). Specimen cited: Japan. s.n.]. Honshu ，Shizuoka Pref. (Prov. Totomi) ， Salix op α ca Andersson [in Tokubuchi in Hamanagun ，cu1 t.， sine 1oco specia1i Bo t. Mag. Tokyo 10: page (1 24) (1 896)] ex indicato. indicato. E. Isikawa s.n. (♂)， Mar. 1940 Seemen ， Sa1ic. Jap.: 51 ， tab. 10 (1 903); (TUS-K)]. Shirasawa ， Icon. Forest Tr. Jap. 2: tab. 9 Distr.: Distr.: J apan: Hokkaido and Honshu (cen- (1 908). tra1 tra1 and northern regions ， cultivated in Salix makinoana Seemen in Reper t. Sp. Honshu ，Shikoku and Kyushu) ，Korea ，NE. Nov. Reg. Veg. 1: 173 (1 905) ， p. p. China and East Russia: Siberia ，Amur ， Salix pilgeriana Seemen in Reper t. Sp. Khabarovsk ， Primorye ， Sakha1in ， Nov. Reg. Veg. 7: 134 (1 909). Kamchatka and Magadan. Salix sachalinensis va r. pilgeri αna (Seemen) C.K.Schneid. in Sarg. ， P l. Wi1s. 3: 28 28 植物研究雑誌第75 巻第l号 平成 12 年2月

159 159 (1 916). Koch ， Syn. F1. Germ. Helv. 2: 650 (1 837); Salix Salix sach α linensis f. latior Kimura in C.K.Schneid. in Sarg. ， P l. Wils. 3: 148 Sci. Sci. Rep. Tohoku Univ. 4 ser. (Bio 1.) 27: 141 (1916) ，p.p. ，as “Capreae Bluff & (1 961) [Ty pe: Japan. Honshu ，Tokyo (Prov. Fingerhuth"; Kitam.. & Murata ，Co 1. Il lus t. Musashi) ，ad fossam Dokanbori in Sede Woody P1. Jap. 2: 305 (1979) ， sub subgen. Imperatoris Augustissimi Tokiensi. A. Salix. Ty pe: Salix caprea L. Kimura 965 (fo 1. intermed.) ，24 May 1957 Salix sec t. Vetrix Dumortier ， Verh. Ges 1. (TUS-K holo.)]. Wilgen: 14 (1 825); Newsholme ， Willows: Salix Salix sachalinensis f. ligulata Kimura in 25 (1 992) ， p. p. inc 1. sec t. Capreae. J. J. Jap. Bo t. 36: 163 (1 961) [Type: Japan. Sα lix subgen. Caprisalix (Dumortier) Hokkaido ， Prov. Kushiro ， in paludosis Nasarov in F1. URSS 5: 57 (1936). Type: Kiritappu ，spontanea. A. Kimura 2000 (str.) ， Salix caprea L. 21 21 May 1931 (TUS-K holo.)] & in Sci. Rep. Tohoku Univ. 4 ser. Bio 1. 27: 142 ，tab. 6-8 28. Salix taraikensis Kimura in J. Fac. (1 96 1)， as “mstr. ligulat αKimura". Agric. Hokkaido Imp. Univ. 26: 419 (1934) Distr.: Distr.: Japan: Hokkaido ，Honshu and [Ty pe: Sachalin australis. Shikka ，Siska. A. Shikoku ，Korea ，NE. China and East Russia: Kimura 6718 (♀ fl.) ，7 June 1927 (TUS-K Siberia ，Amur ，Primorye ，Khabarovsk ，Kuril ， lecto. designated by Kimura ， 1981)]; Ohwi ， Sakhalin ，Kamchatka and Magadan. F1. Jap. ed. Eng 1.: 366 (1965) & F1. Jap. ed. *Two different chemical races were found rev.: 471 (1 965); Skvortsov ， Willows USSR: in in this species in Japan; one composed of 175 (1 968); Kitam. & Murata ，Co l. Il lus t. flavonoids flavonoids and the other phenylpropanoid Woody P1. Jap. 2: 317 ， fig. 771 (1 979); derivatives derivatives (Mizuno & a1. 1989). Kitag. ，Neo-Lineam. F l. Mansh.: 210 (1979); (1979); Chou in Fl. Reipub. Pop. Sin. 20 IV-ix. IV-ix. Sec t. Cinerella Seringe (2): 296 (1 984); Kimura in Wild Flow. Jap. Salix sec t. Cinerella Seringe ， Wood. 1: 43 ， photo 50 ， 1-4 (1 989); Ohwi & Exemplaires Exemplaires desseches de la revision inedit Kitag. ， New F l. Jap.: 536 (1992); du genre Salix ，2nd page [no pagination] Nedoluzhko in Charkevicz ， P 1. Vascu 1. Ori. (1 824); Argus in System. Bo t. Monogr. 52: Extr. Sovie t. 7: 175 (1 995). 74 (1 997) ，sub subgen. Vetrix (Dumortier) Salix taraikensis var. latifolia Kimura in Dumortier. Dumortier. Ty pe: Salix cinerea L. J. Fac. Agric. Hokkaido Imp. Univ. 26: 421 Salix Salix sec t. Vetrix Dumortier ， Verh. Ges 1. (1934) [Type: J apan. Hokkaido ， Prov. Wilgen: 14 (1 825) & in Bijdr. Natuurk. Kushiro ，Hamanaka. A. Kimura 2005 (fo 1.)， Wetensch. Wetensch. 1: 56 (1 826) ， p. p.; Skvortsov in 22 Ju 1. 1931 (TUS-K holo. & iso.)]; Chou in Acad. Sci. URSS Nov. Sys t. P1. 1968: 67 F1. Reipub. Pop. Sin. 20 (2): 297 (1 984). (1 968) & Willows USSR: 157 (1 968) ，sub Salix taraikensis f. latifoli α(Kimura) subgen. subgen. Vetrix (Dumortier) Dumortier; Chou Kimura in Eco 1. Rev. 13: 200 (1953). in in F1. Reipub. Pop. Sin. 20 (2): 293 (1 984). Distr.: Japan: Hokkaido (Kushiro and Ty pe: Sα lix cα prea L. Nemuro Prov.) ，China and East Russia: Salix [unranked] Capreae Bluff & Primorye ，Khabarovsk ，Sakhalin ，Amur and Fingerhuth ，Comp. F1. Germ. 2: 565 (1 825); Siberia. Dumortier ， F 1. Belg. Prodr.: 11 (1 828). Salix Salix cohors Capreae Koch ，Salic. Eur.: 29. Salix caprea L. ， Sp. P1.: 1020 (1753); 11 11 (1 828); Nakai ， F l. Sylv. Kor. 18: 122 Tokubuchi in Bo t. Mag. Tokyo 10: page (1 930). (1 23) (1 896); Miyabe ， F 1. Kuri 1. Is 1.: 260 Salix Salix sec t. Capreae (Bluff & Fingerhuth) (1 890); Seemen ， Salic. Jap.: 33 ， tab. 6 February February 2000 Journal of Japanese Botany Vo l. 75 No. 1 29

