FAUNA OF

Volume 5, Number 2

Syrphidae Ⅱ

Arthropoda: Insecta: Diptera: Brachycera: Syrphidae: Milesiinae

2018

National Institute of Biological Resources Ministry of Environment

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Copyright © 2018 by the National Institute of Biological Resources

Published by the National Institute of Biological Resources Environmental Research Complex, 42, Hwangyeong-ro, Seo-gu Incheon 22689, Republic of Korea www.nibr.go.kr

All rights reserved. No part of this book may be reproduced, stored in a retrieval system, or transmitted, in any form or by any means, electronic, mechanical, photocopying, recording, or otherwise, without the prior permission of the National Institute of Biological Resources.

ISBN: 978-89-6811-320-8(96470), 978-89-94555-00-3(Set) Government Publications Registration Number: 11-1480592-001355-01

Printed by Doohyun Publishing Co. in Korea on acid-free paper

Publisher: National Institute of Biological Resources Authors: Deuk-Soo Choi ( and Plant Quarantine Agency), Sang-Wook Suk (Yonsei Univ.), Su-Bin Lee (Animal and Plant Quarantine Agency), Ho-Yeon Han (Yonsei Univ.) Project Staff: Jung-Sun Yoo, Hong Yul Suh, Jinwhoa YUM, Tae Woo Kim, Seon Yi Kim

Published on February 20, 2018

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Volume 5, Number 2

Syrphidae Ⅱ

Arthropoda: Insecta: Diptera: Brachycera: Syrphidae: Milesiinae

Deuk-Soo Choi1, Sang-Wook Suk2, Su-Bin Lee1 and Ho-Yeon Han2 1Animal and Plant Quarantine Agency, Gimcheon-si, Gyeongsangbuk-do 2Yonsei University, Wonju-si, Gangwon-do

곤충-알락긴꽃등에아과Ⅱ_영문 수정.indd 3 2018-09-12 오후 12:13:01 The Flora and Fauna of Korea logo was designed to represent six major target groups of the project including vertebrates, invertebrates, , algae, fungi, and bacteria.

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The biological resources include all the composition of organisms and genetic resources which possess the practical and potential values essential to human live. Biological resources will be firmed competition of the nation because they will be used as fundamental sources to make highly valued products such as new lines or varieties of biological organisms, new material, and drugs. As the Nagoya Protocol was adopted in 2010 and entered into force in the 12th Conference of Parties of the Convention on Biological Diversity (CBD) in 2014, it is expected that the competition to get biological resources will be much intensive under the rapidly changed circumstance on the access and benefic sharing of the genetic resources (ABS). To cope with a new international paradigm on all kinds of issues related to biological resources, the Ministry of Environment of Korea enforced a new law called ‘An act on access and benefit sharing of genetic resources’ on August 17th, 2017. Each nation in the world is investigating and clearing information of native within its territory in order to secure its sovereignty rights over biological resources. The National Institute of Biological Resources (NIBR) of the Ministry of Environment has published the ‘Flora and Fauna of Korea’ since 2006 to manage biological resources in comprehensive ways and to enhance national competitiveness by building up the foundation for the sovereignty over biological resources. Professional research groups consisting of professors and related experts of examined systematically a total of 14,336 species for the past eight years to publish 173 volumes in both Korean and English versions, and two volumes of World Monograph covering 216 species of invertebrates. This year, 13 volumes of the Flora and Fauna of Korea in both Korean and English versions including 1,407 species of invertebrates, insects and vascular plants are additionally published. Flora and Fauna of Korea are the first professional records to describe all the species of the nation in a comprehensive way, and they would contribute to level up the taxonomic capacity. The NIBR will continue to publish flora and fauna of Korea that will contribute conservation and application of biological resources for successful implementation of the ABS protocol. Finally, I would like to express my sincere appreciation to authors who spared no effort to publish the Flora and Fauna of Korea.

President of the National Institute of Biological Resources

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CONTENTS

List of Taxa 5 Introduction 10 Materials and Methods 11 Taxonomic Notes 14 1. Callicera aenea (Fabricius) 17 2. Monoceromyia pleuralis (Coquillett) 18 3. Primocerioides petri (Hervé-Bazin) 19 4. Sphiximorpha rachmaninovi (Violovitsh) 20 5. abbreviata Shiraki 22 6. Cheilosia aterrima (Sack) 23 7. Cheilosia bombiformis (Matsumura) 24 8. Cheilosia facialis Shiraki 25 9. Cheilosia fuscipennis Shiraki 25 10. Cheilosia illustrata (Harris) 25 11. Cheilosia longiptera Shiraki 26 12. Cheilosia matsumurana (Shiraki) 27 13. Cheilosia motodomariensis Matsumura 28 14. Cheilosia ochripes (Shiraki) 29 15. Ferdinandea cuprea (Scopoli) 30 16. Ferdinandea nigrifrons (Egger) 31 17. Ferdinandea ruficornis (Fabricius) 32 18. Rhingia laevigata Loew 33 19. Brachyopa testacea (Fallén) 34 20. karumaiensis (Matsumura) 35 21. clunipes (Fallén) 36 22. Sphegina japonica Shiraki and Edashige 38 23. aeneus (Scopoli) 39 24. Eristalinus quinquestriatus (Fabricius) 40 25. (Linnaeus) 40 26. Eristalinus tarsalis (Macquart) 41 27. Eristalinus viridis (Coquillett) 42 28. Eristalis arbustorum (Linnaeus) 44 29. Eristalis cerealis Fabricius 46

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30. Eristalis japonica van der Goo 48 31. Eristalis kyokoae (Kimura) 48 32. Eristalis tenax (Linnaeus) 49 33. Helophilus lunulatus Meigen 52 34. Helophilus sapporensis Matsumura 53 35. Helophilus trivittatus (Fabricius) 53 36. Helophilus virgatus Coquillett 54 37. Mallota ambigua (Shiraki) 56 38. Mallota analis (Shiraki) 57 39. Mallota dimorpha (Shiraki) 58 40. Mallota eurasiatica Stackelberg 60 41. Mallota koreana (Enderlein) 60 42. Mallota rossica Portschinsky 60 43. Mallota shatalkini Mutin 61 44. Mallota takasagoensis Matsumura 62 45. Mallota tricolor Loew 62 46. Mesembrius flaviceps (Matsumura) 64 47. Mesembrius niger Shiraki 65 48. Phytomia zonata (Fabricius) 65 49. Pseuderistalis bicolor (Shiraki) 67 50. djakonovi Stackelberg 68 51. Eumerus iidai Shiraki 68 52. Eumerus japonicus Matsumura 69 53. Eumerus kongosanensis Shiraki 70 54. (Fallén) 70 55. equestris (Fabricius) 72 56. Merodon kawamurai Matsumura 72 57. brevicornis Shiraki 73 58. japonica (Shiraki) 74 59. Blera nitens (Stackelberg) 76 60. Criorhina brevipila Loew 76 61. Criorhina trifasciata Shiraki 77 62. Macrozelima hervei (Shiraki) 77 63. Matsumyia jesoensis (Matsumura) 78 64. Matsumyia nigrofacies Shiraki 79 65. Milesia undulata Vollenhoven 80

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66. Narumyia narumii Shiraki 81 67. Pterallastes unicolor (Shiraki) 81 68. Spilomyia permagna Stackelberg 83 69. Spilomyia suzukii Matsumura 83 70. Syritta orientalis Macquart 84 71. Syritta pipiens (Linnaeus) 85 72. bombylans (Fabricius) 87 73. Temnostoma frequense Matsumura 88 74. Temnostoma vespiforme (Linnaeus) 88 75. rostrata Shiraki 90 76. abdominalis (Shiraki and Edashige) 91 77. Pipiza austriaca Meigen 92 78. Pipiza flavimaculata Matsumura 93 79. Pipiza inornata Matsumura 94 80. Pipiza signata Meigen 94 81. Pipizella biglumis (Matsumura) 95 82. Stenopipiza bipunctata Matsumura 95 83. primus Loew 96 84. Pseudovolucella decipiens (Hervé-Bazin) 97 85. Sericomyia dux (Stackelberg) 99 86. Sericomyia sachalinica Stackelberg 99 87. Graptomyza ihai Shiraki 100 88. Volucella bivitta Huo, Ren and Zhemin 103 89. Volucella coreana Shiraki 104 90. Volucella inanoides Hervé-Bazin 105 91. Volucella jeddona Bigot 106 92. Volucella linearis Walker 107 93. Volucella nigricans Coquillett 108 94. Volucella nigropicta Portschinsky 109 95. Volucella pellucens tabanoides Motschulsky 110 96. Volucella suzukii Matsumura 112 97. Volucella thompsoni Choi, Ôhara and Han 113 98. Brachypalpus nipponicus Shiraki 114 99. carbonus (Violovitsh) 117 100. Chalcosyrphus femoratus (Linnaeus) 118 101. Chalcosyrphus laterimaculatus Jeong, Jung and Han 120

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102. Chalcosyrphus longus (Coquillett) 121 103. Chalcosyrphus nemorum (Fabricius) 122 104. Chalcosyrphus nitidus (Portschinsky) 123 105. Chalcosyrphus piger (Fabricius) 124 106. Chalcosyrphus rufipes (Loew) 125 107. Chalcosyrphus unicus (Violovitsh) 126 108. abiens Meigen 128 109. Xylota coquilletti Hervé-Bazin 129 110. Xylota ignava (Panzer) 130 Literatures Cited 131 Plates 135 Index to Scientific Names 146

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LIST OF TAXA

Phylum Arthropoda von Siebold, 1848 Class Insecta Linnaeus, 1758 Order Diptera Linnaeus, 1758 Suborder Brachycera Macquart, 1834 Family Syrphidae Latreille, 1802 Subfamily Milesiinae Bezzi, 1893 Tribe Callicerini Callicera Panzer, 1806 Callicera aenea (Fabricius, 1781) Tribe Ceriodini Genus Monoceromyia Shannon, 1922 Monoceromyia pleuralis (Coquillett, 1898) Genus Primocerioides Shannon, 1927 Primocerioides petri (Hervé-Bazin, 1914) Genus Sphiximorpha Rondani, 1850 Sphiximorpha rachmaninovi (Violovitsh, 1981) Tribe Cheilosiini Genus Cheilosia Meigen, 1822 Cheilosia abbreviata Shiraki, 1953 Cheilosia aterrima (Sack, 1927) Cheilosia bombiformis (Matsumura, 1916) Cheilosia facialis Shiraki, 1968 Cheilosia fuscipennis Shiraki, 1953 Cheilosia illustrata (Harris, 1780) Cheilosia longiptera Shiraki, 1968 Cheilosia matsumurana (Shiraki, 1930) Cheilosia motodomariensis Matsumura, 1916 Cheilosia ochripes (Shiraki, 1930) Genus Ferdinandea Rondani, 1844 Ferdinandea cuprea (Scopoli, 1763) Ferdinandea nigrifrons (Egger, 1860) Ferdinandea ruficornis (Fabricius, 1775) Genus Rhingia Scopoli, 1763

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Rhingia laevigata Loew, 1858 Tribe Chrysogasterini Genus Brachyopa Meigen, 1822 Brachyopa testacea (Fallén, 1817) Genus Orthonevra Macquart, 1829 Orthonevra karumaiensis (Matsumura, 1916) Genus Sphegina Meigen, 1822 (Fallén, 1816) Sphegina japonica Shiraki and Edashige, 1953 Tribe Genus Eristalinus Rondani, 1845 (Scopoli, 1763) Eristalinus quinquestriatus (Fabricius, 1794) Eristalinus sepulchralis (Linnaeus, 1758) Eristalinus tarsalis (Macquart, 1855) Eristalinus viridis (Coquillett, 1898) Genus Eristalis Latreille, 1804 Eristalis arbustorum (Linnaeus, 1758) Eristalis cerealis Fabricius, 1805 Eristalis japonica van der Goot, 1964 Eristalis kyokoae (Kimura, 1986) Eristalis tenax (Linnaeus, 1758) Genus Helophilus Meigen, 1822 Helophilus lunulatus Meigen, 1822 Helophilus sapporensis Matsumura, 1911 Helophilus trivittatus (Fabricius, 1805) Helophilus virgatus Coquillett, 1898 Genus Mallota Meigen, 1822 Mallota ambigua (Shiraki, 1968) Mallota analis (Shiraki, 1968) Mallota dimorpha (Shiraki, 1930) Mallota eurasiatica Stackelberg, 1950 Mallota koreana (Enderlein, 1934) Mallota rossica Portschinsky, 1877 Mallota shatalkini Mutin, 1999 Mallota takasagoensis Matsumura, 1916

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Mallota tricolor Loew, 1871 Genus Mesembrius Rondani, 1857 Mesembrius flaviceps (Matsumura, 1905) Mesembrius niger Shiraki, 1968 Genus Phytomia Guérin-Méneville, 1833 Phytomia zonata (Fabricius, 1787) Genus Pseuderistalis Shiraki, 1930 Pseuderistalis bicolor (Shiraki, 1930) Tribe Genus Eumerus Meigen, 1822 Eumerus djakonovi Stackelberg, 1952 Eumerus iidai Shiraki, 1952 Eumerus japonicus Matsumura, 1916 Eumerus kongosanensis Shiraki, 1930 Eumerus strigatus (Fallén, 1817) Genus Merodon Meigen, 1803 (Fabricius, 1794) Merodon kawamurai Matsumura, 1916 Genus Psilota Meigen, 1822 Psilota brevicornis Shiraki, 1968 Tribe Genus Blera Billberg, 1820 Blera japonica (Shiraki, 1930) Blera nitens (Stackelberg, 1923) Genus Criorhina Meigen, 1822 Criorhina brevipila Loew, 1871 Criorhina trifasciata Shiraki, 1930 Genus Macrozelima Stackelberg, 1930 Macrozelima hervei (Shiraki, 1930) Genus Matsumyia Shiraki, 1949 Matsumyia jesoensis (Matsumura, 1911) Matsumyia nigrofacies Shiraki, 1949 Genus Milesia Latreille, 1804 Milesia undulata Vollenhoven, 1863 Genus Narumyia Shiraki, 1952 Narumyia narumii Shiraki, 1952

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Genus Pterallastes Loew, 1863 Pterallastes unicolor (Shiraki, 1930) Genus Spilomyia Meigen, 1803 Spilomyia permagna Stackelberg, 1958 Spilomyia suzukii Matsumura, 1916 Genus Syritta Lepeletier and Serville, 1828 Syritta orientalis Macquart, 1842 Syritta pipiens (Linnaeus, 1758) Genus Temnostoma Lepeletier and Serville, 1828 Temnostoma bombylans (Fabricius, 1805) Temnostoma frequense Matsumura, 1931 Temnostoma vespiforme (Linnaeus, 1758) Genus Tropidia Meigen, 1822 Tropidia rostrata Shiraki, 1930 Tribe Genus Pipiza Fallén, 1810 Pipiza abdominalis (Shiraki and Edashige, 1953) Pipiza austriaca Meigen, 1822 Pipiza flavimaculata Matsumura, 1918 Pipiza inornata Matsumura, 1916 Pipiza signata Meigen, 1822 Genus Pipizella Rondani, 1856 Pipizella biglumis (Matsumura, 1916) Genus Stenopipiza Matsumura, 1919 Stenopipiza bipunctata Matsumura, 1919 Genus Triglyphus Loew, 1840 Triglyphus primus Loew, 1840 Tribe Sericomyini Genus Pseudovolucella Shiraki, 1930 Pseudovolucella decipiens (Hervé-Bazin, 1914) Genus Sericomyia Meigen, 1803 Sericomyia dux (Stackelberg, 1930) Sericomyia sachalinica Stackelberg, 1925 Tribe Volucellini Genus Graptomyza Wiedemann, 1820 Graptomyza ihai Shiraki, 1968

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Genus Volucella Geoffroy, 1762 Volucella bivitta Huo, Ren and Zhemin, 2007 Volucella coreana Shiraki, 1930 Volucella inanoides Hervé-Bazin, 1923 Volucella jeddona Bigot, 1875 Volucella linearis Walker, 1852 Volucella nigricans Coquillett, 1898 Volucella nigropicta Portschinsky, 1883 Volucella pellucens tabanoides Motschulsky, 1859 Volucella suzukii Matsumura, 1916 Volucella thompsoni Choi, Ôhara and Han, 2006 Tribe Xylotini Genus Brachypalpus Macquart, 1834 Brachypalpus nipponicus Shiraki, 1952 Genus Chalcosyrphus Curran, 1925 Chalcosyrphus carbonus (Violovitsh, 1975) Chalcosyrphus femoratus (Linnaeus, 1758) Chalcosyrphus laterimaculatus Jeong, Jung and Han, 2017 Chalcosyrphus longus (Coquillett, 1898) Chalcosyrphus nemorum (Fabricius, 1805) Chalcosyrphus nitidus (Portschinsky, 1879) Chalcosyrphus piger (Fabricius, 1794) Chalcosyrphus rufipes (Loew, 1873) Chalcosyrphus unicus (Violovitsh, 1977) Genus Xylota Meigen, 1822 Xylota abiens Meigen, 1822 Xylota coquilletti Hervé-Bazin, 1914 Xylota ignava (Panzer, 1798)

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INTRODUCTION

Syrphidae (Diptera: Brachycera) commonly called ‘’ or ‘flowerflies’, with almost 6,000 described species and is a speciose family of Diptera (Miranda et al., 2013; Pape and Thompson, 2013). Syrphid provide crucial ecosystem services as pollinations, biological control agents and in environmental assessment (Kevan, 2002; Ssymank et al, 2008; Mengual and Thompson, 2011). Almost all adult syrphids visit flowers for pollen and nectar. Larvae, on the other hand, are found in a very diverse array of habitatas, including fungal fruiting bodies (mycophagy), dung, nests of social Hymenoptera, decaying wood and water bodies of several types (saprophagy), mining in leafs and stems of numerous plant families (phytophagy), and as predators of (Thompson and Rotheray, 1998). The family Syrphidae is traditionally divided into three subfamilies, Microdontinae, Milesiinae (= ) and Syrphinae, and 15 tribes are currently recognized in the family (Young et al., 2016). Immature stages of Microdontinae are inquilines in ant’s nests feeding on eggs, larvae and pupae, whereas larvae of Milesiinae include saprophages in a wide range of decaying media from dung to dead wood, while some species of the genus Volucella are - and - brood predators (Rotheray, 2003; Mengual et al., 2008). Larvae of Syrphinae are mostly predaceous on soft-bodied Hemiptera such as , coccids and psyllids, but some species feed on Thysanoptera, immature Coleoptera and / or (Rotheray, 1993; Rojo et al., 2003; Mengual et al., 2008).

Historical review of the family Syrphidae in Korea

Eristalis cerealis Fabricius was the first Korean syphid species reported by Ichigawa (1906). Later, Shiraki (1930) for the first time provided a serious revisionary work based on a collection primarily from Taiwan, Korea, Japan, and Sakhalin. In this revision, he described 302 species, among which 27 were from Korea. More recently, Kim, CW (1971) revised Korean Syrphidae, providing Descriptions and illustrations of 82 nominal species. Kim, CW (1980) later provided detailed distribution maps of 34 Korean syrphid species. Kim, JI (1975, 1980) reviewed Korean syrphid references and provided a check list of 109 species with source references. ESK & KSAE (1994) included a total 127 syrphid species without providing source references. Han and Choi (2001) provided a check list of 150 species, including 12 newly recorded species in Korea. Han et al. (2014), in the most updated check list, included a total 175 species. Suk and Han (2014a), Suk et al. (2015) and Jeong et al. (2017) added new records of 13 additional species in Korea. In total, three subfamilies, 70 genera and 188 species of the family Syrphidae are now officially recognized for the Korean fauna.

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MATERIAL AND METHODS

The morphological terminology and interpretations used in this study largely follow Thompson (1999) and McAlpine (1981). The Korean terms used here basically follows KSAE & ESK (1998) and Park et al. (2013). For synonymy, we followed peck (1988), Han and Choi (2001), Pape and Thompson (2013) and Han et al. (2014). All photographs were obtained using Olympus C-2500L, Panasonic DMC FZ50 digital camera and Leica DMS300 camera attached to a stereo microscope, processed using Leica Application Suite software (LAS EZ).

B

A

C

D E

Fig. 1. Mensuration system (A, D, E: Volucella. nigropicta, male, B, C: V. nigricans). A. Body in dorsal view (wings and right legs removed); B. Cells of wing; C. Veins and maculae of wing; D. Head in anterior view; E. Head in lateral view.

Abbreviations: BL-body length, BTL-basotarsomere length, HW-head width, LE-longest eye diameter, ML-mesonotum length, SE-shortest eye diameter, TL-tibia length, WL-wing length, m sctl s-marginal scutellar seta, npl s-notopleural seta, pal s-postalar seta, psct s-prescutellar seta, spal s-supra-alar seta.

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We examined more than 500 specimens of Syrphidae from Korea. Detailed collection data are provided for each species, the acronyms of collecting localities are as follows: Chungcheongbuk-do (CB), Chungcheongnam-do (CN), Gyeongsangbuk-do (GB), Gyeongsangnam-do (GN), Gyeonggi-do (GG), Gangwon-do (GW), Jeollabuk-do (JB), Jeollanam-do (JN), Jeju-do (JJ). The majority of the examined specimens are deposited in the YSUW (Division of Biological Science and Technology, Yonsei University, Wonju-si) and APQA (Animal and Plant Quarantine Agency, Gimcheon-si), Korea. The acronyms of the terms and zoogeographical regions mentioned in the text are as follows: Holotype (HT), Monotypy (MO), Original designation (OD), Type locality (TL), Type species (TS); Afrotropical (AF), Australasian (AU), Nearctic (NE), Neotropical (NT), Oriental (OR), Palaearctic (PA).

mesonotum scutum scutellum A

anepst s sct trn sut trn npl pal cal sctl

pprn wing base anatg halter 1 anepst 4 3 ktg 2 5 1st t anepm 2nd t kepm

cx1 mr kepst 1st s 2nd s

cx2 ms cx3

coxa B C scape femur pedicel trochanter

arista tibia antenna basoflagellomere

basotarsomere

tarsus

Fig. 2. A‒C (Volucella thompsoni). A. Thorax in lateral view (modified from Thompson, 1999); B. Antenna in lateral view; C. Leg in lateral view.

Abbreviations: 1-anterior anepisternum, 2-posterior anepisternum, 3-anterior anepimeron, 4-dorsomedial

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anepimeron, 5-posterior anepimeron, anatg-anatergite, anepim-anepimeron, anepst-anepisternum, anepst s-anepisternal seta, cx-coxa, kepm-katepimeron, kepst-katepisternum, ktg-katatergite, mr-meron, ms- metasternum, npl-notopleuron, pal cal-postalar callus, pprm-postpronotum, sct-scutum, sctl-scutellum, trn sut-transverse suture.

frontal lunule triangle A B frons eye contiguity Male Female

eye eye

vertical ocellar triangle triangle

cerc C D epand

aed apo

hypd b epand lb hypoproct sur C-1 cerc

aed

Fig. 3. A‒D (A and B: Volucella nigropicta, C and D: V. elegans). A. Male head in dorsal view; B. Female head; C. Male genitalia in lateral view; C-1. Sutylus, innerview; D. ditto, posterodorsal view.

Abbreviations: aed-aedeagus, aed apo-aedeagal apodeme, b epand lb-basal epandirial lobe, cerc-cercus, epand-epandrium, hypd-hypandrium, sp lb-superior lobe, sur-surstylus.

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TAXONOMIC NOTES

Phylum Arthropoda von Siebold, 1848

Class Insecta Linnaeus, 1758

Order Diptera Linnaeus, 1758

Suborder Brachycera Macquart, 1834

Family Syrphidae Latreille, 1802

Type: Syrphus Fabricius.

Species: ca. 6,000 (188 in Korea).

Distribution: Worldwide.

Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

Key to subfamilies of Korean Syrphidae (after Vockeroth and Thompson, 1987)

1. Postpronotum bare. Head posteriorly strongly convex and closely appressed to thorax; postpronotum partly or entirely hidden. Male abdomen with tergite 5 visible in dorsal view························ Syrphinae − Postpronotum pilose. Head posteriorly less strongly convex; postpronotum clearly exposed. Male abdomen with tergite 5 not visible in dorsal view································· Milesiinae and Microdontinae

Subfamily Milesiinae Bezzi, 1893

Key to genera of Milesiinae from

1. Antennae longer than head······························································································ 2 − Antennae shorter than head····························································································· 5 2. Eye hairly; frontal prominence absent ······································································ Callicera − Eye bare; frontal prominence present·················································································· 3 3. Abdominal tergite 2 narrow than scutellum··········································································· 4

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− Abdominal tergite 2 broad than scutellum·························································· Primocerioides 4. Antenna attached scape················································································· Monoceromyia − Antenna separated scape················································································· Sphiximorpha 5. Arista plumose············································································································ 6 − Arista not plumose········································································································ 8 6. Wing with cell R1 closed·······················································································Volucella − Wing with cell R1 opened······························································································· 7 7. Wing with black tinge medially ····································································· Pseudovolucella − Wing without black tinge medially········································································ Sericomyia 8. Wing with cell R1 closed································································································ 9 − Wing with cell R1 opend······························································································ 10 9. Scutellum entirely black; eyes spotted····································································· Eristalinus − Scutellum brown or partly black; eyes not spotted (with 2 bands of hair)······························· Eristalis 10. Scutellum concave medially ·············································································· Graptomyza − Scutellum convex medially ··························································································· 11 11. Wing vein R4+5 strongly looped ···················································································· 12 − Wing vein R4+5 straight or slightly looped ········································································ 18 12. Wing with black tinge medially ············································································· Phytomia − Wing without black tinge medially··················································································· 13 13. Eye dichoptic both sexes ···················································································· Helophilus − Eye holoptic in male ··································································································· 14 14. Scutum with 2 distinctly longitudinal bands ······································································· 15 − Scutum without longitudinal band ··················································································· 16 15. Basoflagellomere black· ·················································································· Mesembrius − Basoflagellomere yellow brown· ············································································· Tropidia 16. Hind femur with large triangular plate apically ···························································· Merodon − Hind femur without tooth or triangular plate ······································································· 17 17. Vein r-m right angle with vein R4+5 ········································································· Mallota − Vein r-m curved to basal ··················································································· Pterellastes 18. Scutum covered uniformaly yellow brown long dense hairs ·········································· Matsumyia − Scutum not above ······································································································ 19 19. Wing vein br and r-m with dark tinge ··············································································· 20 − Wing vein br and r-m without dark tinge ··········································································· 22 20. Face not convex ···························································································· Ferdinandea − Face distinctly convex ································································································· 21 21. Hind femur almost black with reddish brown apex; tergite 3 with a pair of corss bands ··········· Criorhina

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− Hind femur black with reddish brown basally; tergite 2 without band ··························· Brachypalpus 22. Abdominal tergite 3 visible in dorsal view ······························································· Triglyphus − Abdominal tergite 4 distinctly visible in dorsal view ····························································· 23 23. Mouth edge elongated into a long snout ····································································· Rhingia − Mouth edge not elongaged into a long snout ······································································· 24 24. Abdominal tergite 2 narrow than scutellum ································································ Sphegina − Abdominal tergite 2 broader than scutellum ······································································· 25 25. Scutella hair fringe present ·················································································· Cheilosia − Scutella hair fringe absent ···························································································· 26 26. Abdominal tergite 2 entirely reddish orange ·························································· Macrozelima − Abdominal tergite 2 entirely black or partly black ································································ 27 27. Vein M2 right angle with M1 ·············································································· Orthonevra − Vein M2 not right angle with M1 ···················································································· 28 28. Postpronotum yellow ·································································································· 29 − Postpronotum black or grey ·························································································· 31 29. Abdominal tergite 2 with unique pattern ····································································· Milesia − Abdominal tergite 2 with cross band ················································································ 30 30. Fore tibia entirely black; hind femur without spine basally ·········································· Temnostoma − Fore tibia entirely yellowish brown; hind femur with spine basally ··································· Spilomyia 31. Metasternum hairly ···································································································· 32 − Metasternum bare ······································································································ 33 32. Hind femur greatly enlarged ····················································································· Syritta − Hind femur not enlarged ··············································································· Chalcosyrphus 33. Wing longer than body size ················································································· Brachyopa − Wing shorter than body size ·························································································· 34 34. Abdomen elongated ······························································································ Xylota − Abdomen oval ·········································································································· 35 35. Abdominal tergite 2‒4 oval ··················································································· Eumerus − Abdominal tergite 2‒4 not above ···················································································· 36 36. Frons convex; face elongated below ············································································ Blera − Frons concave; face not elongated below ·········································································· 37 37. Eye with dense hairs ······························································································ Psilota − Eye bare or sparsely haired ······················································································ Pipiza

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Tribe Callicerini

Genus Callicera Panzer, 1806

Callicera Panzer [1809]: Faunae insect. germ., Fasc. 104: 17. TS: Bibio aenea Fabricius (MO).

1. Callicera aenea (Fabricius, 1781) (Pl. 1A)

Bibio aenea Fabricius, 1781: 413. Syrphus aurata Rossi, 1790 Callicera panzerii Rondani, 1844.

Korean record: Callicera aenea: Han et al. 1998a: 10 (Yonsei Univ. Maeji Campus, first Korean record); Han et al. 1998b: 129 (Korean cat.); Choi et al. 2000: 143 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 120 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 9 (Korean check.).

Description: Body length 13.2‒15.3 mm. Male. Head shiny black ground color; face with long yellow brown pilose laterally; frons shiny black without pile; eyes holoptic, dense pilose; antenna almost entirely black with black setulae except basoflagellomere apically. Thorax shiny black ground color with 4 distinct longitudinal dust-strpies; scutellum shiny black with dense long whitish brown hairs. Wing hyaline with brown tinge anteriorly; halter yellow brown. Legs: coxae and trochanters black, covered with yellowish brown pile; femora entirely black except apical 1/3‒1/4 yellow brown, covered with yellowish brown hairs; tibiae almost entirely whitish yellow. Abdomen almost entirely black with dense yellowish brown hairs. Female. Similar to male except for following non-genitalic characters: eye dichoptic.

Specimen examined: Korea: CB: Chungju-si, Hansu-myeon, Songgye-ri, Mt. Woraksan, 10. v. 2005, DS Choi et al., 1♂; Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 3. vi. 2004, HW Byun et al., 1♂. Danyang- gun, Sobaeksan Observatory, 5. vi. 2016, DS Choi, 1♂. CN: Cheonan-si, Seonggeo-eup, Cheonheung-ri, 21. v. 2006, HW Byun et al., 1♂; Seosan-si, Unsan-myeon, Mt. Gayasan, 28. v. 2011, SW Suk et al., 1♂. GB: Gyeongju-si, Geoncheon-eup, Mt. Danseoksan, 5. v. 2011, DS Choi, 2♂. GN: Miryang-si, Pyochungsa, 7. vi. 1995, SW Lee, 1♀. GG: Gwangju-si, Opo-myeon, Mr. Baengmasan, 27. v. 2001. DS Choi et al., 1♂. GW: Hongcheon-gun, Nae-myeon, Mt. Gachilbong, from Sambongyaksu to 1,240 m peak, 27. v. 2011, YB Lee & HS Lee, 1♂; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 30. v. 2006, DJ Cha, 1♂. JN:

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Gwangju, Dong-gu, Mt. Mudeungsan, 13. v. 2000, DS Choi et al., 1♂. Distribution: , Russia, China, Korea, Taiwan [PA, OR]. Biology: Larvae of this species breed in rot-holes on Fagus (Rotheray, 1993). Korea: CB, CN, GB, GN, GG, GW, JB, JN.

Tribe Ceriodini

Genus Monoceromyia Shannon, 1922

Monoceromyia Shannon, 1922: Bull. Brook. ent. Soc., 17: 41 (as subgenus of Cerioides Rondani). TS: Ceria tricolorLoew (MO). Sphyximorphoides Shiraki, 1930: Mem. Fac. Agric. Taihoku imp. Univ., 1 (1): 6 (as subgenus of Cerioides Rondani).TS: Sphyximorpha pleuralis Coquillett (OD).

2. Monoceromyia pleuralis (Coquillett, 1898) (Pl. 1B)

Sphiximorpha pleuralis Coquillett, 1898: 302.

Korean record: Monoceromyia pleuralis: ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998b: 129 (Korean cat.); Choi et al. 2000: 144 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 143 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 10 (Korean check.). Ceriana pleuralis: Kim, JI 1980: 387 (Korean check.). Sphyximorpha pleuralis: Doi 1938a: 10 (Korean check., presumably first Korean record); ZSK 1968: 180 (Korean check.); Kim, CW 1971: 884 (redescrip.); Kim, JI 1975: 42 (Korean check.).

Description: Body length 17‒18.5 mm. Male. Head a little broader than the thorax; frons blackish brown with a pair of large orange-yellow lateral spots, covered with short pile; antenna almost entirely dark brown, distinctly extended. Thorax black ground color; postpronotum yellowish brown; scutum subquadrate, a little longer than wide. Wing hyaline with anterior 1/3 dark brown tinge; halter whitish brown. Legs reddish brown ground color; basal half of femora dark brown; fore- and midtibia brown; hind tibia dark brown, more or less curved. Abdomen conspicuously longer than thorax and distinctly narrower than the thorax; tergite 2 black with orange yellow bands anteriorly and posteriorly, constricted in the middle; tergite 3 black with orange yellow band posterior margin; tergite 4 entirely black.

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곤충-알락긴꽃등에아과Ⅱ_영문 수정.indd 18 2018-09-12 오후 12:45:16 SYRPHIDAE: Primocerioides

Female. No specimens available.

Specimen examined: Korea: CB: Yeongdong-gun, Yonghwa-myeon, Minjujisan Recreational Forest, 21. v. 2011, DS Oh, 1♂. GG: Namyangju-si, Sudong-myeon, Mt. Cheonmasan, 30. v. 1982, HK Kim, 1♂. GN: Geochang-gun, Gajo-myeon, Seoggang-ri, JS Jeon, 6. vi. 1987, 1♂. GW: Wonju-si, Heungeop-myeon, Maeji- ri, Hoechon, 31. v. 1997. SM Shin & EJ Yeo, 1♂; Yonsei Univ. Campus, 12. vi. 1998, HW Byun, 1♂.

Distribution: Russian Far East, Korea, Japan [PA].

Biology: Unknown.

Korea: CB, GN, GG, GW.

Genus Primocerioides Shannon, 1927

Primocerioides Shannon, 1927: J. Wash. Acad. Sci., 17 (2): 41. TS: Cerioides petri Hervé-Bazin (OD).

3. Primocerioides petri (Hervé-Bazin, 1914) (Pl. 1C)

Cerioides petri Hervé-Bazin, 1914: 414.

Korean record: Primocerioides petri: Han & Choi 2001: 143 (Korean check., first Korean record); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 10 (Korean check.).

Description: Body length 14‒16 mm. Male. Head yellow brown ground color with entirely covered with very fine whitish grey dense pubescence; ocellar triangle reddish brown; antenna long; scape slender, reddish brown; pedicel short and thick, reddish brown; basoflagellomere black, the base broad but gradually narrowed apically. Thorax black ground color; scutum entirely black except postpronotum, transverse suture, lateral longitudinal vittae yellowish brown; scutellum entirely yellowish brown. Wing hyaline with anterior half dark brown tinge but cell C hyaline; halter yellowish brown. Legs reddish brown ground color; femora and tibiae entirely reddish brown; fore- and midtarsi yellowish brown; hind tarsi reddish brown. Abdomen conspicuously longer than head and thorax together; tergite 2 and 3 almost parallel-sided but, tergite 3 posteriorly slightly widened; tergite 2 black with 2 yellow brown lateral spots and yellow brown hind marginal band; tergite 3 with reddish brown marginal band. Female. Similar to male except for following non-genitalic characters: eye dichoptic.

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Specimen examined: Korea: GB: Cheongsong-gun, Budong-myeon, Mt. Juwangsan, 1. vi. 2000, DS Choi et al. 5♂1♀; Yeongju-si, Punggi-eup, Mt. Sobaeksan, 21. iv. 2000, DS Choi & SK Kim, 1♂. GW: Wonju-si, Gwirae-myeon, from Cheoneunsa to Mt. Sipjabong peak, 4. iv. 1998, DS Choi & SK Kim, 2♂; Heungeop- myeon, Maeji-ri, Yonsei Univ. Campus, 31. iii. 1999, SK Kim & MK Choi, 1♂. Distribution: Russian Far East, Korea, Japan [PA]. Biology: Unknown. Korea: GB, GW.

Genus Sphiximorpha Rondani, 1850

Sphiximorpha Rondani, 1850: Ann. Soc. Ent. France, (2) 8: 213. TS: Ceria subsessilis Illiger in Rossi (OD). Shambalia Violovitsh, 1981: Nasekomye i Kleshchi Sibiri (Novye i maloizvestnye vidy fauny Sibiri, 15): 85. TS: Shambalia rachmaninovi Violovitsh (OD).

4. Sphiximorpha rachmaninovi (Violovitsh, 1981) (Pls. 1D, E)

Shambalia rachmaninovi Violovitsh, 1981: 85.

Korean record: Sphiximorpha rachmanimovi: Suk et al. 2015: 260 (new Korean record).

Description: Body length 11.5‒14.3 mm. Male. Head reddish brown ground color; face brown with unique linear markings; eyes holoptic, bare; antenna long; scape slender, reddish brown; pedicel short and thick, dark brown; basoflagellomere yellowish brown, the base broad but gradually narrowed apically. Thorax brown ground color; scutum brown with three thick black longitudinal stripes with lateral stripes interrupted at transverse suture; scutellum black background with large widely semicircular yellow area medially and narrow brown posterior margin. Wing hyaline with anterior half dark brown tinge; halter whitish yellow. Legs reddish brown ground color; hind femur brown with dark brown tinge at posterobasal half. Abdomen conspicuously longer than thorax; abdominal tergite 2 constricted in the middle; tergite 3 and 4 each with pair of elongated yellowish spots (not clearly seen on some specimens due to the orientation as well as the body fluid oozed out). Female. Similar to male except for following non-genitalic characters: eye dichoptic.

Specimen examined: Korea: GG: Namyangju-si, Byeollae-myeon, Mt. Bulamsan, 3. iv. 1987, H. K. S., 1♂; Paju-si, Gwangtan-myeon, Bogwangsa, 25. iv. 1975, DI An, 1♂; Pocheon-si, Sohol-eup, Korea National

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곤충-알락긴꽃등에아과Ⅱ_영문 수정.indd 20 2018-09-12 오후 12:42:31 SYRPHIDAE: Cheilosia

Arboretum, 20. v. 1990, JH Gye, 1♀; , Seocho-gu, Mt. Cheonggyesan, 28. v. 1986, SH Ko, 1♂. GW: Wonju-si, Gwirae-myeon, from Cheoneunsa to Mt. Sipjabong peak, 5. v. 2004, HW Byun et al., 1♂; Wonju- si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 16. v. 1996, HW Byun, 1♂; ditto, 7. vi. 1998, DS Choi et al., 1♂; ditto, 5. v. 2002, HW Byun, 2♀; ditto, 16. v. 2004, HW Byun, 1♀; ditto, 29. v. 2004, HW Byun & HS Lee, 1♂; ditto, 16. v. 2006, DS Choi & HW Byun, 1♂; ditto, 23. vi. 2006, DS Choi, 1♂1♀; ditto, 24. v. 2009, DJ Cha & HS Lee, 1♀. Distribution: Russian Far East, Korea [PA]. Biology: Unknown. Korea: GG, GW.

Tribe Cheilosiini

Genus Cheilosia Meigen, 1822

Cheilosia Meigen, 1822: Syst. Beschr., 3: 296. TS: Eristalis scutellatus Fallén (des. Rondani 1856: Dipt. Ital. Prodromus, 1: 51, as Eristalis scutellatus Fabricius). Chilosia Agassiz, 1846: Nomencl. zool. ind. univ.: 80; unjustified emendation. Cartosyrphus Bigot, 1883: Annls Soc. ent. Fr., (6) 3: 230. TS: Syrphus paganus Meigen (des. Coquillett, 1910: Can.Ent., 42: 376). Nephomyia Matsumura, 1916: Thousand Ins. Japan, Add. 2: 220. TS: Nephomyia bombiformis Matsumura (OD). Stenocheilosia Matsumura, 1916: Thousand Ins. Japan, Add. 2: 242, Stenocheilosia isshikii Matsumura (OD). Taeniochilosia Oldenberg, 1916: Wien. ent. Ztg, 35 (3-4): 101 (as subgenus of Cheilosia Meigen). TS: Chilosiaatriseta Oldenberg (MO). Chaetochilosia Enderlein in Tierwelt Mitteleur., 6 (2), Insekten 3, 1936: 125 (as subgenus of Cheilosia Meigen). TS: Eristalis mutabilis Fallén (OD). Dasychilosia Enderlein in Tierwelt Mitteleur., 6 (2), Insekten 3, 1936: 125 (as subgenus of Cheilosia Meigen). TS: Syrphus variabilis Panzer (OD). Nigrocheilosia Shatalkin, 1975: Ent. Obozr., 54 (4): 901 (as subgenus of Cheilosia Meigen). TS: Eristalis pubera Zetterstedt (OD). Neocheilosia Barkalov, 1983: Diptera (Insecta), their systematics, geographical distribution and ecology, Leningrad: 6.TS: Chilosia scanica Ringdahl (OD). [For discussion of nomenclature problems see Wirth et al. 1965: 583].

