145 a New Species of Dixonius (Sauria: Gekkonidae
Total Page:16
File Type:pdf, Size:1020Kb
Load more
Recommended publications
-
Cyrtodactylus Saiyok Sp. Nov., a New Dry Evergreen Forest-Dwelling Bent-Toed Gecko (Squamata: Gekkonidae) from Kanchanaburi Province, Western Thailand
Zootaxa 3869 (1): 064–074 ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2014 Magnolia Press ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.3869.1.6 http://zoobank.org/urn:lsid:zoobank.org:pub:0334116F-5288-4F5E-9B9B-FD5D71FFB179 Cyrtodactylus saiyok sp. nov., a new dry evergreen forest-dwelling Bent-toed Gecko (Squamata: Gekkonidae) from Kanchanaburi Province, western Thailand NONN PANITVONG1, MONTRI SUMONTHA2, JITTHEP TUNPRASERT3 & OLIVIER S. G. PAUWELS4,5 1siamensis.org, 408/144 Phaholyothin Place Bldg 34FL, Phaholyothin Rd., Phayathai, Bangkok 10400, Thailand. E-mail: [email protected] 2Ranong Marine Fisheries Station, 157 Saphanpla Rd., Paknam, Muang, Ranong 85000, Thailand. E-mail: [email protected] 3Institute for Population and Social Research, Mahidol University, Sai Yok, Kanchanaburi 71150, Thailand 4Département des Vertébrés Récents, Institut Royal des Sciences naturelles de Belgique, Rue Vautier 29, B-1000 Brussels, Belgium 5Corresponding author. E-mail: [email protected] Abstract We describe Cyrtodactylus saiyok sp. nov. from a dry evergreen forest on a limestone hill in Khao Krajae, Sai Yok District, Kanchanaburi Province, western Thailand. It is characterized by a maximal known SVL of 61.0 mm; 18–19 longitudinal rows of dorsal tubercles; 23 or 24 ventral scale rows between ventrolateral skin folds; a continuous series of enlarged fem- oro-precloacal scales, including 5 pore-bearing precloacal scales (males); no precloacal groove or depression; transversely enlarged subcaudal scales; a complete black nuchal loop; a W-shaped band above shoulders and 3–5 irregular, medially interrupted or not, black dorsal bands between limb insertions. Cyrtodactylus saiyok sp. -
Cryptic Taxonomic Diversity in Two Broadly Distributed Lizards Of
The Natural History Journal of Chulalongkorn University 1(1): 1-7, August 2001 ©2001 by Chulalongkorn University Cryptic Taxonomic Diversity in Two Broadly Distributed Lizards of Thailand (Mabuya macularia and Dixonius siamensis) as Revealed by Chromosomal Investigations (Reptilia: Lacertilia) HIDETOSHI OTA1*, TSUTOMU HIKIDA2, JARUJIN NABHITABHATA3 AND SOMSAK PANHA4 1 Tropical Biosphere Research Center, University of the Ryukyus, Nishihara, Okinawa 903-0213, JAPAN 2 Department of Zoology, Graduate School of Science, Kyoto University, Kitashirakawa, Sakyo, Kyoto 606-8502, JAPAN 3 Reference Collection Division, National Science Museum, Klong Luang, Pathumthani 12120, THAILAND 4 Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, THAILAND ABSTRACT.−Karyological surveys were made for two common lizards, Mabuya macularia and Dixonius siamensis, in Thailand. Of the M. macularia specimens karyotyped, the one from Mae Yom, northwestern Thailand, had 2n=38 chromosomes in a graded series as previously reported for a specimen from southwestern Thailand. The other specimens, all from eastern Thailand, however, invariably exhibited a 2n=34 karyotype consisting of four large, and 13 distinctly smaller chromosome pairs. Of the D. siamensis specimens, on the other hand, those from eastern Thailand invariably exhibited a 2n=40 karyotype, whereas three males and one female from Mae Yom had 2n=42 chromosomes. Distinct chromosomal variation was also observed within the Mae Yom sample that may reflect a ZW sex chromosome system. These results strongly suggest the presence of more than one biological species within each of those “broadly distributed species”. This further argues the necessity of genetic surveys for many other morphologically defined “common species” in Thailand and its vicinity in order to evaluate taxonomic diversity and endemicity of herpetofauna in this region appropriately. -
