This is anopenaccess article undertheCCBY-NC-SAThis Universidad NacionalAutónoma de México. Peer Reviewing under the responsability of Manuscript accepted: Corrected manuscript received: Manuscript received: BSGM2021v73n3a030121 73 (3), A030121.http://dx.doi.org/10.18268/ Boletín de la Sociedad Geológica Mexicana, Germany: southern the Upper Jurassic of crabs (Decapoda: Brachyura: Dromiacea) from prosopid Schweigert, G., 2021, A newgenus of How tocitethisarticle: [email protected] * Corresponding author:(G. Schweirgert) Germany stein 1,70191Stuttgart, 1 Günter Superior delsurdeAlemania Un nuevogénerodecangrejosprosópidos(Decapoda:Brachyura:Dromiacea)delJurásico Jurassic ofsouthernGermany A new genus of prosopid crabs(Decapoda: Brachyura: Dromiacea) from the Upper license(https://creativecommons.org/licenses/by-nc-sa/4.0/) Staatliches Museumfür Naturkunde, Rosen Schweiger 1,* Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín - Formation, Bavaria, Germany. palaeoecology,Treuchtlingen diversification,Late Jurassic, Keywords:Homolodromioidea, Formation inspecial. the Treuchtlingen ran faunaof general Ocean in and tothe brachyu the Tethys shelve of the northern sponge-microbial magnafaciesalong early true crabs withinthe hotspot of Petersbuchia Zone). Kimmeridgian, Acanthicum Treuchtlingen Formation (early late the Upper Jurassic lithic limestone of lected from a thick-bedded spongio pletely preserved carapace, wascol Germany. holotype, The anincom sp.,gen. n. described from is southern brachyurans, of genusA newmonotypic and species ABSTRACT adds to the diversification adds Petersbuchia thauckei n. - - - - /73(3)A0301212021 colectado de una gruesa capa espongiolítica capa colectado de unagruesa holotipo,caparazón un incompleto, fue n. sp., es descrito del sur de Alemania. El brachyuros, género yespecie monotípico de nuevo Un RESUMEN Alemania. ción Treuchtlingen, Bavaria, Tardío, palaeoecología,Forma- dea, diversificación, Jurásico Palabras clave: Homolodromioi en particular. de braquiuros de la Formación Treuchtlingen, Océano delTethys general,en y delafauna nortedel dela Plataforma bianas a lolargo verdaderos espongio-micro en magnafacies el punto de diversificación de los cangrejos Zona Acanthicum). Superior (Kimmeridgiano temprano tardío, de la Formación Treuchtlingen delJurásio Petersbuchia thauckei Petersbuchia seañade n.gen. 1 -

New crab of the Jurassic in Germany ABSTRACT New crab of the Jurassic in Germany INTRODUCTION/ GEOLOGICAL SETTING 1. Introduction 2 2 enrcre rmcae aiesrt fEarly been recorded from clayey marinestrata of ones, sometimescontroversiallyhave discussed, species (Davie arerans well represented byapproximately 7,250 marinebenthicenvironments,In modern brachyu n pce fthe Prosopidae. abbrevia The and species of al Robins, 2016; Schweigert, 2019; Klompmaker al and Koppka, 2011;Starzyk Schweigert 2011; Franţescu, 2010a-c; 2009a-d; 2007, 2018;Schweitzer and Feldmann, 2008a, b, ally added (Feldmann differently or discovered Jurassicinterpreted crab material were occasion newly on based taxa old collectionsbecame necessary, andadditional the original type material from revisionsnomic of pod crustaceans (Glaessner, 1969), numerous taxo the ʻTreatiseʼ volume updatededicated of to deca origination (seeKrobicki andZatoń, 2008). Tethys the southern (Förster, dubious 1985)are of Karasawa andHirota, 2019). Early recordsfrom area (Karasawa and Kato, 2007; Kato Pacific western the from come even records ran group, isolated brachyu although geographically this of hotspot diversification a also probably was brachyuran studies(von Meyer, 1842,1860)but areaa classical not only belt, is for early reef Tethyanmany, thisnorthern which waspart of et al (Wehner, 1988;Müller crabs of expansion biogeographical and sification 1996). This probably triggered the firstgreat diver the TethysOcean (e.g., of Leinfelder margin became widespread along the northern crobial magnafacies aswell as coeval coral reefs al Fraaije 2008; Zatoń, and (Krobicki scarce still Brachyuran recordsfrom the Middle Jurassicare Krzemińska 2014; weitzer and Feldmann, 2010d;HaugandHaug, Jurassic(Pliensbachian) age (Förster, 1986;Sch / / ., 2013). In the Late Jurassic, siliceous sponge-mi ., 2020). 