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Key to species of Cheilosia from South Korea

1. Eyes densely pilose ······································································································ 2 − Eyes apparently bare ······································································ C. matsumurana (Shiraki) 2. Wing with brown spot in the middle ·················································································· 3 − Wing with brown apical spot ···································································C. abbreviata Shiraki 3. Scutum entirely covered with yellow brown hairs ·································································· 4 − Posterior 1/2 of scutum covered with black hairs ············································ C. illustrata (Harris) 4. Scutum densely with orange yellow hairs ···········································C. bombiformis (Matsumura) − Scutum with yellowish hairs ······················································ C. motodomariensis Matsumura 5. Legs entirely black ······································································································· 6 − Legs black with some yellow area ····················································································· 7 6. Face protrudes ventrally ············································································ C. facialis Shiraki − Face does not protrudes ventrally ····························································· C. fuscipennis Shiraki 7. Hind femur yellow ·················································································· C. aterrima (Sack) − Hind femur black ········································································································ 8 8. Face protrudes ventrally ········································································ C. longiptera Shiraki − Face does not protrudes ventrally ······························································ C. ochripes (Shiraki)

5. Cheilosia abbreviata Shiraki, 1953 (Pls. 1F, G)

Cheilosia abbreviata Shiraki, 1953: 87.

Korean record: Cheilosia abbreviata: Kim, CW 1971:860 (redescrip., first Korean record); Kim, JI 1975: 39 (Korean check.); Kim, JI 1980: 384 (Korean check.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 126 (Korean cat.); Han & Choi 2001: 111 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 10 (Korean check.).

Description: Body length 7‒11 mm. Male. Body entirely shiny black. Head brownish black with yellow and black hairs; vertex brownish black with yellow grey pruinosity, yellow and black hairs mixed; face black with yellow grey pruinosity; gena brownish black; eyes brownish black with densely yellow brown hairs; antennal scape and pedicel dark brown with hairs and laterally with black bristles; antennal basoflagellomere yellow brown, bare; arista brownish black, bare; length ratio of scape: pedicel: basoflagellomere (measured dorsally) approximately 1: 3: 6. Thorax brownish black with golden tinge; scutum yellow brown hairs and laterally with five bristles;

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scutellum brownish black with posteriorly with yellow grey pruinosity; pleura brownish black with yellow brown and black hairs. Wing mostly hyaline with apical dark area, entirely microtrichose; halter yellow brown, bare. Legs brownish black ground color with some yellow brown area, yellow brown and black hairs. Abdomen brownish black with ivory pruinosity; abdominal tergites with yellow brown hairs; sternites yellow brown hairs and short black hairs. Female. Similar to male except for following non-genitalic characters: eyes dichoptic; abdominal sternite with yellow brown hairs.

Specimen examined: Korea: GG: Gwangju-si, Docheok-myeon, Mt. Taehwasan, from Eungoksa to 612.8 m peak, 25. viii. 2006, HY Han et al., 1♀. GW: Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo- ri to 1,200 m peak, 3. v. 2003, DS Choi et al., 2♂; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, N. Valley, 22. vii. 2006, HY Han & KE Ro, 1♀; Hongcheon-gun, Nae-myeon, Mt. Gachilbong, from Sambongyaksu to 1,240 m peak, 2. v. 2003, HY Han & KE Ro, 1♀.

Distribution: Korea, Japan [PA].

Biology: Unknown.

Korea: GG, GW.

6. Cheilosia aterrima (Sack, 1927)

Chilosia aterrima Sack, 1927: 305.

Korean record: Cheilosia aterrima: Kim, JI & Kim, CW 1972: 8 (new Korean record); Kim, JI 1975: 39 (Korean check.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 126 (Korean cat.); Han & Choi 2001: 112 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 10 (Korean check.). Cheilosia atterima [misspelling]: Kim, JI 1980: 384 (Korean check.).

Distribution: Russian Far East, Korea, Japan, Oriental Region [PA, OR].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim and Kim (1972).

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7. Cheilosia bombiformis (Matsumura, 1916) (Pls. 1H, I)

Nephomyia bombiformis Matsumura, 1916: 220. Cheilosia sachtlebeni Stackelberg, 1963: 513.

Korean record: Cheilosia bombiformis: Kim, JI 1980: 384 (Korean check.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 126 (Korean cat.); Han & Choi 2001: 112 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 10 (Korean check.). Nephomyia bombiformis: Doi 1938a: 9 (Korean check., presumably first Korean record).

Description: Body length 12‒16 mm. Male. Body entirely shiny black. Head brownish black with yellow hairs; vertex brownish black with yellow grey pruinosity, yellow and black hairs mixed; face black with yellow grey pruinosity; gena brownish black with yellow pruinosity; eyes brownish black with yellow brown hairs dorsally; antennal scape and pedicel dark brown with black hairs; antennal basoflagellomere brown, bare; arista brownish black with short black hairs; length ratio of scape: pedicel: basoflagellomere (measured dorsally) approximately 3:4:6 Thorax brownish black; scutum brownish black with grey pruinosity anteriorly, yellow brown hairs; scutellum black with yellow brown hairs, yellow grey pruinositgy posteromarginally; pleura black, bare but some area yellow brown hairs. Wing mostly hyaline with dark area in the middle, entirely microtrichose; halter yellow brown, some specimens dark brown apically, bare. Legs brownish black ground color with some yellow brown area, yellow brown and black hairs. Abdomen brownish black with some area yellow grey pruinosity; abdominal tergite 1 with pale yellow hairs; tergite 2 with yellow brown hairs, some specimens mostly bare; tergite 3 with yellow brown hairs except on posterior area black hairs; tergite 4 with yellow brown hairs except on black hairs posterior 1/3; sternite 1 with yellow brown hairs; sternite 2 with yellow brown hairs and posterior 1/3 black hairs mixed; sternites 3‒5 with black hairs. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: GW: Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 3. ix. 2000, SK Kim et al., 1♂; Inje-gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 21. viii. 1998, HY Han et al., 1♂1♀; Jeongseon-gun, Nam-myeon, Mt. Mindungsan, form Yupyeong-ri to 1,119 m peak, 30. viii. 2000, HY Han et al., 1♂; -si, Mt. Taebaeksan, from Yuilsa to 1,560 m peak, 12. viii. 2000, DS Choi et al., 1♀.

Distribution: Korea, Japan [PA].

Biology: Unknown.

Korea: GW.

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8. Cheilosia facialis Shiraki, 1968

Cheilosia facialis Shiraki, 1968a: 141.

Korean record: Cheilosia facialis: Kim, JI 1980: 384 (Korean check., presumably first Korean record); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 126 (Korean cat.); Han & Choi 2001: 112 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 11 (Korean check.).

Distribution: Korea?, Japan [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim (1980).

9. Cheilosia fuscipennis Shiraki, 1953

Cheilosia fuscipennis Shiraki, 1953: 90.

Korean record: Cheilosia fuscipennis: Kim, CW 1971: 859 (redescrip., first Korean record); Kim, JI 1975: 39 (Korean check.); Kim, JI 1980: 384 (Korean check.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 126 (Korean cat.); Han & Choi 2001: 113 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 11 (Korean check.).

Distribution: Korea?, Japan [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim (1971).

10. Cheilosia illustrata (Harris, 1780) (Pls. 1J, K)

Musca illustrata Harris, 1780: 104. Musca fulva Gmelin, 1790: 2878. Syrphus rupestris Panzer, 1798: 13.

Korean record: Cheilosia illustrata: Shiraki 1968a: 92 (redescrip.); Kim, JI 1980: 384 (Korean check.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 127 (Korean cat.); Han & Choi 2001: 113 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 11 (Korean check.).

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Description: Body length 10‒14 mm. Male. Body entirely black ground color. Head brownish black with yellow hairs; vertex yellow brown with yellow grey pruinosity, brownish black hairs; face black with yellow grey pruinosity; gena brownish black with yellow grey pruinosity; eyes brownish black with yellow brown hairs dorsally; antennal scape and pedicel brown with black hairs; basoflagellomere brown, bare; arista brownish black with short black hairs; length ratio of scape: pedicel: basoflagellomere (measured dorsally) approximately 3:4:6 Thoraxic scutum brownish black with grey pruinosity anteriorly and laterally, yellow brown hairs; scutellum black with yellow grey pruinosity posteromarginally, yellow brown hairs; pleura black with yellow grey pruinosity, bare but some area yellow brown hairs. Wing mostly hyaline with dark area in the middle, entirely microtrichose; halter yellow brown, bare. Legs brownish black ground color with some dark brown area, yellow brown and black hairs. Abdominal tergite 1‒2 brownish black with some area yellow grey pruinosity; tergite 1 with pale yellow hairs; tergite 2 with yellow brown hairs, some specimens mostly bare; tergite 3‒5 brownish black; tergite 3 with yellow brown hairs except on posterior area black hairs; tergite 4 with yellow brown hairs except on black hairs posterior 1/3; sternite 1 with yellow brown hairs; sternite 2 with yellow brown hairs and posterior 1/3 black hairs mixed; sternites 3‒5 with black hairs. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: GN: Namhae-gun, Idong-myeon, 7. xi. 1998, JY Choi, 11♂3♀; Sancheong- gun, Sicheon-myeon, Mt. Jirisan, 30. vii. 1981, HY Han, 1♂. GW: Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 1. viii. 2006, HW Byun, 1♂; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 10. viii. 2009, DJ Cha & HS Lee, 1♂2♀; ditto, 5. viii. 2009, SW Suk et al., 1♂; ditto, 14. viii. 2009, SW Suk et al., 1♀.

Distribution: Europe, Russia to Korea, Japan [PA].

Biology: Unknown.

Korea: GN, GW.

11. Cheilosia longiptera Shiraki, 1968 (Pls. 1L, 2A)

Cheilosia longiptera Shiraki, 1968a: 103.

Korean record: Cheilosia longiptera: Peck 1988: 108 (Palaearctic cat.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 127 (Korean cat.); Han & Choi 2001: 114 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 11 (Korean check.). Cheilosia longiptear [misspelling]: Kim, JI et al. 1983: 206 (new Korean record).

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Description: Body length 9‒12 mm. Male. Body entirely black ground color. Head brownish black with yellowish brown hairs; vertex brownish black with yellow grey pruinosity, yellowish brown hairs; face black with yellow grey pruinosity; gena brown; eyes brownish black with yellow brown hairs densely; antennal scape and pedicel brownish black with yellowish brown hairs except on brownish black hairs laterally; basoflagellomere dark brown except for brown posteriorly, bare; arista brownish black with short black hairs; length ratio of scape: pedicel: basoflagellomere (measured dorsally) approximately 1:5:6 Thoraxic scutum brownish black with grey pruinosity anteriorly and laterally, yellow brown hairs, laterally with 2‒3 black bristles; scutellum brownish black with yellowish brown hairs, some specimens black hairs mixed; pleura black with yellow grey pruinosity, bare but some area yellow brown hairs. Wing mostly hyaline with entirely microtrichose; halter yellow brown, bare. Legs brownish black ground color with some dark brown area, yellowish brown and black hairs; femora brownish black; tibiae yellow brown basally, brownish black apically. Abdominal tergites brownish black with some area yellow grey pruinosity, yellow brown hairs, some specimens mostly bare; sternites with yellow brown hairs. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: GW: Jeongseon-gun, Nam-myeon, Mt. Mindungsan, form Yupyeong-ri to 1,119 m peak, 31. v. 2006, HY Han et al., 1♂2♀; ditto, 26. v. 2007, HY Han et al., 1♀; ditto, 27. v. 2009, HS Lee et al., 1♀; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 15. v. 2008, JM Jung, 1♂.

Distribution: Russian Far East, Korea, Japan [PA].

Biology: Unknown.

Korea: GW.

12. Cheilosia matsumurana (Shiraki, 1930)

Chilosia matsumurana Shiraki, 1930: 316. Cheilosia abdominalis Shiraki, 1968a: 74. Cheilosia moneronica Violovitsh, 1971: 109. Chilosia luteipes Shiraki, 1930: 319.

Korean record: Cheilosia matsumurana: Shiraki 1968a: 63 (redescrip.); Kim, JI 1980: 384 (Korean check.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 127 (Korean cat.); Han & Choi 2001: 114 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 11 (Korean check.). Cheilosia luteipes: Kim, CW 1971: 860 (redescrip.); Kim, JI 1975: 39 (Korean check.). Chilosia luteipes: Doi 1938a: 8 (Korean check., presumably first Korean record); Doi 1938b: 92 (Gaema-plateau); ZSK 1968: 179 (Korean

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check.).

Distribution: Russian Far East, Korea, Japan [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim (1971).

13. Cheilosia motodomariensis Matsumura, 1916 (Pl. 2B)

Cheilosia motodomariensis Matsumura, 1916: 239. Cheilosia portschinskiana Stackelberg, 1960: 441. Cheilosia subillustrata Stackelberg, 1963: 2. Cheilosia unicolor Shiraki, 1968a: 94. Chilosia magnificaHellen, 1930: 29. Chilosia unicolor Sack, 1941: 189.

Korean record: Cheilosia motodomariensis: Kim, CW 1971: 861 (redescrip.); Kim, JI 1975: 39 (Korean check.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 127 (Korean cat.); Han & Choi 2001: 114 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 11 (Korean check.). Chilosia motodomariensis: Doi 1938a: 8 (Korean check., presumably first Korean record); ZSK 1968: 179 (Korean check.).

Description: Body length 9‒11 mm. Male. Body entirely black ground color. Head brownish black with yellow hairs; vertex yellow brown with yellow grey pruinosity, brownish black hairs; face black with yellow grey pruinosity; gena brownish black with yellow grey pruinosity; eyes brownish black with yellow brown hairs densely; antennal scape and pedicel dark brown with black hairs; basoflagellomere yellow brown, bare; arista brownish black with short black hairs; length ratio of scape: pedicel: basoflagellomere (measured dorsally) approximately 3:4:8 Thoraxic scutum brownish black with golden tinge, yellow brown hairs; scutellum black with golden tinge, yellow brown hairs; pleura black with yellow grey pruinosity, bare but some area yellow brown hairs. Wing mostly hyaline with dark area in the middle, entirely microtrichose; halter yellow brown, bare. Legs brownish black ground color with some dark brown area, yellow brown and black hairs. Abdominal tergites brownish black with yellow brown hairs, some specimens mostly bare; sternites with yellow brown hairs. Female. No specimens available.

Specimen examined: Korea: GW: Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to

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1,200 m peak, 6. v. 2001, DS Choi & SK Kim, 3♂; ditto, 1. viii. 2006, HW Byun 1♂; Inje-gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 17. v. 1997, HY Han et al., 1♂.

Distribution: Siberia, Russian Far East, Mongolia, Korea, Japan [PA].

Biology: Unknown.

Korea: GW.

14. Cheilosia ochripes (Shiraki, 1930)

Chilosia ochripes Shiraki, 1930: 292.

Korean record: Cheilosia ochripes: Kim, JI & Kim, CW 1972: 9 (new Korean record); Kim, JI 1975: 39 (Korean check.); Kim, JI 1980: 384 (Korean check.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 128 (Korean cat.); Han & Choi 2001: 115 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 12 (Korean check.).

Distribution: Russian Far East, Korea, Japan [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim and Kim (1972).

Genus Ferdinandea Rondani, 1844

Ferdinandea Rondani, 1844: Nuovi Ann. Sci. nat. Inst. Bologna, (2) 2: 196. TS: Conops cuprea Scopoli (des. Rondani,1856: Dipt. Ital. Prodromus, 1: 51). Chrysoclamys Walker, 1851: Insecta Br., 1: 279 (unjustified new name for Ferdinandea Rondani). TS: Conops cuprea Scopoli. Chrysoclamis Rondani, 1856: 51 error. Chrysochlamys Rondani, 1857: 145 error. Chrysochlamis Schiner, 1857: 431 error.

Key to species of Ferdinandea from South Korea

1. Frons entirely shiny black ······································································ F. nigrifrons (Egger) − Frons entirely yellow brown ···························································································· 2 2. Abdomen mostly shiny brassy color; tibiae with black pilose ······························· F. cuprea (Scopoli)

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− Abdomen mostly shiny bluish-black color; tibiae without black pilose except hind tibia ······················· ··································································································· F. ruficornis(Fabricius)

15. Ferdinandea cuprea (Scopoli, 1763) (Pl. 2C)

Conops cuprea Scopoli, 1763: 355. Chrysochlamys nigrifrons Egger, 1860: 664. Dideoides eizoi Azuma, 2001: 460. Ferdinandea suzukii Matsumura, 1916: 224. Ferdinandea testacicornis Rondani, 1844. Musca nitens Villers, 1789: 549. Musca rutilo Harris, 1780: 80.

Korean record: Ferdinandea cuprea: ZSK 1968: 179 (Korean check.); Kim, CW 1971: 857 (redescrip.); Kim, JI 1975: 41 (Korean check.); Kim, JI 1980: 383 (Korean check.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998a: 10 (Yonsei Univ. Maeji Campus); Han et al. 1998b: 128 (Korean cat.); Choi et al. 2000: 143 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 116 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 12 (Korean check.).

Description: Body length 11.8‒14.3 mm. Male. Head broader than the thorax; vertical triangle shiny black with black pubescence; face shiny yellowish brown with abundant whitish dust near the eyes; eyes holoptic, dense pilose; frons entirely yellow brown; antennal scape and pedicel entirely reddish brown with black setulae; basoflagellomere oval; arista dark brown. Thorax with scutum subshiny black with 4 grey dust longitudinal stripes; scutellum brown with dense long pilose and black long setulae on hind margin. Wing hyaline with pale brownish tinge and dark brown maculae near bm-cu and r-m; halter with yellow brown stem and dark brown knob. Legs almost entirely yellow brown; coxae and trochanters dark brown. Abdomen moderately elongated with dense brown pilose. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: CB: Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 3. vi. 2004, HW Byun et al., 1♂. GB: Dalseong-gun, Yaga-myeon, Mt. Biseulsan, Recreational Forest, 20. v. 2012, DS Choi, 1♀. GN: Sancheong-gun, Sicheon-myeon, Mt. Jirisan, Seseoksanjang (1,458 m), Malaise trap, 26. vii. -11. x. 2001, DS Ku, 1♂. GG: Gapyeong-gun, Buk-myeon, Mt. Myeongjisan, 19. vi. 2006, DS Choi et al., 4♂1♀. GW: Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 9. viii. 2006, HW Byun

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et al., 2♂; Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 1. viii. 2006, HW Byun et al., 1♂; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 13. viii. 2004, OY Lim & HS Lee, 2♂; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 7. viii. 2004, HY Byun et al., 1♀; Pyeongchang-gun, Bangrim-myeon, Ungyo-ri, 29. vii. 2010, HS Lee et al., 1♀; Pyeongchang-gun, Jinbu-myeon, Mt. Odaesan, from Woljeongsa to Birobong (1,563 m) peak, 28. vii. 2004, DS Choi et al., 1♀; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 2. viii. 2000, DS Choi et al., 1♂.

Distribution: Europe, Russia, Korea, Japan [PA].

Biology: In Europe, of this species are found in sap of Populus alba attacked by Cossus cossus (Torp, 1994).

Korea: CB, GB, GN, GG, GW.

16. Ferdinandea nigrifrons (Egger, 1860) (Pl. 2D)

Chrysochlamys nigrifrons Egger, 1860: 664.

Korean record: Ferdinandea nigrifrons: Kim, JI et al. 1983: 204 (new Korean record); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 128 (Korean cat.); Han & Choi 2001: 117 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 12 (Korean check.).

Description: Body length 10.5‒12.5 mm. Male. Head broader than the thorax; vertical triangle shiny black with black pubescence; face shiny yellowish brown with abundant whitish dust near the eyes; eyes holoptic, dense pilose; frons entirely shiny black; antennal scape and pedicel entirely reddish brown with black setulae; basoflagellomere oval, dark brown; arista dark brown. Thorax with scutum subshiny black with 4 grey dust longitudinal stripes; scutellum brown with dense long pilose and black long setulae on hind margin. Wing hyaline with pale brownish tinge and dark brown maculae near bm-cu and r-m; halter with yellow brown stem and dark brown knob. Legs almost entirely yellow brown; coxae and trochanters dark brown; hind femur dark brown 3/4 basally. Abdomen moderately elongated with dense brown pilose. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: CB: Cheongwon-gun, Namseong-myeon, Mt. Ingyeongsan, 15. viii. 1997, HY Han et al., 2♂; Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 15. ix. 2006, HW Byun et al., 1♂. GW: Hwacheon-gun, Sanae-myeon, Mt. Gwangdeoksan, 20. vii. 1997, HY Han et al., 1♂; Wonju-si, Gwirae- myeon, from Cheoneunsa to Mt. Sipjabong peak, 18. viii. 1997, HY Han et al., 1♂; Wonju-si, Heungeop-

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myeon, Maeji-ri, Yonsei Univ. Campus, 14. vi. 1998, HW Byun et al., 1♂1♀; ditto, 11. viii. 1998, HW Byun & DS Choi, 1♀; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 18. ix. 2000, DS Choi et al., 1♂.

Distribution: Europe, Korea, Japan [PA].

Biology: In Korea, This species were often found on the trunk of Quercus spp. (Han and Choi, 2001).

Korea: CB, GW.

17. Ferdinandea ruficornis (Fabricius, 1775) (Pl. 2E)

Syrphus ruficornisFabricius, 1775: 769.

Korean record: Ferdinandea ruficornis: Han et al. 1998a: 10 (Yonsei Univ. Maeji Campus, first Korean record); Han et al. 1998b: 129 (Korean cat.); Choi et al. 2000: 143 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 117 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 12 (Korean check.).

Description: Body length 11.8‒13.2 mm. Male. Head broader than the thorax; vertical triangle shiny black with black pubescence; face shiny yellowish brown with abundant whitish dust near the eyes; eyes holoptic, dense pilose; frons entirely yellow brown; antennal scape and pedicel entirely reddish brown with black setulae; basoflagellomere oval; arista dark brown. Thorax with scutum subshiny black with 4 grey dust longitudinal stripes; scutellum brown with dense long pilose and black long setulae on hind margin. Wing hyaline with pale brownish tinge and dark brown maculae near bm-cu and r-m; halter with yellow brown stem and dark brown knob. Legs almost entirely yellow brown; coxae and trochanters dark brown; hind femur dark brown 2/3 basally. Abdomen moderately elongated with dense brown pilose. Female. Similar to male except for following non-genitalic characters: eyes dichoptic; hind femur entirely yellow brown.

Specimen examined: Korea: CN: Boryeong-si, Cheongna-myeon, Mt. Oseosan, 20. vii. 1999, HY Han et al., 1♀. GG: Namyangju-si, Sudong-myeon, Mt. Cheonmasan, 4. ix. 1965, YC Jeon, 1♀. GW: Hoengseong- gun, Anheung-myeon, Mt. Chiaksan, from Gangrim 4-ri to Cheonjibong (1,086 m) peak, 16. v. 1999, DS Choi et al., 2♂; Hongcheon-gun, Nae-myeon, Mt. Gachilbong, 22. v. 1999, HY Han & KE Ro, 7♂; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 5. vi. 1999, DS Choi et al., 1♂; Wonju-si, Gwirae-myeon, from Cheoneunsa to Mt. Sipjabong peak, 19. iv. 1998, DS Choi & SK Kim, 1♂; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 15. v. 1996, SH Rhee, 1♂. JN:

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Gwangju, Dong-gu, Mt. Mudeungsan, 13. v. 2000, DS Choi et al., 1♂.

Distribution: Europe, Russia, N. China, Korea [PA].

Biology: In Europe, pupa of this species was found in soil at the foot of a Populus alba attacked by Cossus sp. (Torp, 1994).

Korea: CN, GG, GW, JN.

Genus Rhingia Scopoli, 1763

Rhingia Scopoli, 1763: Entom. carniolica: 358. TS: Conops rostrata Linnaeus (MO).

18. Rhingia laevigata Loew, 1858 (Pl. 2F)

Rhingia laevigata Loew, 1858: 107. Rhingia apicalis Matsumura, 1905: 97.

Korean record: Rhingia laevigata: Doi 1938a: 9 (Korean check., presumably first Korean record); Doi 1938b: 92 (Gaema-plateau); ZSK 1968: 180 (Korean check.); Kim, CW 1971: 857 (redescrip.); Kim, JI 1975: 39 (Korean check.); Kim, JI 1980: 384 (Korean check.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 129 (Korean cat.); Choi et al. 2000: 143 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 118 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 13 (Korean check.).

Description: Body length 7.2‒9.5 mm. Male. Head slightly broader than thorax; frons shortly produced, covered with grayish tomentum on the eye-margins; eyes bare; eye contiguity about twice as long as vertical triangle; face entirely yellow brown with lower portion distinctly protrude anteriorly with sharp apex; antenna brownish yellow; scape about as long as thick; pedicel longer than scape. Thorax about as wide as long, shiny black; scutum shiny black with dense yellow brown pilose; scutellum shiny black with dense long black pilose. Wing hyaline with yellow brown tinge; halter yellow brown. Legs yellowish brown ground color except black basal half of femora, middle of hind tibia, hind tarsus. Abdomen shorter than the head and thorax together; tergite 1 yellowish brown with black in hind margin; tergite 2 entirely yellow brown with black spots in middle and black thin band on hind margin; tergite 3 yellow brown anteriorly and black laterally, hind margin; tergite 4 almost entirely shiny black. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

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Specimen examined: Korea: GB: Cheongsong-gun, Budong-myeon, Mt. Juwangsan, 6. vi. 1989, EM Park, 1♀; Uljin-gun, Onjeong-ri, Mt. Baegamsan, 16. viii. 1999, HY Han et al., 2♂. GW: Goseong-gun, Geojin- eup, Mt. Geonbongsan, 18. ix. 1967, JK Oh, 1♀; Hoengseong-gun, Anheung-myeon, Mt. Chiaksan, from Gangrim 4-ri to Cheonjibong (1,086 m) peak, 16. v. 1999, HY Han & KE Ro, 11♂; Hongcheon-gun, Nae- myeon, Mt. Gachilbong, 21. vi. 1984, SS Hwang, 1♂; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 16. vi. 1998, HY Han et al., 1♂; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, N. Valley, 14. viii. 1998, HY Han & KE Ro, 1♀; ditto, 20. viii. 2007, HS Lee et al., 1♀; Hwacheon-gun, Sanae-myeon, Mt. Gwangdeoksan, 29. viii. 2000, CH Park & DW Kim, 3♂; Inje-gun, Buk- myeon, Mt. Maebongsan, E. Valley, 25. viii. 1998, HY Han et al., 1♂; Taebaek-si, Mt. Taebaeksan, from Yuilsa to 1,560 m peak, 29. viii. 1999, DS Choi & CH Park, 1♂1♀; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 3. v. 1998, HY Han et al., 1♂1♀.

Distribution: Russian Far East to Korea, Japan [PA].

Biology: Unknown.

Korea: GB, GW.

Tribe Chrysogasterini

Genus Brachyopa Meigen, 1822

Brachyopa Meigen, 1822: Syst. Beschr., 3: 260. TS: Musca conica Panzer (des. Westwood 1840: Synopsis: 137) [=panzeri Goffe].

19. Brachyopa testacea (Fallén, 1817) (Pl. 2G)

Rhingia testacea Fallén, 1817: 34.

Korean record: Brachyopa testacea: Han & Choi 2001: 133 (Korean check., first Korean record); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 13 (Korean check.).

Description: Body length 6.8‒7.5 mm. Male. Head yellowish brown ground color; frons yellowish brown; vertical triangle dark brown; face entirely yellow brown with lower portion distinctly protrude anteriorly with blunt apex; lunule shiny yellow brown; eyes holoptic, bare; antenna yellowish brown; basoflagellomere oval; arista brown. Thorax: scutum entirely black except lateral margins with pair of dust gray submedial longitudinal vittae; scutellum brown

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with yellow brown pilose. Wing hyaline with yellow brown tinge; halter whitish yellow. Legs almost entirely yellow brown except hind tarsus. Abdomen whitish yellow ground color; tergite 2‒4 each with dark median longitudinal band and hind marginal band. Female. No specimens available.

Specimen examined: Korea: GG: Gapyeong-gun, Buk-myeon, Mt. Myeongjisan, 19. vi. 2006, DS Choi et al., 1♂. GW: Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 20. vi. 1999, DS Choi & SK Kim, 2♂. JJ: Bukjeju-gun, Jocheon-eup, Goepyeongi oreum, 2. v. 2003, DS Choi et al., 1♂.

Distribution: Europe, Russia, Mongolia, Korea [PA].

Biology: Unknown.

Korea: GG, GW, JJ.

Genus Orthonevra Macquart, 1829

Orthonevra Macquart, 1829: Mém. Soc. Sci. Agric. Lille, 1827-1828: 188 and Ins. Dipt., 4: 40. TS: elegans Meigen (MO). Orthoneura Loew, 1843: Stettin. ent. Ztg. 4 (7): 207; unjustified emendation. Campeneura Rondani, 1856: Dipt. Ital. Prodromus, 1: 52. TS: Campeneura venusta Rondani (MO) [= frontalis (Loew)]. Camponeura Verrall in Scudder, Nomencl. Zool.: 1882: 58; unjustified emendation.

20. Orthonevra karumaiensis (Matsumura, 1916) (Pls. 2H, I)

Pipizella karumaiensis Matsumura, 1916: 16. Orthoneura ussuriana Stackelberg, 1929: 248.

Korean record: Orthonevra karumaiensis: Kim, JI et al. 1983: 205 (new Korean record); Peck 1988: 141 (Palaearctic cat.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 132 (Korean cat.); Han & Choi 2001: 133 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 13 (Korean check.).

Description: Body length 5.2‒6.2 mm. Male. Head moderately broader than thorax; frons slightly produced, about 1/3 as long as wide, shiny greenish black; vertical triangle about 1/5 as wide as head, slightly longer than wide, shiny aeneous black,

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clothed with long black brown pubescence; face brightly shiny green; eyes holoptic, bare; eye contiguity shorter than vertical triangle; antenna yellowish brown ground color; scape nearly twice as long as thick with some blackish setae; pedicel about twice as long as scape; basoflagellomere about as long as scape and pedicel together. Thorax shiny greenish black ground color; scutum bright shiny greenish black with five aeneous brown longitudinal vittae; scutellum about 1.5 times as wide as long. Wing hyaline with dark tinge vein r-m and M1; halter whitish yellow. Legs yellowish ground color except black femora and middle of tibiae. Abdomen oval, shorter than head and thorax together, entirely bright shiny greenish black. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: CB: Cheongwon-gun, Namseong-myeon, Mt. Ingyeongsan, 15. viii. 1997, HY Han et al., 1♀. CN: Cheonan-si, Byeongcheon-myeon, Maeseong-ri, 21. viii. 2006, DS Choi et al., 2♀; Taean-gun, Gonam-myeon, Janggok-ri, Haebyeonarae beach, 24. v. 2014, DS Choi, 1♀. GB: Gyeongju-si, Naenam-myeon, Gwaneumsa, 4. vi. 2011, DS Choi, 3♂. GN: Ulsan, Ulju-gun, Onsan-eup, Deoksin-ri, 14. vi. 2008, DS Choi, 1♀. GG: Namyangju-si, Sudong-myeon, Mt. Chungnyeongsan, 6. ix. 1980, HK Park, 1♂1♀; Paju-si, Tanhyeon-myeon, Daedong-ri, 25. vii. 2006, DS Choi et al., 1♂; Yangju-si, Gwangjeok-myeon, Ugo- ri, 2. viii. 2006, DS Choi et al., 1♂. GW: Cheorwon-gun, Naedong-ri, 22. viii. 1967, JK Oh, 1♂; Wonju- si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 7. viii. 1998, HW Byun & DS Choi, 1♀; ditto, 1. v. 2006, DS Choi & HW Byun, 1♂; ditto, 30. vi. 2006, HW Byun & JS Lim, 2♂1♀; Wonju-si, Buron-myeon, Jungsan-ri, 25. vii. 1998, MK Choi, 1♂. JN: Wando-gun, Yaksan-myeon, Is. Joyakdo, 15. vii. 2003. KD Han, 1♀.

Distribution: Russian Far East, Korea, Japan [PA].

Biology: Unknown.

Korea: CB, CN, GB, GN, GG, GW, JN.

Genus Sphegina Meigen, 1822

Sphegina Meigen, 1822: Syst. Beschr., 3: 193. TS: Milesia clunipes Fallén (des. Westwood, 1840: Synopsis: 136). Asiosphegina Stackelberg in Knutson, Thompson and Vockeroth, 1975: Cat. Dipt. Orient. Reg., 2: 338. TS: Spheginasibirica Stackelberg (OD).

21. Sphegina clunipes (Fallén, 1816) (Pls. 2J, K)

Milesia clunipes Fallén, 1816: 12.

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Sphegina flavaMacquart, 1834: 577. Sphegina nigricornis Macquart, 1829: 166.

Korean record: Sphegina clunipes: Hokuryukan 1965: 214 (redescrip., presumably first Korean record); Kim, JI 1975: 39 (Korean check.); Kim, JI 1980: 385 (Korean check.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998a: 10 (Yonsei Univ. Maeji Campus); Han et al. 1998b: 133 (Korean cat.); Choi et al. 2000: 144 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 134 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 13 (Korean check.).

Description: Body length 5.8‒7.2 mm. Male. Head brown ground color; frons shiny black; face dark brown with yellowish brown ventrally; Antenna yellow to yellowish brown; eyes holoptic, bare; arista brown, microscopic pubescent. Thorax almost entirely shiny black; scutum with pair of dusted whitish longitudinal vittae; scutellum shiny black with 2 long yellow setulae. Wing hyaline with brown tinge; halter yellowish brown. Legs; fore and midleg almost entirely yellowish brown; hind femur shiny black except basal 1/4 yellowish brown, distinctly swollen, some black spine ventrally; apical 1/2 of hind tibia and hind tarsus dark brown. Abdomen shiny black ground color, longer than head and thorax together; tergite 2 shiny black; tergite 3 shiny black with pair of brown spots on anterior lateral margin; tergite 4 shiny black. Female. Similar to male except for following non-genitalic characters: eyes dichoptic; abdominal tergite 3 and 4 about 5 times as wide as middle of tergite 2 medially.

Specimen examined: Korea: GB: Yeongju-si, Buseok-myeon, Mt. Seondalsan, from namdae-ri to 1,236 m peak, 29. vi. 1998, HY Han et al., 1♂. GN: Geochang-gun, Buksang-myeon, Mt. Deokyusan, Songgyesa, 7. vii. 2012, DS Choi, 1♀; ditto, Wolseong Valley, 8. vii. 2012, DS Choi, 1♂. GW: Hongcheon-gun, Nae- myeon, Mt. Gachilbong, 9. v. 1998, HY Han & KE Ro, 1♂; ditto, 3. vii. 2000, HY Han et al., 3♂1♀; Inje- gun, Girin-myeon, Mt. Bangtaesan, 6. vii. 2000, HY Han et al., 1♀; Wonju-si, Heungeop-myeon, Maeji-ri, Hoechon, 21. vii. 1997, HW Byun & DS Choi, 1♀; ditto, 23. v. 1998, HW Byun et al., 1♂1♀; ditto, 23. vi. 1998, HY Han & DS Choi, 1♀; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 18. v. 1998, DS Choi & SK Kim, 1♂; ditto, 30. v. 1998, HY Han & DS Choi, 1♀. JB: Jangsu-gun, Gyenam- myeon, Mt. Jangansan, 2. vi. 2013, DS Choi, 2♂; Sunchang-gun, Paldeok-myeon, Mt. Gangcheonsan, 25. v. 2013, DS Choi, 1♀.

Distribution: Europe, Russia, Korea, Japan [PA].

Biology: Unknown.

Korea: GB, GN, GW, JB.

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22. Sphegina japonica Shiraki and Edashige, 1953

Sphegina japonica Shiraki and Edashige, 1953: 108. Sphegina macrocerca Stackelberg, 1956: 938.

Korean record: Sphegina japonica: Han et al. 1998b: 133 (Korean cat.); Han & Choi 2001: 135 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 13 (Korean check.).

Distribution: Russian Far East, Korea, Japan [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Han et al. (1998b).

Tribe Eristalini

Genus Eristalinus Rondani, 1845

Eristalinus Rondani, 1845: Nuovi Ann. Sci. nat. Ist. Bologna, (2) 2: 453 (as subgenus of Eristalis Latreille). TS: Musca sepulchralis Linnaeus (des. Rondani 1857: Dipt. Ital. Prodromus, 2: 38). Metalloeristalis Kanervo, 1938: Annls Univ. turku., (A) 6 (4): 13. TS: Conops aeneus Scopoli (OD). Eristalodes Mik, 1897: Wien. ent. Ztg, 16(2): 114 (as genus). TS: Eristalis taeniops Wiedemann (OD). Lathyrophthalmus Mik, 1897: Wien. ent. Ztg, 16(2): 114 (as genus). TS: Conops aeneus Scopoli (OD).

Key to species of Eristalinus from South Korea

1. Eyes meeting on frons (male) ·························································································· 2 − Eyes separated on frons (female) ······················································································ 4 2. Eye contiguity distinctly longer than vertical triangle length ························ E. tarsalis (Macquart) (♂) − Eye contiguity short than vertical triangle length ··································································· 3 3. Body entirely metallic green ······························································ E. viridis (Coquillett) (♂) − Body entirely shiny aeneous black ························································· E. aeneus (Scopoli) (♂) 4. Abdominal tergite 2‒4 each with distinct dusted whitish bands ····················· E. tarsalis (Macquart) (♀) − Abdominal tergite 2‒4 each without any bands ····································································· 4 5. Body entirely metallic green ································································ E. aeneus (Scopoli) (♀) − Body entirely shiny aeneous black ······················································ E. viridis (Coquillett) (♀)

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23. Eristalinus aeneus (Scopoli, 1763) (Pls. 3A, B)

Conops aeneus Scopoli, 1763: 356. Conops stygius Newman, 1835: 313. Eristalis aenescens Macquart, 1842: 119. Eristalis concolor Philippi, 1865: 743. Eristalis cuprovittatus Wiedemann, 1830: 190. Eristalis sincerus Harris, 1841: 409. Eristalis sincerus Walker, 1849: 611. Eristalis taphicus Wiedemann, 1830: 191. Eristalomyia auricalcia Rondani, 1865: 129. Lathyrophthalmus nigrolineatus Hervé-Bazin, 1923: 134. Musca leucocephala Gmelin, 1790: 2878. Musca macrophthalma Preyssler, 1791: 68. Musca ochroleuca Gmelin, 1790: 2878. Musca punctata Müller, 1764: 85. Syrphus aeneus Fabricius, 1794: 302.

Korean record: Eristalinus aeneus: Knutson et al. 1975: 347 (Orient. cat., presumably first Korean record); Han et al. 1998b: 137 (Korean cat.); Han & Choi 2001: 145 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 14 (Korean check.).

Description: Body length 11.2‒12.8 mm. Male. Body entirely shiny aeneous black. Head entirely shiny black; vertical triangle black with pale yellow pilose; face entirely shiny black, covered dense silver pilose; antennal scape dark brown; pedicel brown; basoflagellomere oval, dark brown dorsally; eyes holoptic, bare; eye contiguity shorter than vertical triangle length. Thorax shiny aeneous black ground color; scutum densely covered with yellowish brown pile, with 5 thin silver grey longitudinal vittae; scutellum with dense yellowish brown pilose. Wing hyaline; halter yellowish brown. Legs: coxae and trochanters black; femora entirely black except apical 1/5 yellowish brown; fore and midtibia apical 1/2 black and basal 1/2 yellowish brown; hind tibia apical 2/3 black and basal 1/3 yellowish brown. Abdomen moderately elongated, aeneous black ground color, densely covered with yellowish brown pile. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: CN: Hongseong-gun, Hongseong-eup, Mt. Baegwolsan, 16. viii. 1997, HY Han et al., 1♀. GB: Gimcheon-si, Jirye-myeon, Gyo-ri, near Gyorigyo, 3. x. 2016, DS Choi, 3♂5♀.

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GG: Paju-si, Paju-eup, Bongam-ri, 25. vii. 2006, DS Choi et al., 5♂; Pocheon-si, Ildong-myeon, Gisan-ri, Daedeogsa, 23. vii. 2006, DS Choi et al., 1♀. JB: Iksan-si, Sin-dong, Wongwang Univ. Campus, 30. x. 1999, HY Han et al., 5♂.

Distribution: Holarctic, Oriental, Australian Regions [PA, NE, OR, AU]. Biology: In Europe, this species occurs along shorelines in rock pools containing large amounts of decaying seaweed (Rotheray, 1993).

Korea: CN, GB, GG, JB.

24. Eristalinus quinquestriatus (Fabricius, 1749)

Syrphus quinquestriatus Fabricius, 1794: 289. Eristalis aesepus Walker, 1849: 625. Eristalomyia picta Bigot, 1880: 219. Lathyrophthalmus basalis Shiraki, 1968b: 175.

Korean record: Eristalinus quinquestriatus: ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998b: 137 (Korean cat.); Han & Choi 2001: 145 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 14 (Korean check.). Eristalis quinquestriatus: Kim, JI 1980: 387 (Korean check.). Lathyrophthalmus quinquestriatus: Kim, CW 1971: 870 (redescrip., first Korean record); Kim, JI 1975: 41 (Korean check.).

Distribution: China, Korea, Japan, Oriental Region [PA, OR].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim (1971).

25. Eristalinus sepulchralis (Linnaeus, 1957)

Eristalinus riki Violovitsh, 1957: 752. Eristalis impunctata Strobl, 1910: 106. Eristalis miki Mutin and Barkalov, 1999: 451. Musca ater Harris, 1776: 58. Musca melanius Harris, 1776: 59. Musca sepulchralis Linnaeus, 1758: 596. Syrphus tristis Fabricius, 1794: 303.

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Korean record: Eristalinus sepulchralis: Han & Choi 2001: 144 (Korean check., first Korean record); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 14 (Korean check.).

Distribution: Widespread in Palaearctic and Oriental Regions [PA, OR].

Biology: In Europe, this species occurs in accumulations of decaying vegetation in ponds, pools and marshes and also in wet manure (Rotheray, 1993). Remarks: We could not find specimens but cited this species based on Han and Choi (2001).

26. Eristalinus tarsalis (Macquart, 1855) (Pls. 3C, D)

Eristalis tarsalis Macquart, 1855: 107. Eristalis ocularius Coquillett, 1898: 325.