(1 903); C.K.Schneid. in Sarg. ， P l. Wils. 3: Kimura in Sc i. Rep. Tohoku Univ. 4 ser. 149 (1 916); Miyabe & Kudo ， Icon. Ess. (Bio l.) 28: 42 (1 962). Forest Forest Tr. Hokkaido 1: 57 ， tab. 15 (1 921); [Salix bαkko f. angustifrons (Koidz.) Rech. f. in F l. Europ. 1: 50 (1964); Sugimoto ，New Keys Jap. Tr.: 101 (1 961) ， Skvortsov ，Willows USSR: 166 (1 968); comb. nud.]. Kitag. ，Neo-Lineam. F l. Mansh.: 204 Salix caprea f. subglabra (Chang & Skv.) (1979); (1979); Sohma in Sci. Rep. Tohoku Univ. 4 Kitag. ，Neo-Lineam. F l. Mansh.: 204 ser. ser. (Bio l.) 40: 108 (1 993); Nedoluzhko in (1 979). Charkevicz ， P l. Vascu l. Ori. Extr. Sovie t. 7: Distr.: J apan: Hokkaido ，Honshu (Kinki 176 (1 995). District and eastward) and Shikoku ，Korea ， Salix Salix hultenii Flod. in Arkiv Bo t.， Stockh. Russia and Europe. 20A: no. 6， 51 (1 926); Kimura in J. Fac. Note: According to Flora Europaea (1 993 ， Agric. Agric. Hokkaido Imp. Univ. Sapporo 26: p. 55) ，Sα lix caprea is characterized in the 414 (1 934); Kitam. & Murata ，Co l. Il lus t. keyas “ Leaves 1.5-2 times as long as wide ， Woody Pl. Jap. 2: 316 (1 979); Kimura in ovate or obovate to ovate-oblong; no ridges Wild Flow. Jap. Wood. 1: 42 (1 989). beneath the bark." In the description (p. 61) ， Salix Salix bakko Kimura in Bo t. Mag. Tokyo S. caprea is characterized as “decorticated 42: 42: 568 ，fig. 1 (1 928) [Ty pe: Japan ，Honshu ， wood without ridges ，and leaves 5-10 cm ， Kanagawa Pref. (Prov. Sagami) ，Hakone ， A. 1. 5-2 times as long as wide ，" and “ leaves Kimura 571 (♀ fl.) ， 12 Apr. 1927 (TUS-K usually broadly ovate to ovate to ovate-ob- holo.)]; holo.)]; Makino & Nemoto ， F l. Jap. ed. 2: long ，rounded at the base ，……; lateral veins 162 (1931); Kimura in J. Fac. Agric. 6-9 pairs ， forming almost a right angle with Hokkaido Imp. Univ. Sapporo 26: 417 the midrib." Kimura (1 928) ， in the original (1 934); Makino ，Il lus t. Fl. Nippon: 673 ，fig. description of Salix bakko ， pointed out that 2017 (1940); Ohwi ， F l. Jap. ed. Eng l.: 365 “Affinis S. capreae L. et S. cinereae L. ， sed (1 965) & F l. Jap. ed. rev.: 470 (1965); a prima praecipue foliis angustioribus Kurata ，Illus t. Imp. For. Tr. Jap. 3: 4， p l. 2 plurinervatis ，ligno nudo vibicibus (1971); (1971); Sugaya in Terasak i. Il lus t.: 101 ，tab. instructis ，…… facile distinguitur." In his de- 390 (1 977); Kitam. & Murata ，Co l. Il lus t. scription ， leaves are 8.7-13 cm long and ， Woody P l. J ap. 2: 317 ， fig. 770 (1 979); 3.5-4.1 cm broad with 10-13 nerves ，and Chou in F l. Reipub. Pop. Sin. 20 (2): 301 showed a clear photo of ridges on decorti- (1 984); Kimura in Wild Flow. Jap. Wood. 1: cated wood. Salix hultenii is intermediate 42 ，photo 47 ，1-5 (1 989); Makino ，New between S. caprea and S. bakko. It is distrib- Il lus t. Fl. Jap. rev. ed.: fig. 97 (1989); Ohwi uted in Hokkaido ，Kamchatka ， Sakhalin and & Kitag. ，New Fl. Jap.: 536 (1 992); Sohma southern Kurile Islands. This species has in in Sci. Rep. Tohoku Univ. 4 ser. (Bio l.) 40: broadly elliptic or elliptic leaves with acute 94 (1 993). or obtuse apex ，rounded or slightly cordate Salix Salix hultenii var.α ngustifolia Kimura in base. Decorticated wood with or without J. J. Fac. Agric. Hokkaido Imp. Univ. 26: 415 ridges. Salix caprea and S. bakko seems to (1934) [Type: Kamtschatka. ad urb. be a geoclinal form of one species. Petropaulowsk. Petropaulowsk. Y. Kobayashi s.n. (fo l.) ， 15 Skvortsov (1 968) treated S. bakko as a syn- Aug. 1931 (TI holo.)]; Ohwi ， F l. Jap.: 403 onym of S. caprea. His treatment is sup- (1 953); Ohwi ， F l. Jap. ed. Eng l.: 365 (1 965) ported in this paper. & F l. Jap. ed. rev.: 470 (1 965); Ohwi & Kitag. ，New Fl. Jap.: 536 (1 992). 30. Salix futura Seemen ，Salic. Jap.: 71 ， Salix Salix hultenii f.α ngustifolia (Kimura) tab. 17~fig. F-G (1 903); Kimura in Sci. Rep. 30 植物研究雑誌第75 巻第l号 平成12 年2月

Tohoku Imp. Univ. 4 ser. (Bio l.) 13: 73 31. Salix vulpina Andersson in Mem. (1 938); Ohwi ， F l. Jap. ed. Eng l.: 366 (1 965) Am. Acad. Arts Sci. n. s. 6: 452 (1 859); & Fl. Jap. ed. rev.: 472 (1 965); Kitam. & Franch. & Sav. ，Enum. P l. Jap. 1: 461 Murata ，Co l. Illus t. Woody Pl. Jap. 2: 322 ， (1 875); Tokubuchi in Bo t. Mag. Tokyo 10: fig. fig. 77 (1 979); Nova Fl. Nasu. Supp l.: 113 ， page (1 28) (1896); Kawakami in Bo t. Mag. figs. figs. 57-60 (1 985); Kimura in Wild Flow. Tokyo 10: 50 (1 896); Seemen ，Salic. Jap.: Jap. Jap. Wood. 1: 44 ，photo 51 ，4-6 (1 989); 37 ， tab. 5 (1 903); Koidz. in Bo t. Mag. To- Makino ，New Il lus t. Fl. Jap. rev. ed.: fig. 96 kyo 27: 89 & 264 (1 913); Schneid. in Sarg. ， (1 989); Ohwi & Kitag. ，New Fl. Jap.: 537 Pl. Wils. 3: 130 (1 916); Kimura in J. Fac. (1 992); Sohma in Sci. Rep. Tohoku Univ. 4 Agr. Hokkaido Imp. Univ. 26: 404 (1 934); ser. ser. (Bio l.) 40: 94 (1 993). Makino ，Il lus t. Fl. Nippon: 672 ，fig. 2016 Salix Salix vulpinoides Koidz. in Bo t. Mag. (1 940); Kimura in Sci. Rep. Tohoku Univ. Tokyo 27: 94 (1 913) [Ty pe: Japan. Honshu ， 4 ser. (Bio l.) 23: 9 ，figs. 2-3 ，tab. 3 (1 957); Gumma Pre f. (Prov. Kozuke). Collector un- Ohwi ， F l. Jap. ed. EngL: 336 (1 965) & Fl. known (仕.)， 4 May 1888 (TI lecto. desig- Jap. ed. rev.: 472 (1 965); Skvortsov ，Wil- nated nated by Kimura 198 1)]. lows USSR: 165 (1 968); Sugaya in Terasak i. Salix ν ulpina var. tomentosa Koidz. in Il lus t.: 102 ， fig. 395 (1977); Kitam. & Bo t. Mag. Tokyo 27: 265 (1 913) [Ty pe: Ja- Murata ，Co l. Il lus t. Woody Pl. Jap. 2: 323 ， pan. pan. Honshu ，Ishikawa Pref. (Prov. Kaga) ， fig. 779 (1 979); Lauener & McKean in Kanazawa [Kanazawa-hukin]. T. Ichimura Notes RBG Edinb. 41: 178 (1983); Kimura s.n. s.n. (仕.)， (TI lecto. designated by Kimura in Wild Flow. Jap. Wood. 1: 45 ，photo 51 ， 1981)]; 1981)]; Sugimoto ，New Keys Jap. Tr.: 106 1-3 (1 989); Makino ，New Il lus t. Fl. Jap. rev. (1 961). ed.: fig. 95 (1 989); Ohwi & Kitag. ，New Fl. Salix Salix futura Seemen var. psilocarpa Jap.: 537 (1992); Sohma in Sc i. Rep. Kimura in Sci. Rep. Tohoku Imp. Univ. 4 Tohoku Univ. 4 ser. (Bio l.) 40: 111 (1 993); r. se r. (Bio l.) 11: 247 (1 936) [Type: Japan. Nedoluzhko in Charkevicz ， P l. Vascu l. Ori. Honshu ，Fukushima Pref. (Prov. Iwaki) ， Extr. Sovie t. 7: 173 (1 995). Kosekimura. A. Kimura 884 (♀日.)， 23 Apr. Salix vulpina var. discolor Seemen ，Salic. 1934 (TUS-K holo.)]; Kitam. & Murata ， Jap.: 40 (1 903); Makino & Nemoto ， F l. Ja- Co l. Il lus t. Woody P l. Jap. 2: 322 (1 979); pan ed. 2: 174 (1 93 1). Ohwi ， F l. Jap. ed. Eng l.: 366 (1 965); Ohwi Salix daiseniensis Seemen ，Salic. Jap.: 66 & Kitagawa ，New Fl. Jap.: 537 (1 992). (1903) ， p. min. p. ，inc l. specim. ex Distr.: Distr.: Endemic to Japan: Honshu (South- Rebunsiri ， Faurie 3712. [白 de Kimura in Sci. ern ern Tohoku to Kinki District). Rep. Tohoku Imp. Univ. 4 ser. (Bio l.) 15: 30-1. 30-1. Salix futura f. futura 413 (1 940)]. 30-2. 30-2. Salix futura f. rufa (Kimura) Salix shiraii var.ν ulcaniana H.Lev l. in H.Ohashi ，sta t. nov. Bul l. Acad. Geog. Bo t. 14: 209 (1904) Salix ルtura Seemen var. rufa Kimura in [Ty pe: Japan. Hokkaido ， volcan de Tarumai ， Sci. Sci. Rep. Tohoku Imp. Univ. 4 ser. (Bio l.) 17 June 1893. U. Faurie 10026 ，U. Faurie 13: 13: 73 ，fig. 2 (1 938) [Ty pe: Japan ，Honshu ， 10027. (E-syntypes)]. Fukui Pref. (Prov. Etizen) ，Shimeimura- Salix ignicoma H.Lev l. in Bul l. Acad. Mizuoti ， Y. Hori 23 (fo l.)， 28 Aug. 1937 Geog. Bo t. 16: 143 (1 906) [Type: Japan. (TUS-K holo.)]. Honshu ， in sylvis Aomori ，May 1904. U. Distr.: Distr.: J apan: Honshu (Toyama and Fukui Faurie 5763 (E holo.)]. Prefectures) Prefectures) . Salix ν ulpina var. coriacea Koidz. in Bo t. Mag. Tokyo 27: 90 (1 913) [Type: Japan. February February 2000 Journal of Japanese Botany Vo l. 75 No. 1 31

