No. 3 1981

SER. B VOL. 28 NO. 2 Norwegian Journal of Entomology

PUBLISHED BY NORSK ZOOLOGISK TIDSSKRIFfSENTRAL Fauna norvegica Ser. B Norwegian Journal of Entomology Norsk Entomologisk Forenings tidsskrift

Appears with one volume (two issues) annually norske biblioteker. Andre ma betale kr. 55, -. Disse Utkommer med to hefter pr. ar. innbetalinger sendes til NZT, Zoologisk museum, Editor-in-Chief (Ansvarlig redaktod Sarsgt. I, Oslo 5, Ole A, Srether, Museum of Zoology, Museplass 3, Postgiro 2 34 83 65. 5014 Bergen/Univ, Editorial Committee (Redaksjonskomite) FAUNA NORVEGICA B publishes original new in­ Arne Nilssen, Zoological Dept., Troms\') Museum, formation generally relevant to Norwegian entomo­ N-9000 Troms\'), John 0, Solem, DKNVS Museet, logy. The journal emphasizes papers which are main­ Erling Skakkes gt. 47B, N-7000 Trondheim, Albert ly faunistical or zoogeographical in scope or con­ Lillehammer. Zoological Museum, Sars gt. I, tent, including checklists, faunallists, type catalogues Oslo 5, and regional keys, Submissions must not have been previously published or copyrighted and must not be Subscription published subsequently except in abstract form or by Members of Norw,Ent.Soc. wiIl receive the journal written consent of the Editor-in-Chief. free, Membership fee N,kr. 50,- should be paid to the Treasurer of NEF: Lise Hofsvang, BrattvoIlveien NORSK ENTOMOWGISK FORENING 107, Oslo 11. Postgiro 5 44 09 20. Questions about ser sin oppgave i a fremme det entomologiske stu­ membership should be directed to the Secretary of dium i Norge, og danne et bindeledd meIlom de in­ NEF. Trond Hofsvang, P.O, Box 70, N-1432 As ­ teresserte. Medlemskontingenten er for tiden kr. 50.­ NLH, Members of NOF receive the journal by pr. if. Henvendelse om medlemskap i NEF sendes paying N,kr. 35,-, non-members by N.kr, 55,- to: sekretreren: Trond Hofsvang, Postboks 70, 1432 As­ NZT, Zoological Museum, Sarsgt. I, N-Oslo 5, Post­ NLH. Medlemmer far tidsslaiftet fritt tilsendt og kan' giro 2 3483 65, Outside Fennoscandia: Additional abonnere til redusert pris pa FAUNA NORVEGICA postage Nkr. 5, - per year (surface mail), serie A (genereIl zoologi, I hefte pr. ad for kr. 15,- og pa serie C (ornitologi, 2 hefter pr. ad for kr. 25,-, Disse innbetalinger sendes til NZT,Zoologisk mu­ Abonnement seum, Sarsgt. 1, Oslo 5, Medlemmer av Norsk Entomologisk Forening far Postgiro 2 34 83 65. tidsslaiftet fritt tilsendt. Medlemskontingent kr. 50,­ innbetales til kassereren i NEF: Lise Hofsvang, Bratt­ voIlveien 107, Oslo 11. Postgiro 5 44 09 20. Medlem­ mer av Norsk Ornitologisk Forening mottar tids­ Trykket med bidrag fra Norges almenvitenskapelige skriftet ved a betale kr. 35, -, dette gjelder ogsa forskningsract. Opplag 800

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Kristiansen & W\')ien, Oslo, ISSN 0332-7698

Fauna (Norsk Zoologisk Forening) har fra na gart ut av Norsk Zoologisk Tidsskriftsentral. Avtalen om gjen­ sidig reduserte abonnementspriser pli foreningenes tidsskrifter vii for fremtiden derfor bare gjelde mellom Norsk Entomologisk Forening og Norsk Ornitologisk Forening.

1948. Ueber die Ausbreitung und die Ausbreitungs­ 1961. Opiliones. Soulh A{riculI Allimal Lile 8. umstande der Wechselkrote (Bul;) I'iridis Laud 1-197. im Ostseegebiet. LUllds ullil'.Arsskr. N.F. 2, 1962. Grod- och kraldjur. Diure!1S Vdrld 7, 1-560. 44(/21, 1-30 Malm0. 1949. Om ni!gra zoologiska iakttagelser pi! Gotland. 1963. Harvestmen 10piliones) from the Azores. Faulla Flora Upp.~. 44, 201-207. Bolll1.Mu.\. mUllie. FUllchal 17 (58). 10-18. 1949. Die Odonaten-Fauna von Estland. Apopl1iJrda 1964. Ober Bremsen (Tabanidae. Diptera) von Slid­ Tarll/ellSia. Soe. Lil/. E\llJIlicu ill Suecia, Stock­ west-Hame, Finland. Loullai.\-Hiimeell Luoll/o holm, 408 -420. 15.39-45. 1949. Einige Odonaten-notizen aus Gotland. Opuse. 1964. KJockgrodan i Ski!ne. Skdll<:\ Na/ur 51. 2p. 1'111. 14, 26. Lund. 1950. Oks kild CraCI<:S illlauslus Jakobsoni loodus­ 1964. The influence of postoptimal temperature on teaduslikus tegevusest. Ober einen Fund van the development and ossification of the cranium eracles illlau.\I<:S in Estland. Tulimuld, of Ralla <:Kul<:ll/a L. LUl1ds UlliI'Ars.\kr. N.F. 2. RCI'E\IUI& Sci. I, 148-152. 59(9),1-18. 1951. Bemerkungen liber schwedische Tabaniden. 1964. Uber die nordskandinavischen Tahallidell ()1)IISe. 1'111 16, 97 -I09. (Dip/era), Opuse. 1'11/ 29, 99 - 108. 1951. Om lovgrodans forekomst pi! Gotland. Faulla 1965. Concernng Haloral<:S repmhu.\ 10.P. Cambrid­ Flora U/'p.\ 46, 244-246. ge) (Araneae. Erigonidae) occurrence in Scandi­ 1953. A few lines from the One Hundred-year-old navia. Norsk 1'111. Tid.\skr, 13. 16. history of the Naturalist Society at the University 1965. Rolf Krogerus in memoriam. Nor.\k m/. of Tartu (Dorpat). Tulillwld. Rel'.EILiI.& Sci. 4 Tidsskr. 13. 130-131. (3). 1-9. 1965. Richard Frey in memoriam. Nor.\k 1'111. 1954. Harvest-spiders from S. W. Australia. Report Tidsskr 13.131-132. from professor T. Gislens expedition to Australia 1966. Ekologiska faktorer och kroppsformen hos in 1951 - 52. I.UIld.\ Ullil,Arsskr. N.F. 2, 50 (j 11, Opiliolles. Norsk ell/ Tids\/..r. 13. 261 - 264. 1-10 1966. En kolleksjon av Araneae og Opiliones fra 1954. Ober die systematische Stellung der europai­ Sogn. Norsk 1'111. Tids.\kr. 13. 394-395. schen grlinen Frosche RllIIU <:.\culell/a L. und R. 1967. Tabanidae et Leptidae. LiIll//lJlalllla Eumllam, ridihullda Pall. LUIld.\ Ullil·.A rs\kr. N.F. 2. 50 (j 2I, 410 -411. Stuttgart. Gustav Fischer verlag. 1-30. 1967. Naturvern og menneskenes milj0. Norsk Nalllr 1954. Bemerkungen liber schwedische Tabaniden 11. 3.26-29. O/IUSe. <:111 19. 239- 244. 1968. Taxonomy-ecology. a mutual relationship. 1955. Das licht als hemmender Faktor bei der Larve­ Opuse.elll. 33, 24- 27. nentwicklung der Wechselkrbte (Bul;1 I'iridis 1968. Uber die norwegischen Tabaniden (Diptera). Laud Oikos 6. 78 - 91. Norsk 1'111. Tidsskr 15,63-64. 1956. En ski!nsk spindellokal. FaullaFlora UJ!f's. 51, 1969. Uber bremsen (Tabanidae, Diptera) von Slid­ 179-181. west-Hame, Finland. 11. Loullais-Hiim<:et1 Ll/IJII/o 1957. Ober die geographische Variabilitat einiger 30,1-4. Merkma~ bei R. eSClllell/a L. A1111. Soe. Tarl.ad. 1970. Faunaen pi! 0.5 m2 i engvegetasjon pi! h0yfjel­ res Nal. /111'. COIlSI. I, 148 -151. let. lHardangervidda) ISP. Non/ell 4. 1957. Zur geschichte der Naturforscher-Gesellschaft 1970. Amfibiene. NorKes D.I'r. 3. 314-333. Oslo. in Tartu (Dorpatl, I. 1853-1868. AIIII. Soe. 1970, Krypdyrene. NorKes Dl'r. 3. 334-353, Oslo. Tarl.ad. res Nal. 1111'. COIISI. I. 177 - 196. 1970. 0kologi - ein viktig vitenskap i dag. SI'II OK 1957. Hundid Eestis. Wolves in Estonia. Tulimuld. SeK", 421-432. ReI'EIlI.LiI. & Sci. 8.310-315. 1970. T. Moldung & T. Solh0y,: Turnbull and Ni­ 1958. Uber zwei fUr Europa und Nord-Amerika ge­ choll's «Quicktrap» for acquiring standning crop meinsame Tabanidenarten. O/JIIse. 1'111. 23. of evertebrates in high mountain grassland com­ 95-104. munities. Norsk 1'111. Tid\skr. 16. 133 -136. 1958. Bromsar och styngflugor. SI'ellska dillr, IIlS<:k­ 1971. Vevkjerringene og moseskorpionene. NorK<:s lama. S/ockholm. 547-553. D.I'r 4.142-147 1959. & T. Gislen. Zoogeography of the Swedish 1971. KJeggene. NorKes D.I'r. 4, 346 - 350. Oslo. amphibians and reptiles with notes on their 1971. Landjorden som milj0. NorKe.\ D.I'r. 5, growth and ecology. Acla Verl<:hr. I. 191-397. 278-288. Oslo, 1959. Die RassellhildulIK hei eumpdischet1 Ralla-ar­ 1971. Jordbunnens dyreliv. NorKes pI'r 5, 288-295. II'll ulld die GulliKkeil der KlimareKelll. Lund. Oslo. 1-172. 1971. & G, Lid, DI'relil' /Ia HardallKerl'idda. ((Har­ 1959. Die Wolfsplage in Estland. Balli.\che Helie, 5, dal1Kerl'idda, llalllrl'em - kralill/h.l'gKillK. Re­ 235-239. dakt0r Olav R. Skage Universitetsforlaget. 1960. Tvarflackig snog fran Kalmar lan. Faulla Flora 34-40. Oslo, Bergen, Troms0. Upp.\. 55, 77-78. 1971. & T. Moldung & T. Solh0Y: Notes on Turnbull­

so Nicholls' Quick-trap method. Acla e/I/. /el1l1. 2R, 1975, T. Solhoy, E. 0stbye, A. Hagen, L. Lien and 106-107. H.-J. Skar. Faunal structure of Hardangervidda. 1971. A drive vassdrag «regnskapsmessig», forun­ . Fennoscandian Tundra Ecosystems, dersokelser. konsekvenser, tiltak. 1 Rolf Vik (red. Part 2. Ecol. Sll/d. 17,29-45. «Vassdrag og samfunn», 151-159. Universitets­ [975. T. J. Moldung & A. Fjellberg: Respiration forlaget. Oslo. Bergen. Tromso. rates, winter and summer activity in Collembola [971. Preface. FiJll/w o{lhe Hardal1gerl'idda I. Un i­ on Hardangervidda. Fennoscandian Tundra Eco­ versitetsforlaget 1972. Oslo, Bergen. Tromso. systems, Part 2. Ecol.Swd 17. 117 - 121. 1971. Notes on Turnbull-Nicholls' Quick-trap met­ 1976. Havfruer som proteinreserve? Nail/reil. hod (summary). Acla e/II. r"'"". 2R. 106-107. 81-84. 1972. Zoologiske samlinger og den nye museumsor­ 1976. The scandinavian populations of H.I'l'0ll1ilra ganisasjonen. Museene. 1I11lSIillil1gfra kOl17ileel1lil polari.\ (Frey) (Diptera, Tabanidae). - EII/.swl1d. IIlredl1il1g 01' I17I1Seelle.~ .\Iillillg illllell IIl1in',-,\ileIS­7.145-[49. slrukluren. 55 - 59. Universitetsforlaget. Oslo, 1976. Salamandre, pattedyr og krypdyr. Nalllrell. Bergen. Tromso. 134-144. 1972. Forord. Redaksjonen av Aurlalldseh'en. Et 1976. & T. Solhoy: Ecological investigations at Stigs­ symposium angaende rennende vanns okologi. tuv on Hardangervidda (Norwegian IBP project>. Zoologisl, MlIseUIl1. Bergell 1972 Fall/la Hardal1gerl'idda 7-11.47-54. 1972. List of at Stigstuv area, Har­ 1977. Mire invertebrate fauna at Eidskog, Norway. dangervidda. Norway. Evertebrata [ (Red.), VII. Opiliones. Norw.J. fil/. ]4. 111-112. 1-14. IBP Nonm\' 197]. (Stensil). 1977. & T. Andersen: Horse (Dipt. Tabanidae) 1973. G.G. Raddum & S. Mossige: Temperatur­ new to Norway. Norw.J.EII/. ]4. 177. forholdene for Strondaelva fm og etter regulering. 1977. Tabanidae. Lil17l1ofiJIIl1a ElIl'Ol'aca. J. Oppl. 3 l.a!Jora/orilll17 .fill' r",rsk IWll/s"kologi Zoologi.\k pp. Stuttgart. Gustav Fisher Verlag. lI1u.\el/lll. Ul1il". i Bergell. 7. 1-7, (stensill. 1979. Om faunan pa fredade omraden pa Johannis­ 1974. Bromsar (Diptera:Tabanidae) i Messaureomra­ hus fideikommiss. Blekillge.\ Nalllr 1979. 62 - 84. det. Norr!JO/lellS Nall/r. I. 71 -72. 1979. Carl H. Lindroth 1905-1979 Fall/lO IIO/T. 197 5. Zoologiske undersokelser pa Hardangervidda Ser. B. J6. 2-4. 1967 - 72. l.a!Joraloriull1/iJr r",rskl'al1l1wkologi og 1980. D, Terrestrial invertebrates of the Faroe Is­ illllelllal1dsfis/,e. 17. 1- 16. lands: II. Harvest-spiders (Opiliones). Falll1a I/Orl·. 1975. Muscel 15() or I Zoologi.\/' II1USel/lll. 34-37. ser B. 27. 72 - 75. Universitetet i Bergen. 1975. & E. 0stbye. Introduction. Fennoscandian [ tillegg tallrike bokanmeldelser i bl.a. Naturen. Tundra Ecosyst. Pan 2. Eeol.Swd. 17. 3-7. Norsk Entomologisk Tidsskrift. Fauna och Flora 1975. Area, size and niche breadth at higher latitu­ og Fauna. Oslo. des. Fennoscandian Tundra ecosystems, Part 2. Em/.SwJ 17. 8-18. Lita Greve

SI Terrestrial invertebrates of the Faroe Islands: Ill. (Coleoptera): Check-list, distribution, and habitats

SVEN-AXEL BENGTSON

Bengtson, S.-A. 1981. Terrestrial invertebrates of the Faroe Islands: [JJ. Beetles (Coleoptera): Check-list, distribution, and habitats. FUlIna 11IJrl'. Se,.. B. 28. 52-82.

During the summers of 1978 -1979 beetles were collected in III localities on 17 of the Far­ oe Islands. A total of 101 species were recorded, 16 of which for the first time in the Faroes; I'iz. Hydrophilidae: He/o{Jhorlls flQ1'i{Jes (Fabriciusl, Staphylinidae: Proteilllls brachY{J/ertls (Fabricius), O/oflhrl/lnfllscl/Il1(Gravenhorst), Les/el'u 1I1011/ico/a Kieswetter, L. heeri Fauvel, Lalhrobillll1 brlll/l1i{Jes (Fabricius), XUllIho/illllS Iil1l!Qris (Olivierl, Qlledills boo{Joides Muns­ ter, My//ama brel'icornis Matthews, /14ycelofJorlls rlljescl!I/s Stephens, Atheta hanf()odi Wil­ liams, A. /lIridi{Jel1l1is Mannerheim, : Oliorrh.n1chm singll/uris L.. Micrcllls er­ icae Gyllenhal, Rhit/ot/clIs {Jericar{Jills L., and Silolla {Jl/Ilc/ico//i\ Stephens. The total coleop­ terous fauna now comprises at least 168 species of which 98 % occur in Scotland, 79 l!(; in Shetland, 90 % in , and 52% in Iceland. About one-forth of the species ap­ pear to be widely distributed and common within the Faroes. Many species have wide ecolo­ gical amplitudes. Differences in species richness between habitats are discussed. Faroese dis­ tribution maps for all 101 species found are presented.

S,-A. Bengtson, Dept of Animal Ecology, University of Bergen, N-5014 Bergen/L!niv., Norway.

INTRODUCTION The Faroe Islands consist of 18 small islands si­ tions made by Danish zoologists in 1924-1928 tuated in the North Atlantic (approx. 62°N) and published records. West's list included some about 675 km W of Norway, 300 km NW of 156 species (some of which are doubtful; see be­ Shetland, and 450 km SE of Iceland, The cli­ low) to which our survey added only 16 (pos­ mate is Atlantic with frequent and strong winds, sibly 17) but greatly increased our knowledge of much rainfall, mild winters, and cool summers, the inter-island distribution of many species, The meanltemperatures of the warmest (August) This paper is one in a series presenting the fau­ and coldest (February) months are I I, I and 4, I nistic results of our field-work, accounts of the 0c, respectively, The landscape is greatly affec­ spiders and opilionids have already appeared ted by farming and sheep-grazing, and the vege­ (Bengtson & Hauge 1979 and Kauri 1980) and is tation is dominated by grass heaths. Natural followed by reports on the lumbricids (Enckell shrubs or woods are completely absent. The & Rundgren in prep.), terrestrial gastropods biota is almost entirely West European, (Solh0y 1981), and possibly a few more taxa. In the summers of 1978 and 1979 a Swedish­ These purely faunistic papers will form the basis Norwegian team made a survey of certain terre­ for forthcoming ecological papers. strial invertebrates on the islands in order to apply modern theories on island ecological bio­ geography, and to assess human influence on MATERIAL AND METHODS the present-day terrestrial fauna (see Bengtson & The islands in the Faroe group range in size Hauge 1979), For various reasons (sampling from 374 km2 (Streymoy) to 0,8 km2 (Litla Di­ methods, etc.l attention was focussed mun) and we worked on 17 of them, i,e. on all on the taxa Coleoptera, Araneae, Lumbricidae, but the smallest, and uninhabited Litla Dimun and terrestrial Gastropoda, Here we describe the (Fig, I). Since the field-work has previously been faunistic results concerning the beetles. described in detail by Bengtson & Hauge (I979; Previously West (I930) has treated Coleopt­ especially in Appendix [) only pertinent infor­ era in the series «The Zoology of The Faroes», mation concerning Coleoptera will be briefly basing his large paper on the extensive collec- outlined here, We collected in I1I localities

52 Fal/I/a I/on·. Sa. B 28· 52-82. Oslo 1981 KALSOl' VIDOY ~ in houses and barns which might have produced I EYSTUROY f',_ KUNOY (' '!,! I some synanthropic species. The possible effects STREYMO , 286 \diil.~ 1-"1',,°' r-,"," FUGLOY, I y 1\, /) I I I,' 11­'~ ofour limited seasonal coverage is difficult to as­ ".. :0 ~, ,', \'.. ' ;':\ oe/ .~ ~ \'\f-J, '<7 sess. We found about 55 % of the species listed r. \ 'I ,.,\1 .,,/,/fD by West (] 930), who included material collected : • \ \•• ,0 <.-~'_'i-' ) , ~. '.' \, ra " v) at all times of the year, but most of the missing VAGAR " " ~., ~ ,'" '~~ ~" his far more extensive seasonal and habitat co­ KOLTUR ;-a .' \ NOlso ?s ~ ­ ,c' y verage. HE~TUR '~ Ue~ The collecting localities were assigned to one of 9 different habitat categories (see Bengtson &

SANDOl' • Hauge 1979: Appendix I), The main division '.'-'~ was between «infields» and «outfields». The in­ , . .­ field localities are the fenced-off areas surroun­ S~uvoy .- _'beetle Helophorus flavipes (Fabrici­ table for pitfalls or sieving (e.g. cliffs). The main us), 11 species of staphylinid beetles Protein liS shortcomings of the field-work were the restric­ brachypterus (Fabricius), Olophrul/l fuscu/Il ted seasonal coverage and the lack, or underrep­ (Gravenhorst), Lestel'a I/lonlicola (Kieswetter), resentation, of some habitat types. For instance. L. heeri (FauveO, Lathrobi/ll/l brunnipes (Fabri­ few localities covered typical sea-shore habitats cius), Xantholinus linearis (Olivier), Quedius (might have yielded Bembidion-species and some boopoides Munster, Myllaena brevicornis Matt­ rove beetles), high mountain-sites (> 400 m), hews, Mycetop0rLIs rl({escens Stephens, Atheta and freshwater localities (we certainly missed se­ harwoodi Williams, and A.luridipennis Manner­ veral species of Dytiscidae). Nor did we collect heim, and 4 species of Curculionid beetles

53 Table 1. List ofspecies ofColeoptera (N =101) collected in 1978 -1979 in the Faroes. The table gives: number of islands in (n = 17), number of localities (in = Ill), total number of individuals, and number of localities in different habitats. The abbreviations for habitats are: (A) mountain sites, (B) dwarf shrub heaths, (C) planta­ tions, (0) cliffs, shelves, and crevices, (E) lowland bogs, (F) grass heaths, (G) infields: outskirts, (H) infields:'set­ tlement, and (I) sand dunes. For further information concerning the habitats (number of localities, vegetation etc) see Bengtson & Hauge (1979: Appendix I) and the text.

Number of Habitat (no. of Iocs) Species Islands IDes Inds A B C D E F G H I (4) (l0) (4) (3) (3) (31) (34) (21) t (1)

Q.A.edius sp. 9 13 60 2 - 1 - 6 3 1 '.f whinuB signatus Gr. 16 47 4061 - 1 2 -- 4 22 18 T Zatico Zlis Gr. 1 3 11 - - -- 2 1 H.upocyptus laeviusculus Mnh. 5 5 8 - - 2 2 1 M./Uaena brevicornis Matth. 1 2 2 1 ---- 1 A::/ischa analis Gr. 9 22 138 2 1 1 - 1 14 SpaUa circeUaris Gr. 7 12 50 -- - - - 2 7 M:'cetopoY'Us rufescens SteFh. 1 1 1 kdaUa puncticoUis Sharp. 1 1 1 A';heta gregaria Er. 5 7 13 A me lanocera '!horn. 3 4 6 A islandica Kr. 9 13 53 A exceUens Kr. 10 19 42 A amicula Steph: 1 1 1 A nigricornis '!horn. 1 1 3 A. trinotata Kr. 2 2 15 ------2 A. graminico Za Gr. 3 5 82 1 - - - - 2 1 1 A. vestita Gr. 3 3 8 - - - - 1 - 1 A atramentaria Gyl1. 10 18 34 1 1 - 1 - 3 la 2 A. celata Er. 2 2 2 ------1 A. fungi Gr. 3 4 8 - - 1 - - - 1 2 A hartJoodi Will. 1 1 1 ------1 A. ZU.Y'idipennis Mnh. 2 2 2 ------2 Atheta sp. 7 12 16 -- 1 1 2 6 2 Oealea picata SteFh. 7 12 53 -- -- - 5 7 '·.Y - O::;ypoda haemorrhoa Mnh. 3 3 8 ------1 1 Aleochara sparsa Heer. 2 2 2 ------1 1 Hypo Zi thus riparius F. 16 51 285 - 5 2 2 - 16 17 9 EL-odes minuta L. 3 4 6 -- - - - 1 1 2 S:.mplocaria semistriata F. 9 14 50 -- -- - 1 8 5 B!)rrhus fasciatus Forst. 2 3 5 1 - -- - 2 A7.omaria apicaUs Er. 2 2 30 Aiomaria sp. 3 3 5 - -- - 1 Ci'yptophagus scanicu8 L. 6 7 247 - 1 - -- 1 C distinguendus Stunn 1 1 1 ------1 i C piZOBUS Gyll. 1 1 62 Lathridius minutus L. 6 6 77

Anobium punctatum D;! G. 1 1 1 ChrysoUna staphyZea L. 6 9 22 - 1 - - - 4 3 1 01;iorrhynchus arcticu8 o. Fabr. 15 33 102 1 5 - 2 - 7 11 6 0 dubius Strm 11 17 23 1 2 3 2 - 4 3 2 O. singuZaris L. 2 2 10 -- 1 - - - - 1 BO.Y'ynotus SquamosU8 Germ. 9 14 20 - 1 -- - - 9 4 Ti>opiphorus obtusus Bonsd. 6 9 12 1 - -- 1 5 2 Micrelus ericae Gyll. 2 2 3 Riiinoncus pericarpius L. 3 4 5 5, tona puncticoZZis SteFh. 1 1 2 Cf·utorrhyn.ehus contractus Mrsh. 3 3 3 Arion haematodes Kirb. 3 5 7

54 Nurrber of Habitat (no. of locs) Species Islands Locs Inds A B C D E F G H (4) (l0) (4) (3) (3) (31) (34) (21) (1)

CaY'ahus pY'ob lematicus Hbst. 7 17 56 4 5 3 NebY'ic: gyllenhali Schh. 16 82 2990 20 27 20 N. salina Fairm. 16 91 908 27 27 20 Notiophi lus aquaticus L. 2 '3 4 3 N. biguttatus F. 15 53 198 26 8 4 LOY'iceY'a pilicoY'nis F. 13 40 168 71 21 9 PatY'obus septentY'ionis Dej. 16 92 3323 26 29 18 P. assimi lis O1aud. 8 39 355 10 14 5 P. atY'oY'ufus StrOm 6 9 300 4 4 TY'echus obtusus Er. 17 80 1254 19 27 19 T. fulvus Dej. 1 1 1 PteY'ostichus adstY'ictus Esch. 7 19 109 6 1 P. nigY'ita F. 8 23 64 10 1 P. diligens Sturm 3 5 33 1 1 Calathus fusc:-ipes Cb2ze 13 48 901 19 14 C. me lanocephalus L. 15 36 455 14 12 AmaY'a aulica Panz. 7 23 167 14 8 TY'ichocellus placidus Gyll. 1 1 Hydr'opoY'us sp. 2 6 Agabus so lieY'i Al.lbe 2 2 1 He lophoY'us aquaticus L. 3 2 H. bY'evipalpis Bed. 8 17 4 H. flavipes F. 6 8 29 5 Anacaena globulus Payk. 11 14 27 3 2 MegasteY'num obscUY'um Mrsh. 15 42 288 18 18 Catops fuliginosus Er. 4 4 7 1 Hydnobius sp. 1 2 21 1 Stenichnus collaris MUll. 5 5 7 4 ACY'OtY'ichis sp. 12 27 171 13 9 PY'oteinus bY'achypteY'us F. 3 2 Oma l i um Y'i vulaY'e Payk. 11 23 137 11 O. excavatum Steph. 9 12 27 10 O. caesum Gr. 1 1 1 Eucneeosum bY'achypteY'wn Gr. 15 36 113 14 OlophY'urn fuscwn Gr. 6 7 187 Xy lodY'omus depressus Gr. 1 1 X. concinnus Mrsp.. 1 1 Lesteva longe ~uJ~Y'a ta Goeze • 14 34 111 13 L. monticola Kiesw. 1 2 L. heeY'i Fauv. 3 MicY'alymma maY'inwn Strom 1 Stenus impY'essus Germ. 12 27 47 S. nitidisculus Steph. 6 9 10 S. bY'unnipes Steph. 7 15 27 LathY'obium fulvipenne Gr. 2 2 2 L. bY'unnipes F. 1 1 Xantho linus l ineaY'is 01. 1 3 1 Othius punctulatus Goeze 8 12 15 1 5 3 O. angustus Steph. 15 60 260 14 24 14 O. myY'mecophi lus Kiesw. 15 44 98 14 13 11 Ph i lon thus fime taY'i us Gr. 4 5 8 2 3 P. cephalotes Gr. 4 5 6 4 P. maY'gi na tus Strom 1 3 1 quedius cUY'tipennis Bernh. 11 33 73 1 13

,~'. ;nt-;jon;l l'(ri.U2 Mrsh. 9 15 63 10 Q. umbrinus Er. 11 28 52 12 q,. ni tipennis Steph. 9 28 46 9 (:2. bocpoide:s Munst. 4 11 35

54 a Otiorrhynchus singularis L., Micrelus ericae (1974), and unpublished data based on large and Gyllenhal, pericarpius L., and Sitona only partly worked-up collections made by my­ puncticollis Stephens. To this list may be added self and co-workers in the 1970's. Information the staphylinid beetle Quedius curtipennis Bern­ on Western Norway is taken from «Catalogus hauer which we found to be fairly widely distri­ Coleopterorum Fennoscandiae et Daniae» (ed. buted, whereas we failed to obtain any Quedius Lindroth 1960). fuliginosus Gravenhorst which is previously re­ The Appendix contains maps giving the dis­ ported to occur «in almost all the isles of the Fa­ tribution in the Faroes for all species recorded in roe group and to be common» (West 1930). For 1978 -1979. Previously published records were the time being I regard the two species as syno­ entered on the maps (using different symbols) nymous. West (1930), on the basis of the exten­ whenever they substantiated the picture. sive Danish collections from 1924-1928 and Family Carabidae previous published records, compiled a list of 156 species including some determined to genus Carablls problemalieus Herbst. Found in 17 localities Oocs 5, 6, 7, 13, 14, 17,27, only. However, 5 species (viz. Hydnobius punc­30,46,47,68,69,76,101,105,106, 109)on Stre­ tatus Sturm, Philonthus politus L. (aeneus Ros­ ymoy, VagaL Mykines, Eysturoy, Bordoy, Vi­ si), Tachinus pallipes Gravenhorst, Atheta debilis doy, and Svinoy. Previously recorded in several Erichson, and Otiorrhynchus rugifrons Gyllen­ localities including on Suduroy (West 1930). Pre­ ha]) are questionable for one reason or another fers dwarf shrub (Callulla) heaths but is found (see below) and in need of confirmation. Anot­ also in infield and outfield grassland sites. Occurs her species (Calathus micropterus Duftschmid) is in Se, Sh, I, and WN. For a discussion on the sub­ only found subfossil. With the addition of Ca­specific rank of the North Atlantic populations see tops (Sciodrepa) watsoni (Spence) (Walker 1938) Lindroth (1968). Nebria gyllenhali Seh()nherr. and Otiorrhynchus rugosostriatus Goeze (Lars­ Found in 82 localities Oocs I. 3, 4, 6, 7, 8, 9, 10, son & Gigja 1959) and our abovementioned 16 11,17,18,21,22,25,26,27,28,29,30,31,32, species the total number of species on the Faroe 34,35,36,37,38,39,41. 42, 43, 44, 48, 51. 52, list now stands at 168, not including the doubt­ 53. 54, 55. 56. 57. 58, 59, 60, 61, 62, 63, 64, 65, ful species and those only identified to genus. 66,67,68,69,70,73,74,75,76,78,79,81,82, For each species is given the localities (the 83, 84, 87, 90, 91, 93, 94, 95, 96, 97, 98, 101, numbers refer to those in the Appendix I 102,103,104,105,106,107,108,109,110, 11]) published by Bengtson & Hauge (1979)) and is­ on all islands except on Skuvoy. Found in all lands on which it was collected in 1978-1979. types of habitats though particularly abundant in infields around settlements and on the sand dunes Habitat preferences, as found by us, are briefly on Sandoy. Occurs in Se, Sh, I, and WN. described (see also information in Tab. I). Habi­ Nebria sa/ilia Fairmaire & Laboulbene ( = N. iberiea tat and community structure of carabid beetles Oliviera). have been descr~bed and discussed in a previous Found in 91 localities Oocs I, 3,4,5, 6, 7,8,9, 10, paper (Bengtson 1980). As to previous records I 11. 12, 13, 16, 17, 19,20,21. 22, 23, 24, 25, 26, mainly refer to West (1930) but only in a very 27, 28, 29, 30, 32, 33, 34, 35, 36, 39,40,41, 42, condensed form and particularly emphasising 43, 44, 45, 46, 47, 48, 49, 50, 51, 52, 53, 54, 55, information on status and occurrence in islands 57,58,59,60,61. 62, 64, 65, 66, 67, 68, 69, 70, where we did not find the species. For details 71, 73, 74, 75, 76, 78, 79, 81, 82,84,87,89,90, 92, 93, 94, 95, 96, 98, 99, 101, 102, 103, 104, It West's original paper should be consulted. 105, 106, 107, 108, 109, 110) on all islands except shold be pointed out that I have not revised any on St6ra Dimun. Found in all types of habitats of the previous records, and although 1 have though particularly abundant in the plantations. tried to modernize the nomenclature the syno­ Occurs in Se, Sh, and WN. nyms of West, and other commonly used syno­ NOliophilus aqua/ieus (L.) nyms, have often been entered in brackets. For Found in 3 localities on Streymoy «ocs I and 28 • each species I also state the occurrence in the ne­ on wet grass heath and grazed grassland, respecti­ ighbouring areas viz. the Scottish mainland (Se), vely) and Eysturoy Ooc. 9 on wet grass heath). Shetland (Sh), Iceland (I), and Western Norway Previously recorded in 2 localities (West 1930) (WN) defined as the counties , Horda­ and evidently not common in the Faroes. Occurs in Se, Sh, I, and WN. land, and Sogn og Fjordane. For Scotland I have NOliophilus bigllt/allls (Fabricius). relied on a number of sources, for Shetland ma­ Found in 53 localities Oocs I. 2, 3, 4,5,7,9,10, inly on the list given by Bacchus in Berry & 13, 16, 19, 20, 21, 22, 23, 24, 25, 27, 28, 29, 30, Johnston (I 980), for Iceland on Larsson & Gigja 31,34,38,39,41,42,47,51,52,53,54,57,61, (I 959), Lindroth et al. (I 973), Sigurjonsdottir 63,65,70,73,76,78,79,84,85,92, 94, 95,100,

