Many Types of Evidence Support Evolution
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Lecture 20 - the History of Life on Earth
Lecture 20 - The History of Life on Earth Lecture 20 The History of Life on Earth Astronomy 141 – Autumn 2012 This lecture reviews the history of life on Earth. Rapid diversification of anaerobic prokaryotes during the Proterozoic Eon Emergence of Photosynthesis and the rise of O2 in the Earth’s atmosphere. Rise of Eukaryotes and the Cambrian Explosion in biodiversity at the start of the Phanerozoic Eon Colonization of land first by plants, then by animals Emergence of primates, then hominids, then humans. A brief digression on notation: “ya” = “years ago” Introduce a simple compact notation for writing the length of time before the present day. For example: “3.5 Billion years ago” “454 Million years ago” Gya = “giga-years ago”, hence 3.5 Gya = 3.5 Billion years ago Mya = “mega-years ago”, hence 454 Mya = 454 Million years ago [Note: some sources use Ga and Ma] Astronomy 141 - Winter 2012 1 Lecture 20 - The History of Life on Earth The four Eons of geological time. Hadean: 4.5 – 3.8 Gya: Formation, oceans & atmosphere Archaean: 3.8 – 2.5 Gya: Stromatolites & fossil bacteria Proterozoic: 2.5 Gya – 454 Mya: Eukarya and Oxygen Phanerozoic: since 454 Mya: Rise of plant and animal life The Archaean Eon began with the end of heavy bombardment ~3.8 Gya. Conditions stabilized. Oceans, but no O2 in the atmosphere. Stromatolites appear in the geological record ~3.5 Gya and thrived for >1 Billion years Rise of anaerobic microbes in the deep ocean & shores using Chemosynthesis. Time of rapid diversification of life driven by Natural Selection. -
Chapter 22 Notes: Introduction to Evolution
NOTES: Ch 22 – Descent With Modification – A Darwinian View of Life Our planet is home to a huge variety of organisms! (Scientists estimate of organisms alive today!) Even more amazing is evidence of organisms that once lived on earth, but are now . Several hundred million species have come and gone during 4.5 billion years life is believed to have existed on earth So…where have they gone… why have they disappeared? EVOLUTION: the process by which have descended from . Central Idea: organisms alive today have been produced by a long process of . FITNESS: refers to traits and behaviors of organisms that enable them to survive and reproduce COMMON DESCENT: species ADAPTATION: any inherited characteristic that enhances an organism’s ability to ~based on variations that are HOW DO WE KNOW THAT EVOLUTION HAS OCCURRED (and is still happening!!!)??? Lines of evidence: 1) So many species! -at least (250,000 beetles!) 2) ADAPTATIONS ● Structural adaptations - - ● Physiological adaptations -change in - to certain toxins 3) Biogeography: - - and -Examples: 13 species of finches on the 13 Galapagos Islands -57 species of Kangaroos…all in Australia 4) Age of Earth: -Rates of motion of tectonic plates - 5) FOSSILS: -Evidence of (shells, casts, bones, teeth, imprints) -Show a -We see progressive changes based on the order they were buried in sedimentary rock: *Few many fossils / species * 6) Applied Genetics: “Artificial Selection” - (cattle, dogs, cats) -insecticide-resistant insects - 7) Homologies: resulting from common ancestry Anatomical Homologies: ● comparative anatomy reveals HOMOLOGOUS STRUCTURES ( , different functions) -EX: ! Vestigial Organs: -“Leftovers” from the evolutionary past -Structures that Embryological Homologies: ● similarities evident in Molecular/Biochemical Homologies: ● DNA is the “universal” genetic code or code of life ● Proteins ( ) Darwin & the Scientists of his time Introduction to Darwin… ● On November 24, 1859, Charles Darwin published On the Origin of Species by Means of Natural Selection. -
Timeline of the Evolutionary History of Life
