Northeast Partners in Amphibian and Reptile Conservation: Northeast Species List

Species Common Name Connecticut Delaware District of Columbia Maine Maryland Massachusetts New Hampshire New Jersey New York Pennsylvania Island Rhode Vermont Virginia West Virginia AMPHIBIA ANURANS FROGS Bufonidae True Toads Anaxyrus americanus American Toad NNNNNNNNNNNNNN Anaxyrus americanus americanus Eastern American Toad NNNNNNNNNNNNNN Anaxyrus fowleri Fowler's Toad N N N NNNNNNNNNN Anaxyrus quericus Oak Toad N Anaxyrus terrestris Southern Toad N

Hylidae Treefrogs Acris blanchardi Blanchard's Cricket Frog X Acris crepitans Eastern Cricket Frog N ? N N N N N N Acris gryllus Southern Cricket Frog ? N Acris gryllus gryllus Coastal Plain Cricket Frog ? N Hyla andersonii Pine Barrens Treefrog N Hyla chrysoscelis Cope’s Gray Treefrog N N N N N N Hyla cinerea Green Treefrog N X N I N Hyla femoralis Pine Woods Treefrog N Hyla gratiosa Barking Treefrog N N N Hyla squirella Squirrel Treefrog N Hyla versicolor Gray Treefrog NNNNNNNNNNNNNN Pseudacris brachyphona Mountain Chorus Frog N N N N Pseudacris brimleyi Brimley’s Chorus Frog N Pseudacris crucifer Spring Peeper NNNNNNNNNNNNNN Pseudacris feriarum Upland Chorus Frog ? N N N N Pseudacris kalmi New Jersey Chorus Frog N N N N N Pseudacris maculata Boreal Chorus Frog N X Pseudacris nigrita Southern Chorus frog N Pseudacris ocularis Little Grass Frog N Pseudacris triseriata Western Chorus Frog N N

Pelobatidae Spadefoot Toads

NEPARC publication 2013-01 Northeast Partners in Amphibian and Reptile Conservation: Northeast Species List

Species Common Name Connecticut Delaware District of Columbia Maine Maryland Massachusetts New Hampshire New Jersey New York Pennsylvania Island Rhode Vermont Virginia West Virginia Scaphiopus holbrookii Eastern Spadefoot N N X N N NNNN NN

Ranidae True Frogs Lithobates catesbeianus American Bullfrog NNNNNNNNNNNNNN Lithobates clamitans Green Frog NNNNNNNNNNNNNN Lithobates palustris Pickerel Frog NNNNNNNNNNNNNN Lithobates pipiens Northern Leopard Frog N N I N N X NNNN N Lithobates septentrionalis Mink Frog N N N N Lithobates sphenocephalus Southern Leopard Frog N N N N N N N Lithobates sphenocephalus utricularius Southern Leopard Frog N N N N N N N Lithobates sylvaticus Wood Frog NNNNNNNNNNNNNN Lithobates virgatipes Carpenter Frog N N N N

Microhylidae Narrow-mouthed Frogs Gastrophryne carolinensis Eastern Narrow-mouthed Toad N N

CAUDATA SALAMANDERS Ambystomidae Mole Salamanders Ambystoma barbouri Streamside Salamander N Ambystoma jeffersonianum Jefferson Salamander N X NNNNNN NNN Ambystoma laterale Blue-spotted Salamander N N NNNN N Ambystoma mabeei Mabee's Salamander N Ambystoma maculatum Spotted Salamander NNNNNNNNNNNNNN Ambystoma opacum Marbled Salamander N N N NNNNNNN NN Ambystoma talpoideum Mole Salamander N Ambystoma texanum Small-mouthed Salamander N Ambystoma tigrinum Eastern Tiger Salamander N ? N N N X N

