Osteological Revision of the Holotype of the Middle
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A Reassessment of the Phylogenetic Position of Cretaceous Sauropod Dinosaurs from Queensland, Australia
Asociación Paleontológica Argentina. Publicación Especial 7 ISSN 0328-347X VII International Symposium on Mesozoic Terrestrial Ecosystems: 139-144.Buenos Aires, 30-6-2001 A ReasSessmenT of the phylogenetic position of CretaceoUS SaUROPOd dinosaURS from Queensland, Australia Ralph E. MOLNAR1 Abstract. The Cretaceous sauropod material from Queensland, Australia, has been regarded as pertaining to a persistently primitive sauropod lineage (e.g., Coombs and Molnar). The specimens derive from the Toolebuc and Allaru (Albian marine) and Winton (Cenomanian continental) Formations. Recent phyloge- netic analyses carried out by workers in Argentina, the USA and England permit a reassessment of this fragmentary material. As far as can be ascertained from the material, there is no indication from the char- acter states that more than a single taxon is represented. Character states diagnostic of the Titanosauriforrnes, the Titanosauria, the Somphospondyli and the Titanosauridae are present. Thus the Queensland material does not pertain to cetiosaurids but belongs to titanosaurs, extending their range in- to Australia Key words. Sauropods. Austrosaurus. Titanosaurs. Cretaceous. Paleozoogeography. IntroductioN (1998),has made it possible to reassess the phyloge- netic affinities of the Australian Cretaceous sauropod By the 1950's titanosaurs were widely recognized material and address the anomalous absence of ti- both as the latest sauropod group to diversify and as tanosaurs. This paper looks specifically at pre-eminently the sauropods of Gondwanaland. -
Valérie Martin, Varavudh Suteethorn & Eric Buffetaut, Description of the Type and Referred Material of Phuwiangosaurus
ORYCTOS, V ol . 2 : 39 - 91, Décembre 1999 DESCRIPTION OF THE TYPE AND REFERRED MATERIAL OF PHUWIANGOSAURUS SIRINDHORNAE MARTIN, BUFFETAUT AND SUTEETHORN, 1994, A SAUROPOD FROM THE LOWER CRETACEOUS OF THAILAND Valérie MARTIN 1, Varavudh SUTEETHORN 2 and Eric BUFFETAUT 3 1 Musée des Dinosaures, 11260 Espéraza, France 2 Geological Survey Division, Department of Mineral Resources, Rama VI Road, 10400 Bangkok, Thailand 3 CNRS (UMR 5561), 16 cour du Liégat, 75013 Paris, France Abstract : The type specimen of P. sirindhornae Martin, Buffetaut and Suteethorn, 1994 is an incomplete, partly articulated, skeleton discovered in the Phu Wiang area of northeastern Thailand). Most of the abundant sauropod material from the Sao Khua Formation (Early Cretaceous), collected on the Khorat Plateau, in northeastern Thailand, is referable to this species. Phuwiangosaurus is a middle-sized sauropod, which is clearly different from the Jurassic Chinese sauropods (Euhelopodidae). On the basis of a few jaw elements and teeth, P. sirindhornae may be considered as an early representative of the family Nemegtosauridae. Key words : Sauropoda, Osteology, Early Cretaceous, Thailand Description du type et du matériel rapporté de Phuwiangosaurus sirindhornae Martin, Buffetaut et Suteethorn, 1994, un sauropode du Crétacé inférieur de Thaïlande Résumé : Le spécimen type de Phuwiangosaurus sirindhornae est un squelette incomplet, partiellement articulé, découvert dans la région de Phu Wiang (Nord-Est de la Thaïlande). Phuwiangosaurus est un sauropode de taille moyenne (15 à 20 m de longueur) très différent des sauropodes du Jurassique chinois. La majeure partie de l’abondant matériel de sauropodes, récolté sur le Plateau de Khorat (Formation Sao Khua, Crétacé inférieur), est rap - portée à cette espèce. -
Dino Cards Project D E F List B
