G. F. BENNEIT, R. A. EARLE, HESTER DU TOIT & F. W. HUCHZERMEYER

TABLE 2 Annotated host-parasite catalogue of Haematozoa in of the sub-Saharan region of Africa

ACCIPITRJDAE fa/lax: Greiner & Mundy, 1979 Trypanosoma mayae: IRCAH Accipiter badius Unknown parasite: IRCAH toddi: Peirce, 1984a Gyps coprotheres Leucocytozoon martyi: Commes 1918a; Tendeiro, 1948c, d Leucocytozoon sp: Ashford et al., 1976; Tendeiro 1947a sp.: IRCAH 1948a ' ' Microfilariae: Cowper, 1969 Gyps rueppellii No parasites seen: Ashford et al., 1976 Accipiter melanoleucus Haliaeetus vocifer Haemoproteus figueiredoi: Travassos Santos Dias, 1953 Leucocytozoon audieri: Laveran & Nattan-Larrier, 1911 Accipiter minul/us (now L. toddi) Leucocytozoon toddi: IRCAH Hieraaetus dubius Leucocytozoon sp.: Ashford eta/., 1976 No parasites seen: Peirce & Cooper, 1977b Accipiter ovampensis Hieraaetus fasciatus No parasites seen: Peirce, 1984a Haemoproteus elani: IRCAH Accipiter rufiventris Hieraaetus pennatus Haemoproteus sp.: Neave, 1906 No parasites seen: IRCAH Accipiter tachiro Kaupifalco monogrammicus Leucocytozoon toddi: Bennett & Herman, 1976 Haemoproteus saviannae: Tendeiro, 1947a, 1948a (species Leucocytozoon sp.: Ashford et al., 1976 incertae sedis) Leucocytozoon bacelari: Tendeiro, 1947a, 1948a Aquila rapax Leucocytozoon toddi: Tendeiro, 1947a, 1948a Haemoproteus buteonis: IRCAH Leucocytozoon ziemanni: Todd, 1907 Haemoproteus elani: IRCAH Leucocytozoon sp.: Dutton eta/., 1907a Haemoproteus figueiredoi: Peirce & Cooper, 1977b Microfilariae: Tendeiro, 1947a, 1948a Haemoproteus sp.: Ashford eta/., 1976 Trypanosoma fiadeiroi: Tendeiro, 1947a, 1948a Leucocytozoon toddi: IRCAH Trypanosoma sp.: Dutton et al., 1907a Plasmodium fa/lax: Peirce & Cooper, 1977b Unknown parasite: Tendeiro, 1947a, 1948a Aquila wahlbergi Lophaetus occipitalis Leucocytozoon toddi: IRCAH Leucocytozoon sp.: Schwetz, 1935c Plasmodium fa/lax: Peirce & Cooper, 1977b Melierax canorus Haemoproteus janovyi: IRCAH A viceda cucu/oides No parasites seen: Ashford et al., 1976 Me/ierax metabates Haemoproteus nisi: IRCAH Butastur liventer Haemoproteus columbae: Rousselot, 1943, 1953 Haemoproteus sp.: IRCAH Haemoproteus sp.: Ashford, et al., 1976 Leucocytozoon sp.: Rousselot, 1953 Buteo brachypterus Micronisus gabar Leucocytozoon toddi: Bennett & Blancou, 1974 Leucocytozoon sp.: Ashford et al., 1976 Buteo buteo Microfilariae: Leger, 1913 Leucocytozoon toddi: IRCAH Milvus aegyptius Buteo rufofuscus Plasmodium sp.: IRCAH Leucocytozoon toddi: Peirce & Cooper, 1977b Milvus migrans Circaetus cinereus sp.: Peirce & Cooper, 1977b Haemoproteus nisi: IRCAH Haemoproteus sp.: Rodhain et al. , 1913a Haemoproteus sp.: Peirce & Cooper, 1977b Leucocytozoon sr. : Rodhain et al. , 1913a; Schwetz, 1935c Leucocytozoon neavei: Neave, 1909 Circaetus gallicus Leucocytozoon sp.: Rodhein et al., 1913a Trypanosoma milvi: Stephens & Christophers, 1908 Haemoproteus elani: IRCAH Necrosyrtes monachus Dryotriorchis spectabilis Haemoproteus janovyi: Greiner & Mundy, 1979 Microfilariae: Plimmer, 1912 Haemoproteus sp.: Balfour, 1908b; Leger & Leger, 1914c; Todd & Wohlbach, 1912 Elanus caeruleus Leucocytozoon neophrontis: Todd & Wohlbach, 1912 (now Leucocytozoon sp.: Ashford et al., 1976 L . toddi) Trypanosoma sp.: IRCAH Microfilariae: Ashford et al. , 1976 Gyps africanus Neophron percnopterus Atoxoplasma sp.: Greiner & Mundy, 1979 (re-identified as Haemoproteus sp.: Rousselot, 1953 sp.) Microfilaliae: Neave, 1906 Haemoproteus elani: IRCAH Trypanosoma sp.: Neave, 1906 Haemoproteusjanovyi: Greiner & Mundy, 1979 Haemoproteus sp.: IRCAH Polemaetus be/licosus Hepatozoon sp.: IRCAH No parasites seen: IRCAH Leucocytozoon toddi: Greiner & Mundy, 1979 Leucocytozoon sp.: IRCAH Polyboroides radiatus Microfilariae: Greiner & Mundy, 1979 No parasites seen: Bennett & Blancou, 1974

39 HOST-PARASITE CATALOGUE OF THE HAEMATOZOA OF THE SUB-SAHARAN BIRDS

Polyboroides typus Alcedo Jeucogaster No parasites seen: Peirce & Cooper, 1977b No parasites seen: Peirce, 1969 Stephanoaetus coronatus Alcedo semitorquata Haemoproteus sp.: Rodhain eta/. , 1913a Microfilariae: IRCAH Terathopius ecaudatus Ceryle maxima No parasites seen: IRCAH No parasites seen: IRCAH Torgostracheliotus Ceryle rudis Haemoproteus ecae: Tendeiro, 1947a, 1948b; Travassos Atoxoplasma sp:: Greiner & Mundy, 1979 (re-identified as Santos Dias, 1953 (syn H. halcyonis) Hepatozoon adiei) Haemoproteus halcyon is: Bennett et al., 1974 Haemoproteus janovyi: Greiner & Mundy, 1979 Haemoproteus sp.: Ashford et al., 1976 Haemoproteus sp.: Archetti, 1946, 1947 Trypanosoma avium: Bennett et al. , 1974 Hepatozoon adiei: IRCAH Leucocytozoon toddi: Greiner & Mundy, 1979 Halcyon albiventris Trigonoceps occipitalis sp.: Bennett & Herman, 1976 Haemoproteus fusca: IRCAH Haemoproteus janovyi: Greiner & Mundy, 1979 Leucocytozoon sp.: IRCAH Haemoproteus sp.: IRCAH Halcyon chelicuti Haemoproteus sp,: Ashford et al. , 1976; Schwetz, 1938 ALAUDIDAE Leucocytozoon sp.: IRCAH Calandrella cinerea Halcyon leucocephala , Haemoproteus alaudae: IRCAH Microfilariae: Ashford et al., 1976 Chersomanes albofasciata Trypanosoma sp.: Ashford et al., 1976 No parasites seen: IRCAH Halcyon malimbica No parasites seen: Bennett et al. , 1974, 1977 Eremopterix australis No parasites seen: IRCAH Halcyon senegalensis Haemoproteus halcyonis: Bennett et al., 1977 Eremopterix Jeucotis Haemoproteus sp.: Schwetz, 1944; Theiler, 1930 Haemoproteus sp. : Ashford et al., 1976 Microfilariae: Ashford eta/., 1976 Plasmodium sp. : Schwetz, 1944 Eremopterix signata lspidina picta Haemoproteussp.: Peirce, 1976b Haemoproteus enucleator: Ashford et al., 1976; Bennett et Galerida magnirostris al., 1977; Peirce, 1984a, i; Williams et al., 1977 Haemoproteus halcyonis: Bennett & Campbell, 1973 No parasites seen: IRCAH Haemoproteus sp.: Cox & Vickerman, 1965; Schwetz, 1938 Mirafra africana Leucocytozoon fringillinarum: IRCAH Leucocytozoon sp.: Bennett et al., 1974; Schwetz, 1938 No parasites seen: Bennett et al., 1974 Microfilariae: Bennett et al., 1977 Mirafra africanoides Plasmodium vaughani: Bennett et al., 1977 Plasmodiumsp.: Bennettetal., 1974 No parasites seen: Bennett & Herman, 1976 Trypanosoma avium: Bennett et al. , 1974, 1977 Mirafra albescens Trypanosoma calmettei: IRCAH Trypanosoma sp.: IRCAH No parasites seen: IRCAH Unidentified parasite: IRCAH Mirafra apiata ANATIDAE Trypanosoma avium: IRCAH Alopochen aegyptiacus Mirafra cheniana Haemoproteus sp.: Ashford et al. , 1976; Rousselot, 1953 Haemoproteus alaudae: IRCAH Anas capensis Mirafra curvirostris No parasites seen: Ashford et al., 1976; Peirce, 1976b No parasites seen: IRCAH Anas erythrorhyncha Mirafra rufocinnamomea Haemoproteus sp.: Thomas & Dobson, 1975 No parasites seen: Ashford et al., 1976 Leucocytozoon sp.: Thomas & Dobson, 1975 (identified in error) Mirafra sabota Unidentified parasite: IRCAH No parasites seen: Oosthuizen & Markus, 1967b Anas hottentotta Spizocorys conirostris No parasites seen: Ashford et al. , 1976; Bennett & Herman, 1976b; Peirce, 1976b; Peirce, 1984i No parasites seen: IRCAH Anas me/Jeri Spizocorys fringillaris No parasites seen: Bennett & Blancou, 1974 No parasites seen: IRCAH Anassparsa No parasites seen: IRCAH ALCEOINIDAE Anas undulata Alcedo cristata No parasites seen: Ashford et al., 1976, Bennett & Herman, Haemoproteus halcyonis: Bennett et al. , 1977 1976 Haemoproteus sp.: Ashford et al., 1976 Microfilariae: Ashford et al., 1976 Cygnus atratus Plasmodiumsp.: IRCAH No parasites seen: IRCAH

40 G. F. BENNETT, R. A. EARLE, HESTER DU TOIT & F. W. HUCHZERMEYER

Dendroeygna viduata Egretta intermedia Lankesterella sp.: Ashford et al. , 1976 No parasites seen: Peirce, 1984i Netta erythrophthalma Gorsachius Ieuconotus Leucocytozoon sp.: Thomas & Dobson, 1975 (identified in No parasites seen: IRCAH error) Microfilariae: Thomas & Dobson, 1975 Ixobrychus minutus Unidentified parasite: IRCAH Haemoproteus sp.: Ashford eta/., 1976 Leucocytozoon sp.: Ashford et al. , 1976 Plectropterus gambensis Plasmodium sp.: Ashford et al., 1976 Haemoproteus nettionis: Williams & Bennett, 1980 Sarkidiornis melanotos BALAENICIPITIDAE Haemoproteus sp.: Rodhain et al., 1913a Balaeniceps rex Haemoproteus sp.: Berson, 1964 ANHINGIDAE ,, • , Leucocytozoon sp: : Rodhain et al., 1913a Anhinga rufa BUBALORNITHIDAE Haemoproteus sp.: Rodhain eta/., 1913a Leucocytozoon vandenbrandeni: Rodhain, 1931 Bubalornis albirostris Leucocytozoon sp.: Rodhain et al. , 1913a Haemoproteus bubalornis: Bennett & Peirce, 1991b Microfilariae: Ashford et at., 1976 Haemoproteus fallisi: Peirce et al., 1977b Haemoproteus sp.: Oosthuizen & Markus, 1967b APODIDAE Leucocytozoon fringillinarum: Peirce et al., 1977b Apusafflnis Bubalornis niger Unknown parasite: IRCAH Haemoproteus bubalornis: Bennett & Peirce, 1991b Haemoproteussp.: Ashford eta/., 1976 Apuscaffer Dinemellia dinemelli Unknown parasite: IRCAH Haemoproteus bubalornis: Bennett & Peirce, 1991b Apushorus Haemoproteus sp.: Peirce, 1984i No parasites seen: IRCAH Microfilariae: Ashford et al., 1976 Apusmelba BUCEROTIDAE No parasites seen: IRCAH Aceros undulatus ColloeaHa francica Microfilariae: IRCAH No parasites seen: Peirce et at., 1977a Anthracoeeros malabaricus Cypsiurus parvus Microfilariae: Peirce, 1969 No parasites seen: Oosthuizen & Markus, 1969b Bucorvus abyssinicus

ARDEIDAE Haemoproteus sp.: Rousselot, 1953 Microfilariae: Tendeiro, 1948a Ardea cinerea Bucorvus Ieadbeateri Haemoproteussp.: Lowery, 1971 Microfilariae: Lowery, 1971 No parasites seen: IRCAH Ardea goliath Bycanistes brevis Haemoproteus sp.: Rousselot, 1953 Haemoproteus sp.: Berson, 1964 Leucocytozoon ardeae: Rodhain et al., 1913a Trypanosoma sp.: Ross, 1911 Trypanosoma sp.: Rodhain et al., 1913a Bycanistes bucinator Ardea melanocephala Haemoproteus sp.: Schwetz, 1931; Schwetz & Fornara, 1929 No parasites seen: IRCAH Leucocytozoon ziemanni: Berson, 1964 Leucocytozoon sp.: Schwetz, 1931; Schwetz & Fornara, 1929 Ardea purpurea Microfilariae: IRCAH Trypanosoma sp.: Rodhain et al. , 1913a; Todd, 1907 Haemoproteus sp.: Rodhain et al., 1913a Leucocytozoon sp.: Schwetz, 1935c Bycanistes sharpii Ardeola ibis Haemoproteus sp.: Schwetz, 1931; Schwetz & Fornara, 1929 Leucocytozoon sp.: Schwetz, 1931, 1935c; Schwetz & For­ Leucocytozoon ardeae: IRCAH nara, 1929 Ardeola idae Microfilariae: Schwetz, 1931, 1935c, 1944; Schwetz & For­ nara, 1929 No parasites seen: Bennett & Blancou, 1974 Trypanosoma bycanistes: Schwetz & Fornara, 1929 Butorides striatus Trypanosomasp.: Schwetz, 1931,1933, 1938 No parasites seen: Lowery, 1971 Bycanistes subcylindricus Egretta alba Microfilariae: Peirce, 1984i Trypanosoma sp.: Peirce, 1984i No parasites seen: Peirce, 1984i Toekus alboterminatus Egretta ardesiaca Microfilariae: Ashford et al., 1976 No parasites seen: Bennett & Blancou, 1974 Trypanosoma sp.: Ashford eta/., 1976 Egretta garzetta Toekus deckeni Microfilariae: Lowery, 1971 Leucocytozoon sp.: Peirce et al., 1977b 41 HOST-PARASITE CATALOGUE OF THE HAEMATOZOA OF THE SUB-SAHARAN BIRDS

Tockus erythrorhynchus Pogoniulus bilineatus Haemoproteussp.: Ashford et a!. , 1976 Haemoproteus thereicerycis: Bennett & Nandi, 1981 Leucocytozoon sp.: Peirce & Backhurst, 1970 Haemoproteus sp.: Bennett et al., 1977 Microfilariae: Leger, 1913; Oosthuizen & Markus, 1967b Trypanosoma sp.: Oosthuizen & Markus, 1967b Pogoniulus chrysoconus No parasites seen: Ashford et al., 1976; Peirce, 1984a Tockus fasciatus Haemoproteus sp.: Schwetz, 1944 Pogoniulus leucolaima Microfilariae: Leger, 1913 No parasites seen: Bennett & Herman, 1976; Bennett et al., Plasmodium sp.: Schwetz, 1944 1977 Trypanosoma sp.: Schwetz, 1944 Pogoniulus pusillus Tockus flavirostris No parasites seen: Ashford et al., 1976 Microfilariae: Oosthuizen & Markus, 1967b Tockus nasutus Trachyphonusdarnaudil Haemoproteus thereicerycis: Bennett & Nandi, 1981; Peirce, Haemoproteus sp.: Cowfer, 1969 1984i Microfilariae: Leger, 19 3; Neave, 1906, Tendeiro, 1948b Haemoproteus sp.: Balfour, 1908b Trypanosoma sp.: Todd & Wolbach, 1912 Leucocytozoon fringillinarum: Peirce et al., 1977b Leucocytozoon sp.: Peirce, 1.976b BURHINIDAE Trachyphonus erythrocephalus Burhinus capensis Haemoproteus sp.: Bennett & Herman, 1976 Plasmodium (Novyella) sp.: IRCAH Leucocytozoon sp.: Peirce & Backhurst, 1970 Burhinus senegalensis Trachyphonus purpuratus Haemoproteus sp.: Rouselot, 1953 Haemoproteus thereicerycis: Peirce, 1984i Leucocytozoon sp.: Peirce, 1984i CAMPEPHAGIDAE Trachyphonus vaillantii Campephaga phoenicea Haemoproteus cornuata: IRCAH Haemoproteus thereicerycis: IRCAH No parasites seen: Ashford et al., 1976; Bennett et al., 1977; Haemoproteus xantholaemae: IRCAH Peirce, 1984a Leucocytozoon sp.: IRCAH Coracina caesia Tricholaema melanocephala No parasites seen: Bennett & Herman, 1976; Peirce, 1984i No parasites seen: Peirce & Backhurst, 1970