55 101. 102, 105, 106. 108. Ill) on all islands except Streymoy, and Eysturoy (West 1930). Occurs in on Hestur and Kunoy. Found in all sorts of habi­ Se, r, and WN. tats but highest abundances recorded in fell-field Bembidioll Reniell/alllln Heer ( = B. redlellbaeheri K. sites and on cliffs and shelves among rich vegeta­ Daniell. tion. Frequently found in outfield grassland. Oc­ Previously recorded in 5 localities on Suduroy, curs in Sc, Sh, Land WN. Sandoy. and Streymoy (West 1930). Occurs in Se. Lorieera pilieomis (Fabricius). Bembidirm bmxel/ense Wesmael (= B. I'llpeslre L.) Found in 40 localities (Iocs 1,3,12,14.17,18,27, Previously found in at least 3 localities on Sudu­ 29,33.36,43.44,46,48,49,50, 51. 52. 53, 55, roy and Streymoy (West 1930). Occurs in Sc, Sh, 58.59.60,61. 62. 64, 69. 70, 73, 79, 82, 87.90, r, and WN. 93,94,95, 101, 102. 103, 110) on all islands ex­ Pterosliehll5 ad51riel1l5 Eschscholtz. cept on Skuvoy, St6ra Dimun. Hestur, and N61­ Found in 19 localities (locs 2, 5. 12. 14, 16. 17, 20, soy. Mostly found in infield localities but also in 27.30,33,39.42,45.46.47.49.65,84, 109) on plantations and outfield grassland. Occurs in Sc. Sandoy, Vagar, Streymoy. Eysturoy, Bordoy, Sh, and WN. Kalsoy. and Vidoy. Previously recorded in many Palroblls 5eplelllrilmi5 (Dejean). localities including on Suduroy and N61soy (West Found in 92 localities (Iocs I. 2. 5. 6, 7, 8, 9. 10, 1930). Found in many different habitats but appa­ I I. 12, 13, 14, 16, 18, 19, 20, 2 J, 22, 23, 24, 25, rently favouring outfield grassland and dwarf 26. 27. 28, 29. 30, 31, 32. 33, 34, 35. 36. 38. 39, shrub heaths but also found in infields and on wet 40,41. 42, 43. 44, 45,46.47.48.49.50,51,52. and boggy ground. Occurs in Sc, Sh, r, and WN. 53. 54. 55. 56, 57, 58. 59, 60. 6I. 62,63. 64, 65, Pterosliehll5 niRrila (Paykull). 66.67,68.69.71,73,75.81,82,83,84,86.87, Found in 23 localities (Iocs 4, 5, 12. 13. 14,20,29, 89.90,91. 92, 93. 94,95,96,97.98. 101, 102, 30, 36, 38, 42, 44, 45, 48, 49, 52. 53, 61, 65, 69. 104.105,106, 108. 109. 110, Ill) on all islands 95,96, 106) on Suduroy, Sandoy, Koltur. Vagar. except on Skuvoy. Common in all habitats. Oc­ Streymoy, Eysturoy. Bordoy,,and Vidoy. Previo­ curs in Sc, Sh. I. and WN. usly recorded in several localities including as Palroblls a55imilis Chaudoir. subfossil on Streymoy (West 1930). Found in dif­ Found in 39 localities (Iocs I, 2, 4. 5, 6, 10, 12, 13, ferent habitats though mostly in infield grassland 14. 16,17, 18.20.21,22,23. 27. 28, 30, 32.33, and in moist sites. Occurs in Sc, Sh, Land WN. 34,36.38.39,41. 42.44.49.50.52.65.66,67. Plerosliehlls di/iRell5 (Sturm). 69,87.90,106. 108) on Suduroy. Sandoy, Strey­ Found in 5 localities (tocs 16, 17, 18, 23, 30) on moy, N6lsoy, Vagar. Eysturoy, Bordoy. and Ku­ Streymoy, N6lsoy. and Eysturoy. Previously re­ noy. Found in a variety of habitats with a slight corded on a few localities including on Suduroy preference for dwarf shrub heaths and outfield (West 1930). Found in outfield and infield grass­ grassland. Occurs in Se. Sh, and WN. land as well as in shrub heaths; the highest abun­ Palrobus all'Omfil5 (Strom) ( =P.exeamllls Paykull). dance was recorded on a very wet and boggy Found in 9 localities !locs 3, 8. 26, 35, 51, 68, 69. grassland (Ioc. 23) on N6lsoy. Occurs in Sc, Sh, r, 90, 107) on Sandoy, N6lsoy. Streymoy, Vagar, and WN. Eysturoy. and Bordoy. Previously recorded in se­ Pteroslichll5 51rellllllS (Panzer). verallocalities including on Suduroy (West 1930). Previously found in one locality on Streymoy Found in infield localities and particularly abun­ (West 1930). Occurs in Sc. Sh. and WN. dant in plantations. Occurs in Sc, Sh. I. and WN. Ca/athu5 filscipe5 (Goeze). Trechus oblusus Erichson (= T. qlladrislrialll5 Found in 48 localities (locs 3, 4. 11, 13, 14, 16, 17, Sehrank). 18.26.29.30.32,33. 35. 36, 37. 39,41. 42.45, Found in 80 localities (Iocs I. 2, 3. 4, 5. 7, 9, 10, 46,47,49.50.52.55,56,57,58,59,60,61.62, 11. 12. 14. 16. 17. 18.19.20,21,22,24,26,27, 65.68,73,75,76,81,87,89,90,101,104.105, 28,29,30,31. 32, 33, 34, 35, 36, 38,39,41. 42. 106, 107, Ill) on all islands except on Skuvoy, 43, 44. 45, 46. 48. 49, 50, 51, 52, 53. 54, 55, 57. Koltur, Hestur, and Kalsoy. Common in several 58,59.60.61. 62,63,64,65,66.67.68,69,73, different habitats, especially in infields around set­ 79, 8I. 82. 84, 87. 89, 90. 91, 92, 94. 95, 98, 99, tlements but also found on sand dunes and dwarf 101, 102, 104. 106, 108, 110. Ill) and on all is­ shrub heaths. Occurs in Sc. Sh, and WN. lands. Commonly met with in all habitats. Occurs Calathlls me/alloeephalus (L.). in Sc, Sh, r, and WN. Found in 36 localities (Iocs 3, 11, 14, 16, 18, 21, Treclllls fir/VIIS. Dejean. 22, 25, 26. 29, 32. 33, 34, 36, 39, 42, 43, 46, 47, Found in one locality on Suduroy !loco 58 on a 52,55.58,59,60, 6i, 73, 81,84,94.95,98,99, grass-herb patch near shore). Previously recorded 101, 102, 104, 11 I) on all islands except on Bor­ in about 6 localities including on Vagar, N61soy, doy and Kunoy. Previously regarded as «common and Streymoy (West 1930). Evidently not com­ in all the islands» (West 1930). Found in a variety mon in the Faroes. Occurs in Se, Sh. r, and WN. of habitats but mostly in infield localities, though Bembidioll bipullelaHlIl1 (L.). also reaching a high abundance on fell-fields. Oc­ Previously found in 4 localities on Sandoy, Vagar, curs in Sc, Sh, r, and WN.

56 Ca/athus micropterus Duftschmid. Hydroporus memnonills Nicolai. Previously found as subfossil on Streymoy (West Previously found once on Streymoy (West 1930). 1930). In all probability extinct in the Faroes. Oc­ Occurs in Sc, Sh, and WN. curs in Sc and WN. Hydroporlls sp. O/isthoplIs rotllndatus (Paykull). Found in 2 localities on Suduroy (loc. 53 on grass Previously found in one locality (22 specimens) heath near brook) and Streymoy (loc. 41 in rich on Suduroy (West 1930). Occurs in Sc, Sh, and herbage near shore). WN. ARablls so/ieri Aube ( = A. bipllstll/atlls Sharp). Amara all/ica (Panzer). Found in 2 localities on Suduroy (loc. 62 in rich Found in 23 localities (locs 6, 11, 26, 27, 29, 32, meadow) and Streymoy (loc. 51 in plantation). 35,41, 42,51,52,55,58,59,60,61,62,66,67, Previously stated to be «widely distributed all 68,69,90, 101) on Suduroy, Sandoy, N6lsoy, Va­ over the Faroe and very common» (West 1930). gar, Mykines, Streymoy, and Eysturoy. Appa­ Occurs in Sc, Sh, I, and WN. rently absent in the northern islands, Found exc­ lusively in infields except for one plantation site. Occurs in Sc, Sh, and WN. Family Hydrophilidae Harpa/lIs quadripllnctatlls Dejean. Previously recorded on Suduroy (?) and Sandoy He/op/lIIrus aqllaticlls (L.). but evidently rare or occasional (West 1930), Oc­ Found in 2 localities on St6ra Dimun (Ioc. III in curs in Sc, Sh. and WN, rich grassland) and Svinoy (loc. 8G in grass mea­ Dicheirotrichlls RlIstal'i Crotch ( =D. pllbescens Pay­ dow along ditches). Previously found in 3 locali­ ku\l). ties on Suduroy, Koltur, and Streymoy (West Previously found on N61soy and Mykines (West 1930). Occurs in Se, Sh, and WN. 1930), Occurs in Se, Sh. and WN. He/ophofllS RlIttll/lIS brel'ipa/pis Bedel. Trichocelllls Wicheirotrichlls) COR/wtus (Gyllenhall. Found in 8 localities (locs 3, 27,35,41. 43, 44, 94, Previously found in several localities on Suduroy, 107) on Sandoy Koltur, Streymoy, Eysturoy, and Sandoy, N61soy, Vagar, Mykines, and Streymoy Bordoy. Previously found in 4 places including on and considered rare in the Faroes (West 1930), N61soy (West 1930). Only found in infield locali­ Occurs in Se, Sh. Land WN. ties. Occurs in Sc, Sh, and WN. Trichoce//lls Wicheirotrichlls) p/acidlls (Gyllenhal), He/ophorus fla\'ipes (Fabricius), Found in one locality on Suduroy (loc, 53 on Found in 8 localities (Iocs 33. 35, 53, 56, 79, 95, grass heath near a brook). Previously recorded 97, IIIl on Suduroy, Sandoy, St6ra Dimun, Kol­ once on Suduroy and Streymoy, respectively tur, Hestur, and Svinoy. Apparently not very • (West 1930). Occurs in Sc and WN. common and mainly confined to the southern is­ lands. All individuals were found in grassland (in­ Family Haliplidae and outfields) and in rich vegetation un cliffs and shelves. Occurs in Sc, Sh, and WN. Ha/ip/lls fit/l'lIs Fabricius). Helophorus I'iridico/lis Stephens. Previously found on a few localities on Suduroy, Previously found in several localities on Suduroy, Sandoy, and Eysturoy (West 1930). Occurs in Se, Sandoy, Koltur, and Streymoy, though apparently Sh, I, and Wl'Q". not in the northern islands (West 1930). Occurs in Sc and Sh. Anacaena R/obll/llS (Paykulll. Family Dytiscidae Found in 14 localities locs 20, 23, 33, 49, 52, 53, Hydroporus Rriseostriatlls (De Geed. 56, 64, 65, 81, 89, 93, 95, 110) on Suduroy, San­ Previously found in a few localities on Vagar, doy, N6lsoy, Koltur, Vagar. Streymoy, Eysturoy, Streymoy, Bordoy, and Vidoy (West 1930). Oc­ Bordoy, Kalsoy, Kunoy, and Svinoy. Usually in curs in Sc and Sh. small numbers but in a variety of habitats inclu­ Hydroporus erythrocep/w/lls (L.). ding in- and outfield grassland, shrub heaths, • Previously found several times on Suduroy and plantations and on cliffs and shelves. Occurs in on Vidoy (West 1930). Occurs in Sc, Sh, and WN. Se, Sh, and WN. Hydroporlls palllstris (L.). Ceryon /itora/is (GyllenhaO. Previously found once on Eysturoy and Bordoy, Previously found twice, but in large numbers, on respectively (West 1930). Occurs in Se, Sh, and Suduroy and Sandoy (West 1930). Occurs in Se, WN. Sh, I, and WN. H\'droporus pllbescens (GyllenhaO. CercI'on haemorr/lOida/is (Fabricius) Previuusly found «in almost all the isles .... ap­ Previously recorded «in most of the isles .... oft­ pears to be the commonest species of the genus» en abundant» (West 1930). Occurs in Sc, Sh, and (West 1930), Occurs in Sc, Sh, and WN. WN. Hydroporus niRrita (Fabricius), CercWI/1 marinlls Thomson. Previously found once on Streymoy (West 1930). p'reviously recorded once Suduroy (West 1930). Occurs in Se, Sh, I, and WN. Occurs in Sc, Sh, and WN,

57 MeKasterl1l1f1l bo/elophagllm obscllrllm (Marsham). Acrolrichis sp. Found in 42 localities (Jocs 8,11, 17,26,27,29, Found in 27 localities (Jocs 27, 33,41,43,44,51, 32, 33, 35, 36, 42. 43, 44, 51, 52, 55, 58, 60, 61. 52,56,61,62,66,67,73,80.82,83,85,87,90, 62, 64, 64, 66. 67, 68. 73, 80, 81. 83, 84, 85. 87, 91, 93, 97, 98, 104, 105, 107, 110) on Suduroy, 91. 93, 94, 95, 97, 98,101, 104, 110. 11\) on all Sandoy, Vagar, Hestur, Streymoy, Eysturoy, Bor­ islands except on Skuvoy and Bordoy. Previously doy, Kalsoy, Kunoy, Vidoy, Svinoy, and Fugloy. recorded on «most of the isles» and found as sub­ Found among rich vegetation i.e. infields, planta­ fossil on Streymoy (West 1930). Almost entirely tions, and on cliffs and shelves. restricted to infields, though also found in 3 plan­ tations and 3 outfields sites. Occurs in Se, Sh, and WN. Family Staphylinidae ProleillllS brachyplerus (Fabricius). Found in 3 localities on Hestur (Joc. 98 among Family Silphidae grass and herbs within settlement), Streymoy (Joc. 51 in Larix and Sorblls plantation), and Eysturoy Ca lOps ./iliiKinoslIs Erichson (= C. borealis Kroge­ (Joc. 44 among wasteland vegetation in village). rus). Occurs in Sc and WN. Found in 4 localities on N61soy ([oc. 26 in grass AI1//lObium minuwlIl (Fabricius). meadow with Rumex). Vagar (loc. 11 in waste­ Previously found once on Streymoy (West 1930). land vegetation on sand), Eysturoy (Ioc. 43 on Occurs in Se and WN. rich grassland), and Kalsoy (loc. 91 in grass mea­ Omalium rivu/are (Paykul\). dow). Previously found in 5 localities including 3 Found in 23 localities (Jocs 11, 17, 18. 22. 26, 29, on Streymoy (West 1930). Occurs in Sc, I, and 32, 33, 35, 39, 43, 44, 51, 58,66, 67, 68, 74, 81, WN. 91, 93, 98, 107) on Suduroy, Sandoy, N6lsoy. Catops (Sciodrepa) watsoni (Spence). Hestur, Vagar, Streymoy, Eygturoy, Bordor. Kal­ Previously found once on Mykines (Walker soy, Svinoy, and Fugloy. Previously stated to be 1938). Occurs in Se, Sh, and WN. «distributed in nearly all the isles ... very com­ mom> (West 1930). Found in infield sites and only once in outfield grass heath, once in a plantation, Family Leiodidae and once on cliffs and shelves. Occurs in Sc, Sh, I, Hydnobills punclaws (Sturm). and WN. • Previously recorded once on Suduroy, although Omaliutn excavaWtn Stephens. the specimen (a female) is damaged and the identi­ Found in 12 localities (Jocs 3, 11, 17, 26, 27, 32, fication is somewhat uncertain (West 1930). Oc­ 55,60,68, 87, 91. 98) on Suduroy, Sandoy, N61­ soy Hestur, Vagar, Streymoy, Eysturoy, Kalsoy, curs in Sc and Sh. and Kunoy. Previously found only in two places Hydnobius seplenlriona/is Thomson. including on Koltur. Apparently fairly common Previously found once on sand dunes on Sandoy (West 1930). Occurs in northern Seandinavia. although mostly in small numbers. Only encoun­ tered in infields and mainly close to settlements. Hydnobius sp. Occurs in Se, Sh, I, and WN. Found in 2 localities on SUduroy ([oc. 58. 19 spe­ Oma/ium caeswn Gravenhorst. cimens among grass and herbs on sand near shore Found in one locality on St6ra Dimun (Joc. I1I and loco 64, 2 specimens on heavily grazed grass­ on rich grassland). Previously found in two locali­ land). ties on N61soy and Streymoy (West 1930). Occurs in Se and WN. Oma/ill/l1 /aeviuscll/um Gyllenhal. Family Scydmaenidae Previously found in 3 localities on Streymoy, Eysturoy, and Bordoy (West 1930). Occurs in Se, Slenichnus col/aris (Muller & Kuntze). Sh, I, and WN. Found in 5 localities (Jocs 42, 52, 66, 79, 105) on Omalium riparium Thomson. Suduroy, Vagar, Streymoy, Vidoy, and Svinoy. Previously found in 2 sites on Suduroy and San­ Four of the sites were on outskirts of infields and doy (West 1930). Occurs in Sc, Sh, I, and WN. one in rich vegetation on cliffs and shelves. Pre­ Eucnecosum (A rpedium) brachyplerum (Graven­ viously found once (West 1930). Occurs in Se. Sh, horst). [, and WN. Found in 36 localities (Jocs 1, 2, 5, 7, 9, 12, 14, 19, 21, 22, 24, 27, 31, 33, 34, 45, 47, 52, 53, 63, 65, 74,78,83,84,85,88,89,91,92,94,95,98,102, Family Ptiliidae 106, 108) on all islands except on St6ra Dimun Acrolrichis atomaria De Geer. and Skuvoy. Previously only recorded in very Previously recorded in 3 localities on Suduroy few places and considered rare in the Faroes and Streymoy (West 1930). Occurs in Sc, I, and (West 1930). Although widely distributed the spe­ WN. cies is usually met with in small numbers. Found

58 in all sorts of habitats and nol particularly bound 91. 94) on Suduroy, Sandoy, Koltur, Vagar, Eys­ to infields; relatively often found in outfields grass turoy, and Kalsoy. Previously recorded in 4 places heaths, on fell-fields, and in dwarf shrub heaths. including on Streymoy and N6lsoy. Found in Occurs in Sc, Sh, and WN. many different habitats ranging from fell-fields to Olophrum fuscum (Gravenhorst). rich infield sites. Occurs in Sc. Sh, and WN. Found in 710calitiesOocs 27,83,91. 97,101,107, SlenLlS c1al'icornis (Scopolil. Ill) on St6ra Dimun, Hestur, Mykines, Strey­ Previously found in one locality on Suduroy moy, Bordoy, and Kalsoy. Found exclusively in (West 1930). Occurs in Se. Sh, and WN. infield sites and 175 of the 187 specimens were Slenus bmnnipes Stephens. obtained on the rich grassland site on St6ra Di­ Found in 15 localities Oocs 19, 27, 32, 33. 4 I, 42, mun Ooc. I 11), possibly indicating that it is 44,47. 52, 54, 56. 60, 62. 66, 107) on Suduroy. «autumn» species since the pitfall trapping exten­ Sandoy, Vagar. Streymoy, Eysturoy, Bordoy. and ded into October which was much later than for Vidoy. Previously found in 3 places (West 1930). the other sites. Occurs in Sc, Sh, and WN. Mostly found in infields but a few records from Xylodromus depressus (Gravenhorst). Found in one outfield grass heath. Occurs in Sc, Sh, and WN. locality on Eysturoy (loc. 43 on rich grassland Oxylelus (Ano/yllls) il1l1s/lIs Gravenhorst. near shore). Previously recorded from a few pla­ Previously reported from Suduroy (West 1930). ces including on SUduroy, Sandoy, and Streymoy Occurs in the British Isles and on the Continent of (West 1930). Occurs in Se, Sh, I, and WN. . Xl'lodroll1us concinnus (Marsham). La/hrobillll1 fiilvipenne (Gravenhorst). Found in one locality on N61soy Ooc. 26 on grass Found in 2 localities on Suduroy Ooc. 55 in grass meadow with Rumex). Previously recorded on and herbage) and Eysturoy (loc. 43 in rich grass­ «isles north of Suduroy» though only Streymoy land); both sites near shore. Previously found in 5 specified (West 1930). Occurs in Sc, Sh, I, and places including on Streymoy (West 1930). Oc­ WN. curs in Sc, Sh, I, and WN. Leslel'a longelylrala (Goeze). Found in 34 localities La/hrobium hrunnipes (Fabricius). (locs I, 2,4,5,6,7,9,10,12,16,17,19,22. 24, Found in one locality on Vagar Ooc. lOon rich 26,28,34,35,38,44,50,53,63,64,67,71,85, grass heath). Occurs in Sc, Sh, and WN. 92, 93, 94, 97. 100, 10 I, Ill) on all islands except Xan/holil1l1s linearis (Olivierl. on Kunoy, Vidoy, and Svinoy. Widely distributed Found in one locality on Suduroy (loc. 62 on rich but not very numerous. Found in a variety of ha­ grass-herb meadow). Occurs in Sc, Sh, and WN. bitats mostly in outfield grass heaths. Occurs in O/hius Pllllcllllallls (Goeze). Sc, Sh, I, and WN. Found in 12 localities (locs 8, 22, 27. 28. 34, 53. Leslel'Q 1I10nlicola Kiesenwetter. 55. 59, 61, 67, 98. 11 [) on Suduroy. St6ra Di­ Found in one locality on Streymoy Ooc. 51 in mun, Sandoy, N6lsoy, Vagar Hestur, Streymoy, mixed Larix and Sorbus plantation). Occurs in Sc. and Eysturoy. Previously reported from very few Sh, and WN. places and apparently not very common and not Lesleva heeri Fauvel. found on the northern islands. Almost entirely re­ Found in 3 localities on Suduroy Ooc. 53 on grass stricted to infield sites and cliffs and shelves and heath near brook), N6lsoy (loc. 24 on grass heath only once recorded in outfield grassland. Occurs with Nardus), and Hestur (loc. 97 on rich mea­ in Sc, Sh, and WN. dow around abandoned farmstead). Occurs in Se O/hiLls angusllls (Stephens) (= O. melal/ocephalus and Sh. Gravenhorstl. Micralymma lI1arinull1 (Strom). Found in 60 localities Oocs 3, 8, 9, 11, 14, 16, 18, Found in one locality on Suduroy Ooc. 58 among 19, 20, 21. 22, 23, 24, 25, 26. 27. 28. 29, 30, 33, grass and herbs on sand near shore). Previously 34,36,38,41,42,43,45,46,47,48,49.50,51, found in one place on Sandoy (West 1930). Oc­ 52,55,57,58,59.60,61, 62, 63, 64, 65,66,68, curs in Se, Sh, I, and WN. 73,81,82,83,87.93,97,98,101,102,104,105, Slenlls ill1presslIs German. 108, I1 I) on all islands except on Skuvoy and Found in 27 localities (locs 2, 10, 14, 19, 22, 4 I. Koltur. Apparently one of the most widely distri­ 51,53,56,65,76,83,84,85,87,88,89,90,91, buted and commonest rove beetles. Found in all 92,93.97,98,101. 108, 110, Ill) on Suduroy, sorts of habitats. Occurs in Se. Sh, I, and WN. St6ra Dimun, N6lsoy, Hestur, Vagar, Mykines, O/hiLls myrll1ecopl1ilLls Kiesenwetter. Streymoy, Eysturoy, Bordoy, Kalsoy, Kunoy, and Found in 44 localities Oocs I, 5, 10, 12, 14, 21, 22, Svinoy. Previously only found in 3 localities and 23,24,27.28,33,34,35,36,41,42,44.46,47, considered very local (West 1930). Found in 48,51.52.53,55,57,59,61,63,66,73,78,83, many different habitats from shrub heaths and 84,87,91,93,94,95,98,101,102, 104)on all is­ outfield grass heaths to infields and plantations; lands except on St6ra Dimun and Skuvoy. Previo­ always in small numbers. Occurs in Sc, Sh, I, and usly recorded only in about 10 places (West 1930) WN. but apparently almost as common and widely dis­ Slenus nilidisculus Stephens. tributed as O. angLls/lis. Found in a variety of ha­ Found in 9 localities (locs 7, 10, 19,34, 35, 52, 56, bitats and relatively more often encountered in

59 ..-..

outfield localities than O. angus/us. Occurs in Sc, tions and dwarf shrub heaths. Occurs in Sc, Sh, I, Sh, and WN. and WN. Phi/on//llIs fimetarius (Gravenhorst). Quedius nitipennis (Stephens) (0= Q. allen!lo/us Gyl­ Found in 5 localities Oocs 43, 55, 67, 68, 98) on lenhaD. Suduroy, Hestur, Vagar, and Eysturoy. Previo­ Found in 28 localities Oocs 5, 6, 10, 13, 16, 18, 26, usly found in 6 places including on Sandoy, Nol­ 27,28,29,30; 41,43,45,50,52,53,56,64,65, soy, and Streymoy (West 1930). Only found in in­ 66,67, 87, 89, 92, 105, 107, 109) on Suduroy, field sites, and apparently not very common and N6lsoy, Vagar, Streymoy, Eysturoy, Bordoy, Kal­ absent from the northern islands. Occurs in Sc, soy, Kunoy, and Vidoy. Previously recorded in Sh, and WN. «nearly all the isles» and common (West 1930), Phi/on/hus cepha/o/es (Gravenhorst). Found in all sorts of habitats. Occurs in Sc, Sh, Found in 5 localities Oocs 29, 35, 43, 55, 58) on and WN. Suduroy, Sandoy, Streymoy, and Eysturoy. Pre­ Q!ledi!lS boops (Gravenhorstl. viously only one specimen found on Streymoy Previously found once (2 specimens) on Streymoy (West 1930). Apparently not common and confi­ (West 1930). Occurs in Sc, Sh, I, and WN. ned to infield sites; mostly on the outskirts. Oc­ Quedi!ls boopoides Munster. curs in Sc, Sh, Land WN. Found in 1I localities Oocs 1,2,5,9, 10, 16,21, Phi/on/hus margina/us (Strom). 22, 23, 24,28) on N6lsoy, Vagar, Streymoy, and Found in one locality on Streymoy Ooc. 3 on hea­ Eysturoy. Mostly found in outfield grass heath vely grazed grassland). Previously found in 4 lo­ but also in shrub heath and on cliffs and shelves. calities including on Suduroy and Sandoy (West Occurs in the British Isles and much of Fenno­ 1930). Occurs in Sc, Sh, and Wn. scandia, including WN. Philonthus chalceus Stephens. Quedi!ls fit/vicol/is (Step hens). Previously recorded once on Streymoy (West Previously found on «Suderoy and other parts of 1930). Occurs in Sc, Sh, and WN. the Faroes» (West 1930). OcQurs in Sc, Sh, I, and Phi/ol1/hus poli/us (L.) (0= P. aeneus Rossil. WN. Previously reported once from Streymoy (West Q!ledi!ls sp. 1930) but seems to require confirmation. Occurs Found in 13 localities Oocs I, 19, 22, 24, 34, 42, in Sc, Sh, I, and WN. 44, 53, 63, 78, 97, 102, Ill) on Suduroy, St6ra Creophiltls maxil/os!ls (L.). Dimun, Sandoy, N61soy Hestur, Mykines, Strey­ Previously found on Suduroy and Sandoy (West moy, Eysturoy, and Svinoy. Mostly on grllSs 1930). Occurs in Se, Sh, I, and WN, heaths and in infields. Quedills cllrtipennis Bernhauer (0= Q. fuliginoslls Tachyporus a/riceps Stephens. Gravenhorst), Previously found on Suduroy (West 1930). Oc­ Found in 33 localities Oocs 5, 6, 8, 10, 14,22,25, curs in Sc, Sh, and WN. 27,28,29,30,34,41,42,45,46,47,48,50,51, Tachinus pa/lipes Gravenhorst. 61,62,65,66,68,69,76,84,91,92,106,107, Previously reported from N6lsoy and the north­ 1I \} on Suduroy, St6ra Dimun, Sandoy, N6lsoy, ern islands but a confirmation is desirable (West Vagar, Streynoy, Eysturoy, Bordoy, Kalsoy, Vi­ 1930). Occurs in Sc and Fennescandia. doy, and Svinoy. Since cllr/ipennis andfitligillosus Tachinus signa/us Gravenhorst (0= T. rufipes De Ge­ are considered synonymous it is previously found er). in almost all the isles (West 1930). Found in seve­ Found in 47 localities (Iocs 3,11,14,25,26,27, ral different habitats from fell-fields and shrub 29,32,33,35,36,41,43,44,45,48,49,50,51, heaths to in- and outfield grasslands. Occurs in Se, 52,55,58,59,60,61,62,64,66,67,73,81,82, Sh, and WN. 87,90, 91, 93, 94,95, 98, 101, 103, 104, 107, Quedi!ls mesome/illus (Marsham). 110, Ill) on all islands except on Skuvoy. Previo­ Found in IS localities Oocs 8, 11, 18, 26, 41, 43, usly reported from most of the islands but consi­ 55, 58, 61, 64,68, 73, 81, 98, 99) on Suduroy, dered not to be numerous (West 1930). In our Skuvoy, N6lsoy, Hestur, Vagar, Streymoy, Eys­ material by far the most numerous member of the turoy, Svinoy, and Fugloy. Previously recorded in family (68 % of all rove beetles). Predominantly several places «in the islands north of Suduroy» found in infield localities and only occasionally in without further details (West 1930). Almost exc­ outfields (grass heaths and shrub heaths) and lusively found in infield localities. Occurs in Sc, plantations. Occurs in Sc, Sh, and WN. Sh, I, and WN. Tachinus lalico/lis Gravenhorst. Quedius umbrinus Erichson. Found in 28 localities Found in 3 localities on Suduroy (Iocs 59, 60, 62; Oocs 2,3,4,6,9, 14,27,28,29,32,33,48,51, all rich infield meadows). Previously recorded 52, 53, 58, 59, 60, 62, 73, 76, 84, 88, 91,92, 107, twice on Suduroy (West 1930), Occurs in Sc, Sh, 110, II \} on Suduroy, St6ra Dimun, Sandoy, Va­ and WN. gar, Streymoy, Eysturoy, Bordoy, Kalsoy, Kunoy, Hypocyp/us laeviusculus (Mannerheim). Svinoy, and Fugloy. Previously found in several Found in 5 localities Oocs 17,21,36,52,95) on localities including on Vidoy (West 1930). Mostly Suduroy, Sandoy, N6lsoy, Koltur, and Eysturoy. found in infields and grassland but also in planta­ Previously found once (1 specimen) on Sandoy,