Hue University College of Education Do Trong Dang
HUE UNIVERSITY COLLEGE OF EDUCATION DO TRONG DANG RESEARCH ON THE SPECIES DIVERSITY, DISTRIBUTION AND CONSERVATION VALUE OF AMPHIBIANS AND REPTILES SPECIES IN SOUTH PART OF CU MONG PASS, PHU YEN PROVINCE Major: Zoology Code: 62 42 01 03 SUMMARY OF Ph.D IN BIOLOGY Instructors PhD. Nguyen Quang Truong Prof. PhD. Ngo Dac Chung HUE, 2017 The work was completed in: The work was completed in: College of Education, Hue University Science instructors: PhD. Nguyen Quang Truong Prof. PhD. Ngo Dac Chung Reviewer 1: Reviewer 2: Reviewer 3: The thesis was defended at the Council of thesis assessment of Hue University Council held at: 4 Le Loi street, Hue city, Thua Thien Hue province, at ……………………………………... on .…../…../2017 Theses can be further referred at: 1. National Library 2. Center for Information and Library of College of Education, Hue University WORKS RELATED TO THE THESIS HAS BEEN PUBLISHED 1. Dang Trong Do, Chung Dac Ngo, Truong Quang Nguyen (2015), Diversity of the narrow-mouth frogs (Amphibia: Anura: Microhylidae) from Phu Yen Province, In Proceedings of the sixth National Scientific Conference on Ecology and Biological Resources, pp. 515-519. 2. Dang Trong Do, Chung Dac Ngo, Truong Quang Nguyen (2016), New records of turtles from Phu Yen Province, Vietnam, In Proceedings of the 2ND National Scientific Conference on Biologycal Research and Teaching in Vietnam, pp. 129-136. 3. Dang Trong Do, Chung Dac Ngo, Truong Quang Nguyen (2016), New records of Colubridae (Squamata: Serpentes) and an updated list of snakes from Phu Yen Province, Vietnam, In Proceedings of third National Scientific Conference on Amphibians and Reptiles in Vietnam, pp. -
A Biogeographic Synthesis of the Amphibians and Reptiles of Indochina
BAIN & HURLEY: AMPHIBIANS OF INDOCHINA & REPTILES & HURLEY: BAIN Scientific Publications of the American Museum of Natural History American Museum Novitates A BIOGEOGRAPHIC SYNTHESIS OF THE Bulletin of the American Museum of Natural History Anthropological Papers of the American Museum of Natural History AMPHIBIANS AND REPTILES OF INDOCHINA Publications Committee Robert S. Voss, Chair Board of Editors Jin Meng, Paleontology Lorenzo Prendini, Invertebrate Zoology RAOUL H. BAIN AND MARTHA M. HURLEY Robert S. Voss, Vertebrate Zoology Peter M. Whiteley, Anthropology Managing Editor Mary Knight Submission procedures can be found at http://research.amnh.org/scipubs All issues of Novitates and Bulletin are available on the web from http://digitallibrary.amnh.org/dspace Order printed copies from http://www.amnhshop.com or via standard mail from: American Museum of Natural History—Scientific Publications Central Park West at 79th Street New York, NY 10024 This paper meets the requirements of ANSI/NISO Z39.48-1992 (permanence of paper). AMNH 360 BULLETIN 2011 On the cover: Leptolalax sungi from Van Ban District, in northwestern Vietnam. Photo by Raoul H. Bain. BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY A BIOGEOGRAPHIC SYNTHESIS OF THE AMPHIBIANS AND REPTILES OF INDOCHINA RAOUL H. BAIN Division of Vertebrate Zoology (Herpetology) and Center for Biodiversity and Conservation, American Museum of Natural History Life Sciences Section Canadian Museum of Nature, Ottawa, ON Canada MARTHA M. HURLEY Center for Biodiversity and Conservation, American Museum of Natural History Global Wildlife Conservation, Austin, TX BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY Number 360, 138 pp., 9 figures, 13 tables Issued November 23, 2011 Copyright E American Museum of Natural History 2011 ISSN 0003-0090 CONTENTS Abstract......................................................... -