Daanosaurus Efraasia Dacentrurus Einiosaurus "Dachongosaurus" – nomen nudum Ekrixinatosaurus Daemonosaurus Elachistosuchus – a rhynchocephalian Dahalokely Elaltitan Dakosaurus – a metriorhynchid crocodilian Elaphrosaurus Dakotadon Elmisaurus Dakotaraptor Elopteryx - nomen dubium Daliansaurus Elosaurus – junior synonym of Brontosaurus "Damalasaurus" – nomen nudum Elrhazosaurus Dandakosaurus - nomen dubium "Elvisaurus" – nomen nudum; Cryolophosaurus Danubiosaurus – junior synonym of Struthiosaurus Emausaurus "Daptosaurus" – nomen nudum; early manuscript name for Deinonychus Embasaurus - theropoda incertae sedis Darwinsaurus - possible junior synonym of Huxleysaurus Enigmosaurus Dashanpusaurus Eoabelisaurus Daspletosaurus Eobrontosaurus – junior synonym of Brontosaurus Dasygnathoides – a non-dinosaurian archosaur, junior synonym Eocarcharia of Ornithosuchus Eoceratops – junior synonym of Chasmosaurus "Dasygnathus" – preoccupied name, now known as Dasygnathoides Eocursor Datanglong Eodromaeus Datonglong "Eohadrosaurus" – nomen nudum; Eolambia Datousaurus Eolambia Daurosaurus – synonym of Kulindadromeus Eomamenchisaurus Daxiatitan Eoplophysis - Dinosauria indet. Deinocheirus Eoraptor Deinodon – possibly Gorgosaurus Eosinopteryx - Avialae Deinonychus Eotrachodon Delapparentia - probable junior synonym of Iguanodon Eotriceratops Deltadromeus Eotyrannus Demandasaurus Eousdryosaurus Denversaurus Epachthosaurus Deuterosaurus – a therapsid Epanterias – may be Allosaurus Diabloceratops "Ephoenosaurus" – nomen nudum; Machimosaurus (a crocodilian) Diamantinasaurus -
The Princeton Field Guide to Dinosaurs, Second Edition
MASS ESTIMATES - DINOSAURS ETC (largely based on models) taxon k model femur length* model volume ml x specific gravity = model mass g specimen (modeled 1st):kilograms:femur(or other long bone length)usually in decameters kg = femur(or other long bone)length(usually in decameters)3 x k k = model volume in ml x specific gravity(usually for whole model) then divided/model femur(or other long bone)length3 (in most models femur in decameters is 0.5253 = 0.145) In sauropods the neck is assigned a distinct specific gravity; in dinosaurs with large feathers their mass is added separately; in dinosaurs with flight ablity the mass of the fight muscles is calculated separately as a range of possiblities SAUROPODS k femur trunk neck tail total neck x 0.6 rest x0.9 & legs & head super titanosaur femur:~55000-60000:~25:00 Argentinosaurus ~4 PVPH-1:~55000:~24.00 Futalognkosaurus ~3.5-4 MUCPv-323:~25000:19.80 (note:downsize correction since 2nd edition) Dreadnoughtus ~3.8 “ ~520 ~75 50 ~645 0.45+.513=.558 MPM-PV 1156:~26000:19.10 Giraffatitan 3.45 .525 480 75 25 580 .045+.455=.500 HMN MB.R.2181:31500(neck 2800):~20.90 “XV2”:~45000:~23.50 Brachiosaurus ~4.15 " ~590 ~75 ~25 ~700 " +.554=~.600 FMNH P25107:~35000:20.30 Europasaurus ~3.2 “ ~465 ~39 ~23 ~527 .023+.440=~.463 composite:~760:~6.20 Camarasaurus 4.0 " 542 51 55 648 .041+.537=.578 CMNH 11393:14200(neck 1000):15.25 AMNH 5761:~23000:18.00 juv 3.5 " 486 40 55 581 .024+.487=.511 CMNH 11338:640:5.67 Chuanjiesaurus ~4.1 “ ~550 ~105 ~38 ~693 .063+.530=.593 Lfch 1001:~10700:13.75 2 M. -
The Origin and Early Evolution of Dinosaurs
Biol. Rev. (2010), 85, pp. 55–110. 55 doi:10.1111/j.1469-185X.2009.00094.x The origin and early evolution of dinosaurs Max C. Langer1∗,MartinD.Ezcurra2, Jonathas S. Bittencourt1 and Fernando E. Novas2,3 1Departamento de Biologia, FFCLRP, Universidade de S˜ao Paulo; Av. Bandeirantes 3900, Ribeir˜ao Preto-SP, Brazil 2Laboratorio de Anatomia Comparada y Evoluci´on de los Vertebrados, Museo Argentino de Ciencias Naturales ‘‘Bernardino Rivadavia’’, Avda. Angel Gallardo 470, Cdad. de Buenos Aires, Argentina 3CONICET (Consejo Nacional de Investigaciones Cient´ıficas y T´ecnicas); Avda. Rivadavia 1917 - Cdad. de Buenos Aires, Argentina (Received 28 November 2008; revised 09 July 2009; accepted 14 July 2009) ABSTRACT The oldest unequivocal records of Dinosauria were unearthed from Late Triassic rocks (approximately 230 Ma) accumulated over extensional rift basins in southwestern Pangea. The better known of these are Herrerasaurus ischigualastensis, Pisanosaurus mertii, Eoraptor lunensis,andPanphagia protos from the Ischigualasto Formation, Argentina, and Staurikosaurus pricei and Saturnalia tupiniquim from the Santa Maria Formation, Brazil. No uncontroversial dinosaur body fossils are known from older strata, but the Middle Triassic origin of the lineage may be inferred from both the footprint record and its sister-group relation to Ladinian basal dinosauromorphs. These include the typical Marasuchus lilloensis, more basal forms such as Lagerpeton and Dromomeron, as well as silesaurids: a possibly monophyletic group composed of Mid-Late Triassic forms that may represent immediate sister taxa to dinosaurs. The first phylogenetic definition to fit the current understanding of Dinosauria as a node-based taxon solely composed of mutually exclusive Saurischia and Ornithischia was given as ‘‘all descendants of the most recent common ancestor of birds and Triceratops’’. -