2015a, b,., 2015;Starzyk, 2020;Schweigert and Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín h i fthisstudyto reportis a newgenus aim of The anextensive revisionand of the course In ., 2013). The Upper Jurassic of southern Ger southern Upper Jurassic., 2013). The of et al ., 2015). The mostprimitive., 2015).The et al et al et al ., 2020;Scholz, 2020). ., 2006;Schweitzer ., 2000;Klompmaker et al http://dx.doi.org/10.18268/BSGM2021v73n3a030121 ., 2012; Hyžný 2012; ., et al et al ., 1994, ., 2010; et al et et et ., /73(3)A0301212021 ------/73(3)A0301212021 2. Geologicalsetting ietnso the Treuchtlingen Formationlimestones of yield and traded as “Treuchtlingen Marble”. The a building stonetheselimestonesare well-known Bavaria (Figure 1). Due to its usage as (Germany) Markt Titting in the township of which ispart of the Petersbuch-Erkertshofen in quarried district, lingen Formation (Niebuhr andPürner, 2014) the Treucht from thick-bedded limestones of hereinThe described specimen wascollected Naturkunde Germany. Stuttgart, tion SMNS used foris Staatliches Museumfür sals fbrachyurans from the Treuchtlingenis a list of and theyare ignored mostly and overlooked. Here Formation these tiny fossilsare not in their focus hunting for fossilsin the Franconian Treuchtlingen reefs (cf. Schweitzer species otherwise restricted to Upper Jurassiccoral relatively low-diverseany and doesnotcomprise small-sized crabs(see below), is of fauna consists et al felder wavebelow the storm-weather basis (e.g., Lein ca.50to 150 metres, certainly within arange of greater water depths put forwardfavour in of thick-shelled bivalves, large gastropods), have been corals,a typicalshallow-water fauna (hermatypic many arguments, especially the total absence of bysuggested Kott (1989); however, subsequently verythe past. A in shallow modelwas depositional havewas deposited been controversiallydiscussed ditions duringwhich the Treuchtlingen Formation Schlampp, environmental 2004,2009).The con the late Kimmeridgian (Zeiss, 1964;Bantz, 1970; these rockswithin fauna allowsa precise dating of characteristic for thislithology. ammonite The wall (“ ish macroscopicallyvisible foraminifera with awhit fromlithology.absent this finer matrixthe rock In coralsarenoids. totally Incontrast, hermatypic and calcareous sponges, brachiopods, andechi abundant ammonites, belemnite rostra, siliceous Although numerous amateur are collectors ., 2005). The occurring decapod crustacean occurring ., 2005).The et al Tubiphytes ., 1994,1996;Schmid, 1996;Schmid ”) are extremelyand common et al ., 2018). - - - - - Figure 1 Germany. Mapbased onOpenStreetMap(www.openstreetmap.org). taxonomy modifiedafterSchweitzer gert and Kuschel, 2018, and own observations; Formation (compiled after Wehner, 1988, Schwei Locality of the holotype. Arrow indicates location of the Petersbuch quarry at Markt Titting-Petersbuch, Bavaria, southern Titting-Petersbuch, Localitythe holotype. ofMarkt the Petersbuch quarry Arrowindicateslocationof at Nodoprosopon ornatum Laeviprosopon laeve Laeviprosopon laeve Eodromites grandis Eodromites grandis (von Meyer, 1857) (von Meyer, 1857) Boletín de la Sociedad Geológica Mexicana Mexicana Mexicana Geológica Geológica Sociedad Sociedad la la de de Boletín Boletín (von Meyer, 1842) http://dx.doi.org/10.18268/BSGM2021v73n3a030121 et al ., 2018): - airmts scheffnerae Tanidromites Pilidromia thiedeae Petersbuchia thauckei Planoprosopon heydeni Tanidromites insignis Verrucarcinus torosus /73(3)A0301212021 / 73(3)A0301212021 (Schweigert andKoppka, 2011) Schweigert and Koppka, 2011. n.gen.,sp. (herein) (von Meyer, 1857) (von Meyer, 1857) (von Meyer, 1857) 3 3

