2
AN ORDINAL The Angiosperm Phylogeny Group Iridaceae Anarthriaceae Ixioliriaceae Centrolepidaceae CLASSIFICATION Lanariaceae Cyperaceae Laxmanniaceae Ecdeiocoleaceae FOR THE FAMILIES Orchidaceae Eriocaulaceae OF FLOWERING PLANTS Tecophilaeaceae Flagellariaceae Themidaceae Hydatellaceae Xanthorrhoeaceae Joinvilleaceae Xeronemataceae Juncaceae Poaceae CLASSIFICATION OF FLOWERING PLANTS Dioscoreales Hook.f. Prioniaceae Amborellaceae Nartheciaceae Burmanniaceae Restionaceae Austrobaileyaceae Petrosaviaceae Dioscoreaceae Sparganiaceae Canellaceae Triuridaceae Taccaceae Thurniaceae Chloranthaceae Thismiaceae Typhaceae Hydnoraceae Acorales Reveal Trichopodaceae Xyridaceae Illiciaceae Acoraceae
Nymphaeaceae Liliales Perleb Zingiberales Griseb. [+Cabombaceae] Alismatales Dumort. Alstroemeriaceae Cannaceae Rafflesiaceae Alismataceae Campynemataceae Costaceae Schisandraceae Aponogetonaceae Colchicaceae Heliconiaceae Trimeniaceae Araceae Liliaceae Lowiaceae Winteraceae Butomaceae Luzuriagaceae Marantaceae Cymodoceaceae Melanthiaceae Musaceae Ceratophyllales Bisch. Hydrocharitaceae Philesiaceae Strelitziaceae Ceratophyllaceae Juncaginaceae Ripogonaceae Zingiberaceae Limnocharitaceae Smilacaceae Laurales Perleb Posidoniaceae
Atherospermataceae Potamogetonaceae Pandanales Lindl. EUDICOTS Calycanthaceae Ruppiaceae Cyclanthaceae Gomortegaceae Scheuchzeriaceae Pandanaceae Buxaceae Hernandiaceae Tofieldiaceae Stemonaceae Didymelaceae Lauraceae Zosteraceae Velloziaceae Sabiaceae Monimiaceae Trochodendraceae Siparunaceae Asparagales Bromhead [+Tetracentraceae] Agapanthaceae COMMELINOIDS Magnoliales Bromhead Agavaceae Proteales Dumort. Annonaceae Alliaceae Abolbodaceae Nelumbonaceae Degeneriaceae Amaryllidaceae Bromeliaceae Platanaceae Eupomatiaceae Anemarrhenaceae Dasypogonaceae Proteaceae Himantandraceae Anthericaceae Hanguanaceae Magnoliaceae Aphyllanthaceae Mayacaceae Ranunculales Dumort. Myristicaceae Asparagaceae Rapateaceae Berberidaceae Asphodelaceae Circaeasteraceae Piperales Dumort. Asteliaceae Arecales Bromhead [+Kingdoniaceae] Aristolochiaceae Behniaceae Arecaceae Eupteleaceae Lactoridaceae Blandfordiaceae Lardizabalaceae Piperaceae Boryaceae Commelinales Dumort. Menispermaceae Saururaceae Convallariaceae Commelinaceae Papaveraceae Doryanthaceae Haemodoraceae [+Fumariaceae] Hemerocallidaceae Philydraceae [+Pteridophyllaceae] MONOCOTS Herreriaceae Pontederiaceae Ranunculaceae Hesperocallidaceae
Corsiaceae Hyacinthaceae Poales Small Japonoliriaceae Hypoxidaceae 3 4
CORE EUDICOTS Penthoraceae Casuarinaceae Dirachmaceae Pterostemonaceae Fagaceae Elaeagnaceae Aextoxicaceae Saxifragaceae Juglandaceae Moraceae Berberidopsidaceae Tetracarpaeaceae Myricaceae Rhamnaceae Dilleniaceae Nothofagaceae Rosaceae Gunneraceae Rhoipteleaceae Ulmaceae Myrothamnaceae ROSIDS Ticodendraceae Urticaceae Vitaceae Aphloiaceae Malpighiales Mart. Caryophyllales Perleb Crossosomataceae Achariaceae EUROSIDS II Achatocarpaceae Ixerbaceae Balanopaceae Aizoaceae Krameriaceae Caryocaraceae Tapisciaceae Amaranthaceae Picramniaceae Chrysobalanaceae Ancistrocladaceae Podostemaceae Clusiaceae Brassicales Bromhead Asteropeiaceae Stachyuraceae Dichapetalaceae Akaniaceae Basellaceae Staphyleaceae Erythroxylaceae [+Bretschneideraceae] Cactaceae Tristichaceae Euphorbiaceae Bataceae Caryophyllaceae Zygophyllaceae Euphroniaceae Brassicaceae Didiereaceae Flacourtiaceae Caricaceae Dioncophyllaceae Geraniales Dumort. Goupiaceae Emblingiaceae Droseraceae Francoaceae Hugoniaceae Gyrostemonaceae Drosophyllaceae Geraniaceae Humiriaceae Koeberliniaceae Frankeniaceae [+Hypseocharitaceae] Irvingiaceae