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Està subjecte a una llicència de Reconeixement-NoComercial- SenseObraDerivada 4.0 de Creative Commons CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

1 Late Quaternary pedogenesis of lacustrine terraces in 2 Gallocanta Lake, NE

3 E. Luna a,*, C. Castañeda a, F.J. Gracia b, R. Rodríguez-Ochoa c

4 a Estación Experimental de Aula Dei, EEAD-CSIC, Av. Montañana 1005, 50059 ,

5 Spain

6 b Departamento de Ciencias de la Tierra, Universidad de Cádiz, 11510 Puerto Real, Cádiz, Spain

7 c Departamento de Medio Ambiente y Ciencias del Suelo, Universitat de Lleida, Av. Rovira

8 Roure 191, 25198 Lleida, Spain

9 *Corresponding author. E-mail address: [email protected] (E. Luna).

10

11 Abstract

12 Transitional areas of lake margins are complex environments whose evolution is strongly

13 controlled by flooding frequency and persistence. The edaphic development of lacustrine

14 marginal environments can be reconstructed by combining detailed geomorphological analysis

15 with a systematic edaphic study of toposequences. This approach has been applied to a set of

16 recent lacustrine terraces in the downwind palustrine area of the Gallocanta saline lake, located

17 in a semiarid area in NE Spain. Up to five terraces, from 1.6 to 4.5 m above the lake bottom,

18 have been identified and mapped using stereo photointerpretation and airborne LiDAR data.

19 Several cycles of water level fluctuations, as part of a general trend towards lake desiccation,

20 have generated stepped terrace levels. The soils of these terraces have different morphological

21 characteristics and provide evidences for the Gallocanta paleolake being larger than that of the

22 present day. The soils have a sandy loam texture with variable clay content (1% to 46%) and a

23 predominantly carbonate composition (mean = 26%). The soils are developed in a sequence of

24 lacustrine carbonate-rich (mean = 37%) fine-grained gray layers overlaying detrital (mean = 51% CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

25 gravels) and frequently erosive, carbonate-poor reddish layers. The pedogenesis of the

26 downwind palustrine area is mainly characterized by poorly-developed carbonate accumulations

27 and common redox mottles associated with water level fluctuations in the lake, which

28 continuously rejuvenate or truncate the soils. Integrating pedological and geomorphological

29 features provides insight into recent complex lacustrine and soil forming processes and facilitates

30 management strategies and plans for this protected saline environment.

31 1. Introduction

32 Little is known about wetland soils developed in lake basins under semiarid climates. These soils

33 frequently become seasonally or intermittently dry due to the limited precipitation and high

34 evapotranspiration rates. Along the margins of arid wetlands, soil formation and properties are

35 closely related to geomorphic position and fluctuations in lake, or playa-lake, water levels

36 (Kolka and Thompson, 2007; Biggs et al., 2010; Farpoor et al., 2012; Shabanova et al., 2015).

37 For this reason the study of wetland soils is always intimately linked to the study of wetland

38 geomorphology and hydrology (Richardson et al., 2001).

39 Lake margins in semiarid climates are complex environments where sedimentation and soil

40 formation are determined by the balance between detrital inputs during wet seasons and salt

41 deposition during dry conditions (Boettinger and Richardson, 2001). In the wetting-drying

42 margins of the lake, water action on soils strongly influences their characteristics such as texture,

43 color, and types of horizons (Richardson et al., 2001). In this context, the study of soils provides

44 evidence of recent and past water level fluctuations in the lake (Castañeda et al., 2015), and may

45 be used for identifying regulatory boundaries (Lichvar et al., 2006). If high lake water periods

46 are long enough, they favor the generation of a morphosedimentary marginal surface of mixed

47 sedimentary-edaphic origin which can be abandoned and left perched once the lake level drops

48 again (Romanovsky, 2002). This is the origin of stepped lacustrine terraces in lakes that

49 experience a progressive desiccation trend (Gracia, 1995; Landmann and Reimer, 1996), as is the CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

50 case of Gallocanta Lake. Water level fluctuations in lakes are common in the arid and semiarid

51 Mediterranean region where most lakes are shallower (Beklioglu et al., 2007) and more sensitive

52 to climate oscillations than in more humid areas. Lake water level fluctuations during the Late

53 Quaternary have received much attention as proxies for identifying past environmental changes,

54 usually based on sedimentology (Ghinassi et al., 2012; De Cort et al., 2013; McGlue et al., 2013)

55 together with paleoecological evidence (e.g., pollen, ostracods, diatoms) (Shuman et al., 2001;

56 Hoffmann et al., 2012).

57 Although classic geomorphological studies of lakes focus on the different lake morphologies in

58 order to understand their origin and general evolution (Timms, 1992), few studies have

59 investigated the geomorphology of lacustrine terraces in shallow lakes and most of these studies

60 have looked at Pleistocene terraces related to major climatic oscillations (Bowman, 1971; Stine,

61 1990; Abu and Kempe, 2009; Ocakoglu et al., 2013). Only isolated contributions relate the

62 distribution and elevation of Holocene lacustrine terraces to recent climate changes

63 (Romanovsky, 2002; Gutiérrez et al., 2013). In fact, when compared to Pleistocene terraces,

64 Holocene historical levels are usually close to present water levels and hence their study requires

65 a very detailed high-resolution topographic analysis to distinguish different historical and recent

66 terrace levels, not often affected by present flooding.

67 Recognition of such lacustrine terraces and associated past flooding events requires

68 geomorphological and topographical techniques. Hence, lake terrace formation and the

69 interaction between lacustrine and pedogenetic processes can be reconstructed by combining

70 detailed geomorphological analysis and a systematic edaphic study of toposequences. This kind

71 of quantitative analysis is feasible with modern topographic techniques like airborne LiDAR

72 surveys and the digital terrain models derived from them, together with GIS software (Jones et

73 al., 2008; Budja and Mlekuž, 2010). High-resolution LiDAR-derived digital elevation models CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

74 have been widely applied in coastal areas (Kim et al., 2013; Matsu’ura, 2015) and fluvial

75 systems with subtle topography (Jones et al., 2008).

76 The present study focuses on soil development in lacustrine terraces that are assumed to have

77 been intermittently exposed during Late Quaternary-historical times, in the Gallocanta saline

78 lake, NE Spain. The aim is to integrate pedological and geomorphological features to reconstruct

79 the lacustrine terraces formation and understand recent lacustrine and soil formation processes

80 associated with water level fluctuations.