New crab of the Jurassic in Germany GEOLOGICAL SETTING New crab of the Jurassic in Germany SYSTEMATIC PALAEONTOLOGY 3. Systematicpalaeontology 4 4 The classification The follows Schweitzer obra u obekg.Sraeo carapace to burial due to breakage. Surface of the carapace partly is lost prior left sideof The the carapace. mensionally preserved inner cast of 70622 (Figure 2)isrepresented bythe three-di Description areasHepatic andgastric notseparated by grooves. margins. spatulate with trifid, Rostrum tubercles. pustulose, anterior part with a few prominent protrusive outer orbital spine; Surface coarsely Diagnosis type locality. Occurrence Studied material close toaquarry. adump with the fossilwas found ex-situ on top of 2014). Exactbednottraceable, becausetherock gian, AcanthicumZone;Niebuhr andPürner, buch Member (Upper Jurassic,late Kimmerid Type horizon (Figure 1). Germany Erkertshofen, Markt Titting, Franconian Alb, Type locality: Holotype sole specimen. whostadt-Mörsdorf), collected and donated the Etymology (monotypic). Diagnosis sp., monotypic.Genderisfeminine. Type species / / Superfamily Homolodromioidea Alcock,1900 Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Infraorder Brachyura Linnaeus, 1758 Family Prosopidae von Meyer, 1860 Section Dromiacea DeHaan,1833 Order DecapodaLatreille, 1802 Petersbuchia thauckei : SpecimenSMNS70622(Figure 2). : Carapace subquadrate, with long the type species : See diagnosis of : Named after Thomas Haucke(Frey: Named after Thomas : Atpresentknown only from the Genus : The small-sized: The holotype SMNS Figures 2Ato2D, 3 : Petersbuchia thauckein.gen., : Treuchtlingen Formation, Peters Qar rao Petersbuch/ area of Quarry : Holotypeonly. Petersbuchia n. gen.,sp. nov. http://dx.doi.org/10.18268/BSGM2021v73n3a030121 et al . (2010). /73(3)A0301212021 - - - - /73(3)A0301212021 with subparallel, spring-like ornamentation. In with subparallel, spring-like ornamentation. plane.its symmetry Posterior muscle gastric scars bylateral grooves, withtwo tubercles coarser in from each other. area Mesogastric well separated areasprotogastric notseparated andepigastric orbital area hidden. Hepatic, Frontal aspects of rostrum andprotrusive outer orbital spine. base of in thecentre. Rounded orbital margin connects with tip marginsspatulate and downturned upturned rostrum Trifid mm. 2.1 height maximum width by plane mirroring3.6 mm, insymmetry including rostrummm, reconstructed 4.6mm; rostrum. trifid carapace excluding rostrumTotal 3.7 length of broad with subquadrate, apace car densely pustulose in all areas. Outline of n. gen.,sp. wasanadultorajuvenile. Petersbuchia thauckei sole small-sized specimenof Remarks a sickle-likeoutline. Free fingerlost. tally broken, but slightly curved imprint pointsto dis smooth, thick, finger Fixed carapace. the in propodus as withsimilarpustuloseornamentation as long as broad and widens distally. Surface of individual (Figure propodus 3,inset).The twice is preserved chela possiblybelonging tothesame thereClose tothespecimen, an incompletely is region the not intestinal preserved.area. Sternal smooth rim, with concavities placed laterally from a carapace forming carapace. Posterior margin of branchial region parallel to longitudinal axis of of margin lateral Outer region. branchial adjacent diac region triangular, but hardly separated from Meta- and urogastric regions fusedaswell. Car developed. Epi- and mesobranchial regions fused. Cervical groove and branchiocardiac groove well hepatic region but a lobe. posteriorwards forming rostrum. Subhepatic region notseparated from the region, atone in the epigastric the base of last the and region hepatic/protogastric joint the cervical groove, the third onenear the centre of the hepatic region, close to the the outer third of groove,close tothe mesogastric another one in the carapace, one oped inthe anterior part of four strong andprominent tubercles are devel addition tothe general pustulose ornamentation, : It is impossible to state whether the - - - - Kimmeridgian, AcanthicumZone,Bavaria, southernTitting-Petersbuch, MarktGermany. Dorsal(A,C),lateral(B)and frontal (D)views.A Figure 2 with artificially enhanced contrast. Scalebar=2mm.

