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Download Download Journal of Threatened Taxa PLATINUM OPEN ACCESS J Building TTevidence for conservaton globally ISSN 0974-7907 (Online) ISSN 0974-7893 (Print) 26 June 2019 (Online & Print) Vol. 11 | No. 8 | 13951–14086 10.11609/jot.2019.11.8.13951-14086 www.threatenedtaxa.org ISSN 0974-7907 (Online); ISSN 0974-7893 (Print) Publisher Host Wildlife Informaton Liaison Development Society Zoo Outreach Organizaton www.wild.zooreach.org www.zooreach.org No. 12, Thiruvannamalai Nagar, Saravanampat - Kalapat Road, Saravanampat, Coimbatore, Tamil Nadu 641035, India Ph: +91 9385339863 | www.threatenedtaxa.org Email: [email protected] EDITORS Typesetng Founder & Chief Editor Mr. Arul Jagadish, ZOO, Coimbatore, India Dr. Sanjay Molur Mrs. Radhika, ZOO, Coimbatore, India Wildlife Informaton Liaison Development (WILD) Society & Zoo Outreach Organizaton (ZOO), Mrs. Geetha, ZOO, Coimbatore India 12 Thiruvannamalai Nagar, Saravanampat, Coimbatore, Tamil Nadu 641035, India Mr. Ravindran, ZOO, Coimbatore India Deputy Chief Editor Fundraising/Communicatons Dr. Neelesh Dahanukar Mrs. Payal B. Molur, Coimbatore, India Indian Insttute of Science Educaton and Research (IISER), Pune, Maharashtra, India Editors/Reviewers Managing Editor Subject Editors 2016-2018 Mr. B. Ravichandran, WILD, Coimbatore, India Fungi Associate Editors Dr. B.A. Daniel, ZOO, Coimbatore, Tamil Nadu 641035, India Dr. B. Shivaraju, Bengaluru, Karnataka, India Ms. Priyanka Iyer, ZOO, Coimbatore, Tamil Nadu 641035, India Prof. Richard Kiprono Mibey, Vice Chancellor, Moi University, Eldoret, Kenya Dr. Mandar Paingankar, Department of Zoology, Government Science College Gadchiroli, Dr. R.K. Verma, Tropical Forest Research Insttute, Jabalpur, India Chamorshi Road, Gadchiroli, Maharashtra 442605, India Dr. V.B. Hosagoudar, Bilagi, Bagalkot, India Dr. Ulrike Streicher, Wildlife Veterinarian, Eugene, Oregon, USA Dr. Vatsavaya S. Raju, Kakatay University, Warangal, Andhra Pradesh, India Dr. D.J. Bhat, Retd. Professor, Goa University, Goa, India Editorial Assistant Dr. G.P. Sinha, Botanical Survey of India, Allahabad, India Vidya Mary George, Bengaluru, India Plants Editorial Board Dr. N.P. Balakrishnan, Ret. Joint Director, BSI, Coimbatore, India Ms. Sally Walker Dr. Shonil Bhagwat, Open University and University of Oxford, UK Founder/Secretary, ZOO, Coimbatore, India Prof. D.J. Bhat, Retd. Professor, Goa University, Goa, India Dr. Ferdinando Boero, Università del Salento, Lecce, Italy Dr. Robert Lacy Dr. Dale R. Calder, Royal Ontaro Museum, Toronto, Ontario, Canada Department of Conservaton Biology, Chicago Zoological Society (also known as the Brookfeld Dr. Cleofas Cervancia, Univ. of Philippines Los Baños College Laguna, Philippines Zoo), Brookfeld, Illinois 60513 USA; and Commitee on Evolutonary Biology, University of Dr. F.B. Vincent Florens, University of Mauritus, Mauritus Chicago Dr. Merlin Franco, Curtn University, Malaysia Dr. Francesco Dal Grande, Senckenberg Gesellschaf für Naturforschung, Frankfurt Dr. Russel Mitermeier Dr. V. Irudayaraj, St. Xavier’s College, Palayamkotai, Tamil Nadu, India Executve Vice Chair, Conservaton