Limnanthaceae Molluginaceae Greyiaceae Ixonanthaceae Moringaceae Nepenthaceae Ledocarpaceae Lacistemataceae Pentadiplandraceae Nyctaginaceae Melianthaceae Linaceae Resedaceae Physenaceae Vivianiaceae Malesherbiaceae Salvadoraceae Phytolaccaceae Malpighiaceae Setchellanthaceae Plumbaginaceae Medusagynaceae Tovariaceae Polygonaceae EUROSIDS I Ochnaceae Tropaeolaceae Portulacaceae Pandaceae Rhabdodendraceae Celastraceae Passifloraceae Malvales Dumort. Sarcobataceae Huaceae Putranjivaceae Bixaceae Simmondsiaceae Parnassiaceae Quiinaceae [+Diegodendraceae] Stegnospermataceae [+Lepuropetalaceae] Rhizophoraceae Cistaceae Tamaricaceae Stackhousiaceae Salicaceae Cochlospermaceae Scyphostegiaceae Dipterocarpaceae Santalales Dumort. Cucurbitales Dumort. Trigoniaceae Malvaceae Olacaceae Anisophylleaceae Turneraceae Muntingiaceae Opiliaceae Begoniaceae Violaceae Neuradaceae Loranthaceae Coriariaceae Sarcolaenaceae Misodendraceae Corynocarpaceae Oxalidales Heinze Sphaerosepalaceae Santalaceae Cucurbitaceae Cephalotaceae Thymelaeaceae Datiscaceae Connaraceae Saxifragales Dumort. Tetramelaceae Cunoniaceae Myrtales Rchb. Altingiaceae Elaeocarpaceae Alzateaceae Cercidiphyllaceae Fabales Bromhead Oxalidaceae Combretaceae Crassulaceae Fabaceae Tremandraceae Crypteroniaceae Daphniphyllaceae Polygalaceae Heteropyxidaceae Grossulariaceae Quillajaceae Rosales Perleb Lythraceae Haloragaceae Surianaceae Barbeyaceae Melastomataceae Hamamelidaceae Cannabaceae Memecylaceae Iteaceae Fagales Engl. Cecropiaceae Myrtaceae Paeoniaceae Betulaceae Celtidaceae Oliniaceae 5 6
Onagraceae EUASTERIDS I EUASTERIDS II FAMILIES OF UNCERTAIN POSITION Penaeaceae Psiloxylaceae Boraginaceae Adoxaceae Balanophoraceae Rhynchocalycaceae Plocospermataceae Bruniaceae Bonnetiaceae Vochysiaceae Vahliaceae Carlemanniaceae Cardiopteridaceae Columelliaceae Ctenolophonaceae Sapindales Dumort. Garryales Lindl. [+Desfontainiaceae] Cynomoriaceae Anacardiaceae Aucubaceae Eremosynaceae Cytinaceae Biebersteiniaceae Eucommiaceae Escalloniaceae Dipentodontaceae Burseraceae Garryaceae Icacinaceae Elatinaceae Kirkiaceae Oncothecaceae Polyosmaceae Geissolomataceae Meliaceae Sphenostemonaceae Hoplestigmataceae Nitrariaceae Gentianales Lindl. Tribelaceae Kaliphoraceae [+Peganaceae] Apocynaceae Lepidobotryaceae Rutaceae Gelsemiaceae Apiales Nakai Lissocarpaceae Sapindaceae Gentianaceae Apiaceae Lophopyxidaceae Simaroubaceae Loganiaceae Araliaceae Medusandraceae Rubiaceae Aralidiaceae Metteniusaceae Griseliniaceae Mitrastemonaceae ASTERIDS Lamiales Bromhead Melanophyllaceae Paracryphiaceae Acanthaceae Pittosporaceae Pentaphylacaceae Cornales Dumort. Avicenniaceae Torricelliaceae Peridiscaceae Cornaceae Bignoniaceae Plagiopteraceae [+Nyssaceae ] Buddlejaceae Aquifoliales Senft Pottingeriaceae Grubbiaceae Byblidaceae Aquifoliaceae Sladeniaceae Hydrangeaceae Cyclocheilaceae Helwingiaceae Strasburgeriaceae Hydrostachyaceae Gesneriaceae Phyllonomaceae Tepuianthaceae Loasaceae Lamiaceae Asterales Lindl. Lentibulariaceae Ericales Dumort. Myoporaceae Alseuosmiaceae Actinidiaceae Oleaceae Argophyllaceae Balsaminaceae Orobanchaceae Asteraceae Clethraceae Paulowniaceae Calyceraceae Cyrillaceae Pedaliaceae Campanulaceae Diapensiaceae [+Martyniaceae] [+Lobeliaceae] Ebenaceae Phrymaceae Carpodetaceae Ericaceae Plantaginaceae Donatiaceae Fouquieriaceae Schlegeliaceae Goodeniaceae Halesiaceae Scrophulariaceae Menyanthaceae Lecythidaceae Stilbaceae Pentaphragmataceae Phellinaceae Marcgraviaceae Tetrachondraceae Myrsinaceae Verbenaceae Rousseaceae
