Crab Predators Are More Important at Higher Latitudes
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Universidad Austral De Chile Facultad De Ciencias Escuela De Biología Marina
Universidad Austral de Chile Facultad de Ciencias Escuela de Biología Marina Profesor Patrocinante: Dr. Dirk Schories. Instituto de Ciencias Marinas y Limnológicas. Facultad de Ciencias – Universidad Austral de Chile. Profesor Co-patrocinante: Dr. Luis M. Pardo. Instituto de Ciencias Marinas y Limnológicas. Facultad de Ciencias – Universidad Austral de Chile. ECOLOGÍA TRÓFICA DEL ASTEROIDEO Cosmasterias lurida (Phillipi, 1858) EN EL SENO DEL RELONCAVÍ (SUR DE CHILE): DISTRIBUCIÓN, ABUNDANCIA, ALIMENTACIÓN Y MOVIMIENTO. Tesis de Grado presentada como parte de los requisitos para optar al grado de Licenciado en Biología Marina y Título Profesional de Biólogo Marino. IGNACIO ANDRÉS GARRIDO IRIONDO VALDIVIA - CHILE 2012. AGRADECIMIENTOS Primero que todo, me siento extremadamente afortunado gracias a tanta gente maravillosa que en estos 25 años se ha cruzado por mi camino. Quiero agradecer especialmente a todos los que aportaron de alguna forma en mi formación como Biólogo Marino: A mi núcleo familiar, Margarita I., Dagoberto G. y Augusto G. (también Gorlak y Ulises) que con sus consejos y apoyo incondicional logre cumplir este sueño que tanto anhelaba. Gracias por todo el cariño y por creer en mí, esto se los dedico a ustedes. Al Dr. Dirk Schories, amigo y profesor, quien me enseño a disfrutar y valorar lo que más admiro en la vida, la naturaleza y el infinito mundo submarino. Asimismo, quien me guió en mi formación como Biólogo Marino y con quien compartí incontables inmersiones fascinantes e inolvidables. Además fue quien financio esta tesis de pregrado. Espero podamos continuar trabajando en el futuro. ¡Muchas gracias por todo! Al Dr. Luis M. Pardo, quien con el tiempo se convirtió en un importante guía profesional y amigo. -
Impacts of Climate Change on the Occurrence of Harmful Algal Blooms
Office of Water EPA 820-S-13-001 MC 4304T May 2013 Impacts of Climate Change on the Occurrence of Harmful Algal Blooms Summary Background Climate change is predicted to change many Algae occur naturally in marine and fresh waters. environmental conditions that could affect the Under favorable conditions that include adequate natural properties of fresh and marine waters both in light availability, warm waters, and high nutrient the US and worldwide. Changes in these factors levels, algae can rapidly grow and multiply causing could favor the growth of harmful algal blooms and “blooms.” Blooms of algae can cause damage to habitat changes such that marine HABs can invade aquatic environments by blocking sunlight and and occur in freshwater. An increase in the depleting oxygen required by other aquatic occurrence and intensity of harmful algal blooms organisms, restricting their growth and survival. may negatively impact the environment, human Some species of algae, including golden and red health, and the economy for communities across the algae and certain types of cyanobacteria, can produce US and around the world. The purpose of this fact potent toxins that can cause adverse health effects to sheet is to provide climate change researchers and wildlife and humans, such as damage to the liver and decision–makers a summary of the potential impacts nervous system. When algal blooms impair aquatic of climate change on harmful algal blooms in ecosystems or have the potential to affect human freshwater and marine ecosystems. Although much health, they are known as harmful algal blooms of the evidence presented in this fact sheet suggests (HABs). -
Ocean Circulation and Climate: an Overview