Internatonal, Arlington, Virginia 22202, USA Dr. B.S. Kholia, Botanical Survey of India, Gangtok, Sikkim, India Dr. Christoph Kuefer, Insttute of Integratve Biology, Zürich, Switzerland Prof. Mewa Singh Ph.D., FASc, FNA, FNASc, FNAPsy Dr. Pankaj Kumar, Kadoorie Farm and Botanic Garden Corporaton, Hong Kong S.A.R., China Ramanna Fellow and Life-Long Distnguished Professor, Biopsychology Laboratory, and Dr. V. Sampath Kumar, Botanical Survey of India, Howrah, West Bengal, India Insttute of Excellence, University of Mysore, Mysuru, Karnataka 570006, India; Honorary Dr. H.C. Nagaveni, Insttute of Wood Science and Technology, Bengaluru, India Professor, Jawaharlal Nehru Centre for Advanced Scientfc Research, Bangalore; and Adjunct Dr. K.S. Negi, NBPGR, Nainital District, Utarakhand, India Professor, Natonal Insttute of Advanced Studies, Bangalore Dr. A.J. Solomon Raju, Andhra University, Visakhapatnam, India Dr. Vijayasankar Raman, University of Mississippi, USA Dr. Ulrike Streicher, DVM Dr. B. Ravi Prasad Rao, Sri Krishnadevaraya University, Anantpur, India Wildlife Veterinarian / Wildlife Management Consultant, 1185 East 39th Place, Eugene, OR Dr. K. Ravikumar, FRLHT, Bengaluru, Karnataka, India 97405, USA Prof. Satsh C. Verma, Panjab University, Chandigarh, Punjab, India Dr. Aparna Watve, Pune, Maharashtra, India Stephen D. Nash Dr. Qiang Liu, Xishuangbanna Tropical Botanical Garden, Yunnan, China Scientfc Illustrator, Conservaton Internatonal, Dept. of Anatomical Sciences, Health Dr. Noor Azhar Mohamed Shazili, Universit Malaysia Terengganu, Kuala Terengganu, Malaysia Sciences Center, T-8, Room 045, Stony Brook University, Stony Brook, NY 11794-8081, USA Dr. M.K. Vasudeva Rao, Shiv Ranjani Housing Society, Pune, Maharashtra, India Prof. A.J. Solomon Raju, Andhra University, Visakhapatnam, India Dr. Fred Pluthero Dr. Mandar Datar, Agharkar Research Insttute, Pune, Maharashtra, India Toronto, Canada Dr. M.K. Janarthanam, Goa University, Goa, India Dr. K. Karthigeyan, Botanical Survey of India, India Dr. Martn Fisher Dr. Errol Vela, University of Montpellier, Montpellier, France Cambridge, UK Dr. P. Lakshminarasimhan, Botanical Survey of India, Howrah, India Dr. Larry R. Noblick, Montgomery Botanical Center, Miami, USA Dr. Ulf Gärdenfors Dr. K. Haridasan, FRLHT, Bengaluru, India Professor, Swedish Species Informaton Center, SLU, Uppsala, Sweden Invertebrates Dr. John Fellowes Honorary Assistant Professor, The Kadoorie Insttute, 8/F, T.T. Tsui Building, The University of Dr. Deepak Apte, Bombay Natural Hisotry Society, Mumbai, India. Hong Kong, Pokfulam Road, Hong Kong Dr. R.K. Avasthi, Rohtak University, Haryana, India (Orthoptera) Dr. D.B. Bastawade, Maharashtra, India (Araneae) Dr. Philip S. Miller Dr. Partha Pratm Bhatacharjee, Tripura University, Suryamaninagar, India Senior Program Ofcer, Conservaton Breeding Specialist Group (SSC/IUCN), 12101 Johnny Dr. Ferdinando Boero, Università del Salento, Lecce, Italy Cake Ridge Road, Apple Valley, MN 55124, USA Dr. Kailash Chandra, Zoological Survey of India, Jabalpur, Madhya Pradesh, India (Lepidoptera) Dr. Arthur Y.C. Chung, Sabah Forestry Department, Sandakan, Sabah, Malaysia Prof. Dr. Mirco Solé Dr. H.C. Paulo