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293 315 Nikitskij.P65 Åâðàçèàòñêèé ýíòîìîë. æóðíàë 4(4): 293315 © EUROASIAN ENTOMOLOGICAL JOURNAL, 2005 Ñèñòåìàòè÷åñêèé ñïèñîê æåñòêîêðûëûõ (Insecta, Coleoptera) Óäìóðòèè Checklist of beetles (Insecta, Coleoptera) of Udmurtia Ñ.Â. Äåäþõèí*, Í.Á. Íèêèòñêèé**, Â.Á. Ñåì¸íîâ*** S.V. Dedyukhin*, N.B. Nikitsky**, V.B. Semenov*** * Êàôåäðà ýêîëîãèè æèâîòíûõ, Óäìóðòñêèé ãîñóäàðñòâåííûé óíèâåðñèòåò, óë. Óíèâåðñèòåòñêàÿ 1, Èæåâñê 426034 Ðîññèÿ. E-mail: [email protected]. * Department of animal ecology, Udmurt State University, Universitetskaya str. 1, Izhevsk 426034 Russia. ** Çîîëîãè÷åñêèé ìóçåé ÌÃÓ èì. Ì.Â. Ëîìîíîñîâà, óë. Áîëüøàÿ Íèêèòñêàÿ 6, Ìîñêâà 125009 Ðîññèÿ. Email: [email protected]. ** Zoological Museum of Moscow Lomonosov State University, Bolshaya Nikitskaya 6, Moscow 125009 Russia. *** Èíñòèòóò ìåäèöèíñêîé ïàðàçèòîëîãèè è òðîïè÷åñêîé ìåäèöèíû èì. Å.È. Ìàðöèíîâñêîãî, óë. Ìàëàÿ Ïèðîãîâñêàÿ 20, Ìîñêâà 119435 Ðîññèÿ. *** Martsinovsky Institute of Medical Parasitology and Tropical Medicine, Malaya Pirogovskaya 20, Moscow 119435 Russia. Êëþ÷åâûå ñëîâà: Coleoptera, ôàóíà, Óäìóðòèÿ. Key words: Coleoptera, fauna, Udmurtia. Ðåçþìå. Ïðèâîäèòñÿ ïîëíûé ñïèñîê æåñòêîêðûëûõ óêàçàííûå â ïåðâîì ñïèñêå, çäåñü îòñóòñòâóþò, ÷òî, íàñåêîìûõ (Coleoptera) Óäìóðòèè, ãäå îòìå÷åíî íåìíî- ïî-âèäèìîìó, ãîâîðèò îá èõ îøèáî÷íîì îïðåäåëåíèè. ãèì áîëåå 2400 âèäîâ æóêîâ èç 91 ñåìåéñòâà. Âïåðâûå Ñ íàèáîëüøåé (ïðîïîðöèîíàëüíî îáùåìó ïîòåíöèàëü- äëÿ ðåãèîíà ïðèâåäåíî 1482 âèäà (ïîìå÷åíû â ñïèñêå íîìó ÷èñëó âèäîâ) ïîëíîòîé áûëè èçó÷åíû ñåìåéñòâà: çâåçäî÷êîé *, ñ ïðàâîé ñòîðîíû îò íàçâàíèÿ âèäà). Carabidae (139 âèäîâ), Cerambycidae (58), Scarabae- Abstract. A complete checklist (2400 species in 91 idae (s. l.) (57 âèäîâ), Elateridae (50 âèäîâ), à òàêæå families) of the beetles (Coleoptera) of Udmurtia is provided. Curculionidae (s. l.) (76), Buprestidae (20), Coccinellidae 1482 species are reported from Udmurtia for the first time, (17), äëÿ îñòàëüíûõ ïðèâåäåíû ëèøü îòäåëüíûå ïðåä- as indicated by an asterisk (*) in the list. ñòàâèòåëè.  