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Acada tolmelea.rirA - floweringbranch, B -se€d,C-pod. AtuomLD. Cowie1434 & C.R.Dunhp (DNA\I B & C from 1.r. Coltie 1844(DNA). (A & C scalebar = 1 cm, B scalebar = 5 mm) Typus:Litchfield National Park, Northern Tenitory, 13'06'5, 130" 51'E,23 Nov. 199O,I.D Cowie 1427 &C.R. Dunl.op(holo DNA; iso: BRI, CANB, K, NSW, PERTH 01851829). Erect shrub to 1.5m high, sometimessuckering, sub-pruinose on young phyllodes,branchlets and inflorescences.Brqnchlets flattened or angular,glabrous. Stipulesearly caducous.Pulvinus 4-7 mm long, glabrous. Phyllodes shallowlyrecurved, dimidiately narrowly elliptic, 110-245mm long, 48-88 mrn wide, 2.1-4.2 times long as wide, glabrous;main longitudinal nerves3 to 4 (rarely to 6), distant, basallyconfluent but remainingfree from abaxialmarginal nerve, anastomoses numerous andforming a somewhatopenand irregularreticulum; baseasymmetric;margin rarely indentedat the glands;apex acuteto obtusewith an obscuremucro. Glands tptomirrcnt, 3-4,lowermost at distal end ofpulvinus, additional glandsscattered along adaxialmargin, narrow andelongated, 1-2mmlong. Inflorescences typically 1-2 axillary racemes,sometimes forming terminal panicles,6-12 headsper raceme;raceme axis 55-150 mrn long, rather stout,shaight, glabrous. Heads glob ar,white to creamor pale yellow, 7-11 mm wide, c. 80-flowered. Peduncles9-26mmlong, glabrous;basalpeduncular bracts persistent, I or on upper pedunclesoften 2, glabrous,ciliate; bracteolespeltate, 1.2-1.4 mm long, with aflattened, glabrous or ciliate stipe, laminae sub-circular,evident in young buds, fimbriolate with white or pale golden hairs. Flowers 5-merous. Calyx 1.2-1.5mmlong, sparselypuberulous at apex and abaxially on prominent midnerve, shallowly lobed, membranousother than midnerve; calyx lobes broadly -l .etsu?d reurloJ eqt uo peJluecEu raq serceds aqt;o uo[nqr4srp aql tuo{ se,rueplaqlrde cgrce ds eqJ-,(Botout]A '3gg 'seru 3urpo3 leUE [[e,,[Jelocer o1srueddu pue 1rud tuuorleNplegqJ]r1 ur peluasardar11e,r sr 'y srsualawlot pauelEerqlpeJeprsuoJlou srl! uorlnqFlsppetcFtserJoqAnorfllv 'sn1D1suoltogasuo) 'peqsqqelseSunuoceq seedecsa puolsecco q1t,r pale^lllnc ,(lepr,ad^\ou s1t1 qSnoqtle uorSor re,rld euolcr^ eql ol palcrJlseJsr rlaunp V lo ecueJ]rrla)o '(u]c l€rnt€u eql ,&olrual urequoN eqt uJrIriA Ll-V x Zi-ZI) sepo ,{qd re8rel pu? slelqru€rq 'ule^ .V eteJal+ aql purSreru eql qlra.rsure,r pupngrtuol eql Jo oJsenuuoc["sBq eqt [q s\uaraulot 'sepollfqd rrroq per{srn8uqslpsr nuunp V pue slelqJueJqsnour1nlal-etellels ,{lalnuru esuep ,u,rou)l ,olocuas .V eql ,{q .{llBdrcuuddnotS eql u! 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Specimenswith narrow phyllodeshave, in the past,generally been attributed to the taxon describedas A. tanurubirinensis by Maiden (1917)but later reducedto a subspeciesofA. plectocarpaby Pedley( 1990). The geographicand morphological limits of subsp.tanumbiri?enri.s were not well known. Pedley (1978) remarkedthat fruiting matedal was unknown. In the absenceof fruits, specimensof A. plectocatpq have often been confused with A. torulosaBenth.ex F. Muell. The sandstoneescarpment of westernArnhem Land hasbeen a particularly difficult areain terms of the variation exhibited within A. plectocarpa ,ren.rlat specimens with somewhat naFow but generally long, curved phyllodes have been consideredas stbsp. tanumbirinensis. Other forms showing differencesofcorolla indumentumand inflorescenceor pod dimensionshave been dealt with variously asbelong\ngtoA. plectocarpaor to undescribedtaxa. In preparingto describeone ofthese segregatetaxa it becameobvious that somecladfication of the complex was required. This task was greatly facilitated following the recentrecognition by Tindale and Kodela (pers.comm.) that one of the No hem Territory entitiesin this complex was A. annitiiF N4uell ex Maiden, previously only known from the type collection in Queensland.Based on the immaturepods ofthe type collection, Pedley ( 1978)considered A. armitii as conspecificwith A. torulosa. However, in recentJreatments he hasreinstated it as a distinct species(Pedley 1987). The podsofA. armitii show no relationshipto A. torulosa(whichaseclearlymoniliform) butcertainlysupportacloserelationship with A. plectocarpa. After further investigationother species may alsobe found to belongwith this group.A. hammondii Maiden and the closely relatedbut poorly known A. mallocladaMaiden &Blakely havePlectocorpa type pods. A. hemsLeyiMaidenis alsosimilar but the pods arenot undulate,nor arethe seedsalternate along the pod which suggestsonly superficial similarity. The complex in the Northern Tenitory is consideredhere to involve 4 closelyrelated taxa namely, A. p lectocarpasnbsp.plectocarpa and sttbsp. tanumbirinensis,A. armitii ^r\dA. echinuliflora, with further workpossibly establishingother species in the group. The latterspecies is describedbelow asnew. Becausesome ofthese taxa have been kRown from incomplete material or few specimensa key and descriptionsare provided.
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