Hokkaido ，Prov. Oshima ， Hakodateyama. E. Illus t. Woody P1. Jap. 2: 334 (1 979); Ohwi Tokubuchi Tokubuchi s.n. (fo 1. & fr.) ，26 Ju 1. 1888 (TI & Kitag. ，New F1. Jap.: 542 (1 992). lecto.)]. lecto.)]. Salix x yamatoensis Koidz. in Bo t. Mag. Salix Salix vulpina var. sub α lpina Koidz .i n Bo t. Tokyo 39: 2 (1 925) [Ty pe: Japan. Honshu ， Mag. Tokyo 28: 285 (1 914) [Type: Japan. Nara Pre f. (Prov. Yamato) ， M t. Honshu ，Yamagata Pre f. (Prov. Uzen) ， Butsuryjigoye (sphalm. Butsuyujigoye) ， Gassan. Gassan. G. Koidzumi s.n. (fr.) ，Aug. 1911 Kongosan. G. Koidzumi s.n. (fo 1.) [cult. in (TI (TI holo.)]; Sugimoto ，New Keys Jap. Tr.: Horto Bo t. Kyoto Univ. ，Ju 1. 20 ， 1928]. 106 106 (1 961). (KYO lecto. desiganated by Kimura 1981)]; [Salix [Salix alopochroa KimuI ・a in Sci. Rep. Kitam. & Murata ，Co l. Il lus t. Woody Pl. Jap. Tohoku Imp. Univ. 4 ser. (Bio 1.) 12: 100 2: 334 (1 979). (1937) ，nom. illeg. sine descrip t.]: Ohwi ， F 1. Salix x ishikawae Kimura in Sci. Rep. Jap. Jap. ed. rev.: 472 (1 965); Kitam. & Murata ， Tohoku Imp. Univ. 4 se r. (Bio 1.) 12: 100 ， Co 1. Il lus t. Woody P1. Jap. 2: 323 ，fig. 778 figs. 3-4 ，tabs. 5-6 (1 937) [Type: Japan. (1 979); Kimura in Wild Flow. Jap. Wood. 1: Honshu ，Hyogo ， Prov. Settu ，monte 45 ， photo 51 ，7-8 (1 989); Ohwi & Kitag. ， Rokkosan. E. Isikawa 2 (♀ f1.)， 23 Apr. New F l. Jap.: 538 (1992) ， as “ S. 1935 (TUS-K holo.)]; Ohwi ， F 1. Jap.: 404 alopechroa ぺSohma in Sc i. Rep. Tohoku (1 953) & F1. Jap. ed. Eng 1.: 368 (1 965); Univ. Univ. 4 ser. (Bio 1.) 40: 131 (1 993). Ohwi & Kitag. ，New F1. Jap.: 542 (1 992). Dist r.: Japan: Hokkaido and Honshu Salix xα samensis Sugimoto: Ohwi & (Tohoku and northern Kanto Districts) and Kitag. ，New F1. Jap.: 542 (1 992). South South Kuriles. =Sα lix udensis Trautv. & Mey. (as S. sachalinensis sachalinensis F.Schmidt) x S. vulpina Putative Putative natural hybrids of Salix Andersson (or as S. alopochroa Kimura). H3. Salix x arakiana Koidz. in Acta Hl. Salix x algista C.K. Schneid. in Phytotax. Geobo t. 1: 176 (1 932) [Ty pe: Ja- Sarg. ， P l. Wils. 3: 174 (1 916) ， pro sp. [Ty pe: pan. Honshu ，Kyoto ， Prov. Tanba [Tango] ， Japan. Japan. Prov. Suruga ， slopes of Fuji-san ， Kasagun ，Katomura ， M t. Karasugadake. Y. abundant abundant above alt. 300 m. E. H. W i1 son Araki 36 (ster.) ，8 Jun. 1930 (KYO lecto. 6637 6637 (♀ fl.) ，7 May 1914 (TI iso.)]; Kimura designated by Kimura 1981 ，TUS-K in in Bo t. Mag. Tokyo 42: 566 (1 928) ， pro sp.; isolecto.)]. Ohwi & Kitag. ，New F1. Jap.: 542 (1 992). = Salix gracilistyla Miq. x S. udensis = Salix. jessoensis Seemen subsp. Trautv. & Mey. (as S. sachalinensis serissaefolia serissaefolia (Kimura) H.Ohashi (as S. F. Schmidt)(?). serissaefolia serissaefolia Kimura) x Salix reinii Seemen. H4. Salix caprea L. (as S. bakko Kimura) H2. H2. Salix x ampherista C.K.Schneid. in x S. futura Seemen: Nova F1. Nasu. 145 Sarg. ， P 1. Wils. 3: 175 (1 916) [Ty pe: Japan. (1 972) & 1. c. Supp 1.: 118 ，fig. 79 (1 985); Hokkaido ，Prov. Osima ，Hakodate. U. Faurie Kitam. & Murata ，Co l. Illus t. Woody Pl. Jap. 5758 (f r.)， 1 Jun. 1904 (A holo.) ， fide 2: 334 (1979). Kimura 1940]; Makino & Nemoto ， F 1. Jap. H5. Salix x cremnophylla Kimura in Sci. ed. ed. 2， 161 (1931); Kimura in J. Fac. Agr. Rep. Tohoku Imp. Univ. 4 ser. (Bio 1.) 12: Hokkaido Imp. Univ. 26: 452 (1934); 97 ，figs. 1-2 ，tabs. 3-4 (1 937) [Ty pe: Japan. Kimura in Sci. Rep. Tohoku Univ. 4 se r. Honshu ，Kanagawa Pref. (Prov. Sagami) ， in (Bio 1.) 15: 410 ，fig. 6 ，tabs. 15-16 (1 940); ripa fl. Sakawagawa ，Hukusawamura- Koit i. Ohwi ， F 1. Jap.: 404 (1 953) & F1. Jap. ed. A. Kimura 2367 (fo 1.)， 16 May 1935 (TUS- Eng 1.: 367 (1 965); Kitam. & Murata ，Co 1. K lecto. designated by Kimura 1981)]; 32 植物研究雑誌第75 巻第1号 平成 12 年2月

Ohwi ， F 1. Jap.: 407 (1 953) & F1. Jap. ed. Trautv. & Mey. (as S. sach α linensis Eng 1.: 368 (1 965); Kitam. & Murata ，Co 1. F. Schmidt): Nova Fl. Nasu: 118 (1 985). Illus t. Woody P1. Jap. 2: 333 (1 979); Ohwi H10. Salix x gracilistyloides Kimura in & Kitag. ，New F1. Jap.: 542 (1 992). Bo t. Mag. Tokyo 40: 8 (1926) ， pro sp. = Salix gracilistyla Miq. x S. japonica [Ty pe: Japan. Honshu ，Hyogo Pref. (Prov. Thunb. Thunb. Settsu) ，monte Rokkosan. A. Kimura 152 H6. Salix x eriocataphylla Kimura in (♀ fl.) ，27 Apr. 1924 (TI holo. ，TUS-K. Bo t. Mag. Tokyo 40: 7 (1 926) ， pro sp. iso.)]; Kitam. & Murata ，Co l. Il lus t. Woody [Type: [Type: Japan. Honshu ，Shizuoka ， Prov. P1. Jap. 2: 333 (1 979); Ohwi & Kitag. ，New Suruga ，prope Subashiri ，pede montis Fl. Jap.: 542 (1 992). Fujisan. Fujisan. A. Kimura 198 (♀ f1.)， 12 May Sa !i x x gracilistyloides mst r. 1924 1924 (TI holo; TUS-K iso)]; Ohwi & Kitag. ， sympolyandra Kimura in Sci. Rep. Tohoku New F1. Jap.: 542 (1992). Univ. 4 ser. (Bio 1.) 18: 549 (1950) [Ty pe: Salix Salix x eriocataphylla var. psilocarpa Japan. Prov. Settsu ， in paludibus graminosis Kimura in Sci. Rep. Tohoku Imp. Univ. 4 montis Rokkosan. A. Kimura 444 (♂ fl.) ， 13 ser. ser. (Bio 1.) 20: 208 (1 954) [Type: Japan. Apr. 1926 (TUS-K)]. Honshu ，Yamanashi Pre f. (Prov. Kai) ， = Salix gracilistyla Miq. x S. vulpina Wadatoge ，Aikawamura. S. Huno s.n. (♀ Andersson. fl.) ， 19 Apr. 1936 (TUS-K holo.)]. H11. Salix x hapala Kimura in Bo t. Mag. = Sa !i x caprea L.(as S. bakko Kimura) x Tokyo 59: 82 (1946) [Type: Japan. S.jα~ponica Thunb. Hokkaido ，Prov. Osima ，Onomura- Kaihotu. H7. H7. Salix x eriocataphylloides Kimura A. Kimura 2875 (♀ fl.) ， 10 May 1941 in in J. Jap. Bo t. 60: 269 ，figs. 1-2 (1 985) (TUS-K lecto. designated by Kimura 198 1)]; [Ty pe: Japan. Honshu ，Miyagi Pref. Prov. Ohwi ， F l. Jap.: 406 (1 953) & Fl. Jap. ed. Rikuzen ，cult. in Hor t. Bo t. Fac. Scien t. Eng 1.: 368 (1 965); Kitam. & Murata ，Co 1. Tohoku Univ. A. Kimura 3284 (♀ fl.) ，5 Illust. Woody P1. Jap. 2: 334 (1 979); Ohwi Apr. Apr. 1961 (TUS-K holo.)]. & Kitag. ，New F1. Jap.: 542 (1 992). Sa !i x bakko Kimura x S. shiraii Seemen: = Sa !i x miy αbeana Seemen subsp. Nova F1. Nasu.: 145 (1 972) & 1. c. Supp l.: gilgiana (Seemen) H.Ohashi (as S. gilgiana 118 ，fig. 79 (1 985). Seemen) x S. integra Thunb. = Sa !i x caprea L. (as S. bakko Kimura) x H12. Salix x hatusimae Kimura in Sci. S. S. shiraii Seemen. Rep. Tohoku Imp. Univ. 4 ser. (Bio l.) 10: H8. H8. Salix x euerata Kimura in Bot. Mag. 560 ，figs. 6-8 ，tab. 9 (1 935) ， as hatusimai Tokyo 59: 81 (1946) [Type: Japan. [Ty pe: Japan. Kyushu ，Hukuoka Pref. (Prov. Hokkaido ，Prov. Oshimo ，Onomura- Kaihotu. Chikuzen) ，Takenoshita- Tokozi in vicinia A. Kimura 2879 (♂ fl.) ， 10 May 1941 urbis Hukuoka. S. Hatusima s.n. (♀ fl.) ，2 (TUS-K lecto. designated by Kimura 1981)]; Aug. 1933 (TUS-K holo.)]; Ohwi ， F l. Jap.: Ohwi ， F 1. Jap.: 406 (1 953) & F1. Jap. ed. 407 (1 953) & Fl. Jap. ed. Eng 1.: 368 (1 965); Eng l.: 367 (1 965); Kitam. & Murata ，Co l. Kitam. & Murata ，Co l. Il lus t. Woody Pl. Jap. Illus t. Woody Pl. Jap. 2: 334 (1 979); Ohwi 2: 333 (1 979); Ohwi & Kitag. ，New F l. Jap.: & Kitag. ，New Fl. Jap.: 542 (1 992). 542 (1 992). = Sa !i x miyabeana Seemen subsp. = Salix gr αcilistyla 恥1iq. x S. integra gilgiana gilgiana (Seemen) H.Ohashi (as S. gilgiana Thunb. Seemen) x S. udensis Trautv. & Mey. (as S. H13. Salix x hayatana Kimura in Bo t. sacha !i nensis F. Schmidt). Mag. Tokyo 42: 572 (1 928) ， pro sp. [Ty pe: H9. Salix futura Seemen x S. udensis J apan. Kyushu ，Saga Pre f. (Prov. Hizen) ， February February 2000 Journal of Japanese Botany Vo l. 75 No. 1 33