60 (West 1930). Confined to infields and grass but the determination is not absolutely certain heaths. Occurs in Se, Sh, and WN. (West 1930) and a confirmation needed. Occurs in Mvl/aena brevicornis (Matthews). the British Isles, Sh, and WN. Found in 2 localities on Kalsoy (Ioc. 85 in a coni­ Atheta (Bessobia) excel/ens (Kraatz). ferous plantation and loc. 93 on a grass meadow). Found in 19 localities (Iocs 5, 9, 13, 16, 21,22,34, Occurs in Se, Sh, and WN. 39,42,54,61,63,64,82,91,104,105,108, I1 I) Amischa (A theta) ana/is Gravenhorst. on Suduroy, St6ra Dimun, Sandoy, N6lsoy, Va­ Found in 22 localities (Iocs 3, 14, 17, 32, 33, 36, gar, Streymoy, Eysturoy, Bordoy, Kalsoy, and Vi­ 42, 43, 45, 46, 50, 51, 52, 56, 59, 60, 61, 65, 66, doy. Previously recorded from 6 places (West 74, 83, 92) on Suduroy, Sandoy, Vagar, Strey­ 1930). One of the most widely distributed species moy, Eysmroy, Bordoy, Kalsoy, Vidoy, and Fug­ of the genus and found in many different habitats; loy. Previously found in 4 places (West 1930). Fo­ mostly in grass heaths. Occurs in Se, Sh, I, and und in many different habitats though mostly on WN. the outskirts of infields. Usually in small numbers Atheta (Traul/1oecia) angusticol/is Thomson. except on the rich grass meadow at loc. 42 (69 of Previously found once (or twice) on Vagar (West the 138 specimens collected). Occurs in Se, Sh, I, 1930). Occurs in Se and WN. and WN. Atheta (Hygroecia) pa/ustris Kiesenwetter. Sipalia circel/aris (Gravenhorst). Previously found once on Suduroy (West 1930). Found in 12 localities (Iocs 3, 17,21,26, 33, 42, Occurs in Se and WN. 43, 52, 55, 74, 83, 84) on Suduroy, Sandoy, N61­ Atheta (Microdota) amiCl//a (Stephens). soy, Streymoy, Eysturoy, Kalsoy, and Fugloy. Found in one locality on Vagar (Ioc. II among Previously found in two places (West 1930). wasteland vegetation on sandy ground). Previo­ Mostly in infields habitats and only twice on grass usly found in about 6 places including on Sudu­ heaths. Neither very common nor numerous. Oc­ roy, Sandoy. Streymoy, Koltur, and Svinoy. Ap­ curs in Se, Sh, I, and WN. parently not common. Occurs in Se, Sh, I, and Mycetoporus rufescens Stephens. WN. Found in one locality on N61soy (Ioc. 22 on cliffs Atheta (s.str.) nigricornis Thomson. and shelves). Occurs on the British Isles and in Found in one locality on Eysturoy (Ioc. 43 on rich WN. grassland near shore). Previously recorded in one Auta/ia puncticollis Sharp. locality on Streymoy (West 1930). Occurs in Se Found in one locality on Streymoy (Ioc. 28 on and WN. heavily grazed grassland). Previously recorded in Atheta (s.str.) coriaria (Kraatz). several places including on Suduroy, Sandoy, Va­ Previously found once on Suduroy (West 1930). • gar, and Vidoy (West 1930). Occurs in Se, Sh, I, Occurs in Se and Seandinavia. and WN. Atheta crassicornis (Fabricius). Atheta (A/oconata) gregaria (Erichson). Previously found once on Suduroy (West 1930). Found in 7 localities (Iocs 3, 8, 26, 29, 32, 43, 60) Occurs in Se and WN. on Suduroy, Sandoy, N6lsoy, Streymoy, and Eys­ Atheta (s.str.) trinotata (Kraatz). turoy. Previously found in 4 localities including Found in 2 localities on Sandoy (Ioc. 32 in rich on Koltur and~ugloy (West 1930). Found only in herbage) and N61soy (loc. 26, 14 specimen on infield localities. Occurs in Se, Sh, and WN. grass meadow with Rumex). Previously found in :4.theta (Hygroecia) me/anocera (Thomson). Found in 4 sites including on Koltur and Streymoy. Occurs 4 localities on Suduroy (Ioc. 53 on grass heath in Se, Sh, I, and WN. near brook and loc. 55 among grass and herbage), Atheta (s.str.) gral/1inico/a (Gravenhorst). Sandoy (loc. 35 among grass and herbage on Found in 5 localities (locs 9, 19,26,27,31) on sandy ground) and Vagar (loc. lOon rich grass N6lsoy, Streymoy, and Eysturoy. Previously re­ heath). Previously recorded in about 6 places inc­ corded once on Streymoy (West 1930). Found in luding on Koltur and Streymoy (West 1930). Oc­ infields, grass heaths and on fell-field. Occurs in curs in Se, Sh, I, and WN. Se, Sh, I, and WN. Atheta (Hygroecia) e/ongatu/a (Gravenhorst). Atheta (Thinobaena) vestita (Gravenhorst). Previously found once (I specimen) on Suduroy Found in 3 localities on Suduroy (loc. 64 on hea­ (West 1930). Occurs in Se, Sh, and WN. vily grazed grassland), Sanooy (loc. 37 on sand Atheta (Dimetrota) is/andica (Kraatz). dunes), and Eysturoy (loc. 44 among wasteland Found in 13 localities (Iocs 9, 12, 22, 23, 24, 27, vegetation near shore). Previously found in 4 pla­ 34, 53, 58, 63, 71, 77, 93) on Suduroy, Sandoy, ces including on Koltur and Streymoy. Occurs in N6lsoy, Vagar, Streymoy, Eysturoy, Kalsoy, Svi­ Se, Sh, I, and WN. noy, and Fugloy. Previously found in two locali­ Atheta (Dimetrota) atramentaria (Gyllenhail. ties on Streymoy (West 1930). Found in a variety Found in 18 localities (Iocs 3, 14, 18, 22, 28, 34, of habitats ranging from fell-fields to infields. Oc­ 42,43,47,50,55,57,61,66,68,82,106,lll)on curs in Se, Sh, I, and northern Scandinavia. Suduroy, St6ra Dimun, Sandoy, N6lsoy, Vagar, Atheta (Hygroecia) debilis (Erichson). Streymoy, Eysturoy, Bordoy, Kalsoy, and Vidoy. Previously reported once (I female) on Eysturoy Previously found in a few places on five of the is­

61 lands (West 1930). Apparently one of the most Sandoy, and Streymoy (West 1930). Occurs in Sc, widely distributed and commonest members of Sh, and WN. the genus. Predominantly found on the outskirts Aleochara obscurella Gravenhorst. of infields but occurs in a variety of other habitats Previously recorded once on Sandoy (West 1930). as well. Occurs in Sc, Sh, I, and WN. Occurs in Sc, Sh, and WN. Atlu'la Is. str,) setigera Sharp. Previously recorded once on Suduroy (West 1930). Occurs in Se and WN. Family Cantharidae Athela (Acrotona) parvu/a Mannerheim. Malthodes atomus (?) Thomson. Previously recorded once on Suduroy (West Previously found in 3 places on Suduroy and 1930). Occurs in Sc and WN. Streymoy, but since only females were collected At17ela lDatol/licra) celata (Erichson) (= A. arenico/a the identity of the species (but not genus) is uncer­ Thomson). tain (West 1930). Occurs in Se, Sh, I, and WN, Found in 2 localities on Suduroy (Joc. 55 among grass and herbage near shore) and Sandoy (Joc. 37 on sand dunes). Previously found in some 5 pla­ Family Elateridae ces including on Koltur, Vagar, Streymoy, and Svinoy (West 1930). Occurs in Se, Sh, I, and WN. Hvpolithus (Crypto17ypnus) riparills (Fabricius). At17eta IAcrotona).fiIflRi (Gravenhorst). Found in 51 localities (Jocs 3, 5, 6, 9, 10, 16, 17, Found in 4 localities on Suduroy (Joc. 59 in rich 18, 19, 20, 21, 22, 27, 28, 29,32, 36,39,42,43, grass meadow and loc, 60 among rich herbage), 45,46,47,50,51,52,56,57,58,59,60,61,64, Streymoy (Joc. 51 in mixed Larix and Sorbus 65,68, 73, 74, 75, 76, 84, 87, 89, 95, 99, 101, plantation), and Vidoy (Joc. 104 in rich grass mea­ 102, 104, 105, 106, 108, Ill) on all islands except dow near shore). Previously found once on Sudu­ on Hestur. Previously stated to be widely distribu­ roy (West 1930). Occurs in Sc, Sh, I, and WN. ted and very common (West) 930). Found in infi­ A/17ela (s.str.Jhanmodi Williams. Found in one loca­ elds, grass heaths as well as plantations and shrub lity on Suduroy (loc. 60 among rich herbage). Oc­ heaths. Occurs in Se, Sh, l. and WN. curs in Sc, Sh, and WN. At17e/a (HvRroecia) Il/ridip(,/{l1is Mannerheim. Found in 2 localities, on Hestur (Joc. 98 in grass Family Helodidae and herbs around abandoned farm) and Suduroy E/odes (Helodes) lIlinll/a (L.). (Joc. 55 among grass and herbage). Occurs in Se, Found in 4 localities on Sandoy (loc. 32 in a rich I, and WN. herb meadow near waterl, Streymoy (Joc. I on Atheta sp. wet grass heath and loc. 29 on grass-herb mea­ Found in 12 localities (locs 9, 35, 41, 49, 52, 53, dow), and Kalsoy (loc. 91 on grass meadow). Pre­ 56,61, 67, 68, ~3, i 10) on Suduroy, Sandoy, Va­ viously found in at least 6 localities including on gar, Streymoy, Eysturoy, Bordoy, and Kalsoy, Suduroy and on Hestur (West 1930). Occurs in Oea/ea picata (Stephens). Se, Sh, and WN. Found in 12 localities (locs 11,17,29,35,41, 55, 67,68, 82, 90, 91,99) on Suduroy, Skuvoy, San­ doy, Vagar, Streymoy, Eysturoy, and Kalsoy. Family Previously found in 5 places including on Bordoy (West 1930), All records from infields localities. Dermestes lardarius (LJ Occurs in Sc, Sh, I, and WN, Previously found once on Streymoy (West 1930). Oxypoda 17aemorr17oa (Mannerheim). Occurs in Sc, I, and WN, Found in 3 localities on Suduroy (Joc. 58 in a pellio (L.). patch of grass and herbs on sandy ground near Previously found once on Suduroy (West 1930). shore), Sandoy (Joc. 37 on sand dunes), and Vagar Occurs in Se and WN. (loc. 11 among wasteland vegetation on sand sho­ Anthrenus verbasci (L.), re). Previously known from a sandy area on San­ Previously found once on Sandoy (West 1930). doy and from N61soy (West 1930). Occurs in Sc, Occurs in the British Isles. Sh, l. and WN. Aleochara sparsa Heer. Found in 2 localities on Sandoy (Joc, 35 among Family Byrrhidae rich grass and herbage) and Vagar (Joc. 68 in rich Simp/ocaria semistriata (Fabricius), vegetation near waterl. Previously recorded from Found in 14 localities (Iocs 3, 11, 20, 29, 52, 59, 5 places including on Suduroy, Koltur, and Strey­ 60,61, 62, 73, 83, 94, 97, 11 I) on Suduroy, St6ra moy. Appear not to be common and even absent Dimun, Koltur, Hestur, Vagar, Streymoy, Eystu­ from the northern islands. Occurs in Sc, Sh, I, and roy, Kalsoy, and Fugloy, Previously found in 6 WN. places including on Sandoy (West 1930). Almost A/eochara lanllginosa Gravenhorst. exclusively found in infield sites and only one re­ Previously known from 6 localities on Suduroy, cord from grass heath. Occurs in Se, Sh, and WN. 62 Cytilus sericeus (Forster). in almost all the isles» (West 1930). Only found in Previously recorded in 2 localities on Suduroy infield sites and usually in small numbers but 68 and Streymoy (West 1930). Occurs in Sc, Sh, I, of 77 specimens were obtained from the Rumex­ and WN. rich meadow on N6lsoy (loc. 26). Occurs in Se, L Byrrhus fasciatus (Forsted. and WN. Found in 3 localities on Streymoy (loc. 31 on a fell-field on 600 m a.s.U and Eysteroy (loc. 9 on wet and patchy grass heath and loco 19. on grass­ Family Coccinellidae land). Previously found once on Suduroy (West Coccinel/a seplempUnClala (L.). 1930). Occurs in Sh, Sc, I, and WN. Previously encountered once on Streymoy (West 1930). Occurs in Sc. Land WN. Family Atomaria apicalis Erichson. Family Anobiidae Found in 2 localities on Sandoy (loc. 32 one indi­ vidual among rich herbage) and N6lsoy Hoc. 26, Anobillll1 pllllctalllm (De Geed (= A. slriatum Olivi­ 29 individuals in grass meadow with RUl11ex). er). Previously found in 5 places including on Sudu­ Found in one locality on Kalsoy (loc. 92 on grass roy, Koltur, Streymoy, and Eysturoy. Occurs in heath). Previously recorded once on Suduroy Se, Sh, I, and WN. (West 1930). Occurs in Sc, Sh, T, and WN. Atol11aria niKripennis (Paykull). Ernobius mol/is (L.). Previously recorded once on Streymoy (West Previously found in 5 places on Suduroy, Strey­ 1930). Occurs in Sc, I, and WN. moy, and Bordoy (West 1930). Occurs in Sc. T, AtOl11a ria rt(ficornis (Marsham). and WN. Previously found twice on Sandoy and Streymoy Sitodrepa panicea (L.). (West 1930). Occurs in Se, Sh, and WN. Previously found once on board a vessel from Atomaria sp. Streymoy to Suduroy (West 1930). Occurs in Sc. Found in 3 localities on Streymoy (loc. 41), Eystu­ T, and WN. roy (loc. 54). and Kunoy (Ioc. 87). CryptophaKus scanicus (L.). Found in 7 localities (locs 11, 26, 32, 67, 98, 101, Family Ptinidae 110)on Sandoy, N6lsoy, Hestur. Vigar, Mykines, Niptus unicolor Piller. and Streymoy. Previously regarded as occuring in Previously found in 3 localities on Suduroy, Vi­ nearly all the isles and to be very common (West gar, and Streymoy (West 1930). Occurs in Sc, I, 1930). In present study found only in infield sites and WN. and in one plantation and usually in small num­ PlillUS filr (L.). bers (total 247 specimens) except in the Ru­ Previously recorded twice on Suduroy and Strey­ mex-dominated grass meadow on N61soy Hoc. moy (West 1930). Occurs in Sc, I, and WN. 26, 239 specimens). Occurs in Sc, Sh, T, and WN. PlillUS tectus Boieldieu. CryptophaKus sa!linalus Sturm. Previously recorded at least twice on Streymoy Previously found once on Suduroy (West 1930). (West 1930). Occurs in Sc, Sh, T, and WN. Occurs in Se. Sh, and WN. CryptophaKus dell/alUS (Herbst). Previously found twice on Suduroy and Strey­ moy (West 1930). Occurs in Sc, Sh, and Scandina­ Family Tenebrionidae via. Triboliul11 confilsull1 Duval. Previously recorded distinKuendus (Sturm). once (I specimen) on Streymoy (West 1930). Oc­ Found in one locality on Kalsoy (loc. 83 in grass curs in Sc, I, and WN. meadow). Previously recorded once on Streymoy (West 1930). Occurs in Se, T, and WN. CryplophaKus pilosus Gyllenhal. Found in one locality on N6lsoy (loc. 26 grass Family Scarabaeidae meadow with plenty of Rumex; 62 individuals; Aphodius aler (De Geer). cf. Atomaria apicalis and CryptophaKus scanicus). Previously found once (in large numbers) on Su­ Occurs in Sc, Sh, I, and WN. duroy (West 1930). Occurs in Se. Sh, and WN. Aphodius foelidus (Fabricius). Previously found once on Suduroy (West 1930). Family Lathridiidae Occurs in Sc and WN. Lathridius (Enicl11l1s) milllllUS (L.). Aphodius lapponul11 Gyllenhal. Found in 6 localities (locs 26, 32, 41, 43, 52, 94) Previously found in about 4 places on Suduroy, on Suduroy, Sandoy, N6lsoy, Koltur, Streymoy, Streymoy and Eysturoy (West 1930). Occurs in and Eysturoy. Previously stated to be «distributed Se, Sh, I, and WN.

63 Family Cerambycidae Oliorrhynehlls singularis (L.) Found in 2 localities on Vagar Ooc. 68 among Graeilia lI1illllla (Fabricius). Previously recorded once on Suduroy (West wasteland vegetation in village) and Streymoy 1930). Occurs in the British Isles. Ooc. 51 in mixed Larix and Sorbus plantation). Occurs in Se, Sh, and WN. Callidilll11 violaeelll11 (L). Previously found in 3 localities (West 1930). Oc­ Barynotus sqllamosus German. curs in Se, Sh, Land WN. Found in 14 localities (Iocs 11, 18, 27, 33, 36, 42, 4.5,52,59,67,75,81,87,91) on Suduroy, San­ doy, Vagar, Streymoy, Eysturoy, Kalsoy, Kunoy, Family Chrysomelidae Vidoy, and Svinoy. Previously found in 4 places (West (930). All but one record from infield loca­ Chrysolilla IChrysol11ela) slaphylea (L.l. lities. Occurs in Sc, Sh, I, and WN. Found in 9 localities Oocs 6, 9, 21, 27, 28, 47, 65. Tropiphort/s OblUSllS (BonsdorfO. 66, 87) on N6lsoy, Vigar, Streymoy, Eysturoy, Found in 9 localities Oocs 7, 19,27,33,36,42,46, Kunoy, and Vidoy. Previously found in at least 5 94,101) on Koltur, Mykines, Sandoy, Streymoy, places including on Suduroy, Sandoy, and Bordoy Eysturoy, and Vidoy. Previously found in 3 loca­ (West 1930). Found in infields and outfield grass lities including on Suduroy and considered to be heaths and once on dwarf shrub heath. Also re­ restricted to this island and the southem parts 0 f corded on Midalfelli (550-600 m a.s.l.) on Eys­ Streymoy (West 1930). Although it clearly has a turoy in June 1977 (colI. T. Solh0Y). Occurs in Se, much wider distribution in the Faroes it is only Sh, I, and WN. met with in small numbers and mostly in infield sites. Occurs in Se, Sh, Land WN. Pissodes pilli (L.). Family Curculionidae Previously found once (I specimen) on Suduroy Oliorrhl'lIehlls areliellS (Fabricius). (West 1930). Occurs in Se, I~ and WN. Fou'nd in 33 localities Oocs 7,9, 11, 18, 25, 30, 35, Micrelus erieae (Gyllenhal). 36,37,39,45,46,47,56,58,59,64,65,67,70, Found in 2 localities on Vagar Ooc. 13 on Calluna 71,73,76,78,83,93,95,96,97,99. 101, 104, heath) and Svinoy Ooc. 76 among tall Calluna). Ill) on all islands except on Bordoy and Kunoy. Occurs in Sc, Sh, and WN. Previously stated to occur in almost all the isles Rhinonells pericarphls (L.). without further details (West 1930). Found in Found in 4 localities on Sandoy Oocs 32 and 36\)n most sorts of habitats ranging from sand dunes, rich grass-herb meadows), Streymoy Ooc. 42 on shrubs heaths to rich infield sites. Occurs in Se, grass herb meadow), and Eysturoy Ooc. 43 in rich Sh, I, and WN. grassland). All sites near water. Occurs in Se, Sh, Oliorrhvllclllls dllbills Str6m (= /lOdoslIs allell,). I, and WN. Found in 17 localities Oocs 5, 6, 21. 22, 28, 36, 51, Rhinoneus ineollspeelus Herbst (= R. gramineus Fa­ 55.71,76,77,84.85,88,97,107, 110)on Sudu­ bricius). roy Sandoy, N6lsoy, Hestur, Streymoy, Eysturoy, Previously found once on Streymoy (West 1930). Bordoy, Kalsoy, Kunoy, Svinoy, and Fugloy. Pre­ Occurs in Se and Seandinavia. viously recorded only from 3 places and assumed NOlaris aelhiops (Fabricius). to be rare (West 1930). Found in all habitats ex­ Previously found once on Sandoy (West 1930). cept wetlands and sand dunes. Occurs in Sc, Sh, I, Occurs in Se and Seandinavia. and WN. Silolla pllnelieollis Stephens). OliorrhYllehus mgi/rolls Gyllenhal. Found in one locality on Streymoy Ooc. 3 on hea­ Previously reported once from Bordoy but ente­ vily grazed grassland). Occurs in Sc and Sh. red on the Faroese list with doubts (West 1930). Ceulorrhynehus eolltraellls (Marsham). Lindroth (1931 :231) assumed the record to be a Found in 3 localities on Suduroy Ooc. 61 on rich confusion with O. dubius. The species should the­ grass meadow), Sandoy Ooc. 34 on wet fell-field refore possibly be deleated from the list; at least a ground), and Vagar Ooc. 66 among grass and confirmation is needed. Occurs in Sc, I, and WN. herbs). Previously recorded twice including on Oliorrhynehlls alroaplerus (De Geerl. Streymoy (West (930). Occurs in Sc, Sh. I, and Previously found once on sand dunes on Sandoy WN. (West 1930). Occurs in Se, Sh, and WN. Apion aelhiops Herbst. Oliorrhynehlls rllgososlriatus (Goeze) (= O. seabro­ Previously found twice on Sandoy and Koltur sus Marsham). (West 1930). Occurs in Sc and Sh. Previously recorded once from the Faroes with­ Apion haemalodes Kirby (= A. frumelllarium (Pay­ out any details as to locality except that it occur­ kull) and West's (1930) A. eruenlalllm Walton). red as a pest in a nursery (Larsson & Gigja Found in 5 localities Oocs 32, 33, 59.61, 97) on 1959: 194). This species does not occur in any of Suduroy, Sandoy, and Hestur. Previously found the neighbouring areas except possibly once in in about 7 places including on Koltur and Strey­ Iceland (Larsson & Gigja Op.eft.}, Occurs in the moy (West (930). Apparently not very common British Isles and in Continental Europe. and restricted to the southern parts of the island

64 group. In the present study only found in infields species found in Shetland (340 species) and (4 on the outskirts and one close to settlements). about 80 % ofthe number in Iceland (214 speci­ The «species» occurs in Sc, Sh, 1, and WN. es). A comparison between the Faroes and Shet­ land seems particularly relevant since the two is­ DISCUSSION land groups are of almost the same size (I,400 km2). There are probably several reasons for the Comparison with neighbouring areas great difference in species richness, one being The coleopterous fauna of the Faroes is entirely that Shetland is situated much closer to a main­ West European and effectively all its species oc­ land species pool which presumably favours co­ cur in the British Isles and/or in the western lonization of new species. Iceland is about 75 parts of Scandinavia; areas from which the Far­ times larger in size but situated more distantly oese beetles are traditionally thought to have im­ from mainland areas and further north. The tax­ migrated (West 1930). The number of species is, onomic composition (by family) of the coleopte­ of course, much lower than that found on the rous faunas of Shetland, Faroes, and Iceland is British mainly or Western Norway. At present very similar (Tab. 2). Within all three areas rove at least 168 identified species have been recorded beetles, ground beetles, and weevils dominate in the Faroes. This is about half the number of (in descending order) in terms of species rich-

Table 2. Comparing the taxonomic composition (number of species in each family) of the Coleoptera of Shet­ land, Faroes, and Iceland.

Family Shetland' Faroes2 Iceland) Carabidae 55(16.2%) 26(15.4%) 22 (10.3%) Haliplidae 3 I I Dytischidae 27 ( 7.9%) 7( 4.1 %) 4( 1.4%) Hydrophilidae 15 ( 4.4%) 9( 5.3 %) 3( 1.4%) Silphidae 3 2 1 Leiodidae 6 1 Scydmaenidae I I 1 Ptiliidae 4 I I Staphylinidae 12807.7%) 73(43.2%) 62 (29.0 %) Cantharidae I : 2 Elateridae 4 I 2 Helodidae I Dermestidae 3 2 Byrrhidae 4 3 2 Cryptophagidae 16 ( 4.7%) 8( 4.7%) 13 6.1 %) Lathridiidae 7 I 5 Coccinellidae 2 I 8 Anobiidae I 3 3 Ptinidae 4 3 Tenebrionidae I 9 Scarabaeidae 9 3 3 Cerambycidae 2 17 Chrysomelidae 7 ( 2.1 %) I ( 0.6%) 4( 1.9%) Curculionidae 31( 9.1%) 16( 9.5%) 18 ( 8.4 %) Others4 12( 3.5%) 18( 8.4%) Total 340 168 214

According to Bacchus' list in Berry & Johnston (1980). Omitting 5 doubtful species (see introduction to the synopsis of species) the apparently extinct Calathlls lIIi­ cropterlls, species only identified to genus, and regarding QlIedills clIrtipennis and Q. fllligilloslIs as syno­ nyms. ) According to Larsson & Gigja (1959), Lindroth et al. (1973), Sigurj6nsd6ttir (1974). and Bengtson et al. (un­ publ.). 4 For Shetland the families Gyrinidae, Pselaphidae, Geotrupidae, Clambidae, Scirtidae, Dryopidae, and Niti­ dulidae and for Iceland Pselaphidae, Nitidulidae, Ostomidae, Monotomidae, Cucujidae, Mycetophagidae, Colydiidae. Endomychidae, Cleridae, Lyctidae, Anthicidae, Bruchidae, Brenthidae, and Ipidae.

65 ness, but for most taxa the number of species into any lengthy discussion West (I 930) conclu­ usually descreases from south to north. The ded that «the greater part of the present coleop­ high proportion of carnivorous species relative terous fauna of the Faroes have had conditions to herbivores is characteristic for many island for surviving the glacial period» referring to the coleopterous faunas (e.g. Becker 1975). Shet­ latest glacial period. Currently there is no strong land has 7 and Iceland 14 families not found in evidence for faunal survival in glacial refugia in the Faroes. At least in the case of Iceland this the Faroes. Assuming that the entire fauna is of may be explained by a higher habitat diver­ post-glacial origin leaves us with the problem of sity and a far more intensive collecting, especi­ identifying routes of immigration. The present ally in and around habitation where accidental distribution of the species involved does not give introductions are most likely to be discovered. us any definite answers. Western Norway and Tab. 3 gives the number of species of Faroese Scotland are the nearest main species pools from beetles nol recorded in the neighbouring areas. which the Faroese beetles could have emmigra­ Only 4 and 17 species are not found in Scotland ted. Only 5 of the 17 Faroese species not found and Western Norway, respectively i.e. 97.6 % of in Western Norway are also absent from Nor­ the Faroese species occur in Scotland and way as a whole (viz. Bernbidion geniculaturn. 89.9 % in Western Norway. Of the Faroese spe­ Lesteva heeri. Oxytelus inustus. Anthrenus ver­ cies 78.7 % have been found in Shetland but basci. and Gracilia minuta; the two latter are, only 51. 5% in Iceland. Thus, proceeding from however, introduced in Scandinavia), and may Shetland to the Faroese there is a marked drop therefore have reached the Faroes from the Bri­ in species richness but in terms of species com­ tish Isles. Three Faroese species not recorded tn position the two faunas have a great affinity. Scotland (viz. Oxvtelus inustus. Gracilia rninuta. Comparing the Faroes with Iceland shows that and Otiorrhynch~s rugosostri&tus) occur further the number of species is somewhat higher in the south on the British Isles, whereas a fourth spe­ latter area, but only about half the number of cies, Hydnobius septentrionalis appears to be ab­ Faroese species have extended their ranges to in­ sent. Thus, the vast majority of Faroese beetles clude Iceland. As to the origin of the Faroese may have arrived from either Scotland or Scan­ coleopterous fauna I will in this context not go dinavia, or both.

Table 3. Number of Faroese Coleoptera /lot recorded in Scotland (Sc), Shetland (Sh), Iceland m. Western Nor­ way (WN), and in combinations of these neighbouring areas, assumed to be the most likely sources from which the Faroese beetles originated.

Family Number of Faroese beetles /lol found in (N) Sh WN Sh + I Sh + [ + WN I + WN Sc + Sh + WN Sc + Sh + [ + Wr-.;

Carabidae (I 2) 1 9 Dytischidae ( 5) - 4 Hydrophilidae ( 8) - 7 Silphidae ( 2) I I Leiodidae ( 2) Ptiliidae (1) Staphylinidae (40) 1 25 10 Scarabaeidae ( 2) - I I Helodidae ( 1) - I Dermestidae ( 3) Byrrhidae ( I) - 1 Cryptophagidae ( 5) L 2 Coccinellidae ( 1) 1 Anobiidae ( 2) 2 2 Ptinidae ( 2) 2 Tenebrionidae ( 1) I Cerambycidae ( I) Curculionidae ( 9) 1 3 2 2 Total (98) 14 54 13 4 7 3

66 Distribution within the Faroes ted since: (a) the number of infield localities was About one-tenth (16) of the Faroese beetle spe­ large and (b) the number of niches and amount cies have a wide distribution in the Faroes and of organic matter is presumably much larger in occur in at least 13 of the 18 islands (viz. Nebria infields than in other habitats. The grass heaths, gyllenhali, Nebria salina, Notiophilus biguttatus, represented by about the same number of sites Loricera pilicornis, Patrobus septentrionis, Tre­ as any of the two infield categories, supported a chus obtusus, Calathus !uscipes, Calathus mela­ total of 58 species which is about 65-70% of nocephalus, Megasternum obscurum, Hypolithus the number found in the infields. The lower 1 riparius, Otiorrhynchus arcticus, Eucnecosum number of staphylinid beetles accounted for brachypterwn, Lesteva longelytrata, Othius ang­ much of the difference between infields and ustus, Othius myrmecophilus, and Tachinus sig­ grass heaths, presumably because the latter have natusJ. Another four species (viz. Hydroporus less foerna. The proportion of carabid beetles pubescens, Agabus solieri, Cercyon haemorrhoi­ was higher and the proportions of cryptopha­ dall's. and Cryptophagus scanicus) are reported gids and curculionids lower in the grass heaths to be widely distributed in almost all the islands than in the infields. The number of species was about the same (33-34) in dwarf shrub heaths (West 1930). Presumably the numbers of species crevice~ occurring in about two-thirds (i.e. 12) of the is­ plantations, and on cliffs shelves and in lands may be in order of 40 since, in addition to but the proportion of carabids decreased and the species mentioned above, 11 species have staphylinids increased successively (Tab. 4). The been found in 10 -12 islands and 20 species in lowest numbers of species were found in the 7- 9 islands all 11 of the latter are staphylinid mountain sites (23), on lowland bogs (I 3), and beetles, a taxa which is probably less well on sand dunes (6). However, it should be kept in known than many others. A large proportion of mind that for these habitats only relatively few the species appear to be rare or accidental; about localities were surveyed. The proportion ofcara­ 48 (28 %) have only been recorded once and 17 bid beetles varied between 68 % (lowland bogs) (IO %) twice. and 35 % (shrubs heaths) to 19 % (infields), alt­ Species which are common, or moderately hough the number of species did not vary mar­ common, usually occur in all parts of the Faroes kedly (cr. Bengtson 1980). The proportion of (see distribution maps in the Appendix). This is staphylinid beetles was about 50 % in most ha­ perhaps not surprising considering the relatively bitats and the lowest values were found on short distance between the southernmost and dwarf shrub heaths (41 %) and on lowland bogs northernmost points of the Faroes (approx. 120 (31 %). In all habitats carnivorous beetles made km) and the small differences in climate and to­ up at least 80 % of the number of coleopterous pography. The northern islands are generally species. more mountainous and have a higher precipita­ The common and most «typical» species (here tion than the isl<{nds further south and Suduroy, defined as those found in at least half the num­ ~md especially Sandoy, have a slightly warmer ber of sites surveyed) in the different habitats and sunnier climate favouring agricultural acti­ were: vities. This may partly explain why some of the In mountain sites: Nebria gyllenhali. N. sa­ less common or rare species often have a lina. Notiophilus biguttatus, Trechus obtusus, southern distribution in the Faroes (e.g. Apion Patrobus septentrionis, Eucnecosum brachypte­ haematodes, Hypocyptus laeviusculus, Tachinus rum, Othius myrmecophilus, and Atheta islan­ la ticollis, Atheta melanocera, Helophorus flavi­ dica. • pesJ. There are no species which have a distinc­ In dwarf shrub heaths: Carabus problemati­ tly northern distribution within the Faroes. cUS, Nebria gyllenhali, N. salina. Notiophilus bi­ guttatus. Trechus obtusus, Patrobus septe/llrio­ nis, P. assimilis, Pterostichus adstrictus, P. nig­ rita, Calathus !uscipes, Encnecosum brachypte­ Comparison of habitats rum, Quedius curtipennis, Q. nitipennis, Hypolit­ The comparison of the taxonomic composition hus riparius, and Otiorrhynchus arcticus. (by family) and number of species in different In plantations: Nebria gyllenhali, N. salina, habitats presented in Tab. 4 is based on the in­ Notiophilus biguttatus, Loricera pilicornis, Tre­ formation contained in Tab. 1. The largest total chus obtusus. Patrobus septentrionis, Anacaena number of species was found in the infield loca­ globulus, Megasternum obscurum, Acrotrichis lities (84 species on the outskirts and 81 species sp., Eucnecosum brachypterum, Stenus impres­ around the settlements). This was not unexpec­ sus, Othius myrmecophilus. Quedius umbrinus.

67 Table 4. Taxonomic composition of Faroese Coleoptera in different habitats: (A) mountain sites, (B) dwarf shrub heaths, (C) plantations, (0) cliffs, shelves, and crevices, (E) lowland bogs, (F) grass heath, (G) outskirts of infields, (H) infield around settlements, and (I) sand dunes. Note the large differences in the number of locali­ ties in different habitats.