Pellicieraceae Stylidiaceae Polemoniaceae Solanales Dumort. Primulaceae Convolvulaceae Dipsacales Dumort. Roridulaceae Hydroleaceae Caprifoliaceae Sapotaceae Montiniaceae Diervillaceae Sarraceniaceae Solanaceae Dipsacaceae Styracaceae Sphenocleaceae Linnaeaceae Symplocaceae Morinaceae Ternstroemiaceae Valerianaceae Tetrameristaceae Theaceae
Theophrastaceae
7 8
ORDINAL SYNONYMS Connarales Takht. ex Reveal Elaeocarpales Takht. Haemodorales Hutch. Acanthales Lindl. Austrobaileyales Takht. ex Calycanthales Mart. = Cunoniales = Oxalidales = Commelinales = Lamiales Reveal = Laurales Convolvulales Dumort. Elatinales Nakai Haloragales Bromhead Acerales Lindl. - not accepted, family at Calycerales Takht. ex Reveal = Solanales - not accepted, family = Saxifragales = Sapindales beginning of system = Asterales Coriariales Lindl. unplaced Hamamelidales Griseb. Actinidiales Takht. ex Reveal Avenales Bromhead Campanulales Rchb. = Cucurbitales Elodeales Nakai = Saxifragales = Ericales = Poales = Asterales Corylales Dumort. = Alismatales Hanguanales R.Dahlgren ex Adoxales Nakai Balanopales Engl. Canellales Cronquist = Fagales Empetrales Raf. Reveal - not accepted, family under = Malpighiales - not accepted, family at Corynocarpales Takht. = Ericales = not accepted, family under euasterids II Balanophorales Dumort. beginning of system = Cucurbitales Eriocaulales Nakai commelinoids Aesculales Bromhead - not accepted, family Cannales Dumort. Crassulales Lindl. = Poales Helleborales Nakai = Sapindales unplaced = Zingiberales = Saxifragales Eucommiales Nemejc ex = Ranunculales Agavales Hutch. Balsaminales Lindl. Capparales Hutch. Crossosomatales Takht. ex Cronquist Helwingiales Takht. = Asparagales = Ericales = Brassicales Reveal = Garryales = Aquifoliales Alliales Traub Barbeyales Takht. & Reveal Caprifoliales Lindl. - not accepted, family under Euphorbiales Lindl. Himantandrales Doweld and = Asparagales = Rosales = Dipsacales rosids = Malpighiales Shevyryova Alstroemeriales Hutch. Batales Engl. Cardiopteridales Takht. Cunoniales Hutch. Eupomatiales Takht. ex Reveal = Magnoliales = Liliales = Brassicales - not accepted, family under = Oxalidales = Magnoliales Hippuridales Pulle ex Reveal Altingiales Doweld Begoniales Dumort. euasterids II Cyclanthales J.H.Schaffn. Eupteleales Hu ex Reveal = Lamiales = Saxifragales = Cucurbitales Carduales Small = Pandanales = Ranunculales Homaliales Bromhead Amaranthales Dumort. Berberidales Dumort. = Asterales Cymodoceales Nakai Euryalales H.L.Li = Malpighiales = Caryophyllales = Ranunculales Caricales L.D.Benson = Alismatales - not accepted, family at Hortensiales Griseb. Amaryllidales Bromhead Betulales Bromhead = Brassicales Cynomoriales Burnett beginning of system = Cornales = Asparagales = Fagales Cassiales Horan. - not accepted, family Ficales Dumort. Hydatellales Cronquist Ambrosiales Dumort. Biebersteiniales Takht. = Fabales unplaced = Rosales = Poales = Asterales = Sapindales Casuarinales Lindl. Cyperales Hutch. Flacourtiales Heinze Hydnorales Takht. ex Reveal Ammiales Small Bignoniales Lindl. = Fagales = Poales = Malpighiales - not accepted, family at = Apiales = Lamiales Celastrales Baskerville Cytinales Dumort. Fouquieriales Takht. ex Reveal beginning of system Amomales Lindl. Bixales Lindl. - not accepted, family under - not accepted, family = Ericales Hydrangeales Nakai = Zingiberales = Malvales eurosids I unplaced Francoales Takht. = Cornales Ancistrocladales Takht. Boraginales Dumort. Centrolepidales Takht. Daphnales Lindl. = Geraniales Hydrastidales Takht. = Caryophyllales - not accepted, family under = Poales = Malvales Frangulales Wirtg. = Ranunculales Annonales Lindl. euasterids I Cephalotales Nakai