Timeline of the evolutionary history of life This timeline of the evolutionary history of life represents the current scientific theory Life timeline Ice Ages outlining the major events during the 0 — Primates Quater nary Flowers ←Earliest apes development of life on planet Earth. In P Birds h Mammals – Plants Dinosaurs biology, evolution is any change across Karo o a n ← Andean Tetrapoda successive generations in the heritable -50 0 — e Arthropods Molluscs r ←Cambrian explosion characteristics of biological populations. o ← Cryoge nian Ediacara biota – z ← Evolutionary processes give rise to diversity o Earliest animals ←Earliest plants at every level of biological organization, i Multicellular -1000 — c from kingdoms to species, and individual life ←Sexual reproduction organisms and molecules, such as DNA and – P proteins. The similarities between all present r -1500 — o day organisms indicate the presence of a t – e common ancestor from which all known r Eukaryotes o species, living and extinct, have diverged -2000 — z o through the process of evolution. More than i Huron ian – c 99 percent of all species, amounting to over ←Oxygen crisis [1] five billion species, that ever lived on -2500 — ←Atmospheric oxygen Earth are estimated to be extinct.[2][3] Estimates on the number of Earth's current – Photosynthesis Pong ola species range from 10 million to 14 -3000 — A million,[4] of which about 1.2 million have r c been documented and over 86 percent have – h [5] e not yet been described. However, a May a -3500 — n ←Earliest oxygen 2016 -
Plant Evolution an Introduction to the History of Life
Plant Evolution An Introduction to the History of Life KARL J. NIKLAS The University of Chicago Press Chicago and London CONTENTS Preface vii Introduction 1 1 Origins and Early Events 29 2 The Invasion of Land and Air 93 3 Population Genetics, Adaptation, and Evolution 153 4 Development and Evolution 217 5 Speciation and Microevolution 271 6 Macroevolution 325 7 The Evolution of Multicellularity 377 8 Biophysics and Evolution 431 9 Ecology and Evolution 483 Glossary 537 Index 547 v Introduction The unpredictable and the predetermined unfold together to make everything the way it is. It’s how nature creates itself, on every scale, the snowflake and the snowstorm. — TOM STOPPARD, Arcadia, Act 1, Scene 4 (1993) Much has been written about evolution from the perspective of the history and biology of animals, but significantly less has been writ- ten about the evolutionary biology of plants. Zoocentricism in the biological literature is understandable to some extent because we are after all animals and not plants and because our self- interest is not entirely egotistical, since no biologist can deny the fact that animals have played significant and important roles as the actors on the stage of evolution come and go. The nearly romantic fascination with di- nosaurs and what caused their extinction is understandable, even though we should be equally fascinated with the monarchs of the Carboniferous, the tree lycopods and calamites, and with what caused their extinction (fig. 0.1). Yet, it must be understood that plants are as fascinating as animals, and that they are just as important to the study of biology in general and to understanding evolutionary theory in particular. -
Biology Assessment Plan Spring 2019
Biological Sciences Department 1 Biology Assessment Plan Spring 2019 Task: Revise the Biology Program Assessment plans with the goal of developing a sustainable continuous improvement plan. In order to revise the program assessment plan, we have been asked by the university assessment committee to revise our Students Learning Outcomes (SLOs) and Program Learning Outcomes (PLOs). Proposed revisions Approach: A large community of biology educators have converged on a set of core biological concepts with five core concepts that all biology majors should master by graduation, namely 1) evolution; 2) structure and function; 3) information flow, exchange, and storage; 4) pathways and transformations of energy and matter; and (5) systems (Vision and Change, AAAS, 2011). Aligning our student learning and program goals with Vision and Change (V&C) provides many advantages. For example, the V&C community has recently published a programmatic assessment to measure student understanding of vision and change core concepts across general biology programs (Couch et al. 2019). They have also carefully outlined student learning conceptual elements (see Appendix A). Using the proposed assessment will allow us to compare our student learning profiles to those of similar institutions across the country. Revised Student Learning Objectives SLO 1. Students will demonstrate an understanding of core concepts spanning scales from molecules to ecosystems, by analyzing biological scenarios and data from scientific studies. Students will correctly identify and explain the core biological concepts involved relative to: biological evolution, structure and function, information flow, exchange, and storage, the pathways and transformations of energy and matter, and biological systems. More detailed statements of the conceptual elements students need to master are presented in appendix A. -