81

82 2. Gallocanta Lake

83 2.1. General setting

84 Gallocanta Lake is the largest well-preserved saline lake in Western Europe and has been

85 included in the Ramsar list since 1994 (Ramsar Convention Secretariat, 2010). The area

86 comprises a 6477 ha natural reserve that is protected and managed by the local government in

87 order to conserve endemism as well as habitats for the overwintering of migratory birds (Leránoz

88 and González, 2009). The lake, formed at the bottom of a karst polje (Gracia et al., 2002), is

89 located in a 543 km2 endorheic basin at approximately 1000 m.a.s.l. in the Iberian Chain, NE

90 Spain. The basin holds more than 20 lakes of karstic origin, Gallocanta Lake being the largest.

91 The Gallocanta Quaternary basin is elongated in the dominant wind direction (NW-SE), parallel

92 with the Valdelacasa mountain range, which runs along the NE side of the basin with peaks of up

93 to 1400 m.a.s.l. (Figure 1). This mountain range is composed of siliceous Ordovician rocks and

94 flanks an extensive outcrop of deformed carbonate units from the Mesozoic (Gracia, 2014). The

95 basin is excavated into Triassic clays and gypsum, as well as other more soluble salts (Gracia,

96 2009) which contribute to the soil and water salinity. The center of the lake contains about 1 m

97 of lacustrine sediments, the oldest of which have been dated as 43 ky BP (Rodó, 1997). Sediment CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

98 cores were analyzed for reconstructing Late Quaternary paleoenvironmental and paleoclimatic

99 changes by Schütt (1998), Rodó et al. (2002) and Luzón et al. (2007), as well as other authors.

100 The climate is dry semiarid (Liso and Ascaso, 1969) and there is a large yearly variation in

101 rainfall; the mean annual precipitation over the last 70 years is 488 mm yr-1 (range = 791 mm in

102 1959 to 232 mm in 2001), and the mean annual temperature is 11.3 °C for the period 1969-2012.

103 The frequent NW winds (Figure 1) exacerbate the hydric deficit (Martínez-Cob et al., 2010) and

104 produce longshore currents to the SE, generating shoreline landforms similar to those of marine

105 coasts (Castañeda et al., 2013). Water level fluctuations constitute the most outstanding feature

106 of the lake. The maximum lake water level, 2.84 m, was registered in 1974 (Pérez-Bujarrabal,

107 2014) and the lake desiccates completely during periods of low rainfall.

108 2.2. Downwind palustrine area

109 The SE sector of the lake, or downwind palustrine area, is approximately 500 ha in size and is

110 the largest area of the lake, where sediments and water accumulate during extraordinary flooding

111 events (Figure 1). This sedimentary plain, though flat in appearance (slope <1%), has preserved

112 lacustrine and coastal landforms which can be seen from aerial photographs due to the vegetation

113 and soil patterns. The plain is dominated by alternating flooding and drying periods that lead to

114 changes in soil salinity and moisture. The soil moisture regime around Gallocanta Lake is xeric

115 but soils subjected to frequent flooding have aquic soil moisture regime (Castañeda et al., 2015).

116 Historically, soils have been subjected to longstanding flooding (Comín et al., 1983; Pérez-

117 Bujarrabal, 2014) though at present most soils are exposed for long periods and are subjected to

118 either erosion or sediment transport under aerial conditions.

119 Figure 1

120 The small topographic variations and mixing of saline groundwater with fresh surface water

121 from runoff favor the preservation of a large area of protected habitats with an intricate

122 distribution of shallow ponds (Figure 1), halophytic and non-saline communities, mainly rushes CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

123 and reed beds. Winter cereal and other subsidized crops provide safe sites for the feeding and

124 nesting of protected birds. Low-lying saline areas stand as bare soils or are colonized by annual

125 and perennial halophytes, some of which include protected species such as Limonium sp. and

126 Puccinellia pungens. Limonium sp. is part of a priority habitat (1510 Mediterranean salt steppes)

127 and P. pungens is endemic (Gómez et al., 1983), being included in Annex II of the Habitat

128 Directive and in Appendix I of the Convention on the Conservation of European Wildlife and

129 Natural Habitats (Moreno, 2013).

130

131 3. Material and methods

132 Geomorphological photointerpretation was performed using aerial photographs from 2006

133 printed at 1:15 000 scale. The aerial photographs were taken in summer, when there was no

134 standing water in the lake. Field inspections were crucial for confirming the geomorphological

135 map, which was then transferred to orthophotographs and managed within the geographic

136 information system ArcGIS©. Orthophotographs from the 2009 and 2012 dry seasons were

137 overlain to contrast the stereo photointerpretation.

138 A digital elevation model (DEM) generated from airborne LiDAR data with an absolute vertical

139 accuracy of 0.20 m and a density of 0.5 points per square meter, was used to complement the

140 geomorphological photointerpretation. The elevation model was managed in ENVI© and

141 ArcGIS© for interactive histogram stretching and elevation data statistics. The average elevation

142 for each terrace was computed using the median value, as this is more robust than the mean

143 value in non-Gaussian distributions.

144 Soil sampling was based on the geomorphological map together with vegetation type and pond

145 distribution. Pits were dug during dry periods (zero lake water level), in May, June and August

146 2013, and August 2014. A total of nine pedons were studied, located along two toposequences

147 oriented NW-SE and NE-SW (Figure 2), which are parallel and perpendicular, respectively, to CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

148 the direction of the prevailing winds and the main axis of Gallocanta Lake (Figure 1). The

149 toposequences have a maximum difference in elevation of 1.2 m and 2.3 m, respectively. A soil

150 sample was collected from each horizon identified in the soil profiles, making a total of 53 soil

151 samples. Groundwater samples were collected from the pits where the water table was reached.

152 Soil profiles were described following Schoeneberger et al. (2012), and genetic and diagnostic

153 horizons and soil classification were based on Soil Taxonomy (Soil Survey Staff, 2014). The soil

154 samples were air-dried and sieved to less than 2 mm for subsequent laboratory analyses. Soil

155 salinity was measured as the electrical conductivity of the saturated paste extract, ECe (MAPA,

156 1994), using a conductivity cell (Orion 013605MD) and expressed in dS m-1 at 25 °C; pH of the

157 1:2.5 soil:water extract of the soil was measured using a pH electrode (Orion 9157BNMD).

158 Calcium carbonate equivalent, CCE, was measured by gasometry (MAPA, 1994). Organic

159 matter, OM, was determined by chromic acid digestion (Heanes, 1984) with a UV/V UNICAM

160 8625 spectrophotometer; and particle size distribution was assessed by laser diffraction with a

161 correction for the clay value following Taubner et al. (2009). The gypsum content was

162 determined using thermogravimetry ( et al., 2006) and confirmed with the qualitative test

163 (Van Reeuwijk, 2002) for gypsum content < 2%. The ionic content (Na, Ca and Mg) of saturated

164 soil-paste extracts was analyzed using an ionic chromatograph (Metrohm 861 Advanced compact

165 IC) (APHA, 1989). The pH and EC of the groundwater samples were measured (MAPA 1994)

166 with a pH electrode (Orion 9157BNMD) and a conductivity cell (Orion 013605MD),

167 respectively. In order to compare different properties of the soil profiles according to depth,

168 proportions of ECe and the sand, silt and clay content were calculated at soil depth intervals of

169 25 cm (Castañeda et al., 2012). The original 53 soil samples resulted in 63 synthetic soil layers

170 of 25 cm thickness. CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

171 Rainfall data recorded since 1944 at Tornos weather station (Figure 1), were complemented with

172 data from nearby weather stations using monthly regressions (Luna et al., 2014). Normal years

173 were identified from mean annual precipitation (Soil Survey Staff, 2014).