Daphniphyllales Pulle ex = Rosales Hydropeltidales (Bartl.) Spenn. = Magnoliales Brexiales Lindl. = Oxalidales Cronquist Galiales Bromhead - not accepted, family Anthobolales Dumort. - not accepted, family under Cercidiphyllales Hu ex Reveal = Saxifragales = Gentianales Nymphaeaceae at beginning = Santalales eurosids I = Saxifragales Datiscales Dumort. Geissolomatales Takht. ex of system Apocynales Bromhead Bromeliales Dumort. Chenopodiales Dumort. = Cucurbitales Reveal Hydrostachyales Diels ex = Gentianales - not accepted, family under = Caryophyllales Desfontainiales Takht. - not accepted, family Reveal Aponogetonales Hutch. commelinoids Chironiales Griseb. - not accepted, family under unplaced = Cornales = Alismatales Bruniales Dumort. = Gentianales euasterids II Gesneriales Dumort. Hypericales Dumort. Arales Dumort. - not accepted, family under Chloranthales A.C.Sm. ex J.- Diapensiales Engl. & Gilg = Lamiales = Malpighiales = Alismatales euasterids II F.Leroy = Ericales Glaucidiales Takht. ex Reveal Hypoxidales Takht. Araliales Reveal Brunoniales Lindl. - not accepted, family at Didymelales Takht. = Ranunculales = Asparagales = Apiales = Asterales beginning of system - not accepted, family under Globulariales Dumort. Icacinales Tiegh. ex Reveal Aralidiales Takht. ex Reveal Burmanniales Heinze Cinchonales Lindl. eudicots = Lamiales - not accepted, family under = Apiales = Dioscoreales = Gentianales Dilleniales Hutch. Goodeniales Lindl. euasterids II Aristolochiales Dumort. Burserales Baskerville Circaeasterales Takht. - not accepted, family under = Asterales Illiciales Hu ex Cronquist = Piperales = Sapindales = Ranunculales core eudicots Greyiales Takht. - not accepted, family at Asarales Horan. Butomales Hutch. Cistales Rchb. Dioncophyllales Takht. ex = Geraniales beginning of system = Piperales = Alismatales = Malvales Reveal Grossulariales Lindl. Iridales Raf. Asclepiadales Dumort. Buxales Takht. ex Reveal Citrales Dumort. = Caryophyllales = Saxifragales = Asparagales = Gentianales - not accepted, family under = Sapindales Diospyrales Prantl Gunnerales Takht. ex Reveal Ixiales Lindl. Asteliales Dumort. eudicots Cocosales Nakai = Ericales - not accepted, family under = Asparagales = Asparagales Byblidales Nakai ex Reveal = Arecales Droserales Griseb. core eudicots Jasminales Dumort. Atriplicales Horan. = Lamiales Colchicales Dumort. = Caryophyllales Gyrocarpales Dumort. = Lamiales = Caryophyllales Cactales Dumort. = Liliales Ebenales Engl. = Laurales Juglandales Dumort. Aucubales Takht. = Caryophyllales Combretales Baskerville = Ericales Gyrostemonales Takht. = Fagales = Garryales Callitrichales Dumort. = Myrtales Elaeagnales Bromhead = Brassicales Julianiales Engl. = Lamiales = Rosales = Sapindales 9 10
Juncaginales Hutch. Mitrastemonales Makino Paridales Dumort. Rhamnales Dumort. Sedales Rchb. Tropaeolales Takht. ex Reveal = Alismatales - not accepted, family = Liliales = Rosales = Saxifragales = Brassicales Juncales Dumort. unplaced Parnassiales Nakai Rhinanthales Dumort. Silenales Lindl. Turnerales Dumort. = Poales Monimiales Dumort. - not accepted, family under = Lamiales = Caryophyllales = Malpighiales Lacistematales Baskerville = Laurales eurosids I Rhizophorales Tiegh. ex Reveal Simmondsiales Reveal Typhales Dumort. = Malpighiales Moringales Nakai Passiflorales Dumort. = Malpighiales = Caryophyllales = Poales Lactoridales Takht. ex Reveal = Brassicales = Malpighiales Rhodorales Horan. Smilacales Lindl. Ulmales Lindl. = Piperales Myricales Engl. Penaeales Lindl. = Ericales = Liliales = Rosales Lardizabalales Loconte = Fagales = Myrtales Rhoipteleales Novák ex Reveal Stellariales Dumort. Urticales Dumort. = Ranunculales Myristicales Thomé Petiveriales Lindl. = Fagales = Caryophyllales = Rosales Lecythidales Cronquist = Magnoliales = Caryophyllales Roridulales Nakai Stylidiales Takht. ex Reveal Vacciniales Dumort. = Ericales Myrothamnales Nakai ex Petrosaviales Takht. = Ericales = Asterales = Ericales Leitneriales Engl. Reveal - not accepted, family under Rubiales Dumort. Styracales Bisch. Vallisneriales Nakai = Sapindales - not accepted, family under monocots = Gentianales = Ericales = Alismatales Lentibulariales Lindl. core eudicots Philydrales Dumort. Ruppiales Nakai Taccales Dumort. Velloziales R.Dahlgren ex = Lamiales Myrsinales Spenn. = Commelinales = Alismatales = Dioscoreales Reveal Ligustrales Bartl. ex Bisch. = Ericales Physenales Takht. Rutales Perleb Tamales Dumort. = Pandanales = Lamiales Najadales Dumort. = Caryophyllales = Sapindales = Dioscoreales Veratrales Dumort. Limnanthales Nakai = Alismatales Pinguiculales Dumort. Sabiales Takht. Tamaricales Hutch. = Liliales = Brassicales Narcissales Dumort. = Lamiales = not accepted, family under = Caryophyllales Verbenales Horan. Linales Baskerville = Asparagales Pittosporales Lindl. eudicots Tecophilaeales Traub ex Reveal = Lamiales = Malpighiales Nartheciales Reveal & Zomlefer = Apiales Salicales Lindl. = Asparagales Viburnales Dumort. Loasales Bessey - not accepted, family under Plantaginales Lindl. = Malpighiales Theales Lindl. - not accepted, family under = Cornales monocots = Lamiales Salvadorales R.Dahlgren ex = Ericales euasterids II Loganiales Lindl. Nelumbonales Reveal Platanales J.H.Schaffn. Reveal Theligonales Nakai Vincales Horan. = Gentianales = Proteales = Proteales = Brassicales = Gentianales = Gentianales Lonicerales T.Liebe Nepenthales Dumort. Plumbaginales Lindl. Samolales Dumort. Thymelaeales Willk. Violales Perleb = Dipsacales = Caryophyllales = Caryophyllales = Ericales = Malvales = Malpighiales Loranthales Dumort. Nolanales Lindl. Podophyllales Dumort. Samydales Dumort. Tiliales Caruel Vitales Reveal = Santalales = Solanales = Ranunculales = Malpighiales = Malvales - not accepted, family under Lythrales Caruel Nyctaginales Dumort. Podostemales Lindl. Sanguisorbales Dumort. Tofieldiales Reveal & Zomlefer core eudicots = Myrtales = Caryophyllales = not accepted, family under = Rosales = Alismatales Vochysiales Dumort. Marathrales Dumort. Nymphaeales Dumort. rosids Sapotales Hook f. Torricelliales Takht. ex Reveal = Myrtales - not accepted, family = not accepted, family at Polemoniales Bromhead = Ericales = Apiales Winterales A.C.Sm. ex Reveal Podostemaceae under rosids beginning of system = Ericales Sarraceniales Bromhead Tovariales Nakai - not accepted, family at Mayacales Nakai Ochnales Hutch. ex Reveal Polygalales Dumort. = Ericales = Brassicales beginning of system - not accepted, family under = Malpighiales = Fabales Scheuchzeriales B.Boivin Trilliales Takht. Xyridales Lindl. commelinoids Oenotherales Bromhead Polygonales Dumort. = Alismatales = Liliales = Poales Medusagynales Takht. = Myrtales = Caryophyllales Scleranthales Dumort. Triuridales Hook.f. Zosterales Nakai = Malpighiales Olacales Benth. Pontederiales Hook f. = Caryophyllales - not accepted, family under = Alismatales Medusandrales Brenan = Santalales = Commelinales Scrophulariales Lindl. monocots Zygophyllales Takht. - not accepted, family Oleales Lindl. Portulacales Dumort. = Lamiales Trochodendrales Takht. ex - not accepted, family under unplaced = Lamiales = Caryophyllales Scyphostegiales Croizat Cronquist rosids Melanthiales