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A Neoceratopsian Dinosaur from the Early Cretaceous of Mongolia And
ARTICLE https://doi.org/10.1038/s42003-020-01222-7 OPEN A neoceratopsian dinosaur from the early Cretaceous of Mongolia and the early evolution of ceratopsia ✉ Congyu Yu 1 , Albert Prieto-Marquez2, Tsogtbaatar Chinzorig 3,4, Zorigt Badamkhatan4,5 & Mark Norell1 1234567890():,; Ceratopsia is a diverse dinosaur clade from the Middle Jurassic to Late Cretaceous with early diversification in East Asia. However, the phylogeny of basal ceratopsians remains unclear. Here we report a new basal neoceratopsian dinosaur Beg tse based on a partial skull from Baruunbayan, Ömnögovi aimag, Mongolia. Beg is diagnosed by a unique combination of primitive and derived characters including a primitively deep premaxilla with four pre- maxillary teeth, a trapezoidal antorbital fossa with a poorly delineated anterior margin, very short dentary with an expanded and shallow groove on lateral surface, the derived presence of a robust jugal having a foramen on its anteromedial surface, and five equally spaced tubercles on the lateral ridge of the surangular. This is to our knowledge the earliest known occurrence of basal neoceratopsian in Mongolia, where this group was previously only known from Late Cretaceous strata. Phylogenetic analysis indicates that it is sister to all other neoceratopsian dinosaurs. 1 Division of Vertebrate Paleontology, American Museum of Natural History, New York 10024, USA. 2 Institut Català de Paleontologia Miquel Crusafont, ICTA-ICP, Edifici Z, c/de les Columnes s/n Campus de la Universitat Autònoma de Barcelona, 08193 Cerdanyola del Vallès Sabadell, Barcelona, Spain. 3 Department of Biological Sciences, North Carolina State University, Raleigh, NC 27695, USA. 4 Institute of Paleontology, Mongolian Academy of Sciences, ✉ Ulaanbaatar 15160, Mongolia. -
Valérie Martin, Varavudh Suteethorn & Eric Buffetaut, Description of the Type and Referred Material of Phuwiangosaurus
ORYCTOS, V ol . 2 : 39 - 91, Décembre 1999 DESCRIPTION OF THE TYPE AND REFERRED MATERIAL OF PHUWIANGOSAURUS SIRINDHORNAE MARTIN, BUFFETAUT AND SUTEETHORN, 1994, A SAUROPOD FROM THE LOWER CRETACEOUS OF THAILAND Valérie MARTIN 1, Varavudh SUTEETHORN 2 and Eric BUFFETAUT 3 1 Musée des Dinosaures, 11260 Espéraza, France 2 Geological Survey Division, Department of Mineral Resources, Rama VI Road, 10400 Bangkok, Thailand 3 CNRS (UMR 5561), 16 cour du Liégat, 75013 Paris, France Abstract : The type specimen of P. sirindhornae Martin, Buffetaut and Suteethorn, 1994 is an incomplete, partly articulated, skeleton discovered in the Phu Wiang area of northeastern Thailand). Most of the abundant sauropod material from the Sao Khua Formation (Early Cretaceous), collected on the Khorat Plateau, in northeastern Thailand, is referable to this species. Phuwiangosaurus is a middle-sized sauropod, which is clearly different from the Jurassic Chinese sauropods (Euhelopodidae). On the basis of a few jaw elements and teeth, P. sirindhornae may be considered as an early representative of the family Nemegtosauridae. Key words : Sauropoda, Osteology, Early Cretaceous, Thailand Description du type et du matériel rapporté de Phuwiangosaurus sirindhornae Martin, Buffetaut et Suteethorn, 1994, un sauropode du Crétacé inférieur de Thaïlande Résumé : Le spécimen type de Phuwiangosaurus sirindhornae est un squelette incomplet, partiellement articulé, découvert dans la région de Phu Wiang (Nord-Est de la Thaïlande). Phuwiangosaurus est un sauropode de taille moyenne (15 à 20 m de longueur) très différent des sauropodes du Jurassique chinois. La majeure partie de l’abondant matériel de sauropodes, récolté sur le Plateau de Khorat (Formation Sao Khua, Crétacé inférieur), est rap - portée à cette espèce. -
