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Flying Dromaeosaurs
A winged, but flightless, Deinonychus by Stephen A. Czerkas FLYING DROMAEOSAURS Stephen A. Czerkas, Dianshuang Zhang, Jinglu Li and Yinxian Li The Dinosaur Museum, 754 South 200 West, Blanding, Utah 84511, USA; Liaoning Provincial Bureau of Land Resources Management, Liaoning Fossil Administration Office, and Liaoning Museum of Paleontology, Left of Nanshan Park, Beipiao, Liaoning Province 122100, People’s Republic of China. The Dinosaur Museum © 2002 Abstract Dromaeosaurs have been regarded as theropod dinosaurs that were among the closest avian ancestors which were strictly terrestrial having not yet evolved the ability to fly. Consequently, phylogenetic analyses have resulted in the claims of birds having evolved from “the ground up” within a dinosaurian ancestry. Though widely accepted, the relationship between birds and dinosaurs has remained highly controversial and disputed by advocates of birds as having been derived from an arboreal, non-dinosaurian type of archosaur. The cladistical interpretation of the dinosaur/bird relationship hinges upon the presumption of the dromaeosaurs inability to fly. Recent discoveries of dromaeosaurs have revealed impressions of feathers and avian characters in the skeleton that nearly equal and even surpass that of Archaeopteryx. Yet despite this, the ability to fly has been discounted due to the shorter length of the forelimbs. Described below are two such dromaeosaurs, but preserved with impressions of primary flight feathers extending from the manus which demonstrate an undeniable correlation towards the ability to fly. This compelling evidence refutes the popular interpretation of birds evolving from dinosaurs by revealing that dromaeosaurs were already birds and not the non-avian theropod dinosaurs as previously believed. -
Mesozoic—Dinos!
MESOZOIC—DINOS! VOLUME 9, ISSUE 8, APRIL 2020 THIS MONTH DINOSAURS! • Dinosaurs ○ What is a Dinosaur? page 2 DINOSAURS! When people think paleontology, ○ Bird / Lizard Hip? page 5 they think of scientists ○ Size Activity 1 page 10 working in the hot sun of ○ Size Activity 2 page 13 Colorado National ○ Size Activity 3 page 43 Monument or the Badlands ○ Diet page 46 of South Dakota and ○ Trackways page 53 Wyoming finding enormous, ○ Colorado Fossils and fierce, and long-gone Dinosaurs page 66 dinosaurs. POWER WORDS Dinosaurs safely evoke • articulated: fossil terror. Better than any bones arranged in scary movie, these were Articulated skeleton of the Tyrannosaurus rex proper order actually living breathing • endothermic: an beasts! from the American Museum of Natural History organism produces body heat through What was the biggest dinosaur? be reviewing the information metabolism What was the smallest about dinosaurs, but there is an • metabolism: chemical dinosaur? What color were interview with him at the end of processes that occur they? Did they live in herds? this issue. Meeting him, you will within a living organism What can their skeletons tell us? know instantly that he loves his in order to maintain life What evidence is there so that job! It doesn’t matter if you we can understand more about become an electrician, auto CAREER CONNECTION how these animals lived. Are mechanic, dancer, computer • Meet Dr. Holtz, any still alive today? programmer, author, or Dinosaur paleontologist, I truly hope that Paleontologist! page 73 To help us really understand you have tremendous job more about dinosaurs, we have satisfaction, like Dr. -
The Geologic Time Scale Is the Eon
Exploring Geologic Time Poster Illustrated Teacher's Guide #35-1145 Paper #35-1146 Laminated Background Geologic Time Scale Basics The history of the Earth covers a vast expanse of time, so scientists divide it into smaller sections that are associ- ated with particular events that have occurred in the past.The approximate time range of each time span is shown on the poster.The largest time span of the geologic time scale is the eon. It is an indefinitely long period of time that contains at least two eras. Geologic time is divided into two eons.The more ancient eon is called the Precambrian, and the more recent is the Phanerozoic. Each eon is subdivided into smaller spans called eras.The Precambrian eon is divided from most ancient into the Hadean