R.Dahlgren ex Onagrales Rchb. Posidoniales Nakai = Malpighiales - not accepted, family under Reveal = Myrtales = Alismatales eudicots = Liliales Opuntiales Willk. Potamogetonales Dumort. Melastomatales Oliv. = Caryophyllales = Alismatales = Myrtales Orchidales Raf. Primulales Dumort. Meliales Lindl. = Asparagales = Ericales = Sapindales Paeoniales Heinze Quercales Burnett Menispermales Bromhead = Saxifragales = Fagales = Ranunculales Pandales Engl. & Gilg Rafflesiales Oliv. Menyanthales T.Yamaz. ex = Malpighiales - not accepted, family at Takht. Papaverales Dumort. beginning of system = Asterales = Ranunculales Resedales Dumort. Metteniusales Takht. Paracryphiales Takht. = Brassicales - not accepted, family - not accepted, family Restionales J.H.Schaffn. unplaced unplaced = Poales 11 12
SELECTED FAMILIAL SYNONYMS Napoleonaceae = Lecythidaceae Scepaceae = Euphorbiaceae Abrophyllaceae = Carpodetaceae Conostylidaceae = Haemodoraceae Gyrocarpaceae = Hernandiaceae Naucleaceae = Rubiaceae Schoepfiaceae = Olacaceae Acanthochlamydaceae = Cordiaceae = Boraginaceae Halophilaceae = Hydrocharitaceae Nectaropetalaceae = Erythroxylaceae Sclerophylacaceae = Solanaceae Velloziaceae Coridaceae = Primulaceae Halophytaceae = Amaranthaceae Nelsoniaceae = Acanthaceae Scoliopaceae = Liliaceae Aceraceae = Sapindaceae Corokiaceae = Argophyllaceae Hectorellaceae = Portulacaceae Nemacladaceae = Campanulaceae Scybaliaceae = Balanophoraceae Achradaceae = Sapotaceae Corylaceae = Betulaceae Heliotropiaceae = Boraginaceae Nesogenaceae = Cyclocheilaceae Scytopetalaceae = Lecythidaceae Aegicerataceae = Myrsinaceae Croomiaceae = Stemonaceae Heloniadaceae = Melanthiaceae Nolanaceae = Solanaceae Selaginaceae = Scrophulariaceae Agdestidaceae = Phytolaccaceae Cruciferae = Brassicaceae Helosidaceae = Balanophoraceae Nolinaceae = Convallariaceae Sesamaceae = Pedaliaceae Aitoniaceae = Meliaceae Curtisiaceae = Cornaceae Henriqueziaceae = Rubiaceae Nupharaceae = Nymphaeaceae Sesuviaceae = Aizoaceae Alangiaceae = Cornaceae Cuscutaceae = Convolvulaceae Hippocastanaceae = Sapindaceae Nyctanthaceae = Oleaceae Simethidaceae = Hemerocallidaceae Aloaceae = Asphodelaceae Cyananthaceae = Campanulaceae Hippocrateaceae = Celastraceae Nyssaceae = Cornaceae Siphonodontaceae = Celastraceae Alsinaceae = Caryophyllaceae Cyanastraceae = Tecophilaeaceae Hippuridaceae = Plantaginaceae Octoknemaceae = Olacaceae Sonneratiaceae = Lythraceae Ambrosiaceae = Asteraceae Cynocrambaceae nom. illeg. = Hortoniaceae = Monimiaceae Oftiaceae = Scrophulariaceae Spigeliaceae = Loganiaceae Amygdalaceae = Rosaceae Rubiaceae Hostaceae = Agavaceae Ophiopogonaceae = Convallariaceae Stenomeridaceae = Dioscoreaceae Androstachyaceae = Euphorbiaceae Cyphiaceae = Campanulaceae Humbertiaceae = Convolvulaceae Osyridaceae = Santalaceae Sterculiaceae = Malvaceae Antoniaceae = Loganiaceae Cyphocarpaceae = Campanulaceae Hydrastidaceae = Ranunculaceae Pachysandraceae = Buxaceae Stilaginaceae = Euphorbiaceae Apodanthaceae = Rafflesiaceae Cypripediaceae = Orchidaceae Hydrocotylaceae = Araliaceae Palmae = Arecaceae Strychnaceae = Loganiaceae Apostasiaceae = Orchidaceae Dactylanthaceae = Balanophoraceae Hydropeltidaceae = Nymphaeaceae Papilionaceae = Fabaceae Stylobasiaceae = Surianaceae Aptandraceae = Olacaceae Davidiaceae = Cornaceae Hydrophyllaceae = Boraginaceae Peganaceae = Nitrariaceae Stylocerataceae = Buxaceae Aristoteliaceae = Elaeocarpaceae Davidsoniaceae = Cunoniaceae Hymenocardiaceae = Euphorbiaceae Pentastemonaceae = Stemonaceae Symphoremataceae = Lamiaceae Asclepiadaceae = Apocynaceae Decaisneaceae = Lardizabalaceae