A New Raptorial Dinosaur with Exceptionally Long Feathering Provides Insights Into Dromaeosaurid flight Performance
ARTICLE Received 11 Apr 2014 | Accepted 11 Jun 2014 | Published 15 Jul 2014 DOI: 10.1038/ncomms5382 A new raptorial dinosaur with exceptionally long feathering provides insights into dromaeosaurid flight performance Gang Han1, Luis M. Chiappe2, Shu-An Ji1,3, Michael Habib4, Alan H. Turner5, Anusuya Chinsamy6, Xueling Liu1 & Lizhuo Han1 Microraptorines are a group of predatory dromaeosaurid theropod dinosaurs with aero- dynamic capacity. These close relatives of birds are essential for testing hypotheses explaining the origin and early evolution of avian flight. Here we describe a new ‘four-winged’ microraptorine, Changyuraptor yangi, from the Early Cretaceous Jehol Biota of China. With tail feathers that are nearly 30 cm long, roughly 30% the length of the skeleton, the new fossil possesses the longest known feathers for any non-avian dinosaur. Furthermore, it is the largest theropod with long, pennaceous feathers attached to the lower hind limbs (that is, ‘hindwings’). The lengthy feathered tail of the new fossil provides insight into the flight performance of microraptorines and how they may have maintained aerial competency at larger body sizes. We demonstrate how the low-aspect-ratio tail of the new fossil would have acted as a pitch control structure reducing descent speed and thus playing a key role in landing. 1 Paleontological Center, Bohai University, 19 Keji Road, New Shongshan District, Jinzhou, Liaoning Province 121013, China. 2 Dinosaur Institute, Natural History Museum of Los Angeles County, 900 Exposition Boulevard, Los Angeles, California 90007, USA. 3 Institute of Geology, Chinese Academy of Geological Sciences, 26 Baiwanzhuang Road, Beijing 100037, China. 4 University of Southern California, Health Sciences Campus, BMT 403, Mail Code 9112, Los Angeles, California 90089, USA. -
The Princeton Field Guide to Dinosaurs, Second Edition
MASS ESTIMATES - DINOSAURS ETC (largely based on models) taxon k model femur length* model volume ml x specific gravity = model mass g specimen (modeled 1st):kilograms:femur(or other long bone length)usually in decameters kg = femur(or other long bone)length(usually in decameters)3 x k k = model volume in ml x specific gravity(usually for whole model) then divided/model femur(or other long bone)length3 (in most models femur in decameters is 0.5253 = 0.145) In sauropods the neck is assigned a distinct specific gravity; in dinosaurs with large feathers their mass is added separately; in dinosaurs with flight ablity the mass of the fight muscles is calculated separately as a range of possiblities SAUROPODS k femur trunk neck tail total neck x 0.6 rest x0.9 & legs & head super titanosaur femur:~55000-60000:~25:00 Argentinosaurus ~4 PVPH-1:~55000:~24.00 Futalognkosaurus ~3.5-4 MUCPv-323:~25000:19.80 (note:downsize correction since 2nd edition) Dreadnoughtus ~3.8 “ ~520 ~75 50 ~645 0.45+.513=.558 MPM-PV 1156:~26000:19.10 Giraffatitan 3.45 .525 480 75 25 580 .045+.455=.500 HMN MB.R.2181:31500(neck 2800):~20.90 “XV2”:~45000:~23.50 Brachiosaurus ~4.15 " ~590 ~75 ~25 ~700 " +.554=~.600 FMNH P25107:~35000:20.30 Europasaurus ~3.2 “ ~465 ~39 ~23 ~527 .023+.440=~.463 composite:~760:~6.20 Camarasaurus 4.0 " 542 51 55 648 .041+.537=.578 CMNH 11393:14200(neck 1000):15.25 AMNH 5761:~23000:18.00 juv 3.5 " 486 40 55 581 .024+.487=.511 CMNH 11338:640:5.67 Chuanjiesaurus ~4.1 “ ~550 ~105 ~38 ~693 .063+.530=.593 Lfch 1001:~10700:13.75 2 M. -
Black Mesa Now Open New Dinosaur Discovered
