Phylocode: a Phylogenetic Code of Biological Nomenclature
Total Page:16
File Type:pdf, Size:1020Kb
Load more
Recommended publications
-
International Code of Zoological Nomenclature
International Commission on Zoological Nomenclature INTERNATIONAL CODE OF ZOOLOGICAL NOMENCLATURE Fourth Edition adopted by the International Union of Biological Sciences The provisions of this Code supersede those of the previous editions with effect from 1 January 2000 ISBN 0 85301 006 4 The author of this Code is the International Commission on Zoological Nomenclature Editorial Committee W.D.L. Ride, Chairman H.G. Cogger C. Dupuis O. Kraus A. Minelli F. C. Thompson P.K. Tubbs All rights reserved. No part of this publication may be reproduced, stored in a retrieval system, or transmitted in any form or by any means (electronic, mechanical, photocopying or otherwise), without the prior written consent of the publisher and copyright holder. Published by The International Trust for Zoological Nomenclature 1999 c/o The Natural History Museum - Cromwell Road - London SW7 5BD - UK © International Trust for Zoological Nomenclature 1999 Explanatory Note This Code has been adopted by the International Commission on Zoological Nomenclature and has been ratified by the Executive Committee of the International Union of Biological Sciences (IUBS) acting on behalf of the Union's General Assembly. The Commission may authorize official texts in any language, and all such texts are equivalent in force and meaning (Article 87). The Code proper comprises the Preamble, 90 Articles (grouped in 18 Chapters) and the Glossary. Each Article consists of one or more mandatory provisions, which are sometimes accompanied by Recommendations and/or illustrative Examples. In interpreting the Code the meaning of a word or expression is to be taken as that given in the Glossary (see Article 89). -
Dinosaur Wars Program Transcript
Page 1 Dinosaur Wars Program Transcript Narrator: For more than a century, Americans have had a love affair with dinosaurs. Extinct for millions of years, they were barely known until giant, fossil bones were discovered in the mid-nineteenth century. Two American scientists, Edward Drinker Cope and Othniel Charles Marsh, led the way to many of these discoveries, at the forefront of the young field of paleontology. Jacques Gauthier, Paleontologist: Every iconic dinosaur every kid grows up with, apatosaurus, triceratops, stegosaurus, allosaurus, these guys went out into the American West and they found that stuff. Narrator: Cope and Marsh shed light on the deep past in a way no one had ever been able to do before. They unearthed more than 130 dinosaur species and some of the first fossil evidence supporting Darwin’s new theory of evolution. Mark Jaffe, Writer: Unfortunately there was a more sordid element, too, which was their insatiable hatred for each other, which often just baffled and exasperated everyone around them. Peter Dodson, Paleontologist: They began life as friends. Then things unraveled… and unraveled in quite a spectacular way. Narrator: Cope and Marsh locked horns for decades, in one of the most bitter scientific rivalries in American history. Constantly vying for leadership in their young field, they competed ruthlessly to secure gigantic bones in the American West. They put American science on the world stage and nearly destroyed one another in the process. Page 2 In the summer of 1868, a small group of scientists boarded a Union Pacific train for a sightseeing excursion through the heart of the newly-opened American West. -
H17 the Descent of Birds < Neornithines, Archaeopteryx
LIFE OF THE MESOZOIC ERA 465 h17 The descent of birds < neornithines, Archaeopteryx, Solnhofen Lms > Alfred, Lord Tennyson / once solved an enigma: When is an / eider [a seaduck] most like a merganser [a lake and river duck]? / He lived long enough to forget the answer. —an anonymous clerihew.1 All eighteen species of penguin live in the Southern Hemisphere. ... The descendants of a common ancestor, sharing common ground. ... ‘This bond, on my [Darwin’s] theory, is simple inheritance.’2 Bird evolution during the Cenozoic has been varied and rapid in the beak but not in the body. For example, Hawaiian Island bird arrivals beginning 5 Ma (million years ago) have evolved to new species with beaks specialized to feed on various flowers and seeds but their bodies have changed so little that the various pioneer-bird stocks are easy to identify. With that