The Sauropodomorph Biostratigraphy of the Elliot Formation of Southern Africa: Tracking the Evolution of Sauropodomorpha Across the Triassic–Jurassic Boundary
Editors' choice The sauropodomorph biostratigraphy of the Elliot Formation of southern Africa: Tracking the evolution of Sauropodomorpha across the Triassic–Jurassic boundary BLAIR W. MCPHEE, EMESE M. BORDY, LARA SCISCIO, and JONAH N. CHOINIERE McPhee, B.W., Bordy, E.M., Sciscio, L., and Choiniere, J.N. 2017. The sauropodomorph biostratigraphy of the Elliot Formation of southern Africa: Tracking the evolution of Sauropodomorpha across the Triassic–Jurassic boundary. Acta Palaeontologica Polonica 62 (3): 441–465. The latest Triassic is notable for coinciding with the dramatic decline of many previously dominant groups, followed by the rapid radiation of Dinosauria in the Early Jurassic. Among the most common terrestrial vertebrates from this time, sauropodomorph dinosaurs provide an important insight into the changing dynamics of the biota across the Triassic–Jurassic boundary. The Elliot Formation of South Africa and Lesotho preserves the richest assemblage of sauropodomorphs known from this age, and is a key index assemblage for biostratigraphic correlations with other simi- larly-aged global terrestrial deposits. Past assessments of Elliot Formation biostratigraphy were hampered by an overly simplistic biozonation scheme which divided it into a lower “Euskelosaurus” Range Zone and an upper Massospondylus Range Zone. Here we revise the zonation of the Elliot Formation by: (i) synthesizing the last three decades’ worth of fossil discoveries, taxonomic revision, and lithostratigraphic investigation; and (ii) systematically reappraising the strati- graphic provenance of important fossil locations. We then use our revised stratigraphic information in conjunction with phylogenetic character data to assess morphological disparity between Late Triassic and Early Jurassic sauropodomorph taxa. Our results demonstrate that the Early Jurassic upper Elliot Formation is considerably more taxonomically and morphologically diverse than previously thought. -
Titanosauriform Teeth from the Cretaceous of Japan
“main” — 2011/2/10 — 15:59 — page 247 — #1 Anais da Academia Brasileira de Ciências (2011) 83(1): 247-265 (Annals of the Brazilian Academy of Sciences) Printed version ISSN 0001-3765 / Online version ISSN 1678-2690 www.scielo.br/aabc Titanosauriform teeth from the Cretaceous of Japan HARUO SAEGUSA1 and YUKIMITSU TOMIDA2 1Museum of Nature and Human Activities, Hyogo, Yayoigaoka 6, Sanda, 669-1546, Japan 2National Museum of Nature and Science, 3-23-1 Hyakunin-cho, Shinjuku-ku, Tokyo 169-0073, Japan Manuscript received on October 25, 2010; accepted for publication on January 7, 2011 ABSTRACT Sauropod teeth from six localities in Japan were reexamined. Basal titanosauriforms were present in Japan during the Early Cretaceous before Aptian, and there is the possibility that the Brachiosauridae may have been included. Basal titanosauriforms with peg-like teeth were present during the “mid” Cretaceous, while the Titanosauria with peg-like teeth was present during the middle of Late Cretaceous. Recent excavations of Cretaceous sauropods in Asia showed that multiple lineages of sauropods lived throughout the Cretaceous in Asia. Japanese fossil records of sauropods are conformable with this hypothesis. Key words: Sauropod, Titanosauriforms, tooth, Cretaceous, Japan. INTRODUCTION humerus from the Upper Cretaceous Miyako Group at Moshi, Iwaizumi Town, Iwate Pref. (Hasegawa et al. Although more than twenty four dinosaur fossil local- 1991), all other localities provided fossil teeth (Tomida ities have been known in Japan (Azuma and Tomida et al. 2001, Tomida and Tsumura 2006, Saegusa et al. 1998, Kobayashi et al. 2006, Saegusa et al. 2008, Ohara 2008, Azuma and Shibata 2010). -
Dinosaurs British Isles