Hypecoaceae = Papaveraceae Peperomiaceae = Piperaceae Syringaceae = Oleaceae Asteranthaceae = Lecythidaceae Desfontainiaceae = Columelliaceae Hypericaceae = Clusiaceae Periplocaceae = Apocynaceae Tetracentraceae = Trochodendraceae Averrhoaceae = Oxalidaceae Dialypetalanthaceae = Rubiaceae Hypseocharitaceae = Geraniaceae Peripterygiaceae = Cardiopteridaceae Tetradiclidaceae = Peganaceae Avetraceae = Dioscoreaceae Dianellaceae = Hemerocallidaceae Idiospermaceae = Calycanthaceae Petermanniaceae = Colchicaceae Tetragoniaceae = Aizoaceae Balanitaceae = Zygophyllaceae Dichondraceae = Convolvulaceae Illecebraceae = Caryophyllaceae Petiveriaceae = Phytolaccaceae Tetrastylidiaceae = Olacaceae Barbeiuaceae = Phytolaccaceae Diclidantheraceae = Polygalaceae Jasionaceae = Campanulaceae Philadelphaceae = Hydrangeaceae Thalassiaceae = Hydrocharitaceae Barclayaceae = Nymphaeaceae Diegodendraceae = Bixaceae Jasminiaceae = Oleaceae Phormiaceae = Hemerocallidaceae Theligonaceae = Rubiaceae Barringtoniaceae = Lecythidaceae Dionaeaceae = Droseraceae Johnsoniaceae = Hemerocallidaceae Phylicaceae = Rhamnaceae Thunbergiaceae = Acanthaceae Baueraceae = Cunoniaceae Dracaenaceae = Convallariaceae Julianiaceae = Anacardiaceae Picrodendraceae = Euphorbiaceae Tiliaceae = Malvaceae Baxteriaceae = Dasypogonaceae Duabangaceae = Lythraceae Kiggelariaceae = Flacourtiaceae Pinguiculaceae = Lentibulariaceae Trapaceae = Lythraceae Bembiciaceae = Flacourtiaceae Duckeodendraceae = Solanaceae Kingdoniaceae = Circaeasteraceae Pistaciaceae = Anacardiaceae Trapellaceae = Pedaliaceae Berzeliaceae = Bruniaceae Dulongiaceae nom. illeg. = Kirengeshomaceae = Hydrangeaceae Pistiaceae = Araceae Tribulaceae = Zygophyllaceae Bischofiaceae = Euphorbiaceae Phyllonomaceae Labiatae = Lamiaceae Platystemonaceae = Papaveraceae Tricyrtidaceae = Liliaceae Blepharocaryaceae = Anacardiaceae Dysphaniaceae = Amaranthaceae Langsdorffiaceae = Balanophoraceae Plumeriaceae = Apocynaceae Trilliaceae = Melanthiaceae Boerlagellaceae = Sapotaceae Ehretiaceae = Boraginaceae Leeaceae = Vitaceae Podoaceae = Anacardiaceae Triplostegiaceae = Valerianaceae Bombacaceae = Malvaceae Ellisiophyllaceae = Scrophulariaceae Leguminosae = Fabaceae Podophyllaceae = Berberidaceae Uapacaceae = Euphorbiaceae Boopidaceae = Calyceraceae Empetraceae = Ericaceae Leitneriaceae = Simaroubaceae Polygonanthaceae = Ullucaceae = Basellaceae Bretschneideraceae = Akaniaceae Epacridaceae = Ericaceae Lemnaceae = Araceae Anisophylleaceae Umbelliferae = Apiaceae Brexiaceae = Celastraceae Eremolepidaceae = Santalaceae Lenoaceae = Boraginaceae Potaliaceae = Gentianaceae Utriculariaceae = Lentibulariaceae Brunelliaceae = Cunoniaceae Eriospermaceae = Convallariaceae Leoniaceae = Violaceae Ptaeroxylaceae = Rutaceae Uvulariaceae = Colchicaceae Brunoniaceae = Goodeniaceae Erycibaceae = Convolvulaceae Lepuropetalaceae = Parnassiaceae Pteridophyllaceae = Papaveraceae Vacciniaceae = Ericaceae Bumeliaceae = Sapotaceae Erythropalaceae = Olacaceae Lilaeaceae = Juncaginaceae Punicaceae = Lythraceae Viburnaceae = Adoxaceae Burchardiaceae = Colchicaceae Eucryphiaceae = Cunoniaceae Limoniaceae = Plumbaginaceae Pyrolaceae = Ericaceae Viscaceae = Santalaceae Byttneriaceae = Malvaceae Euryalaceae = Nymphaeaceae Liriodendraceae = Magnoliaceae Ranzaniaceae = Berberidaceae Viticaceae = Lamiaceae Cabombaceae = Nymphaeaceae Exocarpaceae = Santalaceae Lobeliaceae = Campanulaceae Reaumuriaceae = Tamaricaceae Walleriaceae = Tecophilaeaceae Caesalpiniaceae = Fabaceae Flindersiaceae = Rutaceae Lomandraceae = Laxmanniaceae Retziaceae = Stilbaceae Wellstediaceae = Boraginaceae Calectasiaceae = Dasypogonaceae Foetidiaceae = Lecythidaceae Lophiraceae = Ochnaceae Rhinanthaceae = Orobanchaceae Xanthophyllaceae = Polygalaceae Callitrichaceae = Plantaginaceae Frangulaceae = Rhamnaceae Lophophytaceae = Balanophoraceae Rhodoleiaceae = Hamamelidaceae Xerophyllaceae = Melanthiaceae Calochortaceae = Liliaceae Fumariaceae = Papaveraceae Luxemburgiaceae = Ochnaceae Rhopalocarpaceae = Zannichelliaceae = Camelliaceae = Theaceae Funkiaceae = Agavaceae Malaceae = Rosaceae Sphaerosepalaceae Potamogetonaceae Canotiaceae = Celastraceae Galacaceae = Diapensiaceae Martyniaceae = Pedaliaceae Rhynchothecaceae = Ledocarpaceae Cansjeraceae = Opiliaceae Geitonoplesiaceae = Mastixiaceae = Cornaceae Roxburghiaceae = Stemonaceae Capparaceae = Brassicaceae Hemerocallidaceae Medeolaceae = Liliaceae Ruscaceae = Convallariaceae Carduaceae = Asteraceae Geniostomaceae = Loganiaceae Meliosmaceae = Sabiaceae Saccifoliaceae = Gentianaceae Cassythaceae = Lauraceae Geosiridaceae = Iridaceae Mendonciaceae = Acanthaceae Salaciaceae = Celastraceae Chailletiaceae = Dichapetalaceae Gisekiaceae = Phytolaccaceae Mesembryanthemaceae = Aizoaceae Salicorniaceae = Amaranthaceae Chenopodiaceae = Amaranthaceae Glaucidiaceae = Ranunculaceae Mimosaceae = Fabaceae Salpiglossidaceae = Solanaceae Chionographidaceae = Globulariaceae = Plantaginaceae Monotaceae = Dipterocarpaceae Sambucaceae = Adoxaceae Melanthiaceae Goetzeaceae = Solanaceae Monotropaceae = Ericaceae Samolaceae = Primulaceae Chloanthaceae = Lamiaceae Gonystylaceae = Thymelaeaceae Mouririaceae = Memecylaceae Saniculaceae = Apiaceae Cichoriaceae = Asteraceae Gouaniaceae = Rhamnaceae Moutabeaceae = Polygalaceae Sarcophytaceae = Balanophoraceae Cleomaceae = Brassicaceae Gramineae = Poaceae Myriophyllaceae = Haloragaceae Sarcospermataceae = Sapotaceae Cneoraceae = Rutaceae Gronoviaceae = Loasaceae Mystropetalaceae = Balanophoraceae Sargentodoxaceae = Lardizabalaceae Cobaeaceae = Polemoniaceae Gustaviaceae = Lecythidaceae Najadaceae = Hydrocharitaceae Saurauiaceae = Actinidiaceae Compositae = Asteraceae Guttiferae = Clusiaceae Nandinaceae = Berberidaceae Sauvagesiaceae = Ochnaceae Scaevolaceae = Goodeniaceae 13
Ceratophyllales 100/75 Laurales 100/54 Magnoliales 95/Š Piperales
99/55 Acorales 95/88
monocots Alismatales 58/59 99/66 96/100 Asparagales 99/83 Dioscoreales 100/100 99/68
angiosperms Pandanales 84/87 Liliales 100/99 Arecales 68/Š 67/Š Š/Š 95/Š Poales 82/Š Commelinales commelinoids 100/83 Zingiberales 98/88 85/Š Ranunculales 100/99 Proteales 100/80 100/99 Caryophyllales eudicots Santalales 98/Š 80/Š Saxifragales 59/Š 100/72 Geraniales 100/Š Malpighiales 77/55 Oxalidales 98/95 68/Š 100/77 Fabales 99/58 eurosid I rosids Š/Š Rosales 100/Š 100/94 core eudicots 60/Š Cucurbitales 100/89 Fagales 100/99 Myrtales 100/94 95/Š Brassicales 100/94 eurosid II 51/Š Malvales 100/98 Sapindales 98/Š 98/85 Cornales 82/98 Ericales 100/Š Garryales asterids 56/89 100/99 Gentianales 99/Š 100/96 euasterid I Lamiales 89/Š 96/Š Solanales 100/92 Aquifoliales 88/Š 100/22 Apiales 97/Š 94/85 euasterid II 86/96 Asterales Dipsacales
Figure 1. Phylogenetic interrelationships of the orders of flowering plants, compiled from recent cladistic analyses cited in the text. Jackknife support is given on the branches (a dash for values <50%), first jackknife values from analysis of 545 sequences of the rbcL, atpB, and 18S rDNA genes (D. E. Soltis, M. W. Chase, P. S. Soltis, D. Albach, M. E. Mort, V. Savolainen, M. Zanis, and J. S. Farris, unpublished, in preparation) and second jackknife values from analysis of 2538 rbcL sequences (Källersjö et al., 1998).