ocean-climate.org Bertrand Delorme Ocean Circulation and Yassir Eddebbar and Climate: an Overview Ocean circulation plays a central role in regulating climate and supporting marine life by transporting heat, carbon, oxygen, and nutrients throughout the world’s ocean. As human-emitted greenhouse gases continue to accumulate in the atmosphere, the Meridional Overturning Circulation (MOC) plays an increasingly important role in sequestering anthropogenic heat and carbon into the deep ocean, thus modulating the course of climate change. Anthropogenic warming, in turn, can influence global ocean circulation through enhancing ocean stratification by warming and freshening the high latitude upper oceans, rendering it an integral part in understanding and predicting climate over the 21st century. The interactions between the MOC and climate are poorly understood and underscore the need for enhanced observations, improved process understanding, and proper model representation of ocean circulation on several spatial and temporal scales. The ocean is in perpetual motion. Through its DRIVING MECHANISMS transport of heat, carbon, plankton, nutrients, and oxygen around the world, ocean circulation regulates Global ocean circulation can be divided into global climate and maintains primary productivity and two major components: i) the fast, wind-driven, marine ecosystems, with widespread implications upper ocean circulation, and ii) the slow, deep for global fisheries, tourism, and the shipping ocean circulation. These two components act industry. Surface and subsurface currents, upwelling, simultaneously to drive the MOC, the movement of downwelling, surface and internal waves, mixing, seawater across basins and depths. eddies, convection, and several other forms of motion act jointly to shape the observed circulation As the name suggests, the wind-driven circulation is of the world’s ocean. -
Coastal Upwelling Revisited: Ekman, Bakun, and Improved 10.1029/2018JC014187 Upwelling Indices for the U.S
Journal of Geophysical Research: Oceans RESEARCH ARTICLE Coastal Upwelling Revisited: Ekman, Bakun, and Improved 10.1029/2018JC014187 Upwelling Indices for the U.S. West Coast Key Points: Michael G. Jacox1,2 , Christopher A. Edwards3 , Elliott L. Hazen1 , and Steven J. Bograd1 • New upwelling indices are presented – for the U.S. West Coast (31 47°N) to 1NOAA Southwest Fisheries Science Center, Monterey, CA, USA, 2NOAA Earth System Research Laboratory, Boulder, CO, address shortcomings in historical 3 indices USA, University of California, Santa Cruz, CA, USA • The Coastal Upwelling Transport Index (CUTI) estimates vertical volume transport (i.e., Abstract Coastal upwelling is responsible for thriving marine ecosystems and fisheries that are upwelling/downwelling) disproportionately productive relative to their surface area, particularly in the world’s major eastern • The Biologically Effective Upwelling ’ Transport Index (BEUTI) estimates boundary upwelling systems. Along oceanic eastern boundaries, equatorward wind stress and the Earth s vertical nitrate flux rotation combine to drive a near-surface layer of water offshore, a process called Ekman transport. Similarly, positive wind stress curl drives divergence in the surface Ekman layer and consequently upwelling from Supporting Information: below, a process known as Ekman suction. In both cases, displaced water is replaced by upwelling of relatively • Supporting Information S1 nutrient-rich water from below, which stimulates the growth of microscopic phytoplankton that form the base of the marine food web. Ekman theory is foundational and underlies the calculation of upwelling indices Correspondence to: such as the “Bakun Index” that are ubiquitous in eastern boundary upwelling system studies. While generally M. G. Jacox, fi [email protected] valuable rst-order descriptions, these indices and their underlying theory provide an incomplete picture of coastal upwelling. -
Habs in UPWELLING SYSTEMS