CAPITONIDAE

Buccanodon Jeucotis CAPRIMULGIDAE Leucocytozoon sp.: Bennett & Herman, 1976 Caprimulgus climacurus Plasmodium vaughani.· Bennett & Herman, 1976 No parasites seen: Ashford et al., 1976 Buccanodon olivaceum No parasites seen: Bennett & Herman, 1976 Caprimulgus donaldsoni No parasites seen: Peirce et al., 1977b Buccanodon whytii No parasites seen: Bennett & Herman 1976 Caprimulgus europaeus No parasites seen: Peirce, 1984a Gymnobucco c/avis Haemoproteus thereicerycis: Bennett & Nandi, 1981 Caprimulgus fossil Haemoproteus sp. : Theiler, 1930 Haemoproteussp.: Ashford eta!., 1976 Lybius bidentatus Leucocytozoon caprimulgi: Kerandel, 1913 Leucocytozoon sp.: Kerandel, 1913 Haemoproteus sp. : Ashford et al., 1976 Microfilariae: IRCAH Trypanosoma caprimulgi: Kerandel, 1913 Lybius dubius Trypanosoma sp.: Kerandel, 1913 Haemoproteus sp.: Cheke, 1971 Microfilariae: Cheke, 1971 Caprimulgus madagascariensis Lybius guifsobalito No parasites seen: Lowery, 1971 Haemoproteus sp.: Ashford et al., 1976 Caprimulgus natalensis Lybius Jachrymosus No parasites seen: Bennett et al. , 1977 No parasites seen: Peirce et al. , 1977b Caprimulgus pectoralis Lybius Jeucomelas Leucocytozoon caprimulgi: IRCAH Haemoproteus cornuata: IRCAH Caprimulgus poliocephalus Haemoproteus thereicerycis: IRCAH Haemoproteussp.: Ashford eta[., 1976 No parasites seen: Ashford et al., 1976 Leucocytozoon fringillinarum: IRCAH Caprimulgusruflgena Leucocytozoon sp.: Ashford et al., 197t Microfilariae: Ashford et al., 1976 No parasites seen: Ashford et al. , 1976 Lybius torquatus Caprimulgus tristigma No parasites seen: Peirce, 1984a No parasites seen: IRCAH Lybius undatus Macrodipteryx vexillarius Haemoproteus sp.: Ashford et al., 1976 No parasites seen: Peirce, 1984a 42 G. F. BENNETI,R. A. EARLE,HESTERDUTOIT& F. W. HUCHZERMEYER

CHARADRIIDAE COLUMBIDAE Charadrius marginatus Aplopelialarvata No parasites seen: IRCAH Haemoproteussp.: Ashford eta/., 1976 Leucocytozoon sp.: Ashford et al., 1976 Charadrius mongo/us Microfilariae: Ashford et al., 1976 No parasites seen: Ashford et al., 1976 Columba guinea Charadrius pecuarius Haemoproteus columbae: Bennett & Herman, 1976; Rousse­ lot, 1943 No parasties seen: Ashford et al., 1976 Haemoproteus sacharovi: Berson, 1964 (probably in error as Charadrius tricollaris this species is known only from New World) Haemoproteus sp.: Ashford et al., 1976; Cowper, 1969; Haemoproteus sp.: Peirce & Backhurst, 1970 Rousselot, 1953; Thomas & Dobson, 1975 Leucocytozoon sp.: Peirce & Backhurst, 1970 Microfilariae: Ashford et al., 1976 Vanellus a/biceps Columba iriditorques No parasites seen: IRCAH No parasites seen: Schwetz, 1944 Vanellus armatus Columba Iivia No parasites seen: Peirce, 1976b Atoxoplasma sp.: Peirce et al., 1977a Haemoproteus columbae: Peirce et al., 1977a Vanellus coronatus Plasmodium nucleophilum: IRCAH Microfilariae: Oosthuizen & Markus, 1967b Columba mayeri Vanellus crassirostris Leucocytozoon marchouxi: Peirce, 1984i No parasites seen: Bennett etal., 1974 Geopelia cuneata Vanellus tectus Plasmodium sp.: IRCAH Haemoproteus nascimentoi: Tendeiro, 1947a, 1948a, (re­ Goura victoria quires re-evaluation) No parasites seen: IRCAH Leucocytozoon sousadiasi: Tendeiro, 1947a, 1948a (requires re-evaluation) Oena capensis Haemoproteus columbae: Peirce et al., 1977b CICONIIDAE Haemoproteussp.: Archetti, 1946, 1947; Ashford et al., 1976; Jansen, 1952 Anastomus lamelligerus Leucocytozoon sp.: Jansen, 1952 Haemoproteus sp.: Rodhain et at. , 1913a Microfilariae: Archetti, 1947; Ashford et al., 1976 Trypanosoma oenae: Sergent, Ed, 1941 Ciconia abdimii Trypanosoma sp.: Oosthuizen & Markus, 1969b No parasites seen: IRCAH Streptopelia capicola Ephippiorhynchus senegalensis Haemoproteus columbae: Peirce, 1984a Haemoproteus turtur: Peirce, 1984a Haemoproteus danilewskii: Wenyon, 1908 (now H. peircei) Haemoproteus sp.: Ashford et al., 1976; Jansen, 1952; Oost- huizen & Markus, 1967b; Peirce & Backhurst, 1970 Ibis ibis Leucocytozoon marchouxi: IRCAH Haemoproteus peircei: Peirce, 1984i Leucocytozoon sp.: Jansen, 1952 Plasmodium vaughani.· VRI Leptoptilus crumeniferus Trypanosoma sp.: Oosthuizen & Markus, 1967b Haemoproteus brodkorbi: Forrester et al., 1977 (now H. crumenium) Streptopelia decipiens Haemoproteus crumenium: Hirst, 1905; Peirce & Cooper, Haemoproteus columbae: Rousselot, 1943 1977a Haemoproteus sp.: Archetti, 1946, 1947; Ashford et al., 1976; Haemoproteus peircei: Forrester et al., 1977 Oosthuizen & Markus, 1969b Haemoproteus sp.: Forrester et al., 1977; Neave, 1906; Leucocytozoon sp.: Ashford eta/., 1976 Rousselot, 1953 Trypanosoma sp.: Ashford eta/., 1976 Microfilariae: Neave, 1906 Streptopelia picturata Mycteria ibis Haemoproteus sp.: Lowery, 1971 Haemoproteus peircei: Forrester eta/., 1977 Haemoproteus sp.: Rodhain et al., 1913a Streptopelia semitorquata Haemoproteus columbae: Bennett & Herman, 1976 COLIIDAE Haemoproteus sp.: Ashford et a/., 1976; Garnham, 1950; Rodhain et a[., 1913a Colius colius Leucocytozoon marchouxi: Ashford et al., 1976; Minchin, No parasites seen: IRCAH 1910 Microfilariae: Travassos Santos Dias, 1953 Colius indicus Trypanosoma sp.: Ashford et al. , 1976 Microfilariae: IRCAH Streptopelia senegalensis Colius macrourus Haemoproteus columbae: Peirce et a[., 1977b; Rousselot, 1943 No parasites seen: Ashford et a/., 1976; Peirce, 1984i; Haemoproteus sp. : Ashford et al., 1976; Rousselot, 1953, Williamsetal., 1977 ( Thomas & Dobson, 1975; Todd & Wolbach, 1912 · Colius striilt'us Leucocytozoon marchouxi: Oosthuizen & Markus, 1967b; Peirce et al., 1977b Haemoproteus sp.: Garnham, 1950 Leucocytozoon sp.: Ashford et al., 1976; Leger & Leger, Leucocytozoon sp.: IRCAH 1914a; Rousselot, 1943; Thomas & Dobson, 1975

43 HOST-PARASITE CATALOGUE OF THE HAEMATOZOA OF THE SUB-SAHARAN BIRDS

Microfilariae: Bennett & Herman, 1976; Bennett et al., 1978; Haemoproteus sp.: Archetti, 1946, 1947 Oosthuizen & Markus, 1967b; Peirce et al., 1977b Plasmodium circumflexum: Crewe, 1974 Coracias spatulata Plasmodium vaughani: Crewe, 1972 Haemoproteus coraciae: Bennett & Herman, 1976 Trypanosoma avium: Bennett et al., 1978 Leucocytozoon eurystomi: Bennett & Herman, 1976 Trypanosoma sp.: Oosthuizen & Markus, 1967b Eurystomus glaucurus Streptopelia turtur Haemoproteus coraciae: Bennett & Herman, 1976; Bennett Haemoproteussp.: Ashford eta/., 1976 et al., 1978 Haemoproteus cruzferreirae: Tendeiro, 1947a, 1948a (syn of Streptopelia vinacea H. coraciae) Haemoproteus columbae: Rousselot, 1943, 1953 Haemoproteus sp.: Ashford et al., 1976; Schwetz, 1938 Haemoproteus sp.: Ashford et al., 1976 Leucocytozoon francae: Tendeiro, 1947a, 1948a (syn L. Trypanosoma sp.: Todd & Wolbach, 1912 eurystomi) Leucocytozoon sp.: Schwetz, 1938 Treron australis Trypanosoma fradei: Tendeiro, 1947a, 1948a Haemoproteus sp.: Ashford et al., 1976 Eurystomus gularis Leucocytozoon sp.: Cowper, 1969 Leucocytozoon eurystomi: Kerandel, 1913 Treron calva Leucocytozoon sp.: Kerandel, 1909 Microfilariae: Kerandel, 1909 Haemoproteus sp.: Schwetz, 1944; Todd & Wolbach, 1912 Trypanosoma eurystomi: Kerandel, 1913 Plasmodium sp.: Schwetz, 1944 Trypanosoma sp.: Kerandel, 1909 Trypanosoma sp.: Todd & Wolbach, 1912

Treron waalia CORVIDAE No parasites seen: Ashford et al., 1976; Bennett et al., 1978 Corvus albus Turtur abyssinicus No parasites seen: Lowery, 1971 Haemoproteus columbae: Williams et al., 1977 Haemoproteus sp.: Ashford eta/., 1976 Corvus capensis Leucocytozoon sp.: Ashford eta/., 1976 Haemoproteus sp.: IRCAH Microfilariae: IRCAH Turturafer Haemoproteus sp.: Ashford et al., 1976 Corvus crassirostris Leucocytozoon sp.: Ashford et al., 1976 No parasites seen: Ashford et al. , 1976 Turtur brehmeri ~UCULIDAE Haemoproteus danilewskyi: Plimmer, 1912 (now H. colum­ bae) Centropus senegalensis Turtur cbalcospilos Haemoproteus centropi: Peirce, 1977 Haemoproteus columbae: Rousselot, 1942, 1943 (now H. Haemoproteus columbae: Peirce, 1984a; Peirce et al., 1977b centropi) Haemoproteus sp.: Ashford et al., 1976; Oosthuizen & Mar· Haemoproteus froilanoi: Tendeiro, 1947a, 1948a (syn of H. kus, 1967b; Peirce & Backhurst, 1970 centropi) Leucocytozoon marchouxi: Peirce, 1984i; Peirce et al., 1977b Haemoproteus sp.: Rousselot, 1953 Leucocytozoon sp.: Peirce & Backhurst, 1970 Leucocytozoon sp.: Schwetz, 1931, 1935c; Tendeiro, 1947a, Plasmodium vaughani: Ashford et al., 1976 1948a Trypanosoma sp.: Oosthuizen & Markus, 1967b Centropus superciliosus Turtur tympanistria Haemoproteus centropi: Fantham, 1921; Peirce, 1984i; VRI Haemoproteus columbae: Bennett & Herman, 1976 Leucocytozoon centropi: Peirce, 1984i; VRI Haemoproteus sp.: Ashford et al., 1976; Bennet et al. , 1974 Leucocytozoon marchouxi: Bennett & Herman, 1976 Centropus toulou Leucocytozoon sp.: Ashford eta/., 1976; Bennet et al., 1974 Trypanosoma avium: Bennett et al., 1974 Haemoproteus sp.: Lowery, 1971 . Trypanosoma sp.: Ashford eta/., 1976 Leucocytozoon centropi: Bennett & Blancou, 1974 Cercococcyx mecbowi CORACIIDAE No parasites seen: Bennett et al. , 1974 Coracias caudata Ceutbmochares aereus Haemoproteus coraciae: Bennett & Herman, 1976; Peirce, Haemoproteus sp.: Schwetz, 1938 1984a; Travassos Santos Dias, 1953 Microfilariae : Bennett et al., 1974 Leucocytozoon eurystomi: Peirce, 1984a Trypanosoma sp.: Martinet al., 1909 Leucocytozoon sp.: Bennett & Herman, 1976 Microfilariae: Peirce, 1984a Chrysococcyx caprius Coracias cyanogaster Aegyptianella sp.: Peirce et al., 1977b Haemoproteus sp.: Bennett et al., 1977 Haemoproteus coraciae: Peirce et al. , 1977b Leucocytozoon centropi: IRCAH Haemoproteus fontesi var. cyanogasteri: Tendeiro, 1947a, Leucocytozoon coccyzus: Oosthuizen & Markus, 1969b (syn 1948a (syn H. coraciae) L. centropi) Leucocytozoon sp.: Ashford et al., 1976 Coracias garrulus Plasmodium vaughani: IRCAH Haemoproteus coraciae: Peirce et al. , 1977b Haemoproteus sp.: Oosthuizen & Markus, 1967b Chrysococcyx cupreus Leucocytozoon eurystomi: Peirce et al., 1977b Trypanosoma sp.: Peirce, 1984a Leucocytozoon sp:: Oosthuizen & Markus, 1%7b Chrysococcyx klaas Coracias naevia Microfilariae: Schwetz, 1938 Haemoproteus coraciae: Bennett eta/., 1978 Haemoproteus fontesi: Tendeiro, 1947a, 1948a (syn. of H. Chunator cafer coraciae) Haemoproteus columbae: Rousselot, 1943 (now H. centropi) 44 G. F. BENNETI,R. A. EARLE,HESTERDUTOIT&F. W. HUCHZERMEYER

Haemoproteus sp.: Rousselot, 1953; Schwetz, 1938 Microfilariae: Ashford eta/. , 1976 Plasmodium octamerium: Ashford et al., 1976 (7/aznatorjacobinus Plasmodium vaughani: Ashford et al., 1976 Haemoptroteus centropi: Peirce et al., 1977b; VRI Trypanosoma sp.: Ashford eta[., 1976 Leucocytozoon centropi: IRCAH Leucocytozoon coccyzus: Peirce et al., 1977b (Syn. L. cen­ Estrilda atricapma tropi) Haemoproteus africanus: Bennett & Peirce, 1991b Oaznator levaillanti Estrilda caerulescens Haemoproteus centropi: IRCAH Haemoproteus sp.: Rousselot, 1953 (7uculus solitarius Estrilda erythronotos No parasites seen: IRCAH Plasmodium sp.: IRCAH Estrilda melpoda DICRURIDAE Microfilariae: Cheke, 1971 Dicrurus adsimilis Plasmodium sp.: Cheke, 1971 Babesia sp.: Bennett & Herman, 1976 Estrilda paludicola Haemoproteus dicruri: Peirce, 1984a, g Haemoproteus sp.: Ashford et al., 1976; Cowper, 1969, Haemoproteus africanus: Bennett & Peirce, 1991b Oosthuizen & Markus, 1967b Haemoproteus fringillae: Bennett & Herman, 1976 Leucocytozoon sp.: Bennett & Herman, 1976 Haemoproteus sp.: Ashford et al., 1976 Trypanosoma sp.: Ashford et al., 1976 Plasmodium octamerium: Ashford et al. , 1976 M1crofilariae: Ashford et al., 1976 Trypanosoma sp.: Ashford et al., 1976 Dicrurus aldabranus Estrilda rhodopyga No parasites seen: Lowery, 1971 Haemoproteus sp.: Ashford et al., 1976 Microfilariae: Ashford eta/., 1976 EMBERIZIDAE Estrilda troglodytes Emberiza cabanisi F/aemoproteus sp.: Leger & Leger, 1914c Mjcrofilariae: Leger, 1913 Haemoproteus orizivorae: Peirce, 1984a Trypanosoma bouffardi: Molyneux, 1973b Leucocytozoon majoris: Peirce, 1984a Trypanosoma everetti: Molyneux, 1973a Trypanosoma sp.: Peirce, 1984a Uni9entified parasites: Peirce, 1969 Emberiza capensis Hypargos niveoguttatus No parasites seen: IRCAH Plasmodium sp.: Peirce, 1984a Emberiza flaviventris Microfilariae: Peirce, 1984a Haemoproteus chloriis: IRCAH Lagonosticta Jarvata Haem()proteus sp.: Oosthuizen & Markus, 1967b, Peirce, Haemoproteus sp.: Ashford et al., 1976 1984a Leucocytozoon sp.: Ashford et al., 1976 Leucocytozoon fringillinarum: Peirce, 1984a Microfilariae: Ashford etal., 1976 Microfilariae: Oosthuizen & Markus, 1967b Trypanosoma sp.: Ashford et al., 1976 Emberiza lmpetuani Lagonosticta rara No parasites seen: IRCAH No parasites seen: Bennett et al. , 1977 Emberiza tahapisi J.,agonosticta rhodopareia Haemoproteus columbae: Rousselot, 1953 (now H. orizi- Haemoproteus sp.: Ashford eta/. , 1976 vorae) Microfilariae: IRCAH Haemoproteus sp.: Rousselot, 1953 Leucocytozoon sp.: IRCAH Lagonosticta rubrii:ata Microfilariae: Ashford et al., 1976 Atoxopwsma sp.: Peirce , 1984a Plasmodiumfallax: Rousselot, 1953 Haemoproteus africanus: Bennett & Peirce, 1991b Plasmodium rousseloti: Bray, 1964 Haemoproteus fringillae/orizivorae: Bennet et al., 1977 Trypanosoma sp.: IRCAH Haemoproteus sp.: Ashford eta/., 1976; Peirce, 1984a Microfilariae: Ashford eta/., 1976 ESTRILDIDAE Plasmodium sp.: Peirce, 1984a Trypanosoma bouffardi: Peirce, 1984a Amandava aznandava Lagonosticta rufopicta No parasites seen: IRCAH Haemoproteus africanus: Bennett & Peirce, 1991b Amandava subflava Leucocytozoon fringillinarum : Wink & Bennett, 1976 No parasites seen: Peirce, 1984a Unidentified parasite: Peirce, 1969 Amadina erythrocephala Lagonosticta senega/a Haemoproteus africanus: Bennett & Peirce, 1991b Trypanosoma sp.: IRCAH Haemoproteus fringillae/orizivorae: Bennett et al., 1977 (now Amadina fasciata H. africanus) Haemoproteus sp.: Ashford et al., 1976; Commes, 1918b; Haemoproteus sp.: Oosthuizen & Markus, 1967b Cowper, 1%9; Rousselot, 1953 Trypimosoma avium: Bennett eta[. , 1978 Leucocytozoon fringillinarum: Bennett & Herman, 19?6 Trypanosoma bouffardi: Molyneux, 1973b Microfilariae: Ashford et al., 1976; Leger, 1913; Phmmer, Trypanosoma sp.: Ashford et al. , 1976 1912 Plasmodium vaughani: Ashford et al., 1976 Estriida astrild Plasmodium sp.: Bennett et al., 1978 Atoxoplasma sp.: PeirCe, 1984a Trypanosoma avium: Bennett & Herman, 1976 Haemogregarina sp.: Peirce, 1977 (probably Hepatozoon sp.) Trypanosoma bouffardi: Molyneux, 1973b Haemoproteus sp.: Ashford eta[., 1976 Trypanosoma everetti: Peirce et al., 1977b