Petersbuchia thauckei n. gen. n. sp., holotype, SMNS 70622, UpperJurassic,Treuchtlingen70622, late Petersbuch holotype,SMNS Member; Formation, sp., n. gen. n. Boletín de la Sociedad Geológica Mexicana Mexicana Mexicana Geológica Geológica Sociedad Sociedad la la de de Boletín Boletín http://dx.doi.org/10.18268/BSGM2021v73n3a030121 /73(3)A0301212021 / 73(3)A0301212021 5 5

New crab of the Jurassic in Germany SYSTEMATIC PALAEONTOLOGY New crab of the Jurassic in Germany DISCUSSION 4. Discussion (inset, close-up view).Scalebar =2mm. Figure 3 6 6 ornamentation isusuallyless developedornamentation in inner as anouter orbital spine. Itmust bestressed that the carapace as well of a pustuloseornamentation buchia in as rostrum trifid similar a exhibit species Some species of similarageandlithology. of veryrare taxon reported fromUpper Jurassic beds Laeviprosopon sublaeve thaucke inent tubercles inthe hepatic region of prom of view, arrangement and first number the species assignedtothisnewgenus.the only In Franconia represents the late Kimmeridgian of At present, / /

Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Petersbuchia thauckei n.gen.However, i n.gen.,sp. reminiscentstrikingly to is Petersbuchia thauckei Laeviprosopon (vonMeyer, 1857),another n. gen.sp.,holotype, SMNS70622,withafragmentarychelapossibly belonging tothesameindividual Laeviprosopon n. gen. n., sp. from includingthe type http://dx.doi.org/10.18268/BSGM2021v73n3a030121 sublaeve lacks Petersbuchia Peters /73(3)A0301212021 - - /73(3)A0301212021 ring genus. cially convergent characters with thisco-occur from 2020). Hence, species of 2009a), but this character does not occur within 2020; Schweigert, and Fraaije Starzyk, Klompmaker, (Prosopon von Meyer, 1835s. str.; few other genera safely assignedto Prosopidae ing laterally placed concavities is shared with a pace exhibits atrapezoidal outline. Laeviprosopon sublaeve been even Moreover, stronger ornamented. in of the cuticle casts than on the cuticle. Therefore, if Petersbuchia h otro agno the carapace show posterior marginThe of Laeviprosopon Protuberosa Laeviprosopon n. gen. was preserved n.gen.was itwould have Petersbuchia Schweitzer andFeldmann, and shows only some superfi some only shows and teatro ato the cara the anterior part of (see Klompmaker n. gen. is welln. gen.is distinct Europrosopon et al ., - - - -

References Acknowledgements Bantz, H.U., 1970,DerFossilinhalt des A., 1900, MaterialsAlcock, for acarcinological México, Mexico) for theircareful reviews. de Geología, Universidad NacionalAutónoma de University, USA) and FranciscoVega (Instituto and EarthSciences, Biology LomaLinda of Finally, IthankAlessandro (Deptment Garassino theTreuchtlingen Formation.of ical information who provided Germany), insightfulbiostratigraph imen and Victor Schlampp (Rednitzhembach, who kindly donatedGermany), the studied spec Haucke (Freystadt-Mörsdorf, I thankThomas De Haan,W., 1833, Crustacea, vonin Siebold, Davie, P. J. F., Guinot,D., Ng, P. K.L.,2015, bhl.title.124951 Batavorum, 243p. https://doi.org/10.5962/ Illustravit. J. Müller et Co., Lugduni Observationibuset Adumbrationibus Suscepto, 1823-1830 Annis Collegit, Notis, in IndiaBatava Imperium Tenent, Jussuet Auspiciis Superiorum, qui summum quae in ItinereAnimalium, per Japoniam, P. F. (ed.), Fauna Japonica sive Descriptio org/10.1163/9789004190832_021 II: Leiden, Brill, 