Korean record: Eristalinus tarsalis: Knutson et al. 1975: 349 (Orient. cat.); Peck 1988: 184 (Palaearctic cat.); ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998a: 11 (Yonsei Univ. Maeji Campus); Han et al. 1998b: 138 (Korean cat.); Choi et al. 2000: 144 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 146 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 15 (Korean check.). Eristalis ocularis: Shiraki 1930: 138 (redescrip.); Kim, JI 1980: 387 (Korean check.). Eristalis ocularius [misspelling]: Doi 1938a: 9 (Korean check.). Lathyrophthalmus ocularis: Shiraki 1968b: 182 (redescrip.); Kim, CW 1971: 869 (redescrip.); Kim, JI 1975: 41 (Korean check.). Lathyrophtalmus ocularis [misspelling]: Kim, CW 1980: 279 (distr. Map).

Description: Body length 10.1‒12.2 mm. Male. Body entirely black ground color. Head entirely shiny black; vertical triangle black with black pilose; face entirely shiny black, covered dense black and brown pile mixed; antennal scape and pedicel dark brown; basoflagellomere oval, dark brown dorsally; eyes holoptic, pilose; eye contiguity distinctly longer than vertical triangle length. Thorax shiny black ground color; scutum densely covered with yellowish brown pile; scutellum with dense yellowish brown pilose. Wing hyaline; halter yellowish brown. Legs: coxae and trochanters black; femora entirely black except apical 1/5 yellowish brown; fore and midtibia apical 1/2 black and basal 1/2 yellowish brown; hind tibia apical 2/3 black and basal 1/3 yellowish brown. Abdomen moderately elongated, black ground color, densely covered with yellowish brown pile. Female. Similar to male except for following non-genitalic characters: eyes dichoptic; scutum with 5 thin silver grey longitudinal vittae; abdominal tergite 2‒4 each pair of undulationg band.

Specimen examined: Korea: CB: Cheongwon-gun, Namseong-myeon, Mt. Ingyeongsan, 15. viii. 1997, HY

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Han et al., 5♂5♀; Chungju-si, Sotae-myeon, Deokeun-ri, Namhangang river side, 11. x. 2002, DS Choi et al., 2♂3♀. CN: Boryeong-si, Cheongna-myeon, Mt. Oseosan, 20. vii. 1999, HY Han et al., 2♂2♀; Cheonan- si, Byeongcheon-myeon, Byeongcheon-ri, Sangnok Resort, 6. xi. 1999, HY Han et al., 1♂1♀; Seosan- si, Daesan-eup, Daesan-ri, Mt. Mangwolsan, 9. viii. 2006, DS Choi et al., 1♀. GB: Gyeongju-si, Naenam- myeon, Gwaneumsa, 6. x. 2011, DS Choi, 2♀; Yeongju-si, -myeon, Mt. Sobaeksan, 27. viii. 2001, DS Choi et al., 1♀. GN: Hapcheon-gun, Gaya-myeon, Haeinsa, 9. x. 2009, DS Choi, 1♀. GG: Gapyeong- gun, Buk-myeon, Hwaak-ri, 3. vii. 2006, DS Choi et al., 1♀; Yangju-si, Goam-dong, 2. viii. 2006, DS Choi et al., 3♀; Yeoju-gun, Yeoju-eup, Sinjin-ri, Namhangang river side, 12. x. 2002, DS Choi & MH Lim, 6♂2♀. GW: Hongcheon-gun, Nae-myeon, Bangnae-ri, Mt. Maenghyeonsan, 12. x. 2002, HY Han & KE Ro, 2♂1♀; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, N. Valley, 14. viii. 1998, HY Han & KE Ro, 1♀; Inje-gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 21. viii. 1998, HY Han et al., 14♂20♀; Jeongseon-gun, Nam-myeon, Mt. Mindungsan, form Yupyeong-ri to 1,119 m peak, 3. x. 2005, HY Han et al., 1♀; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 26. ix. 2007, HS Lee et al., 4♂3♀. JB: Iksan-si, Sin-dong, Wongwang Univ. Campus, 30. x. 1999, HY Han et al., 1♀; Wanju-gun, Unju-myeon, Mt. Daedunsan, 1. x. 2000, DS Choi & SK Kim, 8♀. JN: Gurye-gun, Gwangui-myeon, Mt. Jirisan, Nogodan, 11. ix. 1998, HY Han & KE Ro, 1♂1♀; Jangheung-gun, Gwansan-eup, Mt. Cheongwansan, 14. v. 2000, DS Choi et al., 6♂.

Distribution: China, Korea, Japan, Oriental Region [PA, OR].

Biology: Unknown.

Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

27. Eristalinus viridis (Coquillett, 1898) (Pls. 3E, F)

Eristalis viridis Coquillett, 1898: 326.

Korean record: Eristalinus viridis: ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998a: 11 (Yonsei Univ. Maeji Campus); Han et al. 1998b: 138 (Korean cat.); Choi et al. 2000: 144 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 146 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 15 (Korean check.). Eristalis viridis: Doi 1938a: 9 (Korean check.); Kim, JI 1980: 387 (Korean check.). Lathyrophthalmus viridis: Kim, CW 1971: 870 (redescrip.); Kim, JI 1975: 41 (Korean check.). Lathyrophthamus viridis [misspelling]: ZSK 1968: 179 (Korean check.).

Description: Body length 10.5‒12.3 mm. Male. Body entirely metallic green. Head entirely shiny black; vertical triangle black with black pilose; face entirely shiny black, covered dense silver white pilose; antennal scape and pedicel yellowish brown;

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basoflagellomere oval, dark brown dorsally; eyes holoptic, sparsely pilose; eye contiguity shorter than vertical triangle length. Thorax shiny black ground color; scutum densely covered with yellowish brown pile, with 5 metalic green longitudinal vittae; scutellum with dense yellowish brown pilose. Wing hyaline; halter yellowish brown. Legs: coxae and trochanters black; femora entirely black except apical 1/5 yellowish brown; fore and midtibia apical 1/2 black and basal 1/2 yellowish brown; hind tibia apical 2/3 black and basal 1/3 yellowish brown. Abdomen moderately elongated, shiny black ground color, densely covered with yellowish brown pile; tergite 2‒3 each pair of triangular band. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: CB: Jincheon-gun, Iwol-myeon, Naechon-ri, 21. viii. 2006, DS Choi et al., 1♂1♀. CN: Dangjin-gun, Sinpyeong-myeon, Geumcheon-ri, 10. viii. 2006, DS Choi et al., 1♂; Nonsan-si, Yeonsan-myeon, Gojeong-ri, Gyebaekjanggunmyo, 15. vi. 2010, SJ Park, 1♂. GB: Gyeongju-si, Naenam- myeon, Gwaneumsa, 25. ix. 2009, DS Choi, 1♂; Yeongcheon-si, Hwasan-myeon, Yeonjeong-ri, Yeonji Reservoir, 22. vii. 2001, DS Choi, 4♂; Yeongju-si, Sunheung-myeon, Mt. Sobaeksan, 11. vi. 2011, DS Choi, 1♂. GG: Paju-si, Papyeong-myeon, Masan-ri, 1. viii. 2006, DS Choi et al, 1♀; Yangju-si, Goam-dong, 2. viii. 2006, DS Choi et al., 1♂2♀. GW: Donghae-si, Bukpyeong, Samhwasa, 27. vi. 1984, SH Jang, 1♀; Gangneung-si, Jeo-dong, near Gyeongpoho, 18. vi. 2007, HS Lee et al., 2♀; Hongcheon-gun, Hwachon- myeon, Seongsan-ri, 2. vi. 2006, DS Choi et al., 2♂; Wonju-si, Buron-myeon, Beopcheon-ri, Namhangang river side, 20. viii. 2002, DS Choi et al., 1♂; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 19.vi.2006, 1♂; ditto, 11. vii. 2006, SMR Hwang, 2♀; ditto, 19. viii. 2008, JS Lim et al., 1♂. JB: Gimje-si, Hwangsan-dong, Munsusa Temple, 17. vi. 2011, DS Choi, 1♀; Iksan-si, Sin-dong, Wongwang Univ. Campus, 30. x. 1999, HY Han et al., 1♂; Imsil-gun, Imsil-eup, Jeongwol-ri, Jeongwol reservoir, 6. Vii. 2013, DS Choi, 1♂. JN: Gokseong-gun, Gokseong-eup, Wolpyeong-ri, 15. v. 2007, DS Choi, 1♀; Jangseong-gun, Bukil- myeon, Bangjangsan Recreational Forest, 25. viii. 2013, DS Choi, 1♀.

Distribution: China, Korea, Japan [PA].

Biology: Unknown.

Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

Genus Eristalis Latreille, 1804

Eristalis Latreille, 1804: Nouv. dict. hist. nat., 24 (Tabl. méth. Ins.): 194. TS: Musca tenax Linnaeus (des. Curtis, 1832: Brit. Ent. 9: pl. 432). Tubifera Meigen, 1800: Nouve. class.: 34. TS: Musca tenax Linnaeus (des. Coquillett 1910: 618). Suppressed by I.C.Z.N., 1963: Opinion 678: 339. Elophilus Meigen, 1803: Mag. Insektenk., 2: 274. TS: Musca tenax Linnaeus (des. Latreille, 1810: Consid.

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Générales nat. animaux: 443, as Eristalis tenax Fabricius). Helophilus Leach, 1817 in Brewster’s Edinb. Encycl., 12 (1): 159; unjustified emendation Eristaloides Rondani, 1845: Nuovi Ann Sci. nat. Ist. Bologna, (2) 2 (1845): 453(as subgenus of Eristalis Latreille). TS: Musca tenax Linnaeus (des. Coquillett, 1910: Proc. U.S. natn. Mus., 37: 540). Eristalomya Rondani, 1857: dipt. Ital. Prodromus, 2: 38. TS: Musca tenax Linnaeus (OD). Eristalomyia Verrall in Scudder, 1882: Nomencl. Zool.: 127; unjustified emendation. Eriops Lioy, 1864: Atti Ist. veneto Sci., (3) 9: 743. A junior homonym of Eriops Klug. Eoseristalis Kanervo, 1938: Annls Univ. turku., (A) 6 (4): 12. TS: Eristalis cerealis Fabricius (OD). [For discussion of nomenclature problem see Wirth et al., 1965: 622].

Key to species of Eristalis from South Korea

1. Scutum with vertical grey band ························································································ 2 − Scutum without vertical grey band ···················································································· 3 2. Abdominal tergite 2 with pair of yellow brown bands attached posterior margin in male; tergite 2 with pair of triangular bands in female ····································································E. kyokoae (Kimura) − Abdominal tergite 2 with pair of yellow brown bands separated posterior margin in male; tergite 2 with pair of vertical bands in female ································································· E. cerealis Fabricius 3. Abdominal tergite 3 entirely black ·······················································E. arbustorum (Linnaeus) − Abdominal tergite 3 with brown band ·························································· E. tenax (Linnaeus)

28. Eristalis arbustorum (Linnaeus, 1758) (Pls. 3G, H)

Musca arbustorum Linnaeus, 1758: 591. Eristalis bulgarica Szilady, 1934: 147. Eristalis polonica Szilady, 1934: 151. Eristalis sachalinensis Matsumura, 1916: 263. Eristalis strandi Duda, 1940: 225. Eristalomya distincta Shiraki, 1968b: 166. Musca lyra Harris, 1776: 42. Musca nemorum Linnaeus, 1758: 591. Musca parralleli Harris, 1776: 43. Musca parrelleli Harris, 1776: 43. Syrphus deflagratusPreyssler, 1793: 369. Syrphus succinctus Panzer, 1804: 171.

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Volucella tricincta Müller, 1776: 178.

Korean record: Eristalis arbustorum: Kim, CW 1971: 868 (redescrip., first Korean record); Kim, JI 1975: 41 (Korean check.); Kim, CW 1980: 272 (distr. Map); ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998a: 11 (Yonsei Univ. Maeji Campus); Han et al. 1998b: 139 (Korean cat.); Choi et al. 2000: 144 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 147 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 15 (Korean check.). Eoseristalis arbustorum: Kim, JI 1980: 387 (Korean check.).

Description: Body length 10.2‒12.5 mm. Male. Head entirely black ground color; vertical triangle black with black pilose; face entirely shiny black, covered dense silver white pilose; antennal scape dark brown with brown setulae; pedicel black with black setulae; basoflagellomere oval, entirely black; arista dark brown; eyes holoptic, dense pale yellow pilose; eye contiguity more or less than vertical triangle length. Thorax shiny black ground color; scutum densely covered with yellowish brown pile; scutellum with dense yellowish brown pilose. Wing hyaline; halter yellowish brown. Legs: coxae and trochanters black; femora entirely black except apical 1/5 yellowish brown; fore and midtibia apical 1/4 black and basal 3/4 yellowish brown; hind tibia apical 2/3 black and basal 1/3 yellowish brown. Abdomen moderately elongated, shiny black ground color, densely covered with black and yellow brown pile mixed; tergite 2 pair of triangular marking; tergite 3‒4 entirle black. Female. Similar to male except for following non-genitalic characters: eyes dichoptic; wing hyaline with brown tinge medially; abdomen entirely black.

Specimen examined: Korea: CB: Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 8. viii. 2003, OY Lim & HS Lee, 1♀; Chungju-si, Sotae-myeon, Deokeun-ri, Namhangang river side, 11. x. 2002, DS Choi et al., 2♂. CN: Asan-si, Eumbong-myeon, Ssangyong-ri, 31. v. 2006, HS Lee & SW Suk, 1♂2♀; Cheonan-si, Seonggeo- eup, Cheonheung-ri, 21. v. 2006, HW Byun et al., 3♂4♀. GB: Dalseong-gun, Yaga-myeon, Mt. Biseulsan, Recreational Forest, 11. vii. 2000, HY Han et al., 16♂8♀; Gimcheon-si, Jirye-myeon, near Gyorigyo, 3. x. 2016, DS Choi, 1♀; Gyeongju-si, Amgok-dong, Mujangsaji, 22. vii. 2011, DS Choi, 1♂3♀; Yeongju- si, Sunheung-myeon, Mt. Sobaeksan, 27. viii. 2001, DS Choi et al., 2♂. GN: Ulsan, Ulju-gun, Onsan-eup, Deoksin-ri, 14. vi. 2008, DS Choi, 1♀. GG: Gapyeong-gun, Buk-myeon, Hwaak-ri, 3. vii. 2006, DS Choi et al., 1♂; Gwangju-si, Opo-myeon, Mr. Baengmasan, 27. v. 2001. DS Choi et al., 1♂; Paju-si, Papyeong- myeon, Masan-ri, 1. viii. 2006, DS Choi et al, 8♂2♀; Yangju-si, Gwangjeok-myeon, Ugo-ri, 2. viii. 2006, DS Choi et al., 4♂2♀. GW: Donghae-si, Manu-dong, 19. vii. 2010, SW Suk et al., 2♂; Gangneung-si, Gangdong- myeon, Eonbyeol-ri, 7. x. 2010, SW Suk et al., 2♀; Hoengseong-gun, Anheung-myeon, Mt. Chiaksan, from Gangrim 4-ri to Cheonjibong (1,086 m) peak, 13. vi. 1999, DS Choi, 3♂4♀; Hoengseong-gun, Dunnae- myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 1. viii. 2006, HW Byun et al., 1♂8♀; Inje-gun, Girin-

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myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 16. vi. 2004, HW Byun et al., 2♂; Inje-gun, Nam-myeon, Sangsunae-ri, 18. vii. 2015, 1♂; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 3. x. 2006, SW Suk, 5♂3♀. JB: Gimje-si, Hwangsan-dong, Munsusa Temple, 17. vi. 2011, DS Choi, 4♂3♀; Iksan-si, Sin-dong, Wongwang Univ. Campus, 30. x. 1999, HY Han et al., 4♀; Imsil-gun, Gangjin-myeon, Mt. Baekryensan, 6. vii. 2013, DS Choi, 1♀. JN: Damyang-gun, Subuk-myeon, Ojeong-ri, 17. v. 2007, DS Choi, 2♀.

Distribution: Holarctic, Oriental Regions [PA, NE, OR].

Biology: In Europe, larvae of this species have been reported from farmyard drains and temporary pools with cow droppings, so presumably other polluted situations also suffice (Stubbs & Falk, 1983).

Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

29. Eristalis cerealis Fabricius, 1805 (Pls. 4I, J)

Eristalis cerealis Fabricius, 1805: 232. Eristalis barbata Bigot, 1880: 214. Eristalis incisuralis Loew, 1858: 108. Eristalis solitus Walker, 1849: 619.

Korean record: Eristalis cerealis: Ichigawa 1906: 184 (Jeju-do, first Korean record); Shiraki 1930: 130 (redescrip.); Doi 1938a: 8 (Korean check.); ZSK 1968: 179 (Korean check.); Shiraki 1968b: 155 (redescrip.); Kim, CW 1971: 868 (redescrip.); Kim, JI 1975: 41 (Korean check.); Knutson et al. 1975: 351 (Orient. cat.); Kim, CW 1980: 273 (distr. Map); Peck 1988: 186 (Palaearctic cat.); ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998a: 11 (Yonsei Univ. Maeji Campus); Han et al. 1998b: 139 (Korean cat.); Choi et al. 2000: 144 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 148 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 15 (Korean check.). Eoseristalis cerealis: Kim, JI 1980: 387 (Korean check.).

Description: Body length 10.5‒12.8 mm. Male. Head entirely black ground color; frons black covered black pilose; vertical triangle black with black pilose; face entirely shiny black, covered dense silver white pilose; antennal scape dark brown with black setulae; pedicel black with black setulae; basoflagellomere oval, entirely dark brown; arista dark brown; eyes holoptic, dense pale yellow pilose; eye contiguity more or less than vertical triangle length. Thorax shiny black ground color; scutum with vertical grey band densely covered yellowish brown pile; scutellum with dense yellowish brown pilose. Wing hyaline; halter yellowish brown. Legs: coxae and trochanters black; femora entirely black except apical 1/5 yellowish brown; fore and midtibia apical 1/3 black and basal 2/3

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yellowish brown; hind tibia apical 2/3 black and basal 1/3 yellowish brown. Abdomen moderately elongated, shiny black ground color, densely covered with black and yellow brown pile mixed; tergite 2 with pair of yellow brown bands separated posterior margin. Female. Similar to male except for following non-genitalic characters: eyes dichoptic; abdominal tergite 2 with pair of thin bands.

Specimen examined: Korea: CB: Boeun-gun, Naesongni-myeon, Mt. Songnisan, Munjangdae, 21. vii. 2007, EJ Hong & SK Lee, 1♂; Cheongwon-gun, Namseong-myeon, Mt. Ingyeongsan, 15. viii. 1997, HY Han et al., 2♂1♀. CN: Boryeong-si, Cheongna-myeon, Mt. Oseosan, 20. vii. 1999, HY Han et al., 18♂12♀; Cheonan-si, Byeongcheon-myeon, Byeongcheon-ri, Sangnok Resort, 6. xi. 1999, HY Han et al., 3♂4♀; Taean-gun, Wonbuk-myeon, Banggal-ri, 8. viii. 2006, DS Choi et al., 1♂. GB: Bonghwa-gun, Myeongho- myeon, Mt. Cheongnyangsan, 23. x. 2009, SW Suk et al., 1♀; Cheongsong-gun, Budong-myeon, Mt. Juwangsan, 10. ix. 2006, DS Choi, 1♀; Gimcheon-si, Jirye-myeon, near Gyorigyo, 3. x. 2016, DS Choi, 1♀; Gyeongju-si, Amgok-dong, Mujangsaji, 22. vii. 2011, DS Choi, 1♂3♀. GN: Hapcheon-gun, Gaya-myeon, Haeinsa, 9. x. 2009, DS Choi, 2♂; Yangsan-si, Wondong-myeon, Sinheungsa, 12. iv. 2008, DS Choi, 1♀. GG: Dongducheon-si, Soyo-dong, Mt. Soyosan, 1. viii. 2006, DS Choi et al., 1♂1♀; Gapyeong-gun, Buk- myeon, Mt. Myeongjisan, 8. vii. 2001, DS Choi et al., 10♂7♀. GW: Hoengseong-gun, Anheung-myeon, Mt. Chiaksan, from Gangrim 4-ri to Cheonjibong (1,086 m) peak, 16. v. 1999, DS Choi et al., 2♂; Hoengseong- gun, Gangrim-myeon, Bugok Reservoir, 27. vi. 2015, DS Choi, 1♀; ditto, Mt. Maehwasan, 31. v. 2015, DS Choi, 1♀; Inje-gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 21. viii. 1998, HY Han et al., 3♂2♀; Jeongseon-gun, Nam-myeon, Mt. Mindungsan, form Yupyeong-ri to 1,119 m peak, 17. vii. 2008, SW Suk et al., 1♂; Taebaek-si, Mt. Taebaeksan, from Yuilsa to 1,560 m peak, 12. viii. 2000, DS Choi et al., 2♂1♀; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 12. vi. 1998, HW Byun, 1♂; ditto, 23. vii. 2009, HY Han, 1♀; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 16. vii. 2009, SW Suk et al., 1♂. JB: Iksan-si, Sin-dong, Wongwang Univ. Campus, 30. x. 1999, HY Han et al., 2♂. JN: Gwangju, Buk-gu, Mt. Mudeungsan, Neutjaesamgeori, 4. viii. 2013, DS Choi, 2♂; Jangheung- gun, Gwansan-eup, Mt. Cheongwansan, 14. v. 2000, DS Choi et al., 6♂. JJ: Bukjeju-gun, Jocheon-eup, Goepyeongi oreum, 2. v. 2003, DS Choi et al., 1♂.

Distribution: Russian Far East to China, Korea, Japan, Oriental Region [PA, OR].

Biology: Larvae of this species are found in polluted water.

Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

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30. Eristalis japonica van der Goot, 1964

Eristalis (Eoseristalis) japonica van der Goot, 1964: 218. Eristalis nigricans Matsumura, 1905: 93.

Korean record: Eristalis japonica: ESK & KSAE 1994: 292 (Korean check.); Han et al. 1998b: 140 (Korean cat.); Han & Choi 2001: 149 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 16 (Korean check.). Eristalis nigricans: Doi 1938a: 8 (Korean check., presumably first Korean record); Doi 1938b: 92 (Gaema-plateau); ZSK 1968: 179 (Korean check.); Kim, CW 1971: 867 (redescrip.); Kim, JI 1975: 41 (Korean check.). Eoseristalis nigricans: Kim, JI 1980: 387 (Korean check.).

Distribution: Russian Far East to Korea, Japan [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim (1971).

31. Eristalis kyokoae (Kimura, 1986) (Pls. 3K, L)

Eoseristalis kyokoae Kimura, 1986: 124.

Korean record: Eristalis kyokoae: Han et al. 2004: 123 (Mt. Gyemyeongsan, first Korean record); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 16 (Korean check.).

Description: Body length 11.2‒13.5 mm. Male. Head entirely black ground color; frons black covered black pilose; vertical triangle black with black pilose; face entirely shiny black, covered dense silver white pilose; antennal scape dark brown with black setulae; pedicel black with black setulae; basoflagellomere oval, entirely dark brown; arista dark brown; eyes holoptic, dense pale yellow pilose; eye contiguity more or less than vertical triangle length. Thorax shiny black ground color; scutum with vertical grey band densely covered yellowish brown pile; scutellum with dense yellowish brown pilose. Wing hyaline; halter yellowish brown. Legs: coxae and trochanters black; femora entirely black except apical 1/5 yellowish brown; fore and midtibia apical 1/3 black and basal 2/3 yellowish brown; hind tibia apical 2/3 black and basal 1/3 yellowish brown. Abdomen moderately elongated, shiny black ground color, densely covered with black and yellow brown pile mixed; tergite 2 with pair of yellow brown bands attached posterior margin. Female. Similar to male except for following non-genitalic characters: eyes dichoptic; abdominal tergite 2 with pair of triangular bands.

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Specimen examined: Korea: CB: Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 15. ix. 2006, HW Byun et al., 1♂; Chungju-si, Sotae-myeon, Deokeun-ri, Namhangang river side, 11. x. 2002, DS Choi et al., 3♂3♀; Jincheon-gun, Iwol-myeon, Naechon-ri, 21. viii. 2006, DS Choi et al., 1♂1♀. CN: Boryeong-si, Cheongna- myeon, Mt. Oseosan, 20. vii. 1999, HY Han et al., 2♂2♀; Cheonan-si, Seonggeo-eup, Cheonheung-ri, 21. v. 2006, HW Byun et al., 1♂; Taean-gun, Gonam-myeon, Janggok-ri, Haebyeonarae beach, 24. v. 2014, DS Choi, 1♀. GB: Bonghwa-gun, Myeongho-myeon, Mt. Cheongnyangsan, 13. vii. 2007, HS Lee et al., 1♀; Gyeongju-si, Geoncheon-eup, Mt. Danseoksan, 5. v. 2011, DS Choi, 2♂; Yeongju-si, Buseok-myeon, Mt. Seondalsan, from namdae-ri to 1,236 m peak, 29. vi. 1998, HY Han et al., 1♂. GN: Busan, Seodaesin-dong, Gudeok Campground, 30. iv. 2013, DS Choi, 2♂; Hamyang-gun, Anui-myeon, Mt. Gibaeksan, 28. viii. 2008, HS Lee et al., 1♀; Hapcheon-gun, Gaya-myeon, Haeinsa, 9. x. 2009, DS Choi, 1♀; Ulsan, Ulju-gun, Onsan- eup, Deoksin-ri, 14. vi. 2008, DS Choi, 1♀. GG: Gapyeong-gun, Buk-myeon, Dosolcheonsa, 13. ix. 2014, DS Choi, 1♂; Gwangju-si, Docheok-myeon, Mt. Taehwasan, from Eungoksa to 612.8 m peak, 25. viii. 2006, HY Han et al., 1♀; Paju-si, Paju-eup, Bongam-ri, 25. vii. 2006, DS Choi et al., 5♂2♀; Pocheon-si, Ildong-myeon, Gisan-ri, Daedeogsa, 23. vii. 2006, DS Choi et al., 1♀. GW: Hongcheon-gun, Nae-myeon, Mt. Gachilbong, from Sambongyaksu to 1,240 m peak, 2. v. 2003, HY Han & KE Ro, 3♂1♀; ditto, 7. ix. 2008, HS Lee et al., 1♂; Inje-gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 21. viii. 1998, HY Han et al., 1♂1♀; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 12. vi. 1998, HW Byun, 1♂. JB: Iksan-si, Sin-dong, Wongwang Univ. Campus, 30. x. 1999, HY Han et al., 3♂; Imsil-gun, Cheongung-myeon, Hyanggyo-ri, Mt. Baekisan, 6. vii. 2013, DS Choi, 1♀; Jangseong-gun, Bukha-myeon, Mt. Naejangsan, Ipam Wetland, 11. v. 2013, DS Choi, 1♂. JN: Gurye-gun, Gwangui-myeon, Mt. Jirisan, Nogodan, 11. ix. 1998, HY Han & KE Ro, 1♂1♀; Jangheung-gun, Gwansan-eup, Mt. Cheongwansan, 14. v. 2000, DS Choi et al., 3♂. JJ: Bukjeju-gun, Jocheon-eup, Mt. Hallasan, 5. viii. 2003, DS Choi & HW Byun, 2♂4♀.

Distribution: Korea, Japan [PA].

Biology: Unknown.

Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

32. Eristalis tenax (Linnaeus, 1758) (Pls. 4A, B, C)

Musca tenax Linnaeus, 1758: 591. Conops vulgaris Scopoli, 1763: 354. Eristalis alpinus Strobl, 1893: 185. Eristalis campestris Meigen, 1822: 387. Eristalis claripes Abreu, 1924: 104. Eristalis cognatus Wiedemann, 1824: 37. Eristalis columbica Macquart, 1855: 108.

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Eristalis hortorum Meigen, 1822: 387. Eristalis sinensis Wiedemann, 1824: 37. Eristalis sylvaticus Meigen, 1822: 388. Eristalis ventralis Thomson, 1869: 489. Eristalis vulpinus Meigen, 1822: 388. Musca apiformis Geoffroy, 1785: 488.

Korean record: Eristalis tenax: Okamoto 1924: 155 (Jeju-do, presumably first Korean record); Shiraki 1930: 140 (redescrip.); ZSK 1968: 179 (Korean check.); Kim, JI 1980: 387 (Korean check.); ESK & KSAE 1994: 292 (Korean check.); Han et al. 1998a: 11 (Yonsei Univ. Maeji Campus); Han et al. 1998b: 141 (Korean cat.); Choi et al. 2000: 144 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 150 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 16 (Korean check.). Eristalomyia tenax: Doi 1938a: 9 (Korean check.); Doi 1938b: 92 (Gaema-plateau); Kim, CW 1971: 878 (redescrip.); Kim, JI 1975: 41 (Korean check.); Kim, CW 1980: 275 (distr. Map). Eristalomya tenax [misspelling]: Shiraki 1968b: 163 (redescrip.).

Description: Body length 13.5‒18.5 mm. Male. Head entirely black ground color; frons black covered black and brown pilose; vertical triangle black with black pilose; face entirely shiny black, covered dense silver white pilose; antennal scape and pedicel shiny black with dark brown setulae; basoflagellomere oval, entirely black; arista dark brown; eyes holoptic, dense pale yellow pilose; eye contiguity shorter than vertical triangle length. Thorax: scutum shiny black densely covered yellowish brown pile; scutellum yellowish brown with dense yellowish brown pilose. Wing hyaline with brown tinge medially; halter yellowish brown. Legs: coxae and trochanters black; femora entirely black except apical 1/2 yellow brown on hind femur; fore and midtibiae apical 1/2 black and basal 1/2 yellow brown; hind tibiae entirely black. Abdomen moderately elongated, shiny black ground color, densely covered with yellow brown pile; tergite 2‒3 with yellow brown markings. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: CB: Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 3. vi. 2004, HW Byun et al., 1♂; Chungju-si, Sotae-myeon, Deokeun-ri, Namhangang river side, 11. x. 2002, DS Choi et al., 2♂3♀. CN: Boryeong-si, Cheongna-myeon, Mt. Oseosan, 20. vii. 1999, HY Han et al., 1♂2♀; Cheonan- si, Byeongcheon-myeon, Byeongcheon-ri, Sangnok Resort, 6. xi. 1999, HY Han et al., 1♂1♀; Hongseong- gun, Hongseong-eup, Mt. Baegwolsan, 16. viii. 1997, HY Han et al., 3♀. GB: Dalseong-gun, Yaga-myeon, Mt. Biseulsan, Recreational Forest, 20. v. 2012, DS Choi, 2♂3♀; Gyeongju-si, Amgok-dong, Mujangsaji, 22. vii. 2011, DS Choi, 1♂3♀; Yeongju-si, Sunheung-myeon, Mt. Sobaeksan, 27. viii. 2001, DS Choi et al., 1♀. GN: Hapcheon-gun, Gaya-myeon, Haeinsa, 9. x. 2009, DS Choi, 2♂3♀; Sancheong-gun, Sicheon-myeon,

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Mt. Jirisan, 30. vii. 1981, HY Han, 1♂. GG: Anyang-si, Manan-gu, Byeongmokan Park, 13. ix. 2015, DS Choi, 1♀; Gapyeong-gun, Buk-myeon, Mt. Myeongjisan, 19. vi. 2006, DS Choi et al., 4♂1♀; Namyangju- si, Sudong-myeon, Mt. Chungryeongsan, 30. vi. 1999, HY Han et al., 1♀; Paju-si, Paju-eup, Bongam-ri, 25. vii. 2006, DS Choi et al., 5♂; Yangju-si, Gwangjeok-myeon, Ugo-ri, 2. viii. 2006, DS Choi et al., 1♂. GW: Hoengseong-gun, Anheung-myeon, Mt. Chiaksan, from Gangrim 4-ri to Cheonjibong (1,086 m) peak, 16. v. 1999, DS Choi et al., 2♂; Hoengseong-gun, Gangrim-myeon, Bugok Reservoir, 27. vi. 2015, DS Choi, 4♂1♀; ditto, Mt. Maehwasan, 24. x. 2015, DS Choi, 1♂1♀; Inje-gun, Buk-myeon, Mt. Maebongsan, E. Valley, 25. viii. 1998, HY Han et al., 1♂3♀; Inje-gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 21. viii. 1998, HY Han et al., 1♂1♀; Jeongseon-gun, Nam-myeon, Mt. Mindungsan, form Yupyeong-ri to 1,119 m peak, 12. vi. 2009, SW Suk et al., 1♂; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 12. vi. 1998, HW Byun, 1♂; ditto, 15. xi. 1999, CH Park, 1♂; ditto, 5. vii. 2009, DJ Cha et al., 1♀. JB: Gimje-si, Hwangsan-dong, Munsusa Temple, 17. vi. 2011, DS Choi, 1♀; Wanju-gun, Unju-myeon, Mt. Daedunsan, 1. x. 2000, DS Choi & SK Kim, 2♀. JN: Gokseong-gun, Gokseong-eup, Wolpyeong-ri, 15. v. 2007, DS Choi, 1♀; Jangheung-gun, Gwansan-eup, Mt. Cheongwansan, 14. v. 2000, DS Choi et al., 3♂. JJ: Bukjeju-gun, Jocheon-eup, Mt. Hallasan, 5. viii. 2003, DS Choi & HW Byun, 2♂4♀. Distribution: Cosmopolitan. Biology: Larvae of this species are found in polluted water (Han and Choi, 2001). Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

Genus Helophilus Meigen, 1822

Helophilus Meigen, 1822: Syst. Beschr., 3: 368. TS: Musca pendula Linnaeus (des. Curtis, 1832: Brit. Ent., 9: pl. 429). Eurimyia Bigot, 1883: Annls. Soc. ent. Fr., (6) 3, Bull.: XX. TS: Eurimyia rhingioidis Bigot (MO) [=lineata (Fabricius)]. Published in March 1883. Eurhimyia Bigot, 1883: Annls. Soc. ent. Fr., (6) 3: 226 and 230. TS: Rhingia lineata Fabricius (OD). Published inOctober 1883. Eurinomyia Mik, 1897: Wien. ent. Ztg., 16 (2): 115; unjustified emendation. Anasimyia Schiner, 1864: Catal. syst. dipt. Europ.: 108. TS: Musca transfuga Linnaeus (des. Coquillett, 1910: Proc.U.S. natn. Mus., 37: 506). Kirimyia Bigot, 1882: Annls Soc. ent. Fr., (6) 2, Bulol.: CXXXVI (mon.). TS: Kirimyia eristaloidea Bigot. Parhelophilus Girschner, 1897: Illte Wschr. Ent., 2: 604 (as subgenus of Helophilus Meigen). TS: Syrphus frutetorum Fabricius (des. Curran and Fluke, 1926: Trans. Wis. Acad.Sci. Arts Lett., 22: 230).

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Key to species of Helophilus from South Korea

1. Abdominal bands sparsely attached on tergite 2 ············································ H. virgatus Coquillett − Abdominal bands distintly separated on tergite 2 ··································································· 2 2. Hind femur entirely black ··································································· H. trivittatus (Fabricius) − Hind femur black posteriorly and brown anteriorly ········································· H. lunulatus Meigen

33. Helophilus lunulatus Meigen, 1822 (Pls. 4D, E)

Helophilus lunulatus Meigen, 1822: 370. Eurinomyia japonica Shiraki, 1930: 167.

Korean record: Helophilus lunulatus: Han & Choi 2001: 154 (Korean check., first Korean record); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 16 (Korean check.).

Description: Body length 7.2‒8.5 mm. Male. Head almost entirely yellow brown with dark brown post-ocular orbit and vertex, shiny black lower portion of face; ocellar triangle with black pilose; face almost silver white pilose with lower portion distinctly protrude anteriorly; eyes dichoptic, bare; antennal scape and pedicel brown with black setulae; basoflagellomere yellowish brown, oval; arista dark brown. Thorax: scutum with 4 pale longitudinal stripes, almost entirely yellow brown dense pilose; scutellum dark brown, almost short dense yellow brown pilose. Wing hyaline; halter yellow brown. Legs: coxae and trochanters entirely black; fore and midfemora black ventrally and brown dorsally; hind femur swollen with black posteriorly and brown anteriorly; fore and midtibiae yellow brown; hind tibia yellowish brown with black marking apical and basal portion. Abdomen entirely black with pale bands Female. Similar to male except for following non-genitalic characters: abdominal band more expanded medially on tergite 2.

Specimen examined: Korea: GW: Hoengseong-gun, Anheung-myeon, Anheung-ri, Jucheongang riverside, 9. v. 1999, DS Choi et al., 1♀; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 23. v. 2006, DS Choi, 1♂; ditto, 14. v. 2004, HW Byun & OY Lim, 1♀; ditto, 21. iv. 2004, HW Byun, 2♂; ditto, 2. v. 2001, CH Park, 3♂; ditto, 26. iv. 2001, DS Choi & SK Kim, 2♂; ditto, 25. iv. 2001, DS Choi & OY Lim, 4♂; ditto, 24. iv. 2001, DS Choi & SK Kim, 1♂1♀; ditto, 21. iv. 2001, CH Park et al., 1♀; ditto, 8. v. 2000, CH Park, 1♂; ditto, 6. v. 2000, DS Choi, 6♂; ditto, 4. v. 2000, SK Kim, 2♂; ditto, 3. v. 2000, CH Park, 1♂1♀; ditto, 3. v. 2000, HY Han & KE Ro, 5♂2♀; ditto, 1. v. 2000, SK Kim, 2♂; ditto, 30. iv. 2000, DS Choi & SK Kim,

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4♂2♀; ditto, 25. iv. 2001, DS Choi & OY Lim, 1♂.

Distribution: Europe, Korea, Japan, Nearctic Region [PA, NE].

Biology: Unknown.

Korea: GW.

34. Helophilus sapporensis Matsumura, 1911

Helophilus pendulus var. sapporensis Matsumura, 1911: 75. Helophilus lateralis Matsumura, 1916: 247.

Korean record: Helophilus sapporensis: Kim, CW 1971: 846 (redescrip., first Korean record); Kim, JI 1980: 388 (Korean check.); ESK & KSAE 1994: 292 (Korean check.); Han et al. 1998a: 11 (Yonsei Univ. Maeji Campus); Han et al. 1998b: 143 (Korean cat.); Choi et al. 2000: 144 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 155 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 17 (Korean check.). Helophilus saspporensis [misspelling]: Kim, JI 1975: 41 (Korean check.).

Distribution: Russian Far East to Korea, Japan [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim (1971).

35. Helophilus trivittatus (Fabricius, 1805) (Pl. 4F)

Eristalis trivittatus Fabricius, 1805: 235. Helophilus camporum Meigen, 1822: 372. Helophilus solitarius Rondani, 1857: 50. Musca parallelus Harris, 1776: 57.

Korean record: Helophilus trivittatus: Han & Choi 2001: 155 (Korean check., first Korean record); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 17 (Korean check.).

Description: Body length 15.6 mm. Male. Head entirely black ground color; ocellar triangle black; face entirely shiny black, covered dense silver white pilose; antennal scape and pedicel black; basoflagellomere oval, entirely black; arista dark brown; eyes dichoptic. Thorax: scutum black with 4 yellow brown longitudinal vittae; scutellum brown with

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dense yellowish brown pilose. Wing hyaline; halter yellowish brown. Legs: coxae and trochanters black; femora entirely black; fore and midtibiae yellowish brown, hind tibia apical 3/4 black and basal 1/4 yellow brown; hind tarsus entirely black. Abdomen moderately elongated, longer than head and thorax together; tergite 2‒3 with pair of large yellow brown marking not attached hind margin; tergite 4 with pair of ‘v’ type grey marking. Female. No specimens available.

Specimen examined: Korea: GW: Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 5. x. 2000, CH Park, 1♂.

Distribution: Palaearctic Region [PA].

Biology: Unknown.

Korea: GW.

36. Helophilus virgatus Coquillett, 1898 (Pls. 4G, H, I)

Helophilus virgatus Coquillett, 1898: 326. Helophilus frequens Matsumura, 1905: 103. Helophilus virgatus Coquillett, 1898: 326. Helophilus zodinus Matsumura, 1931: 341

Korean record: Helophilus virgatus: Doi 1938a: 9 (Korean check.); Kim, JI 1980: 388 (Korean check.); ESK & KSAE 1994: 292 (Korean check.); Han et al. 1998a: 11 (Yonsei Univ. Maeji Campus); Han et al. 1998b: 129 (Korean cat.); Choi et al. 2000: 143 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 156 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 17 (Korean check.). Tubifera virgatus: ZSK 1968: 179 (Korean check.); Kim, CW 1971: 851 (redescrip.); Kim, JI 1975: 41 (Korean check.); Kim, CW 1980: 292 (distr. Map). Helophilus frequens: Okamoto 1924: 155 (Jeju-do, presumably first Korean record); ZSK 1968: 179 (Korean check.).

Description: Body length 13.5‒16.2 mm. Male. Head almost entirely yellow brown with dark brown post-ocular orbit and vertex, shiny black anterior and lower portion of face; ocellar triangle with black pilose; face almost yellow brown pilose; eyes dichoptic, bare; antennal scape and pedicel black with black setulae; basoflagellomere yellowish brown, oval; arista dark brown. Thorax: scutum with 2 pale longitudinal stripes, almost entirely yellow brown dense pilose; scutellum dark brown, dense yellow brown and black pilose mixed. Wing hyaline; halter yellow brown. Legs: coxae and trochanters entirely black; fore and midfemora apical 3/4 black and basal 1/4 brown;

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hind femur entirely black, swollen; fore and midtibiae yellow brown; fore tibia apical 1/2 black and basal 1/2 brown; midtibia entirely brown; hind tibia entirely black. Abdomen: tergite 2 black with pair of large triangular yellow brown band; tergite 3 black with vertical yellow brown band; tergite 4 entirely black. Female. Similar to male except for following non-genitalic characters: eyes distinctly dichoptic.