45 HOST-PARASITECATALOGUEOFTHEHAEMATOZOAOFTHESUB-SAHARANBIRDS

Trypanoso/1Ul lagonostictae: Marullaz, 1914; Leger & Leger, Uraeginthus angolensis 1914c Atoxoplas/1Ul sp.: Peirce, 1984a Trypanoso/1Ul sp.: Ashford et al., 1976 Haemoproteus africanus: Bennett & Peirce, 1991b Mandingoa nitidula Haemoproteus orizivorae: Peirce, 1984a Haemoproteus sp.: Oosthuizen & Markus, 1967b Haemoproteus africanus: Bennett & Peirce, 1991b Microfilariae: Peirce, 1984a Haemoproteus fringillae/orizivorae: Bennett & Herman, Plasmodium circumflexum: Peirce, 1984a 1976; Bennett eta/. , 1974 (now H. africanus) Plasmodium sp.: Manwell & Rossi, 1975 Leucocytozoon sp.: Ashford eta/., 1976 Trypanosoi1Ul bouffardi: Peirce, 1984a Nigrita bicolor Uraeginthus bengalus Haemoproteus sp.: Schwetz, 1938 Haemoproteus orizivorae: Peirce et a/., 1977b; Williams et Microfilariae: Schwetz, 1938 a/. , 1977 (now H. africanus) Nigrita canicapilla Haemoproteus sp.; Ashford eta/., 1976 Haemoproteus uraeginthus: Bennett & Peirce, 1991b No parasites seen: Peirce, 1984i Leucocytozoon: sp.; Garnham, 1950 Microfilariae: Ashford eta/., 1976; Peirce, 1969 Nigrita fusconota Plasmodium relictum: Marullaz, 1912b Haemoproteus sp.: Schwetz, 1938 Plasmodium sp.; Peirce, 1969 Trypanoso/1Ul bouffardi: Molyneux, 1973b Ortygospiza atricollis Plasmodium sp.: IRCAH Uraeginthus grana tinus Haemoproteus africanus: IRCAH Pyrenestes ostrinus Lecocytozoon sp.: IRCAH Haemoproteus africanus: Bennett & Peirce, 1991b Plasmodium sp.: IRCAH Pytilia afra Uraeginthus ianthinogaster Haemoproteus africanus: Bennett & Peirce, 1991b Haemoproteus sp.: Ashford et al., 1976 Haemoproteus orizivorae: Peirce, 1984a Microfilariae: Ashford etal., 1976 Microfilariae: Bennett & Herman, 1976 Vidua chalybeata Pytilia melba Haemoproteus fringillaelorizivorae: Bennet et a/.,1978 (now Atoxoplasma sp.: Peirce, 1984a H. africanus) Haemoproteus africanus: Bennett & Peirce, 1991b Microfilariae: Ashford et al. , 1976 Haemoproteus fringillae: Peirce et al., 1977b (now H. afri- Plasmodium sp.: Bennett et al., 1978 canus) Trypanoso/1Ul bouffardi: Molyneux, 1973b Haemoproteus orizivorae: Peirce, 1984a Trypanoso/1Ul sp.: Ashford eta/., 1976 Haemoproteus sp.: Oosthuizen & Markus, 1967b; Peirce, Unidentified parasite: Bennett: et al., 1978 1976b Leucocytozoon fringillinarum: Peirce et al., 1977b Vidua Iorenzi Leucocytozoon sp.: Oosthuizen & Markus, 1967b No parasites seen: Wink & Bennett, 1976 Microfilariae: IRCAH Plasmodium relictum: Peirce, 1984a Vidua macroura Trypanoso/1Ul everetti: Peirce, 1976b; Peirce, 1984a Microfilariae: Leger, 1913 Trypanoso/1Ul sp.: Oosthuizen & Markus, 1967b Plasmodium octamerium: Manwell & Rossi, 1975 Trypanoso/1Ul vidua: Kerandel, 1913 Pytilia phoenicoptera Trypanoso/1Ul sp.: Kerandel, 1909; Leger & Leger, 1914c Haemoproteus sp.: Ashford eta/., 1976 Leucocytozoon sp.: Ashford eta/., 1976 Vidua orienta/is Microfilariae: Ashford et al., 1976 No parasites seen: Bennett et at. , 1978; Peirce, 1984a Trypanoso/1Ul sp.: Ashford eta/., 1976 Vidua paradisea Spermophaga rulicapilla Haemoproteussp. : IRCAH No parasites seen: Bennett et al., 1974; Peirce, 1984i Leucocytozoon fringillinarum: Peirce, 1984a Leucocytozoon sp.: Ashford et al., 1976 Spermestes bicolor Haemoproteus africanus: Bennett & Peirce, 1991b Viduaregia Haemoproteus frinsillaelorizovorae: Bennett et al., 1974 No parassites seen: IRCAH (now H. africanus) Spermestes cucullata FALCONIDAE Atoxoplas/1Ul sp.: Rousselot, 1953 Haemoproteus africanus: Bennett & Peirce, 1991b Falco amurensis Haemoproteus fringillae/orizivorae: Bennett & Herman, No parasites seen: VRI 1976; Bennett et al., 1974; Williams et al., 1977 (now H. africanus) Falcoaraea Haemoproteus sp.: Ashford eta/. , 1976; Rousselot, 1953 No parasites seen: Peirce, 1984i Hepatozoon spermesti: Rousselot, 1953 Microfilariae: Williams eta/. , 1977 Falco ardosiaceus Plasmodium re/ictum: Rousselot, 1943, 1953 No parasites seen: Ashford eta/., 1976 Trypanoso/1Ul sp. : Ashford eta/., 1976 Falco biarmicus Spermestes fringilloides Haemoproteus tinnunculi: IRCAH Haemoproteus sp.: Ashford eta/., 1976; Schwetz, 1931 Falco cherrug Leucocytozoon sp.: Schwetz, 1931, 1935c Haemoproteus tinnunculi: Peirce & Cooper, 1977b Spermestes malabarica Falco chicquera No parasites seen: IRCAH No parasites seen: IRCAH Spermestes montana Falco newtoni No parasites seen: IRCAH Haemoproteus tinnunculi: IRCAH 46 G. F. BENNETI, R. A. EARLE,HESTERDUTOIT& F. W. HUCHZERMEYER

Falco rupicoloides Leucocytozoon dubreuili: Peirce, 1984a Haemoproteus tinnunculi: IRCAH Leucocytozoon fringillinarum: Bennett & Herman, 1976; Haemoproteus sp. : IRCAH Peirce, 1984a Leucocytozoon toddi: IRCAH Leucocytozoon majoris: Bennett & Herman, 1976; Peirce, 1984a Falco subbuteo Leucocytozoon sp.: Ashford et al., 1976 Haemoproteus tinnunculi: IRCAH Microfilariae: Ashford eta!., 1976 Plasmodium rouxi: Ashford eta/., 1976 Falco tinnunculus Plasmodium: sp.: Cheke, 1971 Haemoproteus tinnunculi: Peirce et al., 1990 Trypanosoma bouffardi: Peirce, 1984a Leucocytozoon toddi: IRCAH Trypanosoma everetti: Peirce, 1984a Trypanosoma: sp.: Ashford et al., 1976 Polihierax semitorquatus No parasites seen: IRCAH Serinus nigriceps Leucocytozoon sp.: Ashford eta/., 1976 Plasmodium relictum: Ashford eta/., 1976 FREGATIDAE Serinus striolatus Fregata ariel Leucocytozoon sp.: Ashford et al., 1976 Haemoproteus sp.: Lowery, 1971 Serinus sulphuratus Fregata minor Haemoproteus orizivorae: Peirce, 1984a Haemoproteus sp.: Lowery, 1971 Leucocytozoon fringillinarum: Peirce, 1984a Leucocytozoon majoris: Peirce, 1984a FRINGILLIDAE Plasmodium vaughani: Peirce, 1984a Trypanosoma bouffardi: Peirce, 1984a Serinus albogularis Serinus tristriatus Leucocytozoon fringillinarum: IRCAH Leucocytozoon sp.: Ashford eta/., 1976 Serinus atrogularis Haemoproteus fringillae: IRCAH Haemoproteus sp.: Ashford et al., 1976 GLAREOLIDAE Lankesterella sp.: Ashford et al., 1976 Rhinoptilus africanus Leucocytozoon fringillinarum: Bennett & Herman, 1976 Leucocytozoon majoris: Bennett & Herman, 1976 No parasites seen: IRCAH Serinus canicollis Rhinoptilus chalcopterus Microfilariae: IRCAH No parasites seen: IRCAH Trypanosoma sp.: IRCAH Serinus citrinelloides GRUIDAE Haemoproteus sp.: Ashford et al., 1976 Anthropoides paradisea Leucocytozoon fringillinarum: Bennett & Herman, 1976 Leucocytozoon sp.: Ashford eta/., 1976 No parasites seen: IRCAH Microfilariae: Ashford et al., 1976 Trypanosomasp.: Ashford eta/., 1976 Balearica pavonina Haemoproteus antigonis: IRCAH Serinus citrinipectus Haemoproteus balearicae: Peirce, 1973b; Peirce, 1984i No parasites seen: Oosthuizen & Markus, 1967b Haemoproteus sp.: Rodhain et al., 1913a; Todd & Wolbach, 1912 & numerous zoo records from Europe summarized by Serinus dorsostriatus Bennett et al., 1982 Haemoproteus sp.: Peirce & Backhurst, 1970 Lankesterella sp.: Peirce, 1974 Leucocytozoon sp.: Rodhain et al., 1913a Serinus flaviventris Nuttallia balearicae: Peirce, 1973a (now Babesia balearicae Haemoproteus chloris: IRCAH Peirce, 1975) Plasmodium sp.: Numerous European zoo records summa­ Serinus gularis rized by Bennett et al., 1982 Haemoproteus chloris: IRCAH Balearica regulorum Haemoproteus macropigmentatus: IRCAH Leucocytozoon fringillinarum: IRCAH Haemoproteus balearicae: Peirce, 1973b Leucocytozoon majoris: IRCAH Haemoproteus sp. : Rodhain et al., 1913a; Rodhain & Fain, Leucocytozoon sp.: IRCAH 1954; Schwetz, 1944, 1945 Microfilariae: IRCAH Leucocytozoon sp.: Rodhain eta/., 1913a Plasmodium sp.: Schwetz, 1944 Serinus Jeucopygius Haemoproteus columbae: Rousselot, 1943 (now H. afri- canus) HIRUNDINIDAE Haemoproteus sp.: Ashford et al., 1976; Rousselot, 1953 Delichon urbica Leucocytozoon sp.: Rousselot, 1953 Plasmodiumsp.: IRCAH Haemoproteus sp.: Peirce, 1984a Serinus mennelli Hirundo abyssinica Haemoproteus fringillae: Peirce, 1984a Haemoproteus prognei: White & Bennett, 1978 Haemoproteus sp.: Bennett eta/., 1974, 1977 Serinus mozambicus Trypanosoma sp.: Ashford et al., 1976 Haemoproteus fringillae: Peirce, 1984a Hirundo aethiopica Haemoproteus columbae: Rousselot, 1942, 1943 (now H. fringillae) Haemoproteus sp.: Ashford eta/., 1976; Balfour, 1908b Haemoproteus orizivorae: Peirce, 1984a Haemoproteus sp.: Ashford et a/., 1976; Balfour, 1908b; Hirundo albigularis Cheke, 1971; Rousselot, 1953 No parasites seen: IRCAH 47 HOST-PARASITE CATALOGUE OF THE HAEMATOZOA OF THE SUB-SAHARAN BIRDS

Hirundo cucuHata ]ACANIDAE Hepatozoon atticorae: Bennett & Peirce, 1989b Leucocytozoon sp.: IRCAH Actophilornis africana Plasmodium polare: IRCAH Leucocytozoon sp.: Ashford et al., 1976 Microfilariae: Ashford et al. , 1976 Hirundo daurica Trypanosoma sp.: Ashford et al., 1976 Babesia rustica: Peirce et al., 1977 Hirundo dimidiata LANliNAE No parasites seen: IRCAH (LANliDAE) Hirundo fuligula Cort-inella cort-ina Haemoproteus sp.: Bennett & Herman, 1976; White & Ben· nett, 1978 Haemoproteus columbae: Rousselot, 1942, 1943 (now H. lanii) Hirundo rustica Haemoproteus sp.: Leger & Leger, 1914c Leucocytozoon sp.: Leger & Leger, 1914a No parasites seen: Peirce, 1984a; Peirce et al. , 1977b Hirundo semirufa Conine/la melanoleuca Trypanosoma avium: IRCAH Trypanosoma sp.: Oosthuizen & Markus, 1969b Hirundo senegalensis Laniarius aethiopicus Haemoproteus sp.: Ashford et al., 1976; White & Bennett, Haemoproteus sp.: Ashford et al., 1976 1978 Leucocytozoon sp.: Ashford et al. , 1976 Microfilariae: Ashford et al. , 1976 Hirundo smithii Plasmodium circumflexum: Ashford et al., 1976 Microfilariae: Ashford et al., 1976 Plasmodium relictum: Ashford et al., 1967 Plasmodium vaughani: Ashford et al. , 1967 Hirundo spilodera Haemoproteus prognei: IRCAH Laniarius atrococcineus Haemoproteussp.: IRCAH Hepatozoon: IRCAH Hepatozoon atticorae: Bennett & Peirce, 1989b Microfilariae: IRCAH Microfilariae: IRCAH Plasmodium vaughani: IRCAH Laniarius erythrogaster Phedina borbonica Haemoproteus lanii: Peirce, 1977 Haemoproteus sp.: Ashford et al., 1976; Peirce, 1972b Trypanosoma phedinae: Peirce et al., 1977a Leucocytozoon sp.: Ashford et al., 1976 Psalidoprocne holomelas Laniarius ferrugineus No parasites seen: Bennett & Herman, 1976; Bennett et al., 1977 Leucocytozoon fringillinarum: Bennett & Herman, 1976 Leucocytozoon majoris: Bennett & Herman, 1976 (re-identi­ Psalidoprocne orienta/is fied as L. balmorali) Leucocytozoon sp.: Oosthuizen & Markus, 1967b No parasites seen: IRCAH Microfilariae: Peirce, 1984a Psalidoprocne pristoptera Laniarius funebris No parasites seen: Ashford et al., 1976; Peirce, 1984a Haemoproteus sp.: Ashford et al. , 1976 Pseudhirundo griseopyga Leucocytozoon sp.: Ashford et al., 1976 Haemoproteus stellaris: White & Bennett, 1978 Plasmodium sp.: Ashford et al. , 1976 Haemoproteus sp.: Bennett et al., 1977 Lanius collaris Riparia cincta A toxoplasma sp.: Gamham, 1950 Plasmodium nucleophilum: Bennett et al. , 1974 Haemoproteus lanii: Bennett & Herman, 1976 Plasmodium relictum: Bennett et al., 1974 Leucocytozoon fringillinarum : Bennett & Herman, 1976 Leucocytozoon majoris: Bennett & Herman, 1976 (re-identi- Riparia paludicola fied as L. balmorali) Haemoproteus sp.: Ashford et al. , 1976 Plasmodium relictum: Bennett & Herman, 1976 Hepatozoon atticorae: Bennett & Peirce, 1989b Plasmodium vaughani: IRCAH Leucocytozoon sp.: Ashford et al., 1976 Microfilariae: Ashford et al. , 1976 Lanius collurio Trypanosoma sp.: Ashford et al., 1976 Haemoproteus lanii: Bennett & Herman, 1976; Peirce et al. , Riparia riparia 1977b Haemoproteus sp.: Ashford et al., 1976 No parasites seen: Peirce, 1984a Leucocytozoon balmorali: VRI Leucocytozoon fringillinarum: Peirce et al, 1977b lNDICATORIDAE Plasmodium vaughani: Bennett & Herman, 1976 Plasmodium sp.: Peirce et al., 1977b Indicator indicator Haemoproteus indicator: Bennett et al. , 1986 Lanius dorsalis Haemoproteus sp.: Ashford et al., 1976; Bennett et al. , 1974 No parasites seen: Ashford et al., 1976 Leucocytozoon sp.: Ashford et al., 1976 Indicator minor Lanius excubitoroides Leucocytozoon sp.: Ashford et al. , 1976 No parasites seen: Ashford etal., 1976 Indicator t-ariegatus Lanius nubicus Haemoproteus sp.: Ashford et al., 1976 Haemoproteus sp.: Ashford et al., 1976 48 G. F. BENNETT, R. A . EARLE, HESTERDUTOIT& F. W. HUCHZERMEYER