Transfer of the Type Species of the Genus Themobacteroides to the Genus Themoanaerobacter As Themoanaerobacter Acetoethylicus (Ben-Bassat and Zeikus 1981) Comb
INTERNATIONAL JOURNAL OF SYSTEMATICBACTERIOLOGY, Oct. 1993, p. 857-859 Vol. 43, No. 4 0020-7713/93/040857-03$02.00/0 Copyright 0 1993, International Union of Microbiological Societies Transfer of the Type Species of the Genus Themobacteroides to the Genus Themoanaerobacter as Themoanaerobacter acetoethylicus (Ben-Bassat and Zeikus 1981) comb. nov., Description of Coprothemobacter gen. nov., and Reclassification of Themobacteroides proteolyticus as Coprothennobacter proteolyticus (Ollivier et al. 1985) comb. nov. FRED A. RAINEY AND ERKO STACKEBRANDT* DSM-Deutsche Sammlung von Mikroolganisrnen und Zellkulturen, Mascheroder Weg lb, 38124 Braunschweig, Germany Phylogenetic and phenotypic evidence demonstrates the taxonomic heterogeneity of the genus Thennobac- teroides and indicates a close relationship between Thennobacteroides acetoethylicus and members of the genus Thennoanaerobacter. Since T. acetoethylicus is the type species of Thennobacteroides, its removal invalidates the genus. As a consequence, the remaining species Thennobacteroides proteolyticus is proposed as the type species of the new genus Coprothennobacter gen. nov., as Coprothennobacter proteolyticus comb. nov. Recent phylogenetic studies (8, 9) of anaerobic thermo- hybridization studies with all members of Thermoanaero- philic species demonstrated that the majority of strains fall bacter (6, 10) despite the fact that, like Thermoanaerobacter within the phylogenetic confines of the Clostridium-Bacillus species, Thermobacteroides acetoethylicus is an anaerobic, subphylum of gram-positive bacteria. In contrast to the thermophilic, glycolytic bacterium capable of growth above phylog enet ically coherent genera Thermoana erobac ter (6) 70°C that has been isolated from geothermal environments. and Thermoanaerobacterium (6), members of Thermobac- The reclassifications of the recent study of Lee et al. (6) teroides (1,7) belonged to phylogenetically very diverse taxa increased the numbers of species of Thermoanaerobacter to (8). -
Introduction to Botany
Introduction to Botany Jan Zientek Senior Program Coordinator Cooperative Extension of Essex County [email protected] Basic Botany • The study of the growth, structure and function of plants Plant Functions BOTANY • Evolution • Taxonomy • Plant morphology • Plant physiology and cell biology • Plant reproduction • Plant hormones and growth regulators PLANTAE • Eukaryotic (with a • Chloroplasts nucleus) (green) • Cell walls with • Non-motile cellulose • Several phlyum • Food stored as • Development of carbohydrate pollen • Multi-cellular autotrophs DOMAIN KINGDOM PHYLUM CLASS ORDER FAMILY Genus species Plant phylums Mosses Cycads Liverworts Ginkgoes Hornworts Gnetophytes Club mosses Conifers Horsetails Ferns Common Name vs Scientific Name Foxglove Digitalis purpurea • Maybe local name • Universally recognized • General • Specific Common Name vs Scientific Name Fire bush Scarlet bush Texas firecracker Corail (or is it Koray?) Hamelia patens Polly red head Hummingbird bush Ix-canan Plant Types • Monocots: have a single cotyledon, flower parts in multiples of three, parallel venation of leaves, scattered vascular bundles in stems. Dicots: have two cotyledons, flower parts in multiples of 4 or 5, netted veins, and stems which are organized in a ring pattern. Humans sort things many different ways • Plants can be classified by the type of their seed structure – Gymnosperm: “naked seed” – Angiosperm: seed within a fruiting body • Lifecycles help gardeners distinguish between plants: • Annuals • Biennials • Perennials Annuals complete life cycles in one season Biennials live for 2 years, flower, then die Perennials live for 3 or more years, flower each year and usually do not die after flowering. Structures of Plants Roots Stems Leaves Flowers Seeds Examples of Stem Structure Herbaceous monocot and dicot stem Root cross-section • Apical meristem: point of vigorous cell division and growth. -