Phylogenetic Relationships of Geckos of the Genus Nactus and Their Relatives (Squamata: Gekkonidae)
Acta Herpetologica 3(1): 1-18, 2008 ISSN 1827-9643 (online) © 2008 Firenze University Press Phylogenetic relationships of geckos of the genus Nactus and their relatives (Squamata: Gekkonidae) Todd R. Jackman, Aaron M. Bauer, Eli Greenbaum Department of Biology, Villanova University, 800 Lancaster Ave., Villanova, PA 19085, USA. Corre- sponding author. E-mail: [email protected] Submitted on 2007, 16th June; revised on 2007, 27th November; accepted 2008, 29th January. Abstract. We employed nuclear and mitochondrial DNA sequence data to investigate relationships within the gekkonid genus Nactus and between Nactus and other gekkonid genera. Nuclear (RAG-1, PDC) and mitochondrial (ND2) data provide strong support for conflicting patterns of relationship among bisexual New Guinean species of Nac- tus and the unisexual oceanic form N. pelagicus. This may be explained by an ancient mitochondrial introgression event between N. sphaerodactylodes and N. vankampeni, a recent selective sweep of mitochondrial DNA throughout N. vankampeni, and gene conflict stemming from the hybrid event that gave rise to N. pelagicus. Strong support from all data partitions is obtained for the sister group relationship of Nactus to a clade consisting of the Australian Heteronotia and the Southeast Asian Dixonius. Putative synapomorphies of the Nactus/Heteronotia/Dixonius clade include the reduction of the second phalanx of digit IV of the manus and the presence of regular rows of keeled (sometimes multicarinate) dorsal tubercles on the dorsum. Nactus and Heteronotia both include parthenogenetic species formed via hybridogenesis. This is rare among geckos, and vertebrates in general, and at some level may also be synapomorphic. Dixonius is not known to have any all-female species, but “D. -
A New Species of Dixonius from Nui Chua National Park, Ninh Thuan Province, Southern Vietnam (Squamata, Gekkonidae)
Zoosyst. Evol. 85 (1) 2009, 117–125 / DOI 10.1002/zoos.200800018 A new species of Dixonius from Nui Chua National Park, Ninh Thuan Province, southern Vietnam (Squamata, Gekkonidae) Ngo Tri Van1 and Thomas Ziegler*,2 1 Vietnamese Academy for Science and Technology (VAST), Department of Environmental Management and Technology, Institute of Tropical Biology, 85 Tran Quoc Toan Str, Dist. 3, Hochiminh City, Vietnam 2 Cologne Zoo, Riehler Str. 173, 50735 Kln, Germany Abstract Received 4 February 2008 A new species of the geckonid genus Dixonius is described from Nui Chua National Park, Ninh Accepted 29 May 2008 Thuan Province in southern Vietnam. The description is based on four adult specimens, two Published 17 March 2009 males and females each. The new species is distinguished from any other Dixonius by the com- bination of the following characters: maximum SVL 38.6 mm; bold dark canthal stripe terminat- ing at back of head (occiput); more or less dark barred lips; patternless yellowish-orange back Key Words and tail; supranasals in distinct contact; 8–9 supralabials, with supralabials six or seven in mid- orbital position; 18–19 ventral scale rows at midbody; males with 5 precloacal pores. We further Dixonius aaronbaueri sp. n. provide the first record of D. cf. vietnamensis for Ninh Thuan Province, which occurs in sympa- try with the new species in coastal Nui Chua National Park. We provide first data on the natural morphology history of the new species and a key to all five known Dixonius species. taxonomy distribution Introduction gated whether D. melanostictus is continuously distri- buted in the intervening areas or whether the Vietnamese The genus Dixonius was established by Bauer et al. -