THE UNIVERSITY OF OKLAHOMA TracksSpring 2011 Newsletter, Volume 23, Number 1 BLACK MESA NOW OPEN Whitten-Newman Foundation Funds Exhibit NEW DINOSAUR DISCOVERED Meet Brontomerus mcintoshi, “Thunder Thighs” ARCHAEOLOGY CURATOR TO RETIRE A Conversation with Don Wyckoff CONTENTS MUSEUM EVENTS AND HIGHLIGHTS DONATION: PAGE 3 EXHIBIT: PAGES 4-5 COLLECTIONS PAGES: 6-7 COMING UP: PAGE 11 NEWS: PAGES: 12-13 features & departments FROM THE DIRECTOR ………… 2 CURATOR ………………………… 8-9 NEWS ……………………………… 12-13 Thanks to Museum Members and Archaeology Curator Don Wyckoff Old Field Jacket Yields New Donors to Retire Species Wyckoff Receives Citation of Merit DONATION …………………………… 3 EDUCATION ……………………… 10 Redbud Café Welcomes New Whitten-Newman Foundation Funds Education Grants Fund Programs Chef Exhibit Former Curator Sidney DeVere UPCOMING EVENTS ………… 11 Brown Dies EXHIBIT ……………………………… 4-5 Art and the Animal Museum Welcomes 2011 Black Mesa Exhibit Open Summer Explorers Corporate Partners Baby Apatosaurus Longtime Volunteers Pass On COLLECTIONS ………………… 6-7 Bill Miller Named Volunteer of New ‘Thunder-Thighs’ Dinosaur the Year Discovered TRACKS, SPRING 2011 : VOLUME 23 NO. 1 1 FROM THE DIRECTOR Dear Members and Friends, On March 4 we had a very special ribbon cutting reception for our newest permanent exhibit in the museum—Black Mesa. We started this project in the 1990s, but the construction of this exhibit was made possible by a $1 million gift from Reggie and Rachelle Whitten and the Whitten-Newman Foundation. Located in the Hall of Natural Wonders, Black Mesa is a 1,500 square foot diorama of Oklahoma’s Panhandle, home to the highest elevation in the state. When you walk into the Hall of Natural Wonders, you can now view major habitats of our state from the forest to the prairie to the Black Mesa. -
Dinosaur Discovered (W/ Video) 23 February 2011
New 'thunder-thighs' dinosaur discovered (w/ Video) 23 February 2011 speculate that the larger specimen is the mother of the younger and would have weighed around 6 tons, about the size of a large elephant, and measured 14 meters in length. At a third of the size, the smaller specimen would have weighed about 200 kg, the size of a pony, and been 4.5 m long. The authors classified the new genus based on an incomplete skeleton including bones from the shoulder, hip, ribs, vertebrae and some unidentifiable fragments. They used the bones to identify Brontomerus' unique features, primarily the Brontomerus mcintoshi is a newly discovered dinosaur shape of the ilium (hip bone), which, in the case of from the Early Cretaceous of North America. The name Brontomerus means "Thunder thighs" -- a name chosen Brontomerus, is unusually large in comparison to because the peculiar shape of the hip bone shows that it that of similar dinosaurs. The wide, blade-shaped would have had enormously powerful thigh muscles in bone projects forward ahead of the hip socket, life. providing a proportionally massive area for the attachment of muscles. The shape of the bone indicates that the animal (PhysOrg.com) -- A new dinosaur named would likely have had the largest leg muscles of Brontomerus mcintoshi, or "thunder-thighs" after its any dinosaur in the sauropod family. This is enormously powerful thigh muscles, has been reflected in the name Brontomerus, which literally discovered in Utah, USA. The new species is means "thunder-thighs." The dinosaur's species described in a paper recently published in the name, mcintoshi, was chosen in honor of John journal Acta Palaeontologica Polonica by an "Jack" McIntosh, a retired physicist at Wesleyan international team of scientists from the U.K. -
New Tyrannosaur from the Mid-Cretaceous of Uzbekistan Clarifies Evolution of Giant Body Sizes and Advanced Senses in Tyrant Dinosaurs