perspective, bird varieties distinctive in the beak as owls, hawks, penguins, ducks, flamingos, and parrots, are likely to have diverged in very ancient times. Ducks, albeit primitive ones such as web footed Presbyornis, which could wade but not paddle, already existed in the bird diversity of the Eocene when giant “terror birds,” such as Diatrema, patrolled the ground. Feathers are made mostly of beta keratin, a strong light-weight material. Bird feathers are modified scales which extrude from cells that line skin follicles. The shape and color of the feather so formed can change during the birds growth (say, a downy feather in youth and a stiff feather in adulthood) and by seasonal molts (say, a white feather in the winter and a colored feather for times of courting).3 All birds of the Cenozoic (30 orders) are characterized by toothless jaws and have a pair of elongated foot bones that are partially fused from the bottom up. -
A Phylogenomic Analysis of Turtles ⇑ Nicholas G
Molecular Phylogenetics and Evolution 83 (2015) 250–257 Contents lists available at ScienceDirect Molecular Phylogenetics and Evolution journal homepage: www.elsevier.com/locate/ympev A phylogenomic analysis of turtles ⇑ Nicholas G. Crawford a,b,1, James F. Parham c, ,1, Anna B. Sellas a, Brant C. Faircloth d, Travis C. Glenn e, Theodore J. Papenfuss f, James B. Henderson a, Madison H. Hansen a,g, W. Brian Simison a a Center for Comparative Genomics, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA b Department of Genetics, University of Pennsylvania, Philadelphia, PA 19104, USA c John D. Cooper Archaeological and Paleontological Center, Department of Geological Sciences, California State University, Fullerton, CA 92834, USA d Department of Biological Sciences, Louisiana State University, Baton Rouge, LA 70803, USA e Department of Environmental Health Science, University of Georgia, Athens, GA 30602, USA f Museum of Vertebrate Zoology, University of California, Berkeley, CA 94720, USA g Mathematical and Computational Biology Department, Harvey Mudd College, 301 Platt Boulevard, Claremont, CA 9171, USA article info abstract Article history: Molecular analyses of turtle relationships have overturned prevailing morphological hypotheses and Received 11 July 2014 prompted the development of a new taxonomy. Here we provide the first genome-scale analysis of turtle Revised 16 October 2014 phylogeny. We sequenced 2381 ultraconserved element (UCE) loci representing a total of 1,718,154 bp of Accepted 28 October 2014 aligned sequence. Our sampling includes 32 turtle taxa representing all 14 recognized turtle families and Available online 4 November 2014 an additional six outgroups. Maximum likelihood, Bayesian, and species tree methods produce a single resolved phylogeny. -
Phylogenetic Comparative Methods: a User's Guide for Paleontologists
Phylogenetic Comparative Methods: A User’s Guide for Paleontologists Laura C. Soul - Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, DC, USA David F. Wright - Division of Paleontology, American Museum of Natural History, Central Park West at 79th Street, New York, New York 10024, USA and Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, DC, USA Abstract. Recent advances in statistical approaches called Phylogenetic Comparative Methods (PCMs) have provided paleontologists with a powerful set of analytical tools for investigating evolutionary tempo and mode in fossil lineages. However, attempts to integrate PCMs with fossil data often present workers with practical challenges or unfamiliar literature. In this paper, we present guides to the theory behind, and application of, PCMs with fossil taxa. Based on an empirical dataset of Paleozoic crinoids, we present example analyses to illustrate common applications of PCMs to fossil data, including investigating patterns of correlated trait evolution, and macroevolutionary models of morphological change. We emphasize the importance of accounting for sources of uncertainty, and discuss how to evaluate model fit and adequacy. Finally, we discuss several promising methods for modelling heterogenous evolutionary dynamics with fossil phylogenies. Integrating phylogeny-based approaches with the fossil record provides a rigorous, quantitative perspective to understanding key patterns in the history of life. 1. Introduction A fundamental prediction of biological evolution is that a species will most commonly share many characteristics with lineages from which it has recently diverged, and fewer characteristics with lineages from which it diverged further in the past. This principle, which results from descent with modification, is one of the most basic in biology (Darwin 1859). -