Saga ，cul t. A. Kimura 1286 (♀ fl.) ，30 Mar. Suruga) ，M t. F吋isan. K. Hisauchi s.n. (♀ 1927 (TI holo. ，TUS-K iso.)]; Ohwi & f1.)， 19 May 1918 (TI holo.)]; Kimura in Kitag. ，New F1. Jap.: 542 (1992). Bo t. Mag. Tokyo 40: 635 (1 926) ， pro sp.; = Salix eriocarpa Frach. & Sav. or S. Kimura in Ishii ，Engei Da りiten 6: 2439 pierotii pierotii Miq. (as S. koreensis Andersson) x (1 956); Kitam. & Murata ，Co l. Il lus t. Woody S. S. gracilistyla Miq. Pl. Jap. 2: 334 (1 979); Ohwi & Kitag. ，New H 14. Salix x hiraoana Kimura in Bo t. Fl. Jap.: 542 (1 992). Mag. Tokyo 40: 9 (1 926) ， pro sp. [Type: = Salix integra Thunb. x S. japonica Japan. Japan. Honshu ，Hyogo Pre f. (Prov. Settsu) ， Thunb. monte Rokkosan. A. Kimura & G. Hirao H16. Salix x ikenoana Kimura in Sc i. 138 138 (♀ fl.) ，27 Apr. 1924 (TI holo. ，TUS-K Rep. Tohoku Imp. Univ. 4 ser. (Bio 1.) 13: iso.)]; iso.)]; Kimura in Bo t. Mag. Tokyo 40: 633 71 ，fig. 1， tabs. 6-7 (1938) [Type: Japan. (1 926) ，pro sp.; Kimura in Ishii ，Engei Honshu ，Miyagi Pre f. (Prov. Rikuzen) ， Daijiten Daijiten 6: 2438 (1 956); Kitam. & Murata ， Sendai. A. Kimura 2350 (♂ fl.) ， 15 Apr. Co 1. Il lus t. Woody P1. Jap. 2: 334 (1 979); 1937 (TU ふK holo.)]; Kimura in Bo t. Mag. Ohwi & Kitag. ，New F1. Jap.: 542 (1 992). Tokyo 40: 635 (1 926); Ohwi ， F l. Jap.: 406 Salix x hiraoana Kimura va r. (1 953) & F l. Jap. ed. Eng l.: 367 (1 965); brachystachys brachystachys Kimura in Bot. Mag. Tokyo Kimura in Ishii ，Engei Daijiten 6: 2439 40: 40: 634 (1926) [Type: J apan. Honshu ， (1956); Kitam. & Murata ，Co l. Il lus t. Woody Hyogo Pref. (Prov. Settsu) ，monte Pl. Jap. 2: 334 (1 979); Ohwi & Kitag. ，New Rokkosan. A. Kimura 344 (♀ fl.) ， 15 Apr. Fl. Jap.: 542 (1 992). 1926 (TI holo. ，TUS-K iso.)]. Salix x ikenoana f. subcordata Kimura in Sa !i x x tsugaluensis Koidz. ， F 1. Symb. Sci. Rep. Tohoku Univ. 4 ser. (Bio l.) 23: 17 ， Or.-Asia.: Or.-Asia.: 33 (1 930) [Ty pe: Japan. Honshu ， fig. 1 ，tab. 1 (1 957) [Ty pe: Japan. Honshu ， Aomori Pref. ，Hirosak i. U. Faurie 2154 ， Tochigi Pref. (Prov. Shimotsuke) ， in planitie

May 1898 (P holo.)]; Kimura in Sci. Rep. graminoso ・herbosa in Imperiali Praedio Tohoku Univ. 4 ser. (Bio 1.) 19: 187 ，fig. 1， Rustico Nasuensi ，altitudine circa 680 m tabs. tabs. 3-4 (1 952); Kitam. & Murata ，Co 1. s.m. Augustissimus Imperator Japane 4137

Il lus t. Woody Pl. Jap. 2: 334 (1 979); Ohwi (fo 1.)， 3 Sep t. 1956 (TUS 田 K holo.)]. & Kitag. ，New Fl. Jap.: 542 (1 992). = Salix integra Thunb. x S. udensis Salix Salix x tsugaluensis var. 1αtifolia Kimura Trautv. & Mey. (as S. sachalinensis in in Sci. Rep. Tohoku Univ. 4 ser. (Bio 1.) 19: F. Schmidt). 189 ，fig. 2 (1952) [Ty pe: Japan. Honshu ， H17. Salix integra Thunb. x Salix Miyagi Pre f. (Prov. Rikuzen) ， Sendai. A. miyabeana Seemen: Check List Higher P1. Kimura 3078 (fo 1.)， 14 Oc t. 1949 (TUS-K Hokkaido 3: 19 (1994). holo.)]. holo.)]. H18. Salix x iwahisana Kimura in Sc i. Salix Salix x hiraoana f. brachystachys Rep. Tohoku Imp. Univ. 4 ser. (Bio 1.) 12: (Kimura) Kimura in Eco l. Rev. 13: 199 314 ，fig. 3， tabs.12-13 (1 937) [Ty pe: Japan. (1 953). Hokkaido ， Prov. Shiribeshi ，ad ripas fl. 'Salix 'Salix x tsug αluensis f. latifoli α(Kimura) Hugoppe ， in vicino oppidi Yoit i. 1. Kimura in Eco 1. Rev. 13: 201 (1 953). Yamamoto 6641 (♂ fl.)， 25 Ap r. 1937

= Sa !i x integra Thunb. x S. vulpina (TUS 悶 K holo)] & 26: 1 ，fig. 1， tabs. 1-2 Andersson (or S. alopochroa Kimura). (1 960); Ohwi ， F 1. Jap.: 405 (1 953) & F 1. H15. Salix x hisauchiana Koidz. in Bo t. Jap. ed. Eng l.: 368 (1965); Kitam. & Mag. Tokyo 33: 114 (1919) ， pro sp. [Ty pe: Murata ，Co 1. Il lus t. Woody Pl. Jap. 2: 333 Japan. Honshu ，Shizuoka Pref. (Prov. (1 979); Ohwi & Kitag. ，New F1. Jap.: 542 34 植物研究雑誌第75 巻第l号 平成 12 年2月

(1992). (1992). 2282 (♀ fl.) ，28 May 1933 (TUS-K iso.)] & Salix Salix x iwahisana var. psilocarp αKimura in Bo t. Mag. Tokyo 59: 85 (1946); Check in in Sc i. Rep. Tohoku Univ. 4 ser. (Bio 1.) 26: List Higher P1. Hokkaido 3: 20 (1 994). 4 (1 960) [Type: J apan. Hokkaido ， Prov. Salix x praegravis Kimura in J. Jap. Bo t. Tokachi ，Kawanishimati-Inata ， Obihiro. K. 60: 225 ， figs. 1-2 (1 985) [Type: Japan. Togasi 21 (♀ fl.) ，5 Apr. 1959 (TUS-K Honshu ， Tochigi Pref. (Prov. Shimotsuke) ， holo.)]. holo.)]. prope fontem calidum Asahionsen ，N asu. A. = Salix gracilistyla 恥1iq. x S. rorida Kimura 4639 (fo 1.)， 13 Sep t. 1970 (TUS-K Lacksch. Lacksch. holo.)]. H19. Salix x kamikotica Kimura in Sci. = Salix caprea L. (as S. bakko Kimura or Rep. Rep. Tohoku Imp. Univ. 4 ser. Bio 1. 6: 194 S.hultenii Flod. var. angustifolia Kimura) x (1 931) ，pro sp. [Type: J apan. Honshu ， S. udensis Trautv. & Mey. (as S. schalinensis Nagano Pre f. (Prov. Shinano) ， Kamitoti. A. F. Schmid). Kimura 782 (♂ fl.) ， 16 May 1930 (TUS-K. Distributed in J apan: Rishiri and Rebun holo.; holo.; iso. TUS ，TUS-K)]; Nemoto ， F 1. Jap. Islands in Hokkaido. Supp 1.: 109 (1 936) H22. Salix x koiei Kimura in Sci. Rep. x Toisochosenia kamikotica (Kimura) Tohoku Imp. Univ. 4 ser. (Bio 1.) 15: 407 ， Kimura in Sci. Rep. Tohoku Imp. Univ. 4 fig. 4 ， tabs. 13-14 (1940) [Type: Japan. ser. ser. (Bio 1.) 11: 248 ，figs. 2-3 (1 936); Ohwi ， Honshu ， Akita Pref. (Prov. U go) ，Huzikoto- 1. F1. Jap.: 398 (1 953) ，as “kamikochica" ，l. c. ， mura. G. Koie 34 (♂ f1.)， 21 Apr. 1940 ed. ed. Eng 1.: 362 (1 965) ，1. c. ed. rev.: 465 (TUS ・K lecto. designated by Kimura 1981)]; (1 965); Sugimoto ，New Keys Jap. Tr.: 107 Ohwi ， F 1. Jap.: 407 (1 953) & F l. Jap. ed.