Habitat Number of Carabi- Hydro- Staphyli- Crypto- Curculio- Others Total locs dae philidae nidae phagidae nidae surveyed A 4 7 (30) 0 12 (52) 0 3 (13) I ( 4) 23 B 10 12 (35) I (3) 14 (4 I) 0 5 (IS) 2 ( 6) 34 C 4 9 (27) 2 (6) 16 (49) I (3) 2 ( 6) 3 ( 9) 33 0 3 8 (24) 2 (6) 19 (56) 0 2 ( 6) 3 ( 9) 34 E 3 9 (69) 0 4 (3 I) 0 0 0 13 F 31 15 (26) 3 (5) 29 (SOl 0 3 ( 5) 8 (14) 58 G 34 16 (I 9) 5 (6) 42 (59) 3(4) 8 (10) 10(12) 84 H 21 IS (19) 3 (4) 44 (54) 4 (5) 7 ( 9) 8 (10) 81 I I 2 (33) 0 3 (50) 0 1(7) 0 6

Tachinus signatus. Hypolithus riparius. and particular habitat(s). Many of the other species Otiorrhynchus dubius. show low constancies in most habitats and are On cliffs, shelves and in crevices: Nebria gy/­ possibly more specific in their choice of habitat. /enhali. N. salina. Patrobus septentrionis. He/op­ However, it should be kept in mind that carabid hOnls jlavipes. Acrotrichis sp., Lesteva /onge/y­ beetles are much easier to collc!ct than most oth­ trata. Stenus impressus. Othius punctu/atus, At­ er taxa and, for many species, more intensified heta is/andica, Hypo/ithus riparius. Otiorrhyn­ sampling would undoubtedly have increased the chus arcticus, and O. dubius. number of localities. In lowland bogs: Trechus obtusus, Patrobus septentrionis, P. assimilis, Pterostichus adstric­ tus. P. nigrita. Lesteva /onge/ytrata. ACKNOWLEDGEMENTS In grass heaths: Nebria gyllenha/i. N. salina, The data, which provide the basis for this paper, Notiophilus biguttatus. Trechus obtusus, Patro­ were collected in the field by the present author bus septentrionis, Eucnecosum brachypterum, and Drs. P.H. Enckell, Lund, Sweden, and T. Lesteva /onge/ytrata. Othius angustus. O. myr­ Solh0Y, Bergen, Norway, as part of a joint eco­ mecophilus. and Hypo/ithus riparius. logical project supported by grants from the On the outsk irts of infields: Nebria gyllenhali. Norwegian Research Counsil for the Science N. salina. Loricera pilicornis. Trechus obtusus. and Humanities (to S.-A. Bengtson) and the Patrobus septentrionis, Ca/athusfuscipes. C. me­ Nordic Council for Ecology (to T. Solh0Y). /anocepha/us, P. assimilis, Amara aulica. Me­ Drs. A. Fjellberg, Oslo, and K.-E. Erikstad, gasternum obscurum, Othius angusts, Tachinus Bergen, carried our the tedious taxonomic work signatus, Amischa analis, and Hypo/ithus ripa­ and Or. Dorete Bloch Danielsen and Mr. J.K. rius. Brerentsen, Torshavn, assisted in various ways Around settlements in infields: Nebria gyllen­ during the fieldwork. I am also grateful to St6di­ hali. N. salina, Trechus obtusus. Patrobus sep­ sutbugvingin (Academia Freroensis) for provi­ tentrionis, Ca/athus fuscipes. C. me/anocepha/us, ding laboratory facilities and the Royal Danish Loricera pilicornis, Amara au/ica. Megasternum obscurum, Acrotrichis sp., Omalium rivu/are, O. Navy for helicopter support to reach some of the excavatum, Othius angustus, O. myrmecophilus, sampling sites. Quedius mesomelinus. Tachinus signatus. and Hypolithus riparius. On the only sand dune locality the following REFERENCES 6 species were found: Nebria gyllenhali. Ca/at­ Becker, P. 1975. Island colonization by carnivorous hus fuscipes, Atheta vestita. A. arenico/a, Oxy­ and herbivorous Coleoptera. 1. Anim. Eco/. 44. poda haemorrhoa, and Otiorrhynchus arcticus. 893-906. As can be seen from the above lists many of Bengtson, S.-A. 1980. Species assemblages and coex­ the carabid beetles have broad habitat niches istence of Faroe ground beetles (Coleoptera: Cara­ (see Bengtson (I 980) and are not confined to any bidae). En/om. Gen. 6. 251 - 266. 68 Bengtson, S.-A. & Hauge, E. 1979. Terrestrial inver­ CORRECTIONS tebrates of the Faroe Islands: I. Spiders (Araneae): Check-list, distribution, and habitats. Fauna nor1'. In a previous paper (Fauna norv. Ser. B. 26: 59-83 Ser. B. 26, 59-83. ([ 979)) we claimed that 67 species of spiders (Ara­ Berry, RJ. & Johnston, J.L. 1980. The Natural His­ neae) have hitherto been recorded in the Faroe Is­ tory of Shetland. CoBins, London. lands, although we listed only 65 species. The Kauri, H. 1980. Terrestrial invertebrates of the Faroe following two species were left out by mistake. Islands: 11. Harvest-spiders (Opiliones), Fauna Walckenaera oblllsa Blackwall. Previously found nor1'. Ser. B. 27,72-75. once on Streymoy (Bgd 1928). Occurs in Se and Larsson, S.G. & Gigja, G. 1959. Coleoptera I. Synop­ WN. sis of the species. The Zoology of Iceland Vol. 11/ Porr!Jomma montanum Oackson). Previously found (46a), 1-218. Copenhagen & Reykjavik. once on Streymoy (Bgd 1928). Occurs in Se, Sh. I, Lindroth, CH. 1931. Die Insektenfauna Islands und and WN. ihre Probleme. Zool. Bidrag. Bd. 13. 105-599. Furthermore. we wish to draw attention to a foot­ Uppsala. note (added in prooD in Ashmole's paper (1979:92), - (Red. cur,) 1960. Catalogus Coleopterorum Fen­ stating that Or. Ake Holm believes that Faroese spe­ noscandiae et Daniae. Entomologiska Sallskapet i cimens of Centrol/1erita bicolor (Blackwalll should be Lund. 476 pp. referred to as Centromerita concinna (Thorell). - 1968. The Icelandic form of Carabus problemati­ cus Hbst. (Col. Carabidae). A statistic treatment of S.-A. Bengtson & E. Hauge subspecies. Dpuse. Ent. 33, 157 -182. Lindroth, CH., Andersson, H.. B6dvarsson, H. & Richter, S. 1973. Surtsey, Iceland. The develop­ ment of a new fauna, 1963 -1970, terrestrial in­ vertebrates. Ent. scand. Suppl. 5, 1-280. APPENDIX Sigurjonsdottir, H. 1974. Hvemer fara skordyr og att­ Maps showing the distribution of 101 species (and 3 fretlur a kreik a vorin? NdulirufrCEdingurinn 44, identified to genus) of beetles (Coleoptera) recorded 80-94. Reykjavik. in 1978- 1979 in the Faroes. Black circles denote re­ Solh0Y, T. 1981. Terrestrial invertebrates of the Fa­ cords from the present study, stars refer to previously roe Islands: IV. Slugs and snails (Gastropoda): published records (mainly from West (1930)). and as­ Check-list, distribution, and habitats. Fauna nor1'. terisks refer to previous records. where only the Ser. A. 2. name of the island is known. Previous records have Walker, JJ. 1938. On the Coleoptera, etc. of the Fa­ been entered on the maps only when they markely roe Islands. Ent. mon. mag., London, 74, 77-82. supplement our data. For information on position of. West, A. 1940. Coleoptera. ZooIOK)' q{the Faroes 40, sampling efforts at, and vegetation of localities surve­ I -92. Copenhagen. yed see Bengtson & Hauge (1979: Fig. 2 and Appen­ dix I). The distribution maps are presented in the Received 15 Dec. 1980. same order as the species appear in the synopsis.

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q I 1__q_. o_1 ';~ I The Chalcid fauna of the Jostedalen (Hym. Chalcidoidea)

STEVE COMPTON

Compton, S. 1981. The chalcid fauna of the Jostedalen (Hym., Chalcidoidea). Fauna norv. Ser. B. 28, 83-89.

A total of 175 species of Chalcidoid Hymenoptera (excluding Mymaridae and Eulophidae, Tetrastichinae) were collected in the Jostedalen during July and August 1979 as part of the Hull University Biological Expedition to Norway. Of these 124 have been identified and are listed with location and date ofcapture. Fifteen further species, found elsewhere in Norway, but not in the Jostedalen area are also listed. Steve Compton, Unit of Genetics, University of Hull, Hull, Hu6 7RX, England.

This survey of the chalcid fauna of the Josteda­ Several rare or otherwise interesting species len was carried out during the 1979 Hull Uni­ were present, -including Thektogaster subvire­ versity Biological Expedition to Norway. scens (Zetterstedt), known previously only from were collected by sweeping during two nineteenth-century records from Lapland July and August 1979 in the Jostedal Valley, (Graham 1969). Elachertus longipetiolus Bou~­ Inner Sogn, Norway, by myself and other ek and Pediobius coxalis Bourek are species members of the expedition. It has proved pos­ new to the Western European fauna. Similarly, sible to identify most of the species obtained ex­ a single female of a species of Trjapitzinellus cept for the Mymaridae and Tetrastichinae. Of Viggiani collected at Myklemyr on 11 th August those which could not be determined, several is the first record of this genus from North Wes­ may represent species as yet undescribed. tern Europe. Dig(vphus begini (Ashmead) was The species obtained from the Jostedaien are abundant in the Jostedalen at lower altitudes. listed in Table 1, with their abbreviated locali- This well-known North American species • tion and date of capture. The locality abbrevia­ (Gordh and Hendrickson 1979) is probably wi­ tions are explained in Table 2. Nomenclature, despread in Western Europe, but its identity where possible, follows Fitton et al (978) and here has only recently been established (Z. Bo­ Boucek and Askew (968). Species marked urek, personal communication). with an asterisk (*) represent probable new re­ To allow future comparison of the Jostedalen cords for Fennoscandia. chalcid fauna with others elsewhere, Table 3

Table I. Chalcids collected in the Jostedalen during July and August 1979. (For explanation oflocality abbrevi­ ations see Table 3. Species marked with an asterisk are probably new records for Fennoscandia).

EURYTOMIDAE Tetramesa petiolata (Walker) G 16.vii 1m Eurytoma (Bruchophagus) platyptera (Walkerl L 4.viii 8f 3m

TORYMIDAE Megastigmus aculeatus (Swederus) G 16.vii 1f Megastigmus bipunctatus (Swederus) NF 6. viii 1f, VDM 11.viii 2f 4m Torymus (Lioterphusl i'uscicornis (Walker) L 24.vii 2f, NG 19.vii 1f

Fauna norv. Ser. B 28,83-89. Oslo 1981. 83 Table I (cont'd.)

PERILAMPIDA~ Perilampus tristis Mayr VOM 11.viii 1f

PTER OMAL. IDAE Cea pulicaris Walker SO 22.vii 2m Spalangia erythromera Forster VOM 11.viii Sf 1m Asaphes suspensus (Nees) G 19.vii 2f 1m Asaphes vulgaris Walker G 16.vii 2f Chrysolampus thenae (Walker) G 16.vii 1m Panstenon oxylus (Walker) SO 22.vii 2f ·Sphegigaster nigricornis (Nees) G 19.vii 1f Sphegigaster pallicornis (Spinola) NF 6.viii If Cyrtogaster vulgaris Walker F 24.vii 3f L 4.viii 1f NF 6.viii 2f Toxeuma acilius (Walker) NG 19.vii 1f Halticoptera brevicornis F 24.vii 18f 19m. FB 22.vii 2f 3m Thomson G 19.vii 2f 6m, L 24.viiJ 1m L 4.viii 3m NF 6.viii 1f Sm NG 19.vii 2m VOM 11.viii 3f 1m Halticoptera circulus (Walker) F 24.vii 1f FB 22.vii 1m L 4.viii 1m LO 16.vii 1m NF 6.viii 1f 1m NG 19.vii 1m Merismus lasthenes (Walker) F 24.vii 4f 2m, G 19.vii 1m Merismus nitidus (Walker) VOr'1 11. viii 1m Nodisoplata diffinis (Walker) NO 16.vii 1f NO 17. vii 1f SO 22.vii 6f 3m Seladerma geniculatum (letterstedt) F 24.vii 1m ·Seladerma s:oticum (Walker) FB 22.vii 1f G 16.-vii 1f Thektogaster subvirescens (letterstedt) NF 6. viii 1f Sphaeripalpus viridis Forster F 24.vii 1f VD 28.vii 1m VOM 11.viii 1m ·Stictomischus groschKet F 24.vii 1f G 19. vii 2f Oelucchi SO 22.vii 2f VOM 11. viii 1f Stictomischus obscurus (Walker) FB 22.vii 1f VOM 11.viii 1f Stictomischus tumidus (Walker) L 4. viii 1f VOM 11.viii 1m Miscogaster hortensis Walker G 16.vii Sf 1m NG 19. vii 1f VD 28.vii 1f Miscogaster maculata Walker F 24.vii 1f ·Lamprotatus simillimus Oelucchi FS 13.vii 3f NO 17.vii 1f SO 22.vii 6f 7m VSO 19.vii 10f 1m Lamprotatus splendens Westwood FB 22.vii 1f LO 16.vii 1f VD 28.vii 1f ·Skeloceras novickyi Oelucchi NF 6.viii 1f SO 22.vii 1f VOM 11.viii 2f Skeloceras sQcium (letterstedt) G 19.vii 1f 1m Semiotellus diversus (Walker) F 24.vii 1f

84 Table I (eont'd.)

Semiotellus mundua (Walker F 24. vii 2f G 19.vii 1f VDM 11.viii 1f *Gastraneistrus amaboeus Walker NG 19.vii 1f Gastraneistrus eompressus Walker NF 6.viii 1m G 19.vii 1f 1m *Gastraneistrus erassus Walker F 24.vii 2f G 19.vii 5f S 31.vii 1f *Gastraneistrus dispar Graham NF 6.viii 1f *Gastraneistrus fumipennis Walker G 16.vii 1f Gastraneistrus glabellus (Nees) FB 22.vii 2f 1m, G 19.vii 2f VDM 11.viii 1m Gastraneistrus hamillus Walker NO 16.vii 1f NF 6.viii 2f Gastraneistrus punetieollis (Thomson) G 19.vii 1f Eerizotes montieola Forster F 24.vii 2f L 4.viii 1f NF 6.viii 1f *Pirene eonjungens Graham VDM 11.viii 2f Pirene varieornis Haliday F 24.vii 6f 1m, FB 22.vii 1f NF 6.viii 1f NG 19.vii 2f VDM 11 •viii 1f Habritys brevieornis (Ratzeburg)VDM 11.viii 1f Psiloeera atra (Walker) G 16.vii 1f Spintherus dubius (Nees) G 16.vii 1f 19.vii If Holeaeus eompressus (Walker) NG 19.vii 5f Eulonehetron torymoides (Thomson) F 24.vii 1f 'Stenomalina epistena (Walker) L 4.viii 1f Stenomalina illudens (Walker) G 19.vii 1m VD I"! 11.viii 1f *Chloroeytus formosus (Walker) G 19.vii 1f Chloroeytus spieatus (Walker) F 24.vii 1f G 16.vii 3f Psilonotus aehaeus Walker NF 6.viii 1f Psilonotus ~damas Walker L 4. viii 2m I Mesopolobus ineultus (Walker) G 19.vii 1f Pegopus inornatus (Walker) L 24.vii 1f L 4. viii 1f LD 16.vii 1f VDM 11.viii 4f Triehomalus helvipes (Walker) F 24.vii 1f G 16.vii 2f Triehomalus inops (Walker) F 24.vii 1f FB 22.vii 2f G 16.vii 1f G 19.vii 2f L 24.vii 1f NF 6.viii 1f Triehomalus nanus (Walker) NF 6.viii 4f VDM 11.viii 1f VSD 19.vii 2f Triehomalus perfeetus (Walker) VDM 11.viii 1m Triehomalus rufinus (Walker) F 24.vii 1f L 4.viii 1f Digloehis sylvieola (Walker) HS 5.viii 1m Coelopisthia extenta (Walker) NF· 6.viii 1f SKD 25.vii ff

ENCYRTIDAE Doliphoeeras belibus (Walker) F 24. vii 1f Ginsiana earpetana (Mereet) G 19.vii 1f Prionomitus tiliaris (Dalman) VDM 11.viii 2f

85 Table I (cont' d. )

APHELINIDAE Aphelinus abdominalis Oalman VOM 11.viii 1f Aphelinus chaonia Walker F 24.vii 1f VOM 11.viii 4f Aphelinus varipes (Forster) G 16.vii 2f

TETRACAMPIDAE Platynocheilus cuprifrons (Nees) FB 6. viii 1m Foersterella flavipes (Forster) VOM 11.viii 1f

EULOPHIDAE Euplectrus bicolor (Swederus) VOM 11.viii 1f Elachertus artaeus (Walker) G 19.vii 1m L 24.vii 1f NO 16.vii 1f 1m. NO 17.vii 1f VOM 11.viii 1f Elachertus inunctus (Nees) NF 6.viii 1f *Elachertus longipetiolus Bou~ek VOM 11.viii 1f Cirrospilus vittatus Walker G 19.vii 1f J Sympiesis acalle (Walker NF 6.viii 1f VOM 11.viii 1f Sympiesis notata (Zetterstedt) VOM 11 .viii 1m Sympiesis xanthostoma (Nees) VOM 11.viii 1m Hemiptarsenus dropion (Walker) NG 19.vii 1m Oicladocerus westwoodii Westwood VOM 11.viii 2f *Pnigalio agraules (Walker) NO 16.vii 7f SO 22. vii 1f VOM 11.viii 1f Pnigalio longulus (Zetterstedt) NO 17.vii 1f Pnigalio pectinicornis (Linnaeus) L 4.viii 1f G 19.vii 1f NF 6.viii 1f Pnigalio soemius (Walker) L 4.viii 1f SO 22.vii 2f 3m Pnigalio tridentatus (Thomson) VOM 11.viii 1f Eulophus larvarum (Linnaeus) VOM 11.viii 1f Necremnus artynes (Walker) G 19.vii 1m Necremnus cosconius (Walker) G 19.vii 2m Oiglyphus chabrias (Walker) G 19.vii 1f VOM 11.viii 2f *Oiglyphus begini (Ashmead) FS 13.vii 1f G 19.vii 2f LO 16.vii 1f 1m. NF 6.viii 6m VOM 11.viii 7f 2m Euderus albitarsis (Zetterstedt) G 19.vii 2f VOM 11.viii 1f Euderus viridis Thomson F 24.vii 1f

Entedon diotimus Walker F 24.vii 1f G 16.vii 2f 1m G 19.vii 3f 3m. L 4.viii 1f *Entedon fufius Walker G 19. vii 1f Pediobius acantha (Walker) G 19.vii 1f L 4.viii 1f NF 6.viii 1f VD 28.vii 1f VOM 11.viii 2f 1m Pediobius alcaeus (Walker) VOM 11.viii 1f 1m *Pediobius coxalis Bou'gek VOM 11.viii 1f Pediobius epigonus (Walker) F 24.vii 1f NF 6.viii 1f NG 19.vii 1m 86 Table I (cont 'd. )

EULoPHIDAE (cont'd.)

Pediobius eubius (Walker) G 19.vii 1f 1m. NG 19.vii 1f VD 28.vii 1m VoM 11.viii 6f 1m Pediobius saulius (Walker) NO 17. vii 1f Chrysocharis liriomyzae oelucchi G 19.vii 2m Chrysocharis phryne (Walker) NO 16.vii 3f SO 22.vii 2f VDM 11.viii 1f Chrysocharis prodice (Walker) G 19. vii 1f HS 23.vii 2f L 24. vii 1f NG 19. vii 2f 1m VD 2B.vii 2f VDM 11. vii 2f

Chrysocharis pubicornis F 24.vii 1f FB 22.vii 1f 1m (Zetterstedt) L 4.viii 1m NO 16.vii 1f 1m NF 6.viii 2f 1m. SO 22.vii 2f 1m VD 28. vii 2f VDM 22.viii 2f 3m *Chrysocharis truncatula Graham NG 19.vii 1f Achrysocharoides niveipes L 24.vii 2f NG 19. vii 1f (Thomson) Chrysonotomyia formosa (Westwood) FS 13.vii 1f NF 6.viii 1f SO 22.vii 1f 1m *Asecodes coronis (Walker) VDM 11.viii 1f *Asecodes mento (WcHker) F 24.vii 12f G 16.vii 14f 1m G 19.vii 9f 2m. HS 23.vii 1f L 4. viii 1f NO 17.vii 2f NG 19. vii 4f 3m. VD 28.vii 2f VDM 11.viii 2f 3m *Dmphale acamas (Walker) F 24.vii 1f FB 22.vii 1f GO 11.viii 1f NO S.viii 1f VD 28.vii 2f VDM 11.viii 1f *omphale betulicola Graham VDM 11.viii 2f *Omphale coi 1A.Js (Walker) F 24.vii 1f FB 22.vii 1f Lo 16.vii 1f NF 6.viii 1f VD 28.vii 10f VDM 11 •viii 10f *omphale phaola (Walker) SO 22.vii 49f *omphale rubigus (Walker) F 24. vii 1f G 19. vii 1f omphale salicis Haliday F 24.vii 8f FB 22.vii 3f 1m G 19.vii 3f L 4.viii 3f NF 6.viii 1f 1m. VDM 11.viii 8f Sm

omphale theana (Walker) F 24.vii 1f L 4.viii 2f NF 6.viii 4f VDM 11.viii 2f Ionympha carne (Walker) VDM 11.viii 1f Ceranisus menes (Walker) VDM 11.viii 1f *Ceranisus pacuvius (Walker) SO 22.vii 1m

m " male f female

87 Table 2. Details of localities presented in abbreviated form in Tables 1 and 4.

Abbreviation (Jostedalen Area, Table I) Locality F Fossen (MP)925) FB Faberg (MPI139) FS Fabergstolen (MP 1343) G Gaupne (MP0809) GD Geisdalen HS Haugafjellstolen (MP0736) L Leirdal (MP0517) LD Leirdalen Dam (MP041 7) ND Nigardalen NF Nlgard to Faberg Road NG Gaupne to Leirmo Road S Snotun (MP0635) SD Stordalen SKD SkaldaJen VD Vigdalen VDM MykJemyr and Vigdalen hair-pins VSD Vassdalen (Elsewhere in Norway, Table IV) H Hafslo MN0399) K Kaupanger J o Oslo B Paradis, Bergen

Table 3. Summary of the Chalcid fauna collected in the Jostedalen during July and August 1979.

Family Sub-Family Genera Species Specimens Eurytomidae 2 7 22 Torymidae 3 10 68 Perilampidae 1 I I Pteromalidae - Ceinae 1 I 2 - Spalangiinae 1 I 6 - Asaphinae 1 2 5 - Chrysolampinae 1 1 1 - Panstenoninae 1 1 2 - Miscogasterinae 17 40 228 - Pteromalinae 17 37 105 39 - 83 - 349 Encyrtidae 9 16 23 Aphelinidae 2 4 9 Tetracampidae 2 2 2 Eulophidae - Eulophinae 10 20 65 - Tetrastichinae I - 375 - Euderinae 1 2 7 - Entedontinae 9 29 293 21 - 51" - 740 Trichogrammatidae I 1 1 Mymaridae - - 38

"Total Eulophidae excluding Tetrastichinae.

88 Table 4. Chalcids collected in Norway outside the Jostedalen area, 1979. (For explanation of locality abbrevia­ tions see Table 2. Species marked with an asterisk are probably new records for Fennoscandia).

Syntomopus oviceps Thomson H 3.viii Im *Seladerma caledonicum Graham B Il.vii 1 f Seladerma sabbas (Walker) B Il.vii I f Seladerma tarsale (Walker) B 10.vii I f;B Il.vii 2 f Homoporus luniger (Nees) K 27.vii I f,1 m Pteromalus puparum (Linnaeus) H 3.viii Im *Chlorocytus deschampsiae Graham H 3.viii I f Elachertus isadus (Walker) a 16.viii I f Hemiptarsenus unguice//us (Zetterstedt) B Il.vii 2m Entedon hercyna Walker B Il.vii 2 f Chrysocharis melaenis (Walker) B Il.vii I f *Chrysocharis polyzo (Walker) B 11.vii I f *Chrysocharis submutica Graham B Il.vii I f Omphale lugens (Nees) B 10.vii 6 f; B Il.vii If *Ionympha ochus (Walker) B Il.vii I f; K 27.vii Im M = male f= female contains a summary of all species and specimens Norway was given by The Royal Society, The obtained including those for which identifica­ Sir Phillip Reckitt Education Trust, the lames tion is not complete. Bou~k (I 977) has sugges­ Reginald Heslam Trust and the Hull University ted that in Europe the number of chalcid species Expedition Fund. to be found in an area approximates to the num­ ber of plant species growing there. This sample of around two hundred species would be only a small part of the total fauna predicted on this REFERENCES basis for the Jostedalen, but nonetheless provi­ Boucek, Z. 1977. A faunistic review of the Jugosla­ des an indication of the relative importance of vian Chalcidoidea (Parasitic Hymenoptera). Acta particular families and sub-families in terms of ent.jugosl. Suppl. /3. 1-146. abundance of genera, species and individuals. Boucek, Z. & Askew, R.R. 1968. Paleartic Eulop­ For example, Miscogasterinae and Entedontinae hidae (excl. Tetrastichinae). Index entomoph. Ins. are groups especially well represented in the 10s­ 3. 1-254. tedalen. Fitton, M.G., Graham, M.W.R. de V., Bou~ek, Several chalc'id species obtained en route to Z.R.T., Fergusson, N.D.M., Huddleston, T., and from the 10stedalen were not found in the Quinlan J. & Richards, a.w. 1978. A Check List of British Insects: Hymenoptera. Handbk. valley itself. These are listed in Table 4 with ident. Br. Insects II (4). 1-159. locality abbreviations explained in Table 2. Gordh, G. & Hendrickson, R. 1979. New species of Diglyphus. a world list of the species, taxonomic notes and a key to new world species of Diglyphus ACKNOWLEDGEMENTS and Diau/inopsis (Hymenoptera, Eulophidae). Proc. ent. Soc. Wash. 8/ (4). 665-684. I would like to thank Dr. R.R. Askew for confir­ Graham, M.W.R. de V. 1969. The Pteromalidae of ming and correcting my identifications of Nor­ north-western Europe. (Hymenoptera: Chalcidoi­ wegian Chalcidoidea and Drs. Z. Bourek, M.F. deal. BI//I. Br. MI/s. nat. Hist. Ent. SI/ppl. /6. Qaridge, M.W.R. de V. Graham and 1.S. Noyes 1-908. for their help and encouragement. Financial as­ sistance for the Hull University Expedition to Received 4 Nov. 1980.

89 The migrant moth Spodoptera exigua (Hubner) (Lepidoptera, Noctuidae) recorded in Norway

LEIF AARVIK

Aarvik, L. 1981. The migrant moth Spodoptera exigua (Hiibner) (Lepidoptera, Noctuidae) recorded in Norway. Fauna norv. Ser. B. 28. 90-92.

A capture of Spodoptera exigua (Hiibner) at Siljan TEy in August 1964 is recorded. The 1964 invasion to north Europe is reviewed and possible immigration routes to Norway are discussed. Notes on diagnostic characters, distribution, and biology are given.

Leif Aarvik, Tarnveien 6, N-1430 As, Norway.

INTRODUCTION followed a warm wind from southeast. On 7 Among Lepidoptera several migrant species are August the wind turned, and the swarm conti­ known. In the family Noctuidae the pest species nued in a southwestern direction. Two speci­ Mythimna unipuncta (Haworth) is well known. mens were caught in Riga on 8 August. The Another migrant noctuid is Autagrapha gamma swarm passed southeast Sweqen and reached (L.), which is often seen in great numbers in Denmark on 10 August when one specimen Scandinavia. were caught in Copenhagen and four ones were One of the most remarkable moth immigra­ caught in Bornholm. tions to north Europe was the invasion of Spa­ The first Swedish records do not fit in with daptera exigua (Hiibner) to Finland and neigh­ the advance schedule reviewed above. Two spe­ bouring countries in 1964. This invasion was cimens were trapped in Sodermanland near Ny~ analysed by Mikkola & Salmensuu (I965), and koping on the night 6- 7 August, when the van the arrival route to Finland was traced. of the swarm still was in Finland 400 km. away. When examining unidentified Lepidoptera in a third early specimen was captured in Oland the collection of Alf & Sigurd Bakke, I discove­ the following night. This group of moths is beli­ red two females ofSpadaptera exigua. They had eved to have advanced to Sweden in the upper been captured in a light trap at Siljan TEy (EIS atmospheric strata rather than having flown ex­ 18) on 15 August 1964, Alf Bakke leg. This is ceptionally through the day. the first capture of the species in Norway. Altogether 1365 specimens were captured in Finland from 5 August to 12 September. The northernmost locality was Utajiirvi in Ostrobott­ The arrival route of the 1964 immigration to nia borealis southern part. Finland, Sweden, and Denmark In Sweden 19 specimens were caught bet­ The course of the swarm has been thoroughly ween 6 August and I September in seven pro­ described by Mikkola & Salmensuu (1965). Here vinces from Vastmanland and Uppland south­ a brief review only is presented. wards to Skane (Nordstrom et at. 1969). Because of intense collecting activity with In Denmark 41 specimens were caught bet­ light traps in Finland in August I 964 it was pos­ ween 10 and 28 August. Records were made in sible to obtain information on the invasion. Spe­ Bornholm, M0n, Falster, Zealand, and Jutland cimens captured in other countries were also at­ (Kaaber & Norgaard 1965). tracted to light. Meteorological data as well as reports of the species from the USSR indicated The arrival route to Norway that the swarm originated from an area in the Obviously there are two alternatives as to the direction of Kazakstan. The swarm reached moths' arrival route: Either they have come southeast Finland on 6 August, and at that time from the east having crossed central Sweden it probably consisted of several hundred million and the Oslofjord area, or they have come from individuals. During three days it passed Finland, the south, from northern Jutland. These alterna­ decreasing all the time. So far the swarm had tives are indicated on the map (Fig. I).

90 Fauna norv. Ser. B 28, 90-92. Oslo 1981. less filled with yellow. The former has a charac­ mily Solanaceae, the tea plant, and grape-vine. teristic reddish brown dot in the middle. Hind­ The species develops very quickly. Development wings silky, translucent, with veins and termi­ indoors from the egg to the imago stage took lit­ nal line dark. tle more than a month (Mikkola & Salmensuu 1965).

Distribution of Spodoptera exigua ACKNOWLEDGEMENTS The distribution of S. exigua is almost cosmopo­ litan. It occurs in Europe, Australia, tropical and I wish to express my gratitude to L. Imby, K. subtropical Asia, the whole of Africa, and large Larsen, and K. Mikkola for valuable informa­ parts of America (Mikkola & Salmensuu 1965). tion, to G. Wiig for taking the photographs, and The first record in north Europe was made in to A. Moene and J.-E. Johnsen for providing the Bornholm in 1924, the second at Fan0, Den­ meteorological data. Finally I am indebted to Dr. mark in 1947. Danish records were also made in A. Bakke for loan of material and for valuable 1952, 1960, and in 1962 when 60 specimens comments on the manuscript. were captured in Zealand. The first Fennoscan­ dian specimens were recorded in Finland 1958 (two specimens) and 1959 (one specimen) (Mik­ REFERENCES kola & Salmensuu 1965). In Denmark S. exigua Kaaber, S. & Norgaard, 1. 1965. Fund afstorsommer­ was also captured in the years 1966, 1967, fugle fra Danmark i 1964. Flora Fauna Silkeborg 1968, 1969, 1972, 1974, and 1975 (K. Larsen 71, 99 - 194. , pers. comm.). In Sweden S. exigua was captu­ Mikkola, K. & Jalas, 1. 1979. Suomen perhoset: Yak­ kaset 2. Otava, Helsinki. J red in Gotland in 1969 and 1972 (L. Imby pers. Mikkola, K. & Salmensuu, P. 1965. Migration of comm.), and in 1972 further specimens were re­ Laphygma exigua Hb. (Lep., Noctuidae) in Nort­ corded in south Finland (Mikkola & Jalas 1979). hwestern Europe in 1964. Annls. zoo\. fenn. 2, 124-139. Nordstrom, F., Kaaber, S., Opheim, M., Sotavalta, O. Biology of Spodoptera exigua & Douwes, P. (ed.) 1969. De fennoskandiska och' danska nattjlynas utbredning (NoctuidaeJ. CWK According to Mikkola & Salmensuu Cl 965) the Gleerup, Lund. larva of S. exigua is highly polyphagous. They mention cotton, maize, cereals, plants of the fa­ Received 9 Feb. 1981.