Phylogenetic Definitions in the Pre-Phylocode Era; Implications for Naming Clades Under the Phylocode
PaleoBios 27(1):1–6, April 30, 2007 © 2006 University of California Museum of Paleontology Phylogenetic definitions in the pre-PhyloCode era; implications for naming clades under the PhyloCode MiChAel P. TAylor Palaeobiology research Group, School of earth and environmental Sciences, University of Portsmouth, Portsmouth Po1 3Ql, UK; [email protected] The last twenty years of work on phylogenetic nomenclature have given rise to many names and definitions that are now considered suboptimal. in formulating permanent definitions under the PhyloCode when it is implemented, it will be necessary to evaluate the corpus of existing names and make judgements about which to establish and which to discard. This is not straightforward, because early definitions are often inexplicit and ambiguous, generally do not meet the requirements of the PhyloCode, and in some cases may not be easily recognizable as phylogenetic definitions at all. recognition of synonyms is also complicated by the use of different kinds of specifiers (species, specimens, clades, genera, suprageneric rank-based names, and vernacular names) and by definitions whose content changes under different phylogenetic hypotheses. in light of these difficulties, five principles are suggested to guide the interpreta- tion of pre-PhyloCode clade-names and to inform the process of naming clades under the PhyloCode: (1) do not recognize “accidental” definitions; (2) malformed definitions should be interpreted according to the intention of the author when and where this is obvious; (3) apomorphy-based and other definitions must be recognized as well as node-based and stem-based definitions; (4) definitions using any kind of specifier taxon should be recognized; and (5) priority of synonyms and homonyms should guide but not prescribe. -
Biology Assessment Plan Spring 2019
Biological Sciences Department 1 Biology Assessment Plan Spring 2019 Task: Revise the Biology Program Assessment plans with the goal of developing a sustainable continuous improvement plan. In order to revise the program assessment plan, we have been asked by the university assessment committee to revise our Students Learning Outcomes (SLOs) and Program Learning Outcomes (PLOs). Proposed revisions Approach: A large community of biology educators have converged on a set of core biological concepts with five core concepts that all biology majors should master by graduation, namely 1) evolution; 2) structure and function; 3) information flow, exchange, and storage; 4) pathways and transformations of energy and matter; and (5) systems (Vision and Change, AAAS, 2011). Aligning our student learning and program goals with Vision and Change (V&C) provides many advantages. For example, the V&C community has recently published a programmatic assessment to measure student understanding of vision and change core concepts across general biology programs (Couch et al. 2019). They have also carefully outlined student learning conceptual elements (see Appendix A). Using the proposed assessment will allow us to compare our student learning profiles to those of similar institutions across the country. Revised Student Learning Objectives SLO 1. Students will demonstrate an understanding of core concepts spanning scales from molecules to ecosystems, by analyzing biological scenarios and data from scientific studies. Students will correctly identify and explain the core biological concepts involved relative to: biological evolution, structure and function, information flow, exchange, and storage, the pathways and transformations of energy and matter, and biological systems. More detailed statements of the conceptual elements students need to master are presented in appendix A. -
Diversity-Dependent Cladogenesis Throughout Western Mexico: Evolutionary Biogeography of Rattlesnakes (Viperidae: Crotalinae: Crotalus and Sistrurus)