1049-1130. https://doi. taxonomy, biology. Crustacea, 9C- The (eds.), Treatise– anatomy, onzoology Schram, F. R.,von Vaupel Klein, J. C. in Castro, P., Davie, P. J. F., Guinot, D., Brachyura, of classification and Systematics 1-86. Erlanger Geologische Abhandlungen, 82, Kimmeridge der Südlichen Frankenalb): Treuchtlinger (Mittleres Unter- Marmors https://doi.org/10.5962/bhl.title.15343 68, 123-169. Bengal, Asiatic Society of the Primigenia or Dromiacea: Journal of India, No. Brachyura 5.The fauna of Boletín de la Sociedad Geológica Mexicana Mexicana Mexicana Geológica Geológica Sociedad Sociedad la la de de Boletín Boletín http://dx.doi.org/10.18268/BSGM2021v73n3a030121 - - Förster, Nachweis R.,1986,Dererste eines Förster, R.,1985,Frühe Anomuren undBrachyuren E., C. Schweitzer, I., Lazăr, M., R. Feldmann, yn, . Sazk N, ois C, Kočová C., Robins, N., Starzyk, M., Hyžný, Haug, J. T., Haug, C.,2014,Eoprosoponklugi Glaessner,Moore, M.,1969,Decapoda,in R.C. decapods Brachyuran 2011, D., O. Franţescu, Guinot, W. M., B. Bakel, van B., H. R. Fraaije, Paläontologie und Historische Geologie, 26, der Bayerischen Staatssammlung für Pliensbach) Mitteleuropas: Mitteilungen brachyuren Krebses ausdemLias(oberes Historische Geologie, 25,45-60. Staatssammlung für Paläontologie und Mitteilungen der BayerischenDogger: (Decapoda, Crustacea) aus dem mittleren theMizunamiFossil Museum,33,1-20. of Dobrogea, Romania: Bulletin of bioherms Prosopidae) from Jurassic (Oxfordian) sponge 2006, New crabs (Decapoda: Brachyura: kl (oten oai, zc Republic): Czech Moravia, (Southern skála and Stránská of Oxfordian Taxonomy the from assemblage 2015, adecapod crustacean of palaeoecology M., Veselská, Palaeodiversity, 7,149-158. and most“primitive” crabreconsidered: (Brachyura) –the oldest unequivocal Press, R400-R533,R626-628. Kansas AmericaandUniversityof Society of Pt. R4(2). Boulder and Lawrence, Geological (ed.), TreatiseInvertebrate on Paleontology, org/10.1127/0077-7749/2010/0110 Abhandlungen, 259,271-297.https://doi. Jahrbuch für Geologie und Paläontologie, and one from Dobrogea, Romania: Neues limestones Kimmeridgian) coral reef species new new genus) from Jurassic (Oxfordian- five (including doi.org/10.18268/bsgm2013v65n2a6 Geológica Mexicana, 65,249-254. https:// the Tanidromitidae: Boletín de laSociedad crab from northwest France; the earliest record of homolodromioid (Bajocian) Jurassic D., Jagt, J. W.2013, AnewMiddle M., 25-31. /73(3)A0301212021 / 73(3)A0301212021 7 7

ACKNOWLEDGEMENTS/ New crab of the Jurassic in Germany REFERENCES New crab of the Jurassic in Germany REFERENCES 8 8 Karasawa, H.,Hirota, T., 2019,Two newspecies reisa E, try, . Shegr, G., Schweigert, N., Starzyk, E., Krzemińska, and Middle 2008, M., Zatoń, M., Krobicki, Kott, R., 1989,Fazies undGeochemie des H. R. Fraaije, N., Starzyk, A., A. Klompmaker, Klompmaker, A.A.,Schweitzer, C.E.,Feldmann, Kato, H.,Takahashi,T., Taira, M., 2010,Late Karasawa, H.,Kato, H.,2007,Newprosopid crabs / / Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín hkk,Jpn ultno the Mizunami Japan:Shikoku, Bulletin of Group (Upper Jurassic–Lower Cretaceous), Longodromitidae) from the Torinosu Planoprosopon(Decapoda: Brachyura: of https://doi.org/10.3140/bull.geosci.1559 Geosciences, 90,633-650. Bulletin of the Middle Jurassic (Bajocian) of England England of (Bajocian) Jurassic Middle the (Decapoda, Anomura, Gastrodoridae) from Eogastrodorus