era, Archean era, and Proterozoic era. See Figure 1. Precambrian Eon Proterozoic Era 2500 - 550 million years ago Archaean Era 3800 - 2500 million years ago Hadean Era 4600 - 3800 million years ago Figure 1. Eras of the Precambrian Eon Single-celled and simple multicelled organisms first developed during the Precambrian eon. There are many fos- sils from this time because the sea-dwelling creatures were trapped in sediments and preserved. The Phanerozoic eon is subdivided into three eras – the Paleozoic era, Mesozoic era, and Cenozoic era. An era is often divided into several smaller time spans called periods. For example, the Paleozoic era is divided into the Cambrian, Ordovician, Silurian, Devonian, Carboniferous,and Permian periods. Paleozoic Era Permian Period 300 - 250 million years ago Carboniferous Period 350 - 300 million years ago Devonian Period 400 - 350 million years ago Silurian Period 450 - 400 million years ago Ordovician Period 500 - 450 million years ago Cambrian Period 550 - 500 million years ago Figure 2. -
Recent Advances in Studies on Mesozoic and Paleogene Mammals in China
Vol.24 No.2 2010 Paleomammalogy Recent Advances in Studies on Mesozoic and Paleogene Mammals in China WANG Yuanqing* and NI Xijun Institute of Vertebrate Paleontology and Paleoanthropology, CAS, Beijing 10004, China ike in other fields of paleontology, research in from the articular of the lower jaw and the quadrate of the paleomammalogy mainly falls into two aspects. cranium following the process of reduction and detachment LOne is related to the biological nature of fossil from the reptilian mandible, are new elements of the bony mammals, such as their systematics, origin, evolution, chain in the mammalian middle ear. The appearance of phylogenetic relationships, transformation of key features mammalian middle ear allows mammals to hear the sound and paleobiogeography, and the other is related to the of higher frequencies than reptiles do. Generally speaking, geological context, involving biostratigraphy, biochronology, a widely accepted hypothesis is that mammals originated faunal turnover and its relations to the global and regional from a certain extinct reptilian group. The formation and environmental changes. In the last decade, a number of development of the definitive mammalian middle ear well-preserved mammalian specimens were collected from (DMME) has thus become one of the key issues in the study different localities around the country. Such discoveries have of mammalian evolution and has drawn the attention from provided significant information for understanding the early many researchers for many years. evolution of mammals and reconstructing the phylogeny of Developmental biological studies have proven the early mammals. function of the Meckel’s cartilage and its relationship to Studies of Chinese Mesozoic mammals achieved the ear ossicles during the development of mammalian remarkable progress in the past several years. -
Memorial to Carl Owen Dunbar 1891-1979 KARL M
Memorial to Carl Owen Dunbar 1891-1979 KARL M. WAAGE Peabody Museum, Yale University, P.O. Box 6666, New Haven, Connecticut 06511 Carl Owen Dunbar died suddenly at the age of 88 on April 7, 1979, while at the home of his son, just across the street from his own home in Dunedin, Florida. He is survived by his son, Carl Owen Dunbar, Jr.; his daughter, Mrs. Lora Louise Johnson of Atlanta, Geor gia; six grandchildren, and three great-grandchildren. His wife, Lora Beamer Dunbar, whom he married at Lawrence, Kansas, in September of 1914, predeceased him in December 1978. Carl was born January 1, 1891, to David and Emma Thomas Dunbar on their wheat ranch near Hal- lowell, Cherokee County, Kansas. The Dunbars were pioneer wheat farmers in Kansas Territory and among the first in that area to use powered farm machinery. From his early youth, Carl commonly ran the machines and, as he matured, served as foreman of the hired crews. Graduating from Cherokee County High School in 1909, he entered the University of Kansas the following Sep tember. Early in his undergraduate days a visiting uncle asked whether he would study any geology. “What’s geology? I never heard of it,” replied Carl. The uncle confessed that he didn’t know much about it himself, but he had a friend who was a geologist, and he thought it was great. According to Carl, “Three days later, I was enrolled in a geology course, and the instructor was W. H. Twenhofel.” Recruitment through the inspiration and encouragement of one interested individual is a familiar pattern in our profession. -