Specimen examined: Korea: CB: Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 3. vi. 2004, HW Byun et al., 1♂; Chungju-si, Sotae-myeon, Deokeun-ri, Namhangang river side, 11. x. 2002, DS Choi et al., 1♂2♀. CN: Cheonan-si, Byeongcheon-myeon, Maeseong-ri, 21. viii. 2006, DS Choi et al., 2♀; Dangjin-gun, Sinpyeong-myeon, Geumcheon-ri, 10. viii. 2006, DS Choi et al., 1♂; Nonsan-si, Yeonsan-myeon, Gojeong-ri, Gyebaekjanggunmyo, 15. vi. 2010, SJ Park, 1♂2♀. GB: Gimcheon-si, Jirye-myeon, Gyo-ri, near Gyorigyo, 3. x. 2016, DS Choi, 2♂3♀; Gyeongju-si, Geoncheon-eup, Mt. Danseoksan, 5. v. 2011, DS Choi, 2♂; Yeongju- si, Buseok-myeon, Mt. Seondalsan, from namdae-ri to 1,236 m peak, 29. vi. 1998, HY Han et al., 1♂; Yeongju-si, Punggi-eup, Mt. Sobaeksan, Yeonhwabong, 16. viii. 2016, DS Choi, 1♀. GN: Busan, Seodaesin- dong, Gudeok Campground, 7. x. 2011, DS Choi, 2♀; Geochang-gun, Buksang-myeon, Mt. Deokyusan, Wolseong Park, 8. vii. 2012, DS Choi 4♂2♀; Sancheong-gun, Sicheon-myeon, Mt. Jirisan, 30. vii. 1981, HY Han, 1♂. GG: Gapyeong-gun, Buk-myeon, Mt. Myeongjisan, 19. vi. 2006, DS Choi et al., 1♂1♀; Paju-si, Paju-eup, Bongam-ri, 25. vii. 2006, DS Choi et al., 5♂; Yangju-si, Goam-dong, 2. viii. 2006, DS Choi et al., 1♂2♀. GW: Hoengseong-gun, Anheung-myeon, Mt. Chiaksan, from Gangrim 4-ri to Cheonjibong (1,086 m) peak, 16. v. 1999, DS Choi et al., 2♂; Hoengseong-gun, Gangrim-myeon, Mt. Maehwasan, 27. vi. 2015, DS Choi, 2♂; Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 1. viii. 2006, HW Byun et al., 1♂2♀; Hongcheon-gun, Nae-myeon, Mt. Gachilbong, from Sambongyaksu to 1,240 m peak, 25. v. 2009, SW Suk & DJ Cha, 1♀; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 14. viii. 2009, 1♂; Wonju-si, Gwirae-myeon, from Cheoneunsa to Mt. Sipjabong peak, 18. viii. 1997, HY Han et al., 1♂; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 16. vii. 2009, SW Suk et al., 1♂. JB: Iksan-si, Sin-dong, Wongwang Univ. Campus, 30. x. 1999, HY Han et al., 5♂; Jeongeup-si, Mt. Naejangsan, Wonjeok Valley, 11. v. 2013, DS Choi, 1♀; Sunchang-gun, Bokheung-myeon, Mt. Naejangsan, 13. vii. 2013, DS Choi, 1♂; Wanju-gun, Unju-myeon, Mt. Daedunsan, 1. x. 2000, DS Choi & SK Kim, 5♀. JN: Damyang-gun, Subuk-myeon, Ojeong-ri, 17. v. 2007, DS Choi, 2♀; Gwangyang-si, Ongnyong-myeon, Mt. Baegunsan, 2. vii. 2009, DJ Cha & YB Lee, 1♀; Jangheung- gun, Gwansan-eup, Mt. Cheongwansan, 14. v. 2000, DS Choi et al., 3♂. JJ: Bukjeju-gun, Jocheon-eup, Mt. Hallasan, 5. viii. 2003, DS Choi & HW Byun, 2♂1♀.

Distribution: Russian Far East, China, Korea, Japan [PA].

Biology: Unknown.

Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

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Genus Mallota Meigen, 1822

Mallota Meigen, 1822: Syst. Beschr. 3: 377. TS: Syrphus fuciformis Fabricius (des. Rondani, 1845: Nuovi Ann. Sci.nat. Ist. Bologna, (2) 2: 452). Zetterstedtia Rondani, 1845: Nuovi Ann. Sci. nat. Ist. Bologna, (2) 2: 452. TS: Syrphus cimbiciformis Fallén (OD). Paramesembrius Shiraki, 1930: Mem. Fac. Agric. Taihoku imp. Univ., 1 (1): 176 (as subgenus of Mesembrius Rondani, 1857. TS: Tubifera abdominalis Sack (des. Shiraki 1968: Fauna jap. Syrphidae, 3: 227). Pseudomallota Shiraki, 1930: Mem. Fac. Agric. Taihoku imp. Univ., 1 (1): 187 (as subgenus of Mallota Meigen). TS: Mallota tricolor Loew (OD). Pseudomerodon Shiraki, 1930: Mem. Fac. Agric. Taihoku imp. Univ., 1 (1): 189 (as subgenus of Mallota Meigen). TS: Mallota takasagoensis Matsumura (OD). Bombozelosis Enderlein, 1934: Sber. Ges. naturf. Freunde Berl., 1934 (4-7): 186. TS: Bombozelosis koreana Enderlein (OD).

Key to species of Mallota from South Korea

1. Male eyes distintly holoptic ····································································· M. shatalkini Mutin − Male eyes dichoptic ····································································································· 2 2. Thorax clothed with only black pilose ·························································· M. analis (Shiraki) − Thorax chothed with not only black pilose ··········································································· 3 3. Scutum clothed greenish yellow pilose ··············································································· 4 − Scutum clothed not greenish yellow pilose ·········································································· 5 4. Abdomen with conspicuous black band ···················································· M. dimorpha (Shiraki) − Abdomen without conspicuous black band ················································· M. ambigua (Shiraki) 5. Thorax clothed with dense three color (yellow brown, black and white) pilose ············ M. tricolor Loew − Thorax clothed with predominantly yellow brown pilose ······························ M. rossica Portschinsky

37. Mallota ambigua (Shiraki, 1968) (Pls. 4J, K)

Imatisma ambiguum Shiraki, 1968b: 238.

Korean record: Mallota ambigua: Han & Choi 2001: 158 (Korean check., first Korean record); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 17 (Korean check.).

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Description: Body length 17.2‒20.5 mm. Male. Head almost entirely shiny black with shiny brown lunule; vertical triangle brown with black pilose; frons with greenish yellow pilose; face almost silver white pilose; eyes holoptic, dense white pilose; antennal scape and pedicel shiny black with black and white setulae mixed; basoflagellomere black, oval; arista dark brown. Thorax: scutum black with anterior half dense greenish yellow pilose and dense black pilose posteriorly; scutellum brown with greenish yellow pilose. Wing hyaline; halter dark brown. Legs: coxae and trochanters entirely black; femora entirely black; hind femur strongly thickened and curved with ventral spine at about apical 1/4; tibiae entirely dark brown; tarsi entirely brown. Abdomen entirely black with greenish yellow pilose; tergite 2 without conspicuous black band. Female. Similar to male except for following non-genitalic characters: eyes dichoptic; abdomen wider than male.

Specimen examined: Korea: CB: Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 3. vi. 2004, HW Byun et al., 13♂; ditto, 2. vii. 2003, DS Choi et al., 2♂. GB: Yeongju-si, Punggi-eup, Mt. Sobaeksan, 27. v. 2005, HW Byun, 1♂; ditto, 11. vi. 2004, HW Byun et al., 1♂. GN: Jinyang-gun, Nadong-myeon, Eonpyeong-ri, 27. vi. 1992, 1♀; Ulju-gun, Samnam-myeon, Mt. Sinbulsan, 29. vi. 2003, HY Han et al., 1♂. GG: Gwangju-si, Opo-myeon, Mr. Baengmasan, 27. v. 2001, DS Choi et al., 5♂. GW: Hoengseong-gun, Anheung-myeon, Mt. Chiaksan, from Gangrim 4-ri to Cheonjibong (1,086 m) peak, 5. vi. 2005, DS Choi et al., 3♂; Hoengseong- gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 18. vi. 2005, HY Han et al., 1♂; ditto, 5. vii. 2003, DS Choi et al., 4♂; Hongcheon-gun, Nae-myeon, Mt. Gachilbong, from Sambongyaksu to 1,240 m peak, 3. vii. 2000, HY Han et al., 1♂; Jeongseon-gun, Jeongseon-eup, Mt. Gariwangsan, 31. v. 1998, HY Han et al., 1♀; Jeongseon-gun, Nam-myeon, Mt. Mindungsan, form Yupyeong-ri to 1,119 m peak, 6. vi. 2006, HY Han et al., 3♂; Wonju-si, Gwirae-myeon, from Cheoneunsa to Mt. Sipjabong peak, 4. vi. 1999, JY Lim et al., 1♀; ditto, HG Kim et al., 1♂; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 11. vii. 2006, JS Lim, 1♀; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 12. vi. 2006, HS Lee & SMR Hwang, 1♂; ditto, 15. vii. 2007, HS Lee & SW Suk, 1♂.

Distribution: Korea, Japan [PA].

Biology: Unknown.

Korea: CB, GB, GN, GG, GW.

38. Mallota analis (Shiraki, 1968) (Pl. 4L)

Imatisma analis Shiraki, 1968b: 241.

Korean record: Mallota analis: Han et al. 1998a: 11 (Yonsei Univ. Maeji Campus, first Korean record);

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Han et al. 1998b: 144 (Korean cat.); Choi et al. 2000: 144 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 157 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 17 (Korean check.).

Description: Body length 17.5‒20.5 mm. Male. Head almost entirely black with shiny brown lunule; vertical triangle with black pilose; frons with black pilose; face with dense silver white pilose; eyes holoptic, dense pilose; antennal scape and pedicel black with black setulae; basoflagellomere yellowish brown, oval; arista dark brown. Thorax black ground color; scutum black with dense black pilose; scutellum largely black and dark brown posteriorly with dense black plise and dense long white pilose hind margin. Wing hyaline; halter dark brown. Legs: coxae and trochanters entirely black; femora and tibiae almost entirely black; tarsi brown. Abdomen black ground color; tergite 2 with dense black pilose; tergite 3 with dense black pilose but brown pliose near hind margin; tergite 4 with dense brown pilose. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: CB: Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 2. vii. 2003, HW Byun et al., 1♀. GG: Goyang-si, Hyoja-dong, Bukhansanseong, 28. v. 1991, YA Kim, 1♀. GW: Hongcheon-gun, Nae- myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 10. viii. 2009, DJ Cha & HS Lee, 1♀; ditto, 23. vi. 2005, DS Choi et al., 1♂; Inje-gun, Girin-myeon, Mt. Bangtaesan, 6. vii. 2000, HY Han et al., 2♂; Wonju- si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 18. vi. 2009, DJ Cha, 1♀; ditto, 5. vii. 2009, DJ Cha et al., 1♂; Wonju-si, Sillim-myeon, Mt. Chiaksan, from Seongnam-ri to Namdaebong (1,181.5 m) peak, 10. vi. 2010, DS Choi et al., 1♂; Yeongwol-gun, Yeongwol-eup, Mt. Taehwasan, Palgoe-ri, 9. vi. 2001, DS Choi, DS Choi & SK Kim, 1♂1♀.

Distribution: Korea, Japan [PA].

Biology: Unknown.

Korea: CB, GG, GW.

39. Mallota dimorpha (Shiraki, 1930) (Pls. 5A, B)

Imatisma dimorpha Shiraki, 1930: 202. Mallota floreae Violovitsh, 1952: 57.

Korean record: Mallota dimorpha: ESK & KSAE 1994: 292 (Korean check.); Han et al. 1998a: 11 (Yonsei Univ. Maeji Campus); Han et al. 1998b: 144 (Korean cat.); Choi et al. 2000: 144 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 159 (Korean check.); Paek et al. 2010: 232 (Korean check.);

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Han et al. 2014: 18 (Korean check.). Imatisma dimorpha: Morita 1935: 527 (new Korean record); Kim, CW 1971: 845 (redescrip.); Kim, JI 1975: 41 (Korean check.); Kim, CW 1980: 277 (distr. Map); Kim, JI 1980: 388 (Korean check.).

Description: Body length 18.5‒20.5 mm. Male. Head almost entirely shiny black with shiny brown lunule; vertical triangle brown with black pilose; frons with greenish yellow pilose; face almost silver white pilose; eyes holoptic, dense white pilose; antennal scape and pedicel shiny black with black and white setulae mixed; basoflagellomere black, oval; arista dark brown. Thorax: scutum black with anterior half dense greenish yellow pilose and dense black pilose posteriorly; scutellum brown with greenish yellow pilose. Wing hyaline; halter dark brown. Legs: coxae and trochanters entirely black; femora entirely black; hind femur strongly thickened and curved with ventral spine at about apical 1/4; tibiae entirely dark brown; tarsi entirely brown. Abdomen entirely black with greenish yellow pilose; tergite 2 without conspicuous black band. Female. Similar to male except for following non-genitalic characters: eyes dichoptic; abdomen wider than male.

Specimen examined: Korea: CB: Jecheon-si, Baegun-myeon, Mt. Sirangsan, DS Choi et al., 27. vi. 2000, 1♀; Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 3. vi. 2004, HW Byun et al., 4♂; Cheongwon-gun, Namseong-myeon, Mt. Ingyeongsan, 15. viii. 1997, HY Han et al., 1♀. GB: Dalseong-gun, Yuga-myeon, Mt. Biseulsan, 11. vi. 2000, 1♂. GG: Gwangju-si, Opo-myeon, Mr. Baengmasan, 27. v. 2001, DS Choi et al., 2♂; Pocheon-si, Sinbuk-myeon, Yami-dong, Mt. Deokryeongsan, 27. v. 2001, DS Choi et al., 1♀. GW: Hoengseong-gun, Anheung-myeon, Mt. Chiaksan, from Gangrim 4-ri to Cheonjibong (1,086 m) peak, 5. vi. 2005, DS Choi et al., 4♂; Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 6. vii. 2007, HS Lee et al., 1♂; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 5. viii. 2008, SW Suk et al., 1♀; Inje-gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 4. viii. 2006, DS Choi et al., 2♀; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 24. viii. 2006, HW Byun, 1♀; ditto, 9. viii. 2009, DJ Cha et al., 1♂; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 26. vi. 2004, HS Lee et al., 2♂.

Distribution: Russian Far East, China, Korea, Japan [PA].

Biology: Unknown.

Korea: CB, GB, GG, GW.

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40. Mallota eurasiatica Stackelberg, 1950

Mallota eurasiatica Stackelberg, 1950: 292.

Korean record: Mallota eurasiatica: Peck 1988: 201 (Palaearctic cat.); ESK & KSAE 1994: 292 (Korean check.); Han et al. 1998b: 145 (Korean cat.); Han & Choi 2001: 160 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 18 (Korean check.).

Distribution: Russia, Korea?, Japan [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on ESK & KSAE (1994).

41. Mallota koreana (Enderlein, 1934)

Bombozelosis koreana Enderlein, 1934: 186. Bombozelosis coreana Enderlein, 1937: 434.

Korean record: Mallota koreana: Peck 1988: 108 (Palaearctic cat.); Han et al. 1998b: 160 (Korean cat.); Han & Choi 2001: 160 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 18 (Korean check.).

Distribution: Korea [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Peck (1988).

42. Mallota rossica Portschinsky, 1877 (Pls. 5C, D)

Mallota rossica Portschinsky, 1877: 175 Mallota auricoma Sack, 1910: 36 Mallota aino Violovitsh, 1952: 56

Korean record: Mallota rossica: Suk et al. 2015: 258 (new Korean record).

Description: Body length 16.5‒18.1 mm.

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Male. Head almost entirely black ground color; vertical triangle black with brown pilose; face entirely black; eyes holoptic, densely covered with hairs; antennal scape and pedicel shiny black with black pilose; basoflagellomere wider than long; arista dark brown. Thorax: scutum black, anterior half with dense pale yellow pruinosity, contrasting well with distinctly darker area behind; scutellum brown with long dense brown pilose. Wing hyaline with dark brown tinge medially; halter black. Legs: femora black; hind femur moderately thickened; fore tibia with long black hairs posteroventrally. Abdominal tergites densely covered with long brownish hairs; tergite 2 laterally with inner-facing brownish yellow triangular sopts. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: GW: Inje-gun, Buk-myeon, Mt. , Baekdamsa Temple, 26. vi. 2002, HJ Kim, 1♀; Wonju-si, Panbu-myeon, Baegunsan, from Yongsugol to 1,087.1 m peak, 19. vi. 1999, HY Han et al., 1♂; ditto, 5. vi. 2005, HY Han & HS Lee, 1♂; ditto, 14. vi. 2005, HY Han et al., 1♂; Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 9. viii. 2003, OY Lim & HS Lee, 1♂.

Distribution: Russian Far East, Korea [PA].

Biology: Unknown.

Korea: GW.

43. Mallota shatalkini Mutin, 1999 (Pls. 5E, F)

Mallota shatalkini Mutin in Mutin and Barkalov, 1999: 175

Korean record: Mallota shatalkini: Suk et al. 2015: 258 (new Korean record).

Description: Body length 13.2‒15.2 mm. Male. Head black ground color; face with vertically elliptic, brownish black, shiny and bare median stripe; vertex dark brown with dense black pilose; frons black with dense brown pilose; eyes dichoptic, bare; antennal scape and pedicel dark brown with white setulae; basoflagellomere brown, rounded; arista dark brown. Thorax: scutum black with almost entirely yellow brown dense pilose; scutellum brown with yellow brown dense pilose. Wing hyaline with dark brown tinge medially; halter yellow brown. Legs: coxae and trochanters entirely brown except black of hind trochanter posteriorly; femora brown anteriorly and black posteriorly; hind femur strongly thickened and curved with ventral spine at about apical 2/5; tibiae and tarsi entirely black. Abdomen black ground color with dense brown pilose: tergite 2 and 3 with thick transevers, medially interrupted brownish yellow stripe; tergite 3 black with vertical yellow brown band. Female. Similar to male except for following non-genitalic characters: eyes distinctly dichoptic.

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Specimen examined: Korea: CN: Gongju-si, Gyeryong-myeon, Mt. Gyeryongsan, Gapsa Temple, 26. v. 1974, 1♂; Yeongi-gun, Jochiwon-eup, 18. v. 1984, JK Lee, 1♀. GN: Haman-gun, Gunbuk-myeon, Ogok- ri, 31. v. 1987, 1♀; Uiryeong-gun, Garye-myeon, Gabeul-ri, Mt. Jagulsan, 20. v. 1990, 1♀. GG: Gapyeong- gun, Cheongpyeong-myeon, Daeseong-ri, 17. v. 1986, SR Choi, 1♀; Gwangju-si, Jungbu-myeon, eommi- ri, 23. v. 1981, KH Oh, 1♂; Namyangju-si, Paldang, 17. v. 1969, GC Hwang, 1♂; Namyangju-si, Sudong- myeon, Mt. Cheonmasan, 20. iv. 1971, KH Park, 1♂; Pocheon-si, Sohol-eup, Korea National Arboretum, 27. vi. 1981, DW Park, 1♀; Suwon-si, Gwonseon-gu, Seoul National University, 21. v. 1992, BT Kim, 1♂. GW: Chuncheon-si, Namsan-myeon, Gangchon-ri, 6. vi. 1984, YS Han, 1♂; Wonju-si, Heungeop-myeon, Maeji- ri, Yonsei Univ. Campus, 27. v. 1997, HW Byun & DS Choi, 1♂.

Distribution: Russian Far East, Korea, Japan [PA].

Biology: Unknown.

Korea: CN, GN, GG, GW.

Remarks: At a glance this species appears to be honeybee probably as a typical case of Batesian mimicry (Suk et al., 2015).

44. Mallota takasagoensis Matsumura, 1916

Mallota takasagoensis Matsumura, 1916: 203.

Korean record: Mallota takasagoensis: ESK & KSAE 1994: 292 (Korean check.); Han et al. 1998b: 145 (Korean cat.). Pseudomerodon takasagoensis: Kim, JI & Kim, CW 1972: 11 (new Korean record); Kim, JI 1975: 41 (Korean check.); Kim, JI 1980: 388 (Korean check.); Han & Choi 2001: 161 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 18 (Korean check.).

Distribution: Korea, Japan [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim & Kim (1972).

45. Mallota tricolor Loew, 1871 (Pl. 5G)

Mallota tricolor Loew, 1871: 234. Mallota japonica Matsumura, 1916: 200.

Korean record: Mallota tricolor: Han et al. 1998a: 11 (Yonsei Univ. Maeji Campus); Han et al. 1998b:

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145 (Korean cat.); Choi et al. 2000: 144 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 161 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 18 (Korean check.). Pseudomallota tricolor: Shiraki 1968b: 244 (redescrip., presumably first Korean record).

Description: Body length 13.5‒16.2 mm. Male. Head entirely shiny black except lunule shiny brown; vertical triangle black with black pilose; face with almost entirely silver white pilose; eyes holoptic, bare; antennal scape and pedicel black with white setulae; basoflagellomere yellowish brown, oval; arista dark brown. Thorax: scutum entirely black with dense yellow brown pilose anteriorly and dense black pilose posteriorly; scutellum brown with dense white pilose. Wing hyaline with dark brown tinge medially; halter dark brown. Legs: coxae and trochanters entirely black; femora entirely black; hind femur strongly thickened and curved with ventral spine at about apical 2/5; tibiae entirely black; tarsi entirely brown. Abdomen entirely black; tergite 1 dense white pilose; tergite 2 dense yellow brown pilose anteriorly and black pilose posteriorly; tergite 3 entirely dense black pilose; tergite 4 entirely dense orange pilose. Female. Similar to male except for following non-genitalic characters: eyes distinctly dichoptic.

Specimen examined: Korea: GG: Seoul, Dobong-gu, Mt. Dobongsan, 9. v. 1998, SH Lee, 1♂; Seoul, Seocho-gu, Mt. Cheongtaesan, 28. v. 1989, JH Jung, 1♂. GW: Hongcheon-gun, Nae-myeon, Mt. Gachilbong, from Sambongyaksu to 1,240 m peak, 3. vii. 2000, HY Han et al., 1♂; ditto, 6. vi. 1999, HY Han & HW Byun, 1♂1♀; Inje-gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 7. viii. 1997, HY Han et al., 1♂; Taebaek-si, Sodo-dong, Cheongwonsa, 22. v. 1999, TJ Jin et al., 1♂; Wonju-si, Heungeop- myeon, Maeji-ri, Yonsei Univ. Campus, 6. vii. 2004, HW Byun & JS Soh, 1♂; ditto, 28. vi. 2003, HW Byun, 1♀; ditto, 16. vi. 1996, 1♀. JB: Gochang-gun, Seonunsan, 25. vi. 1991, JI Kim, 1♂; Jeongeup-si, Ongdong- myeon, 22. vi. 1973, TH Shin, 1♂. JN: Suncheon-si, Juam-myeon, Yongmoon-ri, Yongnaegyo, 13. vi. 2010, DS Choi, 2♂.

Distribution: Europe, Russia, Turkey, China, Korea, Japan (Honshu, Kyushu) [PA].

Biology: Unknown.

Korea: GG, GW, JB, JN.

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Genus Mesembrius, 1857

Mesembrius Rondani, 1857: Dipt. Ital. prodromus, 2: 50. TS: Helophilus peregrinus Loew (MO).

46. Mesembrius flaviceps (Matsumura, 1905) (Pls. 5H, I)

Helophilus flavicepsMatsumura, 1905: 104. Helophilus akakurensis Matsumura, 1916: 246. Helophilus flavicepsMatsumura, 1905: 104. Mesembrius nigra Shiraki, 1968b: 225. Mesembrius okinawaensis Shiraki, 1968b: 222. Tropidia sinensis Macquart, 1855: 111.

Korean record: Mesembrius flaviceps: Peck 1988: 202 (Palaearctic cat.); ESK & KSAE 1994: 292 (Korean check.); Han et al. 1998b: 146 (Korean cat.); Han & Choi 2001: 162 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 18 (Korean check.). Mesembrius flavipes [misspelling]: Kim, JI et al. 1983: 208 (new Korean record).

Description: Body length 10‒12.2 mm. Male. Head: vertex black with black pilose; occiput with brown pilose; frons yellowish brown with dense yellowish brown pilose; lunule shiny brown; face almost silver white pilose with lower portion shiny black ventrally; eye holoptic, bare; antennal scape and pedicel brown with black setulae; basoflagellomere black, oval; arista dark brown. Thorax: scutum with 4 pale longitudinal stripes, almost entirely brown dense pilose; scutellum brown with dense brown pilose. Wing hyaline; halter yellow brown. Legs: coxae and trochanters entirely black; femora entirely black; hind femur moderately thickened; fore and midtibia entirely yellow brown; hind tibiae entirely brown. Abdomen entirely black with yellow brown bands; tergite 2‒3 each pair of brown quadrate bands. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: CB: Jincheon-gun, Iwol-myeon, Naechon-ri, 21. viii. 2006, DS Choi et al., 1♀. CN: Cheongyang-gun, Namyang-myeon, Geumjeong-ri, 3. v. 2001, HW Byun, 1♂; Seosan-si, Seongyeon- myeon, Gonam-ri, 9. viii. 2006, DS Choi et al., 1♀. GN: Gimhae-si, Daedong-myeon, Suan-ri, 11. v. 2008, DS Choi, 1♀. GG: Gimpo-si, Daegot-myeon, Daebyeok-ri, 1. vi. 2008, DS Choi, 3♂2♀. GW: Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 25. vi. 2006, HW Byun, 1♂; ditto, 3. vi. 2009, HS Lee et al., 1♀; ditto, 3. vii. 2009, HS Lee et al, 1♀. JN: Suncheon-si, Byeollyang-myeon, 27. vi. 1995, HS Lee, 1♂;

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Wando-gun, Wando-eup, 7. vii. 1982, 1♀.

Distribution: Russian Far East, Korea, Japan [PA].

Biology: Unknown.

Korea: CB, CN, GN, GG, GW, JN.

47. Mesembrius niger Shiraki, 1968

Mesembrius niger Shiraki, 1968b: 225.

Korean record: Mesembrius niger: Peck 1988: 203 (Palaearctic cat.); ESK & KSAE 1994: 292 (Korean check.); Han et al. 1998b: 146 (Korean cat.); Han & Choi 2001: 163 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 19 (Korean check.). Mesembrius nigra [misspelling]: Kim, JI et al. 1983: 204 (new Korean record).

Distribution: Korea, Japan [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim et al. (1983).

Genus Phytomia Guérin-Méneville, 1833

Phytomia Guérin-Méneville in Bélanger, 1834: Voyage aux Indes Orient., Zool.: 509. TS: Eristalis chrysopygus Wiedemann (MO). Megaspis Macquart, 1842: Dipt. exot., Suppl. 2 (2): 27. TS: Eristalis chrysopygus Wiedemann (des. Brunetti, 1923: Fauna of India, Diptera, 3: 196).

48. Phytomia zonata (Fabricius, 1787) (Pl. 5J)

Syrphus zonatus Fabricius, 1787: 337. Eristalis andraemon Walker, 1849: 627. Eristalis datamus Walker, 1849: 628. Eristalis exterus Walker, 1852: 248. Eristalis inamames Walker, 1849: 627. Eristalis lata Macquart, 1842: 95.

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Eristalis latus Macquart, 1842: 90. Eristalis rufitarsisMacquart, 1842: 118. Megaspis cingulate Vollenhoven, 1863: 12 Musca sinensis Gmelin, 1790: 2872. Syrphus zonalis Fabricius, 1794: 294.

Korean record: Phytomia zonata: Knutson et al. 1975: 357 (Orient. cat.); Peck 1988: 193 (Palaearctic cat.); ESK & KSAE 1994: 292 (Korean check.); Han et al. 1998b: 142 (Korean cat.); Choi et al. 2000: 144 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 152 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 19 (Korean check.). Megaspis zonata: Shiraki 1930: 152 (redescrip.); Doi 1938a: 9 (Korean check.,); Okada 1941: 262 (Korean record); ZSK 1968: 180 (Korean check.); Kim, CW 1971: 883 (redescrip.); Kim, JI 1975: 41 (Korean check.); Kim, CW 1980: 280 (distr. Map); Kim, JI 1980: 388 (Korean check.); Megaspis zonalis: Okamoto 1924: 155 (Jeju-do, presumably first Korean record).

Description: Body length 15.5‒20.5 mm. Male. Head entirely black ground color; vertex black with black pilose; frons dark brown with black pilose; lunule shiny brown; face covered almost silver white pilose; eye holoptic, bare; antennal scape and pedicel black with white setulae; basoflagellomere black, oval; arista black. Thorax entirely black ground color; scutum dense yellow brown pilose and black pilose posterior portion; scutellum with dense black pilose. Wing hyaline with dark brown tinge medially; halter yellow brown. Legs: coxae and trochanters entirely black; femora entirely black, slightly thickened; tibiae entirely black; hind tibia slightly thickened. Abdomen oval with entirely covered yellow brown dense pilose; tergite 2 mainly yellow brown with black band posterior margin; tergite 3 black with yellow brown band anterior margin; tergite 4 entirely black. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: CB: Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 3. vi. 2004, HW Byun et al., 2♂3♀. CN: Cheonan-si, Byeongcheon-myeon, Byeongcheon-ri, Sangnok Resort, 6. xi. 1999, HY Han et al., 1♂1♀. GB: Gimcheon-si, Jirye-myeon, Gyo-ri, near Gyorigyo, 3. x. 2016, DS Choi, 2♂3♀; Gyeongju- si, Naenam-myeon, Gwaneumsa, 6. x. 2011, DS Choi, 2♀. GN: Hapcheon-gun, Gaya-myeon, Haeinsa, 9. x. 2009, DS Choi, 1♀; Namhae-gun, Idong-myeon, 7. xi. 1998, JY Choi, 2♂3♀. GG: Namyangju-si, Sudong- myeon, Mt. Chungnyeongsan, 6. ix. 1980, HK Park, 1♂1♀; Yangju-si, Gwangjeok-myeon, Ugo-ri, 2. viii. 2006, DS Choi et al., 1♂; Yeoju-gun, Yeoju-eup, Sinjin-ri, Namhangang river side, 12. x. 2002, DS Choi & MH Lim, 3♂2♀. GW: Donghae-si, Samhwa-dong, Mureung valley, 1. ix. 2009, SW Suk & YB Lee, 1♂; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 3. ix. 2000, SK Kim et al., 1♂; Taebaek-si, Mt. Taebaeksan, from Yuilsa to 1,560 m peak, 12. viii. 2000, DS Choi et al., 1♀; Wonju- si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 24. viii. 2006, JS Lim, 1♀; Wonju-si, Panbu-myeon,

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Geumdae-ri, 30. ix. 2010, SW Suk et al., 1♀. JB: Iksan-si, Sin-dong, Wongwang Univ. Campus, 30. x. 1999, HY Han et al., 3♂. JN: Gurye-gun, Gwangui-myeon, Mt. Jirisan, Nogodan, 11. ix. 1998, HY Han & KE Ro, 1♂1♀. JJ: Namjeju-gun, Pyoseon-myeon, Namjoro swimteo, near 1118 road, 19. x. 2000, HY Han et al., 1♂1♀.

Distribution: Russian Far East, China, Korea, Japan, Oriental Region [PA, OR].

Biology: Unknown.

Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

Genus Pseuderistalis Shiraki, 1930

Pseuderistalis Shiraki, 1930: Mem. Fac. Agric. Taihoku imp. Univ. 1: 147 (as subgenus of Eristalis). TS: Pseuderistalis bicolor Shiraki (OD).

49. Pseuderistalis bicolor (Shiraki, 1930)

Eristalis bicolor Shiraki, 1930: 148.

Korean record: Pseuderistalis bicolor: ZSK 1968: 180 (Korean check.); Kim, CW 1971: 883 (redescrip.); Kim, JI 1975: 41 (Korean check.); Han et al. 1998b: 143 (Korean cat.); Han & Choi 2001: 154 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 19 (Korean check.).

Distribution: Taiwan [OR], Korea? [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim (1971).

Tribe Eumerini

Genus Eumerus Meigen, 1822

Eumerus Meigen, 1822: Syst. Beschr., 3: 202. TS: Syrphus tricolor Fabricius (des. Curtis, 1839: Brit. Ent., 16: pl. 749). Paragopsis Matsumura, 1916: Thousand Ins. Japan, Add. 2: 250. TS: Paragopsis griseofasciatus Matsumura (OD) [= strigatus Fallén]. For discussion of nomenclature problems see Wirth et al. 1965: 596.

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Key to species of Eumerus from South Korea

1. Abdominal tergite 2 with yellowish or reddish quadrate spots ························· E. japonicus Matsumura − Abdominal tergite 2 with dust gray bands ·············································································· 2 2. Male eyes nearly touching for a point ································································· E. iidai Shiraki − Male eyes touching for a short space ·····························································E. strigatus (Fallén)

50. Eumerus djakonovi Stackelberg, 1952

Eumerus djakonovi Stackelberg, 1952: 378.

Korean record: Eumerus djakonovi: Peck 1988: 156 (Palaearctic cat.); Han et al. 1998b: 134 (Korean cat.); Han & Choi 2001: 138 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 20 (Korean check.).

Distribution: E. Siberia, Russian Far East, Mongolia, Korea [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Peck (1988).

51. Eumerus iidai Shiraki, 1952 (Pl. 5K)

Eumerus iidai Shiraki, 1952: 9.

Korean record: Eumerus iidai: Kim, JI & Kim, CW 1972: 10 (new Korean record); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 134 (Korean cat.); Han & Choi 2001: 138 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 20 (Korean check.).

Description: Body length 6.8 mm. Male. Head very slightly wider than thorax; frons very small, very little produced, nearly half as long as wide; vertical triangle 1/4 as wide as head; eyes bare, touching for a point; antenna black ground color; basoflagellomere about as long as scape and pedicel together. Thorax: scutum subquadrate, a little longer than wide, shiny aeneous black, with two pairs of grayish tomentose longitudinal strpes; scutellum semicircular. Wing hyaline; cross vein R-M placed in the middle of the cell dm. Legs: hind femur strongly thickend, narrowed toward the both ends. Abdomen black ground color, longer than head and thorax together; tergite

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2‒4 each with pair of dust gray bands. Female. No specimens available.

Specimen examined: Korea: Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 7. vii. 1999, HY Han et al., 1♂.

Distribution: Korea, Japan (Honshu) [PA].

Biology: Unknown.

Korea: GW.

52. Eumerus japonicus Matsumura, 1916 (Pl. 5L)

Eumerus japonicus Matsumura, 1916: 18.

Korean record: Eumerus japonicus: Shiraki 1930: 96 (redescrip., presumably first Korean record); Shiraki 1968a: 16 (redescrip.); Kim, JI 1975: 40 (Korean check.); Kim, JI 1980: 383 (Korean check.); Peck 1988: 158 (Palaearctic cat.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 134 (Korean cat.); Choi et al. 2000: 144 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 139 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 20 (Korean check.).

Description: Body length 5.2‒7.2 mm. Male. Head wider than thorax; frons very small, not produced, about twice as wide as long, densely covered with silver-white pilose; vertical triangle 1/7 as wide as head; eyes bare, touching for a space about half as long as frons; antenna orange yellow ground color; basoflagellomere about as long as scape and pedicel together. Thorax: scutum longer than wide, shiny bluish black, without any longitudinal stripe; scutellum about one and half times as wide as long. Wing hyaline; about as long as thorax and abdomen together. Legs predominantly black except basal half of tibiae reddish yellow or brown and tarsi yellow brown; hind femur straight but thickened; hind tarsi enlarged but gradually narrowed apically, densely covered with silver white pilose. Abdomen black ground color, longer than head and thorax together; tergite 2 with pair of large yellow brown spots; tergite 3 and 4 each with pair of dust gray bands. Female. Similar to male except for following non-genitalic characters: eyes dichoptic; hind tarsi without silver white pilose.

Specimen examined: Korea: CB: Danyang-gun, Yeongchun-myeon, Namcheon-ri, Namcheon Campground, 15. viii. 2016, DS Choi, 1♀. CN: Cheonan-si, Ipjang-myeon, Hodang-ri, 21. v. 2006, DS Choi & HS Lee, 1♂; Taean-gun, Gonam-myeon, Janggok-ri, Haebyeonarae beach, 24. v. 2014, DS Choi, 1♀. GB: Yeongju-

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si, Buseok-myeon, Mt. Seondalsan, from Namdae-ri to 1,236 m peak, 29. vi. 1998, HY Han et al., 1♀. GG: Pocheon-si, Ildong-myeon, Gisan-ri, Daedeogsa, 23. vii. 2006, DS Choi et al., 1♂. GW: Donghae- si, Samhwa-dong, Mt. Dutasan, Mureung valley, 21. viii. 2002, DS Choi et al., 1♂5♀; Hoengseong-gun, Anheung-myeon, Mt. Chiaksan, from Gangrim 4-ri to Cheonjibong (1,086 m) peak, 16. v. 1999, DS Choi et al., 1♀; Inje-gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 16. vi. 2004, HW Byun et al., 1♂; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 25. vii. 2005, HW Byun, 1♀; Wonju- si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 20. vii. 2004, OY Lim & HS Lee, 1♀. JN: Haenam-gun, Samsan-myeon, Mt. Duryunsan, 9. viii. 2003, HY Han et al., 1♂. JJ: Bukjeju-gun, Aewol- eup, 1,100 goji, 7. viii. 2003, HY Han et al., 1♂1♀; Bukjeju-gun, Jocheon-eup, Mt. Hallasan, 5. viii. 2003, DS Choi & HW Byun, 1♂.

Distribution: Russian Far East (Sakhalin, Primorye Territory) to Korea, Japan (Honshu, Shikoku) [PA].

Biology: Unknown.

Korea: CB, CN, GB, GG, GW, JN, JJ.

53. Eumerus kongosanensis Shiraki, 1930

Eumerus kongosanensis Shiraki, 1930: 93.

Korean record: Eumerus kongosanenesis: Shiraki 1930: 93 (orig. desc., first Korean record); Kim, JI 1975: 40 (Korean check.); Kim, JI 1980: 383 (Korean check.); Peck 1988: 158 (Palaearctic cat.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 135 (Korean cat.); Han & Choi 2001: 140 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 20 (Korean check.).

Distribution: Korea [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Shiraki (1930).

54. Eumerus strigatus (Fallén, 1817) (Pl. 6A)

Pipiza strigata Fallén, 1817: 61. Eumerus aeneus Macquart, 1829: 269. Eumerus grandicornis Meigen, 1822: 208. Eumerus lunulatus Meigen, 1822: 209. Eumerus melanopus Rondani, 1857: 90.

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Eumerus planifrons Meigen, 1822: 209. Eumerus rufitarsisStrobl, 1906: 329. Musca lineata Geoffroy, 1785: 485. Syrphus acanthodes Rossi, 1794: 63. Syrphus spinipes Rossi, 1794: 143.

Korean record: Eumerus strigatus: Kim, CW et al. 1974: 246 (DMZ, presumably first Korean record); Kim, JI 1975: 40 (Korean check.); Kim, JI 1980: 383 (Korean check.); Peck 1988: 163 (Palaearctic cat.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 135 (Korean cat.); Han & Choi 2001: 140 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 20 (Korean check.).

Description: Body length 7.2 mm. Male. Head wider than thorax; frons very short, about 1/3 as long as wide, entirely covered with dense white dust pruinosity; vertical triangle 1/5 as wide as head; eyes pilose; eye contiguity as long as frons; antenna black ground color; basoflagellomere a little longer than scape and pedicel together. Thorax: scutum about as wide as long, shiny aeneous black, with a pair of whitish tomentose longitudinal strpes; scutellum about twice as wide as long. Wing hyaline. Legs predominantly black except basal half of tibiae and tarsi reddish yellow; hind tibiae rather curved, conspicuously enlarged. Abdomen black ground color, about as long as head and thorax together; tergite 2‒4 each with pair of dust gray bands. Female. No specimens available.

Specimen examined: Korea: JN: Yeosu-si, Samsan-myeon, Geomundo, 12. vi. 1984, JI Kim, 2♂.

Distribution: Palaearctic Region, Nearctic Region (immigrant) [PA, NE].

Biology: In UK, this species are known in the larval stage where they are sometimes pests of Narcissus bulbs (Amaryllidaceae) and are known as the ‘lesser bulb flies’ on this account (Rotheray, 1993).

Korea: JN.

Genus Merodon Meigen, 1803

Merodon Meigen, 1803: Mag. Insektenk., 2: 274. TS: Sryphus clavipes Fabricius (des. Westwood 1840, Synopsis: 137). Lampetia Meigen, 1800: Nouve. Class.: 34 (originally without included species). TS: Syrphus clavipes Fabricius (des. Coquillett, 1910: Proc. U.S. nat. Mus., 37: 557). Exmerodon Becker, 1913: Annu. Mus. zool. Acad. Sci. St.-Pétersb., 17: 604 (as subgenus of Merodon Meigen). TS: Exmerodon fulcratus Becker (MO).

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55. Merodon equestris (Fabricius, 1794)

Syrphus equestris Fabricius, 1794: 292. Eristalis ferrugineus Fabricius, 1805: 240. Eristalis narcissi Fabricius, 1805: 239. Merodon bulborum Rondani, 1845: 256. Merodon constans Wiedemann, 1822: 354. Merodon nigrithorax Bezzi, 1900: 89. Merodon nobilis Meigen, 1822: 353. Merodon transversalis Wiedemann, 1822: 354. Merodon tuberculatus Rondani, 1845: 256. Merodon validus Wiedemann, 1822: 365. Musca bombyliformis Geoffroy, 1785: 479. Syrphus flavicansFabricius, 1794: 292.

Korean record: The following records are mostly misidentifications of Mallota spp. (see “Remarks”; see also Han et al. 1998b). Merodon equestris: Kim, JI 1980: 388 (Korean check.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 135 (Korean cat.); Han & Choi 2001: 140 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 20 (Korean check.). Lampetia equestris: Kim, CW 1971: 846 (redescrip.); Kim, JI 1975: 41 (Korean check.); Kim, CW 1980: 278 (distr. Map).