MALACONOTINAE Gygasalba No parasites seen: Lowery, 1971; Peirce, 1984i (LANIIDAE) Larus argentatus Dryoscopus cub/a Haemoproteus lari: IRCAH Haemogregarina sp.: Peirce, 1984a (probably Hepatozoon sp.) Larus cirrocephalus Haemoproteus cublae: Peirce, 1984e Haemoproteus /anii: Bennett & Herman, 1976 Haemoproteus sp.: Rousselot, 1953 Haemoproteus sp.: Peirce, 1984a Larus dominicanus Hepatozoon sp.: IRCAH Leucocytozoon ba/morali: Peirce, 1984d No parasites seen: IRCAH Leucocytozoon sp.: Oosthuizen & Markus, 1967b; Peirce, Larus serranus 1984a Microfilariae: Bennett & Herman, 1976; Oosthuizen & Mar- No parasites seen: IRCAH kus, 1967b; Peirce, 1984a Plasmodium vaughani: Peirce, 1984a Sterna fuscata Trypanosoma everetti: Peirce, 1984a No parasites seen: Peirce & Freare, 1978; Peirce 1984i Trypanosoma sp.: Peirce, 1984a Dryoscopus gambensis MEROPIDAE Leucocytozoon sp.: Ashford eta/., 1976 Merops albicollis Microfilariae: Ashford eta/., 1976 Haemoproteus meropis: Bennett et al., 1978 Plasmodium re/ictum: Ashford eta/., 1976 Haemoproteus sp.: Ashford et al., 1976; Bennett et al. , 1974, Plasmodium rowci: Ashford eta/., 1976 1977; Minchin, 1910 Trypanosoma sp.: Ashford eta/., 1976 Microfilariae; Ashford et al., 1976; Bennett et al., 1977; Min­ Dryoscopus pringlii chin, 1910 No parasites seen: Peirce, 1984i Plasmodium vaughani: Ashford et al., 1976 Trypanosoma sp.: Minchin, 1910 Malaconotus blanchoti Merops apiaster Leucocytozoon balmorali: Peirce, 1984d Leucocytozoon sp.: Rousselot, 1953; Peirce, 1984a No parasites seen: IRCAH Microfilariae: Petree, 1984a Merops bullockoides Malaconotus bocagei No parasites seen: Bennett & Herman, 1976; Peirce, 1984i No parasites seen: Bennett eta/., 1977 Merops bulocki Malaconotus cruentus Haemoproteus meropis: Peirce, 1984i Haemoproteus sp.: Neave, 1906 Plasmodium sp.: Cheke, 1971 Trypanosoma sp.: Neave, 1906 Merops gularis Malaconotus sulfuropectus Haemoproteus meropis: Bennett et al., 1978 Haemoproteus /anii: Bennett eta/., 1977 Haemoproteus sp.: Theiler, 1930 Haemoproteus sp.: Ashford eta/., 1976 Merops nubicoides Plasmodium relictum: Ashford et a/., 1976 Haemoproteus sp.: Oosthuizen & Markus, 1967a Nilausafer Leucocytozoon sp.: Oosthuizen & Markus, 1967a Leucocytozoon balmorali: Peirce, 1984i Microfilariae: Oosthuizen & Markus, 1967a Plasmodium relictum: Ashford eta/., 1976 Merops nubicus Tchagra australis Haemoproteus sp.: Ashford et al. , 1976; Balfour, 1908b Leucocytozoon balmorali: Peirce, 1984d Microfilariae: Ashford et al., 1976; Kerandel, 1909 Leucocytozoon sp.: Bennett eta/., 1974; Peirce, 1984a Merops oreobates Microfilariae: Petree, 1984a Plasmodium vaughani: Bennett eta/., 1974 Microfilariae: Ashford et al. , 1976 Trypanosoma avium: Bennett eta/., 1974 Trypanosoma sp.: Ashford et al., 1976 Tchagra minuta Merops pusillus Haemoproteus /anii: Bennett & Herman, 1976 Haemoproteus sp.: Schwetz, 1938 Haemoproteus sp.: Ashford eta/., 1976 Leucocytozoon sp.: Ashford eta/., 1976 Merops superciliosus Microfilariae: Ashford eta/., 1976 Plasmodium vaughani: Bennett et al., 1977 Plasmodium vaughani: Ashford et al., 1976 Trypanosomasp.: Ashford eta/., 1976 Merops variegatus Haemoproteus lairdi: Bennett et al., 1978 Tchagra senega/a Haemoproteus sp.: Bennett et al., 1974, 1977 Leucocytozoon balmorali: Peirce, 1984d Microfilariae: Schwetz, 1938 Leucocytozoon sp.: Ashford eta/., 1976; Peirce, 1984a Microfilariae: Ashford eta/., 1976 MOTACILLIDAE Plasmodium rowci: Ashford et al., 1976; Peirce, 1984a Plasmodium vaughani: Ashford et al., 1976 Anthus crenatus Telophorus zeylonus Haemoproteus sp.: Ashford et al., 1976 Leucocytozoon sp.: Ashford et al., 1976 Leucocytozoon balmorali: IRCAH Plasmodium vaughani: IRCAH Leucocytozoon sp.: IRCAH Unknown parasite: IRCAH Anthus leucophrys Haemoproteus sp. : Bennett et al., 1977 LARIDAE Leucocytozoon sp.: Peirce, 1984a Microfilariae: Bennett et al., 1977 Anous stolidus Plasmodium rouxi: Peirce, 1984a Haemoproteussp.: Lowery, 1971 Plasmodium vaughani: Bennett et al., 1977 49 HOST-PARASITECATALOGUEOFTHEHAEMATOZOAOFTHESUB-SAHARANBIRDS

Anthus not-aeseelandiae Melaenornis edoliodes No parasites seen: Peirce, 1984a Haemoproteus sp.: Ashford et a/., 1976; Commes, 1918b; Rousselot, 1953 Antbus similis Leucocytozoon sp.: Ashford et at. , 1976 Microfilariae: Peirce, 1984a Melaenornis mariquensis Plasmodium relictum: Ashford eta/., 1976; Peirce, 1984a Plasmodium sp.: Ashford eta/., 1976 No parasites seen: IRCAH Trypanosama everetti: Peirce, 1984a Melaenornis pammelaina Anthus trit-ialis Haemoproteus balmorali: Peirce, 1984b No parasites seen: Peirce et al., 1977b Haemoproteus sp.: Peirce, 1984a Plasmodium circumflexum: IRCAH Macronyx aurantiigula Plasmodium rouxi: Peirce, 1984a No parasites seen: Peirce eta/., 1977b Muscicapa adusta Macronyx capensis No parasites seen: Ashford eta/., 1976 No parasites seen: IRCAH Muscicapa caerulescens Macronyx croceus Trypanosoma sp.: Peirce, 1984a Haemoproteus sp.: Bennett eta/., 1977 Muscicapa griseigularis Motacilla aguimp Haemoproteussp. : Bennett eta/., 1974 Haemoproteus sp.: Cowper, 1969 Muscicapa striata Leucocytozoon sp.: Cowper, 1969 Haemoproteus balmorali: Peirce, 1984b Microfilariae: Cowper, 1969 Haemoproteus sp.: Bennett et a/., 1974; Oosthuizen & Markus, 1967b; Peirce, 1984b Motacilla capensis Microfilariae: Ashford eta/., 1976 Haemoproteus anthi: IRCAH Trypanosoma sp.: Peirce, 1984a Leucocytozoon sp.: VRI Plasmodium relictum: IRCAH Myioparus plumbeus Haemoproteussp.: Bennett eta/., 1977 Motacilla flat-a Plasmodium vaughani: Bennett eta/., 1977 Haemoproteus motacillae: Bennett & Peirce, 1990d Platysteira peltata Tmetothylacus tenellus No parasites seen: IRCAH No parasites seen: Peirce, 1976b; Peirce eta/., 1977b Sigelus silens Haemoproteus balmora/i: IRCAH MUSCICAPINAE Tcbitrea plumbeiceps (MUSCICAPIDAE) No parasites seen: Bennett eta/., 1977 Batisminor Tcbitrea suabelica No parasites seen: Ashford eta/., 1976 No parasites seen: IRCAH Batismixta Terpsipbone bourbonnensis No parasites seen: IRCAH Atoxoplasma sp.: Peirce eta/., 1977a Batis molitor Haemogregarina sp.: Peirce, 1977 (probably Hepatozoon sp.) Plasmodium sp.: Peirce, 1984a Terpsipbone mutata Batis orientalis No parasites seen: Peirce & Cheke, 1977 Leucocytozoon sp.: Ashford et at., 1976 Terpsipbone rutit-enter Plasmodium rouxi: Ashford eta/., 1976 Haemoproteus sp.: Bennett eta/., 1977 Batis pririt Terpsipbone riridis No parasites seen: IRCAH Haemoproteus sp.: Archetti, 1946, 1947; Peirce, 1984a, Schwetz, 1938 Bradornis infuscatus Leucocytozoon sp.: Ashford eta/., 1976 No parasites seen: IRCAH Microfilariae: Ashford eta/., 1976 Bradornis microrbyncbus Plasmodium nucleophilum: Ashford eta/., 1976 Plasmodium vaughani: Ashford eta/., 1976; Peirce, 1984a No parasites seen: Ashford eta/., 1976 Trypanosoma sp.: Ashford et at. , 1976 Bradornis pallidus Trocbocercus albonotatus Haemoproteus balmorali: Peirce, 1984b No parasites seen: Cheke, 1972 Haemoproteus sp.: Ashford et a/., 1976; Oosthuizen & Markus, 1969b; Peirce, 1984a Leucocytozoon sp.: Ashford eta/., 1976 MUSOPHAGIDAE Plasmodium rouxi: Peirce, 1984a Corytbaeola cristata Plasmodium vaughani: Peirce, 1984a Haemoproteus sp.: Minchin, 1910; Schwetz, 1931 Trypanosoma sp.: Peirce, 1984a Leucocytozoon sp.: Schwetz, 1931 , 1935c Bradornis pumilus Microfilariae: Minchin, 1910; Schwetz, 1931, 1935c Trypanosoma cristatae: Schwetz, 1931 Haemoproteus sp.: Bennett & Herman, 1976 Trypanosoma sp.: Minchin, 1910; Schwetz, 1933 Leucocytozoonfringillinarum: Bennett & Herman, 1976 Plasmodium vaughani: Bennett & Herman, 1976 Corythaixoides concolor Dioptornis liscberi Haemoproteus montezi: VRI Hepatozoonsp.: VRI Haemoproteus or Plasmodium: Bennett & Herman, 1976 Leucocytozoon dinizi: IRCAH, VRI Erytbrocercus mccallii Corythaixoides leucogaster No parasites seen: Peirce, 1969 Leucocytozoon dinizi: Peirce eta/., 1977b Ficedula albicollis Corythaixoides personata No parasites seen: Smith & Cox, 1972 Leucocytozoon sp.: Gamham, 1950

50 G . F. BENNETI, R. A. EARLE,HESTERDUTOIT&F. W. HUCHZERMEYER

Crinifer piscator Nectarinia cuprea Leucocytozoon dinizi: Tendeiro, 1947a, 1948a Haemoproteus sp.: Bennett et al. , 1974 Plasmodium vaughani: Ashford et al. , 1976 Musophaga violacea Plasmodium sp.: Peirce, 1984a Haemoproteus sp.: Minchin, 1910 Haemoproteus montezi: Bennett & Peirce, 1990c Nectarinia cyanolaema Leucocytozoon fringillinarum: Bennett & Herman, 1976 No parasites seen: IRCAH Plasmodium vaughani: Bennett & Herman, 1976 Nectarinia famosa Tauraco corythaix Haemoproteus sequeirae: IRCAH Leucocytozoon dinizi: VRI Leucocytozoon sp.: IRCAH Microfilariae: IRCAH Tuuraco hartlaubl Plasmodium sp.: IRCAH Haemoproteus montezi: Bennett & Herman, 1976; Bennett Trypanosoma avium: IRCAH & Peirce, 1990c; Cheke, 1978 Haemoproteus sp.: Cheke, 1972; Peirce, 1984i Nectarinia habessinica Leucocytozoon dinizi: Cheke, 1978 Haemoproteus sp.: Ashford et al., 1976 Leucocytozoon sp.: Cheke, 1972 Nectarinia kilimensis Tauraco livingston# Leucocytozoon fringillinarum: Bennett & Herman, 1976 Haemogregarina sp.: Bennett & Herman, 1976 (probably Leucocytozoon majoris: Bennett & Herman, 1976 Hepatozoon sp.) Microfilariae: Bennett & Herman, 1976 Haemoproteus montezi: Bennett & Herman, 1976; Bennett & Peirce, 1991a Nectarinia marquensis No parasites seen: Ashford et al., 1976; Peirce & Backhurst, Tauraco porphyriolophus 1970 Haemoproteus montezi: Travassos Santos Dias, 1953 Nectarinia mediocris Tauraco schuetii Leucocytozoon sp.: Cheke, 1972 Leucocytozoon sp.: Schwetz, 1935c Nectarinia olivacea Haemoproteus montezi: Bennett & Herm;m, 1976 (now H. NECTARINIIDAE sequeirae) Haemoproteus raymundi: Wink & Bennett, 1976 (now of H. Anthreptes coHil.ris sequeirae) Haemoproteus sp.: Bennett & Herman, 1976 Haemoproteus sequeirae: IRCAH Hepatozoon sp.: IRCAH Haemoproteus sp.: Ashford et al., 1976 Leucocytozoon fringillinarum: Bennett & Herman, 1976 Leucocytozoon fringillinarum: Bennett & Herman, 1976 Plasmodium vaughani: Bennett & Herman, 1976 Leucocytozoon majoris: IRCAH Trypanosoma avium: Bennett & Herman, 1976 Leucocytozoon sp.: Ashford et al., 1976; Schwetz, 1938 Trypanosoma sp.: IRCAH Microfilariae: Bennett & Herman, 1976 Nuttallia sp.: Bennett & Herman, 1976 (now Babesia sp.) Anthreptes Jonguemarei Plasmodium relictum: Bennett & Herman, 1976 No parasites seen: Peirce, 1984a Plasmodium rouxi: Bennett & Herman, 1976 Plasmodium vaughani: Bennett & Herman, 1976; Wink & Anthreptes orienta/is Bennett, 1976 Haemoproteus sp.: Ashford eta/., 1976 Plasmodium sp.: Bennett et al. , 1974 Trypanosoma avium: Bennett & Herman, 1976; Wink & Anthreptes platurus Bennett, 1976 Haemoproteus sp.: Ashford et al. , 1976; Leger & Leger, Trypanosoma sp.: IRCAH 1914c Micro filariae: Ashford eta/. , 1976 Nectarinia pulchella Trypanosoma sp.: Leger & Leger, 1914c Haemoproteus columbae: Rousselot, 1943 (now H. sequeirae) Nectarinia afra Haemoproteus montezi: Williams et al., 1977 (now Haemoproteus sequeirae: IRCAH H. sequeirae) Leucocytozoon sp.: IRCAH Haemoproteus sp.: Ashford et al., 1976 Plasmodium relictum: VRI Nectarinia seimundi Nectarinia amethystina Trypanosoma avium: Wink & Bennett, 1976 Haemoproteus sequeirae: Bennett & Herman, 1976; Peirce, Nectarinia senegalensis 1984a Haemoproteus columbae: Rousselot, 1942, 1943 (now Haemoproteus sp.: Oosthuizen & Markus, 1969b H. sequeirae) Leucocytozoon dubreuili: Peirce 1984a Haemoproteus sequeirae: Peirce, 1984a Leucocytozoon fringillinarum: Bennett & Herman, 1976 Haemoproteus sp.: Ashford et al., 1976; Bennett & Herman, Microfilariae: Bennett & Herman, 1976 1976; Bennett et al., 1974, 1977; Commes, 1918b; Cowper, Plasmodium vaughani: IRCAH 1969; Oosthuizen & Markus, 1969b; Rousselot, 1953 Trypanosoma everetti: Peirce, 1984a Leucocytozoon majoris: Peirce, 1984a Trypanosoma sp.: IRCAH Leucocytozoon sp.: Ashford et al., 1976; Rousselot, 1953 Nectarinia bifasciata Microfilariae: Ashford et al. , 1976 Plasmodium sp.: Peirce, 1984a Leucocytozoon fringillinarum: Peirce, 1984a Trypanosoma everetti: Peirce, 1984a Nectarinia chalybea Nectarinia sovimanga No parasites seen: Peirce, 1984a No parasites seen: Lowery, 1971 Nectarinia chloropygia Nectarinia talatala Haemoproteus sp.: Bennett et al., 1977 Leucocytozoon sp: IRCAH Haemoproteus sequeirae: IRCAH Plasmodium elongatum: Schwetz, 1938 Nectarinia venusta Trypanosoma sp.: Leger & Leger, 1914c; Schwetz, 1938 Haemoproteus sp.: Bennett & Herman, 1976 Leucocytozoon sp.: Bennett & Herman, 1976 Nectarinia coccinogaster Plasmodium sp.: Peirce, 1984a Haemoproteus sequeirae: Tendeiro, 1947a, 1948a Trypanosoma sp.: Ashford et al. , 1976 51 HOST-PARASITE CATALOGUE OF THE HAEMATOZOA OF THE SUB-SAHARAN BIRDS