B., 2020,Newrecords of https://doi.org/10.1016/j. H. R. Fraaije, B., Chandler,R. Whicher,J., palaeo.2008.01.025 30-43. Palaeoclimatology, Palaeoecology, 263, their early evolution: Palaeogeography, palaeoenvironmental distribution during brachyuran crabs: Late Jurassic roots of A, 111,1-115. geowissenschaftliche Abhandlungen, Reihe Kimmeridge, Südliche Frankenalb: Berliner Treuchtlinger Unter- und Mittel- Marmors: 23(2), a32.https://doi.org/10.26879/1045 Brachyura): Palaeontologica Electronica, Jurassicand Cretaceous crabs (Decapoda, reef-associated convergent evolution of B., Schweigert, G., 2020,Systematics and https://doi.org/10.1130/G34768.1 crustaceans: Geology, 41,1179-1182. Mesozoic marine reefs onthe rise of of R. M., Kowalewski, M., 2013, influence The doi: 10.1111/j.1475-4983.2010.00960.x Japan: Palaeontology, 53(4), 761-770. https:// Jurassic decapod crustaceansfrom northeast naturale diMilano, 35,62-65. Scienze naturali e del Museo civicodiStoria Japan: Memorie della Società italiana di Upper Jurassic Torinosu Group, Shikoku, (Crustacea, Decapoda, Brachyura) fromthe Fossil Museum,45,27-32. http://dx.doi.org/10.18268/BSGM2021v73n3a030121 /73(3)A0301212021 /73(3)A0301212021 Leinfelder, R.R.,Krautter, M.,Laternser, R., Latreille, P. A.,1802,Histoire naturelle, générale von Meyer, H., 1857, Briefliche Mittheilungen: Briefliche 1857, H., Meyer, von von Meyer, H.,1842,Ueber die in dem dichten Mittheilungen, Briefliche 1835, H., Meyer, von Linnaeus, C., 1758: Systema Naturae per Leinfelder, R.R.,Werner, W., Nose, M., Schmid, rgno Jurassicreefs. Current research origin of Menges, G., Luterbacher, H.,1994, The Zeiss, A.,Schweizer, V., Christmann, H., Schroeder, J. H.,Reinhold, C., Koch, R., R.,Rehfeld-Kiefer,H., Herrmann, U., G., Werner, W., Keupp, H.,Brugger, Nose, M.,Schmid, D. U., Schweigert, v. 3.F. Dufart,Paris. et particulière, desCrustacéset des Insectes, njgpa/2020/0876 2), 147-156. https://doi.org/10.1127/ und Paläontologie Abhandlungen, 296(1- and France: Neues Jahrbuch für Geologie Geologie und Petrefaktenkunde, 1857, 556. Neues Jahrbuchfür Mineralogie, Geognosie, 70-75. Prosopon:Beiträge zur Petrefaktenkunde, 5, vorkommenden Species des Crustaceengenus gelben Jurakalk von Aalen in Würtemberg Petrefaktenkunde, 1835,325. für Mineralogie, Geognosie, Geologie und an Prof. Bronn gerichtet: Neues Jahrbuch org/10.5962/bhl.title.156772 Laurentius Salvius, Holmiae. https://doi. vol1. 10), (ed. locis synonymis, differentiis, ordines, genera, species, cumcharacteribus, Regna tria Naturae, secundumclasses, 227-248. Geologie Paläontologie,und Sonderband, 2, Research Reports: Göttinger Arbeiten zur evolution. on biogenicsedimentation. I.Reef Gunkel, F. (eds.), Global and regional controls Jurassic.In: Reitner, J., Neuweiler, F. & sponge andmicrobolite reefs from the Late coral, growth anddynamics parameters of M., Hartmann, D., 1996,Paleoecology, Takaćs, R., Laternser, M., Krautter, U., D. https://doi.org/10.1007/BF02536932 developments and results: Facies, 31,1-56. Schlampp, V., 2004,DieMalm-Ammoniten Scholz, G., 2020,Eocarcinus Withers, praecursor Niebuhr, B., Pürner, T., 2014,Plattenkalk und Müller, P., Krobicki, M.,Wehner, G., 2000, Jurassic von Meyer, H.,1860,Die Prosoponiden Schweigert, G., Koppka, J., 2011,Decapods Schweigert, G., 2019, An additional representative Schmid, D. U., Leinfelder, R. R., Schweigert, G., Schmid, D. U., 1996,Marine