Re-Description of the Sauropod Dinosaur Amanzia (“Ornithopsis
Schwarz et al. Swiss J Geosci (2020) 113:2 https://doi.org/10.1186/s00015-020-00355-5 Swiss Journal of Geosciences ORIGINAL PAPER Open Access Re-description of the sauropod dinosaur Amanzia (“Ornithopsis/Cetiosauriscus”) greppini n. gen. and other vertebrate remains from the Kimmeridgian (Late Jurassic) Reuchenette Formation of Moutier, Switzerland Daniela Schwarz1* , Philip D. Mannion2 , Oliver Wings3 and Christian A. Meyer4 Abstract Dinosaur remains were discovered in the 1860’s in the Kimmeridgian (Late Jurassic) Reuchenette Formation of Moutier, northwestern Switzerland. In the 1920’s, these were identifed as a new species of sauropod, Ornithopsis greppini, before being reclassifed as a species of Cetiosauriscus (C. greppini), otherwise known from the type species (C. stewarti) from the late Middle Jurassic (Callovian) of the UK. The syntype of “C. greppini” consists of skeletal elements from all body regions, and at least four individuals of diferent sizes can be distinguished. Here we fully re-describe this material, and re-evaluate its taxonomy and systematic placement. The Moutier locality also yielded a theropod tooth, and fragmen- tary cranial and vertebral remains of a crocodylomorph, also re-described here. “C.” greppini is a small-sized (not more than 10 m long) non-neosauropod eusauropod. Cetiosauriscus stewarti and “C.” greppini difer from each other in: (1) size; (2) the neural spine morphology and diapophyseal laminae of the anterior caudal vertebrae; (3) the length-to-height proportion in the middle caudal vertebrae; (4) the presence or absence of ridges and crests on the middle caudal cen- tra; and (5) the shape and proportions of the coracoid, humerus, and femur. -
Darwin's “Tree of Life”
Icons of Evolution? Why Much of What Jonathan Wells Writes about Evolution is Wrong Alan D. Gishlick, National Center for Science Education DARWIN’S “TREE OF LIFE” mon descent. Finally, he demands that text- books treat universal common ancestry as PHYLOGENETIC TREES unproven and refrain from illustrating that n biology, a phylogenetic tree, or phyloge- “theory” with misleading phylogenies. ny, is used to show the genealogic relation- Therefore, according to Wells, textbooks Iships of living things. A phylogeny is not should state that there is no evidence for com- so much evidence for evolution as much as it mon descent and that the most recent research is a codification of data about evolutionary his- refutes the concept entirely. Wells is complete- tory. According to biological evolution, organ- ly wrong on all counts, and his argument is isms share common ancestors; a phylogeny entirely based on misdirection and confusion. shows how organisms are related. The tree of He mixes up these various topics in order to life shows the path evolution took to get to the confuse the reader into thinking that when current diversity of life. It also shows that we combined, they show an endemic failure of can ascertain the genealogy of disparate living evolutionary theory. In effect, Wells plays the organisms. This is evidence for evolution only equivalent of an intellectual shell game, put- in that we can construct such trees at all. If ting so many topics into play that the “ball” of evolution had not happened or common ances- evolution gets lost. try were false, we would not be able to discov- THE CAMBRIAN EXPLOSION er hierarchical branching genealogies for ells claims that the Cambrian organisms (although textbooks do not general- Explosion “presents a serious chal- ly explain this well). -
Phylocode: a Phylogenetic Code of Biological Nomenclature
PhyloCode: A Phylogenetic Code of Biological Nomenclature Philip D. Cantino and Kevin de Queiroz (equal contributors; names listed alphabetically) Advisory Group: William S. Alverson, David A. Baum, Harold N. Bryant, David C. Cannatella, Peter R. Crane, Michael J. Donoghue, Torsten Eriksson*, Jacques Gauthier, Kenneth Halanych, David S. Hibbett, David M. Hillis, Kathleen A. Kron, Michael S. Y. Lee, Alessandro Minelli, Richard G. Olmstead, Fredrik Pleijel*, J. Mark Porter, Heidi E. Robeck, Greg W. Rouse, Timothy Rowe*, Christoffer Schander, Per Sundberg, Mikael Thollesson, and Andre R. Wyss. *Chaired a committee that authored a portion of the current draft. Most recent revision: April 8, 2000 1 Table of Contents Preface Preamble Division I. Principles