Nectarinis verticalis Oriolus lsrvstus Haemoproteus sp.: Bennett et al. , 1977 Leucocytozoon sp.: Ashford et al. , 1976 Plasmodium sp.: Ashford et al., 1976 NUMIDINAE Oriolus nigripennis (PHASIANIDAE) Leucocytozoon sp.: Schwetz, 1938 Gutters edouarcli Oriolus oriolus No parasites seen: IRCAH Haemoproteus orioli: Peirce, 1984a, f Trypanosoma sp.: Peirce, 1984a Gutters plumifers Haemoproteus sp.: Rodhain & Fain, 1954 0TIDIDAE Plasmodium rouxi: Rodhain & Fain, 1954 Ardeotis kori Plasmodium sp.: Rodhain & Fain, 1954 Haemoproteus tendeiroi: Peirce, 1977 Gutters pucherani Leucocytozoon sp.: IRCAH Haemoproteus sp.: Fallis et al., 1973b; Keysselitz & Mayer, Eupodotis cafrs 1909 Haemoproteus tendeiroi: VRl Leucocytozoon caulleryi: Fallis et al., 1973b (possibly identi­ Leucocytozoon sp.: VRI fied in error) Eupodotis hsrtlsubi Leucocytozoon neavei: Bennett & Herman, 1976; Fallis et al., 1973b Haemoproteus sp.: Neave, 1906 Leucocytozoon schoutedeni: Fallis et a/. ,. 1973b (possibly Eupodotis (= Lissotis) melsnogaster identified in error) Haemoproteus telfordi: Bennett et al., 1975; Bennett & Leucocytozoon sp.: Keysselitz & Mayer, 1909 Herman, 1976 Microfilariae: Fallis et al., 1973b; Keysselitz & Mayer, 1909 Haemoproteus tendeiroi: Bennett et al., 1975; Travassos San­ Plasmodium relictum: Bennett & Herman, 1976 tos Dias, 1953 Trypanosoma numidae: Fallis et al., 1973a Haemoproteus sp.: Hoare, 1932; Rodhain et al. , 1913a; Trypanosoma sp.: Keysselitz & Mayer, 1909 Schwetz, 1938; Todd & Wolbach, 1912 Numids melesgris Plasmodium relictum: Rousselot, 1953 Aegyptianella sp.: Huff, Marshbank, Saroff, Scrimshaw & Eupodotis vigorsii Shiroishi, 1950; Huchzermeyer, Horak & Braack, 1991 Haemoproteus tendeiroi: VR1 Haemoproteus pratasi: Tendeiro, 1947a, 1948a; Bennett & Leucocytozoon sp.: VRI Herman, 1976; VRl Haemoproteus silvai: Son, 1960 (now H. pratasi) Neotis Judwigii Haemoproteus sp.: Ashford et al. , 1976; Fallis et al., 1973b; lf.aemoproteus tendeiroi: IRCAH Minchin, 1910; Neave, 1906; Oosthuizen & Markus, 1967a, 1969b; Rodhain et al., 1913b Thomas & Dobson, PARIDAE 1975; Todd & Wolbach, 1912 Parusafer Hepatozoon sp.: VR1 Leucocytozoon caulleryi: Fallis et al., 1973b (possibly identi­ No parasites seen: IRCAH fied in error) Parus cseruleus Leucocytozoon costae: Tendeiro, 1947a, 1948a (syn of Lankesterella sp.: Bennett et al., 1974 L. neavei) Leucocytozoon neavei: Bennett & Herman, 1976; Fallis et Psrus cinserescens al., 1973b; Minchin, 1910; Oosthuizen & Markus, 1967a; Leucocytozoon majoris: IRCAH Rodbain et al. , 1913b; Rousselot, 1953; Son, 1960; Wenyon, 1908; VRl Parus funereus Leucocytozoon numidae: Kerandel, 1913 (syn of L. neavei) No parasites seen: Peirce, 1984i Leucocytozoon sp. : Ashford et al., 1976; Hoare, 1932; Parus griseiventris Kerandel, 1909; Leger & Leger, 1914a; Oosthuizen & Markus, 1967a, 1969b; Rodhain et al., 1913b; Thomas & Leucocytozoon majoris: Peirce, 1984a Dobson, 1975 Plasmodium relictum: Peirce, 1984a Microfilariae: Bennett & Herman, 1976; Kerandel, 1909; Parus leucomelas Neave, 1909; Tendeiro, 1948b; Oosthuizen & Markus, Leucocytozoon majoris: Peirce, 1984a 1967a; VR1 Leucocytozoon sp.: Ashford et al. , 1976 Plasmodium circumflexum: Huchzermeyer & Van der Vyver, 1991; VRl Parusniger Plasmodium durae: Ashford et al., 1976 (probably in error as Leucocytozoon dubreuili: Peirce, 1984a this species cannot be transmitted experimentally to Trypanosoma everetti: Peirce, 1984a guineafowls} Plasmodiumfallax: Huff eta/., 1950 Parus rufiventris Plasmodium relictum: Rousselot, 1953 Leucocytozoon majoris: Peirce, 1984a Plasmodium sp.: Minchin, 1910; Neave, 1906; Peirce & Trypanosoma sp.: Peirce, 1984a Backhurst, 1970; Thomas & Dobson, 1975; Wenyon, 1908; VRl PASSERIDAE Trypanosoma avium: Bennett & Herman, 1976 Passer diffusus Trypanosoma numidae: Fallis et al. , 1973a; Kerandel, 1909, 1913 Haemoproteus sp.: Oosthuizen & Markus, 1967b Trypanosoma sp.: VRI Microfilariae : Leger, 1913 Unknown parasite: VR1 Passer domesticus Atoxoplasma sp.: Peirce et al. , 1977a 0RIOLIDAE Haemoproteus passeris: IRCAH . Oriolus surstus Haemoproteus sp.: Maya & David, 1912 Leucocytozoon Jringillinarum: Peirce, 1979 Haemoproteus pintoi: Tendeiro, 1947a, 1948a (syn of H. Leucocytozoon sp.: Maya & David, 1912; Peirce, 1984a; orioli) Peirce, et al., 1977a Leucocytozoon fringillinarum: Bennett & Herman, 1976 Plasmodium sp.: Peirce, 1979 Microfilariae: Bennett & Herman, 1976 Trypanosoma mayae: Maya & David, 1912; Peirce et al., Trypanosoma avium: Bennett & Herman, 1976 1977a

52 G. F. BENNETI, R. A. EARLE, HESTER DU TOIT & F. W. HUCHZERMEYER

Passer eminibey Plocepasser superciliosus Haemoproteus sp.: Ashford et al., 1976 Haemoproteus sp.: Cowper, 1969 Leucocytozoon sp.: Ashford eta/., 1976 Leucocytozoon sp.: Ashford et al. , 1976 Micro filariae: Asbford et al., 1976 Plasmodium vaughani: Ashford et al., 1976 Plasmodium relictum: Ashford et al., 1976 Plasmodium vaughani: Ashford et al., 1976 Sporopipes frontalis Plasmodium sp.: Ashford et al., 1976 Plasmodium sp.: Manwell & Rossi, 1975 Passer griseus Sporopipes squamifrons Haemoproteus columbae: Rousselot, 1943 (now H. passeris) Haemoproteus passeris: Bennett & Herman, 1976 Haemoproteus fallisi: Peirce et al., 1977b Plasmodium nucleophilum: Manwell & Rossi, 1975 Haemoproteus fringillaelorizivorae: Bennett et al., 1974; Peirce et al., 1977b; Wink & Bennett, 1976 Haemoproteus passeris: Peirce, 1976a; Bennett & . Peirce, PHAETHONTIDAE 1991b Phaethon lepturus Haemoproteus sp.: Ashford et al., 1976; Garnham, 1950; No parasites seen: Lowery, 1971; Peirce, 1979 Peirce & Backhurst, 1970; Rousselot, 1953 Hepatozoon sp.: Rousselot, 1953 Phaethon rubricauda Leucocytozqon fringillinarum: Peirce et al., 1977b No parasites seen: Lowery, 1971; Peirce, 1979 Leucocytozoon monardi: Rodhain, 1931; Rousselot, 1953 (syn of L. fringillinarum) Leucocytozoon sp.: Ashford et al., 1976; Garnham, 1950; PHALACROCORACIDAE Peirce & Backhurst, 1970 Halietor africanus Micro filariae: Ashford et al., 1976; Wink & Bennett, 1976 Plasmodium relictum: Ashford et al., 1976 Haemoproteus sp.: Rodhain et al., 1913a Plasmodium vaughani: Ashford et al., 1976 Plasmodium sp.: Ashford et al., 1976 PHASIANINAE Trypanosoma avium: Wink & Bennett, 1976 (PHASIANIDAE) Trypanosomasp.: Ashford eta/., 1976 Afropavo congensis Passer iogoensis Haemoproteus chapini: et al., Haemoproteus passeris: Peirce, 1976a; Bennett & Peirce, van den Berghe 1963 ( syn of H. rileyi) 1991b Microfilariae: van den Berghe et al. , 1963 Haemoproteus sp.: Garnham, 1950 Leucocytozoon Jringillinarum: Bennett & Herman, 1976 Coturnix delegorguei Leucocytozoon sp.: Garnham, 1950 Plasmodium relictum: Garnham, 1950 Haemoproteus sp.: Rodhain et al., 1913a Microfilariae: Peirce et al., 1977b Passer luteus Plasmodium sp.: Peirce, 1976b Haemoproteus fringillaelorizivorae: Williams et al., 1977 Francolinus adspersus Haemoproteus passeris: Bennett & Peirce, 1990d Leucocytozoon schoutedeni: IRCAH Passer melanurus Microfilariae: IRCAH Atoxoplasma sp.: IRCAH Haemoproteus fringillaelorizivorae: IRCAH Francolinus afer Haemoproteus passeris: Bennet & Peirce, 1991b Leucocytozoon peaolopesi: Travassos Santos Dias, 1951 Heptazoon sp.: IRCAH Leucocytozoon schoutedeni: IRCAH Leucocytozoon sp.: Fantham & Robertson, 1927; Thomas & Microfilariae: Travassos Santos Dias, 1951 Dobson, 1975 Plasmodium juxtanucleare: VRI Leucocytozoon fringillinarum: IRCAH Leucocytozoon majoris: IRCAH Francolinus africanus Plasmodium octamerium: IRCAH Hepatozoon sp.: VRI Plasmodium polare: IRCAH Leucocytozoon macleani: VRI Plasmodium relictum: IRCAH Leucocytozoon peaolopesi: VRI Plasmodium sp.: IRCAH Leucocytozoon sp.: VRI Unknown parasites: IRCAH Microfilariae: VRI Plasmodium juxtanucleare: Earle, Huchzermeyer, Bennett & Petronia dentata Little, 1991 Haemoproteus sp.: Ashford eta/., 1976 Trypanosoma avium: VRI Plasmodium vaughani: Ashford et al., 1976 Francolinus ahantensis Petronia superciliaris Haemoproteus pratasi: Wink & Bennett, 1976 (now H. rileyi) Atoxoplasma sp.: Peirce, 1984a Leucocytozoon neavei: Wink & Bennett, 1976 (now L. Haemoproteus passeris: Bennett & Peirce, 1991b; Peirce, macleani) 1984a Microfilariae: Wink & Bennett, 1976 Leucocytozoon sp.: Peirce, 1984a Trypanosoma avium: Wink & Bennett, 1976 Plasmodiumsp.: Peirce, 1984a Trypanosoma sp.: Peirce, 1984a Francolinus bicalcaratus Petronia xanthocollis Haemoproteus sp.: Bennett et al., 1978 Leucocytozoon francolini: Kerandel, 1913; Leger & Leger, Haemoproteus passeris: Bennett & Peirce, 1991b 1914a; Rousselot, 1953; Tendeiro; 1948a (probably L. Philetairus socius macleani) Haemoproteus passeris: IRCAH Leucocytozoon sp.: Balfour & Wenyon, 1909; Kerandel, Trypanosoma everetti: IRCAH 1909; Leger & Leger, 1914a Microfilariae: Leger, 1913; Tendeiro, 1948a Plocepasser donaldsoni Trypanosoma aguedaferreirae: Tendeiro, 1947b, 1948a No parasites seen: Peirce & Backhurst, 1970 Trypanosoma francolini: Kerandel, 1913 Plocepasser mahali Trypanosoma sp.: Kerandel, 1909; Leger & Leger, 1914a Haemoproteus fringillae/orizivorae: Bennett et al. , 1977 Francolinuscapensis Haemoproteus passeris: Bennett & Peirce, 1991b Haemoproteus sp.: Cowper, 1969 Leucocytozoon sp.: Peirce & Backhurst, 1970 Trypanosoma sp.: Ashford et al., 1976; Peirce & Backhurst, Francolinus clappertoni 1970 Leucocytozoon sp.: Ashford et al. , 1976 53 HOST-PARASITECATALOGUEOFTHEHAEMATOZOAOFTHESUB-SAHARANBIRDS

Francolinus coqui Dendropicos abyssinicus Leucocytozoon sp.: Oosthuizen & Markus, 1969a; Ross, No parasites seen: Ashford et al., 1976 1911 Plasmodium sp.: Markus & Oosthuizen, 1972 Dendropicos fuscescens Trypanosoma sp.: Oosthuizen & Markus, 1969a No parasites seen: Bennett & Herman, 1976; Bennett et al., 1977; Oosthuizen & Markus, 1969b; Peirce, 1984a Francolinus leucoscepus Leucocytozoon neavei: de Jong, 1971b (now L. peaolopesi) Dendropicos xantholophus Leucocytozoon peaolopesi: IRCAH No parasites seen: Peirce, 1984i Leucocytozoon sp.: Ross, 1911 Plasmodium durae: Markus & Oosthuizen, 1972 Geocolaptes olivaceus Francolinus levaillantii Leucocytozoon squamatus: VRI; IRCAH Haemoproteus: Minchin, 1910 Jynx ruficollis Leucocytozoon: Hoare, 1932; Minchin, 1910 Leucocytozoon squamatus: IRCAH Microfilariae: Minchin, 1910 Jynx torquilla Francolin us levaillantoides Haemoproteus sp.: Ashford eta/., 1976 Haemoproteus sp.: Minchin, 1910 Leucocytozoon sp.: Ashford et al., 1976 Leucocytozoon peaolopesi: VRI Leucocytozoon sp.: Hoare, 1932 Mesopicos goertae Microfilariae: Minchin, 1910 Leucocytozoon sp.: Ashford et al., 1976 Trypanosoma avium: VRI Thripias namaquus Francolinus natalensis Haemoproteus sp.: Ashford et al., 1976 Leucocytozoon peaolopesi: VRI

Francolinus sephaena PLOCEIDAE Leucocytozoon sp.: Ashford et al. , 1976 Plasmodium juxtanucleare: Ashford et al., 1976 Amblyospiza albifrons Haemoproteus sp.: Ashford eta/., 1976 Francolinus squamatus Micro filariae: Ashford et al. , 1976 Leucocytozoon sp.: Ashford et al. , 1976; Ross, 1911; Plasmodium vaughani: Ashford et al., 1976 Schwetz, 1935c Euplectes afer Francolinus swainsonii Haemoproteus columbae: Rousselot, 1943 (now H. quelea) Haemoproteus sp.: Thomas & Dobson, 1975; VRI Haemoproteus quelea: IRCAH Leucocytozoon peaolopesi: IRCAH; VRI Haemoproteus sp.: Ashford eta/., 1976; Rousselot, 1953 Leucocytozoon sp.: Oosthuizen & Markus, 1967a; Thomas & Leucocytozoon sp.: Ashford et al. , 1976 Dobson, 1975; VRI Plasmodium sp.: Manwell, 1970; Manwell & Rossi, 1975 Plasmodium durae: de Jong, 1971a; VRI Plasmodium sp.: Markus & Oosthuizen, 1972 Euplectes albonotatus Trypanosoma sp.: VRI Haemoproteus fringillae: Peirce et al., 1977b (now H. quelea) Haemoproteus quelea: Bennett & Peirce, 1991b Lophura nycthemera Haemoproteus sp.: Ashford et al., 1976 No parasites seen: IRCAH Euplectes anomalus Ptilopachus petrosus Leucocytozoon sp.: Schwetz, 1931, 1935c Haemoproteus sp.: Todd & Wolbach, 1912 Plasmodium elongatum: Schwetz, 1935b Microfilariae: Cheke, 1971 Plasmodium relictum: Schwetz, 1935b Plasmodium sp.: Cheke, 1971 Euplectes ardens Haemoproteus fringillae: Bennett & Herman, 1976 (now H. PHOENICULIDAE quelea) Phoeniculus cyanomelas Haemoproteus passeris: Peirce, 1984a Haemoproteus quelea: Bennett & Peirce, 1991b No parasites seen: Oosthuizen & Markus, 1969b Haemoproteus sp.: Ashford eta/., 1976; Cowper, 1969 Phoeniculus purpureus Leucocytozoon sp.: Ashford et al., 1976 Plasmodium sp.: Cheke, 1971 Plasmodium rouxi: Ashford et al., 1976 Plasmodium vaughani: Peirce, 1984a Plasmodium sp.: Ashford et al., 1976 PICIDAE Euplectes axillaris Campethera abingoni Haemoproteus sp.: Ashford et al., 1976 No parasites seen: Oosthuizen & Markus, 1969b Euplectes capensis Campethera cailliautii Atoxoplasma sp.: Peirce, 1984a No parasites seen: Ashford et al., 1976 Haemoproteus passeris: Peirce, 1984a Microfilariae: Peirce, 1969 Campethera nivosa Leucocytozoon sp.: Peirce, 1984a No parasites seen: Bennett et al. , 1974, 1977 Plasmodium rouxi: Peirce, 1984a Campethera nubica Euplectes gierowii Haemoproteus sp.: Ashford eta/., 1976 Haemoproteus sp.: Ashford eta/., 1976 Leucocytozoon sp.: Ashford et al. , 1976 Euplectes hartlaubi Campethera permista Plasmodium vaughani: Bennett & Herman, 1976 Microfilariae: Schwetz, 1938 Euplectes hordeaceus Campethera punctuligera Haemoproteus sp.: Ashford et al., 1976; Leger & Leger, Haemoproteus borgesi: Tendeiro, 1947a, 1948a 1914c; Rousselot, 1953 54 G. F. BENNETI, R. A. EARLE,HESTERDUTOIT& F. W. HUCHZERMEYER