Mikrobolithe und Schlampp, V.,dem aus 2009,Ammoniten taxonomy, andbiogeography: ecology Prosopidae (Decapoda: Brachyura) –their the family and Cretaceous primitive crabs of Palaeontographica, 7,183-222. oder die Familie der Maskenkrebse: Planoprosopon andTanidromites: Neues newspeciesof with descriptionsof andLithuania, Germany Jurassicof (Crustacea: Brachyura) fromthe jcbiol/ruy092 39(1), 36-39. https://doi.org/10.1093/ Crustacean Biology, Journal Germany: of southern fromthe UpperJurassic of Lecythocaridae (Brachyura: Dromiacea) of – areview: Zitteliana,26B, 31-41. Germany Southern the Upper Jurassicof of 2005, Stratigraphy and palaeoenvironments 9, 101-251. Profil, Oberjura: dem aus Mikroinkrustierer 40-48. Treuchtlinger 3, Der Steinkern, Marmor: Fossilien, 21(4),208-219. der Fränkischendelta: 3und Albgamma org/10.1016/j.asd.2020.100991 & Development, 59,100991.https://doi. stem group brachyuran: Arthropod Structure 1932 (Malacostraca, Decapoda, Meiura) is a Bayern): org/10.1127/sdgg/83/2014/5 für Geowissenschaften, 83,5-71. https://doi. Oberjura, Frankenalb Schriftenreihe der Deutschen Gesellschaft der (außeralpiner Weißjura-Gruppe der Frankendolomit – Lithostratigraphie 49-79. Annales Societatis Geologorum Poloniae, 70, Boletín de la Sociedad Geológica Mexicana Mexicana Mexicana Geológica Geológica Sociedad Sociedad la la de de Boletín Boletín http://dx.doi.org/10.18268/BSGM2021v73n3a030121 Schweitzer, C.E.,Feldmann, R.M.,2008a, Schweigert, G., Robins, C., 2016, Earliest Schweigert, G., Kuschel, H., 2018,Schwäbische Schweitzer, C.E., Feldmann, R.M., Schweitzer, C. E., Feldmann, R. M.,2009b, New Schweitzer, C. E., Feldmann, R. M.2009a, Schweitzer, C. E., Feldmann, R. M.,2008b, Revision A new classification for some Jurassic some for classification new A njgpa/2016/0583 280, 325-330. https://doi.org/10.1127/ Geologie und Paläontologie, Abhandlungen, Germany.southern Neues Jahrbuch für Brachyura) from the Upper Jurassicof Lecythocaridae(Crustacea: representatives of Krabben: Fossilien, 35(4),14-23. und Fränkische Alb–ein Paradies fürJura- org/10.1127/0077-7749/2011/0136 Abhandlungen, 260,221-235.https://doi. Jahrbuch für Geologie und Paläontologie, et al Gabriella Collins 2009c, Revision of org/stable/41701785 https://www.jstor.207-224. 111A, Wien, in Annalen desNaturhistorischen Museums Formation, Late Jurassic (Tithonian), Austria: and Feldmann, 2009,from the Ernstbrunn Longodromitidae Schweitzer species of 55-121. Naturhistorischenin Wien, Museums 110A: des Annalen species: new five and genera, fourincluding newfamilies, four new 1969 (Crustacea: Decapoda: Brachyura) the ProsopinaeGlaessner, sensu Revision of org/10.1127/0077-7749/2008/0250-0273 Abhandlungen, 250,273-285.https://doi. Jahrbuch für Geologie und Paläontologie, Prosopidae) including two new species: Neues (Decapoda: Brachyura: Homolodromioidea: the genus Laeviprosopon Glaessner, 1933 of org/10.1007/BF03043911 lethaia, 87, 119-156.https://doi. Tanidromitidae new family:Senckenbergiana Goniodromitidae Beurlen, 1932and Brachyura: Homolodromioidea): Families Brachyura (Crustacea: Decapoda: ., 2006(Decapoda: Brachyura: /73(3)A0301212021 / 73(3)A0301212021 9 9