Uncorking the Bottle: What Triggered the Paleocene/Eocene Thermal Maximum Methane Release? Miriame
PALEOCEANOGRAPHY, VOL. 16, NO. 6, PAGES 549-562, DECEMBER 2001 Uncorking the bottle: What triggered the Paleocene/Eocene thermal maximum methane release? MiriamE. Katz,• BenjaminS. Cramer,Gregory S. Mountain,2 Samuel Katz, 3 and KennethG. Miller,1,2 Abstract. The Paleocene/Eocenethermal maximum (PETM) was a time of rapid global warming in both marine and continentalrealms that has been attributed to a massivemethane (CH4) releasefrom marine gas hydrate reservoirs. Previously proposedmechanisms for thismethane release rely on a changein deepwatersource region(s) to increasewater temperatures rapidly enoughto trigger the massivethermal dissociationof gas hydratereservoirs beneath the seafloor.To establish constraintson thermaldissociation, we modelheat flow throughthe sedimentcolumn and showthe effectof the temperature changeon the gashydrate stability zone throughtime. In addition,we provideseismic evidence tied to boreholedata for methanerelease along portions of the U.S. continentalslope; the releasesites are proximalto a buriedMesozoic reef front. Our modelresults, release site locations, published isotopic records, and oceancirculation models neither confirm nor refute thermaldissociation as the triggerfor the PETM methanerelease. In the absenceof definitiveevidence to confirmthermal dissociation,we investigatean altemativehypothesis in which continentalslope failure resulted in a catastrophicmethane release.Seismic and isotopic evidence indicates that Antarctic source deepwater circulation and seafloor erosion caused slope retreatalong -
The Triassic Period and the Beginning of the Mesozoic Era
Readings and Notes An Introduction to Earth Science 2016 The Triassic Period and the Beginning of the Mesozoic Era John J. Renton Thomas Repine Follow this and additional works at: https://researchrepository.wvu.edu/earthscience_readings Part of the Geology Commons C\.\- \~ THE TRIASSIC PERIOD and the BEGINNING OF THE MESOZOIC ERA Introduction to the Mesozoic Era: The Triassic Period is the first period of the Mesozoic Era, a span of time from 245 million years ago to 66 million years ago. Although the Mesozoic era commonly known as the "Age of the Dinosaurs", it should be pointed out that there were other important evolutionary developments taking place such as the appearance of the first mammal birds and flowering plans. The onset of the Mesozoic Era, the Triassic Period, was also a time of profound tectonic activity affecting the entire North American craton. In the east, the primary event was the breakup of Pangea and the formation of the Atlantic Ocean. In the west, it was the formation ofan Andean-type continental margin as the newly-formed continent of North America rapidly moved westward in response to the opening of the Atlantic Ocean coupled with the addition of exotic terranes to the western margin of the continent.. As the Atlantic oceanic ridge rose, the volume of ocean waters that was displaced was sufficient to result in the most extensive flooding of the continent by an epeiric sea since the Paleozoic; a sea whose presence was recorded by the accumulation of extensive carbonates throughout the continental interior. In the oceans, new life forms evolved to fill the vacancies brought about by the Permian extinction. -
Dinosaurs, by William Diller Matthew
The Project Gutenberg eBook of Dinosaurs, by William Diller Matthew http://www.gutenberg.org/files/19302/19302-h/19302-h.htm The[10][11][12][13][14][15][16][17][18][19][20][21][22][1][2][3][4][5][6][7][8][9] Project Gutenberg eBook, Dinosaurs, by T[100][103][104][105][106][108][109][110][111][112][113][115][117][118][119][120][121][123][124][125][126][127][128][129][130][131][132][133][134][135][136][137][138][139][140][141][142][143][144][145][146][147][148][149][150][151][152][153][154][155][156][157][158][159][161][162][10][11][12][13][14][15][17][18][19][20][21][22][23][24][26][27][28][29][30][31][34][35][36][38][39][40][41][42][43][44][45][46][47][48][49][50][51][52][53][54][55][56][57][58][61][63][64][65][66][67][68][69][70][71][72][73][74][76][77][78][79][80][81][83][84][85][86][87][88][90][91][93][94][95][96][97][98][99]OC