174 4. Results and discussion

175 4.1 Distribution of the lacustrine terraces

176 The downwind palustrine plain, isolated from the main lake bed by natural barriers, is delimited

177 by alluvial fans formed at the foot of the mountain range (Figure 1) and by Pleistocene lacustrine

178 coastal sediments on the eastern fringe (Figure 2). These ancient deposits, studied previously by

179 Gracia and Santos (1992), form a high plain more than 5 m above the study area. The plain is

180 506 ha in area and is primarily composed of a sequence of five stepped lacustrine terraces, T0 to

181 T4, whose elevation ranges between 1.6 and 4.5 m above the lake bottom (Table 1). The five

182 terrace levels show a fairly concentric distribution, decreasing in elevation towards the center of

183 the palustrine area. They form flat to gently undulating surfaces limited by slopes, often forming

184 low subvertical escarpments or microcliffs. From a sedimentological point of view the plain

185 belongs to the functional palustrine area defined by Pérez et al. (2002). Although some terrace

186 levels are recognized in other littoral zones of the lake at equivalent heights, they are usually

187 small and present incomplete sequences, whereas the downwind area shows the most complete

188 succession of terrace levels.

189 Table 1

190 Figure 2

191 The median elevation for each of the five terraces represents a robust marker for the successive

192 infilling steps in the palustrine area (Figure 3). The upper terraces, T4 and T3, are well

193 differentiated from the lower terraces, T1 and T0. T2 is a transitional terrace between the upper

194 and lower levels. T3 is inset in the previous level surrounding the palustrine depression and has CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

195 the most fragmented distribution (Table 1). T2 is much more connected than the upper terraces

196 and, finally, the lower terraces T1 and T0 are fairly continuous, with only six separate patches

197 (Table 1). T1 extends around the youngest terrace, T0, which comprises a cluster of several

198 ponds and bare floors that connect when flooded. T0 corresponds to the current local lake floor

199 of the palustrine area.

200 The coefficient of variation for the elevation confirms higher dispersion for the upper terraces,

201 especially T4 (Table 1). The slightly skewed distribution of the histograms (Figure 4) and the

202 presence of shoulders can be interpreted as terrace degradation or as different subsurfaces

203 corresponding to minor episodes of water level drop. Surface erosion, minor mass movements on

204 escarpments, and agricultural practices on the higher terraces have all contributed to their

205 topographic variability.

206 Figure 3

207 Figure 4

208 4.2 Geomorphological processes of terraces formation

209 The highest terrace level, T4, forms a set of NNE-SSW oriented barriers (Loma de -

210 Los Estrechos - Loma de Bello), which virtually isolate the downwind plain from the main lake

211 body (Figure 2). This terrace level also defines the outermost fringe of the plain and a group of

212 minor NE-SW aligned islands, which have the greatest extent of all terrace levels present in the

213 zone (Table 1). The roughly circular distribution of the T4 level confines a sub-parallel string of

214 depressions (Figure 2) where vegetation denotes the persistence of soil moisture. The generation

215 of the T4 terrace is associated with sediment accumulation due to the phenomenon of wind-

216 generated wave dissipation that occurs in shallow lakes oriented parallel to prevailing winds,

217 leading to lake segmentation (Zenkovich, 1967; Lees, 1989). Lees and Cook (1991) proposed a

218 conceptual model for the generation of transverse lake barriers and downwind lunettes on

219 shallow lakes, which fits Lake Gallocanta fairly well. CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

220 In an initial stage of lake level stillstand, unidirectional winds blowing from the NW would have

221 generated waves that interacted with the lake bed to transport sediments towards the SE. In a

222 second stage, interaction between waves, the shoreline beaches and migrating bedforms would

223 have led to the construction of a growing bank at a given distance downwind. The morphology

224 of the Loma de Berrueco barrier (Figure 2) suggests the prevailing wave-induced currents moved

225 clockwise. Residual currents flowing to the SE would have built the second, Loma de Bello,

226 barrier and the minor spits of Los Estrechos (Figure 2).

227 After the generation of the T4 terrace, the Loma de Berrueco - Loma de Bello barrier semi-

228 enclosed a downwind basin, only connected with the main lake body through the Los Estrechos

229 inlets. The subsequent water level drop during the Holocene produced a progressive base level

230 fall and the generation of the different lacustrine terrace levels, T3 to T0. The distribution of T3

231 parallels that of T4 and denotes continuity of the geomorphic processes promoting terrace

232 formation. Level T3 significantly segments the plain into two main low-lying basins, Los

233 Lagunazos to the NE and the Loma de Bello - Las Casillas basin to the S (Figure 2). Some

234 remnants of the T3 and T4 terraces display a recurved shape associated with the prevailing

235 longshore currents, which flow to the SE, as can be recognized in other littoral zones of the lake

236 (Gracia, 1995).

237 The 160 cm difference in elevation between T4 and T3 is relatively very high indicating a

238 noticeable water level drop in the paleolake. However, the areal extent of T4 is relatively large,

239 34% of the total palustrine zone, suggesting a substantially stable period of high water level

240 leading to widespread sedimentation. Another lake regression occurred after T3 and these

241 deposits were probably more deeply eroded resulting in a lower difference in elevation between

242 consecutive terraces: 50 cm on average. A subsequent lake regression occurred when T1 began

243 to develop, leading to 70 cm of difference in elevation between T2 and T1 (Table 2 and Figure

244 5). Finally, the last lake regression responds to a minor lake level fall of 20 cm, although this last CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

245 step is very decisive because it culminated in the almost complete enclosure of the palustrine

246 area. From this moment onwards, the downwind palustrine area has been almost isolated from

247 the main lake body by terrace T4, with a water inlet at the southernmost point of the Loma de

248 Bello barrier (Figures 2 and 5). The occasional entrance of water through this narrow gap can be

249 demonstrated with Landsat images taken during wet seasons (Figure 1).

250 The five lacustrine terrace levels provide evidence for the larger extent of the preceding

251 Gallocanta paleolake. Other examples of geomorphological evidence for lake retreat in semiarid

252 environments across the globe are mentioned by authors including Bowman (1971), Abuodha

253 (2004), Timms (2006), Abu and Kempe (2009), and Chen et al. (2013). Luzón et al. (2007)

254 studied sediment cores from Gallocanta Lake and deduced a postglacial maximum lacustrine

255 level (4-10 m) at around 8010 yr BP, coinciding with a relatively humid period at the beginning

256 of the Holocene, as has also been recorded in other Spanish lakes. A progressive water level

257 decrease followed this episode, although with important fluctuations. A recent period of

258 increased humidity was identified for the mid-19th century by Schütt (1998) and Luzón et al.

259 (2007).

260 In a situation of progressively falling water levels, the new lower level would impose new

261 dynamic conditions on the abandoned terrace deposits, and would probably involve their partial

262 erosion due to undercutting by waves, as well as runoff. Therefore, as Bowman (1971) and

263 Flower and Foster (1992) deduced from similar lacustrine features in other lakes, the

264 development of a lacustrine terrace also brings about a change in the earlier levels due to the

265 backwearing process.