New Blood: Greater Mekong New Species Discoveries 2009 1 Flooded Forest, Kratie-Stung Treng, Cambodia, in the Mekong River Ecoregion
REPORT New Blood Greater Mekong New Species Discoveries 2009 DRACULA FISH (DANIONELLA DRACULA) Britz, Conway & Rüber, 2009 Myanmar Written by Nancy Gephart, Geoffrey Blate, Colin McQuistan (WWF) and Christian Thompson (the green room) www.greenroomenvironmental.com Designed by Torva Thompson (the green room) Front cover photo: Danionella dracula, dracula minnow species © Natural History Museum, London © Text 2010 WWF WWF is one of the world’s largest and most experienced independent conservation organizations, with over 5 million supporters and a global Network active in more than 100 countries. WWF’s mission is to stop the degradation of the planet’s natural environment and to build a future in which humans live in harmony with nature, by: conserving the world’s biological diversity, ensuring that the use of renewable natural resources is sustainable, and promoting the reduction of pollution and wasteful consumption. A fanged fish and a fangless snake, five new mammal species, a bald bird and a EXECUTIVE frog that sounds like a cricket are among the 145 species newly described by science SUMMARY in the Greater Mekong region during 2009. The Greater Mekong region of Southeast Asia through which the Mekong River flows comprises the countries of Cambodia, Laos, Myanmar, Thailand, Vietnam and Yunnan Province in southern China. The region is home to some of the planet’s most endangered wild species including tiger, Asian elephant, Mekong dolphin and Mekong giant catfish, in addition to hundreds of newly discovered species. Between 1997 and 2008 an incredible 1,231 species were discovered by science across this region alone1,2. While these discoveries reinforce the conclusion that the Greater Mekong is a biodiversity hotspot, with some being considered for IUCN Red List status, they also highlight the fragility of this region’s diverse species and habitats. -
Malayemys Subtrijuga ) ]4 QJP9 (Cuora Amboinensis Kamaroma ) ^DO]4 Q9U&JPF]JDVL& (Hireremys Annandalii) 3
54 )8S3!L&IP4F]DVLA#DQ8`8=V86T:Q]!QJS8:X 5 9 L?BW79 'P&JFPB9WB T SPECIES OF REPTILE IN LIMESTONE FOREST AND RELIGIOUS TERRITORY, KHAOWONG SUBDISTRICT, PHRA PHUTTHABAT DISTRICT, SARABURI PROVINCE B * 1/* TBW7 'S83Q1/ Kanokorn konlek1/* Virayuth Lauhachinda1/ ABSTRACT Reptile species in limestone forest and religious territory, Khaowong Subdistrict, Phra Phutthabat District, Saraburi Province, was carried out monthly during the period of September 2007 to September 2008 by using direct searching method. The study found 50 species, 11 families, 2 orders which were classified into 7 groups, consisting of residing in forest area, residing inside cave, residing around cave vicinity, residing on tree, residing in water, residing in various construction, and residing in agricultural area including urban vicinity. The comparison of reptile species similarity by using Sorensen’s Index show that Lublae and Subchaom were the most similar whereas Thamnarai and Lublae were the most difference. For conservation status, elongated tortoise is categorized as endangered species whereas Malaysian snail-eating turtle, Malayan box turtle, and Yellow-headed temple turtle are categorized as vulnerable species. In addition, Dark-sided Leaf-toed gecko, Chanhome’s gecko, Siamese gecko, are endemic species of Thailand. The study revealed that forest destruction, cave development for religious ceremony, and utilization of limestone mountain area are the threatened factors for reptile species in the study areas. 9P3A QBGU HQ#FQ@JDQ )8S3!L&IP4F]DVLA#DQ8`8=V86T:Q]!QJS8:X8 -