William Diller Matthew This eBook is for the use of anyone anywhere at no cost and with almost no restrictions whatsoever. You may copy it, give it away or re-use it under the terms of the Project Gutenberg License included with this eBook or online at www.gutenberg.org Title: Dinosaurs With Special Reference to the American Museum Collections Author: William Diller Matthew Release Date: September 16, 2006 [eBook #19302] Language: English Character set encoding: ISO-8859-1 ***START OF THE PROJECT GUTENBERG EBOOK DINOSAURS*** E-text prepared by Brian Janes, Suzanne Lybarger, Jeannie Howse, and the Project Gutenberg Online Distributed Proofreading Team http://www.pgdp.net) Transcriber's Note: Click the image to see a larger version. -
Paravian Phylogeny and the Dinosaur-Bird Transition: an Overview
feart-06-00252 February 11, 2019 Time: 17:42 # 1 REVIEW published: 12 February 2019 doi: 10.3389/feart.2018.00252 Paravian Phylogeny and the Dinosaur-Bird Transition: An Overview Federico L. Agnolin1,2,3*, Matias J. Motta1,3, Federico Brissón Egli1,3, Gastón Lo Coco1,3 and Fernando E. Novas1,3 1 Laboratorio de Anatomía Comparada y Evolución de los Vertebrados, Museo Argentino de Ciencias Naturales Bernardino Rivadavia, Buenos Aires, Argentina, 2 Fundación de Historia Natural Félix de Azara, Universidad Maimónides, Buenos Aires, Argentina, 3 Consejo Nacional de Investigaciones Científicas y Técnicas, Buenos Aires, Argentina Recent years witnessed the discovery of a great diversity of early birds as well as closely related non-avian theropods, which modified previous conceptions about the origin of birds and their flight. We here present a review of the taxonomic composition and main anatomical characteristics of those theropod families closely related with early birds, with the aim of analyzing and discussing the main competing hypotheses pertaining to avian origins. We reject the postulated troodontid affinities of anchiornithines, and the Edited by: dromaeosaurid affinities of microraptorians and unenlagiids, and instead place these Corwin Sullivan, University of Alberta, Canada groups as successive sister taxa to Avialae. Aiming to evaluate previous phylogenetic Reviewed by: analyses, we recoded unenlagiids in the traditional TWiG data matrix, which resulted Thomas Alexander Dececchi, in a large polytomy at the base of Pennaraptora. This indicates that the TWiG University of Pittsburgh, United States phylogenetic scheme needs a deep revision. Regarding character evolution, we found Spencer G. Lucas, New Mexico Museum of Natural that: (1) the presence of an ossified sternum goes hand in hand with that of ossified History & Science, United States uncinate processes; (2) the presence of foldable forelimbs in basal archosaurs indicates *Correspondence: widespread distribution of this trait among reptiles, contradicting previous proposals Federico L. -
Written in Stone
112 W RITTENIN S TONE had yet been undertaken (the fossil had only come to the attention of Canadian paleontologist Phil Currie and paleo-artist Michael Skrepnick two weeks earlier), the specimen confirmed the connection between dino- saurs and birds that had been proposed on bones alone. The new dino- saur was dubbed Sinosauropteryx, and it had come from Cretaceous deposits in China that exhibited a quality of preservation that exceeded that of the Solnhofen limestone. Sinosauropteryx was only the first feathered dinosaur to be an- nounced. A panoply of feathered fossils started to turn up in the Jurassic and Cretaceous strata of China, each just as magnificent as the one be- fore. There were early birds that still retained clawed hands (Confuciu- sornis) and teeth (Sapeornis, Jibeinia), while non-flying coelurosaurs such as Caudipteryx, Sinornithosaurus, Jinfengopteryx, Dilong, and Beipiaosaurus wore an array of body coverings from wispy fuzz to full flight feathers. The fossil feathers of the strange, stubby-armed dinosaur Shuvuuia even preserved the biochemical signature of beta-keratin, a protein present in the feathers of living birds, and quill knobs on the forearm of Velociraptor reported in 2007 confirmed that the famous predator was covered in feathers, too. As new discoveries continued to accumulate it became apparent that almost every group of coelurosaurs had feathered representatives, from the weird secondarily herbivorous forms such as Beipiaosaurus to Dilong, an early relative of Tyrannosaurus. It is even possible that, FIGURE 37 - A Velociraptor attempts to catch the early bird Confuciusornis. Both were feathered dinosaurs. 113 Footprints and Feathers during its early life, the most famous of the flesh-tearing dinosaurs may have been covered in a coat of dino-fuzz. -