Distribution: Europe, North Africa (Morocco, Egypt), Nearctic Region (immigrant) [PA, NE]. Biology: Larva of this species living in plant bulbs and attacks a variety of plants in gardens (Rotheray, 1993) Remarks: We could not find specimens but cited this species based on Kim (1971). According to the distribution range of M. equestris, occurrence of this species in Korea is highly doubtful. Many records of this species, as far as we confirmed, are misidentifications of Mallota spp., most commonly M. tricolor. This is probably because incorrect illustration and Description were given in Kim (1971) (Han et al., 1998b; Han and Choi, 2001).

56. Merodon kawamurai Matsumura, 1916 (Pl. 6B)

Merodon kawamurai Matsumura, 1916: 256. Lampetia micromegas Hervé-Bazin, 1929: 111.

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Korean record: Merodon kawamurai: ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 136 (Korean cat.); Han & Choi 2001: 142 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 21 (Korean check.). Merodon kawamura [misspelling]: Kim, JI & Kim, CW 1972: 11 (new Korean record); Kim, JI 1975: 41 (Korean check.); Kim, JI 1980: 388 (Korean check.).

Description: Body length 10.5‒12.3 mm. Male. Head as wide as thorax, entirely black ground color; frons black with dense white pilose; vertical triangle black with brown and black hairs mixed; face black with dense white pilose; lunule shiny black; eyes dense pilose; eye contiguity shorter than vertical triangle; antenna dark brown ground color; scape and pedicel dark brown with brown setulae; basoflagellomere oval, brown; arista brown. Thorax shiny black ground color; scutum entirely black with dense brown pilose; scutellum shiny black with brown pilose and white pilose posterior margin. Wing hyaline; halter brown. Legs almost entirely shiny black except tarsi; hind femur strongly thickened and curved with ventral spine at about apical 1/4. Abdomen black ground color; tergite 2‒4 each pair of dust gray bands; tergite 2 with pair of yellow brown large spots laterally. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: GG: Gwangju-si, Docheok-myeon, Mt. Taehwasan, 5. v. 2001, SK Kim & OY Lim, 1♂; Namyangju-si, Sudong-myeon, Mt. Cheonmasan, 30. iv. 1972, JI Kim, 1♀; ditto, 18. iv. 1981, JI Kim, 1♂; Yangju-gun, Aengmubong, 14. iv. 1974, JI Kim, 1♂. JN: Jangheung-gun, Gwansan-eup, Mt. Cheongwansan, 14. v. 2000, DS Choi et al., 3♂1♀; Jangseong-gun, Bukha-myeon, Mt. Naejangsan, Cheongryuam Temple, 21. iv. 2013, DS Choi, 2♂; Suncheon-si, Songgwang-myeon, Mt. Jogyesan, 23. v. 1988, P.H., 1♀. Distribution: Korea, Japan [PA]. Biology: Unknown. Korea: GG, JN.

Genus Psilota Meigen, 1822

Psilota Meigen, 1822: Syst. Beschr., 3: 256. TS: Psilota anthracina Meigen (MO)

57. Psilota brevicornis Shiraki, 1968

Psilota brevicornis Shiraki, 1968a: 56.

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Korean record: Psilota brevicornis: Kim, JI et al. 1983: 205 (new Korean record); Peck 1988: 176 (Palaearctic cat.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 137 (Korean cat.); Han & Choi 2001: 142 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 21 (Korean check.).

Description (modified from Kim et al., 1983): Body length about 8 mm. Male. Head about as wide as the thorax; frons well produced, with a middle longitudinal sulcus, covered whit white tomentum, clothed with long white hairs; lunule shiny black with narrowly; vertical triangle shiny black, about 1/7 as wide as the head, one and half times as long as wide, with long black hairs; face broadend weakly to the below, with grayish white tomentum and long white hairs. Eyes holoptic, dense pilose; antenna entirely black ground color. Thorax almost entirely shiny black with long blackish yellow hairs densely; scutellum about twice as wide as long. Abodomen shiny black with long yellowish pubescence which was more blackish on the tergite 3 and 4. Female. No specimens available.

Specimen examined: Korea: GG: Yangju-gun, Aengmubong, 14. iv. 1974, JI Kim, 1♂. Distribution: Korea, Japan [PA]. Biology: Unknown. Korea: GG.

Tribe Milesiini

Genus Blera Billberg, 1820

Blera Billberg, 1820: Enum. Insect.: 118. TS: Musca fallax Linnaeus (des. Johnson, 1911: Psyche, 18: 73). Cynorhina Williston, 1887: Bull. U.S. natn. Mus., 31: 209 (as subgenus of Criorhina Meigen). TS: Milesia analis Macquart (OD). Cynorrhina Verrall, 1901: British flies, 8: 589; unjustified emendation.

58. Blera japonica (Shiraki, 1930) (Pls. 6C, D)

Cynorrhina japonica Shiraki, 1930: 60.

Korean record: Blera japonica: Kim, JI et al. 1983: 207 (new Korean record); Peck 1988: 205 (Palaearctic cat.); ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998a: 11 (Yonsei Univ. Maeji Campus); Han et

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al. 1998b: 146 (Korean cat.); Choi et al. 2000: 144 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 163 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 21 (Korean check.).

Description (modified from Kim et al., 1983): Body length 12.8‒15.2 mm. Male. Body medium size, entirely shiny black with three pairs of yellow spots on the shiny cupreous black abdomen. Head very slightly broader than thorax; frons shiny black, greatly produced; clothed with black pubescence except for the whitish eye margins and the frontal process which have a fine longitudinal strip; vertical triangle shorter than frons; antenna black ground color; basoflagellomere brownish black; arista brown. Thorax shiny black ground color with covered yellow brown hairs densely. Wing hyaline with yellow brown tinge; halter yellow brown. Legs: coxae and trochanters black; femora entirely black; fore and midtibia and tarsi yellow brown; Hind tibia yellow with 1/3 of black at middle. Abdomen entirely shiny black ground color with yellow brown pilose; tergite 2‒4 each pair of yellow spots larerally. Female. Similar to male except for following non-genitalic characters: eyes dichoptic; abdominal bands larger than male.

Specimen examined: Korea: CB: Boeun-gun, Naesongni-myeon, Mt. Songnisan, Cheonhwangbong (1,057 m), 19. v. 2007, DS Choi, 1♀. GB: Yeongju-si, Sunheung-myeon, Mt. Sobaeksan, 11. vi. 2004, HY Byun et al., 1♂. GG: Gapyeong-gun, Buk-myeon, Mt. Myeongjisan, 19. vi. 2006, DS Choi et al., 1♀. GW: Hoengseong-gun, Anheung-myeon, Mt. Chiaksan, from Gangrim 4-ri to Cheonjibong (1,086 m) peak, 5. vi. 2005, DS Choi et al., 1♀; Hongcheon-gun, Nae-myeon, Mt. Gachilbong, from Sambongyaksu to 1,240 m peak, 22. v. 1999, HY Han & KE Ro, 3♂8♀; ditto, 19. v. 2005, DS Choi & SMR Hwang, 1♀; Hongcheon- gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 31. v. 2004, HW Byun et al., 1♀; Pyeongchang-gun, Jinbu-myeon, Mt. Odaesan, from Bukdaesa to Sangwangbong (1,491 m) peak, 17. vi. 2006, DS Choi & HW Byun, 1♂; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 22. vi. 2006, JS Lim, 1♀; ditto, 25. vi. 2008, DJ Cha, 1♂; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 23. v. 2008, JS Lim et al., 1♀; Wonju-si, Sillim-myeon, Mt. Chiaksan, from Seongnam- ri to Namdaebong (1,181.5 m) peak, 28. v. 2005, HW Byun et al., 1♀; Yeongwol-gun, Buk-myeon, Mt. Sambangsan, Noron-ri, 20. v. 2001, DS Choi et al., 1♂. Distribution: Russian Far East (Sakhalin, Kuril Is, Primorye Territory), Korea, Japan (Hokkaido, Honshu) [PA]. Biology: Unknown. Korea: CB, GB, GG, GW.

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59. Blera nitens (Stackelberg, 1923)

Cynorrhina nitens Stackelberg, 1923: 22. Cynorrhina pallipes Stackelberg, 1928: 254.

Korean record: Blera nitens: Peck 1988: 205 (Palaearctic cat., presumably first Korean record); Han et al. 1998b: 146 (Korean cat.); Han & Choi 2001: 164 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 22 (Korean check.).

Distribution: Russia, Korea [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Peck (1988).

Genus Criorhina Meigen, 1822

Criorhina Meigen, 1822: Syst. Beschr., 3: 236 (as "Criorhina Hgg."). TS: Syrphus asilicus Fallén, (des. Westwood, 1840: Synopsis: 136). Criorrhina Verrall in Scudder, 1882: Nomencl. Zool.: 90; unjustified emendation. Penthesilea Meigen, 1800: Nouve. Class.: 35. TS: Musca ruficauda de Geer (by subsequent monotypy: Bezzi in Hendel, 1908: Verh. zool.-bot. Ges. Wien, 58: 67; misidentification) [= ranunculi (Panzer)]. Suppressed by I.C.Z.N. 1963: Opinion 678: 339.

60. Criorhina brevipila Loew, 1871

Criorrhina brevipila Loew, 1871: 235. Criorrhina montivaga Violovitsh, 1973: 114.

Korean record: Criorhina brevipila: Peck 1988: 207 (Palaearctic cat., presumably first Korean record); Han et al. 1998b: 147 (Korean cat.); Han & Choi 2001: 165 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 22 (Korean check.).

Distribution: Russia, Mongolia, Korea? [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Peck (1988).

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61. Criorhina trifasciata Shiraki, 1930

Criorhina trifasciata Shiraki, 1930: 56.

Korean record: Criorhina trifasciata: ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998b: 147 (Korean cat.); Han & Choi 2001: 166 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 22 (Korean check.). Criorhina trifasciatus: Kim, JI 1980: 386 (Korean check.). Penthesilea trifasciatus: Kim, JI 1974: 9 (new Korean record); Kim, JI 1975: 40 (Korean check.).

Distribution: Korea?, Taiwan [PA, OR].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim (1974).

Genus Macrozelima Stackelberg, 1930

Macrozelima Stackelberg, 1930: Konowia, 9(3): 229. TS: Zelima hervei Shiraki (as Macrozelima "bidentata sp. nov.", nomen nudum) (OD).

62. Macrozelima hervei (Shiraki, 1930) (Pls. 6E, F)

Zelima hervei Shiraki, 1930: 86. Macrozelima bidentata Stackelberg, 1930: 229.

Korean record: Macrozelima hervei: Choi et al. 2000: 145 (Baegunsan & Yonsei Univ. Maeji Campus, first Korean record); Han & Choi 2001: 167 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 22 (Korean check.).

Description: Body length 18.5‒22.5 mm. Male. Head very slightly broader than thorax; frons entirely yellow brown; vertical triangle dark brown; eyes dichoptic; eye contiguity shorter than vertical triangle; antenna dark brown ground color; basoflagellomere about as long as scape and pedicel together; arista dark brown. Thorax: scutum longer than wide, dark brown ground color with pair of dust gray longitudinal strip; postpronotum yellow brown; scutellum dark brown. Wing hyaline with dark brown tinge; halter yellow brown. Legs predominantly brown to dark brown; femora dark brown; hind femur with spine apical 1/5. Abdomen elongate, longer than head

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and thorax together, dark brown ground color; tergite 4 entirely black. Female. Similar to male except for following non-genitalic characters: eyes dichoptic; abodominal tergite 3 with triangular black band posteriorly.

Specimen examined: Korea: GW: Hoengseong-gun, Anheung-myeon, Mt. Chiaksan, from Gangrim 4-ri to Cheonjibong (1,086 m) peak, 10. viii. 1999, DS Choi et al., 2♂; Inje-gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 21. viii. 1998, HY Han et al., 1♂; Pyeongchang-gun, Jinbu-myeon, Mt. Odaesan, from Woljeongsa to Birobong (1,563 m) peak, 28. vii. 2004, DS Choi et al., 1♂; Pyeongchang-gun, Jinbu-myeon, Mt. Odaesan, from Bukdaesa to Sangwangbong (1,491 m) peak, 20. vii. 2004, DS Choi et al., 1♂; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 7. vii. 1997, HY Han et al., 1♂1♀.

Distribution: Russian Far East, Korea, Japan [PA].

Biology: Unknown.

Korea: GW.

Genus Matsumyia Shiraki, 1949

Matsumyia Shiraki, 1949: Insecta matsum., 17 (1): 1. TS: Priomerus jesoensis Matsumura (OD).

63. Matsumyia jesoensis (Matsumura, 1911)

Priomerus jesoensis Matsumura, 1911: 78.

Korean record: Matsumyia jesoensis: Kim, JI 1981: 353 (Sudong-myeon, presumably first Korean record); Choi et al. 2000: 147 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 167 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 22 (Korean check.).

Distribution: Russian Far East (Sakhalin, Kunashir), Korea, Japan (Hokkaido, Honshu) [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim (1981).

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64. Matsumyia nigrofacies Shiraki, 1949 (Pl. 6G)

Matsumyia nigrofacies Shiraki, 1949: 2.

Korean record: Matsumyia nigrofacies: ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998b: 147 (Korean cat.); Choi et al. 2000: 145 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 167 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 22 (Korean check.). Matsumyia nigrofascies [misspelling]: Kim, JI & Kim, CW 1972: 11 (new Korean record); Kim, JI 1975: 40 (Korean check.). Matsumyia nigrifascies [misspelling]: Kim, JI 1980: 386 (Korean check.).

Description: Body length 16.5‒20.2 mm. Male. Head entirely black ground color; face with grayish white pilose except central knob; eyes holoptic, bare; eye contiguity about 1/3 as long as vertical triangle; antenna yellow brown ground color; pedicel shorter than scape; basoflagellomere as long as pedicel; arista dark brown, bare. Thorax slightly longer than wide with dense yellow brown pilose with a pair of very narrow non-pubescent median longitudinal stripes; scutellum with long grayish yellow pilose densely. Wing hyaline with dark brown tinge medially; vein R4+5 straight. Legs entirely black; femora moderately thickened. Abdomen black ground color, as long as head and thorax together; tergite 2 and 3 black with broad yellowish gray band, covered dense yellow brown pilose; terigite 4 entirely black with dense yellow pilose. Female. Similar to male except for following non-genitalic characters: eyes dichoptic; hind femur distinctly narrower than male.

Specimen examined: Korea: GB: Yeongju-si, Sunheung-myeon, Mt. Sobaeksan, 11. vi. 2004, HW Byun et al., 6♂. GG: Namyangju-si, Sudong-myeon, Mt. Chungnyeongsan, 30. vi. 1999, HY Han et al., 1♂. GW: Hoengseong-gun, Anheung-myeon, Mt. Chiaksan, from Gangrim 4-ri to Cheonjibong (1,086 m) peak, 5. vi. 2005, DS Choi et al., 1♂; Hongcheon-gun, Nae-myeon, Mt. Gachilbong, from Sambongyaksu to 1,240 m peak, 3. vii. 2000, HY Han et al., 7♂1♀; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 7. viii. 2004, HW Byun et al., 1♀; ditto, 23. vi. 2005, 1♂; Inje-gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 16. vi. 2004, HW Byun et al., 9♂; Jeongseon-gun, Nam- myeon, Mt. Mindungsan, form Yupyeong-ri to 1,119 m peak, 10. vi. 2007, HY Han et al., 2♂; Wonju-si, Gwirae-myeon, from Cheoneunsa to Mt. Sipjabong peak, 7. vi. 1998, DS Choi & SK Kim, 2♀. JB: Muju- gun, Mujugucheondong, 11. vi. 1972, JI Kim, 1♀.

Distribution: Russian Far East (Sakhalin, Kunashir), Korea, Japan (Hokkaido, Honshu) [PA].

Biology: Unknown.

Korea: GB, GG, GW, JB.

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Genus Milesia Latreille, 1804

Milesia Latreille, 1804: Nouv. dict. hist. nat., 24 (Tabl. méth. Ins.): 194. TS: Syrphus crabroniformis Fabricius (des. Williston, 1887: Bull. U.S. natn. Mus., 31: 254). Suspension of rules and designation by I.C.Z.N. required (see Thompson, 1972: Archos Zool. Est. S Paulo, 23 (2): 176). Sphixea Rondani, 1845: Nuovi Ann. Sci. nat. Ist. Bologna, (2) 2: 455. TS: Eristalis fulminans Fabricius (OD) [= semiluctifera (Villers)].

65. Milesia undulata Vollenhoven, 1863 (Pls. 6H, I)

Milesia undulata Vollenhoven, 1863: 12.

Korean record: Milesia undulata: Kim, JI 1974: 8 (new Korean record); Kim, JI 1975: 41 (Korean check.); Kim, JI 1980: 386 (Korean check.); ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998b: 148 (Korean cat.); Han & Choi 2001: 168 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 23 (Korean check.).

Description: Body length 20.8‒23.5 mm. Male. Head very slightly broader than thorax; face yellow to yellow orange with yellow pilose; eyes holoptic, bare; eye contiguity about 1/4 as long as vertical triangle; antenna orange yellow ground color; pedicel about 1.5 times as long as scape; basoflagellomere oval, longer than pedicel; arista orange yellow. Thorax black ground color with yellow brown pilose; scutum longer than wide with pair of yellow brown longitudinal strip; postpronotum yellow brown; scutellum entirely black. Wing hyaline with dark brown tinge anterior half; Legs entirely brown to dark brown; femora entirely dark brown except apical 1/4 brown of hind femur; tibiae entirely brown except basal 1/2 dark brown of hind femur. Abdomen longer than head and thorax together; tergite 2‒4 each with black angulately waved middle transverse band (as in figure). Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: CN: Daejeon, Seo-gu, Wolpyeong-dong, Wolpyeong park 2 site (Malaise Trap), 12. v.-24. vi. 2008, JW Lee, 1♂; Geumsan-gun, 26. vii. 1960, JW Lee, 1♂. GN: Gimhae-si, Sangdong- myeon, Mae-ri, Sinchon, 6. vi. 2008, JC Jeong, 1♀; Namhae-gun, Samdong-myeon, Bonghwa-ri, 16. vi. 1999, JH Seol, 1♀; Ulju-gun, Samnam-myeon, Gacheon-ri, Mt. Yeongchuksan, 29. vi. 2003, HY Han et al., 1♀. GW: Hoengseong-gun, Anheung-myeon, Mt. Chiaksan, from Gangrim 4-ri to Cheonjibong (1,086 m) peak, 10. viii. 1999, DS Choi et al., 1♂; -si, Dogye-eup, Sangdeok-ri, 1. viii. 2003, DS Choi et al., 1♀. JB: Namwon-si, Jucheon-myeon, Yonggung-ri, Jangan reservoir, 29. vii. 2013, DS Choi, 1♂.

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Distribution: Russian Far East (?Sakhalin), Korea, Japan (Hokkaido, Honshu, Shikoku, Kyushu) [PA].

Biology: Unknown.

Korea: CN, GN, GW, JB.

Genus Narumyia Shiraki, 1952

Narumyia Shiraki, 1952: Mushi, 23 (1): 10. TS: Narumyia narumii Shiraki (OD)

66. Narumyia narumii Shiraki, 1952

Narumyia narumii Shiraki, 1952: 11.

Korean record: Narumyia narumii: Choo 1987: 118 (collected from peach and plum blossom, presumably first Korean record); Han et al. 1998b: 148 (Korean cat.); Han & Choi 2001: 167 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 23 (Korean check.).

Distribution: Korea?, Japan (Honshu) [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Choo (1987).

Genus Pterallastes Loew, 1863

Pterallastes Loew, 1863: Berl. ent. Z., 7: 317. TS: Pterallastes thoracicus Loew (des. Osten Sacken, 1875: Bull. Buffalo Soc. nat. Sci., 3: 64). Pseudozetterstedtia Shiraki, 1930: mem. Fac. Agric. Taihoku imp. Univ., 1 (1): 199 (as subgenus of Mallota Meigen). TS: Pseudozetterstedtia unicolor Shiraki (OD).

67. Pterallastes unicolor (Shiraki, 1930) (Pl. 6J)

Pseudozetterstedtia unicolor Shiraki, 1930: 200.

Korean record: Pterallastes unicolor: ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998b:

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148 (Korean cat.); Choi et al. 2000: 142 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 169 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 23 (Korean check.). Pseudozetterstedtia unicolor: Kim, CW 1971: 887 (redescrip., first Korean record). Pseudozetterstedt unicolor [misspelling]: Kim, JI 1975: 41 (Korean check.). Pseudozetterstedtia unicorlor [misspelling]: Kim, JI 1980: 388 (Korean check.).

Description: Body length 15.1‒17.5 mm. Male. Head as wide as thorax; vertical triangle entirely black; frons moderately produced with dense yellowish white pilose; face about one-third as wide as head, moderately broadened below with dense yellowish white pilose; eyes holoptic, bare; eye contiguity about 1/5 as long as vertical triangle; antennal scape and pedicel dark brown; basoflagellomere yellow brown, as long as scape and pedicel together. Thorax longer than wide with dense yellow brown pilose; scutellum black with long grayish yellow pilose densely. Wing hyaline with dark brown tinge medially and around vein r-m; vein R4+5 distinctly curved. Legs entirely black except apex of femora, basal portion of tibiae and midtarsi yellow brown; hind femur straight with dorsal margin slightly arcuate. Abdomen black ground color with dense brown pilose, longer than head and thorax together; tergite 2 with a pair of large transverse yellowish gray dusted lateral spots. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: CB: Danyang-gun, Mt. Sobaeksan, Yeonhwa 1 bong peak, 21. vii. 1974, JI Kim, 1♂. GG: Gapyeong-gun, Buk-myeon, Mt. Myeongjisan, Ikgeun-ri, 8. vii. 2001, DS Choi et al., 1♂. GW: Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 1. viii. 2006, HW Byun et al., 3♂3♀; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 7. viii. 2004, HW Byun et al., 3♂2♀; Inje-gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 4. viii. 2006, DS Choi et al., 3♂2♀; Jeongseon-gun, Nam-myeon, Mt. Mindungsan, form Yupyeong- ri to 1,119 m peak, 8. vii. 2005, HY Han et al., 3♂1♀; Wonju-si, Gwirae-myeon, from Cheoneunsa to Mt. Sipjabong peak, 27. vi. 2008, SW Suk & YB Lee, 1♀; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 10. x. 2006, JS Lim, 1♂; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 6. viii. 2008, DJ Cha et al., 1♀; ditto, 2. viii. 2000, DS Choi et al., 1♂.

Distribution: Russian Far East (Kuril Is: Kunashir), Korea, Japan (Hokkaido, Honshu) [PA].

Biology: Unknown.

Korea: CB, GG, GW.

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Genus Spilomyia Meigen, 1803

Spilomyia Meigen, 1803: Mag. Insektenk., 2: 273. TS: Musca diophthalma Linnaeus (as Syrphus diophthalmus Fabricius) (des. Coquillett, 1910: Proc. U.S. natn. Mus., 37: 607).

68. Spilomyia permagna Stackelberg, 1958

Spilomyia permagna Stackelberg, 1958: 764.

Korean record: Spilomyia permagna: ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998b: 149 (Korean cat.); Han & Choi 2001: 171 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 23 (Korean check.).

Distribution: Russian Far East (Sakhalin, Kuril Is), Japan, Korea? [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on ESK & KSAE (1994).

69. Spilomyia suzukii Matsumura, 1916 (Pls. 6K, L)

Spilomyia suzukii Matsumura, 1916: 229.

Korean record: Spilomyia suzukii: Doi 1938a: 10 (Korean check., presumably first Korean record); ZSK 1968: 180 (Korean check.); Kim, CW 1971: 885 (redescrip.); Kim, JI 1975: 41 (Korean check.); ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998b: 149 (Korean cat.); Choi et al. 2000: 145 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 171 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 23 (Korean check.). Spilomyia matsumurai [error]: Kim, JI 1980: 385 (Korean check.); Han et al. 1998b: 149 (Korean cat.).

Description: Body length 16.8‒22.5 mm. Male. Head yellow brown ground color; vertical triangle shiny black posteriorly and yellow brown anteriorly; face entirely shiny yellow brown with silver white pilose; lunule shiny black; eyes holoptic, bare; antenna entirely dark brown ground color; scape and pedicel with black setulae; basoflagellomere shorter than pedicel. Thorax black ground color; scutum a pair of yellow brown round spots anteriorly and diagonal spots posteriorly; postpronotum yellow brown; scutellum black with yellow brown band on hind margin.

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Wing hyaine with dark brown tinge medially; Legs dark brown ground color; femora dark brown; tibia dark brown apical 1/2 and yellow brown basal 1/2; hind femur with spine apical 1/5 portion. Abdomen longer than head and thorax together; tergite 2‒4 each a pair of yellow brown band medially and thin yellow brown band porterior margin. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: CB: Cheongwon-gun, Namseong-myeon, Mt. Ingyeongsan, 15. viii. 1997, 1♂; Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 29. vii. 2004, HW Byun & JS Soh, 13♂. GB: Cheongsong- gun, Budong-myeon, Mt. Juwangsan (721 m), 10. ix. 2006, DS Choi, 1♂. GN: Geoje-si, Dongbu-myeon, Hakdong, 6. viii. 2004, TW Kim, 1♀. GG: Yangpyeong-gun, Danwol-myeon, Saneum Recreation forest, 11. ix. 2009, DS Choi, 1♂; Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 9. viii. 2003, OY Lim & HS Lee, 1♀; Jeongseon-gun, Nam-myeon, Mt. Mindungsan, form Yupyeong-ri to 1,119 m peak, 29. viii. 2005, HY Han et al., 1♂; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 2.-24. viii. 2004, OY Lim, 1♀; ditto, 5. x. 2005, SMR Cha & DJ Cha, 1♀; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 4. ix. 1999, DS Choi & CH Park, 1♀.

Distribution: Russian Far East (Primorye Territory, Kuril Is), Korea, Japan (Hokkaido, Honshu) [PA].

Biology: Unknown.

Korea: CB, GB, GN, GG, GW.

Genus Syritta Lepeletier and Serville, 1828

Syritta Le Peletier et Serville, 1828: Encyclop. méthod. hist. nat. (Ins.), 10 (2): 808 (as subgenus of Xylota Meigen). TS: Musca pipiens Linnaeus (MO). Coprina Zetterstedt, 1838: Insecta Lapp.: 584. TS: Musca pipiens Linnaeus (MO). Xylota Westwood, 1840: Synopsis: 136. TS: Musca pipiens Linnaeus (MO). Invalid type-species Description, cf. p. 224.

70. Syritta orientalis Macquart, 1842

Syritta orientalis Macquart, 1842: 136. Senogaster lutescens Doleschall, 1856: 410. Syritta amboinensis Doleschall, 1858: 97. Syritta illucida Walker, 1859: 121.

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Korean record: Syritta orientalis: Kim, JI 1974: 8 (new Korean record); Kim, JI 1975: 41 (Korean check.); Kim, JI 1980: 386 (Korean check.); ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998b: 150 (Korean cat.); Han & Choi 2001: 175 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 24 (Korean check.).

Distribution: Oriental Regioin [OR]. Biology: Unknown. Remarks: We could not find specimens but cited this species based on Kim (1974). Occurrence of this Oriental species in Korea is highly doubtful (Han and Choi, 2001).

71. Syritta pipiens (Linnaeus, 1758) (Pls. 7A, B)

Musca pipiens Linnaeus, 1758: 594. Musca pipines Fourcroy, 1785: 486. Spheginoides tenofemorus Dzhafarova, 1974: 40. Syritta albicincta Abreu, 1924: 125. Syritta flavicansSzilady, 1940: 68. Syritta obscuripes Strobl, 1899: 146. Syritta vicina Szilady, 1940: 67. Xylota proxima Say, 1824: 8.

Korean record: Syritta pipiens: Han & Choi 2001: 172 (Korean check., first Korean record); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 24 (Korean check.).

Description: Body length 6.5‒8.8 mm. Male. Head black ground color; vertical triangle black with brown pilose; face black with entirely covered silver pruinosity; lunule shiny black; eyes holoptic, bare; eye contiguity shorter than vertical triangle; antennal scape dark brown; pedicel brown with black setulae; basoflagellomere as long as scape and pedicel together, rounded; arista dark brown. Thorax black ground color; scutum a pair of yellow brown longitudinal stipe laterally and white gray spot anterior margin; postpronotum yellow brown; scutellum black with yellow brown pilose. Wing hyaine and clear; Legs black and dark brown; hind femur strongly thickened. Abdomen longer than head and thorax together; tergite 2 black with a pair of yellow brown spot laterally; tergite 3 black with yellow brown band anterior margin; tergite 4 entirely black. Female. Similar to male except for following non-genitalic characters: eyes dichoptic; bands on tergite 3 smaller than male; tergite 4 with a piar of whitish gray bands.

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Specimen examined: Korea: CB: Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 3. vi. 2004, HW Byun et al., 1♂; Chungju-si, Sotae-myeon, Deokeun-ri, Namhangang river side, 11. x. 2002, DS Choi et al., 2♂3♀; Yeongdong-gun, Yonghwa-myeon, Minjujisan Recreational Forest, 21. v. 2011, DS Oh, 1♂. CN: Boryeong- si, Cheongna-myeon, Mt. Oseosan, 20. vii. 1999, HY Han et al., 2♂2♀; Dangjin-gun, Sinpyeong-myeon, Geumcheon-ri, 10. viii. 2006, DS Choi et al., 1♂; Taean-gun, Gonam-myeon, Janggok-ri, Haebyeonarae beach, 24. v. 2014, DS Choi, 1♀. GB: Dalseong-gun, Yaga-myeon, Mt. Biseulsan, Recreational Forest, 20. v. 2012, DS Choi, 1♀; Gimcheon-si, Jirye-myeon, near Gyorigyo, 3. x. 2016, 3♂; Gyeongju-si, Amgok-dong, Mujangsaji, 22. vii. 2011, DS Choi, 1♂2♀; Yeongju-si, Dansan-myeon, Jwaseok-ri, 4. vi. 2016, DS Choi, 1♂; Yeongju-si, Sunheung-myeon, Mt. Sobaeksan, 27. viii. 2001, DS Choi et al., 1♀. GN: Sancheong-gun, Sicheon-myeon, Mt. Jirisan, 30. vii. 1981, HY Han, 1♂. GG: Yeoju-gun, Yeoju-eup, Sinjin-ri, Namhangang river side, 12. x. 2002, DS Choi & MH Lim, 4♂1♀. GW: Gangneung-si, Okgye-myeon, Sangye-ri, 19. vii. 2010, SW Suk et al., 1♀; ditto, 19. vii. 2010, SW Suk & YB Lee, 1♀; Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 1. viii. 2006, HW Byun et al., 1♂1♀; Hongcheon-gun, Nae- myeon, Mt. Gachilbong, from Sambongyaksu to 1,240 m peak, 2. v. 2003, HY Han & KE Ro, 1♀; ditto, 6. vi. 2015, DS Choi & SB Lee, 2♂; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 12. vi. 1998, HW Byun, 1♂; ditto, 23. vii. 2008, JS Lim, 1♂. JB: Iksan-si, Sin-dong, Wongwang Univ. Campus, 30. x. 1999, HY Han et al., 2♂3♀; Imsil-gun, Gangjin-myeon, Mt. Baekyeonsan, 6. vii. 2013, DS Choi, 2♂; Imsil- gun, Gwanchon-myeon, Hoebong-ri, Grassland, 30. vii. 2016, DS Choi, 1♀; Namwon-si, Deokgwa-myeon, Mando-ri, Mt. Gyeryongsan, 3. vi. 2013, 1♀. JN: Gokseong-gun, Gokseong-eup, Wolpyeong-ri, 15. v. 2007, DS Choi, 1♀. JJ: Namjeju-gun, Pyoseon-myeon, Namjoro swimteo, near 1118 road, 19. x. 2000, HY Han et al., 1♂3♀.

Distribution: Holarctic, Oriental Regions [PA, NE, OR].

Biology: Larvae of this species live in compost, manure, silage and other rotting organic matter (Stubbs & Falk, 1983).

Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

Genus Temnostoma Lepeletier and Serville, 1828

Temnostoma Le Peletier et Serville, 1828: Encyclop. méthod. hist. nat. (Ins.), 10 (2): 518 (as subgenus of Milesia Latreille). TS: Milesia bombylans Fabricius (des. Coquillett, 1910: Proc. U.S. natn. Mus., 37: 612). Tritonia Meigen, 1800: Nouve. Class.: 33. TS: Musca vespiformis Linnaeus (des. Coquillett, 1910: Proc. U.S. natn. Mus., 37: 617). Suppressed by I.C.Z.N. 1963, Opinion 678: 339. It would have been a junior homonym of Tritonia Cuvier. Microrhincus Lioy, 1864: Atti Ist. veneto Sci., (3) 9: 751. TS: Milesia bombylans Fabricius (des. Goffe, 1946: J. Soc. Br. Ent., 3: 30).

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72. Temnostoma bombylans (Fabricius, 1805) (Pls. 7C, D)

Milesia bombylans Fabricius, 1805: 189. Milesia zetterstedtii Fallén, 1816: 8. Temnostoma flavifemurKrivosheina, 2002: 1258?. Temnostoma japonicum Hull, 1944: 41. Temnostoma nitobei Matsumura, 1916: 199.

Korean record: Temnostoma bombylans: Doi 1938a: 10 (Korean check., presumably first Korean record); Kim, CW 1971: 886 (redescrip.); Kim, JI 1975: 40 (Korean check.); Kim, JI 1980: 386 (Korean check.); Peck 1988: 216 (Palaearctic cat.); ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998a: 11 (Yonsei Univ. Maeji Campus); Han et al. 1998b: 149 (Korean cat.); Choi et al. 2000: 145 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 176 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 24 (Korean check.). Temnostoma bomlylans [misspelling]: ZSK 1968: 180 (Korean check.). Temnostoma nitobei: Kim, JI et al. 1983: 206 (new Korean record).

Description: Body length 13.5‒17.2 mm. Male. Head black ground color; vertical triangle black with brown hairs; face entirely shiny brown; lunule shiny dark brown; eyes holoptic, bare; eye contiguity shorter than vertical triangle; antenna entirely dark brown ground color; scape and pedicel with black setulae; basoflagellomere as long as scape and pedicel together, rounded; arista dark brown. Thorax black ground color; scutum a pair of yellow brown bands medially; postpronotum yellow brown; scutellum black with black and brown pilose mixed. Wing hyaine with dark brown tinge anteriorly. Legs black ground color; fore leg entirely black; mid and hind femur entirely black; mid and hind tibae apical 1/2 black and basal 1/2 dark brown. Abdomen as long as head and thorax together; tergite 2‒4 each yellow brown band near anterior margin. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: CN: Cheonan-si, Seonggeo-eup, Cheonheung-ri, 21. v. 2006, HW Byun et al., 2♂. GB: Andong-si, Waryong-myeon, near Yesangyo, 17. v. 2008, DS Choi, 1♀. GG: Gapyeong-gun, Buk- myeon, Mt. Myeongjisan, 19. vi. 2006, DS Choi et al., 1♀; Gwangju-si, Opo-myeon, Mr. Baengmasan, 27. v. 2001, DS Choi et al., 1♂. GW: Gangneung-si, Wangsan-myeon, Mt. Seokbyeongsan, Sapdangnyeong to 1,055 m peak, 18. v. 2002, HW Byun & OY Lim, 2♂4♀; Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 5. vi. 2003, DS Choi & HS Lee, 3♂1♀; Hongcheon-gun, Nae-myeon, Mt. Gachilbong, from Sambongyaksu to 1,240 m peak, 5. vi. 2000, DS Choi, 3♂1♀; Inje-gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 29. v. 1999, HY Han et al., 12♂8♀; Pyeongchang-gun, Jinbu-myeon, Mt. Odaesan, from Bukdaesa to Sangwangbong (1,491 m) peak, 20. vii. 2004, DS Choi & HW

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Byun, 1♂1♀; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 19. vi. 2006, DS Choi & HW Byun, 1♂1♀; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 2. vi. 2003, DS Choi et al., 2♂1♀. JB: Jangsu-gun, Gyenam-myeon, Jangan-ri, Jangansa Temple, 2. vi. 2013, DS Choi, 1♀. JJ: Bukjeju-gun, Jocheon-eup, Mt. Hallasan, 26. vi. 2003, DS Choi et al., 1♂.

Distribution: Europe, Russia, Korea, Japan, North Africa (Morocco) [PA].

Biology: Unknown.

Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

73. Temnostoma frequense Matsumura, 1931

Temnostoma frequense Matsumura, 1931: 339.

Korean record: Temnostoma frequense: Doi 1938a: 10 (Korean check., presumably first Korean record); ZSK 1968: 180 (Korean check.); Kim, JI 1975: 41 (Korean check.); ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998b: 150 (Korean cat.); Han & Choi 2001: 177 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 24 (Korean check.).

Distribution: Korea?, Japan [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Doi (1938).

74. Temnostoma vespiforme (Linnaeus, 1758) (Pls. 7E, F)

Musca vespiformis Linnaeus, 1758: 593. Milesia excentrica Harris, 1835: 598. Milesia excentrica Harris, 1841: 409. Milesia wagae Gorski, 1852: 175. Spilomyia sericomyiaeforme Portschinsky, 1886: 8. Spilomyia sibiricum Portschinsky, 1886: 7. Spilomyia vulgare Portschinsky, 1886: 7. Syrphus frequens Matsumura, 1931: 354. Temnostoma aequalis Loew, 1864: 68. Temnostoma meridionale Krivosheina and Mamaev, 1962: 928. Temnostoma strigosum Sack, 1941: 191.

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Korean record: Temnostoma vespiforme: Kim, JI & Kim, CW 1972: 10 (new Korean record); Kim, JI 1975: 41Korean check.); ESK & KSAE 1994: 291(Korean check.); Han et al. 1998b: 150Korean cat.); Han & Choi 2001: 177 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 25 (Korean check.). Temnostoma vespiformis: Kim, JI 1980: 386.

Description: Body length 16.5‒19.2 mm. Male. Head yellow brown ground color; vertical triangle black posteriorly and yellow brown anteriorly; face entirely yellow brown with longitudinal shiny black band medially and ventrally; lunule shiny black; eyes holoptic, bare; antenna entirely brown ground color; scape and pedicel with brown setulae; basoflagellomere as long as scape and pedicel together. Thorax black ground color; scutum a pair of yellow brown bands medially and three yellow brown bands posterior margin; postpronotum yellow brown; scutellum black with dense long yellow brown pilose. Wing hyaine with dark brown tinge anteriorly. Legs yellow brown ground color; fore leg entirely black; mid and hind legs entirely yellow brown. Abdomen as long as head and thorax together; tergite 2‒4 each yellow brown band anterior 1/3 portion and posterior margin. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: GG: Pocheon-gun, Gwangneung, 28. v. 1972, JI Kim, 1♂; Seongnam-si, Mt. Cheonggyesan, 26. vi. 1984, YH Ko, 1♂1♀. GW: Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 25. v. 2004, HW Byun et al., 1♀; ditto, 17. vi. 2003, DS Choi & HW Byun, 1♂; Hongcheon-gun, Nae-myeon, Mt. Gachilbong, from Sambongyaksu to 1,240 m peak, 2. vi. 2007, HS Lee et al., 1♂; ditto, 12. vi. 2004, HW Byun et al., 1♂; ditto, 3. vii. 2000, HY Han et al., 1♀; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, N. Valley, 20. vi. 1999, HY Han & KE Ro, 1♂; Hongcheon-gun, Nae-myeon, Woldungol, 3. viii. 1996, HY Han & HW Byun, 1♀; Inje-gun, Sangnam-myeon, Mt. Bangtaesan, 4. vi. 1999, SS Gwak, 1♂.

Distribution: Holactic Region [PA, NE].

Biology: Larva saprophagous, boring in decaying wood (Torp, 1994).

Korea: GG, GW.

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Genus Tropidia Meigen, 1822

Tropidia Meigen, 1822: 346. TS: Eristalis milesiformis Fallén, 1817 [= Musca scitus Harris, 1780], by subsequent designation of Curtis (1832: pl. 401)

75. Tropidia rostrata Shiraki, 1930 (Pls. 7G, H)

Tropidia rostrata Shiraki, 1930: 90. Parrhyngia quadrimaculata Shiraki, 1968b: 205.

Korean record: Tropidia rostrata: Suk & Han 2014b: 230 (new Korean record); Han et al. 2014: 25 (Korean check.).

Description (modified from Suk and Han, 2014b): Body length 9.5‒11.3 mm. Male. Head largely yellowish brown; vertex brownish black with yellowish brown pruinosity posterolaterally; frons with heavy yellowish brown pruinosity except for dark brown lower median portion and narrow mid-line upward; ocellar triangle brownish black with yellowish brown hairs; gena brownish black with pale yellow hairs; eyes holoptic, bare; antennal scape and pedicel yellowish brown with brownish black hairs dorsally; basoflagellomere yellowish brown, bare; arista yellowish brown to dark brown, bare. Thorax predominatly brownish black with yellowish brown hairs, with pale yellow pruinosity; scutum shiny black with pale yellow pruinosity laterally except on middle area. Wing entirely mitotrichose, mostly hyaline except for pterostigma yellowish brown. Legs predominantly yellowish brown in ground color with some darker areas, with yellowish brown hairs; fore femur dorsally with brownish black longitudinal stripe except for basal and apical areas, ventrally with similar stripe at apical 1/2; apical 1/4 of fore tibia brownish black; fore tarsus with some brownish black hairs apically; midfemur both dorsally and ventrally with brownish black longitudinal strips at apical 1/2; midtarsus with some brownish black hairs apically; apical 3/4 of hind femur brownish black, swollen with large subapical ventral triangular process; hind tibia strongly curved dorsally at basal 1/3, with subbasal dorsal tubercle, with apex ventrally pointed, yellowish brown with apical 1/3 entirely brownish black, with brownish black ventral spot at basal 1/3, also with brownish black hairs ventrally; hind tarsus brownish black dorsally, yellowish brown ventrally. Abdomen conical shape with preabdominal tergites brownish black in ground color with brownish yellow spots larerally, with brownish black hairs except on yellowish brown areas yellowish brown hairs; tergite 2 with pair of elongated square yellowish brown spots; tergite 3 with elongated square spots; Tergite 4 predominantly brownish black with small triangular yellowish brown spot anterolaterally. Female. Similar to male except for the following non-genitalic characters: frons and face with brownish

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black transverse mid-stripe; eyes dichoptic. Abdomen narrowly oval shape.