öåëîì òðóäíî ïåðåîöåíèòü çíà÷åíèå äëÿ ïîçíà- Ðàñïîëîæåíèå Óäìóðòèè â ïðåäåëàõ Âÿòñêî-Êàìñ- íèÿ ðåãèîíàëüíîé êîëåîïòåðîôàóíû 25-ëåòíèõ èññëå- êîãî ìåæäóðå÷üÿ íà ãðàíèöå Ðóññêîé Ðàâíèíû è Ñðåä- äîâàíèé Â.È. Ðîùèíåíêî, âûâåäøèõ èçó÷åííîñòü äàí- íåãî Ïðåäóðàëüÿ (55° 12' 58°10' ñ.ø., 51° 10' 54° 26' íîé ãðóïïû íàñåêîìûõ â ðåñïóáëèêå íà êà÷åñòâåííî â.ä.), âáëèçè ðÿäà ìåðèäèîíàëüíûõ è íà ñòûêå íåñêîëü- íîâûé óðîâåíü. Îäíàêî, ê ñîæàëåíèþ, áîëüøèíñòâî êèõ øèðîòíûõ ðóáåæåé (îò þæíîé òàéãè äî ñåâåðíîé å¸ êîëëåêöèîííûõ ìàòåðèàëîâ ê íàñòîÿùåìó âðåìåíè ãðàíèöû ëåñîñòåïè) îïðåäåëÿåò ñâîåîáðàçèå äàííîãî óòåðÿíî. Ñîõðàíèëèñü ëèøü ñïðàâî÷íûå êîëëåêöèè ðåãèîíà â êîëåîïòåðîëîãè÷åñêîì îòíîøåíèè. Õîòÿ èñ- ïî íåêîòîðûì ñåìåéñòâàì, ýêçåìïëÿðû â êîòîðûõ áîëü- ñëåäîâàíèÿ æåñòêîêðûëûõ â Óäìóðòèè èìåþò äîñòà- øåé ÷àñòüþ áåç ýòèêåòîê, à îïðåäåëåíèå èõ íå âñåãäà òî÷íî äîëãóþ èñòîðèþ, êðóïíûå ìîíîãðàôè÷åñêèå âåðíî. Ïîýòîìó îáèòàíèå â ðåñïóáëèêå ðÿäà âèäîâ ñâîäêè, ïîñâÿù¸ííûå äàííîé ãðóïïå íàñåêîìûõ, åäè- æóêîâ, óêàçûâàåìûõ â å¸ ðàáîòàõ, íî íå îòìå÷åííûõ íè÷íû è î÷åíü íåïîëíû. Ïåðâûå ñâåäåíèÿ î íàõîäêàõ íàìè, òðåáóåò ïîäòâåðæäåíèÿ. Èñêëþ÷åíèå ñîñòàâëÿ- æóêîâ íà òåððèòîðèè ðåñïóáëèêè ïðèâîäÿòñÿ â ðàáîòàõ þò âèäû, ïðàâèëüíîñòü îïðåäåëåíèÿ êîòîðûõ äîñòî- À. Ëåáåäåâà [1906, 1913], À.È. Øåðíèíà [1927], Ë.Ê. Ýñ- âåðíî ïîäòâåðæäåíà ñïåöèàëèñòàìè (íàïðèìåð, ôàêò òåíáåðãà [1935], ïðîâîäèâøèõ èññëåäîâàíèÿ, â îñíîâ- îáíàðóæåíèÿ Crepidophorus rufiventris ïîäòâåðæä¸í íîì, â ñîñåäíèõ ðåãèîíàõ [Ðîùèíåíêî, Êîçëîâà, 1970]. Â.Ã. Äîëèíûì) èëè êîòîðûå ñîõðàíèëèñü â êîëëåêöè-  ïåðèîä ñ êîíöà 1950-õ ïî 1980-å ãîäû ïðîøëîãî ÿõ (Miltotrogus aequinoctialis, Onthophagus vacca, Coc- ñòîëåòèÿ îñóùåñòâëÿëîñü ìíîãîëåòíåå ñèñòåìàòè÷åñ- cinella trifasciata è íåêîòîðûå äð.). êîå èçó÷åíèå ìåñòíîé êîëåîïòåðîôàóíû ïîä ðóêîâîä- Íîâûé ïîäú¸ì êîëåîïòåðîëîãè÷åñêèõ èññëåäîâà- ñòâîì Â.