Edinburgh Research Explorer New tyrannosaur from the mid-Cretaceous of Uzbekistan clarifies evolution of giant body sizes and advanced senses in tyrant dinosaurs Citation for published version: Brusatte, SL, Averianov, A, Sues, H, Muir, A & Butler, IB 2016, 'New tyrannosaur from the mid-Cretaceous of Uzbekistan clarifies evolution of giant body sizes and advanced senses in tyrant dinosaurs', Proceedings of the National Academy of Sciences, pp. 201600140. https://doi.org/10.1073/pnas.1600140113 Digital Object Identifier (DOI): 10.1073/pnas.1600140113 Link: Link to publication record in Edinburgh Research Explorer Document Version: Peer reviewed version Published In: Proceedings of the National Academy of Sciences General rights Copyright for the publications made accessible via the Edinburgh Research Explorer is retained by the author(s) and / or other copyright owners and it is a condition of accessing these publications that users recognise and abide by the legal requirements associated with these rights. Take down policy The University of Edinburgh has made every reasonable effort to ensure that Edinburgh Research Explorer content complies with UK legislation. If you believe that the public display of this file breaches copyright please contact [email protected] providing details, and we will remove access to the work immediately and investigate your claim. Download date: 04. Oct. 2021 Classification: Physical Sciences: Earth, Atmospheric, and Planetary Sciences; Biological Sciences: Evolution New tyrannosaur from the mid-Cretaceous of Uzbekistan clarifies evolution of giant body sizes and advanced senses in tyrant dinosaurs Stephen L. Brusattea,1, Alexander Averianovb,c, Hans-Dieter Suesd, Amy Muir1, Ian B. Butler1 aSchool of GeoSciences, University of Edinburgh, Edinburgh EH9 3FE, UK bZoological Institute, Russian Academy of Sciences, St. -
A Dinosaur Called Tiny Free
FREE A DINOSAUR CALLED TINY PDF Alan Durant,Jo Simpson | 32 pages | 06 Aug 2007 | HarperCollins Publishers | 9780007233908 | English | London, United Kingdom A Dinosaur Called Tiny by Alan Durant Members of its single species Compsognathus longipes could grow to around the size of a turkey. Paleontologists have found two well-preserved fossilsone in Germany in the s and the second in France more than a century later. Today, C. Many presentations still describe Compsognathus as "chicken-sized" dinosaurs because of the size of the German specimen, which is now believed to be a juvenile. Compsognathus longipes is one of the few dinosaur species whose diet is known with certainty: the remains of small, agile lizards are preserved in the bellies of both specimens. Teeth discovered in Portugal may be further fossil remains of the genus. Although not recognized as such at the time of its discovery, Compsognathus is the first theropod dinosaur known from a reasonably complete fossil skeleton. Until the s, it was the smallest-known non- avialan dinosaur, with the preceding centuries incorrectly labelling them as the closest relative of Archaeopteryx. However, dinosaurs discovered later, such as CaenagnathasiaMicroraptor and Parvicursorwere even smaller. The largest Compsognathus specimen is estimated to have weighed somewhere between A Dinosaur Called Tiny. Compsognathus were small, bipedal animals with long hind legs and longer tails, which they used for balance during locomotion. The forelimbs were smaller than the hindlimbs. The hand bore two large, clawed digits and a third, smaller A Dinosaur Called Tiny that may have been non-functional. The skull had five pairs of fenestrae skull openingsthe largest of which was for the orbit eye socket[7] with the eyes being larger in proportion to the rest of the skull. -
Implications for Predatory Dinosaur Macroecology and Ontogeny in Later Late Cretaceous Asiamerica