(Squamata: Gekkonidae) from Phu Quy Island, Vietnam
Zootaxa 4040 (1): 048–058 ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2015 Magnolia Press ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4040.1.4 http://zoobank.org/urn:lsid:zoobank.org:pub:59444939-F3C3-4648-80B0-D9AEABBD2823 A new species of Dixonius (Squamata: Gekkonidae) from Phu Quy Island, Vietnam ANDREAS BOTOV1, TRUNG MY PHUNG2, TRUONG QUANG NGUYEN3, AARON M. BAUER4, IAN G. BRENNAN4,5 & THOMAS ZIEGLER1,6,7 1AG Zoologischer Garten Köln, Riehler Straße 173, D–50735 Köln, Germany. E-mail: [email protected] and [email protected] 2Dong Khoi 9A, Tam Hiep, Bien Hoa, Dong Nai Province, Vietnam. E-mail: [email protected] 3Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam. E–mail: [email protected] 4Department of Biology, Villanova University, 800 Lancaster Avenue, Villanova, Pennsylvania 19085, USA. E-mail: [email protected] 5current address: Evolution, Ecology, and Genetics, Research School of Biology, The Australian National University, Canberra, ACT, Australia 6Zoological Institute, University of Cologne, Zülpicher Strasse 47b, D–50674 Cologne, Germany 7Corresponding author Abstract We describe a new species of Dixonius on the basis of five specimens from Phu Quy Island, Binh Thuan Province, in southern Vietnam. The new species can be distinguished from congeners based on molecular and morphological differ- ences. Diagnostic features are: small size (SVL up to 44 mm); 7 or 8 supralabials; 11 or 12 rows of keeled tubercles on dorsum; 21–23 ventral scale rows; 5 or 6 precloacal pores in males; a canthal stripe running from rostrum through the eye and terminating behind the head; second pair of postmentals about one third to one half size of first pair; ground color of dorsum brown, with one or two rows of light yellow or orange spots in one or two rows along flanks, and irregular bands or a reticulated network of dark marks on dorsum. -
Phyllodactylus Wirshingi)
Original Article Cytogenet Genome Res Published online: January 26, 2019 DOI: 10.1159/000496379 ZZ/ZW Sex Chromosomes in the Endemic Puerto Rican Leaf-Toed Gecko ( Phyllodactylus wirshingi ) a c a a, b, d Stuart V. Nielsen Juan D. Daza Brendan J. Pinto Tony Gamble a b Department of Biological Sciences, Marquette University, and Milwaukee Public Museum, Milwaukee, WI , c d Department of Biological Sciences, Sam Houston State University, Huntsville, TX , and Bell Museum of Natural History, University of Minnesota, Saint Paul, MN , USA Keywords ed sex chromosomes – but can also be used to identify which Gekkota · Lizard · Phyllodactylidae · RADseq · Reptile · chromosomes in the genome are the sex chromosomes. We Squamata here identify a ZZ/ZW sex chromosome system in P. wir- shingi . Furthermore, we show that 4 of the female-specific markers contain fragments of genes found on the avian Z Abstract and discuss homology with P. wirshingi sex chromosomes. Investigating the evolutionary processes influencing the or- © 2019 S. Karger AG, Basel igin, evolution, and turnover of vertebrate sex chromosomes requires the classification of sex chromosome systems in a great diversity of species. Among amniotes, squamates (liz- Investigating the number and directionality of transi- ards and snakes) – and gecko lizards in particular – are wor- tions among sex-determining systems is a vital prerequi- thy of additional study. Geckos possess all major vertebrate site for studying sex chromosome evolution. This in- sex-determining systems, as well as multiple transitions volves not only determining whether a species has a het- among them, yet we still lack data on the sex-determining erogametic male (XX/XY) or female (ZZ/ZW) sex systems for the vast majority of species. -