Dinosaurs.Pdf
DINOSAURS COMPILED BY HOWIE BAUM THIS IS THE FIRST OF 3 PAGES THAT HELP TO PUT THE 165 MILLION YEAR REIGN OF THE DINOSAURS, IN PERSPECTIVE. IT IS A CALENDAR THAT IS DESIGNED SO THAT ALL OF THE TIME, SINCE THE BIG BANG HAPPENED – 13.8 BILLION YEAR AGO, UP TO THE PRESENT TIME, IS COMPRESSED INTO 1 YEAR. period First Dinosaurs Dinosaurs lived for 165 million years !! Humans have only existed for 0.004% of the age of the Earth ! The rocks of the Cincinnatian series were deposited between approximately 451 and 443 million years ago, during the Ordovician period, when all the land was underwater ! BASIC DINOSAUR FACTS •Dinosaurs are a group of reptiles that have lived on Earth for about 165 million years. About 60% of dinosaurs ate plants (herbivores) and 40% ate meat (carnivores) •In 1842, the English naturalist Sir Richard Owen coined the term Dinosauria, derived from the Greek deinos, meaning “fearfully great,” and sauros, meaning “lizard.” •Dinosaur fossils have been found on all 7 continents. •All non-avian (non-bird) dinosaurs went extinct about 66 million years ago (MYA). •There are roughly 700 known species of extinct dinosaurs. •Modern birds are a kind of dinosaur because they share a common ancestor with non-avian dinosaurs. The Archosaurs consist of a diverse group of Triassic living and extinct reptiles that are subdivided into crocodiles, the 2 main types of dinosaurs, and pterosaurs. ORNITHISCHIAN AND SAURISCHIAN DINOSAURS As shown on the previous diagram, there are 2 types of Dinosaurs. Dinosaurs and reptiles have hip girdles, or pelvises, and all of them are composed of three bones: the ilium, ischium, and pubis. -
Testing the Neoflightless Hypothesis: Propatagium Reveals Flying Ancestry
J Ornithol DOI 10.1007/s10336-015-1190-9 ORIGINAL ARTICLE Testing the neoflightless hypothesis: propatagium reveals flying ancestry of oviraptorosaurs 1 2 Alan Feduccia • Stephen A. Czerkas Received: 4 September 2014 / Revised: 31 December 2014 / Accepted: 23 February 2015 Ó Dt. Ornithologen-Gesellschaft e.V. 2015 Abstract Considerable debate surrounds the numerous Zusammenfassung avian-like traits in core maniraptorans (ovirap- torosaurs, troodontids, and dromaeosaurs), especially in the Die ,,Neoflightless‘‘-Hypothese im Test: Halsflughaut Chinese Early Cretaceous oviraptorosaur Caudipteryx, (Propatagium) offenbart flugfa¨hige Vorfahren der which preserves modern avian pennaceous primary remi- Oviraptorosauria ges attached to the manus, as is the case in modern birds. Was Caudipteryx derived from earth-bound theropod di- Es gibt eine ausgiebige Debatte u¨ber die zahlreichen vo- nosaurs, which is the predominant view among palaeon- gela¨hnlichen Eigenheiten der Maniraptora (Oviraptosaurus, tologists, or was it secondarily flightless, with volant avians Troodontidae, Dromaeosaurus), vor allem des (gefiederten) or theropods as ancestors (the neoflightless hypothesis), Oviraptorosauria Caudipteryx aus der fru¨hen chinesischen which is another popular, but minority view. The discovery Kreidezeit, der genau wie rezente Vo¨gel Handschwingen here of an aerodynamic propatagium in several specimens hatte, die an den Handknochen ansetzen. Stammt Cau- provides new evidence that Caudipteryx (and hence ovi- dipteryx von den nur am Erdboden lebenden Theropoda ab - raptorosaurs) represent secondarily derived flightless die unter den Pala¨ontologen vorherrschende Meinung -, oder ground dwellers, whether of theropod or avian affinity, and war er sekunda¨r flugunfa¨hig und stammte von flugfa¨higen that their presence and radiation during the Cretaceous may Theropoden ab - die ,,Neoflightless‘‘-Hypothese, eine alter- have been a factor in the apparent scarcity of many other native, wenn auch nur von Wenigen unterstu¨tzte These.