92 The species of the subgenus Tipula L. (Dipt., Tipulidae) in Norway

TROND HOFSVANG

Hofsvang, T. 1981. The species of the subgenus Tipula L. (Dipt., Tipulidae) in Norway. Fauna norv. Ser. B. 28, 93-95.

The distribution of the two Norwegian species, Tipula (Tipula) paludosa Meigen 1830 and Tipula (Tipula) czizeki de long 1925, is presented. Some comments are given upon the lar­ vae of the two species as agricultural pests.

Trond Hofsvang, Agricultural Entomology, Agricultural University of Norway, p.a. Box 70, N-1432 As-NLH, Norway.

INTRODUCTION cies were especially looked for and collected du­ Larvae of Tipulidae can be serious agricultural ring the years 1975 -1980 in the Oslo-As pests feeding on roots and undergrown stems of area. different crops, such as cereals, root crops and vegetables. On warm, damp nights larvae can be found on the surface, destroying leaves and cut­ RESULTS ting off plants at soil level. Injuries are frequent Fig. I shows the records of adult T. paludosa when arable crops are planted on old pastures of and T. czizeki from Norway on EIS-maps (ef. leys which have been ploughed in. The most se­ 0kland 1977). During the years 1975 -1980 rious pest species are found within the subgenus adults of T. paludosa were captured during the Tipula L. period 24 July- 27 August in the Oslo-As area Only three of the ten European species of the (EIS 28). The total material presented in Fig. I subgenus Tipula) occur in the north-western was recorded between 10 July and 4 September. part of Europe (Den Hollander 1975). The three T. czizeki has a late appearance as imago and species are Tipula (Tipula) paludosa Meigen has probably been overlooked. This is the pos­ 1830, Tipula (Tipula) oleracea L. 1758, and Ti­ sible explanation of the scarce material of this pula (Tipula) czTzeki de Jong 1925. species (Fig. I). The collecting dates of the total • T. oleracea has been reported as far north as material of T. czizeki from Norway are between Skane in Southern Sweden (Tjeder 1953, 1955, 29 August and 7 October. pers. comm.). T. paludosa and T. czizeki are dis­ When one is aware of this species, a distinct, tributed north to Lule Lappmark in Sweden but short flight period can be observed each (Tjeder 1955, 1975, Tobias 1975). The present year. In the 0810-As area adults were found 18 study summarizes the hitherto known distribu­ September-7 October 1979 and 19 Septem­ tion of T. paludosa and T. czizeki in Norway T. ber-23 September 1980 in rather large amo­ oleracea has not been recorded from Norway. unts on lawns.

MATERIAL AND METHODS DISCUSSION Adult specimens of T. paludosa and T. czizeki T. paludosa has a flight period in late July and in from the zoological museums in Norway are in­ August in Norway. The flight periods in other cluded. Additional records published by Lack­ parts of western Europe are from early August schewitz (I 935) and Tjeder (I965) are given. to the middle of September, sometimes extended Specimens have also been collected by T. Ed­ to early October (Robertson 1939, Maercks land, R. Mehl, G. Taksdal and the author from 1951, Laughlin 1967, Den Hollander 1975). In different parts ofthe country. To study the flight England northern populations of T. paludosa in periods more closly, specimens of the two spe- areas with relatively cold wet climate have an

Fauna norv. Ser. B 28, 93-95. Oslo 1981. 93 are most probably caused by last instar larvae of T. paludosa prior to pupation in July T. palu­ dosa overwinters in the third larval stage, altho­ ugh some second instar larvae are present in the population during the winter (Coulson 1962, Laughlin 1967). When the soil temperature rises in the spring, the fourth instar larvae have a burst of feeding and increase the mean weight by 200 - 300 % and reach a peak weight of 300-500 mg (Laughlin 1967). The larvae of the different species of the sub­ genus Tipula are difficult to separate (Theowald 1967). The second Norwegian species of this subgenus, T. czizeki, spends the winter in the egg stage (Sellke 1936, Maercks 1941 b), and has a very late flight season. The last instar larvae of T. czizeki most likely occur in late July and Au­ gust in Norway and are therefore not respon­ sible for the reported injuries. In Sweden only T. paludosa has been reported as an agricultural pest species (Borg 1964). However, one cannot \ disregard that fullgrown larvae of T. czizeki might do some damage locally Jby feeding on roots in late summer.

ACKNOWLEDGEMENTS I am grateful to Kaare Aagaard, Troms0 Mu- • seum, Trorns0; John O. Solem, The Royal Nor­ wegian Society of Sciences, and Letters, The Fig. I. Records of adult Tipula (Tipula) paludosa Me­ Museum, Trondheim; Lita Greve Jensen, Mu­ igen, 1830 (A) and Tipula (Tipula)'czizeki de long, seum of Zoology, Bergen and Albert Lilleham­ 1925 (B) from Norway. mer, Museum of Zoology, Oslo, for information and loan of material. I also thank T. Edland, R. Mehl and G. Taksdal for providing me with ma­ earlier flight season, more like the Norwegian terial. records, as a results of a shortening of prepu­ pal/pupal period (Coulson 1962, Meats 1975). REFERENCES The reports of the Government Entomolo­ gists W.M. Sch0yen (1891-1913) and T.H. Borg, A. 1964. Angrepp av harkrankslarver pa jord­ Sch0yen (1914 - 1941) and inquiries to the Nor­ bruksvaxter i Sverige. - In: Beretning om Nor­ wegian Plant Protection Institute (1947 -1980) diske lordbruksforskeres Forenings 12. kongress. Helsingfors 1963. Nord. Jordbrug~rorskning. show that the most serious injuries oftipulid lar­ Suppl. 8, 366-370. vae, occur in the coastal areas ofSouth Norway, Coulson, l.C. 1962. The biology of Tipula subnodi­ especially along the western coast. Eggs and first comis Zetterstedt, with comparative observations larval instar of T. paludosa are suspectible to de­ on Tipula paludosa Meigen. J. Anim. Ecol. 31, siccation (Maercks 1939, Coulson 1962, Milne 1-21. et al. 1965). In Germany the injuries are severe Hollander, l. Den, 1975. The phenology and habitat in the north-western part with a more coastal of the species of the subgenus Tipula Linnaeus in climate (Maercks 1939, 194Ia). The western the Nederlands (Diptera, Tipulidae). TUdschr. Ent. part of South and Mid-Sweden have a similar 118. 83-97. Lackschewitz, P. 1935. Zur Kenntnis der polyneuren climate condition, and the most serious attacks Nematoceren (Dipt.) des nordlichen Norwegens. by tipulid larvae on crops occur here (Borg Tromso Mus. Arsh. 53, 3-27. 1964). Laughlin, R. 1967. Biology of Tipula paludosa; The injuries on crops by tipulid larvae in Nor­ growth of the larva in the field. Entomologia expo way are reported from May and June, and they appl. 10, 52-68.

94 Maercks, H. 1939. Untersuchungen zur Biologie und retning. TU/egg C Ii/ Landbruksdirekwrens berel­ Bekiimpfung schadlicher Tipuliden. Arb. physio/. nil/g. al/gew. Enl. 6, 222-257. - 1934 og 1941. Statsentomolog T.H. Sch0yens - 1941 a. Das Schadauftreten der Wiesenschnaken melding. TU/egg C IU Landbruksdireklorens drs­ (Tipulidenl in Abhangigkeit von Klima, Witte­ me/dil/g. rung und Boden. Arb. physio/. angew. Enl. 8, Sch0yen, W.M. 1891. Indberetning fra landbrugsen­ 261-275. tomologen. Aarsberelnil/g Ira Lal/dbrugsdireklo­ - 1941 b. Ober Biologie und Schadlichkeit der ren. Herbstschnake (Tipu/a czizeki de J.l. Arb. physio/. - 1892-1894. Landbrugsentomolog Sch0yens be­ al/gew. Enl. 8, 197 - 205. . retning. Aarsberell/ing fra Landbrugsdirekwren. - 1951. Lebenweise und Bekampfung der Wie­ - 1895 -1913. Statsentomolog Sch0yens beretning. senschnaken. F/ugb/. bioi. BundesanSI. Bra­ Aarsberell/ing Fa Landbrugsdirektoren. unschw. CI7, 1-8. Sellke, K. 1936. Biologische und morphologische Meats, A. 1975. The developmental dynamics of Ti­ Studien an schadlichen Wiesenschnaken (Tipuli­ pu/a pa/udosa and the relation of climate to its dae, DipU. Z. wiss. Zoo/., 148, 465-555. growth pattern, flight season and geographical Theowald, Br. 1967. Familie Tipulidae (Diptera, Ne­ distribution. Oec%gia 19, 117 -128. matoceraJ. Larvel/ ul/d Puppen. Akademie-Ver­ Milne, A., Laughlin, R. & Coggins, R.E. 1965. The lag, Berlin. 1955 and 1959 populations crashes in the leather­ Tjeder, B. 1953. The identity of Tipu/a o/eracea L. jacket, Tipu/a pa/udosa Meigen, in Northumber­ Opusc. EI/t. 18, 111-115. land. J. Anim. Eco/. 34, 529-544. - 1955. Catalogus Insectorum Sueciae XIV. Dip­ 0kland, J. 1977. Litt om biogeografiske metoder, og tera: Fam. Tipulidae. OpIlSC. Ent. 20, 229-247. noen nye data om utbredelsen av stavtege, Rana­ - 1965. Faunistic notes on Norwegian Tipulidae Ira Iinearis, og vannskorpion, Nepa cinerea. i (Dipt.l. Norsk en/. Tidsskr. 13, 41-46. Norge. Fauna, Oslo 30, 145-167. - 1975. Harkrankar (Tipulidae, partiml och Glans­ Robertson, A.G. 1939. The nocturnal activity of myggor (Ptychopteridael i Messaureomnidet. crane-flies (Tipulinael as indicated by captures in Norrbottel/s Natursmdskr. I, 1974, 1-5. a light trap at Rothamsted. 1. AI/im. Eco/. 8, Tobias, W. 1975. Zur Faunistik und Tagesperiodik 300-322. einiger Insektenarten Nordschwedens (Tipulidae, Sch0yen, T.H. 1914-1918. Statsentomolog T.H. Typhlocybidae und Geometridael. Ent. Z. Frankf Sch0yens beretning. Aarsberelning fra Land­ A.M. 85, 217-224. bruksdirekwrel/. 1919-1930. Statsentomolog T.H. Sch0yens be­ Received 9 Dec. 1980.

,

95 The genus (Fallen) (Dip., ) in Norway

LITA GREVE

Greve, L. 1981. The genus Opomyza (Fallen) Dipt., Opomyzidae) in Norway. Fauna Ilorv. Ser. B. 28, 96-99.

Material of the genus Opomyza (Fallen) in Museum of Zoology, Univ. Bergen; Museum of Zoology, Univ. Oslo and Royal Society ofSciences.The Museum, Trondheim has been ex­ amined and revised. It includes 107 specimens belonging to 6 species. A key to the six Opo­ myza species known presently in Norway is given, viz. O. jlorum (Fabricius, 1794), O. ger­ minationis (L., 176 I), O. lineatopunctata von Roser, 1840, O. petrei Mesnil, 1934, O. punc­ tata Haliday, 1833 and O. puncte/la Fallen, 1820. O. lineatopunctata von Roser and O. petrei Mesnil are reported from Norway for the first time. Lists of records are given. The distribu­ tion of the species are mapped in the terms of the 50 km squares of the European Inverte­ brate system for Norway.

Uta Greve, Museum of Zoology, University of Bergen, Museplass 3, N-5014 Bergen/Univ., \ Norway. J

INTRODUCfION In the maps EIS squares are used. For material The family Opomyzidae belongs to the group in the different museums the following abbrevi­ in the suborder Cyclorrhapha. ations has been used: DKNVS (Royal Norwe­ Many of species has wing spots, but not all. The gian Society of Sciences, The Museum, Trond-' Opomyzidae are rather small flies with a body heim), ZMO (Museum of Zoology, Oslo), 2MB length of approximately 2.5-4 mm. The head (Museum of Zoology, Bergen). is more or less rounded with one pair of orbital bristles on frons. In some species a prominent peristomal bristle is present which can be mista­ KEY TO THE GENERA OF OPOMYZIDAE IN ken for a vibrissa. In the genus Opomyza the SCANDINAVIA arista is pubescent, in other Opomyzidae, it can I. Postvertical bristles present. Large eyes, formed have longer dorsal hairs. Pre-and post-sutural like a square. Dark wings with white spots. Hit­ dorsoventral bristles are present, I + 2 in some herto not found in Norway..... Anomalochaeta species, I + 3 in others. The wings are relatively ...... Frey, 1921. long and more or less narrow. The subcosta is 2. Without postvertical bristles. Either hyaline partly melted together with radius I. There is a wings or wings with dark spots. Rounded eyes · 2. «break» in the costa shortly before the end of the 3. Wings with prominent hind corners. Arista pube­ subcosta. The scutellum has 2 basal bristles, and scent. The four bristles on scutellum are of the 2 lateral bristles. The legs are lacking dorsal pre­ same size Opomyza Fallen 1820. apical bristles on the tibia, the middle tibiae has Wings without prominent hind corner, usually a long ventral spur. The male genitalia show sy­ more narrow than in Opomyza. Arista with stematic characters. longer dorsal hairs in many species, but not all. The larvae live in the shoots of Gramineae The lateral bristles on scutellum shorter and not and also in cereals, hence the Norwegian name so prominent as the dorsal ones Geomyza «Gressfluer», (Grassflies)...... Fallen 1810. Earlier records used the name Opomyzidae on several genera now placed in other families. Key to the species of Opomyza Fallen recorded from Siebke (877) lists three species of Opomyza Norway from Norway, viz. O. germination is (L.), O. jlo­ This key covers the six species which hitherto have rum (Fabr.) and 0. punctella Fallen. been found in Scandinavia. A seventh species O. nig­ In the list of localities l..0ken's (973) modifi­ riventris Loew 1865 has been found in Finland. For cations of Strand's 0 943) system has been used. the identification of this species see Hackman (1959).

96 Fauna norv. Ser. B 28: 96-99. Oslo 1981. A B Fig. I Male hypopygium and part of cerci seen in profile (A) and from behind (obliquely upwards) (B) O. petrei Mesnii, 1934. Male hypopygium and part of cerci (C) O. lineato­ punctata (V. Roser 1834).

O. nigriventris is easterly in distribution in Finland, REMARKS ~md reported as rare. The male genitalia of O. germinationis and 0. I. The costal margin of the wings infuscated from petrei is very similar. (See Fig. I A and B). The the end of subcosta and all the way to the wing tip ...... 2. hypopygium is of the same form in the two spe­ The costal margin dark only at the wing tip, not cies, but with lower numbers of hairs in O. pe­ all along the lIlargin 3. trei especially near the prominent tooth (I B) on 2. Abdomen dark, brownish grey. Yellow spots late­ the lower end. On the mesolobes of 0. petrei rally near the border of the tergites. Some speci­ there are numerous small black spines (Fig. I B), mens lack yellow spots all together . lacking in O. germinationis. However, the diffe­ ...... Opomyza germinations L., 176 n. rence in colour of the abdomen serves to sepa­ The abdomen yellow with the following dark rate the species. I have seen only two males of O. markings: Dorsally a narrow median line; late­ petrei. One specimen (drawn) lacked the two rally dark borders of the tergites in contrast to the yellow colour O. petrei Mesnil, 1934. large setae found on the top of the hypopygium, 3. Thorax with a dark median line dorsally. Third probably broken off. The other had the two se­ rib with separate dark spots. Occasionally an extra tae intact. Compared to setae in males of O. ger­ crossrib in tha basal cell. Yellow abdomen..... minationis they are distinctly stronger and . . . ' O. Iineatopunctata von Roser, 1840. longer at least in this specimen. Without dark median line on thorax 4. Fig. I C shows a lateral view of the hypopy­ 4. Wings with an extra crossrib in the basal cell. gium of O. lineatopunctata. The form is some­ This rib is always clouded. In some specimens the what like that of O. punctata. O. lineatopunctata cross-rib is not very distinct . lacks the prominent hook found in O. germina­ ...... O. punctella Fallen, 1820 Without extra-crossrib in basal cell...... 5. tionis and 0. petrei shown in Fig. I B. Both setae 5. Third rib without separate clouded spots, someti­ on the top of the hypopygium was broken, one, mes infuscated ..... O. jlorum (Fabricius, 1794) however, is stipled on the Fig. I C drawn from Third rib with separate clouded spots . another specimen...... O. punctata Haliday, 1833.

97 SYSTEMATIC LIST The locality from Ustaoset was 'it 1000 masl in a vegetation of rich grass meadow and herbage. Ot­ I. Opomyza florum (Fabricius. 1794) her specimens from Vollane, Asane, and Brak­ New and revised records: stad, Meland are from gardens. The locality at Ak: Oslo, T0ien EIS 28 I Q leg. H. Siebke 27 July Herdla is a bare grass meadow near the sea and 1847 (ZMO No. 6603). IQ 28 July 1847 (ZMO likewise the locality at Telavag represents the out­ No. 6602), I d 5 Sept. 1849 (ZMO No. 6600), I Q ermost islands on sea west of Bergen. 29 July 1851 (ZMO No. 6601). YE: Sandefjord, 3. Opomyza lineatopunetata von Roser, 1840. Sandefjord EIS 19 I Q leg. H. Siebke (ZMO No. New records: 6616). TEi: Kviteseid, Triset EIS 17 IQ 2 July AAy: Landvik, Skiftenes UTM 32 MK 702747 1975 leg. L. Greve (ZMB). STi: Dovre EIS 79 Id EIS 6 2 d d I Q 6 Aug. 1971 leg. E. Oug. (ZMB). leg. H. Siebke (ZMO No. 6614). HOy: Bergen, Hellenesset EIS 39 I Q I Aug. 1980 The male labelled «Dovre» was according to Si­ leg. A. Fjeldsa (ZMB). O. lineatopunelata is repor­ ebke (1877) taken at: STi: Oppdal Drivstuen. ted from Norway for the first time. The species Unverified records: MRy: Vestnes, Vestnes (Si­ has hitherto not been found in Fennoscandia. It ebke, H., 1877). One specimen is present in has been reported from England (Collin, 1945). DKNVS without any data. (See also under O. Fig. I C male genitalia. punetata. 4. Opomyza petrei Mesnil, 1934. 2. Opolllyza germinationis (L., 1761) New records: New and revised records: Ry: Tysva:r, Karst0 EIS 22 I d 13 July 1981, I Q Ak: Oslo, Oslo EIS 28 Id, I Q leg. H. Siebke 14 July 1981 leg. L. Greve (ZMB). HOy: Ask0y, (ZMO Nos. 6594, 6595), I d leg. L.M. Esmark Herdla EIS 39 1d 29 July 1935 leg. N. Knaben (ZMO Nos. 6597). YE: Larvik, Laurvik (ZMBl. HOi: Strandebarm, Bakke EIS 31 I Q 10, ( = Larvik) EIS 19 I Q leg. H. Siebke (ZMO No. July 1974 leg. L. Greve (ZMB). 6596). Bv: Hol, Ustaoset EIS 42 I Q 3 July 1980 O. petrei is reported from Norway for the first leg. L. Greve (ZMB). Ry: Ha. Brusand EIS 3 I Q 4 time. The species has been reported from Den­ Aug. 1960; Tysva:r, Karst0 EIS 22 8 d d 6 Q Q mark and Southern Sweden (Andersson, 1962) 13 July 1981, Id 3 Q 14 July 1981 leg. L. Greve and England (Coliin, 1945), but not from Finland (ZMB). HOy: Bergen, Bergen EIS 39 I Q 27 June (Hackman, 1980). O. petrei can be distinguished 1874 leg. Schneider (ZMO No. 6598); Bergen (A­ both by the genitalia (Fig. 1 AB) of the male and sane), Vollane EIS 39 I Q 10 July 1972, I Q 6 the colour of the abdomen which is strikingly dif- • July 1981 leg. L. Greve; Os, S0fteland EIS 31 I d ferent from O. germinationis which has the same 25 Sept. 1966 leg. A. L0ken; Telleviki EIS 31 wing colouring. One female was caught together 3 d d I Q 3 Aug. 1980 leg. Arild Fjeldsa; Haus, with O. germinationis in a garden at Bakke. Both Kleppe EIS 40 I d I Q 1I July 1956 leg. species also occurred together at Karst0. Godskeleiren; Sund, Telavag EIS 30 I d 17 July 5. Opomyza punetata Haliday, 1833. 19531eg. A. L0ken; Meland. Brakstad EIS 39 Id New and revised records: 7 July, I d IQ 8 July 1967, IQ 24 Aug. 1968 Ak: Oslo, T0ien EIS 28 I d I Q leg. H. Siebke leg. L. Greve, Fana, Espegrend, (Institute of Ma­ (ZMO Nos. 6604, 6605). HOi: Eidfjord. Simada­ rine Biology) EIS 30 I Q 26 June 1951, I Q 27 len EIS 41 3 Q Q 10 Aug. 1981 leg. L. Greve June 1951 leg. A. L0ken, 5 d d 7 Q Q (Inst. of (ZMB). SFi: Aurland, Aurland EIS 51 2 d d 23 Marine Biology) leg. A. Tj0nneland, Ask0Y, July 1981 leg. L. Greve (ZMB); La:rdal, La:rdals­ Herdla, N. Valen EIS 393 Q Q 26 June 1980 leg. 0yri EIS 51 I Q 24 July 1981 leg. L. Greve Arild Fjeldsa (ZMB). HOi: Strandebarm, Bakke (ZMB). EIS 31 I d 10 July 1974 leg. L. Greve; Jondal, Two specimens from Ak had been determined as T0rrvikbygd EIS 31 I Q 13 July 1971, I d 19 O. florum (Fabr.). Hackman (1959) remarks as to July 1971 leg. H.R. Skjoldal; Kvinnherad, Rosen­ the slight difference in the male genitalia of O. flo­ dal EIS 31 I Q 13 June 1965 leg. student excur­ rum and O. punetata. The difference used in pre­ sion; 4 d d I Q 26 Aug. 1968 leg. student excur­ vious keys (Collin, 1945, Hackman, 1959 and sion, Kvam, Alvik EIS 41 I d 18 July 1964 leg. Czerny 1928) is that in O. punetata the last part of A. L0ken (ZMB); SFi: J0lster, Skei EIS 59 IQ 25 the cubital wing has dark, small spots or clouds. July 1974 leg. L. Greve (ZMB). MRy: Sm0la, However, in O. florum some individuals also have Sm01a EIS 90 I Q leg. H. Siebke (ZMO No. 6593). very weak infuscation. A revision based on a lar­ Vestnes, Vestnes EIS 77 I Q leg. H. Siebke (ZMO ger material should be made. Andersson (J 962) No. 6599). reports O. punetata from southern Sweden and Unverified records: O. germinationis also is Denmark. published by Siebke from 0: Halden, Fredriks­ Siebke (J 877) probably refers to these two speci­ hald, YE: Stavern, Stavern; Sandefjord. Sandef­ mens as var. b. Zett. under O. punete/la Fall. Thus jord and MRy: 0rskog (Siebke, H., 1877). reported as O. punetata Haliday for the first tinw.. Based on the material in this survey O. germi­from Norway in this survey. nationis seems to be common in southern Nor­ 6. Opomyza punete/la Fallen, 1820. way. At least this is proved for western Norway. New and revised records: Ak: Oslo, T0ien EIS 28 Id 24 Aug. 1847 leg. H. 98 ~~ - ~#".. ~r.:- _[>'r ~ \ t'""~;.. ~ r"""~ , -~ , ~ ~ j 'rr , j "~ , ~ ~ "~ ....."'"­ t "I;,-, ­ \ ~~ IW -­ I- - \I( I'" L\ \ Ih , "[,17 -­ If)! ," ~rf Id / ',-> 1'1 .~ [+ I W"'­ PA> '. " , !"~ " ~ " ,l~ ~.:: ~ l \ "- i , \ ". " , I '." ,. t -, L­ \ . H . ­ ~ .! t±t=!-I I A B C L-L_~ 0 Fig. 2, Records of Opomyza Fallen from Norway. ­ von Roser, 1840 - closed circles. 0, punctata Hali­ A) O. jlorum (Fabr., 1794) - closed circles verified day, 1833 - closed stars. D) O. petrei Mesnil, 1934 records, open circle unverified record. B) O. germi­ - closed circles. O. punctelle Fallen, 1820 - closed nationis (L., 1761) - closed circles verified records, stars verified records, open star unverified record. open circles unverified records. C) O. Iineatopunctata

Siebke (ZMO No. 6611), 2 Q Q 30 Aug. 1849 leg, ACKNOWLEDGEMENTS H. Siebke (ZMO Nos. 6609,6610), Id 31 Aug. I am grateful to Albert Lillehammer, Museum of 1849 leg. H. Siebke (ZMO No. 6608), 3 d d IQ leg. H. Siebke (ZMO Nos. 6606, 6612, 6613, Zoology, Oslo and John O. Solem, The Royal 6607), I Q Oslo leg. Esmark (ZMO No, 6615). Norwegian Society of Sciences, The Museum SFi: Aurland, near Tero EIS 51 I Q 21 July 1981, Trondheim for loan of material. I also want to 3 d d 2 Q Q 25 July 1981 leg. L. Greve (ZMB). thank Mr. Knut Rognes, for loan of Unverified records: Storm (I896) mentions 0. specimens from his private collection. puncteffa from STy: Rissa, Stadsbygd. It is not present in the collections of DKNVS.

REFERENCES Geographical distribution Andersson, J., 1962. Siillsynta eller for Sverige nya None of the six species mentioned above have tvavingar. (Diptera). Opusc. ent. 27, 62 -64. been found no~h of southern Tmndelag pro­ Collin, J.E., 1945. The British species of Opomyzidae vince. However, the material is small and very (Diptera). Entomologist's Rec. J. Var., 57, 13 -16, !Surely not representative for the whole country. Czerny, L., 1928, 54c. Opomyzidae. - In: Lindner, (See Fig. 2 A-D). E. (ed.), Die Fliegen der Palaearktischen Region, Hackman (I 959) in his survey of the Finnish 6(1),1-15. material of Opomyzidae reports no finds from Hackman, W., 1959, The Opomyzidae (Diptera of the northernmost parts. He mentions O. Eastern Fennoscandia. Notul, Ent. 38, 1958, florum, 114-126, O. punctella, and O. punctata as fairly widesp­ Hackman, W., 1980. A Checklist of the Finnish Dip­ read in central and southern Finland. 0. germi­ tera. n. Cyclorrhapha. Notul, Ent. 60, 117-162~ nationis has only been found at AJand and L0ken, A., 1973. Studies on Scandivian bumble bees seems to be westernly in distribution. Anders­ (Hymenoptera, Apidae). Norsk ent. Tidsskr, 20, son's (I962) distributional survey reports these 1-218, species as found in central and southern parts of Siebke, H., 1877. Enumeratio /nsectorum Norvegico­ Sweden. In Sweden, however, O. germinationis rum Fasciculum IV. Dipterorum Continentem. is referred to as common which supports my A.W. Bragger, Christiania. K. view that it might be the most common species Storm, V., 1896. Dipterologiske undersogelser. norske Vidensk. Selsk. Skr. /895. 225-241. in Norway. It is found in the mountainous parts Strand, A. 1943. Inndeling av Norge til bruk ved up to 1000 masl at Ustaoset. faunistiske oppgaver. Norsk ent. Tidsskr. 6, O. petrei is not uncommon in southern Swe­ 208-224. den, but few specimens of this species and 0. li­ neatopunctata have been found. Received June 2, 1981.

99 Three species of (Dipt.) new to Norway, with a note on Ectinocera borealis Zetterstedt 1838

LITA GREVE AND RUDOLF ROZKO~NY

Greve, L. & Rozko~ny, R. 1981. Three species of Sciomyzidae (Dipt.) new to Norway, with a note on Ectinocera borealis Zetterstedt 1838. Fauna norv. Ser. B. 28,100-101.

The following species are reported new to Norway: Pherbellia rozkosnyi Verbeke, Pteromi­ cra angustipennis (Staeger) and Tetanocera ornatifrons Frey. Ectinocera borealis Zetterstedt is reported for the first time from Norway since 1846. Pherbellia sCUlellaris (von Roser) must be excluded from the Norwegian fauna.

Lita Greve, Museum of Zoology, University of Bergen, N-50 14 Bergen-Univ. Norway. Ru­ dolfRozkosny, Natural Science Faculty, J.E. Purkyn~ University, Kotlafskii 2, 61137 Brno, Czechoslovakia.

THE SPECIES the base of differently shaped male genitalia. (see Verbeke, 1967). Especially the posterior Ectinocera borealis Zett. 1838. surstyli are almost plane, beating a long slen­ A female was collected by Tore R. Nielsen on 16. der apex in P. rozkosnyi. So far the species is July 1966 at Vlirstigen, Oppdal, southern Trende­ known to occur in Denmark, Germany, Italy tag at 780 masl. The locality was mountainous and Czechoslovakia, recently it was also fo­ birch (Betula pubescens) forest near the old road und in southern Sweden and Finland. In through the forest. Knutson & Berg (I97 I) the species was mis-' E. borealis was described on material from southern Tmndelag in 1838. The locality was determined as P. scutellaris (von Rosed. P. Drivstuen ( = Drivstua) in the Drivdal valley eight scutellaris (von Rosed must be left out from or nine kilometers from the recent locality. Later the Norwegian list of Sciomyzidae species for the species was recorded from northern Tmnde­ the time being. Jag by Zetterstedt under the name Tetanocera angustipennis (Staeger, 1845). sciomyzina Zett. according to Dahl (J 965). Since One male was collected at Fauske, Fauske, Nord­ then, no material has been found (Knutson & land on 27. June 1956 by H. Andersson. The spe· Berg, 197 n Thus the reported specimen is the cimen is deposited at the Zoological Institute, first confirmation of this species from Norway in Lunds University, Sweden. The locality was a Er­ more than I30 years. iophorum bog. This is the same locality where R. E. borealis is distributed from northern­ Dahl collected,a specimen of P. glabricula (Fallen) most Fennoscandia southwards to the Alps on the same day, mentioned by Knutson & Berg (Schwitzerland, Italy and Austria). (1971). However, a misdetermination of the latter Pherbellia rozkosnyi Verbeke, 1967. specimen seems to be hardly possible because both authors knew results of a revision for the One male was collected by N. Knaben at Seim, Rozko~ny Granvin, inner Hordaland on I. June 1936. Anot­ genus Pteromicra (cf. and Knutson, her male was collected in Mlioodalen, Eidfjord, 1970) where P. angustipennis (Staeger) was resur­ inner Hordaland by Tore R. Nielsen on 6. July rected from the synonymy. Moreover, the true P. 1968 at 340 masI. The last locality was a stony glabricula is not too rare, but largely distributed hillside with profuse vegetation of grass, Gera­ all over Scandinavia so that its incidence together nium silvaticum. Valeriana sp., Filipendula ulma­ with P. angustipennis at Fauske is not sUrPrising. ria. Galium boreale. G. verum, Hieracium sp., Hy­ Another specimen of P. angustipennis. female, is pericum sp., Urtica dioica. Trifolium pratense, Vi­ present in the Zoological Museum in Helsingfors cia cracca. Lotus corniculatus with some groups taken at Fokstua, Dovre, northern Oppland by W. of decidious trees mainly Alnus incana. Betula Hellen. pubescens and Sorbus aucuparia. A third male is P. angustipennis is presented here as new to present in the Zoological Museum, Oslo labelled Norway. It is distributed in the Holarctic «T0ien 1845 S». region, ranging in the Palaearctic from Swe­ P. rozkosnyi was separated from the closely dish Lapland to France and Hungary, east­ related P. scutellaris (von Rosed chiefly on wards to Mongolia and the Kurile Islands.

100 Fauna norv. Ser. B 28, 100-101. Oslo 1981. '­ Tetanocera ornatijrons Fre)\ 1924. ACKNOWLEDGEMENTS One male, was caught on 13. July in Leirdalen, We want to thank Mr. Tore R. Nielsen for allo­ Lom, northern Oppland. The collector was Tore wing us to publish this material and for infor­ R. Nielsen. The locality was scattered mountai­ nous birch forest (Betula pubecens) with some mation on his localities. grass bogs in between. The locality was situated approximately at 1000 masl. Another specimen, a female, was caught in Finmark, near Gargia in REFERENCES Alta (UTM: 34 WEe 965468) on 2. July 1979 by Dah!, R., 1965. A revision of the family Sciomyzidae L. Greve. A sweep-net was used and the locality (Diptera, ) in J.W. Zetterstedt's collec­ was a mingled forest of Betula pubescens and tions. Opusc.ent.. 30, 153 -162. Pinus silvestris with some Juniper communis and Knutson, L.V. & Berg, CO., 1971. The malacopha­ Salix ssp. gous flies of Norway (Diptera. Sciomyzidae). T. ornatijrons is reported for the first time Norsk ent. Tidsskr.. 18, 119 -134. from Norway. It occurs in northern Sweden Rozko~ny, R., & Knutson, L.V., 1970. Taxonomy, and Finland, but also is found at severalloca­ biology and immature stages of Palaearctic Ptero­ lities on the Kola peninsula, in the USSR and micra, snail-killing Diptera (Sciomyzidae). Ann. western and eastern Sibiria. ent. Soc. Am., 63,1434-1459. Verbeke, J., 1967. Contribution a I'etude des dipteres Unless otherwise stated, specimens mentio­ malacophages. V. Trois especes palaearctiques ned in this article are deposited in Museum nouvelles du genre Pherbellia Robineau-Desvoidy of Zoology, University of Bergen. et quelques donnees sur l'identite de P. scutellaris (von Rosed (Dipt. SciomyzidaeJ. Inst. R. Sci. nal. Belg. Bull., 43 (J 8), 1-12.