Specimen examined: Korea: CN: Taean-gun, Sowol-myeon, Cheollipo Arboretum, 14. vi. 2007, DS Choi, 1♂. GB: Pohang-si, Buk-gu, Heunghae-eup, Gokgang-ri, 4. vi. 2011, SW Suk et al., 38♂18♀.

Distribution: Russian Far East, China, Korea, Japan [PA].

Biology: Unknown.

Korea: CN, GB.

Tribe Pipizini

Genus Pipiza Fallén, 1810

Pipiza Fallén, 1810: Specim. entom. novam Dipt.: 11. TS: Musca noctiluca Linnaeus (des. Curis, 1837: Brit. Ent., 14: 669).

Key to species of Pipiza from South Korea

1. Abdomen with a pair of yellowish spots ·············································································· 2 − Abdomen without spots in male ································································ P. austriaca Meigen 2. Abdominal spots small, ill-defined· ······························································ P. signata Meigen − Abdominal spots large, well-defined· ················································ P. flavimaculata Matsumura

76. Pipiza abdominalis (Shiraki and Edashige, 1953)

Pipizella abdominalis Shiraki and Edashige, 1953: 84.

Korean record: Pipiza abdominalis: Kim, JI et al. 1983: 204 (new Korean record); Peck 1988: 86 (Palaearctic cat.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 124 (Korean cat.); Han & Choi 2001: 106 (Korean check.); Paek et al. 2010: 230 (Korean check.); Han et al. 2014: 25 (Korean check.).

Distribution: Korea, Japan [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim (1983).

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77. Pipiza austriaca Meigen, 1822 (Pl. 7I)

Pipiza austriaca Meigen, 1822: 252.

Korean record: Pipiza austriaca: Kim, CW 1971: 859 (redescrip.); Kim, JI 1975: 39 (Korean check.); Kim, CW 1980: 289 (distr. Map); Han et al. 1998a: 10 (Yonsei Univ. Maeji Campus); Han et al. 1998b: 124 (Korean cat.); Choi et al. 2000: 143 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 106 (Korean check.); Paek et al. 2010: 230 (Korean check.); Han et al. 2014: 25 (Korean check.).

Description: Body length 8.5‒11.2 mm. Male. Head black ground color; vertical triangle entirely black; face entirely shiny black with dense silver white pilose; lunule shiny black; frons shiny black with black pilose; eyes holoptic, dense pilose; eye contiguity as long as vertical triangle; antenna scape and pedicel entirely black with black setulae; basoflagellomere dark brown, as long as scape and pedicel together; arista dark brown. Thorax entirely shiny black with dense long brown pilose. Wing hyaline with dark brown tinge medially; halter pale yellow. Legs black ground color; coxae entirely shiny black; trochanters shiny black; femora entirely black; tibia black apical 1/2 and brown basal 1/2. Abdomen black ground color, as long as head and thorax together; abdomen without any spots. Female. Similar to male except for following non-genitalic characters: eyes dichoptic; abdominal tergite 2 with a pair of yellow orange spots.

Specimen examined: Korea: GB: Yeongju-si, Buseok-myeon, Mt. Eoraesan, from Namdae-ri to 1,064 m peak, 30. vi. 1998, HY Han et al., 1♀; Yeongju-si, Sunheung-myeon, Mt. Sobaeksan, 27. viii. 2001, DS Choi et al., 2♀. GG: Namyangju-si, Sudong-myeon, Mt. Cheonmasan, 10. ix. 1972, JI Kim, 1♀; Paju-si, Bogwangsa, 17. ix. 1972, SN Choi, 1♀. GW: Chuncheon-si, Seo-myeon, Seosang-ri, 4. vi. 2006, DS Choi et al., 1♂; Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 15. viii. 2001, DS Choi et al., 2♀; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 23. viii. 2001, DS Choi et al., 1♂1♀; Hwacheon-gun, Sanae-myeon, Mt. Gwangdeoksan, 29. viii. 2000, CH Park & DW Kim, 2♀; Inje-gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 21. viii. 1998, HY Han et al., 2♂2♀; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 24. iv. 2003, DS Choi & HW Byun, 1♀. JB: Imsil-gun, Gwanchon-myeon, Hoebong-ri, Grassland, 1. x. 2016, DS Choi, 1♀. JN: Jangseong-gun, Hwangnyong-myeon, Gwandong-ri, 13. v. 2007, DS Choi, 1♀. JJ: Bukjeju-gun, Jocheon- eup, Goepyeongi oreum, 2. v. 2003, DS Choi et al., 1♂2♀.

Distribution: Europe, Russia to East Asia (Korea, Japan) [PA].

Biology: Rotheray (1993) recorded the species as a predator of Caveriella aphids on Heracleum sphlondylium (Umbelliferacae).

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Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

78. Pipiza flavimaculata Matsumura, 1918 (Pl. 7J)

Pipiza flavimaculataMatsumura, 1918: 2.

Korean record: Pipiza flavimaculata: Kim, JI & Kim, CW 1972: 8 (new Korean record); Kim, JI 1975: 39 (Korean check.); Han et al. 1998b: 124 (Korean cat.); Han & Choi 2001: 107 (Korean check.); Paek et al. 2010: 230 (Korean check.); Han et al. 2014: 26 (Korean check.). Pipiza flavomaculata [misspelling]: Kim, JI 1980: 383 (Korean check.); ESK & KSAE 1994: 290 (Korean check.).

Description: Body length 6.8‒8.8 mm. Male. Head black ground color; vertical triangle entirely black; face entirely shiny black with dense silver white pilose; lunule shiny black; frons shiny black with black pilose; eyes holoptic, dense pilose; eye contiguity as long as vertical triangle; antenna scape and pedicel entirely black with black setulae; basoflagellomere dark brown, as long as scape and pedicel together; arista dark brown. Thorax entirely shiny black with dense long brown pilose. Wing hyaline with dark brown tinge medially; halter pale yellow. Legs black ground color; coxae entirely shiny black; trochanters shiny black; femora entirely black; tibia black apical 1/2 and brown basal 1/2. Abdomen black ground color, as long as head and thorax together; terigite 2 with a pair of yellow brown triangular spots; tergite 3 and 4 entirely black without any spots. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: GB: Uljin-gun, Onjeong-ri, Mt. Baegamsan, 16. viii. 1999, HY Han et al., 1♂. GG: Gapyeong-gun, Buk-myeon, Mt. Myeongjisan, Ikgeun-ri, 22. iv. 2001, DS Choi et al., 2♂. GW: Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 11. viii. 1999, DS Choi et al., 5♀; Inje-gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 21. viii. 1998, HY Han et al., 1♀; Taebaek-si, Mt. Taebaeksan, from Yuilsa to 1,560 m peak, 12. viii. 2000, DS Choi et al., 1♂; Wonju-si, Sillim-myeon, Mt. Chiaksan, from Seongnam-ri to Namdaebong (1,181.5 m) peak, 13. viii. 2000, DS Choi et al., 1♂.

Distribution: Korea, Japan [PA].

Biology: Unknown.

Korea: GB, GG, GW.

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79. Pipiza inornata Matsumura, 1916

Pipiza inornata Matsumura, 1916: 15.

Korean record: Pipiza inornata [Probably misidentification ofP. austriaca]: Kim, JI & Kim, CW 1972: 8 (new Korean record); Kim, JI 1975: 39 (Korean check.); Kim, JI 1980: 383 (Korean check.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 124 (Korean cat.); Han & Choi 2001: 108 (Korean check.); Paek et al. 2010: 230 (Korean check.); Han et al. 2014: 26 (Korean check.).

Distribution: Korea?, Japan [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim & Kim (1972).

80. Pipiza signata Meigen, 1822 (Pl. 7K)

Pipiza signata Meigen, 1822: 246.

Korean record: Pipiza signata: Han et al. 1998a: 10 (Yonsei Univ. Maeji Campus, first Korean record); Han et al. 1998b: 125 (Korean cat.); Choi et al. 2000: 143 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 108 (Korean check.); Paek et al. 2010: 230 (Korean check.); Han et al. 2014: 26 (Korean check.).

Description: Body length 7.5‒9.2 mm. Male. Head black ground color; vertical triangle entirely black; face entirely shiny black with dense silver white pilose; lunule shiny black; frons shiny black with black pilose; eyes holoptic, dense pilose; eye contiguity as long as vertical triangle; antenna scape and pedicel entirely black with black setulae; basoflagellomere dark brown, as long as scape and pedicel together; arista dark brown. Thorax entirely shiny black with dense long brown pilose. Wing hyaline with dark brown tinge medially; halter pale yellow. Legs black ground color; coxae entirely shiny black; trochanters shiny black; femora entirely black; tibia black apical 1/2 and brown basal 1/2. Abdomen black ground color, as long as head and thorax together; terigite 2 with a pair of ill-defined spots; tergite 3 and 4 entirely black without any spots. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: GB: Uljin-gun, Onjeong-ri, Mt. Baegamsan, 16. viii. 1999, HY Han et al., 2♂. GG: Pocheon-gun, Mt. Myeongjisan, 3. v. 1997, HY Han, 1♂. GW: Hongcheon-gun, Nae-myeon, Mt.

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Gyebangsan, from Unduryeong to 1,577 m peak, 11. viii. 1999, DS Choi et al., 2♂1♀; Taebaek-si, Mt. Taebaeksan, from Yuilsa to 1,560 m peak, 29. viii. 1999, DS Choi & CH Park, 3♂1♀; Wonju-si, Heungeop- myeon, Maeji-ri, Yonsei Univ. Campus, 23. vi. 1997, HW Byun & DS Choi, 1♂; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 23. viii. 1999, DS Choi & CH Park, 4♂3♀.

Distribution: Europe, Russia, East Asia (Korea, Japan), Oriental Region (Taiwan) [PA, OR].

Biology: Unknown.

Korea: GB, GG, GW.

Genus Pipizella Rondani, 1856

Pipizella Rondani, 1856: Dipt. Ital. Prodromus, 1: 54. TS: Mulio virens Fabricius (OD). Pizopsis Matsumura, 1918: J. Coll. Agric. Hokkaido imp. Univ., 8 (1): 3. TS: Pipiza biglumis Matsumura (OD).

81. Pipizella biglumis (Matsumura, 1916)

Pipiza biglumis Matsumura, 1916: 14.

Korean record: Pipizella biglumis: Han et al. 1998b: 125 (Korean cat.); Han & Choi 2001: 109 (Korean check.); Han et al. 2014: 16 (Korean check.). Pipiza biglumis: Paek et al. 2010: 230 (Korean check.). Heringia biglumis: Kim, JI & Park 1984: 43 (anatomy).

Distribution: Korea?, Japan [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim & Park (1984).

Genus Stenopipiza Matsumura, 1919

Stenopipiza Matsumura, 1919: Ent. Mag., Kyoto, 3 (3-4): 140. TS: Stenopipiza bipunctata Matsumura (OD).

82. Stenopipiza bipunctata Matsumura, 1919

Stenopipiza bipunctata Matsumura, 1919: 141.

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Korean record: Stenopipiza bipunctata: Doi 1938a: 10 (Korean check., presumably first Korean record); Kim, CW 1971: 845 (redescrip.); Kim, JI 1975: 40 (Korean check.); Han et al. 1998b: 125 (Korean cat.); Han & Choi 2001: 109 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 26 (Korean check.). Stenopipija bipunctata [misspelling]: ZSK 1968: 180 (Korean check.).

Distribution: Korea?, Japan [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim (1971).

Genus Triglyphus Loew, 1840

Triglyphus Loew, 1840: Öffentl. Prüf. Schüler K. Friedrich-Wihelms-Gymnasiums zu Posen, 1840: 30 and Isis (Oken's), 1840 (VII-VIII): 565. TS: Triglyphus primus Loew (OD).

83. Triglyphus primus Loew, 1840 (Pl. 7L)

Triglyphus primus Loew, 1840: 30.

Korean record: Triglyphus primus: Shiraki 1930: 138 (redescrip., presumably first Korean record); ZSK 1968: 180 (Korean check.); Kim, CW 1971: 858 (redescrip.); Peck 1988: 83 (Palaearctic cat.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 125 (Korean cat.); Han & Choi 2001: 110 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 26 (Korean check.). Triglypus primus [misspelling]: Kim, JI 1975: 39 (Korean check.); Kim, JI 1980: 383 (Korean check.).

Description: Body length 6.8‒8.2 mm. Male. Body almost entirely black ground color. Vertical triangle entirely black; face entirely shiny black with dense silver white pilose; lunule shiny black; frons shiny black with black pilose; eyes holoptic, dense pilose; eye contiguity longer than vertical triangle; antenna scape and pedicel entirely black with black setulae; basoflagellomere dark brown, as long as scape and pedicel together; arista dark brown. Thorax entirely shiny black with dense long brown pilose. Wing hyaline with dark brown tinge on cell SC; wing length as long as body length; halter yellow brown. Legs black ground color; coxae entirely shiny black; trochanters shiny black; femora entirely black; fore and midtibiae black apical 2/3 and brown basal 1/3; hind tibiae entirely black. Abdomen black ground color, as long as head and thorax together; each tergite with dense pilose; tergite 4 not visible in dorsal view.

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Female. Similar to male except for following non-genitalic characters: eyes dichoptic; abdominal tergite 4 visible in dorsal view.

Specimen examined: Korea: CB: Jecheon-si, Songgye-ri, Jeolgol, 10. v. 1997, HY Han et al., 1♂. GB: Mungyeong-si, Mt. Juheulsan, from Hyeguksa to 1,106 m peak, 11. viii. 2004, HW Byun et al., 1♀. GW: Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 7. viii. 2004, HW Byun et al., 1♂; Wonju-si, Gwirae-myeon, from Cheoneunsa to Mt. Sipjabong peak, 5. v. 2004, HW Byun et al., 1♂; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 5. vii. 2004, DS Choi et al., 1♂1♀; ditto, 28. iv. 2004, HW Byun et al., 2♂1♀; ditto, 5. vii. 2004, DS Choi et al., 1♂; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 12. vii. 1998, DS Choi, 1♀; ditto, 29. vii. 1998, DS Choi & SK Kim, 1♂2♀. JB: Muju-gun, Mujugucheondong, 10. vi. 1972, JI Kim, 1♀.

Distribution: Europe, Russia, Korea, Japan [PA].

Biology: Torp (1994) recorded larva of this species live in the galls of Cryposiphum artemisiae on .

Korea: CB, GB, GW, JB.

Tribe Sericomyini

Genus Pseudovolucella Shiraki, 1930

Pseudovolucella Shiraki, 1930: Mem. Fac. Agric. Taihoku Imp. Univ., 1 (1): 39. TS: Pseudovolucella mimica Shiraki (OD).

84. Pseudovolucella decipiens (Hervé-Bazin, 1914) (Pls. 8A, B)

Arctophila decipiens Hervé-Bazin, 1914: 410. Acrophila jozana Matsumura, 1916: 205.

Korean record: Pseudovolucella decipiens: Choi et al. 2000: 144 (Baegunsan & Yonsei Univ. Maeji Campus, first Korean record); Han & Choi 2001: 135 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 27 (Korean check.).

Description: Body length 14.5‒17.5 mm. Male. Head pale yellow ground color; face pale yellow, almost silver white pilose with lower portion

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distinctly protrude ventrally; vertical triangle black with black pilose; frons whitish gray with black and brown pilose; lunule shiny dark brown; eyes holopic, bare; eye eontiguity shorter than vertical triangle; antennal scape and pedicel dark brown with black setulae; basoflagellomere brown, rounded; arista brown, plumose. Thorax dark brown gound color; scutum with dense yellow brown pilose; scutellum dark brown and brown hind margin with dense yellow brown pilose. Wing hyaline with dark brown maculae on middle portion; halter pale yellow. Fore leg entirely black; mid and hind femur dark brown dorsally and yellow brown ventrally; hind femur strongly thickened. Abdomen suboval; tergite 2 yellow brown anteriorly and black posteriorly with yellow brown pilose; tergite 3 and 4 entirely black with black pilose and yellow pilose hind margin. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: GB: Yeongju-si, Sunheung-myeon, Mt. Sobaeksan, 10. ix. 2005, HS Lee et al., 1♂; ditto, 30. ix. 2004, HW Byun, 1♂. GN: Ulju-gun, Samnam-myeon, Mt. Sinbulsan, 29. vi. 2003, HY Han et al., 1♂. GG: Namyangju-si, Sudong-myeon, Mt. Chungnyeongsan, 30. vi. 1999, HY Han et al., 1♂. GW: Hoengseong-gun, Anheung-myeon, Mt. Chiaksan, from Gangrim 4-ri to Cheonjibong (1,086 m) peak, 10. viii. 1999, DS Choi et al., 4♂; Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 5. vi. 2003, DS Choi & HS Lee, 2♂1♀; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 12. viii. 2003, HW Byun et al., 1♀; Jeongseon-gun, Nam-myeon, Mt. Mindungsan, form Yupyeong-ri to 1,119 m peak, 13. vi. 2006, HY Han et al., 1♂; Wonju-si, Heungeop- myeon, Maeji-ri, Yonsei Univ. Campus, 28. x. 2001, HS Lee, 1♀; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1087.1 m peak, 23. v. 1999, DS Choi & DW Kim, 1♂; ditto, 16. vi. 2006; DS Choi et al., 1♂; ditto, 26. vi. 2007, HS Lee et al., 3♂; ditto, 25. v. 2008, JS Lim & JM Jung, 1♂.

Distribution: Russian Far East (S Sakhalin, Kunashir, Moneron), Korea, Japan [PA].

Biology: Unknown.

Korea: GB, GN, GG, GW.

Genus Sericomyia Meigen, 1803

Sericomyia Meigen, 1803: Mag. Insektenk., 2: 274. TS: Musca lappona Linnaeus (des. Latreille, 1810: Consid. générales, nat. animaux, Paris: 443). Cinxia Meigen, 1800: Nouve. Class.: 35. TS: Musca lappona Linnaeus (des. Coquillett 1910: Proc. U.S. natn. Mus., 37: 524). Supressed by I.C.Z.N., 1963, Opinion 678: 339. Sericomyza Zetterstedt, 1838: Insecta Lapp.: 589; unjustified new name for Sericomyia Meigen. TS: Musca lappona Linnaeus. Bulboscrobia Gaunitz, 1937: Ent. Tidskr., 58 (1-2): 91. TS: Bulboscrobia undulans Gaunitz (MO).

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85. Sericomyia dux (Stackelberg, 1930) (Pl. 8C)

Cinxia dux Stackelberg, 1930: 119.

Korean record: Sericomyia dux: Choi et al. 2000: 144 (Baegunsan & Yonsei Univ. Maeji Campus, first Korean record); Han & Choi 2001: 137 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 27 (Korean check.).

Description: Body length 17.5‒20.8 mm. Male. Head yellow brown ground color; face almost yellow brown with lower portion distinctly protrude ventrally; ocellar triangle black with yellow pilose; frons shiny yellow brown with yellow brown pilose; lunule shiny brown; eye holoptic, bare; eye contiguity shorter than vertical triangle; antenna scape and pedicel entirely dark brown with black setulae; basoflagellomere brown, rounded apically; arista dark brown, plumose. Thorax entirely dark brown with black and brown hairs mixed; scutellum dark brown with mostly black pilose, brown pilose anterior margin. Wing hyaline with brown tinge anteriorly; halter dark brown. Legs: femora entirely dark brown; tibiae and tarsi brown. Abdomen entirely black; tergite 2‒3 each a pair yellow brown transeverse band; tergite 4 entirely yellow brown except black anterior margin. Female. Similar to male except for following non-genitalic characters: eyes dichoptic.

Specimen examined: Korea: GW: Hoengseong-gun, Anheung-myeon, Mt. Chiaksan, from Gangrim 4-ri to Cheonjibong (1,086 m) peak, 5. vi. 2005, DS Choi et al., 3♂; Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 9. viii. 2003, OY Lim & HS Lee, 1♂1♀; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 23. vi. 2005, DS Choi et al., 1♂; Inje- gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 16. vi. 2004, HW Byun et al., 2♂; Pyeongchang-gun, Jinbu-myeon, Mt. Odaesan, from Woljeongsa to Birobong (1,563 m) peak, 28. vii. 2004, DS Choi et al., 1♀; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 22. v. 1998, HY Han & DS Choi, 1♂.

Distribution: Russian Far East (S Primorye), Korea [PA].

Biology: Unknown.

Korea: GW.

86. Sericomyia sachalinica Stackelberg, 1925

Sericomyia sachalinica Stackelberg, 1925: 92. Sericomyia japonica Shiraki, 1930: 45.

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Sericomyia nigriceps Shiraki, 1930: 46.

Korean record: Sericomyia sachalinica: ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 133 (Korean cat.); Han & Choi 2001: 137 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 27 (Korean check.). Sericomyia japonica: Doi 1938a: 9 (Korean check., presumably first Korean record); ZSK 1968: 180 (Korean check.); Kim, CW 1971: 866 (redescrip.); Kim, JI 1975: 40 (Korean check.); Kim, JI 1980: 385 (Korean check.).

Distribution: Russian Far East (Sakhalin, Kunashir), Korea, Japan (Hokkaido) [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim (1971).

Tribe Volucellini

Genus Graptomyza Wiedemann, 1820

Graptomyza Wiedemann, 1820: 16 (on plate). TS: Graptomyza longirostris Wiedemann, 1820 (by original designation). Graptomyza Henning, 1832: 382 (preoccupied by Graptomyza Wiedemann). TS: Graptomyza longicornis Henning, 1832 (by original designation). Baryterocera Walker, 1857: 123. TS: Baryterocera inclusa Walker, 1857 (by monotypy). Ptilostylomyia Bigot, 1883: cxiv. [des. Knutson et al., 1975: 332]. TS: Ptilostylomyia brevirostris Wiedemann, 1820 (by original designation). Protograptomyza Hull, 1949: 351 (as subgenus of Graptomyza Wiedemann). TS: Graptomyza doddi Ferguson, 1926 (by original designation).

87. Graptomyza ihai Shiraki, 1968 (Pl. 8D)

Graptomyza ihai Shiraki, 1968a: 225.

Korean record: Graptomyza ihai: Choi & Han: 2004: 1 (new Korean record); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 27 (Korean check.).

Description (modified from Choi and Han, 2004): Body length 4.5‒5.2 mm.

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Male. Head yellow brown ground color with dark occiput, vertex, anterior portion of basoflagellomere; face entirely yellow brown, pilose with lower portion distinctly protrude anteriorly with blunt apex; lunule shiny yellow brown; eyes densely short pilose, broadly dichoptic; basoflagellomere kidney bean shaped in lateral view. Thorax: scutum slightly longer than wide, largely shiny black except narrow lateral and posterior areas; scutellum semicircular with broad semicircular depression, shiny black pilose; scutellum with 4 marginal seta.Wing almost entirely hyaline with tiny dark spots on cell R1 at middle position between node and apex of vein R1, slightly posterior to node, on r-m; halter pale yellow. Legs largely yellow except apex of hind femur, basal and apical 1/4 of hind tibia, hind sarsus brown. Abdomen oval, yellow brown ground color, short dense pilose; tergite 1 posteriorly with pair of brown sublateral spots; dark brown posterior transverse band on tergite 2 with 3 triangular anterior peaks, mesal peak longest, reaching 3/4 of tergite length and lareral peaks shorter than half as long as middle; dark brown posterior transverse band on tergite 3 with 3 peaks, mesal peak more or less widely oblong, longest, reaching 3/4‒4/5 of tergite length, and lateral peaks triangular, about half as long as middle peak; tergite 4 posteriorly with 3 dark brown spots, mesal spot more or less widely oblong, sublateral spots small, more or less round. Female. Similar to male except for following non-genitalic characters: Protrusion of lower face laterally with clear dark brown stripe while absent or very weak in male; wing with more extensive dark brown pattern than in male; three dark spots about twice as large as spots in male; abdominal color pattern similar to male but lateral peaks on tergite 2 and 3 about as long as mesal peaks.

Specimen examined: Korea: GN: Mt. Jirisan, 10. viii. 1989, YJ Kwon, 1♀. GW: Hoengseong-gun, Dunnae- myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 27. vii. 2002, DS Choi et al., 1♀; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, N. Valley, 26. v. 2006, HS Lee & DJ Cha, 1♀; ditto, 27. v. 2007, HS Lee et al., 1♂; ditto, 22. vii. 2005, HS Lee, 1♀; Hongcheon-gun, Nae-myeon, Mt. Gachilbong, from Sambongyaksu to 1,240 m peak, 22. v. 1999, HY Han & KE Ro, 1♂; ditto, 14. v. 1999, HY Han & KE Ro, 1♀; Jeongseon- gun, Nam-myeon, Mt. Mindungsan, form Yupyeong-ri to 1,119 m peak, 6. vi. 2006, HY Han et al., 1♀. JJ: Bukjeju-gun, Jocheon-eup, Goepyeongi oreum, 6. viii. 2003, DS Choi & HW Byun, 9♂29♀.

Distribution: Korea, Japan [PA].

Biology: Unknown.

Korea: GN, GW, JJ.

Genus Volucella Geoffroy, 1762

Volucella Geoffroy, 1762: Hist. abrégée ins. Paris, 2: 540. TS: Musca pellucens Linnaeus (des. Curtis, 1833: Brit. ent., 10: pl. 452). Validated by I.C.Z.N. 1957, Opinion 441: 85. Pterocera Meigen, 1803: Mag. Insktenk., 2: 275. TS: Musca pellucens Linnaeus (des. Coquillett, 1910: Proc.

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U.S. natn. Mus., 37: 597). Cenogaster Duméril, 1806: Zool. analytique: 282. TS: Musca pellucens Linnaeus (as Syrphus pellucens F.; by subsequent monotypy: Froriep in Duméril, 1806: German transl. of Zool. analytique: 283). Temnocera Le Peletier et Serville in Latreille et al., 1828: Encycl. Méthod. (Ins.), 10: 786. TS: Temnocera violacea Le Peletier et Serville (OD).

Key to species of Volucella from South Korea

1. Scutellum without strong marginal setae; head entirely black; abdomen with long pili ··· V. jeddona Bigot − Scutellum with strong marginal setae; head not entirely black; abdomen with short pili ······················ 2 2. Antenna about broad; eye dichoptic in male ············································ V. inanoides Hervé-Bazin − Antenna at least 2/3 times as long as shortest eye diameter; basoflagellomere narrow; eye holoptic in male· 3 3. 3rd and 4th abdominal tergites entirely black ········································································ 4 − 3rd and 4th abdominal tergites not entirely black ··································································· 8 4. Scutum shiny reddish brown in ground color; legs almost entirely reddish brown ····························· 5 − Scutum shiny black in ground color; legs almost entirely black ·················································· 6 5. Scutellum with reddish brown marginal setae; face without any black stripe; wing crossvein r-m without dark brown marking ············································································· V. suzukii Matsumura − Scutellum with black marginal setae; face with black stripe laterally; wing crossvein r-m with dark brown marking ································································································ V. linearis Walker 6. 2nd abdominal tergite length 2.4 times as long as width; eye contiguity 0.6 times as long as frontal triangle in male; female eye densely pilose ························································· V. thompsoni Choi et al. − 2nd abdominal tergite length at least 2.7 times longer than width; eye contiguity about as long as frontal triangle; female eye bare ································································································ 7 7. 2nd abdominal tergite black with undulate white band at anterior margin; vertex dark brown to black in female; female scutum entirely shiny black ·················································V. nigricans Coquillett − 2nd abdominal tergite mostly white with black longitudinal band at middle portion; vertex yellow brown in female; female scutum shiny black with brown lateral margins ······ pellucens tabanoides Motschulsky 8. Abdomen yellow brown with six black spots ·········································· V. nigropicta Portschinsky − Abdomen yellow brown without six black spots ···································································· 9 9. Wing crossveins bordered with dark brown tinge; female upper post-ocular orbit clearly seen in lateral view; vertex conspicuously swollen in female ···················································V. coreana Shiraki − Wing crossveins clear; female upper post-ocular orbit barely seen in lateral view; vertex slightly swollen in female ·····························································································V. bivitta Huo et al.

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88. Volucella bivitta Huo, Ren and Zhemin, 2007 (Pls. 8E, F)

Volucella bivitta Huo, Ren and Zhemin, 2007: 484

Korean record: Volucella bivitta: Suk et al. 2015: 260 (new Korean record). Volucella sp. near zonaria: Choi et al. 2000: 144 (Baegunsan & Yonsei Univ. Maeji Campus). Volucella sp.-2: Han & Choi 2001: 132 (Korean check.).

Description: Body length 15‒18 mm. Male. Head almost entirely yellow brown with dark brown post-ocular orbit; face almost yellow pilose with lower portion distinctly protrude anteriorly with sharp apex, protrusion about 1/3 of eye height; gena with yellow pilose; lunule shiny yellow brown; post-ocular orbit barely seen in lateral view; frons almost yellow pilose; eye holoptic, dense pilose; eye contiguity 0.12‒0.17 times as long as frontal triangle; antenna almost entirely yellow brown with yellow and black pilose mixed; basoflagellomere transverse elliptic, 1.8‒2.3 times as long as broad at middle. Thorax entirely dark brown except brown laterally; scutellum dark brown, almost short dense yellow pilose. Wing hyaline with brown tinge. Legs almost entirely dark brown densely covered with black setulae. Abdomen obovate; tergite 2‒4 each brown with dark brown marginal band posteriorly. Female. Similar to male except for following non-genitalic characters: eyes dichoptic, sparsely pilose.

Specimen examined: Korea: CB: Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 30. vii. 2003, DS Choi & HW Byun, 1♀. GB: Cheongsong-gun, Budong-myeon, 4. vi. 1989, CM Kim, 1♀. GN: Geoje-si, Dongbu- myeon, Hakdong-ri, 6. viii. 2004, TW Kim, 1♂. GG: Pocheon-gun, Idong-myeon, Mt. Myeongseongsan, 6. vi. 1981, KS Kang, 1♂. GW: Wonju-si, Heungeop-myeon, Maeji-ri, 7. v. 2005, DS Choi, 1♂. GW: Donghae- si, Manu-dong, 19. vii. 2010, SW Suk et al., 1♂; Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 15. viii. 2001, DS Choi et al., 1♂; Hongcheon-gun, Nae-myeon, Mt. Gachilbong, 1♂1♀; Inje-gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 1♂; Jeongseon-gun, Nam-myeon, Mt. Mindungsan, form Yupyeong-ri to 1,119 m peak, 2. viii. 2001, HY Han & KE Ro, 1♂; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 31. viii. 2007, SW Suk, 1♀; Wonju-si, Panbu- myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 25. viii. 2005, SW Suk et al., 1♀. JB: Jangsu- gun, Jangsu-eup, Deoksan-ri, Mt. Jangansan, 25. viii. 2008, HS Lee & JM Jung, 2♂.

Distribution: Russian Far East, Korea, China [PA].

Biology: In Korea, This species were often found on the flowers of Zanthoxylum schinfolim Siebold & Zucc. together with V. linearis in summer (Suk et al., 2015).

Korea: CB, GB, GN, GG, GW, JB.

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89. Volucella coreana Shiraki, 1930 (Pl. 8G)

Volucella coreana Shiraki, 1930: 225.

Korean record: Volucella coreana: Shiraki 1930: 138 (orig. desc., first Korean record); Sack 1931: 243 (redescrip.), Doi 1938a: 10 (Korean check.); ZSK 1968: 179 (Korean check.); Kim, CW 1971: 862 (redescrip.); Kim, JI 1975: 41 (Korean check.); Kim, JI 1980: 385 (Korean check.); Peck 1988: 129 (Palaearctic cat.); ESK & KSAE 1994: 290 (Korean check.); Han et al. 1998b: 130 (Korean cat.); Choi et al. 2000: 143 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 122 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 27 (Korean check.).

Description: Body length 21.5‒24.2 mm. Male. No specimens available. Female. Head almost entirely yellow brown with yellow pilose; face with lower portion distinctly protrude anteriorly with sharp apex, protrusion about 1/3 of eye height; lunule shiny yellow brown; vertex conspicuously swollen; post-ocular orbit clearly seen as narrow area; eye dichoptic, bare; antenna with yellow and black pilose mixed; basoflagellomere transverse elliptic, 2‒2.1 times as long as broad at middle. Thorax with 5‒6 marginal scutellar seta; scutum with whitish pruinose longitudinal stripe on submedial portion, almost entirely yellow and black short dense pilose; scutellum dark brown, almost short dense yellow pilose with black pilose marginally. Wing hyaline with brown tinge. Legs almost entirely dark brown densely covered with black setulae. Abdomen obovate; tergite 2‒4 each brown with dark brown marginal band posteriorly.

Specimen examined: Korea: GB: Yeongyang-gun, Seokbo-myeon, Yowon-ri, 8. viii. 2008, HS Lee, 1♀. GW: Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 10. viii. 1998, HW Byun, 1♀; ditto, 21. viii. 2005, HW Byun, 1♀; ditto, 11. ix. 2005, HW Byun, 1♀; ditto, 30. viii. 2006, DJ Cha, 1♀; ditto, 31. viii. 2006, HW Byun, 1♀.

Distribution: Russian Far East to Korea [PA].

Biology: Unknown.

Korea: GB, GW.

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90. Volucella inanoides Hervé-Bazin, 1923 (Pls. 8H, I)

Volucella inanoides Hervé-Bazin, 1923: 256.

Korean record: This species has been repeatedly misidentified by Korean authors as V. nigropicta (= V. sexmaculata). This is probably because incorrect illustration and Description were given in Kim, CW (1971) (see Remarks of Han et al., 1998b). Volucella inanoides: Suk et al. 2015: 261 (new Korean record). Volucella sp. near inanoides: Choi et al. 2000: 144 (Baegunsan & Yonsei Univ. Maeji Campus). Volucella sp.-1: Han & Choi 2001: 130 (Korean check.). Volucella nigropicta [misidentification]: ESK & KSAE 1994: 291 (Korean check.). Volucella sexmaculata [misidentification]: Kim, CW 1971: 865 (redescrip.); Kim, JI 1975: 41 (Korean check.); Kim, CW 1980: 294 (distr. Map); Kim, JI 1980: 385 (Korean check.).

Description: Body length 13.5‒19.2 mm. Male. Head yellow brown ground color with dark brown facial stripe laterally; face almost yellow pilose with lower portion distinctly protrude anteriorly with sharp apex, protrusion about 1/3 of eye height; gena with yellow pilose; lunule shiny yellow brown; frons almost yellow pilose; eyes narrowly separated, dense pilose; antenna almost entirely yellow brown with yellow setulae; basoflagellomere kidney shape, widend; arista brown, plumose. Thorax with 5‒6 marginal scutellar seta; scutum shiny black ground color except laterally; scutellum entirely yellow brown with black setulae. Wing hyaline with brown tinge anteriorly. Legs almost entirely yellow brown except coxae and trochanters shiny black. Abdomen obovate; tergite 2 black with a pair of yellow brown triangular band; tergite 3 yellow brown anteriorly and black posteriorly; tergite 4 entirely yellow brown with black longitudinal band medially. Female. Similar to male except for following non-genitalic characters: eyes distinct dichoptic.

Specimen examined: Korea: CB: Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 2. vii. 2003, DS Choi et al., 1♂. GB: Andong-si, Giran-myeon, Goran-ri, Mt. Gyemeoyngsan, 1. vi. 2012, SW Suk et al, 1♂; Choengsong-gun, Hyeonseo-myeon, Galcheon-ri, 29. viii. 2011, SW Suk et al., 2♀. GN: Yangsan-si, Ungsang- eup, Daeunsan jayeonhyuyangnim, JC Jeong, 7. vi. 2008, 1♂; Namhae-gun, Samdong-myeon, Bonghwa-ri, Pyeonbaek Recreational Forest, 20. vi. 2000, JS Jeon, 1♀. GG: Namyangju-si, Hwado-eup, Mt. Cheonmasan, 13. ix. 1994, UH Jeong, 1♀. GW: Chuncheon-si, Sindong-myeon, Palmi-ri, 4. vi. 2006, DS Choi et al., 1♂; Samcheok-si, Dogye-eup, from Sinbangtae to Mt. Dohwasan (925 m), 8. vi. 2003, DS Choi et al, 1♂; Wonju- si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 4. vi. 2003, DS Choi, 1♂; ditto, 1. vi. 2009, HW Byun, 1♀; ditto, 21. vi. 2006, DS Choi et al., 1♀; ditto, black + mercury vapor light trap, 21. vi. 2009, HY Han et al., 1♂. JB: Buan-gun, Byeonsan-myeon, 26. vi. 1991, SM Kim, 1♀; Namwon-si, Sannae-myeon, Mr. Jirisan, Baemsagol (646 m), Malaise Trap (70 % EtOH), 26. vi.-11. vii. 1991, DS Ku, 1♂. JN: Jangseong-gun, Bugi- myeon, Baegyangsan, 26. vi. 1991, JI Kim, 1♀.

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Distribution: Korea, China [PA].

Biology: This species were often found on sap runs of oak trees during summer months (Suk et al., 2015).

Korea: CB, GB, GN, GG, GW, JB, JN.

Remarks: This is only species of the Korean Volucella with separated compound eyes in males (see Suk et al., 2015).

91. Volucella jeddona Bigot, 1875 (Pls. 8J, K)

Volucella jeddona Bigot, 1875: 472. Volucella brevipila Portschinsky, 1886: 7.

Korean record: Volucella jeddona: Doi 1938a: 10 (Korean check., presumably first Korean record); Kim, CW 1971: 864 (redescrip.); Kim, JI 1975: 41 (Korean check.); Kim, JI 1980: 385 (Korean check.); ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998b: 130 (Korean cat.); Han & Choi 2001: 122 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 28 (Korean check.). Volucella ioddona [misspelling]: ZSK 1968: 179 (Korean check.).

Description: Body length 17‒18.5 mm. Male. Head almost entirely black ground color; face entirely shiny black with dense short pale yellow pilose, lower portion distinctly protrude anteriorly with sharp apex, protrusion about 1/3 of eye height; lunule shiny black; frons shiny with almost yellow pilose; eyes holoptic, dense pilose; eye contiguity 1.32‒1.64 times as long as frontal triangle; antennal scape and pedicel almost entirely black with black setulae; basoflagellomere dark brown, transverse elliptic, 1.7‒2.2 times as long as broad at middle. Thorax dark brown ground color with yellow brown pilose; scutum dark brown with yellow brown lateral and posterior margins; scutellum shiny yellow brown with dense short yellow brown pilose. Wing hyaline with dark brown maculae medially; halter pale yellow. Legs almost entirely black. Abdomen obovate; tergite 2 black longitudinal black band medially; tergite 3 entirely black with dense short black and yellow pilose mixed. Female. Similar to male except for following non-genitalic characters: eyes dichoptic, moderate pilose.

Specimen examined: Korea: CB: Danyang-gun, Mt.Sobaeksan, Yeonhwa 1 bong peak, 23. vii. 1974, JI Kim, 1♂. GB: Yeongju-si, Punggi-eup, Mt. Sobaeksan, 2. viii. 1994, SM Ryu, 1♂. GW: Donghae-si, Bukpyeong, Samhwasa, 27. vi. 1984, JI Kim, 1♂; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 29. vii. 1997, HY Han et al., 1♂; ditto, 6. viii. 1997, HY Han et al., 1♂1♀; Jeongseon-gun, Nam-myeon, Mt. Mindungsan, form Yupyeong-ri to 1,119 m peak, 24. vi. 2005, HY Han et al., 1♀; Pyeongchang-gun, Jinbu-myeon, Mt. Odaesan, from Bukdaesa to Sangwangbong (1,491 m) peak, 20.

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vii. 2004, DS Choi & HW Byun et al., 2♂1♀; Yanggu-gun, Haean-myeon, Mt. Daeamsan, 28. vii. 1997, JI Kim, 1♂.

Distribution: Russian Far East (Primorye Territory, Sakhalin, Kunashir, Shikotan), Mongolia, Korea, Japan (Hokkaido, Honshu) [PA].

Biology: Unknown.

Korea: CB, GB, GW.

92. Volucella linearis Walker, 1852 (Pls. 8L, 9A)

Volucella linearis Walker, 1852: 251. Volucella nitobei Matsumura, 1916: 210.

Korean record: Volucella linearis: ESK & KSAE 1994: 291 (Korean check.); Han & Choi 2001: 123 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 28 (Korean check.). Volucella nitobei: ZSK 1968: 179 (Korean check.); Kim, CW 1971: 864 (redescrip.); Kim, JI 1975: 41 (Korean check.); Kim, JI 1980: 385 (Korean check.); Han et al. 1998a: 10 (Yonsei Univ. Maeji Campus); Han et al. 1998b: 131 (Korean cat.); Choi et al. 2000: 143 (Baegunsan & Yonsei Univ. Maeji Campus).

Description: Body length 16.5‒20.5 mm. Male. Head reddish brown ground color with dark brown facial stripe laterally; face almost yellow pilose with lower portion distinctly protrude anteriorly with sharp apex, protrusion about 1/3 of eye height; lunule shiny brown; frons shiny with almost yellow pilose; eyes holoptic, dense pilose; eye contiguity 1.56‒1.72 times as long as frontal triangle; antenna almost entirely yellow brown with brown setulae; basoflagellomere transverse elliptic, 1.5‒1.9 times as long as broad at middle. Thorax dark brown ground color with two black longitudinal band medially; scutum and scutellum shiny dark brown with dense yellow brown pilose. Wing hyaline with broad yellow brown tinge along anterior 1/3; discal macular distinctly broad, dark brown on middle portion; halter yellow brown. Legs almost entirely reddish brown. Abdomen obovate, almost black; tergite 2 black with brown band on anterior margin; tergite 3 and 4 shiny black with dense short black pilose. Female. Similar to male except for following non-genitalic characters: eyes dichoptic; anterior band of tergite 2 larger than male.