GEOHAB CORE RESEARCH PROJECT: HABs IN UPWELLING SYSTEMS 1 GEOHAB GLOBAL ECOLOGY AND OCEANOGRAPHY OF HARMFUL ALGAL BLOOMS GEOHAB CORE RESEARCH PROJECT: HABS IN UPWELLING SYSTEMS AN INTERNATIONAL PROGRAMME SPONSORED BY THE SCIENTIFIC COMMITTEE ON OCEANIC RESEARCH (SCOR) AND THE INTERGOVERNMENTAL OCEANOGRAPHIC COMMISSION (IOC) OF UNESCO EDITED BY: G. PITCHER, T. MOITA, V. TRAINER, R. KUDELA, P. FIGUEIRAS, T. PROBYN BASED ON CONTRIBUTIONS BY PARTICIPANTS OF THE GEOHAB OPEN SCIENCE MEETING ON HABS IN UPWELLING SYSTEMS AND THE GEOHAB SCIENTIFIC STEERING COMMITTEE February 2005 3 This report may be cited as: GEOHAB 2005. Global Ecology and Oceanography of Harmful Algal Blooms, GEOHAB Core Research Project: HABs in Upwelling Systems. G. Pitcher, T. Moita, V. Trainer, R. Kudela, P. Figueiras, T. Probyn (Eds.) IOC and SCOR, Paris and Baltimore. 82 pp. This document is GEOHAB Report #3. Copies may be obtained from: Edward R. Urban, Jr. Henrik Enevoldsen Executive Director, SCOR Programme Co-ordinator Department of Earth and Planetary Sciences IOC Science and Communication Centre on The Johns Hopkins University Harmful Algae Baltimore, MD 21218 U.S.A. Botanical Institute, University of Copenhagen Tel: +1-410-516-4070 Øster Farimagsgade 2D Fax: +1-410-516-4019 DK-1353 Copenhagen K, Denmark E-mail: [email protected] Tel: +45 33 13 44 46 Fax: +45 33 13 44 47 E-mail: [email protected] This report is also available on the web at: http://www.jhu.edu/scor/ http://ioc.unesco.org/hab ISSN 1538-182X Cover photos courtesy of: Vera Trainer Teresa Moita Grant Pitcher Copyright © 2005 IOC and SCOR. -
Upwelling As a Source of Nutrients for the Great Barrier Reef Ecosystems: a Solution to Darwin's Question?
Vol. 8: 257-269, 1982 MARINE ECOLOGY - PROGRESS SERIES Published May 28 Mar. Ecol. Prog. Ser. / I Upwelling as a Source of Nutrients for the Great Barrier Reef Ecosystems: A Solution to Darwin's Question? John C. Andrews and Patrick Gentien Australian Institute of Marine Science, Townsville 4810, Queensland, Australia ABSTRACT: The Great Barrier Reef shelf ecosystem is examined for nutrient enrichment from within the seasonal thermocline of the adjacent Coral Sea using moored current and temperature recorders and chemical data from a year of hydrology cruises at 3 to 5 wk intervals. The East Australian Current is found to pulsate in strength over the continental slope with a period near 90 d and to pump cold, saline, nutrient rich water up the slope to the shelf break. The nutrients are then pumped inshore in a bottom Ekman layer forced by periodic reversals in the longshore wind component. The period of this cycle is 12 to 25 d in summer (30 d year round average) and the bottom surges have an alternating onshore- offshore speed up to 10 cm S-'. Upwelling intrusions tend to be confined near the bottom and phytoplankton development quickly takes place inshore of the shelf break. There are return surface flows which preserve the mass budget and carry silicate rich Lagoon water offshore while nitrogen rich shelf break water is carried onshore. Upwelling intrusions penetrate across the entire zone of reefs, but rarely into the Lagoon. Nutrition is del~veredout of the shelf thermocline to the living coral of reefs by localised upwelling induced by the reefs. -
Life on the Coral Reef
Coral Reef Teacher’s Guide Life on the Coral Reef Life on the Coral Reef THE CORAL REEF ECOSYSTEM The muddy silt drifts out to sea, covering the nearby Coral reefs provide the basis for the most productive coral reefs. Some corals can remove the silt, but many shallow water ecosystem in the world. An ecosystem cannot. If the silt is not washed off within a short pe- is a group of living things, such as coral, algae and riod of time by the current, the polyps suffocate and fishes, along with their non-living environment, such die. Not only the rainforest is destroyed, but also the as rocks, water, and sand. Each influences the other, neighboring coral reef. and both are necessary for the successful maintenance of life. If one is thrown out of balance by either natural Reef Zones or human-made causes, then the survival of the other Coral reefs are not uniform, but are shaped by the is seriously threatened. forces of the sea and the structure of the sea floor into DID YOU KNOW? All of the Earth’s ecosystems are a series of different parts or reef zones. Understand- interrelated, forming a shell of life that covers the ing these zones is useful in understanding the ecol- entire planet – the biosphere. For instance, if too many ogy of coral reefs. Keep in mind that these zones can trees are cut down in the rainforest, soil from the for- blend gradually into one another, and that sometimes est is washed by rain into rivers that run to the ocean. -