Received March 8, 1981.

,

101 Recent records of and (Dipt.) from Norway

KNUT ROGNES

Rognes, K. 1981. Recent records of Tachinidae and Rhinophoridae (Dipt.) from Norway. Fauna norv. Ser. B. 28, 102-114.

Detailed records are given for 91 species of tachinid and rhinophorid flies, of which 31 seem previously not recorded from Norway. Phebellia glaucoides Herting, 1961 is reported from Scandinavia possibly for the first time.

Knut Rognes, Hav0rnbrautene 7a, N-4040 , Norway.

INTRODUCTION Below are given detailed records for 91 species space and partly because many records are unre­ of tachinid and rhinophorid flies recently captu­ liable and in need of revision. The presentation I red in Norway. Thirty-one species seem not to of the new faunistic data largely follows pre­ have been reported from the country previously, vious papers (e.g. Rognes 1980).1 a fact indicative of the level of attention the gro­ ups in question have received in the past. One species is probably new to Scandinavia. All spe­ SYSTEMATIC LIST cimens are in my private collection except for a TACHINIDAE few which are deposited in Museum of Zoology, Bergen (ZMB below) or in other private collec­ Subfamily Phasiinae tions. Identifications mostly follow the works Alophora pusilla (Meigen, 1824). of Stein (I 924), Lundbeck (I927), Mesnil Taxonomy: Draber-Monko 1965. (I 944-1975), Ringdahl (I945b), Emden (I 954) Previous records: Phasia pusilla Meig. - Siebke 1877: 92; Hyalomyia pusilla Meig. - Ringdahl and Herting (1961 b). Other works used are cited 1944b: 7; Parallophora pusilla Meig. - Ringdahl under each species. Some of my identifications 1952: 142-143 No. 230. have been verified by Benno Herting, Ludwigs­ Material: - Telemark: TEi: Hjartdal, Gvammen, burg, and Stig Andersen, Copenhagen, (RH.) EIS 26, I cl 12 July 1976, T. Nielsen, (SA). and (S.A,), respectively, below. The latter also The terminalia have been dissected and agree has identified some of the Siphona species. The with Draber-Monko's figures (1965: 95). nomenclature within the subfamilies Gymnosoma globosum (Fabricius, 1775) and Goniinae follows Mesnil (I944-1975). Material: - Aust-Agder: AAy: Trom0Y, Lovstad, Spelling of names, sequence of species treated EIS 6, I cl 31 July 1974, T. Nielsen, (SA). New to Norway. and their subfamilial arrangement largely follow Gymnosoma rotundatum (L.) Crosskey (I 975) and Pont (I 975). Non-British Taxonomy: Rohdendorf 1947; Mesnill952 x. species are inserted at the appropriate place. For Previous records: Gymnosoma rotundata Lin. ­ information on host relations of the early deve­ Siebke 1977: 91; Bidenkap 1892: 236; Gymno­ lopmental stages the reader is referred to Mesnil soma rotundatum L. - Ringdahl I944b: 7; Ring­ (I 944 -1975), Herting (I 960, 1961 b) and Cros­ dah11952: 142-143 No. 235. skey (I 977). For most species references are gi­ Material: - Telemark: TEi: Seljord, Seljord, EIS ven to recent taxonomic treatment, modern re­ 17, I cl 2 Q Q 26 July 1976, T. Nielsen, (SA). ­ visions, sources of listed synonyms, and sources Aust-Agder: AAi: Evje og Hornnes, Bjoni, EIS 5, 3 cl cl 19 June 1977, S. Svendsen; AAy: Ris0r, of other information of taxonomic or nomencla­ 0ysang, EIS 11, I Q 27 June 1975, T. Nielsen. ­ tural importance which I have been able to Vest-Agder: VAy: Kristiansand, Stangenes, EIS 2, trace. Next follow sources of previous records I Q 13 June 1976,1.R. Svendsen; Mandal, Skogs­ from Norway, including the names under fjord, I\IS 2, I Q 6 Aug. 1976, T. Nielsen. which they have been published. I have refrai­ Cylindromyia brassicaria (Fabricius, 1775) ned from citing the localities, partly to save Previous records: Ocyptera brassicaria Fabr.

102 Fauna norv. Ser. B 28, 102-114. Oslo 1981. Zetterstedt 1849: 3257; Siebke 1877: 91; Bidenkap Previous records: Trixa Iimbata Zetterstedt 1838: 1892: 236; Ringdahl 1944b: 7; Ringdahl 1952: 630; Siebke 1877: 90 (synonym according to 142-143 No. 243. Lundbeck 1927: 417-418 and Ringdahl 1945a: Material: - Akershus: AK: Brerum, Steinsh0gda, 30); Trixa a/pina Meig. - Ringdahl I944b: 10; EIS 28, I d 29 July 1977, (S.AJ. - : Trixa a/pina Zett. (sic) (author's name corrected B0: , Komnes, EIS 19, I d 6 Aut by Ringdahl 1953) - Ringdahl 1952: 142-143 1979; Hvittingfoss, EIS 19, I d 6 Aug. 1979. No. 225. Telemark: TEi: Sauherad, Nordagutu, EIS 18. Material: - Oppland: Os: Gausdal, Svatsum 600 I 9 26 July 1979, K. & A. & 0. & T. Rognes. m a.s.l., EIS 53, 2 d d I 9 5 July 1978, T. Niel~ Cy/indromyia interrupta (Meigen, 1824) sen. - Telemark: TEi: Kviteseid, Lislest01 (Vni­ Previous records: Ocyptera interrupta Meig. ­ vatn), EIS 17, 3 d d 24 June 1980; Skredi 400 m Ringdahl1944b: 7; Ringdahl1952: 142-143 No. a.s.l.,EIS 17, I d I 9 3 July 1980; Skredi 245. 400-750 m a.s.l., EIS 17, Id 3 July 1980. ­ Material: - Akershus: AK: Brerum, 0verland, Rogaland: Ry: , Eltravag, EIS 7. 3 d d 3 EIS 28, I 9 23 June 1979. - Oppland; On: Fron, June 1978, (S.AJ - M0re og Romsdal: MRi: Vinstra, EIS 62, I d 10 June 1979, S. Svendsen. Sunndal. Nes, EIS 85, I d 1-15 July 1964, R. - Telemark: TEi: Hjartdal, Gvammen, EIS 26. Mehl, priv. coil.. - Nordland: Nsi: Rana, Vir­ I d 12 July 1976, T. Nielsen, (SA). vassdalen, EIS 124, 2 d d 24 July 1978, (SA). Cy/indromyia pusilla (Meigen, 1824) Trixa variegata Meigen, 1824 Previous records: Ocypteru/a pusilla Meig. ­ Taxonomy: Herting 1972: 4. Ringdahl 1944b: 7; Ringdahl 1952: 142-143 No. Previous records: Trixa R.D. - Ring­ 246. dahl I944b: 10; Ringdahl 1952: 142-143 No. Material: - Telemark: TEi: Kviteseid, Kviteseid 224. gml kirke, EIS 17, I 9 25 June 1980; Skredtveit, Material: - Aust-Agder: AAi: Bygland, Longe­ EIS 17, Id 5 July 1980; Vnidal (Nordb0), EIS rak, EIS 9. Id 4 Aug. 1979, M. Aardal, 17, Id 6 July 1980. priv.coll.. - Vest-Agder: VAy: Flekkefjord, Hi­ Phania thoracica (Meigen, 1824) dra (Litjern), EIS 4, I d July 1979, A.J. Nilsen, Previous records: Phania thoracica Meig. - Si­ priv. coli.. - Hordaland: Hay: Bergen, Kobbelt­ ebke 1877: 90; Weberia thoracica Meig. - Ring­ veit (Isdalen), EIS 40, 3 d d 28 July 1980. dahl 1952: 142-143 No. 249. Material: - Vest-Agder: VAy: Kristiansand, Hamresanden, EIS 2, I d 25 July 1977, T. Niel­ sen. Subfamily Tachininae • The middle tiQiae have a v seta in apical half (cf. Voria ruralis (Fallen, 1810) Stein 1924; Lunabeck 1927: 118). Taxonomy: Crosskey 1973: 163-164 (Ieetotype designation); Mesnil 1974: 1262. Previous records: Tacltina ruralis Fall. - Siebke Subfamily Dexiinae 1877: 81; P/agia ruralis Fall. - Strand 1914: 324; Billaea triangu/(fera (Zetterstedt, 1844) Voria ruralis Fall. - Ringdahl I944b: 11; Ring­ Previous recor

103 nonymized curvinervis with trepida Meigen, 1824 149; Bidenkap 190 I: 55; Eriothrix rufomacu/atus (Herting 1962, 85), for which reason it is listed Deg. - Ringdahl 1944a: 81; Ringdahl 1944b: 11; under that name in his (1952) catalogue (see be­ Ringdahl 1952: 140-141 No. 198; Ard6 1957: low) 162. Material: - Vestfo1d: YE: Hof, Thorrud, EIS 28, Material: - Buskerud: B0: Kongsberg, Gran, EIS 49 9 28 July 1979. - Telemark: TEi: Sauherad, 18, 19 6 Aug. 1979. - Telemark: TEi: Notod­ Nordagutu, EIS 18, 19 26 July 1979. den, EIgsj0, EIS 27, I 9 15 July 1978, T. Nielsen. Athrycia trepida (Meigen, 1824) - Vest-Agder: VAy: Vennesla, Eikeland, EIS 5, Taxonomy: Herting 1973: 11; Mesnil 1974: 1265. 1a 21 Aug. 1975, S. Svendsen; Lindenes, J0r­ Previous records: Tachina subcincta Zetterstedt genstad, EIS 1,1 a 21 July 1977, T. Nielsen; I a 1844: 1026; Siebke 1877: 82; Voria trepida Meig. 20 July 1979, T. Nielsen; I a 16 July 1980, T. Ni­ - Ringdahl 1944a: 81; Ringdahl 1952: 138 -139 elsen. - Rogaland Ry: Lund, Moi, EIS 4, 1a 23 No. 146. Note that Ringdahl erroneously inclu­ June 1978; Refsland, EIS 3, 3 a a 17 July 1977; ded curvinervis Zett. within his concept of trepida Eigersund, Eigersund, EIS 3, I a 29 July 1979, (Ringdahl 1945a: 27, 29; Herting 1962: 85). L.M. Ramsland, priv.coll.; Klepp, 0ksnevad, EIS Material: - Telemark: TEi: Tokke, Dalen, EIS 7, I a 1 Aug. 1979, T. Nielsen; 19 1 Sept. 1979; 16, 19 I July 1980. Sandnes, Bogafjell, EIS 7, 19 26. Aug. 1979; Ramonda spathu/ata (Fallen, 1820) Sola, Sola, EIS 7, I a 29 9 25 Aug. 1979; Sta­ Taxonomy: Mesnil 1974: 1300. vanger, Krossberg, EIS 7, I a 15 July 1977; 19 Material: - Vest-Agder: VAy: Lindesnes, J0r­ 13 Aug. 1977; 19 9 Aug. 1978; 3 a a 2 Sept. genstad, EIS 1, 1 a 18 July 1980, T. Nielsen. ­ 1979; Revheim, EIS 7,19 11 Aug. 1977;Sunde, Rogaland: Ry: Stavanger, Mad1alia, EIS 7, I a 24 EIS 7, 89 9 7 Aug. 1978, (SA); I a 17 July May 1980; Godalen, EIS 7, 1a 1 June 1980. 1979; I 9 I -15 Aug. 1979; 1a 12 July 1980; • New to Norway. Tjensvoll, EIS 7, 1a 13 JUly 1977, (SA); 19 10 Aphe/ogaster a/pina (Villeneuve, 1910) Aug. 1977; I a II Aug. 1977; Ullandhaug, EIS Taxonomy: Mesnil 1974: 1291. 7, 4a a 8 Aug. 1978; 5a a 2 99 10 Aug. Material: - Oppland: Os: Ringebu, Ringebu stav­ 1978; I 9 26 Aug. 1979; Randaberg, B0raunen, kirke, EIS 63, 1a 9 July 1978, T. Nielsen, (S.AJ EIS 7, I 9 3 Sept. 1977. - Hordaland: HOy: Ber­ New to Norway. gen (Fana), Storetveit, EIS 30, 1a 19 18 July Phyllomya vo/vulus (Fabricius, 1794) 1980; Radal - Stend, EIS 30, 19 25 July 1980. Taxonomy: Mesnil 1975: 1353. - M0re og Romsdal: MRi: Rauma, Lerheim, EIS • Previous records: Musca volvu/us Fabr. - Zetter­ 77, 19 29 July 1978. stedt 1838: 653; Dexia volvulus Fabr. - Zetter­ Campy/ocheta praecox (Meigen, 1824) stedt 1844: 1272; Siebke 1877: 93; Phyllomyia vo/­ Taxonomy: Mesnil 1974: 1254. vu/us Fabr. - Ringdahl I944a: 81; Ringdahl Material: - Vest-Agder: VAy: Flekkefjord, Ras­ I944b: 11; Ringdahl1952: 138-139 No. 171. vag, EIS 4, I a 10-11 May 1979. - Rogaland: Material: - Akershus: AK: , Eidsvoll, Ry: Stavanger, Ullandhaug, EIS 7, I a 18 April EIS 36, 19 4 Aug. 1979. - Buskerud: B0: Hu­ 1979. rum, Kongsdelene, EIS 28, 19 2 Aug. 1979; New to Norway. , Drammen, EIS 28, I 9 31 July 1979; The/aira nigripes (Fabricius, 1794) 0vre Eiker, Burud, EIS 27, 29 9 I Aug. 1979; Taxonomy: Mesnil 1975: 1339. Mesnil (1975: Kongsberg, Hvittingfoss, EIS 19, Id 6 Aug. 13370 has recently shown that three species, viz. 1979; Komnes, EIS 19, 19 6 Aug. 1979; Kongs­ /eucozona Panzer, 1809, nigripes Fabricius, 1794 berg, EIS 27, 1a 28 July 1979. - Telemark: and solivaga Harris, 1776, exist within the genus TEi: B0, B0, EIS 18, 3 9 9 25 July 1979; 0vreb0, The/aira Robineau-Desvoidy, 1830 in Europe in­ EIS 17, 19 25 July 1979; 3 a a 6 July 1980; stead of one as previously believed. Hjartdal, 0veroo, EIS 26, I a 12 July 1976, T. Previous records: Dexia /eucozona Panz. - Si­ Nielsen, (S.A.l; Seljord, Ulvenes, EIS 17, 1a 19 ebke 1877: 93; The/aira /eucozona Panz. - Bi­ 25 July 1979; Fyresdal, Aslestad - Slyst0yl 650 denkap 1892: 237; The/aira nigripes Fabr. ­ m a.s.l., EIS 16, 3 a a 7 July 1980. - Horda­ Ringdahl 1944b: 11; Ringdah1 1952: 140 - 141 land: HOy: Bergen (Fana), Fantoft, EIS 30, 19 27 No. 207. According to Mesnil (I.c.)the true /eu­ July 1980; Fj0sanger, EIS 30, 4 a a 19 22 July cozona is a more southern species in Europe. I 1980; Gf0nnest0len, EIS 30, 39 9 19 July 1980. therefore assume that nigripes is the species refer­ Eriothrix rufomacu/atus (Degeer, 1776) red to by Siebke and Bidenkap. It is possible that Taxonomy: Mesnil 1975: 1322. solivaga also will be found among the above re­ Previous records: Tachina /ateralis Fall. (sic) ­ cords (see below). Siebke 1877: 88. Siebke refers to Zetterstedt 1844: Material: - Telemark: TEy: Bamble, Bamb1e, 1188 where /ateralis Fallen, 1810 = lateralis Fa­ EIS 11, 3 a a 27 July 1979,0. Rognes. - Roga­ bricius, 1781 is treated (cf. also Lundbeck 1927: land: Ry: Sandnes, Lura, EIS 7, 2 a a 19 July 525). T. lateralis Fallen, 1816 = nigripes Fabri­ 1979. cius, 1794 (The/aira), however (cf. Mesnil 1975: The/aira solivaga (Harris, 1776) 1339); Olivieria lateralis Fabr. - Bidenkap 1898: Taxonomy: Mesnil 1975: 1340. 104 '" Material: - Telemark: TEi: Kviteseid, Kvitsund The record from 0vre Alta (69°55'N) is probably skole, EIS 17, I 0 29 June 1980; Kviteseid, EIS the northernmost from Scandinavia. 17, I 9 29 June 1980. Zophomyia temu/a (Scopoli, 1763) New to Norway. Mesnil (J.c.J reports the species Taxonomy: Mesnil 1972: 1083. from «Skandinavien». It is not listed by Hackman Previous records: Tachina tremu/a Lin. - Siebke (1980) from Finland. 1877: 87; Zophomyia temu/a Scop. - Ringdahl My specimens have I or 2 pd setae on the middle 1952: 138-139 No. 170. tibiae, and in the male the hind fifth or sixth of the Material: - 0stfold: 0: Fredrikstad, 0ra, EIS 20, fourth tergite is black as in the British specimens 20 0 24 June 1979. - Vest-Agder: VAy: Lin­ cited by Mesnil (I.c.J. The basal bands of dusting desnes, J0rgenstad, EIS 1, I 0 I 9 20 July 1979, on the intermediate abdominal segments of the T. Nielsen. female are not interrupted at the midline, i.e. op­ Pseudopachysty/um goniaeoides (Zetterstedt, 1838) posite to the condition in my (Danish) female spe­ Taxonomy: Mesnil 1972: 1088. cimens of T. nigripes. Material: - Buskerud: B0: Ringerike, L0vlia 570 Dexiosoma caninum (Fabricius, 1781) m a.s.l., EIS 36, I 9 31 July 1977, (SA); F1es­ Taxonomy: Mesnil 1974: 1233. berg, Belgen, EIS 27, 10 27 July 1979. - Tele­ Previous records: Dexia canina Fabr. - Zetter­ mark: TEi: Fyresdal, Aslestad - SIyst0yl 650 m stedt 1849: 3263; Siebke 1877: 92; Bidenkap a.s.l., EIS 16, I 9 7 July 1980. 1892: 237; Dexiosoma caninum Fabr. - Ringdahl New to Norway. 1944b: 9; Riagdah11952: 140-141 No. 21\. aurea (Fallen, 1820) Material: - Akershus: AK: Brerum, Fossum, EIS Taxonomy: Mesnil 1973: 1129. 28, I 0 3 Aug. 1979; Nordby gard, EIS 28, I 9 3 Previous records: Tachina aurea Fall. - Siebke Aug. 1979. - Buskerud: B0: Drammen, Dram­ 1877: 89; Leskia aurea Fall. - Ringdahl 1952: men, EIS 28, 90 0 49 9 31 July 1979; 0vre Ei­ 140-141 No. 206. ker, Burud, EIS 27, 10 19 I Aug. 1979. - Te­ Material: - Telemark: TEi: Seljord, Seljord, EIS lemark: Tokke, Dalen, EIS 16, 20 0 30 June 17, 19 26 July 1976, T. Nielsen, (S.AJ 1980; 20 0 I 9 I July 1980. - Vest-Agder: Lypha dubia (Fallen, 1810) VAy: Lindesnes, Gasestein, EIS I, I 9 22 July Taxonomy: Mesnil 1971: 1000. 1980, T. Nielsen. - Hordaland: HOy: Bergen Previous records: Tachina dubia Fall. - Siebke (Fana), Trollhaugen, EIS 30, I 9 21 July 1980; 1877: 85; /:'xorista dubia Fall. - Bidenkap 1898: Fj0sanger, EIS 30, 10 22 July 1980; Fantoft, EIS 149; Lypha dubia Fall. - Ringdahl I944a: 80; 30,200 19 24 July 1980; 200 29927 July Ringdahl I944b: 10; Ringdahl 1952: 136-137 1980; Stend (Mj0Ikevika), EIS 30, 20 0 25 July No. 90; Tachina umbrinervis Zetterstedt, 1844: • 1980. 1146 (described from a female specimen taken at C/eonice callida (Meigen, 1824) Nord-Tf0ndelag: NTi: Verdal, Games 30 June Taxonomy: Ringdahl 1934: 270; Ringdahl 1945a: 1840); Siebke 1877: 86. 29; Mesnil 1972: 1090. Material: - Vest-Agder: VAy: Kristiansand, Previous records: Tachina rotundicornis Zetter­ Stangenes, EIS 2, 20 0 3 May 1980, S. Svend­ stedt 1838: 641 (described from a female speci­ sen; I 0 I 9 4 May 1980, S. Svendsen; I 0 6 men taken by,Boheman in the Dovre mountains, May 1980, S. Svendsen; F1ekkefjord, Rasvag (Hi­ Norway); Zetterstedt 1844: 1156; Siebke 1877: dra), EIS 4, 260 0 29 9 I 0 I 9 (in cop.) 4 87; Steiniella eallida Meig. - Ringdahl I944a: May 1979; 500 19 10-11 May 1979. - Ro­ 81; Ringdahl 1944b: 11; Ringdahl 1952: galand: Ry: Stavanger, Byhaugen, EIS 7, 40 0 138-139 No. 167. 29 April 1978; 39 9 9 June 1979; Godalen, EIS Material: - Telemark: TEi: Kviteseid, Skredi, 7, I 9 I June 1980; I 9 3 June 1980; Krossberg, EIS 17, 19 22 June 1980, (B.HJ EIS 7, 19 24 April 1977; 10 30 April 1977; The specimen has only one lateral scutellar seta 800 I May 1977,(SA); 19 14 May 1978; 10 on each side, i.e. a total of 3 pairs of marginal scu­ I 9 27 May 1978, (SA); 20 0 24 April 1979; tellar setae. It was caught on leaves of Popu/us 40 0 29 9 28 April 1979; 10 0 0 I May 1979; tremu/a. 30 0 39 9 24 May 1979; 40 0 I May 1980; Pe/atachina tibialis (Fallen, 1810) 5 0 0 29 9 1I May 1980; Ullandhaug, EIS 7, Taxonomy: Mesnil 1973: 1113. I 0 18 April 1979; I 0 I May 1979; 40 0 20 Previous records: Tachina tibialis Fall. - Siebke April 1980; I 0 24 April 1980; I 9 22 May 1980. 1877: 82; Pe/atachina tibialis Fall. - Ringdahl Lypha ru.fleauda (Zetterstedt, 1838) 1944a: 81; Ringdahl 1944b: 11; Ringdahl 1952: Taxonomy: Mesnil 1971: lOO\. 138-139 No. 169. Previous records: Tachina rufieauda Zetterstedt Material: - Akershus: AK: Bcerum, Steinsh0gda, 1838: 643; Zetterstedt 1844: 1111; Siebke 1877: EIS 28 19 25 June 1979, A. & K. Rognes. - Ro­ 85; Eversmannia rufieauda Zett. - Ringdahl galand: Ry: Stavanger, Byhaugen, EIS 7, 1 9 9 I944a: 80; Ringdahl I944b: 10; Micronychia rufi­ June 1979; Tjensvoll, EIS 7, 19 19 June 1979. ­ cauda Zett. - Ringdahl 1952: 136-137 No. 91; Finnmark: Fi: Alta, 0vre Alta, EIS 173, I 9 20 Tachina maculipennis Zetterstedt, 1844: 1092 July 1978, (S.AJ (described from a single male specimen captured 105 «ad diversorium Thymes prope urbem Levanger 2 d d I 9 28 July 1979. - Telemark: TEi: Sau­ in paroecia Skogn» (Nord-Tr0ndelag: NTi: Le­ herad, Nordagutu, EIS 18, 19 26 July 1979. vanger) 4 July 1840); Siebke 1877: 84. Eurithia vivida (Zetterstedt, 1838) Material: - Hordaland: HOi: Eidfjord, Mflbooa­ Taxonomy: Mesnil 1972: 1066. len 270 m a.s.l., EIS 33?, I d 19, K.E. J0rstad et Previous records: Tachina vivida Zett. - Siebke aI., 2MB. 1877: 84; Nemoraea vivida Zett. - Bidenkap The specimens are on a single pin. In the female, 1898: 149; Emeslia vivida Zett. - Ringdahl which is lacking the abdomen, the prosternum 1944b: 11; Ringdahl 1952: 138-139 No. 162. has three or four hairs on each side. Material: - Telemark: TEi: Seljord, Ulvenes, EIS Linnaemya vu/pina (Fallen, 1810) 17, 19 25 July 1979. Taxonomy: Herting 1961: 1Of; Mesnil I971: The hosts of this species are unknown (Herting 1016. 1960: 113; Mesnil 1972: 1067). Siebke (j .c.J re­ Previous records: Tachina vu/pina Fall. - Siebke ports to have bred his specimens from Arclia caja 1877: 84; Micropa/pus vu/pinus Fall. - Ringdahl L. (Lep., Arctiidae) «

106 July-Aug. 1979, S. Bnlten, priv. coil.. - Horda­ Siphona flaviff'ons (Staeger in Zetterstedt, 1849) land: HOy: Bergen, Ask0y, EIS 30, I 9 July Taxonomy: Mesnil 1965: 865. 1896, 0.1. Lie-Pettersen, 2MB. Material: - Aust-Agder: AAy: Birkenes, Amli, Tachina vernalis (Robineau-Uesvoidy, 1830) EIS 6, I d 25 July 1976, T. Nielsen, (S. Andersen Taxonomy: Mesnil 1966: 924; Herting 1974: 3; deU. Crosskey 1975: 99; Mesnil 1975: 1400. There is Stig Andersen, Copenhagen, (j .c.) also reports to some disagreement as to whether vernalis or have seen material from this country. magnicornis Zetterstedt, 1844 is the correct name New to Norway. for this species. I follow the latest opinion of Mes­ Siphona geniculata (Oegeer, 1776) nil cited above. Taxonomy: Mesnil 1965: 866. Previous records: Echinomyia magnicornis Zett. Previous records: Siphona geniculata OeGeer ­ - Strand 1914: 324; Ringdahl 1952: 136-137 Zetterstedt 1838: 620; Zetterstedt 1844: 989; Si­ No. 101. ebke 1877: 80; Bidenkap 1901: 54; Bucentes geni­ Material: - Akershus: AK: Brerum, Steinsh0gda, culata Oeg. - Ringdahl 1952: 138-139 No. EIS 28, 8 d d 29 July 1977, (SA). - O,pland: 139. On: Fron, Kvam, EIS 62, I d 22 July 1979, S. Material: - Rogaland: Ry: Stavanger, Byhaugen, Svendsen. - Buskerud: B0: 0vre Eiker, Burud, EIS 7, 129 9 6 Sept. 1980, JS.A.l; Krossberg, EIS 27, I d I Aug. 1979; Kongsberg, Eftel0t, EIS EIS 7, I 9 I Det. 1978, (SA); 39 9 19 Aug. 27, I 9 6 Aug. 1979; Gran, EIS 18, I 9 6 Aug. 1980, (S.A.); Sunde (indoors), EIS 7, 19 10 Sept. 1979; Hvittingfoss, EIS 19, I 9 6 Aug. 1979. ­ 1980, (S.A.); Ullandhaug, EIS 7, I d 5 June 1980, Vestfold: YE: Hof, Thorrud, EIS 28, 19 28 July (S. Andersen deU I d 3 9 9 7 Sept. 1980, (SA). 1979. - Telemark: TEi: Sauherad, Nordagutu, Two of the females from Byhaugen lack the med­ EIS 18, 3 d d 26 July 1979; B0, 0vreb0, EIS 17, ian marginal setae of second tergite. I d 25 July 1979; Seljord, Seljord, EIS 17, I d 26 Siphona setosa Mesnil, 1960 July 1976, T. Nielsen; Ulvenes, EIS 17, 4 d d Taxonomy: Mesnil 1965: 876. I 9 25 July 1979. - Aust-Agder: AAy: Grim­ Material: - Telemark: TEi: Tokke, Amdalsverk, stad, Metveit, EIS 6, 29 9 25 July 1976, T. Niel­ EIS 16, I 9 I July 1980, (SA). sen, (S.AJ AAi: Amli, Bjorevja, EIS 10, I d 1I Stig Andersen, Copenhagen, (J .c.J also reports to July 1980, T. & K. Rognes. have seen material from this country. None of my male specimens have 2 prodinate or­ New to Norway. bital setae, most have only one, a few none at all Belida angelicae (Meigen. 1824) (cf. Mesnil 1966: 925). The claws of the front tarsi Taxonomy: Lundbeck 1927: 296, 297; Ringdahl are much longer than the last tarsal segment. 1945a: 28; Mesnil 1962: 749; Herting 1974: 33; •Triarthria spinipennis (Meigen, 1824) Mesnil 1975: 1398. Taxonomy: Mesnil 1973: 1215; Crosskey 1974; Previous records: Tachina fittilis Zett. - Siebke Mesnil 1975: 1401. 1877: 82 (synonym according to Lundbeck and Material: - Telemark: TEi: Vinje, RauJand 700 Ringdahl, not mentioned by Mesnil); Sarcophaga m a.s.l., EIS 25, Id 19 June 1977, (SA). ­ albiceps Meig. - Bidenkap 1901: 58 (misidentifi­ Vest-Agder: VAy: Flekkefjord, Rasvag (Hidra), cation revised by Ringdahl I944b); Lydella ange­ EIS 4, 2 d d ,24 June 1978, (SA). lime Meig. - Ringdahl I944a: 80; Ringdahl The specimen from Rauland lacks the prst ia on 1944b: 9; Aporotachina angelicae Meig. - Ring­ both sides. It has 3 + 3 de. however, and the pat­ dah11952: 132-133 No. 11. tern of pollinosity before the suture is as described Material: - Akershus: AK: Brerum, Steinsh0gda, by Emden (1954). New to Norway. EIS 28, I d 29 July 1977, (S.AJ - Buskerud: B0: Ringerike, L0vlia 570 m a.s.l., EIS 36, I d 31 July 1977, (S.AJ - Telemark: TEi: Kviteseid, Subfamily Goniinae Skredtveit, EIS 17, 19 23 June 1980; Skredi, EIS 17, 19 8 July 1980. Actia pilipennis (Fallen, 1810) Blondelia nigripes (Fallen, 1820) Taxonomy: Mesnil 1963: 826; Crosskey 1974: Taxonomy: Mesnil 1962: 755. 283 - 284, 302 - 303 (Iectotype designation). Previous records: Tachina nigripes Fall. - Zetter­ Material: - Hordaland: HOy: Bergen (Fana), Fj0­ stedt 1838: 636-637; Zetterstedt 1844: 1059; Si­ sanger, EIS 30, I d 28 July 1980. ebke 1877: 82; Blondelia nigripes Fall. - Ring­ New to Norway. dahl 1952: 132-133 No. 7. Siphona con/usa Mesnil, 1961 Material: - Telemark: TEi: Seljord, Ulvenes, EIS Taxonomy: Mesnil 1964: 859. 17, 13d d 299 25 July 1979. Material: - Telemark: TEi: Tokke, Oalen, EIS All the specimens were captured on Angelica flo­ 16, I 9 30 June 1980, (SA). wers. Stig Andersen, Copenhagen, (in litt. 14 Det. 1980) Medina collaris (Fallen, 1820) also reports to have seen material from this coun­ Taxonomy: Mesnil 1962: 727. try. Previous records: Tachina collaris Fall. - Zetter­ New to Norway. stedt 1838: 637; Zetterstedt 1844: 1072; Siebke