Canadian Journal of Earth Sciences Theropod Guild Structure and the Tyrannosaurid Niche Assimilation Hypothesis: Implications for Predatory Dinosaur Macroecology and Ontogeny in later Late Cretaceous Asiamerica Journal: Canadian Journal of Earth Sciences Manuscript ID cjes-2020-0174.R1 Manuscript Type: Article Date Submitted by the 04-Jan-2021 Author: Complete List of Authors: Holtz, Thomas; University of Maryland at College Park, Department of Geology; NationalDraft Museum of Natural History, Department of Geology Keyword: Dinosaur, Ontogeny, Theropod, Paleocology, Mesozoic, Tyrannosauridae Is the invited manuscript for consideration in a Special Tribute to Dale Russell Issue? : © The Author(s) or their Institution(s) Page 1 of 91 Canadian Journal of Earth Sciences 1 Theropod Guild Structure and the Tyrannosaurid Niche Assimilation Hypothesis: 2 Implications for Predatory Dinosaur Macroecology and Ontogeny in later Late Cretaceous 3 Asiamerica 4 5 6 Thomas R. Holtz, Jr. 7 8 Department of Geology, University of Maryland, College Park, MD 20742 USA 9 Department of Paleobiology, National Museum of Natural History, Washington, DC 20013 USA 10 Email address: [email protected] 11 ORCID: 0000-0002-2906-4900 Draft 12 13 Thomas R. Holtz, Jr. 14 Department of Geology 15 8000 Regents Drive 16 University of Maryland 17 College Park, MD 20742 18 USA 19 Phone: 1-301-405-4084 20 Fax: 1-301-314-9661 21 Email address: [email protected] 22 23 1 © The Author(s) or their Institution(s) Canadian Journal of Earth Sciences Page 2 of 91 24 ABSTRACT 25 Well-sampled dinosaur communities from the Jurassic through the early Late Cretaceous show 26 greater taxonomic diversity among larger (>50kg) theropod taxa than communities of the 27 Campano-Maastrichtian, particularly to those of eastern/central Asia and Laramidia. -
A Re-Evaluation of the Enigmatic Dinosauriform Caseosaurus Crosbyensis from the Late Triassic of Texas, USA and Its Implications for Early Dinosaur Evolution
A re-evaluation of the enigmatic dinosauriform Caseosaurus crosbyensis from the Late Triassic of Texas, USA and its implications for early dinosaur evolution MATTHEW G. BARON and MEGAN E. WILLIAMS Baron, M.G. and Williams, M.E. 2018. A re-evaluation of the enigmatic dinosauriform Caseosaurus crosbyensis from the Late Triassic of Texas, USA and its implications for early dinosaur evolution. Acta Palaeontologica Polonica 63 (1): 129–145. The holotype specimen of the Late Triassic dinosauriform Caseosaurus crosbyensis is redescribed and evaluated phylogenetically for the first time, providing new anatomical information and data on the earliest dinosaurs and their evolution within the dinosauromorph lineage. Historically, Caseosaurus crosbyensis has been considered to represent an early saurischian dinosaur, and often a herrerasaur. More recent work on Triassic dinosaurs has cast doubt over its supposed dinosaurian affinities and uncertainty about particular features in the holotype and only known specimen has led to the species being regarded as a dinosauriform of indeterminate position. Here, we present a new diagnosis for Caseosaurus crosbyensis and refer additional material to the taxon—a partial right ilium from Snyder Quarry. Our com- parisons and phylogenetic analyses suggest that Caseosaurus crosbyensis belongs in a clade with herrerasaurs and that this clade is the sister taxon of Dinosauria, rather than positioned within it. This result, along with other recent analyses of early dinosaurs, pulls apart what remains of the “traditional” group of dinosaurs collectively termed saurischians into a polyphyletic assemblage and implies that Dinosauria should be regarded as composed exclusively of Ornithoscelida (Ornithischia + Theropoda) and Sauropodomorpha. In addition, our analysis recovers the enigmatic European taxon Saltopus elginensis among herrerasaurs for the first time. -