Specimen examined: Korea: CB: Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 8. viii. 2003, OY Lim & HS Lee, 2♀. CN: Boryeong-si, Misan-myeon, Mt. Wolmyeongsan, 23. viii. 2000, DS Choi & DW Kim, 1♀. GB: Andong-si, Gilan-myeon, Gyemyeongsan Recreational Forest, 7. vii. 2010, DS Choi, 1♂1♀. GN: Geochang-gun, Buksang-myeon, Mt. Deogyusan, Wolseong Valley (700 m), 2. viii. 2007, DS Choi, 1♂.

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GG: Gwangju-si, Docheok-myeon, Mt. Taehwasan, from Eungoksa to 612.8 m peak, 25. viii. 2006, HY Han et al., 2♀; Paju-si, Papyeong-myeon, Masan-ri, 1. viii. 2006, DS Choi et al, , 1♂1♀. GW: Hoengseong- gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 1. viii. 2006, HW Byun et al., 1♂; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, N. Valley, 14. viii. 1998, HY Han & KE Ro, 1♀; Inje-gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 21. viii. 1998, HY Han et al., 1♂1♀; Pyeongchang-gun, Jinbu-myeon, Mt. Odaesan, from Woljeongsa to Birobong (1,563 m) peak, 28. vii. 2004, DS Choi et al., 1♀; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 20. vii. 2006, SMR Hwang, 1♂1♀; ditto, 19. vii. 2009, SW Suk, 1♂. JB: Jeonju-si, Geumsan-myeon, Mt. Moaksan (793.5 m), Geumsan- ri, 10. ix. 2003, HS Lee, 3♀; Namwon-si, Jucheon-myeon, Yonggung-ri, Jangan reservoir, 29. vii. 2013, DS Choi, 1♂. JN: Gwangju, Buk-gu, Mt. Mudeungsan, Neutjaesamgeori, 4. viii. 2013, DS Choi, 4♀.

Distribution: Korea, Japan [PA].

Biology: This species is associated with Vespula flaviceps lewisii (Cameron) in immature stages (Ôhara, 1985a, b).

Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

93. Volucella nigricans Coquillett, 1898 (Pls. 9B, C)

Volucella nigricans Coquillett, 1898: 324.

Korean record: Volucella nigricans: Okamoto 1924: 155 (Jeju-do, presumably first Korean record); Shiraki 1930: 216 (redescrip.); Doi 1938a: 10 (Korean check.,); ZSK 1968: 179 (Korean check.); Shiraki 1968a: 215 (redescrip.); Kim, CW 1971: 863 (redescrip.); Kim, JI 1975: 41 (Korean check.); Knutson et al. 1975: 336 (Orient. cat.); Kim, CW 1980: 293 (distr. Map); Kim, JI 1980: 385 (Korean check.); Peck 1988: 130 (Palaearctic cat.); ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998a: 10 (Yonsei Univ. Maeji Campus); Han et al. 1998b: 130 (Korean cat.); Choi et al. 2000: 143 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 125 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 28 (Korean check.).

Description: Body length 15.5‒18.6 mm. Male. Head yellow brown ground color with black vertical triangle, post-ocular orbit, posteroventral portion of face, gena; face yellow brown and black, almost yellow pilose with lower portion distinctly protrude anterior with sharp apex, protrusion about 1/3 of eye height; gena shiny black with yellow and black pilose; lunule shiny yellow brown; frons almost black or yellow pilose or mixed; eyes holoptic, dense pilose; eye contiguity 1‒1.2 times as long as frontal triangle; antenna almost entirely yellow brown with almost black pilose; basoflagellomere transverse elliptic, 1.5‒1.7 times as long as broad at middle. Thorax with 6‒7

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marginal scutellar seta; scutum shiny black except dark brown on postalar callus, short dense black pilose; postpronotum pale yellow, almost black pilose; scutellum shiny black, almost short dense black pilose. Wing hyaline with broad yellow brown tinge along anterior 1/3; discal macular distinctly broad, dark brown on middle portion; halter brown to dark brown except white on knob. Legs almost entirely black, black pilose. Abdomen suboval, largerly shiny black except white semicircular bands or undulate band on tergite 2 anteriorly, almost black pilose. Female. Similar to male except for following non-genitalic characters: eyes dichoptic, bare; abdomen with more extensive undulate white band on tergite 2 than in male.

Specimen examined: Korea: CB: Cheongwon-gun, Namseong-myeon, Mt. Ingyeongsan, 15. viii. 1997, HY Han et al., 1♂3♀; Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 18. vii. 2007, JS Lim et al., 1♀. CN: Daejeon, Seo-gu, Wolpyeong-dong, Wolpyeong park 2 site (Malase trap), 12. v~24. vi. 2008, JW Lee, 1♀. GB: Gyeongju-si, Naenam-myeon, Gwaneumsa, 25. ix. 2010, DS Choi, 1♀; Uljin-gun, Onjeong-myeon, Jogeum-ri, 18. viii. 2008, DS Choi, 2♂; Yeongyang-gun, Ibam-myeon, Samsan-ri, 15. viii. 2008, DS Choi, 1♀. GN: Busan, Gijang-gun, Cheolma-myeon, Gomnaejae, 7. vi. 2012, JG Shin, 1♂. GG: Incheon, Ongjin- gun, Jawol-myeon, Seungbong-ri, 18. Vi. 2009, JK Choi & DH Lee, 1♂. GW: Jeongseon-gun, Nam- myeon, Mt. Mindungsan, form Yupyeong-ri to 1,119 m peak, 16. vii. 2005, HY Han et al., 12♂1♀; Wonju- si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 7. vi. 2006, JS Lim, 2♂1♀; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 12. vi. 2006, HS Lee & SMR Hwang, 1♂. JN: Gwangju, Buk-gu, Mt. Mudeungsan, Neutjaesamgeori, 4. viii. 2013, DS Choi, 1♂; Gwangyang-si, Okryong-myeon, Donggok-ri, Mt. Baekunsan, 14. vi. 2014, DS Choi 2♂.

Distribution: Korea, Japan, Taiwan [PA, OR].

Biology: This species is associated with Vespula flaviceps lewisii (Cameron) in immature stages (Okuno, 1970; Ôhara, 1985a, b).

Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

94. Volucella nigropicta Portschinsky, 1883 (Pls. 9D, E)

Volucella nigropicta Portschinsky, 1883: 127. Volucella sexmaculata Matsumura, 1916: 212.

Korean record: There are a large number of Korean recoreds for this species, but they are mostly misidentification of V. inanoides. Volucella nigropicta: Han et al. 1998a: 10 (Yonsei Univ. Maeji Campus); Han et al. 1998b: 131 (Korean cat.); Choi et al. 2000: 143 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 127 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 28 (Korean check.).

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Description: Body length 18.5‒19.4 mm. Male. Head yellow brown ground color with brown to dark brown vertical triangle; face yellow brown, almost yellow pilose with lower portion distinctly protrude anterior with sharp apex, protrusion about 1/3 of eye height; lunule shiny black; frons almost pale yellow pilose; eyes holoptic, dense pilose; eye contiguity longer than vertical triangle; antenna almost entirely brown with almost brown pilose; basoflagellomere transverse elliptic, 1.8‒2.4 times as long as broad at middle. Thorax with 6‒7 marginal scutellar seta; scutum dark brown except yellow brown on postalar callus and supra-alar area, short dense brown pilose; postpronotum pale yellow; scutellum shiny dark brown, almost dense black pilose. Wing hyaline with broad yellow brown tinge anterobasally; halter pale yellow. Legs almost entirely brown ground color. Abdomen suboval, yellow brown with six dark brown spots. Female. Similar to male except for following non-genitalic characters: eyes dichoptic; upper post-ocular orbit clearly seen as narrow area in laternal view.

Specimen examined: Korea: GG: Pocheon-gun, Sohol-eup, Jikdong-ri, Gwangneung, 4. ix. 1956, WJ Kim, 1♀; Seoul, Gwanak-gu, Mt. Gwanaksan, 20. viii. 1992, JY Kim, 1♀. GW: Pyeongchang-gun, Sogeumgang, 17. ix. 1971, JI Kim, 1♀; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 18. viii. 1998, HW Byun & SK Kim, 1♀; ditto, 19. vii. 1997, HW Byun, 1♂. JN: Jangseong-gun, Bugi-myeon, Mt. Baegyangsan, 4. viii. 1974, CW Kim, 1♂1♀.

Distribution: Russian Far East (Primorye Territory), Korea, Japan (Honshu) [PA].

Biology: Unknown.

Korea: GG, GW, JN.

95. Volucella pellucens tabanoides Motschulsky, 1859 (Pls. 9F, G, H)

Volucella tabanoides Motschulsky, 1859: 504. Volucella japonica Bigot, 1875: 473. Volucella pellucens var. japonica Matsumura, 1916: 238. Volucella matsumurai Han et Choi, 2001: 125.

Korean record: Volucella pellucens tabanoides: Peck 1988: 131 (Palaearctic cat.); ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998a: 11 (Yonsei Univ. Maeji Campus); Han et al. 1998b: 132 (Korean cat.); Choi et al. 2000: 144 (Baegunsan & Yonsei Univ. Maeji Campus); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 28 (Korean check.). Volucella tabanoides: Shiraki 1930: 216 (redescrip.); Doi 1938a: 11 (Korean check.); ZSK 1968: 179 (Korean check.); Shiraki 1968a: 213 (redescrip.); Kim, CW 1971: 863 (redescrip.); Kim, JI 1975: 41 (Korean check.); Kim, JI 1980: 385 (Korean check.); Han & Choi 2001: 127

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(Korean check.). Volucella matsumurai [misidentification]: Han & Choi 2001: 125 (Korean check.).

Description: Body length 13.7‒19 mm. Male. Head yellow brown ground color with brown to dark brown vertical triangle, post-ocular orbit, posteroventral portion of face; face yellow brown and black, almost yellow pilose with lower portion distinctly protrude anterior with sharp apex, protrusion about 1/3 of eye height; gena shiny black with yellow and black pilose; lunule shiny yellow brown; frons almost black pilose; eyes holoptic, dense pilose; eye contiguity 1.14‒1.46 times as long as frontal triangle; antenna almost entirely yellow brown with almost black pilose; basoflagellomere transverse elliptic, 1.5‒1.7 times as long as broad at middle. Thorax with 6‒7 marginal scutellar seta; scutum shiny black except dark brown on postalar callus and supra-alar area, short dense black or yellow pilose mixed; postpronotum pale yellow, almost yellow and black pilose mixed; scutellum shiny dark brown, almost dense black pilose. Wing hyaline with broad yellow brown tinge anterobasally; discal macular distinctly broad, dark brown on middle portion; halter brown to dark brown except white on knob. Legs almost entirely black except brown basal 1/3 for and midtibia, black pilose. Abdomen suboval, largerly shiny black with almost black pilose except tergite 2; tergite 2 white with brown to dark brown longitudinal tinge or band medially, almost yellow pilose except posterior and lateral margin black. Female. Similar to male except for following non-genitalic characters: frons almost black or yellow pilose, or mixed; eyes dichoptic, bare; scutum with more extensive brown portion than in male (lateral margin and triangular spot posterior margin); scutellum shiny brown, almost short dense yellow pilose or black mixed. Abdomen with almost entirely white or more narrow brown tinge on tergite 2 than in male.

Specimen examined: Korea: CB: Cheongwon-gun, Namseong-myeon, Mt. Ingyeongsan, 15. viii. 1997, HY Han et al., 4♀; Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 30. vii. 2003, DS Choi & HW Byun, 1♂; ditto, 29. vii. 2004, HW Byun & JS Lim, 1♂. CN: Boryeong-si, Misan-myeon, Mt. Wolmyeongsan, 23. viii. 2000, DS Choi & DW Kim, 1♀. GB: Cheongsong-gun, Budong-myeon, Mt. Juwangsan (721 m), 10. ix. 2006, DS Choi, 1♀. GG: Gwangju-si, Chowol-myeon, Mt. Mugapsan, 9. viii. 2001, DS Choi & HS Lee, 1♂. GW: Hoengseong-gun, Anheung-myeon, Mt. Chiaksan, from Gangrim 4-ri to Cheonjibong (1,086 m) peak, 14. vii. 2003, DS Choi & HW Byun, 2♂; Hongcheon-gun, Nae-myeon, Mt. Gachilbong, from Sambongyaksu to 1,240 m peak, 24. viii. 1997, HY Han et al., 1♂4♀; Hwacheon-gun, Sanae-myeon, Mt. Gwangdeoksan, 20. vii. 1997, 1♀; Jeongseon-gun, Gohan-eup, Mt. Hambaeksan, 9. viii. 2000, HY Han et al., 1♂2♀; Jeongseon- gun, Nam-myeon, Mt. Mindungsan, form Yupyeong-ri to 1,119 m peak, 16. vii. 2005, HY Han et al., 1♂2♀; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 5. x. 2007, YB Lee, 1♀.

Distribution: Russia (Siberia, Far East), Mongolia, China, Korea, Japan [PA].

Biology: This species is associated with Vespa and Vespula species (Hymenoptera) during immature

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stages. The larvae of three Volucella species including this species were collected from soil under the nest of Vespula flaviceps lewisii (Cameron) and studied morphologically in Japan (Ôhara, 1985a).

Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

96. Volucella suzukii Matsumura, 1916 (Pl. 9I)

Volucella suzukii Matsumura, 1916: 236.

Korean record: Volucella suzukii: Kim, JI & Kim, CW 1972: 12 (new Korean record); Kim, JI 1975: 41 (Korean check.); Kim, JI 1980: 385 (Korean check.); ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998b: 132 (Korean cat.); Han & Choi 2001: 129 (Korean check.); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 28 (Korean check.).

Description: Body length 12.2‒15.7 mm. Male. Head almost entirely yellow brown; face almost yellow pilose with lower portion distinctly protrude anteriorly with blunt apex, protrusion about 1/5 of eye height; gena yellow pilose; lunule shiny yellow brown; frons almost yellow pilose; eyes sparsely dichoptic, dense pilose; eye contiguity 1.88‒2.14 times as long as frontal triangle; antenna almost entirely yellow brown with brown; basoflagellomere transverse elliptic, 1.5‒1.6 times as long as broad at middle; arista plumose. Thorax dark brown ground color with dense reddish brown pilose. Wing hyaline with black tinge only small area of cell SC medially; halter yellow brown. Legs almost entirely brown except hind femur dark brown. Abdomen entirely black ground color with dense short pilose. Female. Similar to male except for following non-genitalic characters: eyes distinctly dichoptic.

Specimen examined: Korea: GB: Gyeongju-si, Sannae-myeon, Mt. Munboksan, Simwonsa, 18. viii. 2001, TH Ahn, 1♀. GN: Goseong-gun, Sangni-myeon, Mt. Hyangnobong, 21. vii. 2001, JS Jeon, 1♀. GG: Incheon, Deokjeok-myeon, Deokjeokdo, 31. vii. 1998, MK Paek & NH Ahn, 2♀; Inchon, Jung-gu, Unbuk-dong, Is. Yeongjongdo, 22. vii. 2007, DS Choi, 1♂. GW: Wonju-si, Heungeop-myeon, Maeji-ri, Maesannongwon, 14. vii. 2006, HS Lee et al., 1♂. JN: Jangseong-gun, Bugi-myeon, Mt. Baegyangsan, Baegyangsa, 14. viii. 1974, CW Kim, 1♂; ditto, JI Kim, 1♂; SH Nam, 1♂.

Distribution: Korea, Japan (Honshu) [PA].

Biology: This species is associated with Vespa simillima xanthooptera in immature stages (Ôhara, 1985a, b).

Korea: GB, GN, GG, GW, JN.

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97. Volucella thompsoni Choi, Ôhara and Han, 2006 (Pls. 9J, K)

Volucella thompsoni Choi, Ôhara and Han, 2006: 4.

Korean record: Volucella thompsoni: Choi et al. 2006: 4 (orig. desc., first Korean record); Paek et al. 2010: 231 (Korean check.); Han et al. 2014: 29 (Korean check.).

Description (modified from Choi et al., 2006): Body length 11‒16 mm. Male. Head yellow brown ground color with dark brown to black vertical triangle, post-ocular orbit, posteroventral portion of face, and gena; face yellow brown pilose with varying size of black macula posteroventrally; ventral portion of face distinctly protrude with more or less pointed apex; gena shiny black with short yellow setulae; lunule shiny yellow brown; frons yellow brown with black and yellow setulae mixed; eyes holoptic, dense pilose; antenna almost entirely yellow brown with black setulae; basoflagellomere 2.1‒2.6 times longer than wide, slightly constricted in middle. Thorax with black setae and dense black to yellow brown setulae; scutum shiny black with brown to dark brown lateral margins; scutellum shiny brown densely with short black setulae. Wing hyaline with broad discal and anterior apical dark brown maculae; halter yellow brown. Legs almost entirely black except basal 1/3‒1/2 of fore and midtibiae dark brown; densely covered with black setulae. Abdomen 1.3‒1.7 times longer than wide; tergite 2 ivory white medially with narrow yellow brown longitudinal stripe, densely with pale setulae; tergite 3 and 4 shiny black with black setulae. Female. Similar to male except for following non-genitalic characters: frons almost entirely yellow brown; eyes dichoptic; basoflagellomere transversely elliptic (1.9‒2.3 times longer than wide) without medial constriction; scutum with more extensive brownish portion than in male.

Specimen examined: Korea: CB: Jecheon-si, Songgye-ri, Jeolgol, 29. vi. 1997, HY Han et al., 1♂2♀. CN: Boryeong-si, Cheongna-myeon, Mt. Oseosan, 20. vii. 1999, HY Han et al., 1♀. GB: Yeongju-si, Sunheung- myeon, Mt. Sobaeksan, 11. vi. 2004, HY Byun et al., 1♂. GN: Ulju-gun, Samnam-myeon, Gacheon-ri, Mt. Yeongchuksan, 29. vi. 2003, HY Han et al., 1♀; Yangsan-si, Ungsang-eup, Simyeongsa, 6. vii. 2008, DS Choi, 1♀. GG: Yangpyeong-gun, Yongmun-myeon, Mt. Yongmunsan, Yeonsu-ri, 30. vii. 1998, HW Byun et al., 1♀. GW: Jeongseon-gun, Nam-myeon, Mt. Mindungsan, form Yupyeong-ri to 1,119 m peak, 16. vii. 2005, HY Han et al., 1♀; ditto, 19. vii. 2005, HY Han et al., 4♀; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 14. vii. 2006, JS Lim, 1♀; ditto, 19. vii. 2009, SW Suk, 2♀. JB: Muju-gun, Sangbuk- myeon, Mt. Deogyusan, 24. vii. 1999, KH Kang, 1♀.

Distribution: Korea, Japan [PA].

Biology: Unknown.

Korea: CB, CN, GB, GN, GG, GW, JB.

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Tribe Xylotini

Genus Brachypalpus Macquart, 1834

Brachypalpus Macquart, 1834: Hist. nat. Ins., Dipt., 1: 523. TS: Brachypalpus tuberculatus Macquart (des. Rondani, 1845: Nuovi Ann. Sci. nat. Ist. Bologna, (2) 2: 456) [= valgus (Panzer)]. Crioprora Osten-Sacken, 1878: 136, 251. TS: Pocota alopex Osten-Sacken, 1877; subsequent designation by Williston, 1887: 217.

98. Brachypalpus nipponicus Shiraki, 1952 (Pl. 9L)

Brachypalpus nipponicus Shiraki, 1952: 7 Brachypalpus dentitibia Violovitsh, 1960: 249 [nomen nudum]

Korean record: Brachypalpus nipponicus: Jeong et al. 2017: 1046 (redescrip.). Brachypalpus bimaculatus [misidentification]: Shiraki 1930: 62 (redescrip., presumably first Korean record); ZSK 1968: 179 (Korean check.); Kim, CW 1971: 874 (redescrip.); Kim, JI 1975: 41 (Korean check.); Kim, JI 1980: 386 (Korean check.); ESK & KSAE 1994: 291 (Korean check.). Brachypalpus laphriformis [misidentification]: Han et al. 1998b: 151 (Korean cat.); Han & Choi 2001: 178 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 29 (Korean check.).

Description (modified from Jeong et al., 2017): Body length 10.3‒14.6 mm. Male. Head subtriangular in frontal view; vertical triangle black with long yellow pile except for shiny black bare area anterior to ocellar triangle; face dark brown with transverse brownish yellow area lateral to eye margins under antenal sockets; gena shiny black with high genal depth; frons matte black with yellowish pile at posterior 2/5; eyes narrowly dichoptic; antennae brown with scape and pedicel relatively elongated to form a straight forward antennal shape; basoflagellomere yellowish brown pubescent; arista as long as face width and brownish yellow. Thorax covered with yellowish pile; scutum subshiny black, 3/4 of scutum with pair of purplish tinged submedial longitudinal vittae. Wing hyaline with pale brown tinge on spurious vein especially near cross veins bm-cu and r-m; halter yellow to reddish yellow. Legs densely covered with long yellow pile but with some short dark pile mixed in especially on blackish colord areas; fore- and midfemora black with apices brownish yellow; fore- and midtibiae black with short proximal portions brownish yellow; fore- and midtarsi dark brown; posterior femeral and tibial pile longer than anterior pile; hind femur shiny black with basal 1/3 brownish yellow, swollen with dorso-medially bent strongly, with two vental ridges at basal 1/3 and 3/4, equipped with 2‒3 rows of strong setae. Abdomen moderately elongated and slightly

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widened between tergite 2 and 3; tergite 3 about 3/4 length of tergite 2; tergite 4 approximately as long as tergite 3. Female. Similar to male except for following non-genitalic characters: frons with anterior 1/3 bare and subshiny black, posterior with yellowish pile and yellowish white pruinosity; legs with hind femur dorso- medially bent slightly, with single ventral ridge at basal 3/4; pile sparser than in male; hind tibia with apico- ventral process short and obtuse; abdominal tergite 2 posteriorly widened.

Specimen examined: Korea: CB: Cheongwon-gun, Hyeondo-myeon, Jugam-ri, Jugam Service Area, 12. v. 2000, DS Choi et al., 1♂. GG: Gwangju-si, Chowol-myeon, Mt. Mugapsan, 5. v. 2001, DS Choi & DJ Cha, 1♂; Seongnam-si, Namhansanseong, 12. v. 1991, KY Yoon, 1♀. GW: Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 13. v. 2001, DS Choi et al., 1♂; Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 26. v. 2004, HW Byun & OY Lim, 1♂; ditto, 19. v.-6. vi. 2011, HY Han et al., 1♀; ditto, 6. vi. 2011, HY Han, 2♀; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 17. v. 2006, DS Choi & HW Byun, 2♀. JB: Muju-gun, Mujugucheondong, 11. vi. 1972, JI Kim, 1♀.

Distribution: Russian Far East, China, Korea, Japan [PA].

Biology: Unknown.

Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

Remarks: Brachypalpus nipponicus is one of the most common xylotine species in Korea. They are often found in Malaise traps, and frequently collected by insect nets sitting on logs, leaves and barks of trees, or rocks. In Korea, this species has been misidentification as Brachypalpus laphriformis, which is a European species. These two species are similar in morphology but there are some distinct differences. This species can be distinguished from the closely resembling European B. laphriformis based on the following characteristics: 1) tergum 1 without any lighter colored spot vs. with lateral pair of reddish yellow spots; 2) male eyes narrowly dichoptic vs. closely holoptic; and 3) mela head slightly subtriangular vs. almost rounded (after Jeong et al., 2017).

Genus Chalcosyrphus Curran, 1925

Chalcosyrphus Curran, 1925: Kans. Univ. Sci. Bull., 15: 122 (as subgenus of Chalcomyia Williston). TS: Chalcomyia (Chalcosyrphus) atra Curran (OD) [= depressus (Shannon)]. Hardrimyia Ferguson, 1926: 533. TS: Chrysotoxum elongatum, Hardy, 1921 (OD). Xylotodes Shannon, 1926: Proc. U.S. natn. Mus., 69: 22. TS: Brachypalpus inarmatus Hunter (OD). Xylotomima Shannon, 1926: Proc. U.S. natn. Mus., 69: 15. TS: Xylota vecors Osten Sacken (OD). Neplas Porter, 1927 96 (as subgenus of Chalcosyrphus Curran; new name for Planes Rondani, 1864: 9). Spheginoides Szilady, 1939: 138 (as subgenus of Chalcosyrphus Curran). TS: Chalcosyrphus obscura

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Szilady, 1939). Cheiroxylota Hull, 1949: 361 (as subgenus of Xylota Meigen). TS: Xylota dimidiata Brunetti, 1923). Dimorphoxylota Hippa, 1978: Acta zool. fenn., 156: 114 (as subgenus of Chalcosyrphus Curran). TS: Xylota eumera Loew (OD). Neploneura Hippa, 1978: 125 (as subgenus of Chalcosyrphus Curran). TS: Chalcosyrphus melanocephala Hippa, 1978). Syrittoxylota Hippa, 1978: Acta zool. fenn., 156: 116 (as subgenus of Chalcosyrphus Curran). TS: Xylota annulata Brunetti (OD). Xylotina Hippa, 1978: Acta zool. fenn., 156: 117 (as subgenus of Chalcosyrphus Curran). TS: Milesia nemorum Fabricius (OD).

Key to species of Chalcosyrphus from South Korea (after Jeong et al., 2017)

1. Abdominal terga almost completely black with some pattern in same black color but distinguished by pile of differently oriented microtrichiae ·················································································· 4 − Abdominal terga with some pattern in different color ······························································ 2 2. Abdominal tergite 2 reddish brown with antero-medially inverted triangular black spot, and tergite 3 entirely reddish brown ··········································································· C. piger (Fabricius) − Abdominal tergite 2 and 3 black, each with pair of brownish yellow to reddish yellow spots ················ 3 3. Abdominal tergite 2 and 3 each with pair of brownish yellow triangular spots (smaller and more reddish in female); fore- and midfemora brownish yellow; gena relatively wide (gena-eye ratio over 0.1) ··············· ·························································································· C. laterimaculatus Jeong et al. − Abdominal tergite 2 with pair of trapezoidal orange yellow markings and tergite 3 with pair of more or less rectangular orange yellow markings; fore- and midfemora entirely black; gena narrow (gena-eye ratio about 0.05) ···················································································· C. nemorum (Fabricius) 4. Hind femur entirely black ······························································································· 7 − Hind femur basally red to yellowish red and apically black ······················································· 5 5. Fore leg black; antennal pedicel and basoflagellomere dark brown to black· ········· C. unicus (Violovitsh) − Fore leg reddish yellow; antennal pedicel and basoflagellomere reddish yellow· ······························ 6 6. Arista brownish yellow; halter yellow; midfemur basally black and apically reddish yellow; hind tibia with apico-ventral process strong and sharply pointed; male abdominal tergite 4 distinctly longer than tergite 3 ························································································ C. femoratus (Linnaeus) − Arista reddish brown; halter dark brown; midfemur entirely reddish yellow; hind tibia with apico-ventral process weak; male abdominal tergite 4 about as long as tergite 3 ··························· C. rufipes (Loew) 7. Basal 1/4 of hind tibia yellow; abdominal tergite 4 with metallic greenish tinge ·· C. nitidus (Portschinsky) − Hind tibia entirely black; abdominal tergite 4 shiny black without metallic tinge ······························ 8

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8. Pale brownish wing pattern roughly confined to anterior-apical half; male abdominal tergite 4 about as long as tergite 3 ·············································································· C. carbonus (Violovitsh) − Pale brownish wing pattern more extensive, covering almost entire apical half of wing; male abdominal tergite 4 distinctly longer than tergite 3 ······················································ C. longus (Coquillett)

99. Chalcosyrphus carbonus (Violovitsh, 1975) (Pl. 10A)

Xylota carbona Violovitsh, 1975: 88.

Korean record: Chalcosyrphus carbonus: Jeong et al. 2017: 1049 (redescrip., new Korean record).

Description (modified from Jeong et al., 2017): Body length 9.4‒11.9 mm. Male. Head: Face black with heavy yellowish white pruinosity (pruinosity weaker under antenna); vertical triangle black with dark brown pile, subshiny black anterior to ocellar triangle but with some white pruinosity and dark brown and some yellow pile mixed posteriorly; frons almost bare except for narrow yellowish pruinosity close to eyes; eye contiguity length relatively short, shorter than half length of vertical triangle; antennal scape and pedicel black; basoflagellomere dark brown pubescent; arista brown. Thorax: scutum subshiny black, with dense appressed short black pile and some erect yellowish brown pile mixed; anterior pair of red-purplish tinged submedial longitudinal vittae. Wing with pale brownish wing pattern roughly confined to anterior-apical area; halter with brown stem and yellow knob. Legs entirely black; femora with long yellowish white pile, mid- and hind femora with short brownish pile mixed, posterior femoral pile longer than anterior pile; tibiae dorsally with short appressed dark brown pile and some yellowish wihte pile mixed; tarsi dorsally with yellowish brown pile and ventrally with short erect yellow pile; hind femur swollen with apico-ventral 1/3 forming ridge equipped with 2‒3 with irregular rows of strong setae, short brown pile and yellowish white pile mixed; hind tibia with apico-ventral process sharply pointed. Abdomen moderately elongated; tergite 2 and 3 almost parallel-sided but, together posteriorly slightly widened; tergite 2 and 3 each with pair of large shiny triangular areas covered with long erect pale pile (other areas subshiny, covered with appressed short black pile); tergite 4 shiny, apically narrowed, as long as tergite 3, with pair of shallow humps in middle. Female. Similar to male except for following non-genitalic characters: head with anterior 1/3 of frons bare and subshiny black; posterior 2/3 of frons with yellowish pile and heavy yellwish white pruinosity; legs with hind tibia with short and obtuse apico-ventral process; abdominal tergite 2 and 3 both slightly widened posteriorly.

Specimen examined: Korea: GW: Inje-gun, Mt. Bangtaesan, 4-6. vi. 1999, SS Kwak, 1♂; Pyeongchang-

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gun, Jinbu-myeon, Tapdong-ri, Mt. Odaesan, 27. v.-4. vii. 2013, SJ Park and SB Oh, 1♂; ditto, 28. vi.-31. vii. 2012, SJ Park and JJ Huh, 1♀; Hongcheon-gun, Nae-myeon, Mt. Gachilbong, from Sambongyaksu to 1,240 m peak, 16. vi. 2013, DS Choi, 1♂; Jeongseon-gun, Nam-myeon, Mt. Mindungsan, form Yupyeong-ri to 1,119 m peak, 28. vi. 2012, HY Han et al., 1♂; Wonju-si, Panbu-myeon, Seogok-ri, Mt. Baegunsan, 26. vii. 2012, HY Han et al., 1♂. JB: Namwon-si, Sannae-myeon, Mt. Jirisan, Baemsagol (646 m), 11. vii.-5. ix. 2001, DS Ku, 1♀. Distribution: Russian Far East, Korea [PA]. Biology: Unknown. Korea: GW, JB.

100. Chalcosyrphus femoratus (Linnaeus, 1758) (Pl. 10B)

Musca femorata Linnaeus, 1758: 595. Syrphus volvulus Fabricius, 1794: 295. Xylota curvipes Loew, 1854: 19. Xylota femorata Matsumura 1911: 74. Zelima sapporoensis Shiraki, 1930: 76.

Korean record: Chalcosyrphus femoratus: Jeong et al. 2017: 1049 (redescrip.). Chalcosyrphus sapporoensis: Peck 1988: 223 (Palaearctic cat.); ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998b: 151 (Korean cat.); Choi et al. 2000: 145 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 180 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 29 (Korean check.). Chalcosyrphus sapporensis [misspelling]: Han et al. 1998a: 11 (Yonsei Univ. Maeji Campus). Xylota sapporensis [misspelling]: Kim, JI 1980: 386 (Korean check.). Zelima sapporoensis: Shiraki 1930: 76 (orig. desc., presumably first Korean record); ZSK 1968: 180 (Korean check.); Kim, CW 1971: 872 (redescrip.).Zelima spporoensis [misspelling]: Kim, JI 1975: 41 (Korean check.).

Description (modified from Jeong et al., 2017): Body length 16.0‒21.1 mm. Male. Head almost entirely black; face with whitish pruinosity except for shiny area under antennae; vertical triangle shiny and bare anterior to ocellar triangle but with some dark brown pile and yellow pile mixed posteriorly; frons largely shiny except for lateral yellowish pruinose areas; antennal scape black; pedicel reddish yellow; basoflagellomere reddish yellow pubescent; arista brownish yellow. Thorax with scutum subshiny black, densely covered with short yellow and brown pile mixed; anterior half of scutum with pair of purplish tinged submedial longitudinal vittae. Wing hyaline with pale brownish tinge roughly conflined to anterior-apical area; halter yellow to brown. Legs: coxae and trochanters black, densely covered

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with dark brown pile; posterior femoral and tibaial pile longer than anterior pile; fore leg entirely reddish yellow with yellow pile; midleg predominantly reddish yellow with yellow pile; midfemur coloration highly variable from almost entirely reddish yellow to almost black except for short apical reddish yellow area; midtarsus reddish yellow with appressed yellow pile; hind femur swollen with apico-ventral 1/3 forming ridge equipped with 2‒3 irregular rows of strong spinose setae; basal half of hind femur reddigh yellow in ground color with mostly yellowish pile, but apical half black with blackish pile; hind tibia black, dorsally with appressed dark brown pile and ventrally with erect long black pile; strongly curved inward with apico- ventral process strong and shrply pointed. Abdomen moderately elongated; tergite 2 and 3 almost parallel- sided but, tergite 3 posteriorly slightly widened; anterior 2/3 of tergite 2 with laterally with pair of shiny areas covered by long erect pale whitish pile (other areas of tergite 2 with appressed short black pile); tergite 3 about 4/5 as long as tergite 2, subshiny, anterior half laterally with pair of large shiny areas covered by long erect pale whitish pile; tergite 4 about 1.5 times as long as tergite 3, subshiny black with short black pile, with distinct pair of humps on apical 1/3. Female. Similar to male except for following non-genitalic characters: frons with anterior 1/3 bare and shiny black, medial area subshiny black with dark brown pile, laterally with whitish pruinosity, posterior with dark brown pile; legs with hind femur less stronger than in males; abdominal tergite 2 and 3 together posteriorly slightly widened.

Specimen examined: Korea: GN: Sancheong-gun, Jungsan-ri, Beopgyesa Temple, 17.-18. viii. 1992, JS Jeon, 1♂. GW: Bukpyeong, Samhwasa Temple, 26. vi. 1984, HS Choi, 1♂; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 10. vi. 1999, HY Han et al., 1♂; Hongcheon-gun, Nae- myeon, Mt. Gachilbong, from Sambongyaksu to 1,240 m peak, 17. vi. 2005, DS Choi et al., 2♂; ditto, 12. vi. 2004, HW Byun et al., 1♂; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, N. Valley, 20. vi. 1999, HY Han & KE Ro, 2♂; Jeongseon-gun, Jeongseon-eup, Hoedong-ri, Mt. Gariwangsan, 28. vii. 2009, JD Yeo, 1♂; Mt. Seoraksan, Baekdamsanjang, 27. vi. 2002, HS Lee, 1♂.

Distribution: Widely distributed in Europe and Northeast Asia [PA].

Biology: Unknown.

Korea: GN, GW.

Remarks: In Korean fauna, C. femoratus, C. rufipes and C. unicus are very similar and this similarity might have caused many misidentifications in the past (after Jeong et al., 2017).

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101. Chalcosyrphus laterimaculatus Jeong, Jung and Han, 2017 (Pls. 10C, D)

Chalcosyrphus laterimaculatus Jeong, Jung and Han, 2017: 1051. Zelima frontalis Shiraki and Edashige, 1953: 116. Xylota okadomei Violovitsh, 1976: 127.

Korean record: Chalcosyrphus laterimaculatus: Jeong et al. 2017: 1051 (redescrip.). Xylota frontalis: Kim, JI 1980: 386 (Korean check.); ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998a: 11 (Yonsei Univ. Maeji Campus); Han et al. 1998b: 153 (Korean cat.); Choi et al. 2000: 145 (Baegunsan & Yonsei Univ. Maeji Campus); Han & Choi 2001: 183 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 30 (Korean check.). Zelima frontalis: Kim, JI & Kim, CW 1972: 11 (new Korean record); Kim, JI 1975: 41 (Korean check.); Kim, CW 1980: 297 (distr. Map).

Description (modified from Jeong et al., 2017): Body length 10.9‒11.9 mm. Male. Head: face brownish yellow with yellowish white pruinosity except for shiny black area under antenna; vertical triangle black with dark brown pile, shiny black anterior to ocellar triangle but with yellow pile posteriorly; frons predominantly pale brownish yellow with heavy yellow pruinosity, with lateral blackish area near antennal socket; antennal scape, pedicel yellowish brown; basoflagellomere yellowish brown to reddish brown pubescent; arista basally yellow and apically darker. Thorax with scutum subshiny black with heavy short yellow pile and some brownish pile mixed; anterior pair of submedial longitudinal vittae purplish tinged. Wing hyaline with pale brown tinge; halter pale yellow. Legs: coxae black; trochanters dark brown to black with whitish pile; fore- and midlegs brownish yellow with yellowish pile, posterior pile longer than anterior pile; fore- and midtarsi with tarsomeres 1‒3 brownish yellow and tarsomere 4 and 5 dark brown; hind femur shiny black, basal 1/5 brownish yellow with short whitish yellow pile, swollen with apico- ventral 1/3 forming ridge equipped with 2‒3 irregular rows of strong spinose setae; hind tibia dark brown, dorsally with appressed yellow pile and ventrally with erect short black pile; hind tibia with apico-ventral process short and obtuse. Abdomen moderately elongated; tergite 2 and 3 more or less parallel-sided with slight constriction in between; tergite 2 and 3 black, each with pair of large laterally facing brownish yellow triangular markings, areas of brownish yellow marking and side of tergites with long erect pale yellow pile but blackish area mostly have short dark pile; tergite 4 shiny black with short yellow pile; tergite 4 as long as tergite 3 with pair of shallow humps on apical 1/3. Female. Similar to male except for following non-genitalic characters (please note extreme secual dimorphism): head black with frons anterior 1/3 bare and shiny, about medial 1/3 subshiny with heavy pruinosity; posterior 1/3 subshiny with yellow pile and white pruinosity; legs with basal 4/5 of hind femur reddish yellow with strong ventral ridge; abdominal tergite 2 posteriorly slightly widened; tergite 2 and 3 with reddish yellow triangular spots smaller and sharper than male; tergite 4 with small lateral reddish yellow

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triangular spots, without any hump.

Specimen examined: Korea: CB: Cheongwon-gun, Namseong-myeon, Mt. Ingyeongsan, 15. viii. 1997, HY Han et al., 1♂2♀; Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 2. vii. 2003, DS Choi et al., 1♂. GB: Yeongju-si, Buseok-myeon, Mt. Eoraesan, from Namdae-ri to 1,064 m peak, 30. vi. 1998, HY Han et al., 1♂. GG: Namyangju-si, Sudong-myeon, Mt. Chungnyeongsan, 30. vi. 1999, HY Han et al., 1♀. GW: Hoengseong-gun, Anheung-myeon, Mt. Chiaksan, from Gangrim 4-ri to Cheonjibong (1,086 m) peak, 14. vi. 1999, HY Han et al., 1♂; Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 5. vi. 2003, DS Choi & HS Lee, 1♂; Pyeongchang-gun, Bangrim-myeon, Ungyo-ri, 24. ix. 2010, SW Suk, 1♀; Taebaek-si, Hyeol-dong, Mt. Taebaeksan, from Yuilsa to 1,560 m peak, 12. viii. 2000, DS Choi et al., 1♀; Wonju-si, Gwirae-myeon, from Cheoneunsa to Mt. Sipjabong peak, 17. v. 2003, DS Choi et al., 1♂; Wonju- si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 26. v. 2005, SW Suk & S Hwang, 1♂; ditto, 26. v. 2006, SMR Hwang & JS Lim, 9♂2♀; 21. v. - 27. vi. 2014, HY Han et al., 1♀; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 17. viii. 1999, HY Han et al., 2♂1♀; Wonju-si, Sillim-myeon, Mt. Chiaksan, from Seongnam-ri to Namdaebong (1,181.5 m) peak, 30. vii. 2003, OY Lim & HS Lee, 3♂.

Distribution: Korea, Japan (Shikoku) [PA].

Biology: Unknown.

Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

Remarks: This species has a distinct sexual dimorphism. Due to this, different sexes of this species were regarded as different Species: Zelima frontalis Shiraki and Edashige, 1953, for males and Xylota okadomei Violovitsh, 1976, for females, until Kimura (1998) recognized them as being conspecific. Recognizing this species under the genus Chalcosyrphus creates a homonymy problem, since the specific name,frontalis , was preoccupied by a Neotropical species, Chalcosyrphus (Neplas) frontalis Curran, 1941. Jeong et al. (2017) provided a new name for this species as Chalcosyrphus laterimaculatus Jeong, Jung and Han, new name. This species is one of the most commonly collected Chalcosyrphus specis in Korea by Malaise trap or insect net (after Jeong et al., 2017).

102. Chalcosyrphus longus (Coquillett, 1898)

Xylota longa Coquillett, 1898: 327.

Korean record: Chalcosyrphus longus: ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998b: 151 (Korean cat.); Han & Choi 2001: 179 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 29 (Korean check.); Jeong et al. 2017: 1051 (redescrip.). Xylota longa: Kim, JI 1980: 386 (Korean check.). Zelima longa: Doi 1938a: 10 (Korean check., presumably first Korean record); ZSK 1968: 180

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(Korean check.); Kim, CW 1971: 872 (redescrip.); Kim, JI 1975: 41 (Korean check.).

Distribution: Russian Far East (Kuril Is, Primorye Territory), Korea, Japan (Hokkaido, Honshu) [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim (1971).