(1 96 1); Hatusima ，Nihon-no 圃 jyumoku: 690 Eng 1.: 368 (1 965); Kitam. & Murata ，Co 1. (1 976); Kitam. & Murata ，Co 1. Il lus t. Wood. Illus t. Woody Pl. Jap. 2: 333 (1 979); Ohwi Jap. Jap. 2: 337 (1 979); Ohwi & Kitagawa ，New & Kitag. ，New F1. Jap.: 542 (1 992). 1. F 1. Jap.: 530 (1 992) ，as “kamikochica" ; = Salix gracilistyla Miq. x S. miyabeana Sohma in Sci. Rep. Tohoku Univ. 4th ser. Seemen subsp. gilgiana (Seemen) H.Ohashi (Bio l.) 40: 137 ，figs. 296-301 (1 993) [pol- (as S. gilgiana Seemen). len]. len]. H23. Salix x lasiogyne Seemen ， Salic. =Sα lix arbutifolia Pall. (as Chosenia Jap.: 32 ， tab. 4. figs. A-C (1 903) ， pro sp. ， bracteosa bracteosa (Turcz. ex Trautv.) Nakai) x S. exc l. specim. ex Korea [Type: J apan. cardiophylla cardiophylla Trautv. &恥 1ey. var. urbaniana Kanagawa Pref. (Prov. Sagami) ， Yamakita. (Seemen) Kudo (as Toisusu urbaniana u. Faurie 3702 (♀)， 8 May 1899 (KYO (Seemen) Kimura). lecto. designated by Kimura 1984)]; Koidz. H20. Salix x kawamurae Kimura in J. in Bo t. Mag. Tokyo 27: 265 (1 913); Kimura Jap. Jap. Bo t. 46: 303 (1 971) [Ty pe: Japan. Prov. in 1. Jap. Bo t. 59: 271 (1 984). Shimotuke ，Myozintoge ，Kurobanemat i. A. = Salix babylonica L. x S. jessoensis Kimura 4636 (fo l.)， 11 Sep t. 1970 (TUS-K Seemen subsp. serissaefolia (Kimura) holo.)]; holo.)]; Kitam. & Murata ，Co 1. Il lus t. H.Ohashi (as S. serissaefolia Kimura). Woody P1. Jap. 2: 333 (1 979). H23-1. Salix x lasiogyne nothosubsp. = Salix futura Seemen x S. gracilis η la lasiogyne Miq. H23-2. Salix x lasiogyne nothosubsp. H21. Salix x koidzumii Kimura in J. Fac. yuhkii (Kimura) H.Ohashi ，comb. nov. Agr. Agr. Hokkaido Imp. Univ. 26: 440 (1 934) Salix x yuhkii Kimura in Sc i. Rep. [Type: [Type: Sachalin. Dist r. Toyohara ，prope Tohoku Univ. 4 ser. (Bio 1.) 18: 544 ，fig. 1 oppid. oppid. Toyohara. A. Kimura & S. Sugawara (1 950) [Type: Japan. Honshu ，Yamagata February February 2000 Journal of Japanese Botany Vo l. 75 No. 1 35

Pref. Pref. (Prov. Uzen) ，Yonezawa ， cult. A. (1 916); Nova Fl. Nasu.: 145 (1 972). Kimura 3046 (♀ fl.) ，24 Apr. 1950 (TUS-K Salix futura Seemen x S. vulpina lecto. lecto. designated by Kimura 198 1)]; Kitam. Andersson: Nova F l. Nasu: 145 (1 972); & Murata ，Co l. Il lus t. Woody Pl. Jap. 2: 334 Kitam. & Murata ，Co l. Il lus t. Woody Pl. Jap. (1 979); Ohwi & Kitag. ，New F l. Jap.: 542 2: 334 (1 979). (1 992). = Salix futur αSeemen x S. vulpina = Salix babylonica L. (as S. babylonica Andersson. var. var. lavallei Dode) (cult.) x S. jessoensis Kimura (1981) commented on two Seemen subsp. jessoensis (as S. hondoensis syntypes of S. matsumurae as follows: Koidz.). Koidz.). “Adnotatio: ut ex explicatione Seemenii (l. c. H24. Salix x leucopithecia Kimura in p. 72) facile videtur ，specimina Wrightii Bo t. Mag. Tokyo 40: 11 (1 926) ，pro sp. (No. 901 ，212-441 ，Herb. Lugd. Ba t.) ex [Ty pe: Japan. Honshu ，Tokyo Pre f. (Prov. Simoda verisimiliter ad Salicem japonicam Musashi) ， in sepulcris Somei ，prope Tokyo. Thunb. s. l. sunt referenda ，quamquam C. K. A. Kimura 59 (♂ fl.) ，28 Mar. 1924 (TI Schneider (l. c. p. 132) haec pro typo habe t. holo.)]; holo.)]; Ohwi ， F l. Jap.: 403 (1 953) & F l. Specimen autem Matsumurai n. 20 ex Jap. Jap. ed. Eng l.: 365 (1 965); Kimura in Ishii ， Shimizutoge cum figuris Seemenii (tab. 18 ， Engei Da りiten 6: 2438 (1 956); Sugaya in figs. A-BI) maxime congruit; itaque hoc pro Terasaki. Terasaki. Il lus t.: 101 ， fig. 391 (1 977); isolectotypo habeo." Kitam. Kitam. & Murata ，Co l. Il lus t. Woody Pl. Jap. H26. Salix x microstemon Kimura in 2: 2: 333 (1 979); Kimura in Wild Flow. Jap. Bot. Mag. Tokyo 40: 636 (1926) ，pro sp. Wood. 1: 50 ， photo 57 ，6-7 (1 989); Makino ， [Ty pe: J apan. Honshu ，Okayama Pref. (Prov. New Il lus t. Fl. Jap. rev. ed.: fig. 98 (1 989); Bizen) ，ad ripas fluminis Yoshiigawa ，prope Kimura in J. Jap. Bo t. 59: 353 (1 984); Nova Wake. A. Kimura 461 (♂ fl.) ，20 Apr. 1926 l. F l. Nasu Supp l.: 117 ，figs. 76-78 (1 985); (TI holo. ， TUS-iso.)]; Ohwi & Kitag. ，New Ohwi & Kitag. ，New Fl. Jap.: 542 (1 992). Fl. Jap.: 542 (1 992). Salix Salix x yoitiana Kimura in Bo t. Mag. To- = Salix eriocarpa Frach. & Sav. x S. kyo 59: 83 (1946) [Ty pe: J apan. Hokkaido ， miy αbeana Seemen subsp. gilgiana Prov. Prov. Shiribeshi ， in ripa fl. Hugoppe ， prope (Seemen) H.Ohash i. Yoiti ，1. Yamamoto 8167 (♀ fl.) ，22 Apr. H27. Salix x nasuensis Kimura in Sci. 1938 (TU ふK lecto. designated by Kimura Rep. Tohoku Univ. 4 ser. (Bio l.) 28: 37 ，fig. 198 1)]; Ohwi ， F l. Jap.: 403 (1 953) & F l. 1， tabs. 1-5 (1 962) [Ty pe: Japan. Honshu ， Jap. Jap. ed. Eng l.: 366 (1965); Kitam. & Miyagi Pre f. (Prov. Rikuzen) ，Sendai ，cul t. Murata ，Co l. Il lus t. Woody Pl. Jap. 2: 333 A. Kimura 3295 (♀ fl.) ，6 Apr. 1961 (TUS- (1 979); Ohwi & Kitag. ，New Fl. Jap.: 542 K holo.)]; Kitam. & Murata ，Co l. Il lus t. (1992). (1992). Woody P l. Jap. 2: 334 (1 979); Ohwi & = Salix caprea L. (as S. bakko Kimura or Kitag. ，New Fl. Jap.: 542 (1 992). S. S. hultenii Flod. var. angustifolia Kimura) x = Salix futura Seemen x S. shiraii S. S. gracilistyla Miq. Seemen. H25. Salix x matsumurae Seemen ，Salic. Original collection: “ Prov. Shimotsuke: Jap.: Jap.: 71 ， tab. 18 A-B' (1 903) ，pro sp. ， ut ad saxa ， prope fontem calidum Kitaonsen ， in “ matsumuraei" [Ty pe: Japan. Honshu ，Mt. Imperiali Praedio Rustico Nasuensi ，A. Shimizutoge ， inter Gumma Pref. et Niigata Kimura 3307 ， 14 May 1959." Pref. Pref. 1. Matsumura 20 (fr.) ， 19 Ju l. 1886 (TI H28. Salix x pedionoma Kimura in Sci. isolecto. isolecto. designated by Kimura 1981)]; Rep. Tohoku Imp. Univ. 4 ser. (Bio l.) 10: Schneider in Sargent ， P l. Wilson. 3: 131 558 ， fig. 5 A-D ， tab. 18-1 (1 935) [Type: 36 植物研究雑誌第75 巻第1号 平成12 年2月