New crab of the Jurassic in Germany REFERENCES New crab of the Jurassic in Germany REFERENCES 10 10 Schweitzer, C.E.,Feldmann, R.M.,2009d,Revision cwizr C E, edan R M, Lazăr, M., R. Feldmann, E., C. Schweitzer, Schweitzer, C.E.,Feldmann,Garassino, R.M., Schweitzer, C.E.,Feldmann, R.M.,2010d, Schweitzer, C.E.,Feldmann, R.M., Schweitzer, C. E., Feldmann, R. M., 2010b, Schweitzer, C.E.,Feldmann, R.M.,2010a, / / Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín fthe genus Cyclothyreus Remeš, 1895 of org/10.1127/0077-7749/2009/0252-0001 Abhandlungen, 252,1-16.https://doi. Jahrbuch für Geologie und Paläontologie, with newJurassic species:Neues Homolodromioidea: Tanidromitidae) I., 2007, Decapods from Jurassic doi.org/10.1163/ej.9789004178915.i-222 Crustaceana Monographs, 10,222p. https:// fossilcrustacean species: Systematic listof A., Karasawa, H.,Schweigert, G., 2010, doi.org/10.1651/09-3231.1 Crustacean Biology, 30,251-256. https:// knownto date (Jurassic): Journal of Homolodromioidea: Glaessneropsoidea) oldest BrachyuraThe (Decapoda: org/10.1127/0077-7749/2010/0084 Abhandlungen, 258, 51-60. https://doi. Jahrbuch fürGeologie und Paläontologie Longodromitidae) with newspecies. Neues 1861 (Brachyura: Glaessneropsoidea: genus2010c, The Coelopus Étallon, Wien, SerieA,112,169-194. Annalen des Naturhistorischen Museumsin Homolodromioidea: Goniodromitidae): notes on Eodromites (Brachyura: Cycloprosoponand additional Revisionof org/10.1127/0077-7749/2010/0059 Abhandlungen, 256,363-380.https://doi. JahrbuchNeues Paläontologie fürGeologieund (Homolodromioidea: Goniodromitidae): Distefania Checchia-Rispoli, 1917 of (Glaessneropsoidea) and reevaluation Mesozoic Brachyura A newfamilyof org/10.1127/0077-7749/2009/0253-0357 Abhandlungen, 253,357-372.https://doi. Jahrbuch fürGeologie und Paläontologie (Decapoda: Brachyura: Dromioidea): Neues http://dx.doi.org/10.18268/BSGM2021v73n3a030121 /73(3)A0301212021 /73(3)A0301212021 try,N,21a epria fthe primitive Starzyk, N., 2015a, Reappraisal of Lazăr, M., R. Feldmann, E., C. Schweitzer, Zeiss, A., 1964, Zur Malm Gamma/Delta- Wehner, G., 1988,Über die Prosoponiden Krzemiński, E., Krzemińska, N., Starzyk, the genus Starzyk, N., oldest species of 2020,The the genus Starzyk, N., 2015b, Newspeciesof rbEdoie,wt ecito fcrab Eodromites, withdescriptionof https://doi.org/10.1127/njgpa/2018/0744 Paläontologie, Abhandlungen,288, 307-341. Romania. Neues Jahrbuchfür Geologie und the Carpathians, the Late Jurassicof 2018, O., Franţescu,Decapoda (Anomura; Brachyura) from G., Schweigert, I., org/10.1127/0077-7749/2007/0244-0099 Abhandlungen, 244,99-113. https://doi. Jahrbuch für Geologie und Paläontologie, Nodoprosopon Beurlen, 1928:Neues of Dobrogea, Romania and reconsideration (Oxfordian) of sponge megafacies 104-115. NO-Bayernund angrenzende Gebiete, 14, GrenzeFranken: in Geologische Blätter für München, Germany. zu Ludwig-Maximilians-Universität (Crustacea, Decapoda): Doctoral thesis, org/10.1127/0077-7749/2012/0219 Abhandlungen, 263,143-153.https://doi. Jahrbuch fürGeologie und Paläontologie Brachyura: Goniodromitidae): Neues Poland (Decapoda: the Oxfordianof W., 2012,Anewcrabspeciesfrom https://doi.org/10.3409/azc.63.02 Acta Zoologica Cracoviensia, 63, 9-13. Poland: Prosopidae) fromthe Oxfordian of Laeviprosopon(Decapoda: Brachyura: azc.58_2.129 58, 129-136. https://doi.org/10.3409/ Poland: Acta Zoologica Cracoviensia, Tanidromitidae) from the Oxfordian of Tanidromites (Decapoda: Brachyura: https://doi.org/10.26879/513 Palaeontologica Electronica, 18.3.50A,1-19. (Decapoda: Brachyura: Goniodromitidae): three new speciesfrom European localities