Plethonontidae Lungless Salamanders Aneides aeneus Green Salamander N N N N Desmognathus auriculatus Southern Dusky Salamander N Desmognathus fuscus Northern Dusky Salamander NNNNNNNNNNNNNN

NEPARC publication 2013-01 Northeast Partners in Amphibian and Reptile Conservation: Northeast Species List

Species Common Name Connecticut Delaware District of Columbia Maine Maryland Massachusetts New Hampshire New Jersey New York Pennsylvania Island Rhode Vermont Virginia West Virginia Desmognathus marmoratus Shovel-nosed Salamander N Desmognathus monticola Seal Salamander N N N N Desmognathus ochrophaeus Allegheny Mountain Dusky Salamander N N N N N N Desmognathus orestes Blue Ridge Dusky Salamander N Desmognathus organi Northern Pygmy Salamander N Desmognathus planiceps Flat-headed Salamander N Desmognathus quadramaculatus Black-bellied Salamander N N Desmognathus welteri Black Mountain Salamander N N Eurycea bislineata Northern Two-lined Salamander NNNNNNNNNNNNNN Eurycea cirrigera Southern Two-lined Salamander N N Eurycea guttolineata Three-lined Salamander N Eurycea longicauda Long-tailed salamander N X N N N N N N Eurycea longicauda longicauda Eastern Long-tailed salamander N X N N N N N N Eurycea lucifuga Cave Salamander NN Eurycea wilderae Blue Ridge Two-lined Salamander N Gyrinophilus porphyriticus Spring Salamander N NNNNNNNNNNN Gyrinophilus porphyriticus duryi Kentucky Spring Salamander N N Gyrinophilus porphyriticus porphyriticus Northern Spring Salamander N NNNNNNNNNNN Gyrinophilus subterraneus West Virginia Spring Salamander D Hemidactylium scutatum Four-toed Salamander N N X NNNNNNNNNNN Plethodon chlorobryonis Atlantic Coast Slimy Salamander N Plethodon cinereus Eastern Red-backed Salamander NNNNNNNNNNNNNN Plethodon cylindraceus White-spotted Slimy Salamander N N Plethodon electromorphus Northern Ravine Salamander N N Plethodon glutinosus Northern Slimy Salamander N X N NNNN NN Plethodon hoffmani Valley and Ridge Salamander N N N N Plethodon hubrichti Peaks of Otter Salamander D Plethodon kentucki Cumberland Plateau Salamander N N Plethodon montanus Northern Gray-cheeked Salamander N Plethodon nettingi Cheat Mountain Salamander D Plethodon punctatus Cow Knob Salamander N N Plethodon richmondi Southern Ravine Salamander N N

NEPARC publication 2013-01 Northeast Partners in Amphibian and Reptile Conservation: Northeast Species List

Species Common Name Connecticut Delaware District of Columbia Maine Maryland Massachusetts New Hampshire New Jersey New York Pennsylvania Island Rhode Vermont Virginia West Virginia Plethodon shenandoah Shenandoah Salamander D Plethodon sherando Big Levels Salamander D Plethodon ventralis Southern Zigzag Salamander N Plethodon virginia Shenandoah Mountain Salamander N N Plethodon wehrlei Wehrle's Salamander N N N N N Plethodon welleri Weller's Salamander N Plethodon yonahlossee Yonahlossee Salamander N Pseudotriton montanus Mud Salamander N N N N N N N Pseudotriton montanus montanus Eastern Mud Salamander N N N N N N Pseudotriton montanus diastictus Midland Mud Salamander N N Pseudotriton ruber Red Salamander N N N N N N N N Pseudotriton ruber ruber Northern Red Salamander N N N N N N N N Pseudotriton ruber nitidus Blue Ridge Red Salamander N Stereochilus marginatus Many-lined Salamander N

Cryptobranchidae Giant Salamander Cryptobranchus alleganiensis Hellbender N N N N N Cryptobranchus alleganiensis alleganiensis Eastern Hellbender N N N N N