Sakhalin. Sakhalin. Distr. Toyohara ， prope oppidum Yubisogawa ，Yubiso ，altitudine circa 560 m Toyohara. Toyohara. S. Sugawara 1 (♀ fl.) ， 18 May s.m. A. Kimura & S. Hukao 4674 (fo l. 1934 1934 (TUS-K holo)]; Ohwi & Kitag. ，New adul t.)， 20c t. 1973 (TUS-K holo.)]. l. Fl. Jap.: 542 (1992). = Salix gracilistyl αMiq. x S. hukaoana = Salix caprea L. (as S. hultenii Flod. var. Kimura. angust ザolium Kimura) x S. schwerinii E. H32. Salix x sirakawensis Kimura in Wolf (as S. yezoensis (C.K.Schneid.) Sci. Rep. Tohoku Imp. Univ. 4 ser. (Bio l.) Kimura). Kimura). 12: 102 ，fig. 5 ，tabs. 7-8 (1 937) ， as x Salix Distributed Distributed in South Sakhalin and sir αkawensis [Type: J apan. Honshu ， Hokkaido. Hokkaido. Fukushima Pref. (Prov. Iwaki) ， Kosekimura. H29. H29. Salix x pseudopaludicola Kimura A. Kimura 875 (♂ fl.) ， 23 Apr. 1934 (TU ふ in in J. Fac. Agr. Hokkaido Imp. Univ. 26 (4): K holo.)]; Ohwi ， F l. Jap.: 404 (1953) & Fl. 445 (1934) [Ty pe: J apan. Hokkaido ， Prov. Jap. ed. Eng l.: 366 (1 965); Kimura in Sci. Ishikari ，Takanegahara ， in montibus Rep. Tohoku Univ. 4 ser. (Bio l.) 23: 7 ，fig. Nutakkamushupe. Nutakkamushupe. A. Kimura 1797 (♀ fl.) ， 1 ，tabs. 1-2 (1 957); Kitamura & Murata ，

13 13 Ju l. 1928 (TUS 聞 K holo.)]; Ohwi ， F l. Jap.: Co l. Illus t. Woody Pl. Jap. 2: 334 (1 979); 404 (1 953) & Fl. Jap. ed. Eng l.: 366 (1965); Ohwi & Kitag. ，New F l. Jap.: 542 (1992) ， as Kitam. Kitam. & Murata ，Co l. Illus t. Woody Pl. Jap. “shirakawaensis" . 2: 2: 334 (1979); Ohwi & Kitag. ，New F l. Jap.: Salix x sirakawensis f. pubescens Kimura 542 (1 992). in Sci. Rep. Tohoku Imp. Univ. 4 ser. (Bio l.) = Salix fuscescens Andersson (as S. 12: 103 (1 937) [Type: Japan. Honshu ， paludicola paludicola Koidz.) x S. reinii Seemen (as S. N agano Pref. (Prov. Shinano) ，N ojir i. K. hide 下1J oi Koidz.). Hisauchi 1722 (fo l.)， 12 Aug. 1936 (TUS-K H30. Salix x sendaica Kimura in Sci. holo.)] ， syn. nov. Rep. Rep. Tohoku Imp. Univ. 4 ser. (Bio l.) 6: 196 = Salix futura Seemen x S. integra Thunb. (1 931) [Ty pe: Japan. Honshu ， Miyagi Pref. H33. Salix x sugayana Kimura in Sc i. (Prov. (Prov. Rikuzen) ，colle Mukaiyama (sphalm. Rep. Tohoku Univ. 4 ser. (Bio l.) 20: 202 ， Mukoyama) ，Senda i. A. Kimura 809 (♀ fl.) ， figs. 1-3 ，tabs. 1-4 (1 954) [Ty pe: Japan. 28 Ap r. 1929 (TUS-K holo.)]; Kimura in Honshu ，Miyagi Pref. (Prov. Rikuzen) ， Sci. Sci. Rep. Tohoku Univ. 4 ser. (Bio l.) 18: 552 Sendai ，cul t. A. Kimura 3113 (♂ fl.) ，8 Apr. (1 950); Kitam. & Murata ，Co l. Il lus t. Woody 1953 (TUS- K lecto. designated by Kimura l. Pl. Jap. 2: 334 (1 979); Ohwi & Kitag. ，New 1981)]; Kitam. & Murata ，Co l. Il lus t. Woody l. Fl. Jap.: 542 (1 992). Pl. Jap. 2: 334 (1 979). Salix Salix x ultima Koidz. in Acta Phytotax. Salix x sugayana mstr. ternata Kimura in Geobo t. 8: 111 (1939) [Type: J apan. Sci. Rep. Tohoku Univ. 4 ser. (Bio l.) 20: Honshu ，Nara Pref. (Prov. Yamato) ， Mt. 206 ，fig. 4 (1 954) [Ty pe: Japan. Iwate Pre f. Kongosan. Kongosan. G. Koidzumi s.n. (fo l.)， Ju l. 24 ， (Prov. Rikutyu) ， secus fl. Sekigutigawa in 1930 (KYO lecto. designated by Kimura viculo Sekiya prope oppidulum Yamada. S.

1981)]; 1981)]; Kitam. & Murata ，Co l. Illus t. Woody Sasamura 1188 (fr.) ， 10 July 1949 (TUS 司 l. Pl. Jap. 2: 334 (1 979). K)]. = Salix caprea L. (as S.bakko Kimura) x = Salix integra Thunb. x S. rupifraga S. S. vulpina Andersson (or as S. alopochroa Koidz. Kimura). Kimura). Originally found in “Prov. Shinano: inter H3 1. Salix x sigemitui Kimura in J. Jap. saxa in ripis torrentis Azusagawa ， prope Bo t. 51: 1， fig. 1 (1 976) [Type: Japan. fontem calidum Sakamakionsen (collected Honshu ， Prov. Kozuke ， secus torrentem by A. Kimura and S. Sugaya on 14 May February February 2000 Journal of Japanese Botany Vo l. 75 No. 1 37

1950)". 1950)". Kitag. ，New F 1. Jap.: 542 (1992) ， as S. H34. Salix x sumiyosensis Kimura in Kimura. Bo t. Mag. Tokyo 40: 12 (1 926) [Ty pe: Ja- = *Salix japonica Thunb. (cu 1t.) x S. pan. pan. Honshu ，Hyogo Pref. (Prov. Settsu) ， vulpin αAndersson. co l1 e Kamokogahara (errore Osigaoka) in Sumiyoshi-mura. Sumiyoshi-mura. A. Kimura 228 (♀ fl.) ， 11 1 wish to express my sincere thanks to my Apr. Apr. 1925 (TI holo. ，TUS-K iso.)]; Ohwi & teachers from my beginning of tax- Kitag. ，New F1. Jap.: 542 (1 992). onomy: the late Professors Dr. Fumio =Sα lix gracilistyla 恥1iq. x S. yoshinoi Maekawa and Dr. Hiroshi Hara of the Uni- Koidz.(?). Koidz.(?). versity of Tokyo ， the late Professors Dr. H35. Salix x thaymasta Kimura in Bo t. Arika Kimura ， Dr. Sadao Sugaya and Dr. Mag. Tokyo 40: 641 (1 926) [Ty pe: Japan. Kankichi Sohma of the Tohoku University Honshu ，Hyogo Pref. (Prov. Settsu) ，monte for their invaluable suggestions for my stud- Rokkosan. E. Ishikawa s.n. (♂ fl.) ，8 Apr. ies on Salix. D r. Maekawa introduced me 1940 (TUS-K holo.)]; Kimura in Sci. Rep. Dr. Kimura for my desire to study Salix Tohoku Imp. Univ. 4 ser. (Bio 1.) 15: 409 ， when 1 was a high school studen t. Ms Emily fig. fig. 5 (1940) & in Bot. 孔1ag. Tokyo 59: 84 Wood of Harvard University kindly read the (1946) ， ut thaumast α; Ohwi ， F 1. Jap.: 405 manuscnpt. (1 953) & F1. Jap. ed. Eng 1.: 367 (1965); Kitam. Kitam. & Murata ，Co 1. Illus t. Woody P1. Jap. References 2: 2: 333 (1 979); NovaF 1. Nasu. Supp 1.: 117 ， Argus G. W. 1986. The genus Salix (Salicaceae) in the figs. figs. 73-75 (1 985); Ohwi & Kitag. ，New F1. Southeastern United States. Sys t. Bo t. Monog. 9. Jap.: Jap.: 542 (1 992). pp. 170. American Society of Plant Taxonomists ， US A. = Salix gracilistyla Miq. x Salix 一一一 1997. Infrageneric classification of Sα lix schwerinii E. Wolf (as S. kinuyan α gi (Salicaceae) in the New World. Sys t. Bo t. Monog. Kimura). Kimura). 52. pp. 121. American Society of Plant Taxono- H36. Salix x turumatii Kimura in Sc i. mists ，USA. Rep. Rep. Tohoku Imp. Univ. 4 ser. (Bio 1.) 11: Azuma T. ，K 吋ita T. ，Yokoyama J. and Ohashi H. 2000. Phylogenetic relationships of Sα lix 243 ，fig. 1. tabs. 7-8 (1 936) [Ty pe: Japan. (Salicaceae) based on rbcL data. Amer. J. Bo t. 86: Honshu ，Ibaraki Pre f. (Prov. Hitati) ， 67-75. Midorigaokamura. Midorigaokamura. H. Turumati s.n. (♂ fl.) ， Hatusima S. 1971. Flora of the Ryukyus (including 29 Mar. 1935 (TUS-K)] & 1. c. 12: 107 Amami Islands ，Okinawa Islands ，and Sakishima (1 937); Ohwi ， F 1. Jap.: 407 (1 953) & F l. Archipelago) (in Japanese). Okinawa Seibutu- Jap. Jap. ed. Eng l.: 368 (1965); Kitam. & kyouiku Kenkyuukai. Naha. 一一一 1978. Kagoshima-ken Shokubutsu Mokuroku. Murata ，Co 1. Il1 us t. Woody P1. Jap. 2: 333 234 pp. Kagoshima-ken shokubutsu dokokai. (1 979); Ohwi & Kitag. ，New F l. Jap.: 542 Kagoshima. (1 992). Kimura A. 1926. Contributiones ad Salicologiam = Salix caprea L. (as S.bakko Kimura) x Japonicam I. Bo t. Mag. Tokyo 40: 7-14. S. S. futura Seemen x S. gracilistyla Miq. 一一一 1928. Uber Toisusu ， eine neue Salicaceen- Gattung Gattung und die systematische Stellung derselben. H37. *Salix x japopina Kimura in J. Jap. Bo t. Mag. Tokyo 42: 287-290. Bo t. 43: 90 ， fig. 1 (1968) [Type: J apan. 一一一 1934. Salicaceae in Miyabe & Kudo ， Flora of Honshu ，Miyagi Pref. (Prov. Rikuzen) ， in Hokkaido & Saghalien IV. J. Fac. Ag r. Hokkaido horto horto Ins t. Bio l. Tohoku Univ. ，Senda i. A. Imp. Univ. 26 (4): 391-452. Kimura 3341 (fo 1.)， 11 Sep t. 1967 (TUS-K 一一一 1938. Symbolae It eologicae V I. 72) Systema novum Salicacearum ， sive emendationes et holo.)]; holo.)]; Kitam. & Murata ，Co l. I1 1us t. additamenta additamenta ad systema Salicacearum anno 1928 Woody P l. Jap. 2: 334 (1 979); Ohwi & ab auctore propositum. Sci. Rep. Tohoku Imp. 38 38 植物研究雑誌第75 巻第l号 平成 12 年2月