107 1877: 83; Degeeria col/aris Fall. - Ringdahl seid, Skredi, EIS 17, I Q 8 July 1980; Tokke, Da­ I944b: 11; Medina col/aris Fall. - Ringdahl len, EIS 16, I Q I July 1980. - Vest-Agder: 1952: 136-137 No. 117. VAy: F1ekkefjord, Kvellandstrand, EIS 4, 4 cl cl Material: - Telemark: TEi: Sauherad, Norda­ I Q 3 June 1979; Rasvag (Hidra), EIS 4, 13 cl cl 4 gutu, EIS 18, I Q 26 July 1979; Seljord, Ulvenes, June 1979. - Sogn og Fjordane: SFi: Stryn, Inn­ EIS 17, I cl 6 July 1980; Kviteseid, Kviteseid, EIS vik, EIS 68, I Q 30 July 1978, (SA). 17, I Q 29 June 1980. - Rogaland: Ry: Sta­ Exoris/a rus/ica (Fallen, 1810) vanger, Ullandhaug, EIS 7, IQ 10 June 1979. Taxonomy: Mesnil 1960: 569; Mesnil 1975: 1396. Medina separa/a (Meigen, 1824) For treatment of related species see Herting 1967: Taxonomy: Herting 1966: 2; Herting 1971: 4; 2f; Mesnil 1970x: 112-115; Herting 1971: I; Herting 1972: 12. Herting 1972: 10, 14; Herting 1975b: 1. Material: - Vest-Agder: VAy: Lindesnes, Gases­ Previous records: Tachina rus/ica Fall. - Ring­ tein, EIS I, I Q 22 July 1980, T. Nielsen. - Hor­ dahl I944a: 80; Ringdahl I 944b: 10; Ringdahl daland: HOy: Bergen (Fana), Fantoft, EIS 30, I cl 1952: 134-135 No. 73. 27 July 1980; Fj0sanger, EIS 30, 5 Q Q 22 July Material: - Akershus: AK: Bcerum, 0verland, 1980; Gf0nnest0Ien, EIS 30, I cl 19 July 1980; EIS 28, I cl 23 June 1979. Stend (Mj0Ikevika), EIS 30, I Q 25 July 1980. All The terminalia and fifth sternite have been exami­ the specimens from Hordaland have been exami­ ned. ned by B. Herting. Phorocera obscura (Fallen, 1810) In Scandinavia this species has been reported Taxonomy: Emden 1954: 73, foot-note (Jectotype from Sourthern Sweden (Ringdahl leg.) by Her­ designation); Mesnil 1960: 641; Crosskey 1974: ting (1971). Hackman (1980) does not report it 30!. from Finland. Previous records: Phorocera obscura Fall. -~ New to Norway. Ringdahl 1952: 134-135 No. 78. assimi/is Fall. Meigenia mu/abi/is (Fallen, 1810) of Siebke 1877: 89 and Bidenkai> 1898: 149 may Taxonomy: Mesnil 1962: 707; Ringdahl 1945a: contain some obscura. 27, 29; Mesnil 1967x: 44; Herting 1972: 4; Mesnil Material: - Vest-Agder: VAy: F1ekkefjord, Kvel­ 1975: 1398. landstrand, EIS 4, I cl I Q 3 June 1979. - Roga­ Previous records: Tachina mu/abi/is Fall. - Zet­ land: Ry: Sola, Haga (Hansaberget), EIS 7, I cl 7 terstedt 1844: 1059-1061 (var. b); Siebke 1877: June 1980. 82 - 83; Meigenia mu/abi/is Fall. - Ringdahl Timavia amoena (Meigen, 1824) I944a: 80; Ringdahl 1944b: 9; Ringdahl 1952: Taxonomy: Mesni11949: 74; Mesnil1963x: 4-5; 132 -133 No. I; Meigenia bisigna/a Meig. - Bi­ Herting 1974: 8; Mesnill975: 1389. denkap 1898: 149; Tachina disc%r Zett. - Si­ Previous records: Win/hemia amoena Meig. ­ ebke 1877: 83 (synonym according to Ringdahl Ringdahl 1944b: 10; Chae/o/yga amoena Meig. ­ 1945a: 27 and Mesnil 1967x: 44); Tachina pyg­ Ringdahl 1952: 134-135 No. 64. maea Zetterstedt 1838: 641 (synonym according Material: - Aust-Agder: AAi: Evje og Hornnes, to Zetterstedt 1844: 1060 and Ringdahl 1945a: 29). Hornnes, EIS 5, I Q 28 June 1978, (S.AJ - Ro­ Material: - Akershus: AK: Bcerum, Nordby galand: Ry: Sandnes, St0lsvik, EIS 7, I cl 11 June gard, EIS 28, I cl 21 June 1979; Steinsh0gda, EIS 1980. 28, I cl 29 July 1977, (SA); Oslo, Bygd0Y, EIS Win/hemia ery/hrura (Meigen, 1838) 28, I cl 25 June 1979, A. & K. Rognes; Sogns­ Taxonomy: Stein 1924: 60; Mesnil 1949: 91. vatn, EIS 28, 2 cl cl 21 June 1979. - Buskerud: Material: - 0stfold: 0: Fredrikstad, 0ra, EIS 20, B0: 0vre Eiker, Burud, EIS 27, I cl I Aug. 1979; 2 cl cl 24 June 1979, A. Rognes. - Akershus: Ulleland, EIS 28, I Q 5 Aug. 1979; Kongsberg, AK: Bcerum, 0verland, EIS 28, 4 cl cl 23 June Kongsberg, EIS 27, I cl 28 July 1979. - Vest­ 1979. fold: YE: Hof,Thorrud, EIS 28, 1 cl 28 July B. Herting has seen all my specimens. He expres­ 1979. - Telemark: TEi: Tokke, Dalen, EIS 16, ses some doubt as to the status of this species, but I cl 30 June 1980; LArdal, EIS 17, 2 Q Q 4 July says that «at least your smaller specimens with 1980. - Hordaland: HOy: Bergen, Svartediket, darker mesonotum are determinable as ery/hru­ EIS 39, 2 cl cl 28 July 1980; Stend, EIS 30, 2 cl cl ra» (Herting in litt. 18 Sept. 1980). He is then re­ 25 July 1980. - Sogn og Fjordane: SFi: Stryn, In­ ferring to 3 small males from 0verland. nvik, EIS 68, 2 cl cl 30 July 1978, (SA). Important characteristics of my specimens are: Oswa/dia muscaria (Fallen, 1810) Frons at narrowest point 0.225-0.246 (mean Taxonomy: Mesnil 1962: 763. 0.237) of head width,anterior two thirds of para­ Previous records: Degeeria muscaria Fall. frontals and upper two thirds of parafacials with Sch0yen 1889: 7; Oswa/dia muscaria Fall. golden yellow pollinosity, uppermost parts of oc­ Ringdahl1952: 136-137No.113;Herting 1962: ciput with yellow or yellow-white hairs. Third 81. Vibrissina sordidisquama Zett. - Ringdahl tergite with 4-6 marginal setae. Fifth tergite on I944a: 80. ventral side of abdomen with a black basal band Material: - Akershus: AK: Oslo, S0rkedalen, EIS occupying one fourth to one half of the length of 36, I Q 22 June 1979. - Telemark: TEi: Kvite­ the tergite. Fifth sternite with black lobes. Other­

108 wise the abdomen has much less yellow colour Cyzenis albicans (Fallen, 1810) than my specimens of W. quadripustulata (see be­ Taxonomy: Mesnil 1954: 338. low). Previous records: Tachina albicans Fall. - Si­ Hackman (1980) reports erythrura from Finland. ebke 1877: 86; Cyzenis albicans Fall. - Ringdahl I have not been able to trace further records from 1944b: 9; Ringdahl 1952: 132-133 No. 5. .Scandinavia. Material: - Vest-Agder: VAy: Flekkefjord, Ras­ New to Norway. vag, EIS 4, 2d d 4 May 1979; 14d d 10-11 Winthernia quadripustulata (Fabricius, 1794) May 1979. - Rogaland: Ry: Sandnes, Dale, EIS Taxonomy: Mesnil 1949: 93. 7, 4 d d 1 Q 15 May 1980; Sola, Haga (Hansa­ Previous records: Tachina 4-pustulata Fabr. berget), EIS 7, 5 Q Q' 7 June 1980; Stavanger, By­ Zetterstedt 1838: 644; Zetterstedt 1844: 1104; Zet­ haugen, EIS 7, 3 Q Q 9 June 1979; Madlalia, EIS terstedt 1849: 3243; Siebke 1877: 85; Nernoraea 7, 1 Q 24 May 1980; Mosvannsparken, EIS 7, 4-pustulata Fabr. - Bidenkap 1898: 149; Wint­ 1 d 5 May 1980, Students leg. hernia quadripustulata Fabr. - Ringdahl 1944a: Frontina laeta (Meigen, 1824) 80; Ringdahl 1944b: 10; Ringdahl 1952: Taxonomy: Mesnil 1954: 343. 134-135 No. 60. Previous records: Tachina laetabilis Zett. - Si­ Material: - Buskerud: B0: 0vre Eiker, Burud, ebke 1877: 88; Frontina laeta Meig. - Ringdahl EIS 27,1 d 1 Aug. 1979. - Vestfold: YE: Hof, 1952: 134-135 No. 50. Thorrud, EIS 28, 3 d d 28 July 1979. - Tele­ Material: - Telemark: TEi: Seljord, Seljord, EIS mark: TEi: Seljord, To, EIS 17, 1d 24 July 1979. 17, 1 d 26 July 1976, T. Nielsen, (S.AJ; Ulvenes, My specimens differ from the erythrura speci­ EIS 17, 3 d d 25 July 1979. mens cited above in the following characters: Gonia ornata Meigen, 1826. Frons at narrowest 0.197 -0.215 (mean 0.205) of Taxonomy: Mesnil 1956: 527; Herting 1963: head width, parafrontals and parafacials with 106-107. pure white pollinosity, occiput with white hairs Previous records: Gonia lateralis Zell. - Siebke (one specimen with slightly yellowish hairs in the 1877: 89 (synonym according to Lundbeck 1927: uppermost part). Third tergite with 2- 3 marginal 393); Gonia ornata Meig. - Ringdahl 1944b: 10; setae (one specimen with 6). Fifth tergite on ven­ Ringdahl 1952: 134-135 No. 81. tral side of abdomen yellow. Fifth sternite with Material: - Rogaland: Ry: Klepp, Boresanden, yellow lobes. EIS 7, 6 d d 1 Q 20 May 1979; 11 d d 1 May Erycilla ferruginea (Meigen, 1824) 1980; Orresanden, EIS 7, 10 d d 1 Q 1 May Taxonomy: Mesnil 1955: 440 (as Erycina rutila 1978, (S.AJ; 1d 6 May 1978, (S.AJ; Selesanden, MeigJ; Mesnil 1957x: 20, foot-note; Herting EIS 7, 1 Q 20 May 1978, (S.A.); Sandnes, Mykle­ 1968: 1-3; Herting 1971: 8; Herting 1972: 6, 12; bostad, EIS 7, 1 d 26 May 1979; 1 Q 26 May Mesnil 1975: 1394. 1979, A. Vignes; Randaberg, Sandestranden, EIS Previous records: Cerotnasia ferruginea Meig. ­ 7, 1 Q 1 June 1978, (S.AJ. Ringdahl 1954: 49. All specimens except the two from Sandnes were Material: - Hordaland: HOy: Bergen (Fana), Fj0­ caught on or behind the large sand dunes borde­ sanger, EIS 30, 1 Q 22 July 1980, (B.HJ ring the coast along the northern part of Jreren. Aplon1ya confinis (Fallen, 1820) Gonia picea (Robineau-Desvoidy, 1830) Taxonomy: Mesnil 1954: 330; Herting 1974: 12. Taxonomy: Mesnil 1956: 528 (as sicula); Herting Previous records: Tachina confinis Fall. - Zetter­ 1974: 20; Herting 1975a: 4. stedt 1838: 644; Zetterstedt 1844: 1141; Siebke Previous records: Gonia fasciata Meig. - Siebke 1877: 86; Exorista confinis Fall. - Ringdahl 1877: 89; Ringdahl 1952: 134-135 No. 82. 1952: 132-133 No. 24. Material: - Rogaland: Ry: Sandnes, Foss-Eike­ Material: - Buskerud: B0: 0vre Eiker, Burud, land, EIS 7, 1d 28 May 1979, T. Figved; Sta­ EIS 27, Id 1 Aug. 1979. vanger, Krossberg, EIS 7, 1 d 19 May 1979, T. Bothria subalpina Villeneuve, 1910 Nyb0; Stokkavatn, EIS 7, 1 Q 12 May 1980, L.G. Taxonomy: Mesnil 1954: 336. Waage; Vindafjord, Vormestrand, EIS 14, 1 d 1 Material: - Vest-Agder: VAy: Vennesla, Eike­ May 1980, L.G. Waage. land, EIS 5, 1 d 12 April 1976, S. Svendsen. Onychogonia fZaviceps (Zetterstedt, 1838) Previously recorded from Sweden (Ringdahl Taxonomy: Mesnil 1956: 539 (as interrupta); Her­ 1937, 1952; Herting 1962) and Finland (Hackman ting 1973: 7-9; Mesnil 1975: 1395; Herting 1980) as far as Scandinavia is concerned. 1975a: 10. New to Norway. Previous records: Gonia flaviceps Zetterstedt, Brachicheta strigata (Meigen, 1824) 1838: 632 (described from a single male captured Taxonomy: Mesnil 1956: 499. between Kautokeino and Alta in Finnmark, Nort­ Material: - Rogaland: Ry: Klepp, Orresanden, hern Norway, 14 Aug. 1821) (Mesnil (1956: 542) EIS 7, 1 d 1 May 1978, (S.AJ erroneously refers the type locality to Sweden); Found on dry open heath-land behind sand dunes Zetterstedt 1844: 1196; Siebke 1877: 90; Bidenkap along coast of Jreren. 1901: 57. New to Norway. Material: - Oppland: Os: Gausdal, Svatsum 600

109 m a.s.\., EIS 53, I d 5 July 1978, T. Nielsen, Material: - Rogaland: Ry: Stavanger, Ulland­ (SA). haug, EIS 7, I d I Q 10 Aug. 1978, (SA). The abdomen is all black without red lateral mar­ Cadurciella tritaeniata (Rondani, 1859) kings. Taxonomy: Mesnil 1955: 426; Herting 1969: 202. Myxexoristops blondeli (Robineau-Desvoidy, 1830) Material: - Telemark: TEi: Sauherad, Norda­ Taxonomy: Mesnil 1955: 446; Herting 1964: gutu, EIS 18, I Q 26 July 1979, K. & A. & 0, & T. 59-65; Mesnil 1975: 1394. Rognes, (B.HJ; Kviteseid, Lundeberg (Vravatn), Previous records: Tachina arctica Zetterstedt, EIS 17, I Q 26 June 1980, (B.HJ. 1838: 645 (described from a single male captured The specimens from Lundeberg were caught on at the mountain Skaaddavara close to Alta, Finn­ Vaccinium myrtillus and V. uliginosum. New to mark, Northern Norway 7 Aug. 1820; Zetter­ Norway. stedt 1844: 1187; Siebke 1877: 88; Exorista arc­ Carcelia atricosta Herting, 1961 tica Zett. - Bidenkap 1901: 57. M. blondeli is lis­ Taxonomy: Herting 1961a: 7; Mesnil 1975: 1388; ted in Ringdahl's (1952) catalogue under the name Herting 1977: 5. Zenillia porcula Zett. (cr. Herting 1962: 83), but is Material: - Telemark: TEy: Nome, Varbu, EIS not recorded from Norway. This is probably due 18, Id 7 Aug. 1979,(B.H.). to the fact that Ringdahl (J 945a: 27) (er. Herting Mesnil (f.d regards this species as very close to 1964: 60) erroneously synonymized arctica with the Nearetic Carcelia perplexa Sellers, 1943. Her­ mitis Meigen, 1824. The latter species is recorded ting (in litt. 14 Sept. 1979) reports to have seen from Norway (RingdahI1952: 132-133 No. 19). material (unpublished) from Skane and Smaland Note that Ringdahl 0934: 267) correctly synony­ in Sweden. mized arctica with pexops Brauer & Bergens­ New to Norway. tamm, 1891 (= blonde/i). Eucarcelia pollinosa (Mesnil, 1940 Material: - Telemark: TEi: Fyresdal, Aslestad ­ Taxonomy: Mesnil 1944: 35 (as obesa); Ringdahl SIyst01 650 m a.s.\., EIS 16, Id 7 July 1980, 1934: 269; Ringdahl 1937: 38; -Ringdahl 1945a: (B.H.). 28; Mesnil 1963x: 3; Mesnil 1975: 1388. I follow The cerci agree with the figure given by Herting Mesnil (1963x: 4; 1975: 1388) in using the name (J 964: 60, fig. I a). Eucarcelia Baranov, 1934 instead of Senoll1etopia Myxexoristops stolida (Stein, 1924) Macquart, 1834 (ef. Crosskey 1973: 147; 1975: Taxonomy: Mesnil 1955: 446; Herting 1964: 103; Herting 1974: 7). • 59 -65; Mesnil 1975: 1394. Previous records: Carcelia rutilla B. & B. ­ Material: - Telemark: TEi: Tokke, Dalen, EIS Ringdahl I944b: 10; Ringdahl 1952: 134-135 16, I Q I July 1980, (B.HJ. No. 70. This species seem not to have been recorded from Material: - Akershus: AK: Brerum, Fossum, EIS Scandinavia before, not even by Hackman (1980), 28, 2 Q Q 3 Aug. 1979. - Vestfold: YE: Hof, but this is probably due to it having been regarded Thorrud, EIS 28, I Q 28 July 1979. - Buskerud: in the past as identical to M. blondeli (see above). B0: 0vre Eiker, Ulleland, EIS 28, I Q 5 Aug. New to Norway. 1979; Flesberg, Belgen, EIS 27, I Q 27 July 1979; Ocytata pallipes (Fallen, 1820) Hvila, EIS 27, 1I d d IQ 27 July 1979; Kongs­ Taxonomy: Mesnil 1952: 241; Herting 1974: 17. berg, Skollenborg, EIS 27, I d IQ 5 Aug. 1979. Previous records: Tachina pallipes Fall. - Siebke - Telemark: TEy: Nome, Varbu, EIS 18, 1 d 1977: 89; Rhacodineura pallipes Fall. - Ringdahl 2 Q Q 7 Aug. 1979, (B.HJ. TEi: Sauherad, Nord­ 1952: 136-137 No. 87. agutu, EIS 18, I Q 26 July 1979, K. & A. & 0. & Material: - Vest-Agder: VAy: Flekkefjord, Ras­ T. Rognes; Seljord, Ulvenes, EIS 17, 7 d d 2 Q Q vag (Hidra), EIS 4, I Q 24 June 1978, (SA). ­ 25 July 1979; Kviteseid, Skredi, EIS 17, I d 2 Rogaland: Ry: Lund, Moi, EIS 4, I Q 23 June July 1980; 2 d d 8 July 1980; Tokke, Dalen, EIS 1978, (SA); Sola, Gimra, EIS 7, I Q 19 July 16, IQ 30 June 1980. 1979. Almost all my specimens have been caught on Platymya fimbriata (Meigen, 1824) leaves of Populus tremula. Taxonomy: Mesnil 1954: 373 (as nemestrina); Drino Iota (Meigen, 1824) Herting 1972: 10. Taxonomy: Mesnil 1951: 164. Previous records: Exorista ./imbriata Meig. ­ Material: - Telemark: TEi: Sauherad, Norda­ Sch0yen 1889: 7; Ringdahl I944b: 9; Tachina hy­ gutu, EIS 18, I Q 26 July 1979, K. & A. & 0. & T. alinipennis Zetterstedt, 1838: 645 (misspelled hya­ Rognes. Iipennis by later authors, even Zetterstedt) (descri­ New to Norway. bed from specimens captured at Dovre mounta­ Drino vicina (Zetterstedt, 1848) ins, Norway, by Boheman); Tachina hvalipennis Taxonomy: Mesnil 1951: 166; Herting 1961a: 6; Zetterstedt 1844: 1145; Siebke 1877: 86; Tachina Mesnill975: 1391. tricingulata Zetterstedt, 1838: 646 (synonym ac­ Material: - Oppland: Os: Ringebu, Ringebu stav­ cording to Ringdahl 1934: 271; 1945a: 29); Zet­ kirke, EIS 63, I Q 9 July 1978, T. Nielsen, (S.AJ terstedt 1844: 1146; Siebke 1877: 86; Nemoraea New to Norway. 3-cingulata Zett. - Bidenkap 1898: 149.

110 Epicampocera succincta (Meigen, 1824) Phebellia glauca (Meigen, 1824) Taxonomy: Mesnil 1954: 318. Taxonomy: Mesnil 1955: 464. As for the taxo­ Material: - Akershus: AK: Brerum, Nordby, EIS nomy of the genus Phebellia Robineau-Desvoidy, 28, 1 Q 21 June 1979, (B. HJ. 1846, see Herting 1961a: 1-6; Mesnil 1970x: The record from Finland by Hedstrom (1964: 84) 111; Herting 1973: 4; Herting 1974: 8; Herting is probably the first from Scandinavia. The spe­ 1977: 2. cies is listed by Hackman (1980), but not by Lund­ Previous records: Exorista glauca Meig. - Ring­ beck (1927) or Ringdahl (1952). dahl 1952: 132-133 No. 16. See below. New to Norway. Material: - Telemark: TEi: Seljord, Ulvenes, EIS affinis (Fallen, 1810) 17, 1 Q 25 July 1979, (B.HJ. Taxonomy: Mesnil 1954: 391. I have not been able to trace any further records Previous records: Tachina affinis Fall. - Siebke from Norway, so Ringdahl's reasons for recor­ 1877: 85; Exorista affinis Fall. - Ringdahl ding the species from this country are not clear. 1944b: 10; Ringdahl 1952: 132-133 No. 25. However, among the names cited erroneously (cf. Material: - Vest-Agder: VAy: Lindesnes, Lin­ Herting 1961a; 1977) as synonyms to glauca by desnes, EIS 1, 1 d 20 July 1980, T. Nielsen; Flek­ Ringdahl (1945a: 29), namely strig(frons Zetter­ kefjord, Lauvnes (Hidra), EIS 4, 1 d 4 June 1979. stedt, 1838: 645 and stulta Zetterstedt, 1844: Lydella stabulans (Meigen, 1824) 1109, the former was based on a single male cap­ Taxonon1y: Mesnil 1955: 436; Herting 1959: tured at Alteidet (Troms: TRi: Kvrenangen), Nort­ 425 -429; Mesnil 1975: 1393. hern Norway. This record is repeated by Zetter­ Material: - 0stfold: 0: Fredrikstad, 0ra, EIS 20, stedt (1844: 1136) and Siebke 0877: 86). stulta is 1 Q 24 June 1979, (B. HJ. not reported from Norway. Ringdahl's (1952) re­ Known fron1 Denmark (Lundbeck 1927: 301), cord may therefore have been based on the speci­ Sweden (Ringdahl 1952: 132-133 No. 10) and men from Alteidet only, in which case the above Finland (Hackman 1980) as far as Scandinavia is record from Telemark is the first one from Nor­ concerned. way. New to Norway. Phebellia glaucoides Herting, 1961 Ni/ea hortulana (Meigen, 1924) Taxonomy: Herting 1961a: 1-2,6; Mesnil 1975: Taxonomy: Mesnil 1954: 382; Herting 1972: 8. 1394. On the limits of Ni/ea Robineau-Desvoidy, 1863 Material: - Buskerud: B0: 0vre Eiker, Burud, see Mesnil 1975: 1386. EIS 27, 1 d 1 Aug. 1979, (B.HJ. Material: - Buskerud: B0: 0vre Eiker, Burud, The terminalia agree with the figure published by EIS 27, 1 Q 1 Aug. 1979, (B.HJ. Herting (1961 a: 2, fig. 1). Mesnil (/.c.) supposes P. As regards Scandinavia reported from Denmark glaucoides to be identical with the Nearctic P. (Lundbeck 1927: 314), Sweden (Ringdahl 1952: itnitator Sellers, 1943. First Scandinavian record. 132-133 No. 21; Herting 1962: 82) and Finland Phryxe Inagnicornis (Zetterstedt, 1838) (Hackman 1980). Taxonomy: Mesnil 1954: 405 (as longicauda Wa­ New to Norway. inwright, 1940); Herting 1973: 4; Mesnil 1975: Ni/ea innoxia Robineau-Desvoidy, 1863 1393. Taxonomy: Mesnil 1954: 365; Herting 1974: 8. Previous records: Tachina magnicornis Zetter­ Material: - Buskerud: B0: 0vre Eiker, Burud, stedt, 1838: 644 (type locality Dovre, Norway); EIS 27, 1Q 1 Aug. 1979, (B.HJ. - Telemark: Zetterstedt 1844: 1141; Siebke 1877: 86; Strand TEi: Seljord, Ulvenes, EIS 17, 1 Q 25 July 1979, 1903: 7. (B.HJ. Material: - Hordaland: HOy: Bergen (Fana), Fj0­ In Scandinavia reported only from Sweden (Ring­ sanger, EIS 30, 1 d 1 Q 22 July 1980. dahl 1952: 132 -133 No. 29, as leth({era PandJ. The genitalia of the male agree with the figure in New to Norway. Mesnil (1954: 404, fig. 40). In Scandinavia repor­ Ni/ea ru.fiscutellaris (Zetterstedt, 1859) ted from Sweden (Zetterstedt I.e... Herting 1962: Taxonomy: Ringdahl 1945a: 29, 33; Mesnil 1954: 82, as longicauda), but not from Finland (Hack­ 366 (as temeraria Robineau-Desvoidy, 1863); man 1980). Herting 1973: 4; Herting 1974: 15; Mesnil 1975: Phryxe vulgaris (Fallen, 1810) 1393. Taxonomy: Mesnil 1955: 419. Material: - Telemark: TEi: Tokke, Dalen, EIS Previous records: Tachina vulgaris Fall. - Zet­ 16, 1Q 1 July 1980, (B.HJ. terstedt 1838: 644; Zetterstedt 1844: 1139; Siebke In Scandinavia reported from Sweden (Zetterstedt 1877: 86; Exorista vulgaris Fall. - Bidenkap 1859: 6115, Tachina; Ringdahl 1945b: 198, Pales; 1898: 149; Phryxe vulgaris Fall. - Ringdahl Ringdahl 1952: 132-133 No. 39; Herting 1962: 1944b: 10; Ringdahl1952: 132-133 No. 35; Ta­ 82, as temeraria .. Herting 1973: 4) and Denmark china mobi/is Zett. - Siebke 1877: 87 (synonym (Lundbeck 1927: 343, as Zenillia Bottcheri Ville­ according to Ringdahl 1934: 269; 1945a: 28). neuve, 1919). Material: - Akershus: AK: Oslo, S0rkedalen, EIS New to Norway. 36, 1 Q 22 June 1979. - Buskerud: B0: 0vre Ei­

111

• ker, Burud, EIS 27, I cJ I Q I Aug. 1979. - Hor­ Paykullia maculata (Fallen, 1820) daland: HOy: Bergen (Ask0Y), K1eppest0, EIS 30, Taxonomy: Herting 1961b: 29; Herting 1974: 31. I cJ 20 Sept. 1936, N. Knaben, 2MB. Material: - Rogaland: Ry: Sola, 200 m south of The male from Burud has been dissected and its Hafrsfjord bru, EIS 7, 1 cJ 13 July 1980; S4t­ terminalia agree with the figure in Mesnil (I 954: vanger, Sunde, EIS 7, IQ 15 July 1979; IQ 5 404, fig. 38). Sept. 1979, A. Rognes. Tlephusa cincinna (Rondani, 1859) The male has two pairs of rather distinct prst acr, Taxonomy: Ringdahl 1934: 268; Mesnil 1954: one half and one third as long as the neighbouring 328 (as diligens Zett.); Herting 1969: 192; Herting prst dc, respectively. Only two humeral setae are 1974: 9. present, the male has a weak third one on the left Previous records: Exorista diligens Zett. - Ring­ humeral callus. dahl I 944a: 80; Ringdahl I 944b: 9; Exorista ho­ New to Norway. nesta R.-D. - Ringdahl1952: 132-133 No. 17. Material: - Telemark: TEi: Seljord, To, EIS 17, I cJ 24 July 1979. ACKNOWLEDGEMENTS This species has for a long time been known un­ der the name diligens Zett. However, B. Herting I warmly thank Tore R. Nielsen, Sandnes, Svein informs me (in litt. 4 May 198 I) that none of the Svendsen, Kristiansand, and Torbj0rn Figved, two female syntypes of Tachina diligens Zetter­ lone Hugo Larsen, Trud Nyb0, lames Nystf0m, stedt, 1844: 1122-1123, which he has recently Asmund Vignes and Lars Gunnar Waage, all examined, belong to the species so named by Stavanger, for the gift of precious material; Lita Ringdahl and Mesnil. The latter species therefore Greve lensen, Museum of Zoology, 'Bergen, for must retain the name cincinna Rondani, 1859. Si­ ebke (1877: 85) only cites the data of the diligens the loan of specimens; and Benno Herting, Lud- l syntype captured in Norway. wigsburg, and Stig Andersen, Gopenhagen, for most kindly having verified my identifications of a number of specimens and in this way having greatly encouraged my studies. RHINOPHORIDAE Phyto cingulata (Zetterstedt, 1844) Taxonomy: Herting 1961 b: 16. Material: - 0stfold: 0: Kfiiker0Y; Glombo, EIS REFERENCES 20 I Q 24 June 1979. Ardo, P. I 957. Studies in the marine shore dune eco­ New to Norway. system with special reference to the dipterous Tricogena rubricosa (Meigen, 1824) fauna. Opusc. ent. Suppl. 14, 1-255. Taxonomy: Herting 1961 b: 20. Bidenkap, O. 1892. Unders0gelser over Diptera Bra­ Previous records: Fraue/!feldia trilineata Meig. ­ chycera i Jarlsberg og Laurvigs amt sommeren Ringdahl1944b: 9; Ringdahl1952: 144-145 No. 1891. Ent. Tidskr. 13, 225-246. 269. - 1898. Diptera Brachycera fra Jarlsberg. Tromsi1 Material: - Rogaland: Ry: K1epp, 0ksnevad, EIS Mus. Aarsh. 19 (/896), 147-150. 7, I cJ I Aug. 1979, T. Nielsen. - 190 I. Forel0big oversigt over de i det arktiske Stevenia atramentaria (Meigen, 1824) Norge hidtil bemrerkede Diptera Brachycera. Taxonomy: Herting 1961b: 23; Herting 1974: 33; Tromsi1 Mus. Aarsh. 23 (/900). 13-112. Crosskey 1977: 49. Crosskey, R.W. 1973. A conspectus of the Tachini­ Material: - Akershus: AK: Brerum, Nordby dae (Diptera) of Australia, including keys to the gilrd, EIS 28, I cJ 21 June 1979; 0verland, EIS supraspecific taxa and taxonomic and host catalo­ 28, II cJ cJ 5 Q Q 23 June 1979; Oslo, S0rkeda­ gues. Bull. Br. Mus. nat. Hist. (Ent). Suppl. 21. len, EIS 36, I cJ I Q 22 June 1979. 1-221. All my specimens have well developed external - 1974. The British Tachinidae of Walker and Ste­ vertical setae (vte). Many males are without an av vens (Diptera). Bull. Br. Mus. nat. Hist. (Ent.) 30, seta on the middle tibia (assymmetrically develo­ 267-308. ped in one specimen). The anterior half of the or­ - 1975. Tachinidae. - In: K1oet, G.S. & Hincks, bit has small proclinate setulae in varying number W.D. (ed.l, A check list of British insects. Second outside the frontal setae. The abdomen is black. edition. Part 5: Diptera and Siphonaptera. Hanbk. New to Norway. Ident. Br. Insects 9 (5), 95-105. Stevenia umbratica (Fallen, 1820) - 1977. A review of the Rhinophoridae (Diptera) Taxonomy: Herting 1961b: 27. and a revision of the Afrotropical species. Bull. Previous records: Rhinophora umbratica Fall.­ Br. Mus. nat. Hist. (Ent.) 36. 1-66. Siebke 1877: 91; Stevenia umbratica Fall. ­ Draber-Morilco, A. 1965. Monographie der paliiarkti­ Ringdahl 1952: 144-145 No. 263. schen Arten der Gattung Alophora R. - D. (Dip­ Material: - Vest-Agder: VAy: F1ekkefjord, Ras­ tera, Larvaevoridae). Annls zool. Warsz. 23, vilg (Hidra), EIS 4, I cJ 24 June 1978. 69- 194.