Division II. Rules Chapter I. Taxa Article 1. The Nature of Taxa Article 2. Clades Article 3. Hierarchy and Rank Chapter II. Publication Article 4. Publication Requirements Article 5. Publication Date Chapter III. Names Section 1. Status Article 6 Section 2. Establishment Article 7. General Requirements Article 8. Registration Chapter IV. Clade Names Article 9. General Requirements for Establishment of Clade Names Article 10. Selection of Clade Names for Establishment Article 11. Specifiers and Qualifying Clauses Chapter V. Selection of Accepted Names Article 12. Precedence Article 13. Homonymy Article 14. Synonymy Article 15. Conservation Chapter VI. Provisions for Hybrids Article 16. Chapter VII. Orthography Article 17. Orthographic Requirements for Establishment Article 18. Subsequent Use and Correction of Established Names Chapter VIII. Authorship of Names Article 19. Chapter IX. Citation of Authors and Registration Numbers Article 20. Chapter X. Governance Article 21. Glossary Table 1. Equivalence of Nomenclatural Terms Appendix A. -
Species Change Over Time
KEY CONCEPT Species change over time. BEFORE, you learned NOW, you will learn • Fossils are evidence of earlier • About early ideas and observa- life tions on evolution • More complex organisms have • How Darwin developed his developed over time theory of natural selection • Mass extinctions contributed to • How new species arise from the development of Earth’s older species history VOCABULARY THINK ABOUT evolution p. 797 How have telephones changed over time? natural selection p. 801 adaptation p. 802 Today people across the world can speciation p. 804 communicate in many different ways. One of the most common ways is over the telephone. Looking at the two pictures, can you describe how this form of communication has changed over time? Scientists explore the concept of evolution. MAIN IDEA AND DETAILS In a general sense, evolution involves a change over time. You could Make a chart for the main say that the way humans communicate has evolved. Certainly idea scientists explore the concept of evolution. telephones have changed over time. The first telephones were the size Include details about scien- of a shoebox. Today a telephone can fit in the palm of your hand and tists’ observations. can send images as well as sound. In biology,evolution refers to the process though which species change over time. The change results from a change in the genetic material of an organism and is passed from one generation to the next. Check Your Reading What is evolution? Chapter 23: History of Life 797 Early Ideas reading tip In the early 1800s, a French scientist named Jean Baptiste de Lamarck The word acquire comes was the first scientist to propose a model of how life evolves. -
Recent Developments in International Biological Nomenclature
TurkJBot 28(2004)17-26 ©TÜB‹TAK RecentDevelopmentsinInternationalBiologicalNomenclature WernerGREUTER BotanischerGarten&BotanischesMuseumBerlin-Dahlem,Königin-Luise-Str.6-8,D-14191Berlin,Germany. Received:08.11.2002 Accepted:29.01.2003 Abstract: Afteraperiodofseveraldecadesofgeneralneglect,biologicalsystematicsandtaxonomyareagainsurfacingtothe awarenessofthescientificcommunityandofpolicymakers.Thisweowetothefactthattheyarethecoredisciplinesofbiodiversity research,nowperceivedtobeaprioritytaskinviewoftheimpendingthreatoflossthroughextinctionofagreatnumberofspecies, andthewholesaledestructionofecosystems.Yettherenaissanceofsystematicsandtaxonomyishamperedbytheinadequacyof thetraditionalrulesgoverningtheformationanduseofscientificnamesoforganisms.Namesareacoreaspectofhuman communication.Theyarethemeansbywhichscientistsandlaymenalikerefertoorganisms,theindispensablelabelsthatenablethe storage,retrievalandcommunicationofanyandallorganism-linkeddata.Thispaperhighlightsthefundamentalinadequacyof traditionalnomenclatural Codes tofulfiltheirrole.Pastattemptstointroducenewconceptsintobotanicalnomenclature,suchas theregistrationofnewnamesandthestabilisationoftheexistingoneswhicharepresentlyused,arementioned–aswellast heir failuretogainacceptancebythesmallandhighlyself-centredcommunityofspecialistsinthefield.Recentattemptstolaunc h competingsetsofrules,the“ Phylocode”inparticular,arealsodiscussed.Whiletheymaybeasuitablemeansofbreakingthedire monopolyoftheCodes,theyalsohaveadangerousdestabilisingpotential.Newsolutions,whicharedoubtlessneeded,requiregreat