OCN 201 El Nino
OCN 201 El Nino El Nino theme page http://www.pmel.noaa.gov/tao/elnino/nino-home-low.html Reports to the Nation http://www.pmel.noaa.gov/tao/elnino/report/el-nino-report.html This page has all the text and figures and also how to get the booklet 1 El Nino is a major reorganisation of the equatorial climate system that affects regions far from its point of origin in the western Equatorial Pacific Occurs roughly every 6 years around Xmas-time Onset recognised by climatic effects --warm surface waters -- collapse of fisheries -- heavy rains in Peru/Ecuador/central Pacific -- droughts in Indonesia -- change in typhoon tracks Is a good example of how the ocean and atmosphere interact 2 What phase do you think we are in now? A El Nino B La Nina C Normal D I don’t know! A = El Nino 2014 2015 Southern Oscillation Atmospheric pressure differential between Tahiti and Darwin Normally low pressure in Darwin, high in Tahiti Low pressure High pressure Normal El Nino El Nino high pressure in Darwin, low in Tahiti Change in pressure differential results in weakening of easterly equatorial winds 3 Normal conditions in the Equatorial Pacific Strong easterly winds: Pile up warm water in the western Pacific -- thermocline deep in western Pacific, shallow in eastern Pacific Winds drive equatorial upwelling How much higher do you think that sea level is in the western Pacific? A 10cm B 50 cm C 1 metres D 5 metres E More! About 40 cm 4 Satellite image of chlorophyll abundance As thermocline is shallow in eastern Pacific upwelling brings nutrients to surface waters along -
Description and Mechanisms of the Mid-Year Upwelling in the Southern Caribbean Sea from Remote Sensing and Local Data
Journal of Marine Science and Engineering Article Description and Mechanisms of the Mid-Year Upwelling in the Southern Caribbean Sea from Remote Sensing and Local Data Digna T. Rueda-Roa 1,* ID , Tal Ezer 2 ID and Frank E. Muller-Karger 1 ID 1 Institute for Marine Remote Sensing, University of South Florida, College of Marine Science, 140 7th Ave. S., St. Petersburg, FL 33701, USA; [email protected] 2 Old Dominion University, Center for Coastal Physical Oceanography, 4111 Monarch Way, Norfolk, VA 23508, USA; [email protected] * Correspondence: [email protected] Received: 6 February 2018; Accepted: 27 March 2018; Published: 5 April 2018 Abstract: The southern Caribbean Sea experiences strong coastal upwelling between December and April due to the seasonal strengthening of the trade winds. A second upwelling was recently detected in the southeastern Caribbean during June–August, when local coastal wind intensities weaken. Using synoptic satellite measurements and in situ data, this mid-year upwelling was characterized in terms of surface and subsurface temperature structures, and its mechanisms were explored. The mid-year upwelling lasts 6–9 weeks with satellite sea surface temperature (SST) ~1–2◦ C warmer than the primary upwelling. Three possible upwelling mechanisms were analyzed: cross-shore Ekman transport (csET) due to alongshore winds, wind curl (Ekman pumping/suction) due to wind spatial gradients, and dynamic uplift caused by variations in the strength/position of the Caribbean Current. These parameters were derived from satellite wind and altimeter observations. The principal and the mid-year upwelling were driven primarily by csET (78–86%). However, SST had similar or better correlations with the Ekman pumping/suction integrated up to 100 km offshore (WE100) than with csET, possibly due to its influence on the isopycnal depth of the source waters for the coastal upwelling. -
UNIVERSIDAD NACIONAL DE SAN AGUSTÍN DE AREQUIPA FACULTAD DE CIENCIAS BIOLÓGICAS ESCUELA PROFESIONAL DE BIOLOGÍA Riqueza Y