City University of New York (CUNY) CUNY Academic Works Publications and Research New York City College of Technology 2016 Diversity-dependent cladogenesis throughout western Mexico: Evolutionary biogeography of rattlesnakes (Viperidae: Crotalinae: Crotalus and Sistrurus) Christopher Blair CUNY New York City College of Technology Santiago Sánchez-Ramírez University of Toronto How does access to this work benefit ou?y Let us know! More information about this work at: https://academicworks.cuny.edu/ny_pubs/344 Discover additional works at: https://academicworks.cuny.edu This work is made publicly available by the City University of New York (CUNY). Contact: [email protected] 1Blair, C., Sánchez-Ramírez, S., 2016. Diversity-dependent cladogenesis throughout 2 western Mexico: Evolutionary biogeography of rattlesnakes (Viperidae: Crotalinae: 3 Crotalus and Sistrurus ). Molecular Phylogenetics and Evolution 97, 145–154. 4 https://doi.org/10.1016/j.ympev.2015.12.020. © 2016. This manuscript version is made 5 available under the CC-BY-NC-ND 4.0 license. 6 7 8 Diversity-dependent cladogenesis throughout western Mexico: evolutionary 9 biogeography of rattlesnakes (Viperidae: Crotalinae: Crotalus and Sistrurus) 10 11 12 CHRISTOPHER BLAIR1*, SANTIAGO SÁNCHEZ-RAMÍREZ2,3,4 13 14 15 1Department of Biological Sciences, New York City College of Technology, Biology PhD 16 Program, Graduate Center, The City University of New York, 300 Jay Street, Brooklyn, 17 NY 11201, USA. 18 2Department of Ecology and Evolutionary Biology, University of Toronto, 25 Willcocks 19 Street, Toronto, ON, M5S 3B2, Canada. 20 3Department of Natural History, Royal Ontario Museum, 100 Queen’s Park, Toronto, 21 ON, M5S 2C6, Canada. 22 4Present address: Environmental Genomics Group, Max Planck Institute for 23 Evolutionary Biology, August-Thienemann-Str. -
A Phylogenetic Analysis of the Basal Ornithischia (Reptilia, Dinosauria)
A PHYLOGENETIC ANALYSIS OF THE BASAL ORNITHISCHIA (REPTILIA, DINOSAURIA) Marc Richard Spencer A Thesis Submitted to the Graduate College of Bowling Green State University in partial fulfillment of the requirements of the degree of MASTER OF SCIENCE December 2007 Committee: Margaret M. Yacobucci, Advisor Don C. Steinker Daniel M. Pavuk © 2007 Marc Richard Spencer All Rights Reserved iii ABSTRACT Margaret M. Yacobucci, Advisor The placement of Lesothosaurus diagnosticus and the Heterodontosauridae within the Ornithischia has been problematic. Historically, Lesothosaurus has been regarded as a basal ornithischian dinosaur, the sister taxon to the Genasauria. Recent phylogenetic analyses, however, have placed Lesothosaurus as a more derived ornithischian within the Genasauria. The Fabrosauridae, of which Lesothosaurus was considered a member, has never been phylogenetically corroborated and has been considered a paraphyletic assemblage. Prior to recent phylogenetic analyses, the problematic Heterodontosauridae was placed within the Ornithopoda as the sister taxon to the Euornithopoda. The heterodontosaurids have also been considered as the basal member of the Cerapoda (Ornithopoda + Marginocephalia), the sister taxon to the Marginocephalia, and as the sister taxon to the Genasauria. To reevaluate the placement of these taxa, along with other basal ornithischians and more derived subclades, a phylogenetic analysis of 19 taxonomic units, including two outgroup taxa, was performed. Analysis of 97 characters and their associated character states culled, modified, and/or rescored from published literature based on published descriptions, produced four most parsimonious trees. Consistency and retention indices were calculated and a bootstrap analysis was performed to determine the relative support for the resultant phylogeny. The Ornithischia was recovered with Pisanosaurus as its basalmost member. -
Dieter Thomas Tietze Editor How They Arise, Modify and Vanish
Fascinating Life Sciences Dieter Thomas Tietze Editor Bird Species How They Arise, Modify and Vanish Fascinating Life Sciences This interdisciplinary series brings together the most essential and captivating topics in the life sciences. They range from the plant sciences to zoology, from the microbiome to macrobiome, and from basic biology to biotechnology. The series not only highlights fascinating research; it also discusses major challenges associated with the life sciences and related disciplines and outlines future research directions. Individual volumes provide in-depth information, are richly illustrated with photographs, illustrations, and maps, and feature suggestions for further reading or glossaries where appropriate. Interested researchers in all areas of the life sciences, as well as biology enthusiasts, will find the series’ interdisciplinary focus and highly readable volumes