266 Figure 5

267 4.3 Main characteristics of the soils and groundwater

268 The soil depth ranges from 120 cm to 210 cm and is mostly limited by the presence of

269 groundwater. The general sequence of horizons is A-B-C with the exception of GA33 and GA34. CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

270 In these cases the A-C sequence is probably related to their proximity to the water and sediment

271 inlet from the main lake (Figure 2). The thickness of the A horizon ranges from 4 to 45 cm and

272 weathered B horizons occur only in densely vegetated areas or in crop fields. The main

273 diagnostic horizons are Ochric, Calcic, Salic, and Cambic (Soil Survey Staff, 2014). There is a

274 progression from Inceptisols in the intermediate and upper terraces, formed under a xeric soil

275 moisture regime, to Aridisols in the lower terraces, where both strong salinity and aquic

276 conditions prevail (Soil Survey Staff, 2014). In general the soil profiles show two main distinct

277 matrix colors, gray (mostly 2.5Y) and reddish (10YR, 7.5YR and 5YR, Table 3); these colors

278 principally correspond to surface and subsurface horizons, respectively. Most soils in the area,

279 based on the sequence of horizons (Birkeland, 1999), are moderately developed.

280 Groundwater was reached at depths of between 90 and 200 cm (Table 2), and displayed no

281 relationship with the terrace levels or distance from the depocenter. The shallowest water table

282 was found at the outermost site of the SW-NE transect (GA33), suggesting influence of lateral

283 water flowing from the adjacent alluvial fan and mountain ranges. In general, the groundwater is

284 very saline in the lower terraces, with EC up to 106 dS m-1 in GA58, three times saltier than the

285 sea. At the outermost fringe of the plain (GA36 and GA55) the groundwater is non-saline (Table

286 2).

287 Saline groundwater is magnesium chloride type, similar to the surface water of the lake during

288 low water level periods (Comín et al., 1983), whereas fresh groundwater is magnesium

289 bicarbonate type. Saline and non-saline groundwater are largely enriched in magnesium, with an

290 Mg/Ca ratio of 30.8 in saline water and 8.7 in fresh water (Table 2). This Mg enrichment is

291 probably related to the precipitation of carbonates, as found by Renaut (1990) in semiarid lakes.

292 The predominantly bicarbonate composition of the groundwater preserves the moderately and

293 strongly alkaline pH of soils.

294 Table 2 CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

295 4.3 Soil composition and texture

296 The soils of the palustrine plain are neutral to strongly alkaline and have less than 2% organic

297 matter (OM) (Table 3). The minimum content of OM corresponds to the scarcely vegetated

298 youngest terrace, T0. Carbonate composition (mean CCE of 32.5%) predominates down to a

299 depth of 50 cm in the lower terraces and down to a depth of 100 cm in the upper terraces (Table

300 3). Below these depths the soil horizons are generally carbonate-free. Considering only

301 carbonate horizons, the soils at the center of the plain have a lower carbonate content (mean CCE

302 24.1%) than soils in the outermost area (mean CCE 34.7%). Previous studies focusing on the soil

303 surface, have given similar carbonate contents (Aranzadi, 1980; Calvo et al., 1978).

304 Unlike the soils and sediments at the bottom of the main lake (Comín et al., 1990; Luzón et al.,

305 2007; Castañeda et al., 2015), the palustrine soils are low in gypsum, with usually less than 5%,

306 with the exception of the T0 topsoil (gypsum 9%) (Table 3). Remarkably, 16% gypsum was

307 found below 2 m at Loma de Bello. This deep gypsum-rich layer could be associated with an

308 evaporitic environment suggesting a predominantly lacustrine origin for this barrier.

309 Table 3

310 The salinity of the soil samples, measured as ECe, shows a broad range, from 0.2 dS m-1 to 70.7

311 dS m-1. In general, the maximum salinity within the soil profiles is found mainly below 100 cm,

312 evidencing the influence of the saline groundwater. Comparing the different soils using the ECe

313 values estimated for the 25 cm synthetic layers (Figure 6), the 0-25 cm soil layer is usually less

314 saline than the subsurface layers with the exception of the lowest terrace T0. Following the

315 salinity phases established for irrigated agricultural soils by the NRCS (Soil Survey Division

316 Staff, 1993) and modified by Nogués et al. (2006), and taking into account the greater ECe of the

317 0-50 cm soil samples, we obtain a saline soil distribution that depends on the terrace level and

318 the distance to the depression depocenter (i.e., GA58). Very strongly saline soils occur at the

319 innermost fringes of the lower terraces whereas non-saline soils are found at the outermost CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

320 fringes of the upper terraces. Paralleling soil salinity, the maximum Mg/Ca ratio (> 14) occurs in

321 the very strongly saline soils of the lowest terraces (Table 3).

322 Even though the downwind palustrine area of Gallocanta Lake is quasi isolated from the main

323 lake bed, soil salinity is still a major feature associated with the occurrence of saline groundwater

324 and the evapoconcentration caused by capillary rise. However, the leaching of soluble salts down

325 the soil profile by rainwater results in decreased salinity in the surface layers except for the

326 lowest terrace T0, where the upward movement of saline groundwater predominates. The lateral

327 surface and subsurface flows of fresh water that enter the palustrine area probably cause the local

328 differences in soil salinity.

329 Figure 6

330 The soil samples have a predominantly sandy loam texture with a mean sand content of 59%,

331 ranging from 27% to 86% (Table 3). These sandy soils contrast with the clayey materials of the

332 main lake floor previously described by González-López et al. (1983) and Mayayo et al. (2003).

333 Based on the particle size distribution estimated for the 25 cm synthetic layers, the surface

334 horizons are sandier than the subsurface horizons (Figure 7). The upper terraces show increased

335 sand content at a depth of about 75-100 cm. This sand increase consistently correlates with the

336 sandy layer identified in the soils of the main lakebed margins at a similar depth (Castañeda et

337 al., 2015). The exceptionally high silt/clay ratios (up to 43.6) at a soil depth of approximately

338 100 cm and at various terrace levels (Table 3) is probably related to the occurrence of that sandy

339 layer. A rise in the lake water level could be inferred from this increase in grain size, something

340 which is also seen at a shallower level in GA33, which has a similarly high silt/clay ratio (18.7)

341 (Table 3). The Loma de Bello barrier presents three consecutive fining downwards sequences

342 with their boundaries at depths of 100 cm and 250 cm (Figure 7). These cycles are marked by

343 abrupt changes in the sand and clay content (Table 3), probably revealing a pattern of successive

344 lake water level fluctuations. CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

345 Figure 7

346 Based on the mean sand content per profile, the lowest sand content corresponds to GA59, in the

347 northern sector of the palustrine plain, whereas the highest sand content occurs at GA55, the

348 easternmost point of the NW-SE transect (Figure 2). These extreme values probably reflect the

349 effect of the prevailing NW wind which promotes the accumulation of sandy sediments towards

350 the SE, i.e., GA55. Wind action is also inferred from the relatively high percentage of quartz

351 gravels in the topsoil at GA58 (Table 3); this suggests the prevalence of aeolian deflation and

352 subsequent downwind accumulation. In this regard, an aeolian supply of sand to this lee zone of

353 the lake cannot be discounted, as has been recorded in other lacustrine lunette deposits (Lees,

354 1989).