A New Species of Dixonius (Squamata: Gekkonidae) from Phu Quy Island, Vietnam
Zootaxa 4040 (1): 048–058 ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2015 Magnolia Press ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4040.1.4 http://zoobank.org/urn:lsid:zoobank.org:pub:59444939-F3C3-4648-80B0-D9AEABBD2823 A new species of Dixonius (Squamata: Gekkonidae) from Phu Quy Island, Vietnam ANDREAS BOTOV1, TRUNG MY PHUNG2, TRUONG QUANG NGUYEN3, AARON M. BAUER4, IAN G. BRENNAN4,5 & THOMAS ZIEGLER1,6,7 1AG Zoologischer Garten Köln, Riehler Straße 173, D–50735 Köln, Germany. E-mail: [email protected] and [email protected] 2Dong Khoi 9A, Tam Hiep, Bien Hoa, Dong Nai Province, Vietnam. E-mail: [email protected] 3Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam. E–mail: [email protected] 4Department of Biology, Villanova University, 800 Lancaster Avenue, Villanova, Pennsylvania 19085, USA. E-mail: [email protected] 5current address: Evolution, Ecology, and Genetics, Research School of Biology, The Australian National University, Canberra, ACT, Australia 6Zoological Institute, University of Cologne, Zülpicher Strasse 47b, D–50674 Cologne, Germany 7Corresponding author Abstract We describe a new species of Dixonius on the basis of five specimens from Phu Quy Island, Binh Thuan Province, in southern Vietnam. The new species can be distinguished from congeners based on molecular and morphological differ- ences. Diagnostic features are: small size (SVL up to 44 mm); 7 or 8 supralabials; 11 or 12 rows of keeled tubercles on dorsum; 21–23 ventral scale rows; 5 or 6 precloacal pores in males; a canthal stripe running from rostrum through the eye and terminating behind the head; second pair of postmentals about one third to one half size of first pair; ground color of dorsum brown, with one or two rows of light yellow or orange spots in one or two rows along flanks, and irregular bands or a reticulated network of dark marks on dorsum. -
Nghiên Cứu Đa Dạng Loài, Đặc Điểm Phân Bố Và Giá Trị Bảo Tồn Của Khu Hệ Lƣỡng Cƣ Và Bò Sát Ở Vùng Phía Nam Đèo Cù Mông, Tỉnh Phú Yên
ĐẠI HỌC HUẾ TRƢ ỜNG ĐẠI HỌC SƢ PHẠM ĐỖ TRỌNG ĐĂNG NGHIÊN CỨU ĐA DẠNG LOÀI, ĐẶC ĐIỂM PHÂN BỐ VÀ GIÁ TRỊ BẢO TỒN CỦA KHU HỆ LƢỠNG CƢ VÀ BÒ SÁT Ở VÙNG PHÍA NAM ĐÈO CÙ MÔNG, TỈNH PHÚ YÊN LUẬN ÁN TIẾN SĨ SINH HỌC HUẾ - 2017 ĐẠI HỌC HUẾ TRƢ ỜNG ĐẠI HỌC SƢ PHẠM ĐỖ TRỌNG ĐĂNG NGHIÊN CỨU ĐA DẠNG LOÀI, ĐẶC ĐIỂM PHÂN BỐ VÀ GIÁ TRỊ BẢO TỒN CỦA KHU HỆ LƢỠNG CƢ VÀ BÒ SÁT Ở VÙNG PHÍA NAM ĐÈO CÙ MÔNG, TỈNH PHÚ YÊN Chuyên ngành: Động vật học Mã số: 62 42 01 03 LUẬN ÁN TIẾN SĨ SINH HỌC Ngƣời hƣớng dẫn khoa học TS. NGUYỄN QUẢNG TRƢỜNG GS. TS. NGÔ ĐẮC CHỨNG HUẾ - 2017 LỜI CAM ĐOAN Tôi xin cam đoan đây là công trình nghiên cứu của riêng tôi. Các kết quả nêu trong luận án hoàn toàn trung thực và chƣa từng đƣợc bảo vệ trƣớc bất kỳ hội đồng để nhận học vị nào trƣớc đây. Tác giả Đỗ Trọng Đăng LỜI CẢM ƠN Để hoàn thành luận án này, trƣớc tiên tôi xin bày tỏ lòng biết ơn sâu sắc đến TS. Nguyễn Quảng Trƣờng và GS.TS. Ngô Đắc Chứng, những ngƣời thầy đã tận tình hƣớng dẫn và giúp đỡ tôi từ khâu định hƣớng nghiên cứu, phân tích số liệu, công bố các công trình khoa học và hoàn thiện luận án. Xin trân trọng cám ơn các thầy cô trong Bộ môn Động vật học và Khoa Sinh học, Trƣờng Đại học Sƣ phạm Huế; Phòng Động vật có xƣơng sống, Viện Sinh thái và Tài nguyên Sinh vật đã giúp đỡ và trao đổi những kinh nghiệm quý báu trong nghiên cứu.