Leucocytozoon sp.: Leger & Leger, 1914a; Rousselot, 1953 Ploceus baglafecht Plasmodium nucleophilum: Manwell & Rossi, 1975 Haemoproteus fringillae/orizivorae: Bennett & Herman, Trypanosoma sp.: Ashford et al., 1976 1976 (now H. que/ea) Haemoproteus quelea: Bennett & Peirce, 1991b Euplectes macrourus Haemoproteus sp.: Ashford eta/., 1976; Gamham, 1950 Haemoproteus quelea: Bennett & Peirce, 1991b Leucocytozoon fringillinarum: Bennett & Herman, 1976 Haemoproteus sp.: Ashford et al., 1976 Leucocytozoon sp.: Ashford eta/., 1976; Gamham, 1950 Microfilariae: Ashford et al., 1976 Plasmodium relictum: Gamham, 1950 Plasmodium octamerium: Ashford et al., 1976 Plasmodium rouxi: Ashford eta/., 1976 Plasmodium rouxi: Ashford et al., 1976 Plasmodium vaughani: Bennett & Herman, 1976 Plasmodium vaughani: Ashford et al., 1976 Ploceus bojeri Euplectes orix Microfilaria sp.: Bennett eta/., 1977 Atoxoplasma sp.: Williams et al.; 1977 Haemoproteus columbae: Rousselot, 1943 (now H. quelea) Ploceus capensis Haemoproteus fringillae/orizivorae: Williams et al., 1977 (now H. quelea) Haemoproteus quelea: Bennett & Peirce, 1991b Haemoproteus quelea: Bennett & Peirce, 1991b Haemoproteus sp.: IRCAH Haemoproteus sp.: Ashford et al., 1976;' Commes, 1918b; Leucocytozoon majoris: IRCAH Cowper, 1969; Rousselot, 1953 Leucocytozoon sp.: IRCAH Microfilariae: Williams et al., 1977 Plasmodium relictum: Williams et al., 1977 Ploceus cucullatus Plasmodium rouxi: Ashford et al., 1976; Williams eta/., 1977 Atoxoplasma sp.: Peirce et al., 1977b Plasmodium sp.: Manwell & Rossi, 1975 Haemoproteus columbae: Rousselot, 1942, 1943 (now H. quelea) Euplectes progne Haemoproteus fringillae/orizivorae: Bennett et al., 1974, No parasites seen: IRCAH 1977, 1978; Peirce et al., 1977b Wink & Bennett, 1976 (now H. quelea) Foudia eminentissima Haemoproteus quelea: Bennett & Peirce, 1991b Haemoproteus sp.: Ashford et al., 1976; Leger & Leger, No parasites seen: Lowery, 1971 1914c; Peirce, 1984a; Rousselot, 1953; Schwetz, 1931, Foudia flavicans 1935c Leucocytozoon fringillinarum: Bennett et al., 1974; Peirce et Leucocytozoon fringillinarum: Peirce eta/., 1977a al., 1977b Leucocytozoon sp.: Peirce eta/., 1977a Leucocytozoon sp.: Ashford et al., 1976; Leger & Leger, Plasmodium sp.: Peirce eta/., 1977 a 1914a; Rousselot, 1953; Schwetz, 1931, 1938 Microfilariae: Ashford et al., 1976; Bennett et al. , 1977; Wil­ Foudia madagascariensis liams et al., 1977; Wink & Bennett, 1976 A toxoplasma sp.: Peirce eta/., 1977a Plasmodium nucleophilum: Ashford et al., 1976 Haemogregarina sp.: Peirce eta/., 1977a (probably Hepato­ Plasmodium relictum: Ashford et al., 1976; Bennett eta[., zoon sp.) 1978 Haemoproteus sp.: Maya & David, 1912 Plasmodium vaughani: Ashford et al., 1976; Bennett et al., Leucocytozoon fringillinarum: Bennett & Blancou, 1974; 1978; Peirce et al., 1977b Peirce, 1977; Peirce eta/., 1977a; Peirce, 1979 Trypanosoma avium: Bennett et a/., 1974, 1978; Wink & Leucocytozoon sp.: Maya & David, 1912 Bennett, 1976 Plasmodium sp.: Peirce, 1977; Peirce eta/., 1977a Trypanosoma sp. : Ashford et al., 1976; Leger & Leger, Trypanosoma sp.: Maya & David, 1912; Peirce eta/., 1977a 1914c; Peirce et al., 1977b; Peirce et al., 1977a; Williams et al., 1977 Foudia rubra Atoxoplasma sp.: Peirce eta/., 1977a Ploceus dorsomaculatus Leucocytozoonfringillinarum: Peirce eta/., 1977a No parasites seen· Peirce, 1969 Plasmodium sp.: Peirce eta/., 1977a Trypanosoma mayae: Peirce et al., 1977a .PIOC'f'US ~albula Haemoproteus sp.: Ashford eta/., 1976 Malimbus melanotia Plasmodium nucleophilum: Ashford et al. , 1976 Haemoproteus quelea: Bennett & Peirce, 1991b Plasmodium relictum: Ashford et al., 1976 Haemoproteus sp.: Oosthuizen & Markus, l967b L~uco~ytozoon sp.: Oosthuizen & Markus, 1967b Ploceusintermedius Mtcrofilanae: Oosthuizen & Markus 1967b Haemoproteus fringillae/orizivorae: IRCAH (now H. quelea) Trypanosoma sp.: Oosthuizen & Ma;kus, 1967b Haemoproteus orizivorae: Peirce et al., 1977b (now H. quelea) Malimbus rubriceps Haemoproteus quelea: Bennett & Peirce, 1991b Haemoproteus sp.: Ashford eta/., 1976 Haemoproteus sp.: Ashford et al., 1976; Peirce & Backhurst, Leucocytozoon sp.: Ashford et al., 1976 1970 Microfilariae: Ashford eta/., 1976 Leucocytozoon fringillinarum: Peirce et al, 1977b Leucocytozoon sp.: Peirce & Backhurst, 1970 Ploceus atrogularis Plasmodium (Novye/la) sp.: IRCAH Haemoproteus sp.: Wink & Bennett, 1976 Plasmodium nucleophilum: Ashford et al., 1976 Microfilariae: Wink & Bennett, 1976 Plasmodium sp.: Peirce et al., 1977b Ploceus aurantius Ploceus jacksoni Haemoproteus fringillae/orizivorae: Bennett eta/., 1974, 1977 Haemoproteus sp.: Garnham, 1950 (now H. quelea) Leucocytozoon sp.: Garnham, 1950 Haemoproteus quelea: Bennett & Peirce, 1991b Haemoproteus sp.: Peirce, 1969 Ploceus luteolus Leucocytozoon fringillinarum: Bennett eta/., 1974, 1977 Haemoproteus fringillae/orizivorae: Bennett et al., 1977, Microfilariae: Bennett eta/., 1977 1978 (now H. quelea) Plasmodium circumflexum: IRCAH Haemoproteus quelea: Bennett & Peirce, 1991b Plasmodium sp.: Peirce, 1969 Haemoproteus sp.: Ashford et al., 1976 Trypanosoma avium: Bennett eta/., 1974 Microfilariae: Ashford et al., 1976; Bennett et al., 1978 55 HOST-PARASITE CATALOGUE OF THE HAEMATOZOA OF THE SUB-SAHARAN BIRDS

Plasmodium sp.: Bennett eta/., 1978 Ploeeus velatus Trypanosomasp.: Ashford eta/., 1976 Haemoproteus quelea: Bennett & Peirce, 1991b Haemoproteus sp.: Ashford et al., 1976; Oosthuizen & Ploeeus melanoeephalus Markus, 1967b Leucocytozoon fringillinarum: IRCAH Haemoproteus fringillae/orizivorae: Bennett eta/. , 1974; Wil­ Leucocytozoon sp.: IRCAH liams eta/., 1977 (now H. quelea) Microfilariae: Ashford et al., 1976; Oosthuizen & Markus, Haemoprot~us quelea: Bennett & Peirce, 1991b 1967b Leucocytozoon bouffardi: Leger & Blanchard, 1911; Rousse­ Plasmodium relictum: IRCAH lot, 1953 (now L. fringillinarum) Plasmodium rouxi: Ashford et al., 1976 Leucocytozoon sp.: Gamham, 1950 Plasmodium sp.: Ashford eta/., 1976 Microfilariae: Leger, 1913; Leger & Blanchard, 1911; Wil- Trypanosoma sp.: IRCAH liams eta/., 1977 · Plasmodium cathemerium: Williams et al., 1977 Ploeeus vitellinus Plasmodium sp.-: Leger & Blanchard, 1911 Trypanosoma bouffardi: Leger & Blanchard, 1911 Haemoproteus fringillaelorizivorae: Bennett et al., 1978 (now H. quelea) Ploeeus nigerrimus Plasmodium polare: Bennett eta/., 1978 Plasmodium vaughani: Bennett eta/., 1978 Haemoproteus · fringillae/orizivorae: Bennett et a/. , 1974, Trypanosoma calmettei: Bennett et al. , 1978· 1977; Wink & Bennett, 1976 (now H. quelea) Haemoproteus quelea: IRCAH Ploeeus weynsi Leucocytozoon fringillinarum: Bennett eta/., 1977; Wink & Haemoproteus fringillae/orizivorae: Bennett et al., 1974, Bennett, 1976 1975 (now H. quelea) Leucocytozoon majoris: IRCAH Leucocytozoon fringillinarum: Bennett et al. , 1974, 1977 Microfilariae: IRCAH Microfilariae: Bennett eta/. , 1977 Plasmodium circumflexum: Crewe, 1974 Plasmodium vaughani: Bennett eta/., 1977 Trypanosoma avium: Bennett eta/., 1974, 1977 No parasites seen: Bennett & Herman, 1976; Oosthuizen & Markus, 1969b Ploeeus nigricollis Ploeeus xanthops Haemoproteus fringillaelorizivorae: Benett eta/., 1974, 1977 (now H. quelea) No parasites seen: Bennett & Herman, 1976; Oosthuizen & Haemoproteus quelea: Bennett & Peirce, 1991b Markus, 1969b Leucocytozoon fringillinarum: Bennett et al., 1977 Microfilariae: Wink & Bennett, 1976 Quelea erythrops Plasmodium nucleophilum: Bennett et al. , 1974 Haemoproteus fringillae/orizivorae: Bennett et a/., 1974, Trypanosoma sp.: Ashford et al., 1976 1977 (now H. quelea) Haemoproteus quelea: Bennett & Peirce, 1991b Ploeeus oeularis Haemoproteus sp.: Ashford eta/., 1976 · Leucocytozoon Jringillinarum: Bennett eta/., 1977 Haemoproteus fringillae/oriz"ivorae: Bennett & Herman, Leucocytozoon majoris: IRCAH 1976; Bennett eta/. , 1974, 1977 (now H. quelea) Leucocytozoon sp.: Ashford eta/., 1976 Haemoproteus quelea: Bennett & Peirce, 1991b Microfilariae: IRCAH Leucocytozoon ftingillinarum: Bennett & Herman, 1976 Plasmodium quelea: Marullaz, 1912b Leucocytozoon sp.: Ashford et al. , 1976 Plasmodium relictum: Ashford et at., 1976; Marullaz, 1912b Microfilariae: Ashford eta/., 1976 Plasmodium rouxi: Bennett & Herman, 1976 Plasmodium rouxi: Bennett & Herman, 1976 Plasmodium vaughani: Bennett et a/., 1977 Plasmodium vaughani: Ashford et al., 1976 Plasmodiumsp.: Ashford eta/., 1976 Plasmodium sp.: Manwell & Rossi, 1975 Trypanosoma sp.: Ashford et al., 1976 Trypanosoma avium: Bennett & Herman, 1976 Trypanosoma everetti: IRCAH Quelea quelea Trypanosoma sp.: Ashford eta/., 1976 Haemoproteus quelea: Bennett & Peirce, 1991b Leucocytozoon fringillinarum: Peirce eta/., 1977b Ploeeus pelzelni Plasmodium sp.: Peirce et al. , 1977b Haemoproteus fringillae/orizivorae: Bennett et al. , 1974 (now H. quelea) POEPHILINAE Haemoproteus quelea: Bennett & Peirce, 1991b (ESTRILDIDAE) Leucocytozoon sp.: Berson, 1964 Lonchura bicolor Ploeeus rubiginosus No parasites seen: Peirce, 1984i; Peirce et al., 1977b Haemoproteus fringillae/orizivorae: Peirce eta/., 1977b (now H. quelea) Lonchura cucullata Haemoproteus quelea: Bennett & Peirce, 1991b A toxoplasma sp.: Peirce, 1984a Leucocytozoon fringillinarum: Peirce eta/. , 1977b Haemoproteus orizivorae: Peirce, 1984a Leucocytozoon sp.: Peirce, 1984i Microfilariae: Peirce, 1984a Plasmodium sp.: Peirce et al., 1977b Trypanosoma bouffardi: Peirce, 1984a Trypanosoma sp.: Ashford eta/. , 1976; Peirce et al., 1977b PRIONOPIDAE Ploeeus spekei Euroeephalus anguitimens Leucocytozoon fringillinarum: Bennett & Herman, 1976 Haemoproteus sp.: Oosthuizen & Markus, 1967b Ploeeussubaureus Prionops plumata Haemoproteus passeris: Peirce, 1984i Hepatozoon sp.: IRCAH Leucocytozoon sp.: IRCAH Ploeeus taeniopterus Microfilariae: IRCAH Trypanosoma everetti: IRCAH Haemoproteus sp.: Ashford eta/. , 1976 Microfilariae: Ashford et al., 1976 Prionops retzii Trypanosoma sp.: Ashford et al., 1976 Hepatozoon sp.: IRCAH

56 G. F. BENNETT, R. A. EARLE, HESTERDUTOIT& F. W. HUCHZERMEYER

PROCELLARIDAE Hypsipetes borbonica Pterodroma arminjoniana No parasites seen: Peirce eta/., 1977a No parasites seen: Peirce, 1979; Peirce eta/., 1977a Hypsipetes crassirostris Puffin us diomedea Haemoproteus sanguinis: Peirce & Cheke, 1977; Rahal eta/. 1987 No parasites seen: IRCAH Trypanosoma everetti: Peirce & Chc>.ke, 1977 Puffmus pacificus Hypsipetes madagascariensis No parasites.seen: Peirce, 1979 Haemoproteus otoc_ompsae: Rahal eta/., 1987 Haemoproteus sanguinis: Rahal eta/., 1987 PROMEROPIDAE Nicator gularis Promerops cater No parasites seen: Bennett eta/. , 1974, 1977; Schwetz, 1944 No parasites seen: Bennett & de Swardt, 1989 Phyllastrephus albigularis Promerops gurneyi No parasites seen: Bennett eta/. , 1974, 1977 Leucocytozoon ane/lobiae: Bennett & de Swardt, 1989 Microfilariae: Bennett & de Swardt, 1989 Phyllastrephus cabanisi Plasmodium cathemerium: IRCAH No parasites seen: Peirce, 1984a Trypanosoma avium: Bennett & de Swartdt, 1989 Phyllastrephus cerviniventris

PSIITACIDAE No parasites seen: Peirce, 1984a Phyllastrephus fischeri Agapornis personata No parasites seen: IRCAH Haemoproteus sp.: Bennett & Herman, 1976 Trypanosoma everetti: Peirce, 1976b Agapornis pullaria Phyllastrephus strepitans No parasites seen: IRCAH Haemoproteus sp.: Ashford eta/. , 1976; Garnham, 1950 Agapornis roseicollis No parasites seen: IRCAH Phyllastrephus terrestris Haemoproteus otocompsae: IRCAH Agapornis taranta Leucocytozoon brimonti: IRCAH Microfilariae: Peirce, 1969 Pycnonotus barbatus Poicephalus meyeri Haemoproteus otocompsae: Peirce, 1984a, h; Rahal et a/., No parasites seen: Oosthuizen & Markus, 1969b 1987 Haemoproteus philippinensis: Rahal eta/., 1987 Poicephalus rufiventris Haemoproteus sanguinis: Peirce et a/., 1977b; Wink & No parasites seen: Peirce, 1984i Bennett, 1976; Rahal eta/. , 1987 Haemoproteus sp.: Ashford eta/. , 1976; Bennett & Herman, Psittacus erithacus 1976; Bennett eta/. , 1974; Cowper, 1969; Garnham, 1950; Haemoproteus sp.: Peirce & Bevan, 1977; Zupitza, 1909 Leger & Leger, 1914c; Rousselot, 1953 Trypanosoma avium: Zupitza, 1909 Leucocytozoon brimonti: Bennett & Herman, 1976; Peirce, Unidentified parasites: Peirce, 1969 eta/. , 1977b; Wink & Bennett, 1976 Leucocytozoon fringillinarum: Peirce, 1984a Leucocytozoon sp. : Ashford eta/., 1976; Bennett eta/., 1974; PTEROCLIDAE Leger & Leger, 1914a; Peirce, 1976b; Peirce & Backhurst, 1970; Rousselot, 1953 Pteroeles bicinctus Microfilariae: Ashford et a/., 1976; Bennett et a/., 1974; No parasites seen: Oosthuizen & Markus, 1969b Cowper, 1969; Oosthu,izen & Markus, 1967b; Peirce, 1984a; Peirce & Backhurst, 1970 Pteroeles burchelli Plasmodium circumflexum : Bennett eta/. , 1974 No parasites seen: IRCAH Plasmodium relictum: Ashford eta/. , 1976 Plasmodium rouxi: Ashford eta/., 1976; Bennett & Herman, Pteroeles lichtensteinii 1976; Peirce, 1984a No parasites seen: IRCAH Plasmodium sp.: Ashford et a/., 1976; Peirce & Backhurst, 1970 Pteroeles namaqua Trypanosoma avium: Wink & Bennett, 1976 No parasites seen: IRCAH Trypanosomapycnonoti: Kerandel, 1913; Peirce, 1984a Trypanosoma sp. : Ashford et a/., 1976; Kerandel, 1909; Peirce & Backhurst, 1970 PYCNONOTIDAE Pycnonotus cafer Andropadus importunus No parasites seen: IRCAH Leucocytozoon brimonti: Peirce eta/., 1977b Pycnonotus curvirostris Chloroeichla falkensteini Microfilariae: Bennett eta/. , 1977 Leucocytozoon sp.: Schwetz, 1938 Pycnonotus gracilirostris Microfilariae: Schwetz, 1938 Haemoproteus sp. Schwetz, 1938 Chloroeichla navicollis Pycnonotus gracilis Leucocytozoon sp.: Ashford eta/. , 1976 Trypanosoma sp.: Ashford eta/., 1976 Haemoproteus sp. : Schwetz, 1938 Chloroeichla naviventris Pycnonotus importunus Trypanosoma avium: Bennett eta/. , 1977 Haemoproteus sanguinis: Rahal eta/., 1987 Leucocytozoon brimonti: Bennett & Herman, 1976; Peirce, Criniger calurus 1977 No parasites seen: Bennett eta/. , 1977 Nuttallia sp.: Bennett & Herman, 1976 (now Babesia)