355 Romeo-Gamarra et al. (2011) determined the mineralogical composition of the clays in soils of

356 the north and south lake margins subjected to similar intermittent flooding conditions. Illite

357 predominates, up to 73%, whereas dolomite ranges from 3 to 9%, and quartz varies from 8 to

358 15%. Smectite also occurs in strongly saline soils, with percentages up to 15%. Qualitative

359 determinations of clay minerals by Calvo et al. (1978) and Aranzadi (1980) also mention the

360 illite as predominating, with a lesser quantity of kaolinite, and even including small proportions

361 of smectite.

362

363 4.4. Pedogenic accumulations and redoximorphic features

364 Carbonate accumulations are common at all terrace levels (with the exception of GA35) and

365 include gravel coatings and pendants, as well as soft and friable nodules from 5 to 15 mm. These

366 accumulations, together with redox mottles, are best developed at the bottom of the B and C

367 horizons (Figure 8). The non-saline soils contain the highest content of carbonate nodules,

368 >40%, and even a non-cemented carbonate crust (Figure 8). Carbonate coatings, pendants and

369 banding are much less frequent in the strongly saline soils. The morphologies of the carbonate CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

370 accumulations at all terrace levels correspond to stage II of pedogenic carbonate development

371 (Machette, 1985; Schoeneberger et al., 2012), in contrast with the stage IV carbonate

372 development found in the older soils of the alluvial fans surrounding the lake (Castañeda et al.,

373 2015). The widespread occurrence of non-cemented carbonate nodules and bands evidences

374 present day mobilization of carbonates, suggesting a close relationship with the fluctuating water

375 table.

376 Figure 8

377 Gypsum accumulations are very scarce (Figure 8). Gypsum crystals occur in surface (GA33) and

378 subsurface (GA35) saline layers together with friable gypsum nodules. Vermiform gypsum can

379 be seen in the upper horizons of GA57. Salt crystals are visible in subsurface horizons at the

380 lowest terrace level (GA58).

381 Redoximorphic features resulting from prolonged soil saturation and related to alternating

382 wetting and drying cycles, such as seasonally high groundwater or flooding, are widespread in

383 the palustrine area. Small iron and/or manganese oxidation mottles are frequent, though sparse,

384 in the subsurface horizons of all the pedons studied. The oxidation mottles are either dark (mean

385 value = 2 and chroma = 1) or light (mean value and chroma = 6)(see Table 3 and Figure 8), and

386 are sometimes associated with pores, rock fragments, and root channels. Occasional black,

387 rounded or banded manganese oxide mottles (10YR 3/1) are seen in soils with contrasted salinity

388 (GA34 and GA36).

389 Gray reduction mottles are frequent or abundant in some of the studied soils; they are 2.5Y or 5Y

390 in color with a value of ≥5 and chroma of ≤4. This occurs in subsurface horizons at a soil depth

391 which increases with the terrace level. Reduction mottles can form vertical tongues up to 20 cm

392 in length, probably related to preferential circulation of water, revealing the intensity and extent

393 of the reducing conditions in the palustrine area. Accumulations of manganese bands (5PB 2.5/1) CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

394 are also present in subsurface horizons where a high concentration of reduction mottling occurs

395 (Figure 8).

396 Redoximorphic features in other soils of Gallocanta Lake have been described from macro to

397 microscale in different landscape positions, under either oxidizing or reducing conditions

398 (Castañeda et al., 2015). In general, soils of the palustrine area are under predominantly

399 oxidizing conditions, probably favored by the high porosity of the sandy materials and the

400 significant percentage of gravels in subsurface horizons that allow air and water to circulate. The

401 only exceptions are the surface horizons of GA34 and GA59, from the lower T1 and T2 terraces.

402 There, reducing conditions are preserved probably due to a higher flooding frequency because of

403 their proximity to intermittently ponded areas (Figure 2). Another noticeable redox feature is the

404 presence of depleted matrix in the surface layers of the GA34, GA35, and GA57 soils.

405 According to the criteria of Richardson and Vepraskas (2001) and the USDA-NRCS (2010),

406 these soils are hydric.

407 Figure 9

408 Figure 10

409 4.5 Soil genesis under alternating conditions

410 The palustrine soils of the lowest terraces were submerged in the past century (Pérez-Bujarrabal,

411 2014), covered by macrophytes (Comín et al., 1983), and most likely subjected to subaqueous

412 pedogenesis (Demas et al., 1996). These soils have been exposed during periods of low water

413 level in the lake, which have been longer and/or more frequent in recent decades (CHE, 2003).

414 The formation of these soils results from alternating episodes of flooding with fresh to

415 hypersaline water, and subsequent drying, analogous to a tidal environment.

416 Soils of the palustrine downwind plain are regularly truncated and subjected to a constant

417 process of rejuvenation, as shown by the buried horizon and the grain size sequences of GA35;

418 the truncated sequence of GA57, which is common in subaqueous soils (Demas and Rabenhorst, CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

419 1999); and the frequent lithological discontinuities (Figure 8). The presence of hiatuses in the

420 sediments of Gallocanta Lake has been previously identified from limnological data (Rodó et al.,

421 2002).

422 Two main discontinuities affecting the palustrine plain are identified based on the presence of

423 wavy boundaries. These discontinuities are associated with different episodes of lacustrine and

424 detrital material sedimentation. A deep limit, at about 100-120 cm in the upper terraces, suggests

425 a relationship with the predominant alluvial-littoral sediments underlying the lake bottom. This

426 erosive episode is more evident in soil profiles at the outermost fringe of the palustrine area,

427 GA55 and GA56 (Figure 8). A shallower discontinuity, at a depth of about 30-40 cm in the

428 lowest terraces (Figure 8) suggests episodes of renewed flooding due to a rise in the lake water

429 level and the input of detrital material from the main lake.

430 A simplified pedogenetic model of the palustrine area consists of a sequence of gray lacustrine

431 layers overlaying reddish detrital layers. The lacustrine layers have a high CCE content (mean =

432 37%) and low gravel content (mean = 3%). Their thickness increases with terrace level. The

433 detrital layers, which in places are capped with sandy channels and bars, have a low CCE content

434 (mean = 6%) and consist of quartz gravels (51%) probably with an alluvial-littoral origin.

435 Lacustrine fine-grained gray layers overlay the detrital and frequently erosive reddish layers

436 (Figures 9 and 10). Locally, semi-lacustrine layers with a high content of both CCE and gravels

437 occur at the base of the lacustrine horizons, usually with a wavy lower boundary, or are

438 intercalated with lacustrine materials (e.g., at GA33, Figure 10).