Re-Description of the Sauropod Dinosaur Amanzia (“Ornithopsis
Schwarz et al. Swiss J Geosci (2020) 113:2 https://doi.org/10.1186/s00015-020-00355-5 Swiss Journal of Geosciences ORIGINAL PAPER Open Access Re-description of the sauropod dinosaur Amanzia (“Ornithopsis/Cetiosauriscus”) greppini n. gen. and other vertebrate remains from the Kimmeridgian (Late Jurassic) Reuchenette Formation of Moutier, Switzerland Daniela Schwarz1* , Philip D. Mannion2 , Oliver Wings3 and Christian A. Meyer4 Abstract Dinosaur remains were discovered in the 1860’s in the Kimmeridgian (Late Jurassic) Reuchenette Formation of Moutier, northwestern Switzerland. In the 1920’s, these were identifed as a new species of sauropod, Ornithopsis greppini, before being reclassifed as a species of Cetiosauriscus (C. greppini), otherwise known from the type species (C. stewarti) from the late Middle Jurassic (Callovian) of the UK. The syntype of “C. greppini” consists of skeletal elements from all body regions, and at least four individuals of diferent sizes can be distinguished. Here we fully re-describe this material, and re-evaluate its taxonomy and systematic placement. The Moutier locality also yielded a theropod tooth, and fragmen- tary cranial and vertebral remains of a crocodylomorph, also re-described here. “C.” greppini is a small-sized (not more than 10 m long) non-neosauropod eusauropod. Cetiosauriscus stewarti and “C.” greppini difer from each other in: (1) size; (2) the neural spine morphology and diapophyseal laminae of the anterior caudal vertebrae; (3) the length-to-height proportion in the middle caudal vertebrae; (4) the presence or absence of ridges and crests on the middle caudal cen- tra; and (5) the shape and proportions of the coracoid, humerus, and femur. -
A Diplodocid Sauropod Survivor from the Early Cretaceous of South America
A Diplodocid Sauropod Survivor from the Early Cretaceous of South America Pablo A. Gallina1,2*, Sebastia´n Apesteguı´a1,2, Alejandro Haluza3, Juan I. Canale1,3 1 CONICET, Buenos Aires, Argentina, 2 Fundacio´n de Historia Natural Fe´lix de Azara, Universidad Maimo´nides, Buenos Aires, Argentina, 3 Museo Paleontolo´gico Ernesto Bachmann, Villa El Choco´n, Neuque´n, Argentina Abstract Diplodocids are by far the most emblematic sauropod dinosaurs. They are part of Diplodocoidea, a vast clade whose other members are well-known from Jurassic and Cretaceous strata in Africa, Europe, North and South America. However, Diplodocids were never certainly recognized from the Cretaceous or in any other southern land mass besides Africa. Here we report a new sauropod, Leikupal laticauda gen. et sp. nov., from the early Lower Cretaceous (Bajada Colorada Formation) of Neuque´n Province, Patagonia, Argentina. This taxon differs from any other sauropod by the presence of anterior caudal transverse process extremely developed with lateroventral expansions reinforced by robust dorsal and ventral bars, very robust centroprezygapophyseal lamina in anterior caudal vertebra and paired pneumatic fossae on the postzygapophyses in anterior-most caudal vertebra. The phylogenetic analyses support its position not only within Diplodocidae but also as a member of Diplodocinae, clustering together with the African form Tornieria, pushing the origin of Diplodocoidea to the Middle Jurassic or even earlier. The new discovery represents the first record of a diplodocid for South America and the stratigraphically youngest record of this clade anywhere. Citation: Gallina PA, Apesteguı´a S, Haluza A, Canale JI (2014) A Diplodocid Sauropod Survivor from the Early Cretaceous of South America.