DINOSAURS of the BRITISH ISLES Dean R. Lomax & Nobumichi Tamura Foreword by Dr Paul M. Barrett (Natural History Museum, London) Skeletal reconstructions by Scott Hartman, Jaime A. Headden & Gregory S. Paul Life and scene reconstructions by Nobumichi Tamura & James McKay CONTENTS Foreword by Dr Paul M. Barrett.............................................................................10 Foreword by the authors........................................................................................11 Acknowledgements................................................................................................12 Museum and institutional abbreviations...............................................................13 Introduction: An age-old interest..........................................................................16 What is a dinosaur?................................................................................................18 The question of birds and the ‘extinction’ of the dinosaurs..................................25 The age of dinosaurs..............................................................................................30 Taxonomy: The naming of species.......................................................................34 Dinosaur classification...........................................................................................37 Saurischian dinosaurs............................................................................................39 Theropoda............................................................................................................39 -
Re-Description of the Sauropod Dinosaur Amanzia (“Ornithopsis
Schwarz et al. Swiss J Geosci (2020) 113:2 https://doi.org/10.1186/s00015-020-00355-5 Swiss Journal of Geosciences ORIGINAL PAPER Open Access Re-description of the sauropod dinosaur Amanzia (“Ornithopsis/Cetiosauriscus”) greppini n. gen. and other vertebrate remains from the Kimmeridgian (Late Jurassic) Reuchenette Formation of Moutier, Switzerland Daniela Schwarz1* , Philip D. Mannion2 , Oliver Wings3 and Christian A. Meyer4 Abstract Dinosaur remains were discovered in the 1860’s in the Kimmeridgian (Late Jurassic) Reuchenette Formation of Moutier, northwestern Switzerland. In the 1920’s, these were identifed as a new species of sauropod, Ornithopsis greppini, before being reclassifed as a species of Cetiosauriscus (C. greppini), otherwise known from the type species (C. stewarti) from the late Middle Jurassic (Callovian) of the UK. The syntype of “C. greppini” consists of skeletal elements from all body regions, and at least four individuals of diferent sizes can be distinguished. Here we fully re-describe this material, and re-evaluate its taxonomy and systematic placement. The Moutier locality also yielded a theropod tooth, and fragmen- tary cranial and vertebral remains of a crocodylomorph, also re-described here. “C.” greppini is a small-sized (not more than 10 m long) non-neosauropod eusauropod. Cetiosauriscus stewarti and “C.” greppini difer from each other in: (1) size; (2) the neural spine morphology and diapophyseal laminae of the anterior caudal vertebrae; (3) the length-to-height proportion in the middle caudal vertebrae; (4) the presence or absence of ridges and crests on the middle caudal cen- tra; and (5) the shape and proportions of the coracoid, humerus, and femur. -
And Early Jurassic Sediments, and Patterns of the Triassic-Jurassic
and Early Jurassic sediments, and patterns of the Triassic-Jurassic PAUL E. OLSEN AND tetrapod transition HANS-DIETER SUES Introduction parent answer was that the supposed mass extinc- The Late Triassic-Early Jurassic boundary is fre- tions in the tetrapod record were largely an artifact quently cited as one of the thirteen or so episodes of incorrect or questionable biostratigraphic corre- of major extinctions that punctuate Phanerozoic his- lations. On reexamining the problem, we have come tory (Colbert 1958; Newell 1967; Hallam 1981; Raup to realize that the kinds of patterns revealed by look- and Sepkoski 1982, 1984). These times of apparent ing at the change in taxonomic composition through decimation stand out as one class of the great events time also profoundly depend on the taxonomic levels in the history of life. and the sampling intervals examined. We address Renewed interest in the pattern of mass ex- those problems in this chapter. We have now found tinctions through time has stimulated novel and com- that there does indeed appear to be some sort of prehensive attempts to relate these patterns to other extinction event, but it cannot be examined at the terrestrial and extraterrestrial phenomena (see usual coarse levels of resolution. It requires new fine- Chapter 24). The Triassic-Jurassic boundary takes scaled documentation of specific faunal and floral on special significance in this light. First, the faunal transitions. transitions have been cited as even greater in mag- Stratigraphic correlation of geographically dis- nitude than those of the Cretaceous or the Permian junct rocks and assemblages predetermines our per- (Colbert 1958; Hallam 1981; see also Chapter 24). -