57 HOST-PARASITE CATALOGUE OF THE HAEMATOZOA OF THE SUB-SAHARAN BIRDS

Pycnonotus jocosus SALPORNI1HINAE Atoxoplasma sp.: Peirce etal., 1977a (SITTIDAE) Leucocytozoon sp.: Peirce et al. , 1977a Salpornis spilonotus Pycnonotus latirostris Leucocytozoon sp.: Peirce, 1984a No parasites seen: Bennett et al. , 1974 Microfilariae: Peirce, 1984a Pycnonotus nigricans Haemoproteus otocompsae: Rahal et al. , 1987 SCOLOPACIDAE Haemoproteus sanguinis: Rahal et al., 1987 Leucocytozoon brimonti: IRCAH Actitis hypoleucos Leucocytozoon sp.: IRCAH Plasmodiumsp.: IRCAH Microfilariae: IRCAH Trypanosoma sp.: IRCAH Arenaria interpres Pycnonotus tephrolaemus No parasites seen: Lowery, 1971 Leucocytozoon sp.: Cheke, 1972 Calidris minuta Pycnonotus tricolor No parasites seen: IRCAH Haemoproteussp. : Garnham, 1950 Leucocytozoon sp.: Schwetz, 1931, 1935c Calidris temminckii Trypanosoma pycnonoti: Kerandel, 1913 No parasites seen: Ashford et al. , 1976; Bennett & Herman, Trypanosoma sp.: Kerandel, 1909 1976; Bennett, 1979 Pycnonotus virens Gallinago gallinago Haemoproteus sanguinis: Wink & Bennett, 1976; Rahal et Haemoproteus sp.: Ashford et al., 1976 al. , 1987 Plasmodium sp.: Ashford et al., 1976 Haemoproteus sp.: Bennett et al., 1977 Leucocytozoon brimonti: Bennett et al., 1977 Gallinago macrodacatyla Microfilariae: Bennett et al., 1974, 1977 No parasites seen: Bennett & Blancou, 1974 Pycnonotus xanthopygos Gallinago nigripennis Haemoproteus otocompsae: Peirce, 1984i; Rahal et al., 1987 No parasites seen: Ashford et al., 1976; Bennett & Herman, Haemoproteus sanguinis: Rahal etal., 1987 1976; Peirce, 1984i Haemoproteus sp.: Bennett & Herman, 1976 Lankesterella sp.: Bennett & Herman, 1976 Philomachus pugnax Leucocytozoon brimonti: Bennett & Herman, 1976 Micro filariae: Bennett & Herman, 1976 No parasites seen: Peirce, 1984a, i Nuttallia sp.: Bennett & Herman, 1976 (now Babesia) Tringa glareola Plasmodium rouxi: Bennett & Herman, 1976 Trypanosoma avium: Bennett & Herman, 1976 Haemoproteus sp.: Ashford eta/., 1976 Tylas eduardi Tringa stagnatilis Leucocytozoon brimonti: IRCAH No parasites seen: Peirce, 1984i

RALLIDAE SCOPIDAE Dryolimnas cuvieri Scopus umbretta No parasites seen: Lowery, 1971 No parasites seen: IRCAH Fulica cristata SPHENISCIDAE No parasites seen: Bennett, 1980; Bennett & Herman, 1976 Limnocorax flavirostra Spheniscus demersus No parasites seen: IRCAH Aeygyptianella: Coles, 1941 (re-identified as Babesia) Babesia sp.: VRI Porphyrula alieni Plasmodium relictum: VRI Haemoproteus sp.: Ashford et al., 1976 Plasmodium sp.: VRI NOTE: All parasites referred to this species by Bennett eta/., Prophyrula madagascariensis (1982) represent zoo birds which acquired their infections in No parasites seen: IRCAH North America and Europe and thus do not represent African records) Sarothrura pulchra No parasites seen: Bennett, 1980; Bennett et al., 1974 STRIGIDAE Sarothrura rufa Asio capensis No parasites seen: IRCAH Haemoproteus syrnii: VRI Leucocytozoon ziemanni: VRI R.ECURV1ROSTRIDAE Plasmodium fa/lax: VRI Himantopus himantopus Asio flammeus No parasites seen: IRCAH Haemoproteus sp.: Kerandel, 1909; Leger & Leger, 1914c; Rousselot, 1953 Recurvirostra avosetta Leucocytozoon sp.: Ashford et a/., 1976; Kerandel, 1909, No parasites seen:Ashford et al., 1976; Peirce, 1976b 1913 Trypanosoma sp.: Kerandel, 1909, 1913

SAGITTARIIDAE Bubo africanus Sagittarius serpentarius Haemoproteus noctuae: Bennett & Herman, 1976 Haemoproteus syrnii: Bishop & Bennett, 1989 Leucocytozoon beaurepairei: Travassos Santos Dias, 1954 Haemoproteus sp.: Balfour, 1908b

58 G. F. BENNETI,R. A. EARLE, HESTERDUTOIT&F. W. HUCHZERMEYER

Leucocytozoon ziemanni: Bennett & Herman, 1976 Cinnyricinclus leucogaster Plasmodium sp.: IRCAH Haemoproteus sturni: Bennett & Herman, 1976 Trypanosoma avium: IRCAH Haemoproteus sp.: Oosthuizen & Markus, 1967b Unidentified parasite: IRCAH Microfilariae: Peirce, 1984a Bubo capensis Plasmodium circumflexum: Manwell & Rossi, 1975 Haemoproteus sp.: Rodhain et al., 1913a Cinnyricinclus sharpii Leucocytozoon sp.: Rodhain et al. , 1913a No parasites seen: IRCAH Trypanosoma sp.: Rodhain et al., 1913a Cosmopsarus regius Bubo lacteus Plasmodium nucleophilum: Manwell & Rossi, 1975 Haemoproteus celli: Peirce & Cooper, 1977b (now H. syrnii) Haemoproteus noctuae: Bishop & Bennett, 1989 Creatophora cinerea Haemoproteus syrnii: Bishop & Bennett, 1989 Haemoproteus sp.: Leger & Leger, 1914c; Rousselot, 1953 Haemoproteus morneti: Peirce et al., 1977b (syn of H . pasto- Leucycotyzoon sp.: Leger & Leger, 1914a; Rousselot, 1953 ris) Haemoproteus pastoris: IRCAH Bubo poensis Haemoproteus sp.: Ashford et al. , 1976 Microfilariae: Ashford etal., 1976 Haemoproteus syrnii: Bennett & Herman, 1976; Bishop & Plasmodium sp.: Ashford etal., 1976 Bennett, 1989 Trypanosoma sp.: Ashford et al., 1976 Glaucidium perlatum Lamprotomis australis Haemoproteus columbae: Rousselot, 1943 (now H. syrnii) Haemoproteus sp.: Oosthuizen & Markus, 1969b Haemoproteus noctuae: Bishop & Bennett, 1989 Haemoproteus sp.: Leger & Leger; 1914c; Rousselot, 1953 Lamprotomis caudatus Leucocytozoon ziemanni: IRCAH Microfilariae: Leger, 1913 Leucocytozoon sp.: Rousselot, 1953 Microfilariae: Leger, 1913 Lamprotornis chalybaeus Trypanosoma sp.: Leger & Leger, 1914c Haemoproteus sturni: Bennett & Herman, 1976; Bennett et Otus leucotis al., 1978; Williams et al., 1977 Haemoproteus sp.: Ashford et al., 1976; Oosthuizen & Haemoproteus noctuae: Bishop & Bennett, 1989 Markus, 1967b Haemoproteus syrnii: Bishop & Bennett, 1989 Leucocytozoon fringillinarum: Bennett & Herman, 1976 Haemoproteus sp.: Ashford et al., 1976 Leucocytozoon majoris: Bennett & Herman, 1976 Microfilariae: IRCAH Leucocytozoon sp.: Ashford et al. , 1976 Microfilariae: Ashford et al. , 1976; Oosthuizen & Markus, Otusscops 1967b Haemoproteus syrnii: IRCAH Plasmodium circumflexum: Manwell & Rossi, 1975 Haemoproteussp.: Peirce, 1984i Plasmodium nucleophilum: Manwell & Rossi, 1975 Trypanosoma avium: IRCAH Plasmodium octamerium: Manwell & Rossi, 1975 Trypanosoma sp.: Peirce, 1984i Plasmodium relictum: Ashford et al., 1976; Bennett & Herman, 1976 Scotopelia peli Plasmodium vaughani: Bennett eta[., 1978. Haemoproteus sp.: Rodhain et al., 1913a Leucocytozoon sp.: Rodhain et al., 1913a Lamprotomis chloropterus Haemoproteus morneti: Travassos Santos Dias, 1953 (syn of Strix woodfordii H. pastoris) Haemoproteus noctuae: Bishop & Bennett, 1989 Haemoproteus sturni: Bennett & Herman, 1976 Haemoproteus syrnii: Schwetz, 1935b; Bishop & Bennett, Haemoproteus sp.: Oosthuizen & Markus, 1967b, 1969b 1989 Trypanosoma sp.: Oosthuizen & Markus, 1967b Haemoproteus sp.: Bennett et al. , 1974; Bray, 1962; Schwetz, Lamprotomis corruscus 1931, 19.35b Leucocytozoon sp.: Schwetz, 1931, 1933, 1935c Haemoproteus morneti orienta/is: Travassos Santos Dias, Microfilariae: Schwetz, 1933 1953 (syn of H. pastoris) Plasmodiumfallax: Schwetz, 1931, 1935b Leycocytozoonfringillinarum: Bennett & Herman, 1976 Plasmodiumgundersi: Bray, 1962 Leucocytozoon majoris: Bennett & Herman, 1976 Plasmodium relictum: Schwetz, 1935b Leucocytozoon sp.: IRCAH Plasmodium sp.: Bennett et al., 1974; Schwetz, 1933 Trypanosoma sp.: Schwetz, 1931 Lamprotomis nitens Haemoproteus pastoris: IRCAH STRUTHIONIDAE Lamprotomis purpureus Struthio camelus Haemoproteus morneti: Tendeiro, 1947a, 1948a (syn of H. pastoris) Aegyptianella pullorum: Receveur & Thome, 1948 Haemoproteus sp.: Leger & Leger, 1914c Leucocytozoon struthionis: Walker, 1912, 1913 Microfilariae: Leger, 1913; Tendeiro, 1948b Plasmodium struthionis: Fantham & Porter, 1943 (species Trypanosoma abelsalazari: Tendeiro, 1947a, 1948a incertae sedis) Plasmodium sp.: IRCAH Lamprotomis purpuropterus No parasites seen: Ashford et al., 1976 STURNIDAE Onycbognathus fulgidus Acridotheres tristis Leucocytozoon sp.: Schwetz, 1944 Haemoproteus sp.: Maya & David, 1912 Onycbognathus morio Lankesterella sp.: IRCAH Leucocytozoon sp.: Maya & David, 1912 Haemoproteus pastoris: IRCAH Trypanosoma sp.: Maya & David, 1912 Onycbognatbus waHeri Bupbagus erytbrorbyncbus Haemoproteus sturni: Bennett & Herman, 1976 (syn of H. Leucocytozoon sp.: Peirce & Backhurst, 1970 pastoris) 59 HOST-PARASITE CATALOGUE OF THE HAEMATOZOA OF THE SUB-SAHARAN BIRDS

Spreo hildebrandti Camaroptera brevicaudata No parasites seen: Peirce, 1984i Haemoproteus fringillae/orizivorae: Bennett et al. , 1974, 1977 Spreo bicolor Haemoproteussp.: Ashfordetal., 1976 Plasmodium circumflexum: Bennett et al., 1974 Leucocytozoon sp.: IRCAH Plasmodium sp.: IRCAH Plasmodium vaughani: Ashford et al., 1976 Spreo fischeri Trypanosoma avium: Bennett et al. , 1977 No parasites seen: Peirce, 1984i Trypanosoma sp.: IRCAH Spreo superbus Camaroptera chloronata No parasites seen: Bennett et al. , 1977 Haemoproteus sp.: Ashford et al., 1976; Peirce & Backhurst, 1970 Camaroptera fasciolata Leucocytozoon dubreuili: Peirce et al., 1977b No parasites seen: Oosthuizen & Markus, 1967b Plasmodium nucleophilum: Manwell & Rossi, 1975 Camaroptera simplex SULIDAE Haemoproteus: sp.: IRCAH Leucocytozoon sp.: Peirce & Backhurst, 1970 Sula dactylatra Chloropeta natalensis Babesia sp.: Peirce & Freare, 1978 Haemoproteussp.: Ashford eta/., 1976 Sulasula Cisticola aridula No parasites seen: Lowery, 1971 No parasites seen: IRCAH Cisticola ayresii SYLVIIDAE No parasites seen: IRCAH Acroeephalus arundinaceus Cisticola brachyptera Leucocytozoon fringillinarum: Peirce, et al., 1977b Haemoproteus sp.: Ashford et al., 1976 Microfilariae: Ashford et al., 1976 Acroeephalus baeticatus Plasmodium relictum: Ashford et al., 1976 Haemoproteus sp.: Ashford et al. , 1976 Plasmodium sp.: IRCAH Haemoproteus wenyoni: IRCAH Trypanosoma sp.: Ashford etal., 1976 Acroeephalus gracilirostris Cisticola chiniana Haemoproteussp.: Ashford eta/., 1976 Leucocytozoon sp.: Peirce, 1984a Microfilariae: Ashford et al., 1976 Plasmodium polare: IRCAH Plasmodium vaughani: Ashford et al., 1976 Plasmodium relictum: IRCAH Trypanosoma sp.: Ashford et al., 1976 Plasmodium rouxi: Peirce et al., 1977b Plasmodium sp.: Ashford et al., 1976 Acroeephalus palustris Haemoproteus fringillae: Peirce et al., 1977b Cisticola erythrops Haemoproteus orizivorae: Peirce, 1984a; Peirce et al. , 1977b Haemoproteus wenyoni: Peirce, 1984a, e Haemoproteus sp.: IRCAH Haemoproteus sp.: Ashford et al., 1976 Leucocytozoon fringillinarum: Peirce et al., 1977b Hepatozoon sp.: IRCAH Microfilariae: Peirce et al., 1977b Leucocytozoon sp.: Ashford et al., 1976 Microfilariae: Ashford et al., 1976 Acroeephalus schoenobaenus Plasmodium circumflexum: Ashford et al., 1976 Haemoproteus fringillae: Bennett et al., 1974 Trypanosoma sp.: Ashford et al. , 1976 Haemoproteus orizivorae: Peirce, 1984a Cisticola fulvicapilla Haemoproteus sp.: Peirce, 1984i Leucocytyzoon sp.: Ashford etal., 1976; Peirce, 1984i No parasites seen: Peirce, 1984a Trypanosoma sp.: Peirce, 1984i Cisticola galactotes Acroeephalus scirpaceus Haemoproteus wenyoni: IRCAH Haemoproteus sp.: Ashford et al., 1976 Haemoproteus sp.: Peirce, 1984a Leucocytozoon sp.: Ashford et al. , 1976 Apalis flaYida Plasmodium rouxi: Ashford et al. , 1976 No parasites seen: IRCAH Cisticola juncidis No parasites seen: IRCAH Apalis moreaui No parasites seen: Bennett et al., 1977 Cisticola lais Trypanosoma avium: IRCAH Apalis rufogularis Cisticola latera/is No parasites seen: Bennett et al., 1977 No parasites seen: Wink & Bennett, 1976 Apalis thoracica Cisticola natalensis No parasites seen: IRCAH Haemoproteus sp.: Ashford et al. , 1976 Bradypterus baboecala Plasmodium cathemerium: Ashford et al., 1976 Plasmodium vaughani: Ashford et al., 1976 No parasites seen: Ashford et al. , 1976 Cisticola nigriloris Calamonestes simplex No parasites seen: Bennett et al., 1977 No parasites seen: Peirce, 1984i Cisticola robusta Camaroptera brachyura Leucocytozoon fringillinarium: Bennett & Herman, 1976 Plasmodium sp.: Peirce, 1984a Microfilariae: Bennett & Herman, 1976