Sirenidae Sirens Siren intermedia Lesser Siren N Siren intermedia Intermedia Eastern Lesser Siren N Siren lacertina Greater Siren X N

Amphiumidae Congo Eels Amphiuma means Two-toed Amphiuma N

Proteidae Waterdogs Necturus maculosus Mudpuppy N I X I I NNNNNN Necturus maculosus maculosus Common Mudpuppy N I X I I NNNNNN Necturus punctatus Dwarf Waterdog N

NEPARC publication 2013-01 Northeast Partners in Amphibian and Reptile Conservation: Northeast Species List

Species Common Name Connecticut Delaware District of Columbia Maine Maryland Massachusetts New Hampshire New Jersey New York Pennsylvania Island Rhode Vermont Virginia West Virginia Salamandridae True Salamanders Notophthalmus viridescens Eastern Newt NNNNNNNNNNNNNN Notophthalmus viridescens viridescens Red-spotted Newt NNNNNNNNNNNNNN REPTILIA TESTUDINES TURTLES Chelonidae Sea Turtles Caretta caretta Loggerhead Sea Turtle N N NNNNNN N N Chelonia mydas Green Sea Turtle N N N N N N N N Eretmochelys imbricata Hawksbill Sea Turtle N N N N N N N Eretmochelys imbricata imbricata Atlantic Hawksbill Sea Turtle N N N N N N N Lepidochelys kempii Kemp’s Ridley Sea Turtle N N N N N N N N N

Dermochelyidae Leatherback Sea Turtle Dermochelys coriacea Leatherback Sea Turtle N N N N N N N N N

Chelydridae Snapping Turtles Chelydra serpentina Snapping Turtle NNNNNNNNNNNNNN

Emydidae Pond Turtles Chrysemys picta Painted Turtle NNNNNNNNNNNNNN Chrysemys picta picta Eastern Painted Turtle NNNNNNNNNNNNNN Chrysemys picta marginata Midland Painted Turtle N N N N N N N Clemmys guttata Spotted Turtle NNNNNNNNNNNNNN Deirochelys reticularia Chicken Turtle N Deirochelys reticularia reticularia Eastern Chicken Turtle N Emydoidea blandingii Blanding’s Turtle N N N N N Glyptemys insculpta Wood Turtle N X NNNNNNNNNNN Glyptemys muhlenbergii Bog Turtle N N X N N N N N N Graptemys geographica Northern Map Turtle N I N N N N N Graptemys ouachitensis Southern Map Turtle N Graptemys ouachitensis ouachitensis Ouachita Map Turtle N Graptemys pseudogeographica False Map Turtle I

NEPARC publication 2013-01 Northeast Partners in Amphibian and Reptile Conservation: Northeast Species List

Species Common Name Connecticut Delaware District of Columbia Maine Maryland Massachusetts New Hampshire New Jersey New York Pennsylvania Island Rhode Vermont Virginia West Virginia Graptemys pseudogeographica kohnii Mississippi Map Turtle I Malaclemys terrapin Diamond-backed Terrapin N N X N N N N N N Malaclemys terrapin terrapin Northern Diamond-backed Terrapin N N X N N N N N N Pseudemys concinna River Cooter X N N N Pseudemys concinna concinna Eastern River Cooter X N N N Pseudemys concinna floridana Coastal Plain Cooter N Pseudemys rubriventris Northern Red-bellied Cooter N N N N N I N N N Terrapene carolina Eastern Box Turtle NNNNNNNNNNN?NN Terrapene carolina carolina Woodland Box Turtle NNNNNNNNNNN?NN Trachemys scripta Pond Slider I IIII IIII NN Trachemys scripta scripta Yellow-bellied Slider I I N Trachemys scripta elegans Red-eared Slider I IIII IIII IN Trachemys scripta troosti Cumberland Slider N