È. Ðîùèíåíêî [Ðîùèíåíêî, 1965, 1969, 1970, íèé â Óäìóðòèè íà÷àëñÿ ñ 90-õ ãîäîâ ïðîøëîãî âåêà è 1971, 1973, 1977, 1978, 1979, 1981, 1988; Ðîùèíåíêî, ñâÿçàí, â ÷àñòíîñòè, ñ àêòèâèçàöèåé ïðèðîäîîõðàííîé Êîçëîâà, 1963, 1970; Êîçëîâà, 1965].  1972 ãîäó áûë äåÿòåëüíîñòè [Áîðèñîâñêèé, Àäàõîâñêèé, 1998, 1999; îïóáëèêîâàí ñïèñîê áåñïîçâîíî÷íûõ, âêëþ÷àþùèé Äåäþõèí, 2001; Äåäþõèí, Áîðèñîâñêèé, 2001]. Êî- 522 âèäà æåñòêîêðûëûõ èç 15 ñåìåéñòâ [Ðîùèíåíêî, íå÷íûì ðåçóëüòàòîì äàííûõ èññëåäîâàíèé ñòàë âû- 1972]. Áîëåå ïîëíûé ñïèñîê æóêîâ Óäìóðòèè, ñîñòîÿ- õîä Êðàñíîé Êíèãè ÓÐ, â êîòîðóþ âêëþ÷åíû 14 âèäîâ ùèé èç 675 âèäîâ, îòíîñÿùèõñÿ ê 35 ñåìåéñòâàì, æåñòêîêðûëûõ [Àäàõîâñêèé è äð., 2001].  ïðèëîæåíèè ïðèâîäèòñÿ â ïðèëîæåíèè ê êàíäèäàòñêîé äèññåðòà- [Òðåâîæíûé ñïèñîê], óêàçûâàþòñÿ åù¸ 18 âèäîâ æóêîâ. öèè Â.È. Ðîùèíåíêî [1975]. Ïðè÷¸ì íåêîòîðûå âèäû, Íåñêîëüêî âèäîâ ïðèâîäÿòñÿ â ðàáîòàõ, ïîñâÿù¸ííûõ 294 Ñ.Â. Äåäþõèí è äð. èçó÷åíèþ ýêîëîãèè íàñåêîìûõ-ôèòîôàãîâ [Åðìîëàåâ, ñïåöèàëèñòîâ-êîëåîïòåðîëîãîâ (À.Î. Áåíüêîâñêîãî, 2002; Åðìîëàåâ, Ñèäîðîâà, 2002], â òîì ÷èñëå Acantho- À.À. Ãóñàêîâà, À.À. Ëåãàëîâà è íåêîòîðûõ äð.).  cinus carinulatus (Cerambycidae) è Polydrusus inustus ñïèñêå ïðèâîäÿòñÿ íàçâàíèÿ ïîäîòðÿäîâ, íàäñåìåéñòâ, (Curculionidae), íàìè íå çàðåãèñòðèðîâàííûå. ñåìåéñòâ, ïîäñåìåéñòâ, ðîäîâ, âèäîâ (èíîãäà è ïîäâè- Ñðåäè êîëåîïòåðîëîãè÷åñêèõ ðàáîò, îñíîâàííûõ äîâ) æåñòêîêðûëûõ. Äëÿ êàæäîãî òàêñîíà ïðèâåäåíû íà íàøèõ äàííûõ, èìåþòñÿ ïóáëèêàöèè, ïîñâÿù¸í- ñîâðåìåííûå íàçâàíèÿ, äëÿ ÷àñòè òàêæå ñèíîíèìû. íûå îòäåëüíûì ãðóïïàì æåñòêîêðûëûõ Óäìóðòèè  ïðåäñòàâëåííûõ íèæå ñïèñêàõ íàäâèäîâûå òàêñîíû ìåðòâîåäàì (Silphidae) [Åðìîëàåâ, Äåäþõèí, 2000; äàíû áîëåå èëè ìåíåå â ñèñòåìàòè÷åñêîì ïîðÿäêå, Äåäþõèí, Åðìîëàåâ, 2001], ðîãà÷àì (Lucanidae) è òðîê- âèäû â àëôàâèòíîì. ñàì (Trogidae) [Äåäþõèí, Áîðèñîâñêèé, 2001], â öåëîì Âñåãî íà òåððèòîðèè Óäìóðòèè, íà ñåãîäíÿøíèé íàäñåìåéñòâó Scarabaeoidea [Äåäþõèí, 2003à], à òàê- äåíü, îòìå÷åíî íåìíîãèì áîëåå 2400 âèäîâ æóêîâ èç æå æåñòêîêðûëûì, îáèòàþùèì â íîðàõ ñóðêîâ [Äå- 91 ñåìåéñòâà. Âñå îíè ïðèâåäåíû â äàííîé ðàáîòå. äþõèí, Êàïèòîíîâ, 2004]. Âûøëà îáîáùàþùàÿ ñòà- 16 âèäîâ, â òî÷íîì îïðåäåëåíèè êîòîðûõ ïî òåì èëè òüÿ, ïîñâÿù¸ííàÿ îñîáåííîñòÿì ðåãèîíàëüíîé èíûì ïðè÷èíàì íåò óâåðåííîñòè, îòìå÷åíû çíàêîì ?. êîëåîïòåðîôàóíû (ñ ðàññìîòðåíèåì âåðîÿòíûõ ïóòåé Âïåðâûå äëÿ ðåãèîíà ïðèâåäåíî 1482 âèäà (ïîìå÷åíû å¸ ôîðìèðîâàíèÿ) è çàêîíîìåðíîñòÿì ïðîñòðàíñòâåí- â ñïèñêå çâåçäî÷êîé (*), ñ ïðàâîé ñòîðîíû îò íàçâàíèÿ íîãî ðàñïðåäåëåíèÿ æåñòêîêðûëûõ â ðåñïóáëèêå [Äå- âèäà). Äâóìÿ çâåçäî÷êàìè (**) îáîçíà÷åíû âèäû (âñå- äþõèí, 2003á], â êîòîðîé óêàçàíû îêîëî 160 âèäîâ ãî 63), ïðèâîäèìûå â îôèöèàëüíî îïóáëèêîâàííûõ æóêîâ (ñâûøå 90 âèäîâ âïåðâûå äëÿ ðåñïóáëèêè). Íà ëèòåðàòóðíûõ èñòî÷íèêàõ è â ðóêîïèñè äèññåðòàöèè îñíîâå äàííîé ñòàòüè ñ íåêîòîðûìè äîïîëíåíèÿìè â Â.È. Ðîùèíåíêî [1975] (â äàííîé ðàáîòå óêàçàíû ñ å¸ èíòåðíåò-âàðèàíòå îïóáëèêîâàíû òåçèñû ê òåëåêîí- ëè÷íîãî ñîãëàñèÿ), îäíàêî íå íàéäåííûå àâòîðàìè è ôåðåíöèè «Ñîâðåìåííàÿ áèîãåîãðàôèÿ» [Äåäþõèí, íå ñîõðàíèâøèåñÿ â êîëëåêöèÿõ, ïîýòîìó îáèòàíèå èõ 2003â]. Îòäåëüíûå âèäû ëèñòîåäîâ ïðèâîäÿòñÿ äëÿ â ðåñïóáëèêå òðåáóåò ïîäòâåðæäåíèÿ. Óäìóðòèè òàêæå â îïðåäåëèòåëå À.Î. Áåíüêîâñêîãî Ñëåäóþùèå 19 âèäîâ óêàçàíû â ëèòåðàòóðå äëÿ [Bieñkowski, 2004] íà îñíîâàíèè íàøèõ ìàòåðèàëîâ è Óäìóðòèè íà îñíîâàíèè îøèáî÷íûõ îïðåäåëåíèé, ÷òî ðàáîòû Â.È. Ðîùèíåíêî [1973]. ïîäòâåðæäàåòñÿ íàëè÷åì ýêçåìïëÿðîâ ñ îïðåäåëèòåëü-  îñíîâó äàííîãî ñïèñêà (êîòîðûé â ïåðñïåêòèâå íûìè ýòèêåòêàìè (õîòÿ ýòî íå èñêëþ÷àåò âîçìîæíîñòü ïðîâîäèìûõ ðàáîò, êîíå÷íî æå, áóäåò èçðÿäíî äîïîë- íàõîæäåíèÿ ïî êðàéíåé ìåðå íåêîòîðûõ èç íèõ íà íåí) ïîëîæåíû ìàòåðèàëû, ïîëó÷åííûå â ðåçóëüòàòå àíàëèçèðóåìîé òåððèòîðèè): Pterostichus magus Man- ìíîãîëåòíèõ èññëåäîâàíèé æåñòêîêðûëûõ Óäìóðòèè nerheim, 1825, P. ovoideus Sturm, 1824, Harpalus (1989-2004) Ñ.