439

440 4.5. Lacustrine terraces and historical records of water occurrence

441 Figure 11 shows that only the lowest terraces are susceptible to flooding from the maximum

442 water levels recorded over recent decades. The lake water level needed to cover the median

443 elevation of the uppermost terrace is 4.5 m. This elevation corresponds to a surface extent of CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

444 about 2300 ha, comparable with the 1800 ha and 4 m depth estimated by pioneering studies in

445 the 19th century (Pérez and Roc, 1999). In 1974 the surface extent of the lake water was

446 estimated to be 1505 ha and, recently, it has decreased to 500 ha (CHE, 2003). Evidence for the

447 extent of the water surface during flooding events includes the limited records of water level

448 measurements from the 1970s recorded by CHE (2003), occasional aerial photographs from the

449 past century (Pérez-Bujarrabal, 2014), and remote sensing data (Díaz de Arcaya et al., 2005;

450 Castañeda and Herrero, 2009).

451 The oldest reference evidencing water level fluctuation is the shoreline retreat of up to 200 m

452 mentioned by Hernández-Pacheco and Aranegui (1926). The variation in water level recorded

453 from 1977 to 1988 by Comín et al. (1983, 1990) is associated with annual and seasonal rainfall

454 changes. Seasonal changes in lake water level are from 20 to 60 cm every year (Comín et al.,

455 1990). At the beginning of the eighties the lake totally desiccated (Figure 11), whereas the

456 wettest period identified was 1970-1977, when the lake reached its maximum water level, 2.84

457 m, according to the scale monitored by Pérez-Bujarrabal (2014). Figure 11 summarizes the

458 quantitative and qualitative information available on Gallocanta Lake water level fluctuations

459 compiled from several authors (Aranzadi, 1980; Gracia, 1990; Comín et al., 1990; Rodó et al.,

460 2002; CHE, 2003) from 1944 to the present (Luna et al., 2014). A rough correlation is made

461 between wet/dry years obtained by applying the standard definition of normal years (Soil Survey

462 Staff, 2014) (A, Figure 11) and by compiling the observations from the literature regarding the

463 presence of water in the lake (B, Figure 11). The number of years with rainfall above or below

464 the normal year (Soil Survey Staff, 2014) is fewer than the number of dry, very dry, or wet years

465 compiled from the literature. The best correspondence between the two data sources are for the

466 wet period 1957 - 1977, and the dry period 1982 – 1984, as well as the following wet years

467 (1987-1991). In the last two dry decades, only five years have really been dry based on annual

468 rainfall estimates. CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

469 Figure 11

470 6. Conclusions

471 Photointerpretation based on aerial photopraphs taken in the summer of 2006 was crucial in

472 identifying the subtle landforms of the downwind palustrine area. Integrating LiDAR high-

473 resolution DEM with geomorphological photointerpretation provides consistency when

474 delineating the recent lacustrine terrace levels. The pronounced flatness of Gallocanta Lake,

475 together with intense and persistent unidirectional winds paralleling the marked elongation of the

476 lake, have produced predominant water and sediment transport towards the lee zone where

477 sedimentation processes have generated a set of lunettes forming a complex lacustrine barrier. A

478 palustrine area was therefore generated beyond the barrier, but which was still connected to the

479 main lake body through small inlets. As a consequence, the flat downwind palustrine area was

480 protected against the erosional action of waves, but was particularly sensitive to lake water

481 fluctuations. Several cycles of water fluctuations as part of a general trend towards desiccation

482 gave way to the generation of 5 stepped lacustrine terrace levels that display different

483 pedogenetic properties. At lake scale, the distribution, shape and topography of the successive

484 lacustrine terraces in the palustrine zone evidence a sustained retraction of the lake area which

485 seems to be a result of climatic drying, although brief re-flooding episodes have also been

486 identified. At a finer scale, the pedogenesis of soil terraces reveals morphological and

487 sedimentary changes including truncations and discontinuities in the soils. Pedogenesis has

488 resulted in a sequence of fine-grained, gray lacustrine layers with a thickness that increases

489 according to the terrace level, overlaying detrital and erosive carbonate-poor reddish layers. Our

490 findings show that there was a scant development of lacustrine sedimentation following an

491 intense period of predominantly detrital sedimentation, meaning an episode of high water level

492 in the lake is required to explain the distribution and surface expression of these deposits. The

493 downwind palustrine area of Gallocanta Lake provides a record of enormous edaphodiversity, CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

494 constituted by a variety of complex lacustrine environments where pedogenesis has been

495 strongly controlled by flooding episodes. The lacustrine materials found in the soils studied

496 confirm that the downwind area was part of the submerged floor of a much larger paleolake, and

497 in addition they indicate past palustrine conditions. The next step in the research will be the

498 dating of the different terraces in order to evaluate the rate at which pedogenetic processes have

499 acted in this fluctuating environment.

500

501 Acknowledgements

502 This article has been funded by the Spanish Ministry of Economy and Competitiveness under

503 project AGL2012-40100 and supported by the Andalusian PAI Research Group no. RNM-328.

504 E. Luna was financed by a fellowship from Aragón Government, Spain. Orthophotographs and

505 LIDAR data were supplied by the Spanish National Geographic Institute (Instituto Geográfico

506 Nacional). Rainfall data from Tornos were provided by the Spanish Meteorological Agency

507 (AEMET) after contract no. L2 990130734.

508

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737 CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

738 Table 1. Selected metrics of each of the five terraces as obtained from LiDAR data. C.V: 739 coefficient of variation.

Elevation difference Elevation, m a.s.l. Surface m No. Between From lake Terrace patches Max Median Min CV ha % terraces bottom T4 12 1000.2 995.8 992.9 0.12 173.7 34 1.6 4.5 T3 21 997.2 994.2 992.8 0.06 128.8 25 0.5 3.0 T2 14 995.7 993.7 992.4 0.04 90.3 18 0.7 2.5 T1 6 994.6 993.0 992.2 0.03 60.9 12 0.2 1.8 T0 6 993.7 992.8 992.1 0.03 52.3 10 1.6 740

741 Table 2. Physical properties and main ions of the groundwater in the downwind palustrine area 742 of Gallocanta Lake.

2+ 2+ + + 2- - - - Depth EC Mg Ca Na K SO4 Cl HCO3 NO3 Sample pH -1 cm dS m meq L-1 GA33 155 7.3 87.1 989.6 32.1 756.7 3.6 721.8 991.2 10.0 63.2 GA34 130 7.3 83.0 736.2 34.2 709.4 5.7 526.5 1029.6 2.0 6.9 GA36 90 8.2 4.9 32.0 3.7 29.5 0.6 31.8 18.9 16.0 0.0 GA55 168 8.5 1.3 8.2 4.4 2.9 0.1 5.3 3.3 12.0 0.7 GA57 200 7.5 66.9 611.8 41.9 553.0 4.9 474.4 795.4 7.0 2.0 GA58 180 7.1 106.0 1083.4 36.0 1058.7 12.2 696.5 1545.4 5.0 4.7 743 CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

Table 3. Physical and chemical properties of the nine soils studied. *: Auger sampling. L: lacustrine, D: detritic, SL: semi-lacustrine; ECe: electrical conductivity of the saturation extract; SAR: sodium adsorption ratio; Mg/Ca: ratio in equivalents; pH: measured on the saturated paste; CCE: calcium carbonate equivalent; OM: organic matter; Rock fragments: weight percent. - Not determined.