Univ. Univ. 4th se r. Bio l. 13: 390-394. Ohashi H. 1995. Salicaceae. In: Asahi Encyclopedia: 一一一 1944. De Salice subfragili Anderssonii. Acta The World of Plants 6: 226. Asahi-shinbun-sha ， Phytotax. Phytotax. Geobot. 13: 184-189 (in Japanese). Tokyo (in Japanese). 一一一 1981. List of the type specimens in the herbaria 一一一 1999. The genera ，tribes and subfamilies of Japa- of of Japan. A Working Group on Determination of nese Leguminosae. Sci. Rep. Tohoku Univ. 4 se r. Types and Authentic Specimens of Vascular (Bio l.) 40: 187-268. Plants. Plants. Ohwi J. 1953. Salicaceae. Flora of Japan. pp. 395- 一一一 1988. De Salicis subgenere Pleuradenia 408. Shibundo ，Tokyo (in Japanese). commentatio. commentatio. Sci. Rep. Tohoku Univ. 4th ser. Bio l. 一一一 1965a. Salicaceae. In: Flora of Japan (in En- 39 (3): 143-147. glish). pp. 361-369. Smithsonian Institution ，Wash- 一一一 1989. Salicaceae. In: Satake Y. et al. (eds.) ，Wild ington D.C. Flowers Flowers of Japan Woody Plants 1: 31-51 ，pls. 39- 一一一 1965b. Salicaceae. In: Flora of Japan ， Revised

58. 58. Heibonsha Ltd. ，Publishe r. Tokyo (i n Japa- Edition. pp. 462 -4 76. Shibundo ，Tokyo (i n Japa 司 nese). nese). nese). Kitagawa Kitagawa M. 1939. Lineamenta Florae Manshuricae. 一一- and Kitagawa M. 1992. Salicaceae. In: New Rep. Rep. Ins t. Sci. Res. Manchoukuo 3， Append. 1. Fl ora of Japan ，revised ed. pp. 527-542. Shibundo ， 一一一 1979. Neo- Li neamenta Florae Manshuricae. J. Tokyo (in Japanese). Crame r. Vaduz. Rechinger K. H. (edition 1) revised by Akeroyd J. R. Kitamura S. and Murata G. 1979. Salicaceae. In: 1993. Salix. In: Tutin T. G. et al. (eds.). Flora Kitamura Kitamura S. and Murata G. ， Coloured Ill ustrations Europaea 2nd edition. Cambridge Univ. Press ， of of Woody Plants of Japan 2: 303-340. Hoikusha Cambridge. Publishing Publishing Co. ， Ltd. Osaka (in Japanese). Schneider C. K. 1916. Salix. In: Sargent ， C. S. ， Koidzumi G. 1913. Spiciligium Salicum Japonensium Plantae Wilsonianae 3: 40-179. Publication of novarum aut imperfecte cognitarum. 1. Bo t. Mag. Arnold Arboretum No. 4. Cambridge ，Massachu- Tokyo 27: 87-97. II. Bot. Mag. Tokyo 27: 264- setts. 267. 267. Seemen O. von 1903. Salices Japonicae. Gebruder Leskinen Leskinen E. and Alstrom-Rapaport C. 1999. Molecu- Borntraege r. Leipzig. lar lar phylogeny of Salicaceae and closely related Shimabuku K. 1997. Check List Vascular Flora of the Flacourtiaceae: Flacourtiaceae: evidence from 5.8 S， ITS 1 and ITS Ryukyu Islands. Revised Edition. Kyushu Univer- 2 of the rDNA. Pl. Syst. Evo l. 215: 209-227. sity Press ， Fukuoka. Mizuno M. ， Kato M. ， Iinuma M. ，Tanaka T. ，Kimura Skvortsov A. K. 1968a. Nomenclature and typification A.， A.， Ohashi H. ， Sakai H. and K 吋ita T. 1989. Two in the genus Salix L. Subgenera ，sections and sub- chemical chemical races in Salix sachalinensis F.Schmidt sections pertaining to the flora of the USS R. Acad. (Salicaceae). (Salicaceae). Bo t. Mag. Tokyo 102: 403 -4 11. Sci. URSS Nov. Sys t. Plant Vasc. 1968: 62-72 (in Nakai T. 1928. Une nouve l1 e systematique des Russian). Salicacees Salicacees de Coree. Bull. Soc. Dendro l. France 一一一 1968b. Willows of the USS R. A taxonomic and 66: 66: 37-5 1. geographic revision. Nauka. Moscow. 262 pp. (in 一一一 1930. Salicaceae. In: Flora Sylvatica Koreana Russian).

18: 18: 19-218. The Forest Experiment Station ，Gov- Walker E. H. 1976. Flora of Okinawa and the south 国 ernment ernment General of Chosen. Keijyo. ern Ryukyu islands. Smithsonian Institution. Wash- Newsholme C. 1992. Willows ， the genus Salix. 224 ington D. C. pp. pp. Timber Press. Portland.

大橋広好:日本産ヤナギ属植物集覧 日本産ヤナギ属，ケショウヤナギ属およびオオ (栽培品種 l 変種 1 品種を含む) .これらの日本産 バヤナギ属の分類と学名および分類体系を再検討 種を新たに l 属 4 亜属 17 節に分類した.各分類群 した.本研究では，これら 3 属をヤナギ属にまと について，正名，シノニム，それらの文献をあげ， めた.日本産種としておよそ 40 種が知られていた 種以下には分布域を加えた.学名の多くについて が， 31 種 4 亜種 3 変種 16 品種に整理した.この はタイプを併記した.なお，各分類群の区別，記載 中にはシダレヤナギ，ウンリュウヤナギ，コリヤナ などは Iwatsuki et al. (eds.): Flora of Japan の ギなど3種2品種の外来種が含まれているので，日 Salicaceae に記述したので ここに省略した.また， 本の在来種は 28 種 4 亜種 3 変種'1 4 品種となった ヤナギ科と主な種については朝日百科・植物の世 February February 2000 Journal of Japanese Botany Vo l. 75 No. 1 39