Â. Äåäþõèíà. Êðîìå òîãî, â ðàáîòå solitaris Dejean, 1829 (fuliginosus (Duftschmid, 1812), èñïîëüçîâàíû ñáîðû æóêîâ, ïðåäîñòàâëåííûå Â.È. Ðî- nec (Panzer, 1809)), Stenolophus proximus Dejean, 1829 ùèíåíêî, À.Ã. Áîðèñîâñêèì, Ä.À. Àäàõîâñêèì, Ê.À. Êà- (Carabidae), Saprinus cribellatus Marseul, 1862, S. im- ïèòîíîâûì (ÓäÃÓ), Í.Þ. Ïîïîâîé (Óäìóðòñêèé íàöè- mundus (Gyllenhal, 1827), Hypocaccus metallicus (Herbst, îíàëüíûé êðàåâåä÷åñêèé ìóçåé). 1792) (Histeridae), Aphodius merdarius (Fabricius, 1775), Èññëåäîâàíèÿìè óäàëîñü îõâàòèòü âåñü ñïåêòð çî- A. paykulli Bedel, 1908 (tessulatus (Paykull, 1798), nec íàëüíî-ëàíäøàôòíûõ óñëîâèé ðåñïóáëèêè: ïîäçîíó (Moll, 1782)), Omaloplia nigromarginata (Herbst, 1785) þæíîé òàéãè (ñåâåð Óäìóðòèè); ïîëîñó ïîäòà¸æíûõ (ruricola auct. nec (Fabricius, 1775)), Amphimallon õâîéíî-øèðîêîëèñòâåííûõ ëåñîâ ñ ìàññèâàìè ñîñíî- assimile (Herbst, 1790), Anisoplia agricola (Poda, 1761), âûõ áîðîâ íà íàäïîéìåííûõ òåððàñàõ ðåê è ìàòåðèêî- A. segetum (Herbst, 1783) (Scarabaeidae), Melanotus âûõ ïåñ÷àíûõ äþíàõ (â öåíòðàëüíîé ÷àñòè); ïîëîñó villosus (Geoffroy, 1785) (rufipes (Herbst, 1784), nec øèðîêîëèñòâåííî-õâîéíûõ ëåñîâ ñ ïðèñóòñòâèåì äóáà (Goeze, 1777)) (Elateridae), Cryptocephalus quatuor- è íàëè÷èåì ó÷àñòêîâ îñòåïí¸ííûõ ëóãîâ íà ñêëîíàõ è decimmaculatus Schneider, 1792, Chrysolina hyperici â äîëèíàõ êðóïíûõ ðåê (Êàìà è Âÿòêà) (â þæíîé òðå- (Forster, 1771) (Chrysomelidae), Anastrangalia dubia òè Óäìóðòèè). (Scopoli, 1763) (Cerambycidae), Dorytomus dejeani Ïðåäâàðèòåëüíîå îïðåäåëåíèå æåñòêîêðûëûõ ïðî- Faust, 1882, Phyllobius jacobsoni Smirnov, 1913, Acalles âåäåíî Ñ.Â. Äåäþõèíûì. Ïðîâåðêà îïðåäåëåíèÿ æó- lemur (Germar, 1824) (Curculionidae). êîâ ïî áîëüøèíñòâó ñåìåéñòâ è èäåíòèôèêàöèÿ ðÿäà  ðàáîòå íå ïðèâîäèòñÿ äàííûõ î ðàñïðîñòðàíå- îñîáî ñëîæíûõ äëÿ äèàãíîñòèðîâàíèÿ òàêñîíîâ íèè æåñòêîêðûëûõ â ðåñïóáëèêå1 . Îäíàêî, ñëåäóåò (Leiodidae, Scirtidae, Scaphidiinae,
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