Kinosternidae Mud and Musk Turtles Kinosternon subrubrum Eastern Mud Turtle N N N N N N N Kinosternon subrubrum subrubrum Southeastern Mud Turtle N N N N N N N Kinosternon bauri Striped Mud Turtle N Sternotherus minor Loggerhead Musk Turtle N Sternotherus minor peltifer Stripe-necked Musk Turtle N Sternotherus odoratus Eastern Musk Turtle NNNNNNNNNNNNNN

Trionchidae Softshell Turtles Apalone mutica Smooth Softshell N N Apalone mutica mutica Midland Smooth Softshell N N Apalone spinifera Spiny Softshell N I N N N N N Apalone spinifera spinifera Eastern Spiny Softshell N I N N N N N Pelodiscus sinensis Chinese Softshell I

SQUAMATA Lizards / Snakes Sauria Lizards Anguidae Anguid Lizards

NEPARC publication 2013-01 Northeast Partners in Amphibian and Reptile Conservation: Northeast Species List

Species Common Name Connecticut Delaware District of Columbia Maine Maryland Massachusetts New Hampshire New Jersey New York Pennsylvania Island Rhode Vermont Virginia West Virginia Ophisaurus attenuatus Slender Glass Lizard N Ophisaurus attenuatus longicaudus Eastern Slender Glass Lizard N Ophisaurus ventralis Eastern Glass Lizard N

Gekkonidae Geckos Hemidactylus turcicus Mediterranean Gecko I I

Lacertidae Wall Lizards Podarcis sicula Italian Wall Lizard I I

Phrynosomatidae Sceloporine Lizards Sceloporus undulatus Eastern Fence Lizard N N N N N N N N

Scinidae Skinks Plestiodon anthracinus Coal Skink N N N N N Plestiodon anthracinus anthracinus Northern Coal Skink N N N N N Plestiodon fasciatus Common Five-lined Skink N N N N X N N N N N N Plestiodon inexpectatus Southeastern Five-lined Skink N Plestiodon laticeps Broad-headed Skink N N N N N N Scincella lateralis Little Brown Skink N N N N N N

Teiidae Whiptails Aspidocelis sexlineata Six-lined Racerunner X N N N Aspidocelis sexlineata sexlineata Eastern Six-lined Racerunner X N N N

SQUAMATA SNAKES Coluberidae Colubrids Carphophis amoenus Common Wormsnake N N N N N NNNN NN Carphophis amoenus amoenus Eastern Wormsnake N N N N N NNNN NN Cemophora coccinea Scarletsnake N ? N N N Cemophora coccinea copei Northern Scarletsnake N ? N N N Clonophis kirtlandii Kirtland's Snake N

NEPARC publication 2013-01 Northeast Partners in Amphibian and Reptile Conservation: Northeast Species List