Rock USDA Depth Munsell Color ECe CCE Gypsum OM Sand Silt Clay Silt/ Horizon Sediment SAR Mg/Ca pH fragments Textural Clay cm Matrix Mottles dS m-1 % class GA33 Typic Aquisalid 2.5Y 0-21 Az L 27.2 18.6 6.0 8.0 34.6 2.7 1.4 15.3 70.8 20.2 9.0 2.2 Sandy loam 5.5/3 21-28/39 Cz SL 2.5Y 6/3 7.5YR 5/8 38.6 22.6 14.1 8.2 32.0 <2 0.2 52.3 87.2 12.2 0.6 20.3 Sand 7.5YR Sandy Clay 28/39-54 top 2Cz L 51.1 25.3 16.0 8.2 32.4 <2 0.1 15.3 56.8 17.9 25.3 0.7 7/4 loam 28/39-54 down 2Cz L 2.5Y 8/3 44.7 23.5 14.0 8.2 48.4 2.4 0.2 6.3 50.2 7.2 42.6 0.2 Sandy Clay 7.5YR 54-160 3Cgkz D 10YR 6/6 2.5/1 and 40.7 22.3 13.9 8.0 4.7 2.0 0.1 40.7 59.4 24.5 16.1 1.5 Sandy loam 5Y 7/4 GA34 Typic Aquisalid 0-8/10 Az L 5Y 5.5/2 37.8 23.5 5.5 8.2 27.5 3.0 0.5 3.0 62.3 29.4 8.3 3.5 Sandy loam 2.5Y 8/10-25/30 Cz L 2.5YR 7/4 37.8 24.1 12.1 8.4 36.2 <2 0.3 5.6 49.0 34.0 17.0 2.0 Loam 7.5/2 2.5Y 7.5/4, 2.5Y 6/3 25/30-70 2Cgkz D 10YR 6/6 45.4 27.7 16.1 7.9 12.4 <2 0.1 41.0 70.8 18.4 10.8 1.7 Sandy loam and 10YR 6.5/6 7.5YR 70-117/120 3Cgz D 44.7 26.2 13.5 7.6 <2 <2 0.1 72.6 70.5 21.4 8.1 2.6 Sandy loam 5.5/6 117/120-135 4Cz D 5YR 4/6 59.7 31.8 12.8 7.5 <2 <2 0.0 28.1 91.5 8.3 0.2 41.5 Sand GA35 Typic Haploxerept Ap1 and 0-20 L 2.5Y 5/2 0.5 0.3 1.4 8.0 42.7 3.4 1.3 0.8 61.3 22.5 16.2 1.4 Sandy loam 2 20-37 Bw1 L 2.5Y 6/2 0.4 1.0 2.3 8.5 43.2 3.4 0.7 0.4 45.8 30.2 24.0 1.3 Loam CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

37-70 Bw2 L 2.5Y 5/3 2.5 4.8 4.0 8.2 35.0 3.0 0.6 0.3 52.5 34.9 12.6 2.8 Sandy loam 10YR 4/1 70-100 2Aby L 2.5Y 4/2 and 10YR 10.4 10.4 4.8 8.1 25.0 2.4 0.4 0.8 50.4 35.7 13.9 2.6 Loam 7/5 10YR7/7 100-150 2Cgy1 L 2.5Y 6/6 16.5 11.7 3.4 8.1 39.6 2.1 0.1 0.1 41.6 32.6 25.8 1.3 Loam and 2.5Y 150-210 2Cgy2 L 10YR 7/7 22.6 14.2 3.8 7.9 33.2 2.4 0.1 0.6 40.8 39.3 19.9 2.0 Loam 7/3.5 210-250 * L - 24.1 15.1 3.7 7.8 25.3 16.0 0.1 0.0 65.0 26.7 8.3 3.2 Sandy loam 250-270 * L - 20.1 12.9 3.4 7.9 37.8 7.5 0.1 9.9 46.5 15.9 37.6 0.4 Sandy Clay Sandy Clay 270-290 * SL - 16.4 11.2 3.0 7.8 34.8 <2 0.1 20.9 55.1 20.6 24.3 0.8 loam 290-320 * L - 15.0 10.8 3.0 7.8 37.7 <2 0.1 3.1 43.9 25.1 31.0 0.8 Clay loam Sandy Clay 320-350 * L - 15.0 10.5 2.8 7.5 38.7 3.1 0.1 9.0 52.3 23.7 24.0 1.0 loam GA36 Typic Calcixerept 0-45 Ah L 2.5Y 4/2 1.0 2.7 4.3 8.5 26.7 4.0 2.5 0.9 49.7 30.4 19.9 1.5 Loam 45-57 AB L 2.5Y 5/1 10YR 6/6 2.7 10.3 12.8 8.7 49.5 2.2 0.4 0.0 35.3 37.3 27.4 1.4 Clay loam 57-80 Bwgk L 10YR 6/4 2.5YR 5/1 2.4 0.0 7.2 8.5 47.9 <2 0.3 0.1 26.6 37.1 36.3 1.0 Clay loam 10YR 80-120 Ck L 10YR 6/6 3.4 7.4 5.8 8.4 63.8 <2 0.1 1.6 76.0 11.5 12.5 0.9 Sandy loam 7.5/3 GA55 Typic Calcixerept 7.5YR 0-23 A1 L 1.7 1.0 12.3 8.6 30.8 4.0 3.2 0.0 82.2 14.0 3.8 3.7 Loamy sand 4/2 10YR 23-40 A2 L 1.7 2.8 11.9 8.5 30.3 0.0 1.1 5.5 71.5 22.4 6.1 3.7 Sandy loam 4.5/2 2.5Y Sandy clay 40-75 2Bgk L 2.5Y 7/8 0.9 1.3 1.9 8.2 50.9 0.0 0.6 0.5 64.9 13.7 21.4 0.6 7.5/3 loam 10YR 75-95/105 3Bk SL 0.9 0.8 1.2 8.1 29.9 0.0 0.2 38.9 83.7 8.1 8.2 1.0 Loamy sand 6.5/6 7.5YR 95/105-118/127 4C D 5Y 6/2 0.2 0.6 1.0 8.3 <2 0.0 0.1 35.0 85.5 13.6 0.9 15.1 Sand 5/8 2.5Y 5/4 118/127-168 5Cg D 10YR 5/8 0.4 0.6 0.8 8.1 2.2 0.0 0.1 53.6 67.4 26.9 5.7 4.7 Sandy loam and 5Y 5/3 CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