112 Emden, F.I. van 1954. Diptera Cyclorrhapha Calypt­ - 1975b. Neue palaarktischen Tachiniden (Diptera). rata (I). Section (a) Tachinidae and Calliphoridae. Stuttg. Beilr. Naturk. (A) 287, 1-7. Handbk Ident. Br. Insects 10 (4) (a), 1-133. - 1976. Revision der von Macquart beschriebenen Forsslund, K.-H. 1951. Insektfynd pa fjallet Harje­ palaarktischen Tachiniden und Rhinophorinen hagna. Norsk ent. Tidsskr. 8, 196 - 202. (Diptera). Stuttg. Beilr. Naturk. (A) 289, 1-10. Hackman, W. 1980. A check list of the Finnish Dip­ - 1977. Betrage zur Kenntnis der europaischen Rau­ tera 11. Cyclorrhapha. Notu/. Ent. 60, 117 -162. penfliegen (Dipt. Tachinidae). XIV. Stuttg. Beilr. Hedstrom, L. 1964. Lounais-Hameen karpasia, Bra­ Naturk. (A) 295, 1-16. chycera (Diptera). Lounais -Hiimeen Luonto 15, Lundbeck, W. 1927. Diptera Danica. Part VII. P/aty­ 77-89. pezidae, Tachinidae, G.E.C. Gad, Copenhagen. Herting, B. 1959. Revision einiger europaischer Rau­ Mesnil, L.P. 1944-1975. Larvaevorinae - In: penfliegen (Dipt., Tachinidae). Ann/n naturh. Lindner, E. (ed,), Die Fliegen der Pa/aearktischen Mus. Wien 63, 423-429. Region 10 (64g), 1-1435. - 1960. Biologie der westpalaarktischen Raupenflie­ - 1952x. Notes detachees sur quelques Tachinaires gen (Dipt., Tachinidae). Monogrn angew. Ent. 16, palearctiques. Bull. Ann/s Soc. r. ent. Be/g. 88, 1-188. 149-158. - 1961 a. Beitrage zur Kenntnis der europaischen - 1957x. Nouveaux Tachinaires d'Orient (Deuxi­ Raupenfliegen (Dipt. Tachinidae). III - VI. Stuttg. erne serie). Mem. Soc. r. ent. Be/g. 28, 1-80. Beilr. Naturk. 65, I -12. - 1963x. Nouveaux Tachinaires de la region pa­ - 1961 b. Rhinophorinae - In: Lindner, E. (ed,), learctique principalement de I'URSS et du Japon. Die Fliegen der Pa/aearktischen Region 9 (64e), Bull. Inst. r. Sci. nat. Be/g. 39 (24), I-56. 1-36. - 1967x. Tachinarires Palearctiques inedits (Dipte­ - 1962. Neue faunistische und biologische Daten ra). Mushi 41, 37-57. iiber schwedische Tachiniden (Dipt,). Opusc. ent. - 1970x. Description de nouveaux Tachinaires de 27, 80-86. I'ancien monde, et notes synonymiques (Diptera). - 1963. Beitrage zur Kenntnis der europaischen Ra­ Mushi 44, 89-123. upenfliegen (Dipt. Tachinidae) VII. Mitt. Schweiz. Pont, A.C. 1975. Rhinophoridae. - In: Kloet, G.S. & ent. Ges 35, 105-112. Hincks, W.O. (ed,), A check list of British insects. - 1964: Beitrage zur Kenntnis der europaischen Ra­ Second edition. Part 5: Diptera and Siphonaptera. upenfliegen (Dipt. Tachinidae). VIII. Entolnop­ Handbk. Ident. Br. Insects. 9 (5), 105 - 106. haga 9, 59-65. Ringdahl, O. 1934. Bidrag til kannedomen om en del - 1966. Beitrage zur Kenntnis der europaischen Ra­ av Zetterstedts tachinid-typer (Diptera). Ent. upenfliegen (Dipt. Tachinidae). IX. Stuttg. Beilr. Tidskr. 55, 266-272. Naturk. 146,1-12. - 1937. Bidrag til kannedomen om de svenska ta­ - 1967. Beitrage zur Kenntnis der europaischen Ra­ chinidernas utbredning. Ent. Tidskr. 58, 31-38. upenfliegen (Dipt. Tachinidae). X. Stuttg. Beilr. - 1944a. Revision av Vilhelm Storms Diptersam­ Naturk. 173, 1-11. ling. I. Familjen Tachinidae. K. Norske Vidensk. - 1968. Beitrage zur Kenntnis der europaischen Ra­ Se/sk. Forh. 17 (19),79-81. upenfliegen (Dipt. Tachinidae). XI. Stuttg. Beilr. - 1944b. Contributions to the knowledge of the ta­ Naturk. 196, 1/-8. chinids and muscids of Norway. Troms0 Mus. - 1969. Notes on European Tachinidae (Dipt,) desc­ Aarsh. 65 (1942), (4), 1- 27. ribed by Rondani (I 856-1868). Mem. Soc. ent. - 1945a. Forteckning over de av Zetterstedt i Insecta Ita/. 48, 189 - 204. Lapponica och Diptera Scandinaviae beskrivna ta­ - 1971. Beitrage zur Kenntnis der europaischen Ra­ chiniderna med synonymer jamte anteckningar upenfliegen (Dipt. Tachinidae). XII. Stuttg. Beilr. over en del arter. Opusc. ent. 10, 26-35. Naturk. 237.1-18. - 1945b. Oversikt over de hittils fran Sverige kanda - 1972. Die Typenexemplare der von Meigen arterna av familjen Tachinidae (Diptera). Ent. (1824 -1838) beschriebenen Raupenfliegen Tidskr. 66, 177 - 2to. (Dipt. Tachinidae). Stuttg. Beilr. Naturk. 243, - 1952. Catalogus Insectorum Sueciae XI Diptera 1-15. Cyclorrapha: Muscaria Schizometopa. Opusc. ent. - 1973. Beitrage zur Kenntnis der europaischen Ra­ 17,129-186. upenfliegen (Dipt. Tachinidae). XIII. Stuttg. Beilr. - 1953. Corrigenda ad pars XI: Muscaria Schizome­ Naturk. (A) 254, 1-18. topa. Opusc. ent. 18, 109. - 1974. ,Revision der von Robineau-Desvoidy besc­ - 1954. Nya fyndorter for norska Diptera. Norsk hriebenen europaischen Tachiniden und Rhinop­ ent. Tidsskr. 9, 46 - 54. horinen (Diptera). Stuttg. Beilr. Naturk. (A) 264, Rognes, K. 1980. The blow- genus Lucilia Robi­ 1-46. neau-Desvoidy (Diptera, Calliphoridae) in Nor­ - 1975a. Nachtrage und Korrekturen zu den von way. Fauna norv. Ser. B. 27, 39-52. Meigen und Rondani beschriebenen Raupenflie­ Rohdendorf, B.B. 1947. Kurzer Bestim­ gen (Dipt. Tachinidae). Stuttg. Beilr. Naturk. (A) mungsschliissel der parasitischen Dipteren der 27}, 1-13. Getreidewanze und verwandter Hemipteren der

113 Fam. Pentatomidae. Vrednaya tsherepashka 2. sektfauna. Nyt Mag. Naturvid. 37, 46-72. 75-88. (In Russian, title cited from Herting - 1903. Norske lokaliteter for Diptera, Christiania 1962). Vidensk. Selsk. Forh. 1900 (3), 1-11. Sch0yen, W.M. 1889. Supplement til H. Siebke's - 1914. Neue Beitrage zur Arthropodenfauna Nor­ Enumeratio Insectorum Norvegicorum, Fasc. IV. wegens. XVIII. Weiteres iiber von mir gesam­ (Diptera). Christiania Vidensk. Selsk. Forh. 1889 melte Diptera. Nyt Mag. Naturvid. 51,323-329. (l2), 1-15. Zetterstedt, l.W. 1838. Insecta Lapponica. Sectio Siebke, H. 1877. Enumeratio Insectorum Norvegico­ Tertia. Diptera. Lipsiae ( = Leipzig). rum Fasciculum IV. Catalogum Dipterorum Con­ - 1844. Diptera Scandinaviae disposita et descripta. tinentem. A. W. BT0gger, Christiania. 255 pp. 4, 895-1280. Lundae (= Lund) Stein, P. 1924. Die verbreitetsten Tachiniden Mittele­ - 1849. Diptera Scandinaviae disposita et deseripta. uropas nach ihren Gattungen und Arten. Arch. 8, 2935-3366. Lundae (= Lund). Naturgesch. 90 (A) (6). 1-271. Strand. E. 1900. Bidrag til Hallingdals og Lyng0rs in­ Received 19 Mar. 1981.

J

114 gin. Further limnological data are given by Faa­ feng et al. (1977). ( Short communications) The investigation was carried out during the ice free period. Monthly samples of nymphs at FIRST RECORD OF NEMOURA 15 localities around the lake were taken using A VICULARIS MORTON, 1894 (PLEC.,: the kick method and by picking up stones from NEMOURIDAE) IN WESTERN NORWAY 1 the bottom (Brittain and Lillehammer 1978). In addition, shore emergence traps (box-type, 1 LEIF MAGNUS SJ£TTEM m 3) (Davies 1950) were enlptied daily. The benthic samples contained 285 nymphs A total of 285 nymphs and 372 (213 dd, 159 Q Q) and the traps caugth 372 (213 dd, 159 Q Q) adults of N. avicularis were collected from Myrkdals­ adults of N. avicularis. In Myrkdalsvatn N. avi­ vatn, a lake in western Norway. It has not previously cularis had a one year life cycle with the main been recorded from western Norway. In Myrkdals­ growth period druing the autumn. Emergence vatn it had a univoltine life cycle with emergence at commenced just after ice break during the latter the end of May and the beginning of June, shortly af­ part of May and continued until 19 June (1977) ter ice break. Autumn was the main nymphal growth and 20 June (1978) with a peak at the end of period. Other plecopteran species found are mentio­ ned. May and beginning of June. N. avicularis was numerically the dominant species of Plecoptera Leif Magnus Srettem, Zoo!. Museum, University of in the lake. Other stone fly species recorded Oslo, Sarsgt. 1, N-Oslo 5, Norway. were: benthic samples- Nemoura cinerea (Retzi­ us), Nemurella picteti Klapelek, Leuctra sp., Previous studies (Lillehammer 1974) have indi­ Capnia sp.; emergence traps- N. cinerea, Sipho­ cated that the distribution of Nemoura avicularis noperla burmeistri (Pictet), Brachyptera risi Morton was restricted to the eastern parts of (Morton), Amphinemura sulcicollis (Stephens), Southern Norway. Previously it has been found Protonemura meyeri (Pictet), Capnia pygmaea further west than 0stre Slidre, Svenkerud in Zetterstedt, Leuctra fusca (L.) and L. nigra (Oli­ Hallingdal, Seljord in Telemark and Byglands­ ver). fjord in Aust-Agder (Lillehammer 1974). In With this new record of N. avicularis, two Norway N. avicularis has been found in small species of the genus Nemoura, N. avicularis and and large streams, rivers and lakes from coastal N. cinerea, have been recorded from Western areas up to the low-alpine belt. It occurs at low Norway. current velocities and shows a significant prefe­ rence for unstable substrata and large quantities of allochthonous matter (Lillehammer 1978). ACKNOWLEDGEMENTS The life cycle o(N. avicularis has been described I am indebted to John E. Brittain and Albert Lil­ as univoltine emerging in the spring (Brinck lehammer, both Zoological Museum, Oslo and 1949, Brittain 1973, Lillehammer 1978). Bror Jonsson, University of Oslo, for helping During 1977 and 1978 I investigated the litto­ with this article. ral fauna of the lake Myrkdalsvatn (60 0 49'N, 6°28'E, UTM: 626447). It has a surface area of 1,66 km2, a maximum depth of 107 m and forms a part of the Voss River System, western REFERENCES Norway. It is situated 25 km north of Voss at Brillck, P. 1949. Studies on Swedish stoneflies (Ple­ 230 m a.s.!. The lake is oligotrophic coptera) Opusc. ent. Suppl. 11, 1- 250. (~18:7-20 uS/cm, Ca++: 0.7-1.1 mg/l, Brittain, J.E. 1973. The biology and life cycle of Ne­ pH: 5.9-6.4) and dimictic with a maximum. moura avicularis Morton (Plecoptera). Freshwat. surface temperature of about 20°C in July. It is BioI. 3, 199-210. ice-covered from late November to mid May. Brittain, J.E. & Lillehammer, A. 1978. The fauna of Due to rapid, usually short term, water level the exposed zone of 0vre Heimdalsvatn: Methods, fluctuations the littoral zone is stony with very sampling stations and general results. Holarct. Ecol. 1, 221-228. little vegetation. A significant part ofthe organic Davies, D.M. 1950. A Study of the blackfly popula­ matter occurring in the lake is of terrestrial ori­ tion of a stream in Algonquin Park, Ontario. Trans. R. Can. Inst. 59, 121-160. 1Contribution from the Voss Project, Zoo!. inst., Faafeng, B., Brettum, P., Kristoffersen, T., Lind­ University of Oslo. str0m, E., Matzow, D., Nilsen, lP. & Tjomsland,

Fauna norv. Ser. B 28: Oslo 1981. 115 T. I977 . En unders0kelse av Vossevasdraget buried mouldy carcasses and sometimes under 1977. NIVA 0-76088. the bark of deCiduous trees. It is often found to­ Lilleharnmer, A. 1974. Norwegian stoneflies. n. Dis­ gether with R. parallelocollis. tribution and relationship to the environment. Unless they emerge to the surface, subterra­ Norsk ent. Tidsskr. 21. 195 - 250. nean beetles are rarely taken by the coleoptero­ - 1978. The Plecoptera of0vre Heimdalsvatn. Ho­ larct. Ecol. I, 232-238. logists' conventional collecting methods, and many species may prove to be more numerous Received 15 Sept. 1980. than the few records indicate. Baranowski (I978) reported an original and highly successful method for collecting R. perforatus: Potatoes were buried 10-40 cm below the ground and RHICOPHAGUS PERFORATUS ERICHSON covered with soil and bricks. After some months (COL., RHISOPHAGIDAE) NEW TO large numbers of the beetle were found in the NORWAY potatoes. By using this same method R. perfora­ tus may be discovered to be more common than PREBEN S. OTIESEN formerly assumed. Rhizophagus perforatus Erichson is reported new to Norway. One specimen was found under a rotten log on brick-mixed garden soil in the urban area of Kam­ ACKNOWLEDGEMENT pen, Oslo on 16 April 1979. I would like to thank Enid R. Peacock, British Museum (Natural History) for checking the Preben S. Ottesen, Zoological institute, University of identification of the species. I Oslo, P.O. Box, 1050 Blindern, Oslo 3, Norway.

The fourteen Fennoscandian members of the REFERENCES beetle family Rhizophagidae are usually found under bark where they feed on mould or sap. Baranowski, R. 1978. Interessanta skalbaggsfynd 3 Exceptions are the two closely related species (Coleoptera). Ent. Tidskr. 99. 53-60. • Rhizophagus parallelocollis Gyllenhal and R. Lindroth, C.H.(ed.) I960.Catalogus Coleopterorum Fennoscandia et Dania. Entomologiska perforatus Erichson which are mostly subterra­ Siillskapet, Lund. nean. The former is known as the «graveyard Peacock, E.R. 1977. Coleoptera Rhizophagidae. bettle» as it frequently is found in buried coffins Hanbk Ident. Br. Insects 5 (5aJ, 1-23. or swarming over the graveyards (Peacock Silfverberg, H. (ed.) 1979. Enumeratio 1977). It is apparently common in Oslo where I Coleopterorum Fennoscandia et Dania. have taken several specimens in pit-fall traps in Helsingfors Entomologiska Bytesforening, a graveyard close to downtown. Helsingfors. In a garden partly destroyed by bricks, woo­ den materials and inorganic garbage from a Received 12 Feb. 1981. torn-down house in the urban area of Kampen, Oslo, a specimen of R. perforatus was found on the underside ofa piece of rotten log on 16 April A RECORD OF XYLEBORUS D1SPAR F. 1979. There was still some winter ice under­ (COL., SCOLYfIDAE) IN WESTERN neath the log, and the beetle may thus have been NORWAY at its overwintering site. The species has previo­ KARE HESJEDAL usly not been reported found in Norway (Silf­ verberg 1979). The specimen is deposited in the A female specimen of Xyleborus dispar Fabricius was author's collection. collected in primo July, 1979, in Etne, Hordaland, EIS 23. This is the first record of the species in West­ R. perforatus is considered rare in Fennoscan­ ern Norway. dia and Denmark. Most records are from Swe­ den where it occurs north to ca. 62° latitude. Kare Hesjedal, Ullensvang Research Station, N-5774 The records closest to Oslo are from the regions Lofthus, Norway. of Dalsland and Dalarna (Lindroth 1960) which both border to Norway. Xyleborus dispar Fabricius is distributed According to Peacock (I977) the species is fo­ throughout the eastern and south-eastern parts und in exposed or buried decaying vegetable of Norway. According to the private archive of matter, in fungi, mouldy bones, moles' nests, the late Andreas Strand, records are reported

116 Fallna 1/0'1'. Ser. B 18.. Oslo 1981. from the provinces of 0stfold, Vestfold, Akers­ den, indicating that the specimens found represent a hus, Hedemark, Oppland, Buskerud, Telemark, relict population. Aust-Agder and Vest-Agder. No records, how­ ever, have been reported from the western part AIf Harald Dragseth, HoeIsandvegen, 6600 Sunn­ of Norway. dals0ra:Oddvar Hanssen, Langslagt, 8, 6600 Sunn­ dals0ra. In primo July, 1979, one single female of X. dispar was trapped in a pitfall trap for ground I Fennoskandia er Oedemera femorata Scopoli beetles in a strawberry field in Etne, Hordaland, hittil kjent fra Danmark (untatt Bornholm), Syd­ EIS 23. This was a rather unexpected locality Sverige nord tit og med BohusHin og Ostergot­ for trapping this species, which are associated land, og fra Finland hvor arten finnes syd for with weakened, deciduous trees. 64°N (Lindroth 1960). In East Norway the populations of X. dispar Imago er oppgitt a ha tithold pa skjermplan­ were high in 1978 due to the many weakened ter, hvor den spiser pollen (Hansen & Larson trees after the dry summers of 1975-77 (Ed­ 1945). land 1979). The males of this species cannot fly. Den 7. juni 1979 ble to eksemplarer (1 d 9 Accordingly the mating has to take place before mm og 1 Q 8 mm) av denne art funnet ved Hoel the females leave their adolescence tree, which i Sunndalen pa Nordmere (UTM 32V also are their hibernating site. The females can MQ823478). Den 25. juli 1980 ble ett eksemplar probably migrate over a long distance to lay (1 Q 9 mm) funnet pa Litle-Fale i Sunndalen their eggs (Edland 1979). (UTM 32V MQ937441). Dette individet satt pa The question arises if the species has become en blomst av kvann (Angelica archangelica) i established in Western Norway as a result of the apen krattskog av gnior (Alnus incdnaJ. Sist­ population growth in the eastern part of the co­ nevnte funnsted ligger 12,5 km unna stedet der untry. arten ferst ble funnel. Disse funnehe er gjort langt fra artens ncer­ meste tidligere kjente funnstetl, som er Bohus­ ACKNOWLEDGEMENTS lan. Da arten ikke er registrert i Oslofjordomra­ I thank A. Fjellberg and T. Kvamme for veryfy­ det, som er relativt godt undersekt, er det liten ing my determination of X. dispar. The inv~sti­ sannsynlighet for at den er oversett og har en gation has been financially supported by the Ag­ kontinuerlig utbredelse langs kysten opp tit ricultural Research Council of Norway. Nordmere. Det kan saledes vcere snakk om en relikt-populasjon fra en varmeperiode i postgla­ sial tid. REFERENCES Tit tross for sin nordlige beliggenhet har Strand, A.- Priva~e archive about the distribution of Sunndalen bade gunstig klima og yegetasjon Coleoptera in Norway. Unpubl. rep.. Museum of som kan gi grunnlag for varmekjcere insektarter. Zoology, University of Bergen. Edland, T.- 1979. Barkbiller pa frukttre. Gartneryrket 69, 233-235.

Received 19 Jan. 1981

BILLEN OEDEMERA FEMORA TA SCOPOLI FUNNET I SUNNDALEN pA NORDM0RE, MRi ALF HARALD DRAGSETH OG ODDVAR HANSSEN The beetle Oedemera femorata Scopoli is reported fo­ und in Norway for the first time. Two specimens Fig. 1. Oedemera femorata har gule harete dekk­ were found in Sunndalen, M0re og Romsdal pro­ vinger, men er ellers svart med et blagr0nt metalls­ vince 7 June 1979, and one specimen 12.5 km from kjrer. Tegningen viser en hunn; hannen har til for­ the first place 25 July 1980. This location is a long skjell fortykkede baklar. Eksemplarene fra Sunndal distance away from the nearest locality where the malte 8-9 mm, og litteraturen oppgir 8-10 mm species previously has been found, BohusUin in Swe- (Hansen & Larson 1945).

Fauna nor\.'. Ser. B 28: Oslo 1981. 117 REFERENCES the ground among low vegetation on a sunny, Hansen, V. & Larson, S. G. 1945. Heteromerer, Bil­ moderately warm day, and could be picked with ler Vo!. XII. Danm. Fauna Entom. 50, 1-293. the hands. The wings are slightly worn. The lo­ Lindroth, c.H. (Ed.) 1960. Catalogus Coleopterorum cality is situated in a suburban area which is sur­ Fennoscandiae et Daniae, Entom. sallsk., Lund. rounded by open cultivated farmland and which neighbours the small town of Sandnes in the Received 29 Jan. 1981. northern part of Jreren, south-western Norway. D. sylvestris, its host, is common in this part of the country (see e.g. l..0ken 1964: 203-204). The female of P. omissa closely resembles that of D. sylvestris, especially as regards the colour PSEUDOVESPULA OMISSA

118 Fauna no"". SeT. B 28, Oslo 1981. A RECENT RECORD OF LUCILIA narrowest point are 0.09-0.11 (mean 0.10) and BUFONIVORA MONIEZ, 1876 FROM 0.27 -0.3! (mean 0.30) times head width, re­ NORWAY (DIPT., CALLIPHORIDAE) spectively. In all the specimens the distance in front of the suture between the acrostichal rows KNUTROGNES of setae is distinctly less than the distance bet­ ween acrostichal and dorsocentral rows, as was 24 specimens of Lucilia bufonivora Moniez, 1876 also the case with the two male specimens on have recently been captured in Telemark, Southern which the description in my key (Rognes 1980) Norway. was based. One male has 3 post acr on the right side, but Knut Rognes, Hav0rnbrautene 7a, N-4040 Madla, the most anterior one is situated as far behind Norway. the suture as the front one normally does in this species. One female also has 3 post acr on the The only known captures of Lucilia bufonivora right side, but the front one, which is close to the Moniez, 1876 in Norway were made about 130 suture, is rather weak. In four males the beret is years ago by H. Siebke (Rognes 1980). The follo­ naked, and in one male the prosternum is devoid wing record of this species is therefore of some of hairs, which is quite abnormal for a Lucilia interest. species. The male ternlinalia agree with the figu­ TELEMARK: TEi: Tokke, Dalen, EIS 16, 30 res by Lehrer (1972). June 3 d d 3 Q Q; 1 July 1980 15 d d 3 Q Q. The specimens were caught by the author at low vegetation, mostly ferns, in the border-zone REFERENCES between a small forest, consisting mainly of Alnus sp. and Populus tremula, and an untended Lehrer, A.Z. 1972. Diptera Familia Calliphoridae. Fauna Rep'~ Soc. Romania, Insecta 11 (12), lawn some 100 m from the bus station in the 1-245. centre of the small city of Dalen. Some Rana Rognes, K. 1980. The blow-fly genus Lucilia Robi­ specimens were observed in the area. The loca­ neau-Desvoidy (Diptera, Calliphoridae) in Nor­ lity is not far from the Tokke river and the wes­ way. Fauna norv. ser. B, 27, 39-52. tern end of the lake Bandak (72 m a.s.1.). The male and fenlale frons measured at the Received 8 Sept. 1980.

process of sending the manuscript to the editor, how to deal with editors and printers. Next chapter descri­ (...... __800_'_k_re_V_ie_W__-",) bes how you order your reprints and how you use them. The three following chapters deal with writing review papers, conference reports and thesis. Ethics, rights and permissions are the topics of the next three Day, R.A. 1979. How to Write and Publish a Scienti­ chapters. The last chapter is the authors personal fic Paper. Philadelphia, ISI Press xi + 160 pp., 10 view on writing scientific papers, as well as a sum­ figs., 11 tab., $8.95 (paperback), $15,- (hardbound). mary of the book. The book contains three appenda­ ges of abbreviations: titles of journals, in table hea­ Writing good scientific papers is an art you either dings and general accepted ones. Two appendages have from birth or you must learn it step by step. deal with linguistic errors and words which should This process of learning is illustrated excellently in be avoided, and another with the SI-system of units. this little guidebook. An index of contents as well as a reference list also The book is divided into 26 short and easily read are included. chapters. "The first chapter defines what a scientific The book is well written in a clear and consise paper really is. The next 13 chapters deal thouro­ English. The text contains numerous examples of ughly with all parts of a paper, e.g. the title, the na­ various errors to avoid. me(s) of the author(s), the author' adresses, the ab­ With the aid of this book every student of science stract, the introduction, material and methods, the re­ will easily convert new laboratory findings to a sci­ sults, the discussion, acknowledgements, references, entific paper. For already established authors of sci­ design of tables and illustrations and typing the ma­ ence this book will be an excellent reference book. nuscript. The following three chapters deal with the U. Carlberg

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Arnett, Jr., Oxy­ phone number; 4) group of main interest, area of spe­ copis Pond Research Station, 90 Wallace Road, Kin­ cialization (e.g. family group, name, etc.) and kind of derhook, NY 12106 USA. interest (e.g., collecting, exchanging, systematics, ecology, etc.); 5) geographical area of greatest inter­ est; 6) short statement giving interest details (not over 25 words); 7) if a taxonomist, list groups willing to Announcing a new series published by the Norwegian identify for others; 8) signature and date. These date Entomological Society in association with the Royal will be coded and entered into the file. No person is Norwegian Society of Sciences and Letters, the Mu­ listed unless these data are supplied by that person. seum, Trondheitn. The right to edit all listings p~blished in the Directory is reserved. A new series, Insecta No rvegia, publishing catalo­ The first part of the new 44th edition of the Natu­ gues, and distribution maps will appear as required at ralists' Directory is entitled «The Naturalists' Direc­ irregular intervals. The series can be regarded as a tory of Collectors and Identifiers (Internatio­ supplement to Fauna norvegica ser. B, Norvegian naI)>>. This part includes persons interested in insects, Journal of Entomology. The editor is: John O. Solem, spiders, and other terrestrial invertebrates. The dead­ DKNVS Museet, Erling Skakkes gt. 47B, 7000 line for the receipt of new listings is OCTOBER 1981. Trondheim. The series is distributed by the Norwe­ Hundreds of listings are now in the PIFON file. gian Entomological Society through the secretary: The additional parts will list plant collectors and Trond Hofsvang, Postboks 70, 1432 As-NLH. The identifiers, bird watchers, field biologists (including first issue has been distributed for free to all members those interested in vertebrates other than birds, non­ of the society: insect aquatic , mollusks, ecologists, conser­ vationists, etc.), and rock, mineral, and fossil collec­ Refset, D. 1980. Atlas of the Coleoptera of Norway. tors. The final part will consist of subject, geographi­ 1. Silphidae, Catopidae, Colonidae, Leptinidae. cal, and name indexes for the entire edition. Insecta Norv. 1, 1-44. GUIDE TO AUTHORS. References. In the text: Black (979), Black & Blue 0973:100), or «as noted by Green (1978) and Black FAUNA NORVEGICA Ser. B. publishes papers in (1979)>>. Multiple references should be given in chro­ English, occasionally in Norwegian and German nological order, i.e. (Black & Blue, 1973, Green 1976, with an extensive English abstract. When preparing 1979, Black 1978). manuscripts for submission, authors should consult List of references are to be unnumbered and in inter­ current copies of Fauna norvegica and follow its national alphabetical order (i.e. = AA, JE and style as closely as possible. Manuscripts not confer­ A A. = Ae, 0 and = Oe). Titles of journals should be ring to the guide to authors will be returned for revi­ 6 sion. abbreviated according to the World List of Scientific Periodicals. Do not refer to papers «in prep.» among the references. Manuscripts should be submitted to one of the mem­ bers of the editorial committee or directly to the Edi­ Examples: tor-in-Chief. Send two copies. They must be type­ Journal: written, double spaced throughout, on one side of the L0ken,A. 1962. Social wasps in Norway (Hymenop­ paper, and with wide margins, 5-6 cm on the left. Se­ tera, Vespidae). Norsk ent. Tidsskr. 12, 191 - 218. parate sheets should be used for the following: I) Book~ Title page, with author's name. 2) An abstract, with the name and full postal address of the author under­ Mayr, E. 1913. Animal species and evolution. Har­ yard University Press. Cambridge, Mass. neath. 3) Tables with their headings. 4) Legends to fi­ gures. Fittkau,E.J. 1962. Die Tanypodinae (Diptera, Chiro­ nomidae). Die Tribus Anatopyniini, Macropelo­ Dates should be referred to as 10-20 Aug. 1970. poni und Pentaneurini. Abh. Larvalsyst. Insekten Only Latin names should be underlined. Other 6,453 pp. underlinings should be left to the editor. Approxi­ mate position of figures and tables in the text should Chapter: J be indicated in the margin. All acknowledgements Whitman,L. 1951. The vectors of yellow should be given under a single heading at the end of fever.- In: Strode,K. (ed.), Yellow Fever. Mc. the text, but before the references. Graw - Hill, New York & London, pp 229 - 298. Figures and Tables. Send two copies. All illustra­ tions should be identified lightly with the author's Proofs. Two copies of the first proof will be sent to name and the figure number. The figures and tables the author. One corrected copy should be returned to should be constructed in proportion to either the en­ the editor without delay. Alterations should be limi­ tire width of the typed area (140 mm) or to the co­ ted to correcting typesetting errors. Extensive altera­ lumn width (67 mm). tions will be charged to the author. Nomenclature. The first time a binomen is used in the text the name of its author should be included. Reprints. Twentyfive reprints are supplied free (fifty Author names should be written in full except L. for with multiple authorships). Additional reprints can Linneaus. Dates can be included when considered be ordered at a charge (an order form is sent with the necessary, i.e. Ryacophita nubila (Zetterstedt, 1840). proofs).

FAUNA NORVEGICA Ser. A, B, C viI normalt ut­ Referansemessig skal en aldri ta hensyn tit numme­ komme med til sammen 5 hefter i 10pet av en argang ret i fJvre hfJyre hjfJrne pd oms[aget Gnne i firkanten). (ett volum). For at heftene skal komme inn undet Det en skal ta hensyn t1l er de oppgitte data for re­ Postverkets regler for billig serie-utsendelse, for­ spektive serier. De f0rste heftene i hver serie vii i ar­ langes det at Fauna norvegica i hvert kalenderar gis gang 1981 hete henholdsvis Ser. BVo!. 28 No. 1. Ser. fortl0pende nummer fra I og oppover (altsa i argang C Cinclus Vo!. 4 No. I, Ser. A Vo!. 2 No. l. Det er 1981: 1-5). Det vii kunne bli noe tilfeldig hvilke disse data som gir den korrekte Iitteraturreferansen. hefter som blir gitt de respektive nummer pa grunn og det er disse forkortelsene som star oppf0rt i Ab­ av uregelmessigheter med rekkef01gen i 10pet av aret. stract til hver artikkel og pa srertrykkene. Content

Fauna norv. Ser. B. Vol. 28, No. 2. Dedicated to prof. emer. H. Kauri on his 75th birthday.

Greve, L. Prof. emer. Hans Kauri 75 ar ...... 49 Bengtson, S.A. Terrestrial invertebrates of the Faroe Islands: Ill. Beetles (Coleoptera): Checklist, distribu­ tion, and habitats...... 52 Compton, C. The chalcid fauna of the Jostedalen (Hym., Chalcidoidea)...... 83 Aarvik, L. The migrant moth Spodoptera exigua (Hlibnerl (Lep., Noctuidael recorded in Norway...... 90 Hofsvang, T. The species of the subgenus Tipula L. (Dipt., Tipulidae) in Norway...... 93 Greve, L. The genus Opomyza (Fallen) Dipt., Opomyzidael in Norway...... 96 Greve, L. & R. Rozko~nY. Three species of Sciomyzidae (Dipt.) new to Norway, with a note on Ectinocera bort'alis Zetterstedt 1838 100 Rognes, R. Recent records of Tachinidae and Rhinophoridae (Dipt.) from Norway 102

Short communications Srettem. L.M. First record of Nemoura aricularis Morton, 1894 (Plec. Nemouridael in western Norway 115 Ottesen, P.S. Rhizophagus per!oratus Erichson (Col., Rhizophagidae) new to Norway 116 Hesjedal, K. A record of Xylogorus dispar Fabricius (Col., Scolytidae) in western Norway 116 Dragseth, A.H. & O. Hansen. Billen Oedemea !emorata Scopoli funnet i Sunndalen pa Nordm0re, MRi 117 Rognes, K. & K. Mohn. Pseudovespula omissa (Bischoff, 1931) (Hym., Vespidae) recorded from Norway 118 Rognes, K. A recent record of Luci/ia bujonivora Moniez, 1876 from Norway (Dipt., Calliphoridae) ... 119

Book review , .. ,...... I 19

Announcement ...... 120

The Third European Congress of Lepldopterology will be held in Churchill ColIege, Camb­ ridge 13 -16 April 1982. The main theme of this Congress will be: Lepidoptera ecology and biogeography with Conservation of the Lepidoptera as a subsidiary theme. Provisional offers of papers on these and other LepidopterologicaI topics should be sent as soon as possible to: ­ 1. Heath Monks Wood Experimental Station Abbots Ripton Huntingdon, Cambs. PEI7 2LS Further details of the Congress will be issued in due course.

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