Species Common Name Connecticut Delaware District of Columbia Maine Maryland Massachusetts New Hampshire New Jersey New York Pennsylvania Island Rhode Vermont Virginia West Virginia Coluber constrictor North American Racer NNNNNNNNNNNNNN Coluber constrictor constrictor Northern Black Racer NNNNNNNNNNNNNN Diadophis punctatus Ring-necked Snake NNNNNNNNNNNNNN Diadophis punctatus edwardsii Northern Ring-necked Snake NNNNNNNNNNNNNN Diadophis punctatus punctatus Southern Ring-necked Snake N N N N Farancia abacura Red-bellied Mudsnake N Farancia abacura abacura Eastern Mudsnake N Farancia erytrogramma Rainbow Snake N N Farancia erytrogramma erytrogramma Common Rainbow Snake N N Heterodon platirhinos Eastern Hog-nosed Snake N N X NNNNNNN?NN Lampropeltis calligaster Yellow-bellied Kingsnake X N N Lampropeltis calligaster rhombomaculata Mole Kingsnake X N N Lampropeltis getula Eastern Kingsnake N ? N N N N N Lampropeltis nigra Eastern Black Kingsnake N N Lampropeltis triangulum Milksnake N N X NNNNNNNNNNN Lampropeltis triangulum triangulum Eastern Milksnake N N X NNNNNNNNNNN Nerodia erythrogaster Plain-bellied Watersnake N N N Nerodia sipedon Northern Watersnake NNNNNNNNNNNNNN Nerodia sipedon sipedon Northern Watersnake NNNNNNNNNNNNNN Nerodia taxispilota Brown Watersnake N Opheodrys aestivus Rough Greensnake N N N N N N N Opheodrys aestivus aestivus Northern Rough Greensnake N N N N N N N Opheodrys vernalis Smooth Greensnake N X NNNNNNNNNNN Pantherophis alleganiensis Eastern Ratsnake N N N N N NNNNNNN Pantherophis guttatus Red Cornsnake N X N N N N Pantherophis spiloides Gray Ratsnake ????? Pituophis melanoleucus Pinesnake X N N N Pituophis melanoleucus melanoleucus Northern Pinesnake X N N N Regina rigida Glossy Crayfish Snake N Regina rigida rigida Glossy Crayfish Snake N Regina septemvittata Queensnake N N N N N N N N Storeria dekayi Dekay's Brownsnake NNNNNNNNNNNNNN

NEPARC publication 2013-01 Northeast Partners in Amphibian and Reptile Conservation: Northeast Species List

Species Common Name Connecticut Delaware District of Columbia Maine Maryland Massachusetts New Hampshire New Jersey New York Pennsylvania Island Rhode Vermont Virginia West Virginia Storeria dekayi dekayi Northern Brownsnake NNNNNNNNNNNNNN Storeria occipitomaculata Red-bellied Snake N N ? NNNNNNNNNNN Storeria occipitomaculata occipitomaculata Northern Red-bellied Snake N N ? NNNNNNNNNNN Tantilla coronata Southeastern Crowned Snake N Thamnophis brachystoma Short-headed Gartersnake N N Thamnophis sauritus Eastern Ribbonsnake NNNNNNNNNNNNNN Thamnophis sauritus septentrionalis Northern Ribbonsnake N N N Thamnophis sauritus sauritus Common Ribbonsnake NNNNNNNNNNNNNN Thamnophis sirtalis Common Gartersnake NNNNNNNNNNNNNN Thamnophis sirtalis pallidulus Maritime Gartersnake N N Thamnophis sirtalis sirtalis Eastern Gartersnake NNNNNNNNNNNNNN Virginia striatula Rough Earthsnake N Virginia valeriae Smooth Earthsnake N ? N N N N N Virginia valeriae pulchra Mountain Earthsnake N N N Virginia valeriae valeriae Eastern Smooth Earthsnake N ? N N N N N

Typhlopidae Blindsnakes Ramphotyphlops braminus Brahminy Blind Snake I I

Viperidae Pitvipers Agkistrodon contortrix Copperhead N N N N N N N N N N Agkistrodon contortrix contortrix Southern Copperhead N N Agkistrodon controtrix mokasen Northern Copperhead N N N N N N N N N N Agkistrodon piscivorus Cottonmouth N Agkistrodon piscivorus piscivorus Eastern Cottonmouth N Crotalus horridus Timber Rattlesnake N X X NNNNNNXNNN Sistrurus catenatus Massasauga N N Sistrurus catenatus catenatus Eastern Massasauga N N Key to species designations N = native species, extant D = endemic to state X = extirpated native species

NEPARC publication 2013-01 Northeast Partners in Amphibian and Reptile Conservation: Northeast Species List

Species Common Name Connecticut Delaware District of Columbia Maine Maryland Massachusetts New Hampshire New Jersey New York Pennsylvania Island Rhode Vermont Virginia West Virginia I = introduced, non-native species ? = status unknown

NEPARC publication 2013-01