GA56 Typic Calcixerept 0-35 A L 2.5Y 5/3 0.8 0.6 1.1 8.0 23.9 2.2 1.5 0.9 64.7 25.3 10.0 2.5 Sandy loam Sandy clay 35-85 2Bk1 L 2.5Y 6/5 7.5 YR 4/6 11.8 4.9 5.4 8.1 38.4 <2 0.5 0.0 50.5 27.1 22.4 1.2 loam 10YR 85-100/105 2Bk2 L 7.5 YR 4/6 2.1 3.0 6.9 8.2 50.9 0.0 0.2 0.3 44.1 23.0 32.9 0.7 Clay loam 6.5/4 5Y 5.5/2 100/105- 10YR 3C SL and 7.5YR 1.2 1.8 5.5 8.2 16.5 0.0 0.1 15.4 85.0 11.9 3.1 3.8 Loamy sand 110/117 6.5/5 6/8 7.5 YR 6/8 110/117- 4C D 10YR 5/6 and 7.5 YR 1.1 1.5 6.7 8.2 6.9 0.0 0.1 33.7 67.9 25.8 6.3 4.1 Sandy loam 135/145 2.5/3 7.5YR 135/145-170 4C2 D 7.5 YR 6/8 - - - - <2 2.6 0.0 62.7 73.8 20.9 5.3 3.9 Sandy loam 3.5/4 GA57 Sodic Calcixerept 0-22 A L 2.5Y 6/2 0.9 1.3 0.8 8.0 27.7 2.9 1.8 1.5 71.1 21.7 7.2 3.0 Sandy loam 10YR 22-42 Cy L 20.3 17.9 3.2 8.1 29.5 3.2 0.8 2.1 59.6 29.4 11.0 2.7 Sandy loam 5.5/2 10YR 10R 2/1 42-80 2Bgk D 22.6 19.1 5.9 8.1 14.9 <2 0.3 61.3 62.1 27.6 10.3 2.7 Sandy loam 5.5/8 and 5Y 6/2 80-110/120 2C D 5YR 4/6 5Y 6/2 19.1 18.3 7.9 7.9 8.1 <2 0.3 63.1 72.2 21.1 6.7 3.1 Sandy loam 110/120-135 3C D 5YR 4/6 10YR 6/3 25.2 20.0 10.5 7.9 <2 <2 0.1 4.0 70.7 24.4 4.9 5.0 Sandy loam 135-205 4C D 5YR 4/6 23.9 19.4 9.3 7.9 <2 <2 0.1 42.5 76.1 18.7 5.2 3.6 Loamy sand GA58 Calcic Aquisalid 0-4 Az L 10Y 5.5/1 5PR 2/1 70.7 33.4 5.6 8.0 23.0 9.3 1.6 14.2 56.3 32.7 11.0 3.0 Sandy loam 7.5YR 4-43/48 2Bkz SL 5PR 2/1 37.6 25.5 7.7 8.2 23.2 2.5 0.5 41.7 62.3 22.1 15.6 1.4 Sandy loam 4.5/6 43/48-85/97 3Ckz D 5YR 5/7 5PR 2/1 41.5 26.7 16.3 7.9 <2 2.5 0.2 57.8 63.6 29.4 7.0 4.2 Sandy loam 85/97-105 4Cgz D 10YR 5/8 38.7 25.4 24.1 7.6 <2 2.1 0.2 5.9 55.9 35.0 9.1 3.8 Sandy loam 5YR 105-127 5Cz D 47.7 27.0 20.0 7.4 <2 <2 0.1 64.9 72.9 21.4 5.7 3.8 Sandy loam 4.5/6 127-180 6Cz D 5YR 5/7 53.1 28.8 23.4 7.2 <2 <2 0.1 62.3 85.5 13.3 1.2 11.1 Loamy sand CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

GA59 Typic Calcixerept 0-20 A L 10YR 5/2 5.0 3.7 1.3 7.6 41.5 3.7 3.6 0.0 37.7 40.8 21.5 1.9 Loam 20-55 Bwg L 2.5Y 5/1 6.1 7.2 2.6 8.0 28.1 3.5 1.5 0.8 35.8 43.4 20.8 2.1 Loam 55-100/160 2Bgk SL 10YR 5/8 10YR 2.5/1 14.1 10.7 2.9 8.0 27.5 3.7 0.3 46.0 32.3 43.1 24.6 1.8 Loam 7.5YR 100/160-190 3Cg D 10YR 2.5/1 15.1 13.6 5.5 7.8 2.9 3.0 0.1 50.0 37.3 44.4 18.3 2.4 Loam 4.5/6 190-200 4Cg - 5YR 4/6 10YR 2.5/1 ------

CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

Figure captions

Figure 1. False color composition (RGB 543) of a Landsat 5TM image (from the U.S.

Geological Survey) acquired on 14/04/1987 showing Gallocanta Lake and its

downwind palustrine area partially flooded. The nearest weather stations, Los

Picos and Tornos, are marked. The wind rose shows the relative frequency and

direction of the moderate (2.0 to 5.0 m s-1) winter winds measured at a height of 2

m (modified from Martínez-Cob et al., 2010).

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CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046

Figure 2. Geomorphological map of the downwind palustrine area of Gallocanta

Lake (see Figure 1 for general location). The nine soil profiles of the two soil toposequences studied, NW-SE and NE-SW, are marked.

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Figure 3. Boxplots of elevation for the five stepped terraces forming the downwind

palustrine area of Gallocanta Lake, obtained from LiDAR data. Interquartile range

box, medians and their confidence intervals, and outliers are represented.

Figure 4. Terrace topography extracted from LiDAR DEM (see Figure 2 for scale) and

the corresponding histograms showing the median (red vertical line).

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Figure 5. Photographs taken during dry seasons showing the terraces of the palustrine

area of Gallocanta Lake at different scales.

Figure 6. Soil salinity (ECe) of the studied profiles displayed for 25 cm-thick layers.

Profiles are colored according to their mean salinity: very strongly saline and

strongly saline soils are red; moderately saline soil is purple; and non-saline and

slightly saline soils are green.

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Figure 7. Soil profiles studied through the two perpendicular toposequences with

the particle size distribution and ECe values (in red), calculated for the 25-cm

thick synthetic soil layers.

Figure 8. Main morphological features of the soils studied along the NE-SW and NW-

SE toposequences.

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Figure 9. Scheme of the lacustrine, semi-lacustrine, and detrital materials along the two

toposequences studied.

Figure 10. Sequence of lacustrine, semi-lacustrine and detrital materials in four selected

profiles (GA33, GA34, GA56 and GA55).

Figure 11. The available water level records of Gallocanta Lake, including continuous

(line) and point (dots) measurements, and the mean elevation of the five lacustrine

41

CATENA 147: 372–385 (2016) http://dx.doi.org/10.1016/j.catena.2016.07.046 terraces in the downwind palustrine area of Gallocanta Lake. The colored horizontal lines along the bottom represent: A) annual rainfall from 1944 above

(blue) and below (orange) the normal year (Soil Survey Staff, 2014); and B) dry

(orange), very dry (dark orange), and wet periods (blue and dark blue) as described in the literature (see references in the text).

42