Virginia Tidwell, Kenneth Carpenter & Williams Brooks, New
ORYCTOS, V ol . 2 : 21 - 37, Décembre 1999 NEW SAUROPOD FROM THE LOWER CRETACEOUS OF UTAH, USA Virginia TIDWELL, Kenneth CARPENTER and William BROOKS Department of Earth and Space Sciences, Denver Museum of Natural History, 2001 Colorado Blvd., Denver, CO 80205, USA. Abstract : The sauropod record for the Lower Cretaceous is poor in North America and consists mostly of iso - lated bones. Recently, however, a partial semiarticulated skeleton of a brachiosaurid, Cedarosaurus weiskopfae n.g., n.sp, was recovered from the Yellow Cat Member of the Cedar Mountain Formation, Utah, USA. The humeral leng - th is almost the same as the femur, while the dorsal and caudal vertebrae, and the metacarpal all display characters that identify the specimen as brachiosaurid. The forelimb and caudal vertebrae distinctly set it apart from all cur - rently described genera in that family. A brief review of Early to Middle Cretaceous brachiosaurs sorts through the confusing jumble of taxa that has developed over the years. In North America, most brachiosaurids found in Lower or Middle Cretaceous strata have historically been referred to the genus Pleurocoelus. The review illustrates the need for a reexamination of Pleurocoelus type material, as well as several specimens referred to that genus. Other material previously assigned to Pleurocoelus may yet prove to be the same as Cedarosaurus weiskopfae . Key words: Lower Cretaceous, brachiosaurid, new taxon, South-central United States. INTRODUCTION and several manus and pes elements. Only the proxi - mal end of the femur was recovered from the left side Until recently remains of Cretaceous sauropods and it is likely that all other elements were eroded in North America have been limited to the advanced away. -
Postcranial Skeletal Pneumaticity in Sauropods and Its
Postcranial Pneumaticity in Dinosaurs and the Origin of the Avian Lung by Mathew John Wedel B.S. (University of Oklahoma) 1997 A dissertation submitted in partial satisfaction of the requirements for the degree of Doctor of Philosophy in Integrative Biology in the Graduate Division of the University of California, Berkeley Committee in charge: Professor Kevin Padian, Co-chair Professor William Clemens, Co-chair Professor Marvalee Wake Professor David Wake Professor John Gerhart Spring 2007 1 The dissertation of Mathew John Wedel is approved: Co-chair Date Co-chair Date Date Date Date University of California, Berkeley Spring 2007 2 Postcranial Pneumaticity in Dinosaurs and the Origin of the Avian Lung © 2007 by Mathew John Wedel 3 Abstract Postcranial Pneumaticity in Dinosaurs and the Origin of the Avian Lung by Mathew John Wedel Doctor of Philosophy in Integrative Biology University of California, Berkeley Professor Kevin Padian, Co-chair Professor William Clemens, Co-chair Among extant vertebrates, postcranial skeletal pneumaticity is present only in birds. In birds, diverticula of the lungs and air sacs pneumatize specific regions of the postcranial skeleton. The relationships among pulmonary components and the regions of the skeleton that they pneumatize form the basis for inferences about the pulmonary anatomy of non-avian dinosaurs. Fossae, foramina and chambers in the postcranial skeletons of pterosaurs and saurischian dinosaurs are diagnostic for pneumaticity. In basal saurischians only the cervical skeleton is pneumatized. Pneumatization by cervical air sacs is the most consilient explanation for this pattern. In more derived sauropods and theropods pneumatization of the posterior dorsal, sacral, and caudal vertebrae indicates that abdominal air sacs were also present.