103. Chalcosyrphus nemorum (Fabricius, 1805) (Pls. 10E, F)

Milesia nemorum Fabricius, 1805: 192. Xylota bifasciata Meigen, 1822: 219. Xylota pallidipes Stackelberg, 1922: 27. Syrphus interruptus Panzer, 1804: 168. Xylota baton Walker, 1849: 554. Xylota fraudulosa Loew, 1864: 71. Xylota arsenjevi Violovitsh, 1980: 127.

Korean record: Chalcosyrphus nemorum: Jeong et al. 2017: 1052 (redescrip., new Korean record).

Description (modified from Jeong et al., 2017): Body length 8.5‒8.8 mm. Male. Head: face and frons with heavy brownish yellow pruinosity; vertical triangle black with brownish yellow pile, subshiny black anterior to ocellar triangle but with heavy yellowish brown pruinosity posteriorly, with reddish brown ocelli; antennal scape, pedicel black; basoflagellomere dark brown pubescent; arista brown with darker apex. Thorax with scutum subshiny black with dense short yellow pile, with anterior pair of submedial longitudinal vittae purplish tinged. Wing hyaline with pterostigma pale yellowish brown; halter pale yellow. Legs predominantly black with some yellowish brown portions, with short pale yellowish brown to white pile; fore- and midfemora black with apices brownish yellow; posterior femoral pile longer than anterior pile; fore- and midtibiae black with short proximal portions brownish yellow with short appressed pile; fore- and midtarsomeres 1‒3 yellow; tarsomeres 4 and 5 black; hind femur shiny black, swollen with apico-ventral 1/3 forming ridge equipped with 2‒3 irregular rows of strong spinose setae; hind tibia dark brown, dorsally with appressed pale yellowish brown pile and ventrally with erect black pile; hind tibia with apico-ventral process short and obtuse; hind tarsus black with yellowish appressed pile. Abdomen moderately elongated; tergite 2 black with pair of large trapezoidal orange yellow markings, areas of orange yellow marking and side of tergites with pale yellowish brown pile but blackish area mostly have dark pile; tergite 3 black with pair of more or less rectangular orange yellow markings, pile colors similar to those of tergite 2 but lateral pile much shorter; tergite 4 shiny black, as long as tergite 3, apically narrowed, with indistinct pair

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of humps on apical 1/3. Female. Similar to male except for following non-genitalic characters: head with frons anteriorly bare, medially with yellowish white pruinosity; abdominal tergite 2 and 3 together posteriorly slightly widened.

Specimen examined: Korea: GW: Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 1. v. 2016, SS Euo et al., 1♂; ditto, 1. - 21. v. 2006, HY Han et al., 1♀. JJ: Bukjeju-gun, Aewol-eup, Mt. Hallasan, 1,100 goji, 7. viii. 2003, HY Han et al., 1♀.

Distribution: Widely distributed in Holarctic Region [PA, NE].

Biology: Unknown.

Korea: GW, JJ.

104. Chalcosyrphus nitidus (Portschinsky, 1879) (Pls. 10G, H)

Xylota nitida Portschinsky, 1879: 157.

Korean record: Chalcosyrphus nitidus: Jeong et al. 2017: 1053 (redescrip., new Korean record).

Description (modified from Jeong et al., 2017): Body length 8.9‒12.0 mm. Male. Head: face with heavy yellowish white pruinosity; vertical triangle subshiny black with yellowish pruinosity, bare anterior to ocellar triangle but with yellowish pile posterior to it; frons anteriorly subshiny black but with yellowish white pruinosity on posterior half and lateral areas; antennal scape and pedicel dark brown; basoflagellomere brown to dark brown pubescent; arista dark brown. Thorax with scutum subshiny black, densely covered with short yellow pile; anteriorly with pair of purplish tinged submedial longitudinal vittae. Wing hyaline with pterostigma dark brown; halter yellow. Legs predominantly black with about basal 1/4 of tibiae yellow with short pale yellowish white pile; fore- and midfemora black with apices brownish yellow; posterior femoral pile longer than anterior pile; fore- and midtibiae with appressed short pile; fore tarsus dark brown with appressed short yellow pile; midtarsomeres 1 and 2, each with basal half brown and apical half yellow; midtarsomeres 3‒5 dark brown to black with appressed brownish yellow pile; hind femur shiny black, swollen with apico-ventral 1/3 forming weak ridge equipped with 2‒3 irregular rows of strong spinose setae; hind tibiae predominantly black with basal 1/3 yellow (areas of yellow marking covered with pale yellow pile but blackish area mostly covered with dark pile), without apico-ventral process, dorsally with appressed pile, ventrally with erect short dark brown pile; hind tarsus dark brown, dorsally covered with dark brown pile and ventrally with appressed yellowish pile. Abdomen moderately elongated, about as long as head and body together; more or less paralled-sided, laterally with long pale whitish pile; tergite 2 and 3 shiny with purplish tinge, medially with short black pile, laterally with pale whitish pile; tergite 4 shiny with

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greenish tinge, with short yellow pile, as long as tergite 3, apically narrowed, with indistinct pair of humps on apical 1/3. Female. Similar to male except for following non-genitalic characters: head black in ground color with anterior 1/3 of head bare and subshiny, about medial 1/3 with yellowish white pruinosity and yellowish pile, about posterior 1/3 subshiny with yellowish pile; abdominal tergite 2 and 3 together posteriorly slightly widened; tegite 4 widened and rounded apically.

Specimen examined: Korea: GB: Bonghwa-gun, Murya-myeon, Mt. Seondalsan, from Ojeon-ri to 1,236 m peak, 4. v. 1998, HY Han et al., 1♂; Uljin-gun, Onjeong-ri, Mt. Baegamsan, 16. viii. 1999, HY Han et al., 1♀; Yeongju-si, Sunheung-myeon, Mt. Sobaeksan, 11. vi. 2004, HW Byun et al., 1♂1♀. GG: Gwangju- si, Docheok-myeon, Mt. Taehwasan, 5. v. 2001, SK Kim & OY Lim, 1♀; Pocheon-gun, Mt. Myeongjisan, 3. v. 1997, HY Han, 1♂. GW: Hoengseong-gun, Anheung-myeon, Mt. Chiaksan, from Gangrim 4-ri to Cheonjibong (1,086 m) peak, 16. v. 1999, DS Choi et al., 3♂1♀; Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 25. v. 2004, HW Byun et al., 2♂1♀; Hongcheon-gun, Nae-myeon, Mt. Gyebangsan, from Unduryeong to 1,577 m peak, 25. viii. 2002, HW Byun & OY Lim, 3♂10♀; Inje- gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 29. v. 1999, HY Han et al., 5♂3♀; Jeongseon-gun, Gohan-eup, Mt. Hambaeksan, 14. viii. 1999, HY Han et al., 59♂82♀; Taebaek-si, Mt. Taebaeksan, from Yuilsa to 1,560 m peak, 29. viii. 1999, DS Choi & CH Park, 14♂11♀; Wonju-si, Panbu- myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 5. v. 2003, HW Byun, 31♂12♀. JB: Muju-gun, Seolcheon-myeon, Daebul-ri, 26. v. 1993, 2♂. JN: Gurye-gun, Sandong-myeon, Mt. Jirisan, Nogodan (1,507 m), 8. ix. 2006, DS Choi, 1♀. JJ: Bukjeju-gun, Jocheon-eup, Mt. Hallasan, 26. vi. 2003, DS Choi et al., 8♂3♀; Seogwipo-si, Mt. Hallasan, 3. v. 2003, OY Lim & HS Lee, 1♀.

Distribution: Russia, Korea, China, Japan [PA].

Biology: Unknown.

Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

105. Chalcosyrphus piger (Fabricius, 1794) (Pl. 10I)

Syrphus piger Fabricius, 1794: 295. Milesia haematodes Fabricius, 1805: 103 Xylota crassipes Wahlberg, 1839: 15 Xylota nigra Walker, 1849: 554. Xylota pini Perris, 1870: 330. Xylota rubbiginigaster Bigot, 1884: 543.

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Korean record: Chalcosyrphus piger: Jeong et al. 2017: 1053 (redescrip., new Korean record).

Description (modified from Jeong et al., 2017): Body length 12.0‒12.9 mm. Male. Head: face black with slight white pruinosity except for shiny area under antenna; vertical triangle black, bare anterior to ocellar triangle but with few white pruinosity and yellowish white pile posteriorly; frons largely subshiny black except for lateral slightly yellowish pruinose areas; antennal scape and pedicel black; basoflagellomere dark brown pubescent; arista dark brown. Thorax with scutum matte black, covered with short strong black pile and some short yellowish brown pile mixed; anterior half with pair of yellowish tinged submedial longitudinal vittae. Wing hyaline but almost entirely pale brown tinged with pterostigma brown; halter with brown stem and yellow knob. Legs entirely shiny black, covered with relatively short and sometimes stout pile; fore- and midfemora with black and reddish yellow pile mixed; fore- and midtibiae with brown and yellow pile mixed; fore- and midtarsi black to dark brown with appressed short reddish yellow pile; hind tibia with apico-ventral process sharply pointed; hind tarsus dorsally with short dark brown pile and ventrally densely with appressed yellow pile. Abdomen moderately elongated, about 1.3 times as long as head and body together; tergite 2 and 3 paralled-sided; tergite 2 shiny reddish brown antero-medially with inverted triangular black spot, antero-laterally with erect short whitish pile; tergite 3 and 4 similar in length, shiny reddish brown with short black pile. Female. No specimens available.

Specimen examined: Korea: CB: Jecheon-si, Songhak-myeon, Mt. Songhaksan, 22. v. 2015, HY Han et al., 1♂; ditto, 4. vi. 2012, YB Lee et al., 2♂. GB: Andong-si, Girin-myeon, Songsa-ri, Mt. Cheonjigapsan, 2. vi. 2012, SW Suk et al., 1♂. GW: Jeongseon-gun, Jeongseon-eup, Mt. Gariwangsan, 4. vi. 2013, HS Lee et al., 1♂.

Distribution: Holarctic Region [PA, NE].

Biology: Unknown.

Korea: CB, GB, GW.

106. Chalcosyrphus rufipes (Loew, 1873) (Pl. 10J)

Xylota rufipesLoew, 1873: 212.

Korean record: Chalcosyrphus rufipes: Jeong et al. 2017: 1055 (redescrip., new Korean record).

Description (modified from Jeong et al., 2017): Body length 12.8‒14.1 mm. Male. Head almost entirely black with largely brown antennae; face with yellowish white pruinosity except

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for shiny area under antenna; vertical triangle subshiny bare anterior to ocellar triangle but posteriorly with black pile and few yellow pile mixed; frons largely shiny except for lateral yellowish pruinose areas; antennal scape, pedicel reddish brown; basoflagellomere reddish brown pubescent; arista reddish brown. Thorax with scutum subshiny black with short appressed yellow pile and erect yellow to brown pile mixed; anterior half of scutum with pair of faint purplish tinged submedial longitudinal vittae. Wing hyaline with pale brown tinge; halter dark brown. Legs: coxae black with yellowish pile; trochanters black; fore- and midfemora reddish yellow with yellow to reddish yellow pile; posterior femoral pile longer than anterior pile; fore- and midtibiae reddish yellow with short appressed yellowish pile; fore- and midtarsi yellow with yellowish white pile; hind femur shiny black with basal 1/3‒1/2 reddish yellow with long yellowish pile, areas of reddish yellow markings of hind femur with yellowish pile but blackish area mostly with black pile, swollen with apico- ventral 1/3 forming weak ridge equipped with short spinose setae; hind tibia black, dorsally with appressed dark brown pile and ventrally with erect short black pile, with apico-ventral process sharply pointed; hind tarsus black to dark brown, dorsally with short dark brown pile, ventrally with appressed yellow pile, ventro- subapically with short yellow spinose setae. Abdomen, elongated with tergite 2 and 3 more or less parallel- sided but tergite 3 posteriorly slightly widened; Anterior half of tergite 2 and 3 each with pair of large shiny trianglular areas covered with long erect pale whitish pile (other areas subshiny, covered with appressed short black pile); tergite 3 with indistinct pair of humps in middle; tergite 4 as long as tergite 3, with distinct pair of humps on basal 1/3. Female. Similar to male except for following non-genitalic characters: head black in ground color with anterior 1/3 of head bare and subshiny, about medial 1/3 with yellowish white pruinosity and yellowish pile, about posterior 1/3 subshiny with yellowish pile; legs with hind femur slightly less swollen; abdominal tergite 2 and 3 together posteriorly slightly widened.

Specimen examined: Korea: GW: Inje-gun, Girin-myeon, Mt. Jeombongsan, from Jindong-ri to 1,424 m peak, 29. v. 1999, HY Han et al., 1♂; Pyeongchang-gun, Yongpyeong-myeon, Mt. Gyebangsan, 17. vii. - 12. viii. 2008, HR Seo and TW Kim, 1♂; Hongcheon-gun, Nae-myeon, Mt. Gachilbong, from Sambongyaksu to 1,240 m peak, 2. vi. 2007, HS Lee et al., 1♀.

Distribution: Europe, Russia, Mongolia, Korea, Japan [PA].

Biology: Unknown.

Korea: GW.

107. Chalcosyrphus unicus (Violovitsh, 1977) (Pl. 10K)

Xylota unica Violovitsh, 1977: 82.

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Korean record: Chalcosyrphus unicus: Jeong et al. 2017: 1058 (redescrip., new Korean record). Chalcosyrphus sapporoensis [misidentification]: Han & Choi 2001: 214 (Korean check.).

Description (modified from Jeong et al., 2017): Body length 12.1‒18.8 mm. Male. Head almost entirely black; face with whitish pruinosity except for shiny area under antenna; vertical triangle shiny and bare anterior to ocellar triangle but with black and some yellow pile mixed posteriorly; frons largely shiny except for lateral yellowish pruinose areas; antennal scape and pedicel black; basoflagellomere brownish black pubescent; arista dark brown. Thorax with scutum subshiny black, predominantly with appressed yellowish pile and erect brown pile mixed; anterior half of scutum with pair of purplish tinged submedial longitudinal vittae. Wing hyaline with pale brown tinge roughly confined to anterior-apical area; halter yellow to reddish yellow. Legs: fore- and midlegs black; fore- and midfemora and tibiae covered with whitish yellow curly pile; posterior femoral and tibial pile longer than anterior pile; fore- and midtarsi dorsally with appressed whitish yellow pile and ventrally with erect brown pile; basal 1/3 of hind femur reddish orange areas of blackish area with long and curly blackish pile but reddish orange area with mostly yellowish pile, swollen with apico-ventral 1/2 forming ridge equipped with 2‒3 irregular rows of strong setae; hind tibia black, dorsally with short appressed black pile and ventrally with erect long and curly black pile, with apico-ventral process strong and sharply pointed; hind tarsus black with dorsally with short erect black pile and ventrally with appressed brown pile. Abdomen moderately elongated, about 1.5 times as long as head and boty together; tergite 2 and 3 almost parallel-sided but tergite 3 posteriorly slightly widened; anterior half of tergite 2 and 3 each with pair of large shiny trianglular areas covered with long erect pale whitish pile (other areas subshiny, covered with appressed short black pile); tergite 3 with indistinct pair of humps on anterior 1/3; tergite 3 about 4/5 as long as tergite 2; tergite 4 subshiny with short black to dark brown pile, 1.7 times as long as tergite 3, with distinct pair of humps in middle. Female. Similar to male except for following non-genitalic characters: head black in ground color; anterior 1/3 of frons almost bare and subshiny, about medial 1/3 with whitish yellowish pruinosity, posterior 1/3 subshiny with yellowish pile; legs with hind femoral pile sparser than in males; abdominal tergite 2 and 3 together posteriorly slightly widened.

Specimen examined: Korea: CB: Seosan-si, Haemi-myeon, Daedok-ri, Hanseo Univ., 22. v. - 1. vi. 2013, JK Kim 1♀; ditto, 1-25. vi. 2013, JW Lee et al., 2♀. GB: Gyeongsan-si, Dae-dong, Yeungnam Univ., 1-8. v. 2009, JW Lee et al., 1♂; Cheongsong-gun, Hyunseo-myeon, Baksanjae, 23. ix. 2011, JC Jeong, 1♂; Yeongyang-gun, Subi-myeon, Suha-ri, Songbang, 11. vi. 2012, JW Lee et al., 1♀; Cheongdo-gun, Unmun- myeon, Munsuseowon, 10. vii. 2012, JW Lee et al., 1♀. GN: Ulsan, Ulju-gun, -ri, 25. vi. - 15. vii. 1987, ??, 1♀; Namhae-gun, Sangju-myeon, 23. v. 2004, HJ Lee, 1♀. GG: Seoul, Ui-dong, 8. vii. 1975, SH Yu; Seoul, Seocho-gu, Heoninleung, 11. vi. 1983, 1♂. GW: Wonju-si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 8. vii. 1996, HW Byun, 1♂; ditto, 16. vi. 2003, HW Byun, 1♀; ditto, 9. vii. 2007, JS Lim,

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1♀; ditto, 5. vii. 2009, DJ Cha, 1♂. JB: Jeongeup-si, Yongsan-dong, 21. iv. - 19. v. 2004, JW Lee et al., 1♀; Wanju-gun, Dongsang-myeon, Daea-ri, Daea arboretum, 24. v. - 17. vi. 2011, JW Lee et al., 1♀; Muju-gun, Mupung-myeon, Hyeonnae-ri, Mt. Bakseoksan, 7-20. v. 2015, JW Lee, 1♀.

Distribution: Russian Far East, Korea [PA].

Biology: Unknown.

Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

Genus Xylota Meigen, 1822

Xylota Meigen, 1822: Syst. beschr., 3: 211. TS: Musca sylvarum Linnaeus (Unjustified new name for Heliophilus Meigen) Zelima Meigen, 1800: Nouve. Class.: 34. TS: Musca segnis Linnaeus (des. Coquillett 1910: Proc. U.S. natn. Mus., 37: 621). Suppressed by I.C.Z.N. 1963, Opinion 678: 339. Eumeros Meigen, 1803: Mag. Insektenk., 2: 273. TS: Musca segnis Linnaeus (des. Coquillett 1910: Proc. U.S. natn. Mus., 37: 541). Heliophilus Meigen, 1803: Mag. Insektenk., 2: 273. TS: Musca sylvarum Linnaeus (MO) (as Syrphus sylvarum Fabricius). Micraptoma Westwood, 1840: Synopsis, 2: 136. TS: Musca segnis Linnaeus (MO) [For discussion of nomenclature problem see Wirth, et al. 1965: 604].

108. Xylota abiens Meigen, 1822

Xylota abiens Meigen, 1822: 218. Musca semulator Harris, 1780: 112. Zelima subabiens Stackelberg, 1952: 326.

Korean record: Xylota abiens: ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998b: 152 (Korean cat.); Han & Choi 2001: 181 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 29 (Korean check.). Zelima abiens: Kim, CW & Nam 1982: 130 (Gyebangsan, Sogyebangsan & Gachilbong, first Korean record).

Distribution: Europe, Russian Far East, Korea?, Japan [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim & Nam (1982).

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109. Xylota coquilletti Hervé-Bazin, 1914 (Pl. 10L)

Xylota coquilletti Hervé-Bazin, 1914: 409. Xylota amamiensisi Shiraki, 1968b: 122. Xylota cuprina Coquillett, 1898: 327. Xylota huangshanensis He and Chu, 1992: 6. Xylota silvicola Mutin, 1991: 103.

Korean record: Xylota coquilletti: Kim, JI 1980: 386 (Korean check.); ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998b: 152 (Korean cat.); Han & Choi 2001: 182 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 30 (Korean check.). Zelima coquilletti: Doi 1938a: 11 (Korean check., presumably first Korean record); ZSK 1968: 180 (Korean check.); Kim, CW 1971: 873 (redescrip.); Kim, JI 1975: 41 (Korean check.); Kim, CW 1980: 296 (distr. Map).

Description: Body length 10.5‒12.3 mm. Male. Head: face black with yellow pruinosity; vertical triangle with yellow hair; antennal scape and pedicel black; basoflagellomere brown with brown pruinosity; arista brown. Thorax: scutum black with metallic greening tinge; anterior a pair of distinct medial longitudinal vittae. Wing hyaline and clear; halter pale yellow. Legs: coxae dark brown to black with white hair; trochanters dark brown with yellow hair; hind trochanter with sharp and short spur; fore and midfemora black; fore and midtibiae yellow with subapico- ventral 1/2 dark brown. Abdomen as long as head and thorax together; tergite 2 almost parallel each side to slightly narrow posterior; tergite 3 slightly widened posteriorly; tergite 4 black with lateral yellow hair. Female. Similar to male except for following non-genitalic characters: eyes distinctly dichoptic; hind trochanter without spur; abdomen with slightly ovate shaped with tergite 2 and 3 posteriorly widened.

Specimen examined: Korea: CB: Cheongwon-gun, Namseong-myeon, Mt. Ingyeongsan, 15. viii. 1997, HY Han et al., 1♂1♀; Chungju-si, Jongmin-dong, Mt. Gyemyeongsan, 2. vii. 2003, DS Choi et al., 1♂. GB: Yeongju-si, Buseok-myeon, Mt. Eoraesan, from Namdae-ri to 1,064 m peak, 30. vi. 1998, HY Han et al., 1♂. GG: Namyangju-si, Sudong-myeon, Mt. Chungnyeongsan, 30. vi. 1999, HY Han et al., 1♀. GW: Hoengseong-gun, Anheung-myeon, Mt. Chiaksan, from Gangrim 4-ri to Cheonjibong (1,086 m) peak, 14. vi. 1999, HY Han et al., 1♂; Hoengseong-gun, Dunnae-myeon, Mt. Cheongtaesan, Sapgyo-ri to 1,200 m peak, 5. vi. 2003, DS Choi & HS Lee, 1♂; Pyeongchang-gun, Bangrim-myeon, Ungyo-ri, 24. ix. 2010, SW Suk, 1♀; Taebaek-si, Hyeol-dong, Mt. Taebaeksan, from Yuilsa to 1,560 m peak, 12. viii. 2000, DS Choi et al., 1♀; Wonju-si, Gwirae-myeon, from Cheoneunsa to Mt. Sipjabong peak, 17. v. 2003, DS Choi et al., 1♂; Wonju- si, Heungeop-myeon, Maeji-ri, Yonsei Univ. Campus, 26. vi. 2003, HW Byun, 1♀; Wonju-si, Panbu-myeon, Mt. Baegunsan, from Yongsugol to 1,087.1 m peak, 17. viii. 1999, HY Han et al., 1♂1♀; ditto, 25. vii. 2002,

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1♂; Wonju-si, Sillim-myeon, Mt. Chiaksan, from Seongnam-ri to Namdaebong (1,181.5 m) peak, 30. vii. 2003, OY Lim & HS Lee, 1♂.

Distribution: W. Siberia (Altai, Sayan, Tuba), Russian Far East (Khabarovsk and Primorye Territories, Sakhalin, Kuril Is), Korea, Japan, Oriental Region [PA, OR].

Biology: Unknown.

Korea: CB, CN, GB, GN, GG, GW, JB, JN, JJ.

110. Xylota ignava (Panzer, 1798)

Syrphus ignavus Panzer, 1798: 4. Musca pigra Schrank, 1803: 114. Xylota basalis Matsumura, 1911: 74. Xylota inermis Becker, 1921: 86.

Korean record: Xylota ignava: Kim, JI 1980: 386 (Korean check.); Peck 1988: 225 (Palaearctic cat.); ESK & KSAE 1994: 291 (Korean check.); Han et al. 1998b: 153 (Korean cat.); Han & Choi 2001: 184 (Korean check.); Paek et al. 2010: 232 (Korean check.); Han et al. 2014: 30 (Korean check.). Zelima ignava: Doi 1938a: 11 (Korean check., presumably first Korean record); ZSK 1968: 180 (Korean check.); Kim, CW 1971: 873 (redescrip.); Kim, JI 1975: 41 (Korean check.).

Distribution: Palaearctic Region [PA].

Biology: Unknown.

Remarks: We could not find specimens but cited this species based on Kim (1971).

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Chalcosyrphus Curran (Insecta: Diptera: Syrphidae) in Korea. Journal of Asia-Pacific Entomology 20(2017): 1043-1064. Kevan, P. G. 2002. Flowers, pollination, and the associated diversity of flies. Biodiversity 3(4): 16–18. Kim, C. W. 1971. Family 26. Syrphidae. pp. 827-887 +plts 47-58 in Kim, C. W. et al. [co-authors], Illustrated Encyclopedia of Fauna and Flora of Korea, Vol. 12, Insecta (IV). Ministry of Education, Korea. [in Korean] Kim, C. W. 1980. Distribution atlas of insects of Korea. Series 3 Hymenoptera and Diptera. Korea University Press, Seoul, Korea. [in Korean] Kim, C.-W. and Nam, S.-H. 1978. Insect fauna of Imgye-myeon area in summer season. Report of the Korean Association for Conservation of Nature 13: 125-142. [in Korean] Kim, C.-W., Kim, J.-I., Oh, J.-K., Noh, Y.-T. and Shin, Y.-H. 1974. Faunistic study of insects near the DMZ. Report of the Korean Association for Conservation of Nature 7: 182-257. [in Korean] Kim, J.-I. 1974. On the four unrecorded Syrphidae from Korea (Diptera). Korean Journal of Entomology 4(1): 8-9. Kim, J.-I. 1975. A list of Syrphidae (Diptera) from Korea. Korean Journal of Entomology 5(1): 38-42. [in Korean] Kim, J.-I. 1980. The historical review and the tentative list of the Korean Syrphidae. Journal of Sungshin University 13: 365-389. [in Korean] Kim, J.-I. 1981. The faunistic study on the insects from Sudong-myeon, Namyangju-gun, Gyeonggi-do, Korea. Bulletin of the Korean Association for Conservation of Nature (3): 329-367. [in Korean] Kim, J.-I. and Kim, C. W. 1972. On the 17 unrecorded Syrphidae (Dipt.) from Korea. Korean Journal of Entomology 2(1): 7-12. [in Korean] Kim, J.-I. and Park, H.-K. 1984. Taxonomical study on the Syrphidae (Diptera) from the anatomical characters of the digestive organ. Journal of Basic Science (Sungshin Women's University) 1: 41-49. [in Korean] Kim, J.-I., Park, H.-K. and Kim, C.-W. 1983. On the 9 unrecorded Syrphidae (Insecta, Diptera) from Korea. Korean J. of Zoology 26(3): 203-209. [in Korean] Kimura, T. 1986. A new species of Eoseristalis (Diptera, Syrphidae) from Japan. Kontyu 54: 124-127. Kimura, T. 1998. New synonym of Chalcosyrphus (Xylotomima) frontalis (Shiraki et Edashige, 1953). Hana Abu 5: 3. [in Japanese] Knutson, L. V., Thompson, F. C. and Vockeroth, J. R. 1975. Family Syrphidae, pp. 307-374. In: Delfinado M.D., Hardy D.E. (eds.), A Catalog of the Diptera of the Oriental Region (Volume 2). The University Press of Hawaii. Korean Society of Applied Entomology (KSAE) and The Enromological Society of Korea (ESK). 1998. English-Korean, Korean-English Entomological Terminology. Korean Society of Applied Entomology. 366 pp.

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Rotheray, G. E. 2003. The predatory larvae of two Nepenthosyrphus species living in pitcher plants (Diptera, Syrphidae). Stud. Dipt. 10, 219–226. Sack, P. 31. Syrphidae. In Lindner, E. [ed], Die Fliegen der paläarktischen Region. Band IV/6: 1-451, Taf. I-XVIII. Shiraki, T. 1930. Die Syrphiden des Japanischen Kaiserreichs, mit Berucksichtigung benachbarter Gebiete. Mem. Fac. Agric. Taihoku imp. Univ. 1(1): 1-446. Shiraki, T. 1968a. Fauna Japonica, Syrphidae (Insecta). Vol. II. Biogeograph. soc. Japan, Tokyo. [x] + 243 pp., 40 pls. Shiraki, T. 1968b. Fauna Japonica, Syrphidae (Insecta). Vol. III. Biogeograph. soc. Japan, Tokyo. [x] +272 pp., 47 pls. Ssymank, A., Kearns, C. A., Pape, T. & Thompson, F. C. 2008. Pollinating flies (Diptera): A major contribution to plant diversity and agricultural production. Biodiversity 9(1–2): 86–89. Suk, S.-W. and Han, H.-Y. 2014a. Three Species of Syrphidae (Diptera) Recorded by the Korean Indigenous Species Survey of the National Institute of Biological Resources (2011-2013). Entomological Research Bulletin, 30(2): 180-181. Suk, S.-W. and Han, H.-Y. 2014b. Tropidia rostrata (Diptera, Syrphidae), first recorded genus and species in Korea. Animal Systematics, Evolution and Diversity 30(3): 230-234. Suk, S.-W., Choi, D.-S. and Han, H.-Y. 2015. Five species of Syrphidae (Insecta: Diptera) newly recorded in Korea. Animal Systematics, Evolution and Diversity. 31(4): 257-265. Thompson, F. C. 1999. A key to the genera of the flower flies (Diptera: Syrphidae) of the Neotropical Region including descriptions of new genera and species and a glossary of taxonomic terms. Contributions on Entomology, International, 3, 321–378. Thompson, F. C., Rotheray, G. E. 1998. Family Syrphidae. In: Papp, L., Darvas, B. (Eds.), Manual of Palaearctic Diptera, Vol. 3. Science Herald, Budapest, pp. 81–139. Torp, E. 1994. Danmarks svirrefluer (Diptera: Syrphidae). Danmarks Dyreliv, Bind 6. Appollo Books, Stenstrup. 490 pp. Vockeroth, J. R. and Thompson, F. C. 1987. Syrphidae. Vol. 2, pages 713-743, in McAlpine, J. F. et al. (eds): Manual of Nearctic Diptera. Research Branch, Agriculture Canada, Ottawa. Agric. Can. Monograph No. 28, vi + 675-1332 pp. Young, A. D., Lemmon, A. R., Skevington, J. H., Mengual, X., Ståhls, G., Reemer, M., Jordaens, K., Kelso, S., Lemmon, E. M., Hauser, M., De Meyer, M., Misof, B. and Wiegmann, B. M. 2016. BMC Evolutionary Biology 16(143): 1-13. Zoological Society of Korea (ZSK). 1968. Syrphidae. pp. 178-180 in Nomina Animalium Koreanorum, (2) Insecta. [in Korean]

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PLATES

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A B C

D E F

G H I

J K L

Plate 1. A. Callicera aenea ♂, 14.1 mm; B. Monoceromyia pleuralis ♂, 17.8 mm (from Han and Choi, 2001); C. Primocerioides petri ♂, 15.2 mm; D. Sphiximorpha rachmaninovi ♂, 15.1 mm (from Suk et al., 2015); E. ditto ♀, 13.4 mm (from Suk et al., 2015); F. Cheilosia abbreviata ♂, 9.2 mm; G. ditto ♀, 10.1 mm; H. Cheilosia bombiformis ♂, 14.1 mm; I. ditto ♀, 14.8 mm; J. Cheilosia illustrata ♂, 12 mm; K. ditto ♀, 12.3 mm; L. Cheilosia longiptera ♂, 10.2 mm.

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곤충-알락긴꽃등에아과Ⅱ_영문 수정.indd 136 2018-09-12 오후 12:04:37 PLATES

A B C

D E F

G H I

J K

Plate 2. A. Cheilosia longiptera ♀, 10.5 mm; B. Cheilosia motodomariensis ♂, 10.2 mm; C. Ferdinandea cuprea ♂, 13.2 mm; D. Ferdinandea nigrifrons ♂, 11.8 mm; E. Ferdinandea ruficornis ♂, 12.2 mm; F. Rhingia laevigata ♂, 8.9 mm; G. Brachyopa testacea ♂, 7.1 mm; H. Orthonevra karumaiensis ♂, 5.6 mm; I. ditto ♀, 5.8 mm; J. Sphegina clunipes ♂, 6.2 mm; K. ditto ♀, 6.5 mm.

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A B C

D E F

G H I

J K L

Plate 3. A. Eristalinus aeneus ♂, 12.2 mm; B. ditto ♀, 11.8 mm; C. Eristalinus tarsalis ♂, 11.1 mm; D. ditto ♀, 10.9 mm; E. Eristalinus viridis ♂, 11.2 mm; F. ditto ♀, 11.2 mm; G. Eristalis arbustorum ♂, 10.8 mm (from Han and Choi, 2001); H. ditto ♀, 11.5 mm (from Han and Choi, 2001); I. Eristalis cerealis ♂, 11.3 mm (from Han and Choi, 2001); J. ditto ♀, 11.6 mm (from Han and Choi, 2001); K. Eristalis kyokoae ♂, 12.5 mm (from Han and Choi, 2001); L. ditto ♀, 12.8 mm (from Han and Choi, 2001).

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A B C

D E F

G H I

J K L

Plate 4. A. Eristalis tenax ♂, 15.2 mm (from Han and Choi, 2001); B. ditto ♂, 16.1 mm (from Han and Choi, 2001); C. ditto ♀, 15.8 mm (from Han and Choi, 2001); D. Helophilus lunulatus ♂, 7.8 mm (from Han and Choi, 2001); E. ditto ♀, 8.2 mm (from Han and Choi, 2001); F. Helophilus trivittatus ♂, 15.6 mm; G. Helophilus virgatus ♂, 14.5 mm (from Han and Choi, 2001); H. ditto ♂, 14.8 mm (from Han and Choi, 2001); I. ditto ♀, 15.5 mm (from Han and Choi, 2001); J. Mallota ambigua ♂, 18.3 mm; K. ditto ♀, 19.5 mm; L. Mallota analis ♂, 19.2 mm.

139

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A B C

D E F

G H I

J K L

Plate 5. A. Mallota dimorpha ♂, 19.2 mm; B. ditto ♀, 19.8 mm; C. Mallota rossica ♂, 17.5 mm (from Suk et al., 2015); D. ditto ♀, 17.6 mm (from Suk et al., 2015); E. Mallota shatalkini ♂, 14.5 mm (from Suk et al., 2015); F. ditto ♀, 13.6 mm (from Suk et al., 2015); G. Mallota tricolor ♀, 15.5 mm; H. Mesembrius flaviceps ♂, 10.8 mm; I. ditto ♀, 11.5 mm (from Han and Choi, 2001); J. Phytomia zonata ♂, 16.2 mm (from Han and Choi, 2001); K. Eumerus iidai ♂, 6.8mm; L. Eumerus japonicus ♂, 5.8 mm.

140

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A B C

D E F

G H I

J K L

Plate 6. A. Eumerus strigatus ♂, 7.2 mm; B. Merodon kawamurai ♂, 11.8 mm; C. Blera japonica ♂, 13.9 mm; D. ditto ♀, 12.8 mm; E. Macrozelima hervei ♂, 19.5 mm; F. ditto ♀, 21.2 mm; G. Matsumyia nigrofacies ♂, 17.8 mm (from Han and Choi, 2001); H. Milesia undulata ♂, 21.5 mm (from Han and Choi, 2001); I. ditto ♀, 22.4 mm; J. Pterallastes unicolor ♂, 16.2 mm (from Han and Choi, 2001); K. Spilomyia suzukii ♂, 18.2 mm; L. ditto ♀, 20.1 mm.

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A B C

D E F

G H I

J K L

Plate 7. A. Syritta pipiens ♂, 7.8 mm; B. ditto ♀, 8.2 mm; C. Temnostoma bombylans ♂, 15.2 mm (from Han and Choi, 2001); D. ditto ♀, 15.8 mm (from Han and Choi, 2001); E. Temnostoma vespiforme ♂, 17.2 mm; F. ditto ♀, 18.1 mm; G. Tropidia rostrata ♂, 10.2 mm (from Suk and Han, 2014b); H. ditto ♀, 10.8 mm (from Suk and Han, 2014b); I. Pipiza austriaca ♂, 10.2 mm (from Han and Choi, 2001); J. Pipiza flavimaculata ♂, 8.5 mm (from Han and Choi, 2001); K. Pipiza signata ♂, 8.9 mm (from Han and Choi, 2001); L. Triglyphus primus ♂, 7.8 mm.

142

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A B C

D E F

G H I

J K L

Plate 8. A. Pseudovolucella decipiens ♂, 16.2 mm (from Han and Choi, 2001); B. ditto ♂, 16.8 mm (from Han and Choi, 2001); C. Sericomyia dux ♂, 19.2 mm; D. Graptomyza ihai ♂, 4.9 mm; E. Volucella bivitta ♂, 17.2 mm (from Suk et al., 2015); F. ditto ♀, 17.8 mm (from Suk et al., 2015); G. Volucella coreana ♀, 23.6 mm; H. Volucella inanoides ♂, 16.5 mm (from Suk et al., 2015); I. ditto ♀, 17.6 mm (from Suk et al., 2015); J. Volucella jeddona ♂, 17.5 mm; K. ditto ♀, 18.2 mm; L. Volucella linearis ♂, 18.2 mm.

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A B C

D E F

G H I

J K L

Plate 9. A. Volucella linearis ♀, 19.1 mm; B. Volucella nigricans ♂, 17.6 mm (from Choi et al., 2006); C. ditto ♀, 17.8 mm (from Choi et al., 2006); D. Volucella nigropicta ♂, 18.8 mm; E. ditto ♀, 19.4 mm; F. Volucella pellucens tabanoides ♂, 17.3 mm; G. ditto ♂, 17.5 mm (from Choi et al., 2006); H. ditto ♀, 18.5 mm (from Choi et al., 2006); I. Volucella suzukii ♂, 12.7 mm; J. Volucella thompsoni ♂, 12.5 mm (from Choi et al., 2006); K. ditto ♀, 13.4 mm (from Choi et al., 2006); L. Brachypalpus nipponicus ♂, 13.5 mm.

144

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A B C

D E F

G H I

J K L

Plate 10. A. Chalcosyrphus carbonus ♂, 10.5 mm (from Jeong et al., 2017); B. Chalcosyrphus femoratus ♂, 19.2 mm (from Jeong et al., 2017); C. Chalcosyrphus laterimaculatus ♂, 11.8 mm; D. ditto ♀, 12.1 mm; E. Chalcosyrphus nemorum ♂, 8.8 mm (from Jeong et al., 2017); F. ditto ♀, 8.5 mm; G. Chalcosyrphus nitidus ♂, 10.8 mm (from Jeong et al., 2017); H. ditto ♀, 11.2 mm; I. Chalcosyrphus piger ♂, 12.5 mm (from Jeong et al., 2017); J. Chalcosyrphus rufipes ♀, 14.1 mm (from Jeong et al., 2017); K. Chalcosyrphus unicus ♀, 13.5 mm; L. Xylota coquilletti ♂, 11.2 mm (from Han and Choi, 2001).

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INDEX TO SCIENTIFIC NAMES

A aterrima 23 bombiformis 24 Arthropoda 14 facialis 25 fuscipennis 25 B illustrata 25 longiptera 26 Blera 74 matsumurana 27 japonica 74 motodomariensis 28 nitens 76 ochripes 29 Brachycera 14 Cheilosiini 21 Brachyopa 34 Chrysogasterini 34 testacea 34 Criorhina 76 Brachypalpus 114 brevipila 76 nipponicus 114 trifasciata 77

C D

Callicera 17 Diptera 14 aenea 17 Callicerini 17 E Ceriodini 18 Chalcosyrphus 115 Eristalini 38 carbonus 117 Eristalinus 38 femoratus 118 aeneus 39 laterimaculatus 120 quinquestriatus 40 longus 121 sepulchralis 40 nemorum 122 tarsalis 41 nitidus 123 viridis 42 piger 124 Eristalis 43 rufipes 125 arbustorum 44 unicus 126 cerealis 46 Cheilosia 22 japonica 48 abbreviata 22 kyokoae 48

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tenax 49 Mallota 56 Eumerini 67 ambigua 56 Eumerus 67 analis 57 djakonovi 68 dimorpha 58 iidai 68 eurasiatica 60 japonicus 69 koreana 60 kongosanensis 70 rossica 60 strigatus 70 shatalkini 61 takasagoensis 62 F tricolor 62 Matsumyia 78 Ferdinandea 29 jesoensis 78 cuprea 30 nigrofacies 79 nigrifrons 31 Merodon 71 ruficornis 32 equestris 72 G kawamurai 72 Mesembrius 64 Graptomyza 100 flaviceps 64 ihai 100 niger 65 Milesia 80 H undulata 80 Milesiinae 14 Helophilus 51 Milesiini 74 lunulatus 52 Monoceromyia 18 sapporensis 53 pleuralis 18 trivittatus 53 virgatus 54 N

I Narumyia 81 narumii 81 Insecta 14 O M Orthonevra 35 Macrozelima 77 karumaiensis 35 hervei 77

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P clunipes 36 japonica 38 Phytomia 65 Sphiximorpha 20 zonata 65 rachmaninovi 20 Pipiza 91 Spilomyia 83 abdominalis 91 permagna 83 austriaca 92 suzukii 83 flavimaculata 93 Stenopipiza 95 inornata 94 bipunctata 95 signata 94 Syritta 84 Pipizella 95 orientalis 84 biglumis 95 pipiens 85 Pipizini 91 Syrphidae 14 Primocerioides 19 petri 19 T Pseuderistalis 67 bicolor 67 Temnostoma 86 Pseudovolucella 97 bombylans 87 decipiens 97 frequense 88 Psilota 73 vespiforme 88 brevicornis 73 Triglyphus 96 Pterallastes 81 primus 96 unicolor 81 Tropidia 90 rostrata 90 R V Rhingia 33 laevigata 33 Volucella 101 bivitta 103 S coreana 104 inanoides 105 Sericomyia 98 jeddona 106 dux 99 linearis 107 sachalinica 99 nigricans 108 Sericomyini 97 nigropicta 109 Sphegina 36 pellucens tabanoides 110

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suzukii 112 thompsoni 113 Volucellini 100

X

Xylota 128 abiens 128 coquilletti 129 ignava 130 Xylotini 114

149

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