Corgosinho, Bairro Universitário, Frutal, Brazil Universidade Estadual de Santa Cruz, Departamento de Ciências Biológicas, Vice-coordenador Dr. B.A. Daniel, ZOO/WILD, Coimbatore, India (Lepidoptera) do Programa de Pós-Graduação em Zoologia, Rodovia Ilhéus/Itabuna, Km 16 (45662-000) Dr. Ansie Dippenaar-Schoeman, University of Pretoria, Queenswood, South Africa (Aran) Salobrinho, Ilhéus - Bahia - Brasil Dr. Rory Dow, Natonal Museum of natural History Naturalis, The Netherlands (Odonata) Dr. Alexander Ereskovsky, IMBE, Marseille, France (Spongillidae) Dr. Brian Fisher, California Academy of Sciences, USA English Editors Dr. Richard Gallon, llandudno, North Wales, LL30 1UP Mrs. Mira Bhojwani, Pune, India Dr. Hemant V. Ghate, Modern College, Pune, India Dr. Fred Pluthero, Toronto, Canada Dr. Mohammad Hayat, Aligarh Muslim University, Aligarh, India (Hymenoptera) Mr. P. Ilangovan, Chennai, India Dr. Tadashi Kawai, Wakkanai Fisheries Research Insttute, Hokkaido, Japan (Crustacea) Dr. S. Ajmal Khan, Annamalai University, Parangipetai, India (Corals) Web Design Dr. Ragnar Kinzelbach, University of Rostock, Rostock, Germany Mrs. Latha G. Ravikumar, ZOO/WILD, Coimbatore, India Dr. Ian J. Kitching, Natural History Museum, Cromwell Road, UK contnued on the back inside cover Front cover—Balanophora polyandra Grif. in Arunachal Pradesh © Umeshkumar L. Tiwari; Back cover—Monochoria hastata (L.) Solms in Litle Andaman © M.V. Ramana. Journal of Threatened Taxa | www.threatenedtaxa.org | 26 June 2019 | 11(8): 13951–13959 T (C, C) V C Michael Hofmann 1 , Alexei Abramov 2 , Hoang Minh Duc 3 , Le Trong Trai 4 , 5 6 7 8 9 Barney Long , An Nguyen , Nguyen Truong Son , Ben Rawson , Robert Timmins , ISSN 0974-7907 (Online) Tran Van Bang 10 & Daniel Willcox 11 ISSN 0974-7893 (Print) 1 Conservaton and Policy, Zoological Society of London, Regent’s Park, London, NW1 4RY, United Kingdom. PLATINUM 2 Zoological Insttute, Russian Academy of Sciences, Universitetskaya nab. 1, Saint Petersburg 199034, Russian OPEN ACCESS Federaton. 3,10 Southern Insttute of Ecology, Vietnam Academy of Science and Technology. 01 Mac Dinh Chi St., District 1, Ho Chi Minh City, Vietnam. 4 Viet Nature Conservaton Centre, Apt. 202, building 18T2, Le Van Luong Street, Hanoi, Vietnam. 5,6 Global Wildlife Conservaton, PO Box 129, Austn, TX 78767, USA. 6 Leibniz Insttute for Zoo and Wildlife Research, Alfred-Kowalke-Strae 17, 10315, Berlin, Germany. 7 Department of Vertebrate Zoology, Insttute of Ecology and Biological Resources, Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Caugiay, Hanoi, Vietnam. 8 WWF-Vietnam, Nam Tu Liem District, Hanoi, Vietnam. 9 51123 Monroe Street, Evanston, IL 60202, USA. 11 Save Vietnam’s Wildlife, Cuc Phuong Natonal Park, Nho Quan, Ninh Binh, Vietnam. 1 [email protected] (corresponding author), 2 [email protected], 3 [email protected], 4 trai.letrong@ thiennhienviet.org.vn, 5 [email protected], 6 [email protected], 7 [email protected], 8 [email protected], 9 [email protected], 10 [email protected], 11 [email protected] Abstract: Four species of wild canids are documented as occurring in Vietnam: Dhole Cuon alpinus, Eurasian
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