界 6: 226-523 (1995) に解説した.雑種について 群について，その「学名+出典(+タイプ標本) + は， 37 雑種を認め，各学名について出典，タイプ， 文献引用;+シノニム j という形の表示方法は古く 文献を整理し，推定両親種の雑種式 hybrid formula から続いている分類学に特有のものである.この を示した. 形式によって，積み重ねてきた成果を含む文献を ヤナギ属の分類体系は Andersson (1868) の体系 学名の下に引用し その分類群についての情報を 以外には世界の全種類を基礎として設立されたも 整理して，過去の研究とのつながりを明らかにし のがなく，その後の研究は全て地域的な種類に基 てきた.引用文献の取捨選択によってその研究者 づく体系である.しかし，旧ソ連のヤナギ属をまと の当該分類群にたいする見解を同時に示すことも めた Skvortsov (1968a ，1968b) と北アメリカの できる. Argus (1988 ，1997) の研究が今日のヤナギ属分類 このような，いわゆる「集覧J Systematic Enu- 体系の基礎となっている.本論文では基本的にこ meration は分類群の研究にとって最も基本的な研 れらの体系と一致するように日本の種類の分類体 究成果であるといえよう.日本産種子植物に関し 系をまとめたが，冬芽鱗片のへりが離生している ては，そのおよそ半分の科について，含まれる属， 群は独立群と考え， トカチヤナギ亜属 subgen. 種，種内分類群の全てを扱った原寛:日本種子植 Urbanianae (Seemen) H.Ohashi とした. 物集覧 1-3 があり，今日でも非常に有用である. 今回の研究によって使い慣れた学名を変更した シダ植物では田川基二: Index Pteridophytorum ものもあるので，本文中に学名と和名の対照リス J aponicarum (原色日本羊歯植物図鑑)と中池敏之: トを示した.例えば，ケショウヤナギ，オオパヤナ Enumeratio Pteridophytorum Japonicarum ，Pilicales ギは属の変更に伴つ変更，タチヤナギ，オノエヤナ がある. ギ，キヌヤナギ(エゾノキヌヤナギを含む) ，パッ 日本のヤナギ科は原先生の集覧に含まれておら コヤナギ(マルバノバッコヤナギを含む)などは種 ず，それに代わるものもない.ヤナギ属の研究を始 を大きく範囲づけたための変更，カワヤナギはエ めるにあたって， Seemen (1903): Salices Japonicae ゾノカワヤナギの亜種としたための変更などであ から後の学名と過去の研究を整理することがまず る.また， ミネヤナギの著者は Prachet & Savatier 必要であった.日本のヤナギ属は木村有香先生が ではなく， Seemen であるとした. 最近まで長い間研究を続けておられ，その成果を 日本産ヤナギ属，ケショウヤナギ属，オオパヤナ 主として東北(帝国)大学理科報告と本誌に発表し ギ属は大井 (1953 ，1965a ， 1965b) ，北村・村田 てこられた.このため ヤナギ属(広義)の学名や (1979) (1979) ，木村 (1989) ，あるいは大井・北川( 1992) 論文などを探し出し易かった.また，日本産ヤナギ によって，詳しく記述されている. しかし，これら 属の多くのタイプ標本が東北大学大学院理学研究 の著作は日本フロラの一部としてこれらの属をま 科生物学教室植物標本館木村ヤナギ科標本室 とめたものであり，属，種の同定を目的としている (TUS-K) にあるので，学名と実物の対応も容易に ため，検索表，記載，分布などの記述が主であり， 検討できた.しかし，本研究では Andersson ， 属内分類群，個々の分類群の学名の出典と研究論 Blume ，Miquel ，Seemen ，Pranchet & Savatier ， 文，タイプ標本などについての研究結果は省略さ Schneider などの学名のタイプについては一部を検 れている. 討したに過ぎない. Leiden と Kew でも， Blume ， 分類学の研究の中でも分類群の研究はこれまで Miquel のタイプは見つからないものが多い. に行われた研究の結果を基礎として新しい成果を 属内分類群と各学名のタイプ標本を加えたこと 付け加えてゆくことが特に必要である.分類学で は異なっているが，本論文は原先生の日本種子植 は学名を基準としてその分類群についてこれまで 物集覧を補充するつもりで用意したので，和文の 何が研究されてきたのかを一覧で示すことのでき 表題をヤナギ属植物集覧とした.不十分の点が多 る発表の形式も確立されている.学名は「その群を いが，今後改訂してより完全なものにしていきた 引用することの手段を与えること J (国際植物命名 いと考えている. 規約前文)が基本的な機能の一つである.ある分類

和名と学名の対照表は以下の通りである. アカゲオオキツネヤナギ Salix futura Seemen f. rufa (Kimura) H.Ohashi イケノヤナギ Salix x ikenoana Kimura イヌコリヤナギ Salix integra Thunb. f. integra イヌコリヤナギ×エゾノカワヤナギ Salix integra Thunb. X S. miyabeana Seemen 40 40 植物研究雑誌第75 巻第l号 平成12 年2月

イヌマルパヤナギ Salix nakamurana Koidz. subsp. yezoalpina (Koidz.) H.Ohashi f. f. neoreticulata (Nakai) H.Ohashi イワヒサヤナギ Salix x iwahisana Kimura ウンリュウヤナギ、 Salix matsudana Koidz. va r. tortuosa Vilm. エゾタカネヤナギ Salix nakamurana Koidz. subsp. yezoalpina (Koidz.) H.Ohashi f. f. yezoalpina (Koidz.) H.Ohashi エゾノカワヤナギ Salix miyabeana Seemen subsp. miyabeana エゾマメヤナギ Salix nummularia Andersson f. nummularia エゾヤナギ Salix rorida Lacksch. f. rorida オオキツネヤナギ Salix futura Seemen f. futura オオキツネヤナギ×オノエヤナギ Salixfutura Seemen x S. udensis Trautv. & Mey. オオタチヤナギ Salix pierotii Miq. f. pierotii オオパヤナギ Salix cardiophylla Trautv. & Mey. va r. urbanian α(Seemen) Kudo オオミヤマヤチヤナギ Salix x psudopaludicola Kimura オクヤマサルコ Salix x matsumurae Seemen オノエヤナギ Sα lix udensis Trautv. & Mey. カミコウチヤナギ Salix x kamikotica Kimura カワイヌコリヤナギ Salix x hα~pala Kimura カワオノエヤナギ Salix x euerata Kimura カワヤナギ Salix miyabeana Seemen subsp. gilgiana (Seemen) H.Ohashi f. f. gilgiana (Seemen) H.Ohashi キツネヤナギ Sα lix vulpina Andersson キヌゲエゾマメヤナギ Salix nummularia Andersson f. hebecarpa (Kimura) H.Ohashi キヌヤナギ Salix schwerinii E. Wolf クロヤナギ Salix gracilistyla Miq. va r. melanostachys (Makino) C. K. Schneid. ケコマイワヤナギ Salix rup 俳句a Koidz. f. eriocarpa (Kimura) Kimura ex H.Ohashi ケショウヤナギ Salix arbut 約 lia Pall. ケタカネイワヤナギ Salix nakamurana Koidz. subsp. nakamurana f. f. eriocarp α(Kimura) T. Shimizu コイエヤナギ Salix x koiei Kimura コエゾヤナギ Salix rorida Lacksch. f. roridaeformis (Nakai) Kimura ex H.Ohashi コガネヤナギ Salix pierotii Miq. f. auricoma Kimura コゴメヤナギ Salix jessoensis Seemen subsp. serissaefolia (Kimura) H.Ohashi コセキヤナギ Salix x sirakawensis Kimura コマイワヤナギ Salix rup 俳句。 Koidz. f. rupifraga コリヤナギ Salix koriyanagi Kimura ex Goerz シグレヤナギ Salix x eriocataphylla Kimura シダレイヌコリヤナギ、 Salix integra Thunb. f. pendula Kimura シダレエゾヤナギ Salix rorida Lacksch. f. pendula Kimura シダレカワヤナギ Salix miyabeana Seemen subsp. gilgi αna (Seemen) H.Ohashi f. f. pendula (Kimura) H.Ohashi シダレネコヤナギ Salix gracilistyla Miq. va r. gracilistyla f. pendul α(Kimura) H.Ohashi シダレミネヤナギ Salix reinii Seemen f. pendula Kimura シダレヤナギ Salix babylonica L. f. babylonica シバキツネヤナギ Salix x japopina Kimura シパヤナギ Salix japonica Thunb. ジャヤナギ Salix eriocarpa Franch. & Sav. シライヤナギ Sα lix shiraii Seemen シロシダレヤナギ Salix x lasiogyne Seemen nothosubsp. lasiogyne February February 2000 Journal of Japanese Botany Vo l. 75 No. 1 41

シロヤナギ Salix jessoensis Seemen subsp. jessoensis スガヤヤナギ Salix x sugayana Kimura スミヨシヤナギ Salix x sumiyosensis Kimura セイコヤナギ Salix babylonica L. f. seiko Kimura センダイヤナギ Sα lix x sendaica Kimura タカネイワヤナギ Salix nakamurana Koidz. subsp. nakamurana f. nakamurana タチネコヤナギ Salix gracilis η la Miq. var. gracilistyla f. f. adscendens (Kimura) H.Ohashi タチヤナギ Sα lix triandra L. タライカヤナギ Salix taraikensis Kimura チクゼンヤナギ Salix x hatusimai Kimura リュウゾウジヤナギ Salix x hayatana Kimura チョウセンネコヤナギ Salix gracilistyla Miq. var. graciliglans (Nakai) H.Ohashi トウゲヤナギ Salix xα 19ista Schneid. トカチヤナギ Salix cardiophylla Trautv. & Mey. var. cardiophylla トヨハラヤナギ Salix x koidzumii Kirnura ナガバノネコヤナギ Salix xα rakiana Koidz. ナスノイワヤナギ Salix x nasuensis Kimura ナスノシグレヤナギ Salix x eriocataphylloides Kimura ネコシノ Tヤナギ Salix x cremnophylla Kimura ネコヤナギ Salix gracilistyla Miq. var. gracilistyla f. gracilistyla ノヤナギ Salix subopposita Miq. ハコダテヤナギ Salix x ampherista Schneid. パッコキヌヤナギ Salix x pedionoma Kimura バッコヤナギ Salix caprea L. バッコヤナギ×オオキツネヤナギ Salix caprea L. x S. futura Seemen ヒダカミネヤナギ Salix nakamurana Koidz. subsp. kurilensis (Koidz.) H.Ohashi ヒタチヤナギ Salix x turumatii Kirnura ヒラオヤナギ Salix x hiraoana Kimura フイリネコヤナギ Salix graci !i styla Miq. va r. gracilistyla f. νariegata (Kimura) Kimura フカオヤナギ Salix x sigemitui Kimura フジヤナギ Sa !i x x hisauchiana Koidz. フリソデヤナギ Salix x leucopithecia Kimura マルパヤナギ Sa !i x chaenomeloides Kirnura ミネヤナギ Salix reinii Seernen f. reinii ミヤコヤナギ Salix x thaymasta Kimura ミヤマヤチヤナギ Salix fuscescens Andersson ミョウジンヤナギ Salix x kawamurae Kimura ヤマヤナギ Salix sieboldiana Bl urne ユウキシダレ Sa !i x x lasiogyne nothosubsp. yuhkii (Kimura) H.Ohashi ユピソヤナギ Salix hukaoana Kimura ヨシノヤナギ Salix yoshinoi Koidz. ロッカクヤナギ Salix babylonica L. f. rokkaku Kimura ロッコウヤナギ Salix x graci !i styloides Kimura ワケノカワヤナギ Salix x microstemon Kimura (東北大学大学院理学研究科生物学教室)