60 G. F. BENNETT, R. A. EARLE,HESTERDUTOIT&F. W. HUCHZERMEYER

Cisticola subrulicapilla Phyllolais collybeta No parasites seen: IRCAH Haemoproteus sp.: IRCAH Leucocytozoon sp.: Ashford eta/., 1976 Cisticola textrix Trypanosoma sp.: Ashford et al., 1976 No parasites seen: IRCAH Phyllolais pulchella Citsticola tinniens Haemoproteus orizivorae: IRCAH Haemoproteus sp.: Ashford et al., 1976 No parasites seen: IRCAH Trypanosoma sp.: Ashford et al., 1976 Eminialepida Phylloscopus trochilus No parasites seen: Bennett et al., 1974 Haemoproteus fringillae: Bennett et al., 1974; Peirce, 1984a Haemoproteus orizivorae: Bennett et al. , 1974 Eremomela canescens Haemoproteus sp.: Ashford et al., 1976; Bennett & Hemian Haemoproteus sp.: Ashford et al., 1976 1976 Leucocytozoon sp.: Ashford et al., 1976 Eremomela gregalis Plasmodium relictum.' IRCAH No parasites seen: IRCAH Plasmodium sp.: B~nnett & Herman, 1976 Trypanosoma sp.: Ashford eta/., 1976 Eremomela icteropygialis Prinia naricans Haemoproteus sp.: Ashford eta[., 1976 Leucocytozoon sp.: IRCAH Eremomela scotops Trypanosoma everetti: IRCAH No parasites seen: Peirce, 1984a frinia inornata Eremomela usticollis Atoxopklsma sp.: IRCAH Microfilariae: Oosthuizen & Markus, 1967b Leucocytozoon fringillinarum: IRCAH Plasmodium rou.xi: IRCAH Hippolais icterina Plgsmodium sp.: IRCAH Haemoproteus sp.: Oosthuizen & Markus, 1967b; Smith & Prinialeucopogon Cox, 1972 Plasmodium sp.: Smith & Cox, 1972 No p~rasites seen: Bennett et al., 1974, 1977 Hippolais languida Prinia mifculosa Haemogregarina sp. Peirce et al., 1977b (probably Hepato- Leucocytozoon sp.: IRCAH zoon sp.) Trypanqsoma everetti: IRCAH Haemoproteus orizivorae: Peirce et al., 1977b Prinia sublfava Haemoproteus sp.: IRCAH Plasmodium sp.: Peirce et al., 1977b Atoxoplasma sp.: Peirce, 1984a Haemoproteus sp.: Ashford eta/., 1976 Hippolais olivetorum Leucocytozoon Jringillinarum: Peirce, 1984a Haemoproteus belopolskyi: Valkiunas, 1989 Leucocytozoon sp.: Ashford et al., 1976 Haemoproteus orizivorae: Peirce et al., 1977b (now H. Plasmodium relictum: Ashford eta/., 1976 belopolskyi) Plasmodium rou.xi: Ashford eta/., 1976; Peirce, 1984a Plasmodiumsp.: Peirce eta/., 1977b Trypanosoma sp.: Ashford et al., 1976 fflppolais pallida Sericornis perpicillatus Haemoproteus orizivorae: Peirce et al., 1977b Haemoproteus orizivorae: IRCAH Haemoproteus sp.: Ashford et al., 1976 Plasmodium sp.: IRCAH Leucocytozoon sp.: Ashford et al., 1976 Stenostira scita Hippolais polyglotta No parasites seen: IRCAH Haemoproteus wenyoni: IRCAH Sylria atricapilla Hylia prasima Atoxoplasma sp.: Peirce et al., 1977b Haemoproteus sp.: Bennett et al., 1974 Haemoproteus sp.: Ashford eta/., 1976; Smith & Cox, 1972 Leucocytozoon dubreuili: Peirce et al., 1977b Locustella fluriatillis Leucocytozoon fringillinarum: Peirce eta/., 1977b No parasites seen: Peirce, 1984a; Peirce et al., 1977b Leucocytozoon sp.: Ashford et al., 1976 Micro filariae: Ashford eta/., 1976 Locustellaluscinioides Trypanosoma avium: IRCAH Plasmodium relictum: Ashford eta/., 1976 Sylvia borin Orthotomus ruliceps Haemoproteus fringillae: Bennett et al., 1974, 1977; Peirce, 1984a No parasites seen: IRCAH Haemoproteus orizivorae: Bennett et al., 1974, 1977; Peirce Parisoma Jayardi et al., 1977b Haemoproteus sp.: Ashford et al., 1976; Smith & Cox, 1972 Hepatozoon sp.: IRCAH Leucocytozoon fringillinarum: Peirce et al., 1977b Parisoma lugens Leucocytozoon sp.: Ashford et al., 1976 Plasmodium rouxi: IRCAH No parasites seen: IRCAH Plasmodium vaughani: Bennett eta/., 1974 Plasmodium sp.: Smith & Cox, 1972 Parisoma subcaeruleum Haemoproteus sylvae: IRCAH Sylria communis , Haemoproteus wenyoni: IRCAH Aegyptianella sp.: Peirce et al., 1977b Hepatozoon sp.: IRCAH Haemogregarina sp.: Peirce, 1977 (probably Hepatozoon sp.) Leucocytozoon fringillinarum: IRCAH Haemoproteus fringillae: Peirce eta/., 1977b Leucocytozoon sp.: IRCAH Haemoproteus orizivorae: Peirce et al. , 1977b Trypanosoma sp.: IRCAH Haemoproteus sp.: Ashford eta/., 1976 Trypanosoma everetti: IRCAH Leucocytozoon fringillinarum: Peirce et al., 1977b 61 HOST-PARASITE CATALOGUE OF THE HAEMATOZOA OF TIIE SUB-SAHARAN BIRDS

Leucocytozoon sp.: IRCAH Turdoides rubiginosus Plasmodium (Novyella) sp.: IRCAH Atoxoplasma argya: Gamham, 1950 Plasmodium relictum: Ashford et al., 1976; Peirce et al., Haemoproteus lima/us: IRCAH 1977b Haemoproteus sp.: Ashford et al., 1976; Gamham, 1950; Plasmodium sp.: Ashford eta/., 1976 Peirce & Backhurst, 1970 Trypanosoma avium: IRCAH Leucocytozoon sp.: Gamham, 1950; Peirce & Backhurst, Sylvia curruca 1970 Microfilariae: Ashford eta/., 1976 Haemoproteus sp.: Ashford eta/., 1976 Plasmodium sp.: Peirce & Backhurst, 1970 Microfilariae: Ashford eta/., 1976 Trypanosoma sp.: Ashford eta/., 1976 Plasmodium sp.: Ashford eta/., 1976

Sylvia nisoria TROGONIDAE Haemoproteus sp.: Peirce, 1976b Trypanosoma sp.: Ashford eta/., 1976 Apaloderma narina No parasites seen: Peirce, 1984i Sylvietta bracbyura Haemoproteus sp.: Ashford eta/., 1976 Leucocytozoon sp.: Ashford el al., 1976 TuRDIDAE Plasmodium sp.: Ashford et al. , 1976 Alethe diademata Trypanosoma sp.: Ashford et al. , 1976 No parasites seen: IRCAH Sylvietta rufescens Cercomela familiaris Haemoproteus sp.: Oosthuizen & Markus, 1967b Leucocytozoon sp.: Oosthuizen & Markus, 1967b Leucocytozoon sp.: Peirce, 1984a Trypanosoma everetti: IRCAH Plasmodium rou.xi: Peirce, 1984a Plasmodium vauglumi: IRCAH Sylvietta ruficapilla Trypanosoma sp.: Ashford et al., 1976 No parasites seen: Peirce, 1984a Cercomela sinuata Sylvietta virens No parasites seen: IRCAH Plasmodium (Novyella) sp.: IRCAH Cichladusa guttata Sylvietta whytil No parasites seen: Peirce, 1984i Haemoproteus sp.: Ashford eta/., 1976 Leucocytozoon sp.: Ashford eta/., 1976 Cossypha caffra Haemoproteussp.: IRCAH THRESKIORNITillDAE Microfilariae: IRCAH Plasmodium matulinum: IRCAH Bostrychia carunculata Trypanosoma avium: IRCAH No parasites seen: Ashford et al., 1976 Cossypha cyanocampter Bostrychia hagedash Microfilariae: Peirce, 1984i Haemoproteus pelouroi: Peirce, 1984i Cossypha dichroa Geronticus calvus No parasites seen: IRCAH No parasites seen: IRCAH Cossypha heuglini Platalea alba Microfilariae: Ashford eta/., 1976 Plasmodium sp.: IRCAH Cossypha humeralis Threskiornis aethiopicus No parasites seen: IRCAH Haemoproteus pelouroi: Bennett et al., 1975 Cossypha natalensis Haemoproteus sp.: Archetti, 1947; Minchin, 1910; Thomas & Dobson, 1975 Haemoproteus fallisi: Bennett eta/., 1977 Leucocytozoon dubreuili: IRCAH Microfilariae: Peirce, 1984a TIMALIIDAE Cossypha niveicapilla Akippe abyssinica No parasites seen: Ashford eta/., 1976; Bennet et al., 1974, No parasites seen: Ashford et al. , 1976 1977 madopsis albipectus Cossypha semirufa No parasites seen: Peirce, 1984i Haemoproteussp.: Ashford eta/., 1976 Leucocytozoon sp.: Ashford et al., 1976 Tricbastoma fulvescens Plasmodium rou.xi: Ashford eta/., 1976 Microfilariae: IRCAH Trypanosoma sp.: Ashford et al., 1976 Plasmodium rou.xi: IRCAH Erithacus luscinia Trichastoma rufipennis Haemoproteus fallisi: Peirce, 1977 No parasites seen: Cheke, 1972 Haemoproteus fringillae: Peirce, 1977 Haemoproteus sp.: Ashford eta/., 1976; Peirce, 1977 Turdoides jardineli Leucocytozoon fringillinarum: Peirce, 1977 Haemoproteus lima/us: IRCAH Plasmodium sp.: Peirce, 1976b; Peirce et al. , 1977b Leucocytozoon sp.: IRCAH Erithacus megarhynchos Turdoides leucopygius Haemoproteus sp.: Ashford et al., 1976 Haemoproteus sp.: Ashford eta/., 1976 Microfilariae: Ashford eta/., 1976 Leucocytozoon sp.: Ashford et al., 1976 Plasmodium relictum: Ashford eta/., 1976 Erythropygia coryphaeus Trypanosoma sp.: Ashford eta/., 1976 No parasites seen: IRCAH

62 G. F. BENNETI,R. A. EARLE,HESTERDUTOIT&F. W. HUCHZERMEYER

Erythropygia hartlaubi Saxicola rubetra No parasites seen: Bennett et al., 1977 Haemoproteus sp.: Smith & Cox, 1972 Erythropygia leucophrys Saxicola tectes Haemoproteus fallisi: IRCAH No parasites seen: Peirce et al., 1977a Haemoproteus orizivorae: Peirce et al., 1977b Leucocytozoon dubreuili: IRCAH Saxicola torquats Haemoproteus fallisi: Bennett & Herman, 1976 Erythropygia paena PU:lsmodium sp.: Bennett & Blancou, 1974 Haemoproteus fallisi: IRCAH Thamnolaea cinnamomeiventris Irania gutturalis Haemoproteus sp.: Rousselot, 1953 Microfilariae: Peirce et al., 1977b Thamnolaea semirufa Luscinia luscinia Haemoproteus sp.: Ashford et al. , 1976 Haemoproteus fallisi: Peirce et al., 1977b Leucocytozoon sp.: Ashford eta/. , 1976 Haemoproteus fringillae: Peirce et al., 1977b Plasmodium sp.: Peirce et al., 1977b Turdus abyssinicus Leucocytozoon sp.: Peirce & Backhurst, 1970 Luscinia megarhynchas Microfilariae: Peirce & Backhurst, 1970 No parasites seen: Peirce eta/., 1977b Plasmodium sp.: Peirce & Backhurst, 1970 Monticola angolensis Turdus albicoms Haemoproteus balmorali: Peirce, 1984b No parasites seen: IRCAH Haemoproteus fallisi: IRCAH Haemoproteus sp.: Peirce, 1984a Turdus bewsheri Leucocytozoon tringillinarum: Peirce, 1984a Microfilariae: Peirce & Cheke, 1977 Leucocytozoon sp.: IRCAH Microfilariae: IRCAH Turdus iliacus Plasmodium relictum: Peirce, 1984a Leucocytozoon giovannoU:li: Travassos Santos Dias, 1954 Plasmodium sp.: IRCAH (syn of L. dubreuili) Trypanosoma everetti: Peirce, 1984a Trypanosoma sp.: IRCAH Turdus libonyana Unknown parasite: IRCAH AtoxopU:lsma sp.: Peirce, 1984a Hepatozoon sp.: IRCAH Monticola brevipes Leucocytozoon dubreuili: Peirce, 1984a No parasites seen: IRCAH Microfilariae: Schwetz, 1938 Plasmodium rouxi: Peirce, 1984a Monticola rufoeinereus: Plasmodium sp.: IRCAH Haemoproteus sp.: Ashford et al., 1976 Trypanosoma everetti: Peirce, 1984a Trypanosoma sp.: IRCAH Monticola rupestris Unknown parasite: IRCAH No parasites seen: IRCAH Turdus olivaceus Monticola saxatilis Haemoproteus fallisi: IRCAH Haemoproteus faillisi: Bennett & Herman, 1976 Leucocytozoon dubreuili: Peirce, 1984a Plasmodium sp.: Smith & Cox, 1972 Microfilariae: Wink & Bennett, 1976 Plasmodium sp.: IRCAH Myrmecocichla arnotti Plasmodium vaughani: Bennett et al., 1974 No parasites seen: Peirce, 1984i Trypanosoma sp.: IRCAH Trypanosoma avium: Bennett & Herman, 1976 Myrmecocichla aethiops Turdus pallidus Leucocytozoon dubreuili: Bennett & Herman, 1976 Microfilariae: IRCAH Myrmecocichla formicivora Plasmodium sp.: IRCAH No parasites seen: IRCAH Turdus pelios Myrmecocichla nigra Leucocytozoon sp.: Ashford et al., 1976 Microfilariae: Ashford et al. , 1976 No parasites seen: Bennett et al., 1977 Plasmodium relictum: Ashford et al., 1976 Neoeossyphus rufus Plasmodium rouxi: Ashford et al., 1976 Plasmodium vaughani: Ashford et al., 1976 No parasites seen: Bennett et al. , 1977 Plasmodium sp.: Ashford eta/., 1976 Oenanthe bifasciats Trypanosoma sp.: Ashford et al. , 1976 No parasites seen: IRCAH Zoothera oberlaenderi Microfilariae: Bennett & Herman, 1976 Oenanthe bottae No parasites seen: Ashford et al. , 1976 TuRNICIDAE Oenanthe deserti Turnix hottentotta No parasites seen: IRCAH No parasites seen: IRCAH Oenanthe monticola Turnix sylvatica No parasites seen: IRCAH No parasites seen: IRCAH Oenanthe pHeats No parasites seen: IRCAH TYTONIDAE Pboenlcurus phoenicurus Tytoalba Haemoproteus sp.: Ashford et al., 1976 Haemoproteus tytoni: Bishop & Bennett, 1989 Plasmodium sp.: Smith & Cox, 1972 Leucocytozoon ziemanni: IRCAH

63 HOST-PARASITECATALOGUEOFTHEHAEMATOZOAOFTHESUB-SAHARANBIRDS

UPUPIDAE Zosterops pallidus Haemoproteus killangoi: IRCAH Upupaepops Haemoproteus zosteropis: IRCAH Leucocytozoon sp.: IRCAH Hepatozoon sp.: IRCAH Microfilariae: Ashford et al., 1976 Leucocytozoon zosteropis: IRCAH Plasmodium garnhami: Ashford et al., 1976 Leucocytozoon sp.: IRCAH Microfilariae: IRCAH Plasmodium (Novyella) sp.: IRCAH ZOSTEROPIDAE Trypanosoma avium: IRCAH Trypanosoma everetti: IRCAH Zosterops abyssinica Trypanosoma sp.: IRCAH Haemoproteus killangoi: Bennett & peirce, 1981 HaemoproteUs zosteropis: Peirce et al., 1977b Zosterops senegalensis Haemoproteus sp. : Ashford eta/., 1976 Haemoproteus killangoi: Bennett & Peirce, 1981 Trypanosoma sp.: Ashford et al., 1976 Haemoproteus zosteropis: Bennett & Pierce, 1981; Cheke, 1978; Peirce, 1984a Zosterops borbonica Haemoproteus sp.: Ashford et al., 1976; Bennett et al. , 1974 Atoxoplasma sp.: Peirce et al. , 1977a 1977; Cowper, 1969 Leucocytozoon zosteropis: Peirce, 1979; Peirce et al., 1977a Lankesterella sp.: Ashford et al. , 1976 Plasmodium relictum: Peirce et al., 1977a Leucocytozoon sp.: Bennett et al. , 1974; Peirce, 1984a Plasmodiumsp.: Peirce eta/., 1977a Microfilariae: Ashford et al., 1976; Bennett et al., 1974 Plasmodium relictum: IRCAH Zosterops maderaspatana Plasmodium vaughani: IRCAH Plasmodium sp.: Peirce, 1984a Haemoproteus killangoi: IRCAH Trypanosoma avium: Bennett et al. , 1974 Haemoproteus sp.: Lowery, 1971 Trypanosoma calmettei: IRCAH Plasmodium sp.: Lowery, 1971 Trypanosoma sp.: Ashford et al., 1976; Peirce, 1984a Unknown parasite: IRCAH Zosterops oli-vacea Atoxoplasma sp.: Peirce et al., 1977a Zosterops -virens Leucocytozoon zosteropis: Peirce, 1979; Peirce eta/., 1977a Haemoproteus zosteropis: Bennett & Herman, 1976 Trypanosoma sp.: Peirce, 1979; Peirce et al. , 1977a Plasmodjum relictum: IRCAH

ACKNOWLEDGEMENTS The production of a host-parasite catalogue of this nature requires the assistance of many contributors. It is a pleasure to acknowledge the major contribution of avian blood smears by Dawid de Swardt of the National Museum in Bloemfontein and Kotie Herholdt from the Kalahari Gemsbok National Park as well as Tony Harris, Tamar Casidy and Walter Neser of the Transvaal Museum. Mrs Margaret Harris of the International Reference Centre for Avian Haematozoa spent long hours in abstracting records from the literature. Professor Banie Penzhorn of the Department of Parasitology, Faculty of Veterinary Science of the Univer­ sity of Pretoria was instrumental in obtaining funding from the Foundation for Research Development to bring the first author to South Africa as a FRO Visiting Fellow, during which time this host-parasite catalogue was essentially completed. Dr D. W. Verwoerd, Director, Veterinary Research Institute, and Dr L. Prozesky, Assistant Director and Head, Patholqgy Section, kindly provided office and laboratory facilities to the first author during his three-month stay in Pretoria. An operating grant from the Natural Sciences and Engineering Research Council of Canada to the first author supported the initial phases of this endeavour. Mrs Mara Stoltz is especially thanked for her hours of labour in translating the rough draft of the manuscript into its final, pohshed appearance.

64