especially appealing. More information about this series at http://www.springer.com/series/15408 Dieter Thomas Tietze Editor Bird Species How They Arise, Modify and Vanish Editor Dieter Thomas Tietze Natural History Museum Basel Basel, Switzerland ISSN 2509-6745 ISSN 2509-6753 (electronic) Fascinating Life Sciences ISBN 978-3-319-91688-0 ISBN 978-3-319-91689-7 (eBook) https://doi.org/10.1007/978-3-319-91689-7 Library of Congress Control Number: 2018948152 © The Editor(s) (if applicable) and The Author(s) 2018. This book is an open access publication. Open Access This book is licensed under the terms of the Creative Commons Attribution 4.0 International License (http://creativecommons.org/licenses/by/4.0/), which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license and indicate if changes were made. -
Species Names in Phylogenetic Nomenclature
Syst. Biol. 48(4):790–807, 1999 Species Names in Phylogenetic Nomenclature PHILIP D. CANTINO,1* HAROLD N. BRYANT,2 KEVIN DE QUEIROZ,3 MICHAEL J. DONOGHUE,4 TORSTEN ERIKSSON,5 DAVID M. HILLIS,6 AND MICHAEL S. Y. LEE7 1Department of Environmental and Plant Biology, Ohio University, Athens, Ohio 45701, USA; E-mail: [email protected] 2Royal Saskatchewan Museum, 2340 Albert Street, Regina, Saskatchewan S4P 3V7, Canada; E-mail: [email protected] 3Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20560, USA; E-mail: [email protected] 4Harvard University Herbaria, 22 Divinity Ave., Cambridge, Massachusetts 02138, USA; E-mail: [email protected] 5Bergius Foundation, Royal Swedish Academy of Sciences, Box 50017, 104 05 Stockholm, Sweden; E-mail: [email protected] 6Section of Integrative Biology, School of Biological Sciences, University of Texas, Austin, Texas 78712, USA; E-mail: [email protected] 7Department of Zoology, University of Queensland, Brisbane, Queensland 4072, Australia; E-mail: [email protected] Abstract.—Linnaean binomial nomenclature is logically incompatible with the phylogenetic nomenclature of de Queiroz and Gauthier (1992, Annu. Rev. Ecol. Syst. 23:449–480): The former is based on the concept of genus, thus making this rank mandatory, while the latter is based on phylo- genetic definitions and requires the abandonment of mandatory ranks. Thus, if species are to re- ceive names under phylogenetic nomenclature, a different method must be devised to name them. Here, 13 methods for naming species in the context of phylogenetic nomenclature are contrasted with each other and with Linnaean binomials. -
Towards Resolving Lamiales Relationships
Schäferhoff et al. BMC Evolutionary Biology 2010, 10:352 http://www.biomedcentral.com/1471-2148/10/352 RESEARCH ARTICLE Open Access Towards resolving Lamiales relationships: insights from rapidly evolving chloroplast sequences Bastian Schäferhoff1*, Andreas Fleischmann2, Eberhard Fischer3, Dirk C Albach4, Thomas Borsch5, Günther Heubl2, Kai F Müller1 Abstract Background: In the large angiosperm order Lamiales, a diverse array of highly specialized life strategies such as carnivory, parasitism, epiphytism, and desiccation tolerance occur, and some lineages possess drastically accelerated DNA substitutional rates or miniaturized genomes. However, understanding the evolution of these phenomena in the order, and clarifying borders of and relationships among lamialean families, has been hindered by largely unresolved trees in the past. Results: Our analysis of the rapidly evolving trnK/matK, trnL-F and rps16 chloroplast regions enabled us to infer more precise phylogenetic hypotheses for the Lamiales. Relationships among the nine first-branching families in the Lamiales tree are now resolved with very strong support. Subsequent to Plocospermataceae, a clade consisting of Carlemanniaceae plus Oleaceae branches, followed by Tetrachondraceae and a newly inferred clade composed of Gesneriaceae plus Calceolariaceae, which is also supported by morphological characters. Plantaginaceae (incl. Gratioleae) and Scrophulariaceae are well separated in the backbone grade; Lamiaceae and Verbenaceae appear in distant clades, while the recently described Linderniaceae are confirmed to be monophyletic and in an isolated position. Conclusions: Confidence about deep nodes of the Lamiales tree is an important step towards understanding the evolutionary diversification of a major clade of flowering plants. The degree of resolution obtained here now provides a first opportunity to discuss the evolution of morphological and biochemical traits in Lamiales.