UNIVERSIDAD NACIONAL DE SAN AGUSTÍN DE AREQUIPA FACULTAD DE CIENCIAS BIOLÓGICAS ESCUELA PROFESIONAL DE BIOLOGÍA Riqueza y tipos de hábitat de equinodermos en la Región Arequipa al 2017 Tesis para optar el título profesional de Biólogo presentado por el Bachiller en Ciencias Biológicas: Michael Leopoldo Espinoza Roque Asesor: Blgo. Dr. Graciano Alberto Del Carpio Tejada AREQUIPA – PERÚ 2018 1 ____________________________________ Blgo. Dr. Graciano Alberto Del Carpio Tejada ASESOR 2 DEDICATORIA A la memoria de mi padre, Leopoldo Espinoza Ramos, por todo su cariño, comprensión y sacrificio, sé que estaría feliz al verme cumplir esta meta. 3 AGRADECIMIENTOS A la Universidad Nacional de San Agustín de Arequipa (UNSA INVESTIGA), por el soporte financiero, con recursos del canon de la UNSA, para que se realice el presente proyecto de investigación (Contrato de financiamiento N° 156 – 2016 – UNSA), y un especial agradecimiento al Blgo. Luis Alberto Ponce Soto por la paciencia y apoyo en el acompañamiento y monitoreo de mi proyecto. A mi asesor de tesis, Blgo. Dr. Graciano Alberto Del Carpio Tejada, por su apoyo incondicional durante el desarrollo del presente trabajo de investigación. A la Blga. Rosaura Gonzales Juárez, por su contribución al inicio de este proyecto y sus enseñanzas y consejos que han aportado en mi formación profesional. Al Blgo. Franz Cardoso Pacheco, por permitirme consultar material de la colección científica del Laboratorio de Biología y Sistemática de Invertebrados Marinos de la Facultad de Ciencias Biológicas (LaBSIM), de la Universidad Nacional Mayor de San Marcos. A Gustavo Robles Fernández, Por permitirme consultar material del Instituto de Investigación y Desarrollo Hidrobiológico de la Universidad Nacional de San Agustín (INDEHI – UNSA). -
Upwelling-Downwelling Sequences in the Generation of Red Tides in a Coastal Upwelling System
MARINE ECOLOGY PROGRESS SERIES Vol. 112: 241-253,1994 Published September 29 Mar. Ecol. Prog. Ser. Upwelling-downwelling sequences in the generation of red tides in a coastal upwelling system G. H. Tilstone, F. G. Figueiras, F. Fraga Instituto de Investigaci6ns Marinas, Eduardo Cabello 6, CSIC, E-36208 Vigo, Spain ABSTRACT: Differences in temporal and spatial hydrographic conditions, water circulation patterns derived from temperature-salinity properties, phytoplankton community composition and distribution were studied in 4 Ria systems (flooded tectonic valleys) in Galicia, NW Spain, from 18 to 21 September 1986. The Rias are affected by upwelling cycles which introduce nutrient-rich Eastern North Atlantic Water (ENAW). During upwelling relaxation periods, the Rias are prone to red tide outbreaks, espe- cially during autumn. In the northern most Ria (Muros), after an upwelling event on 18 September followed by a weak downwelling, a low chlorophyll a (chl a) maximum occurred over the shelf which corresponded to the distribution of a large dinoflagellate/red tide species community identified by principal component analysis (PCA) and cluster analysis of species. This community was identified in all of the other Rias studied, but at different locations. With stronger downwelling on 21 September in the Ria de Vigo. Ria water and the chl a maximum were confined to the Ria interior, which corre- sponded to a shift in the large dinoflagellate / red tide community. The chl a maximum in all Rias was predominantly due to Heterosjqma carterae. The increase in Gymnodinium catenatum cell numbers, from the northern to the southern Rias, corresponded to stronger downwelling events. It is proposed that upwelling-downwelling sequences, enhanced by the presence of inlets and embayments acting as catchment concentration zones, are important mechanisms for generating red tide blooms in coastal upwelling systems. -
Cold Tongue Upwelling / Mixing Coupled Equatorial Feedbacks Engage the Global Climate
Cold tongue upwelling / mixing Coupled equatorial feedbacks engage the global climate. The thermocline carries the essential ocean memory. Upwelling/mixing enables surface-thermocline communication. Remote consequences of these feedbacks are the main reason people outside the tropical Pacific care about what we do. Upwelling controls property transports (BGC/nutrients/CO2). The vertical motion and exchanges produced by this circulation are now largely unconstrained by observations ... We must get this right. Walker Circulation El Niño precip teleconnections Up Trade Easterlies Winds East New WarmWarm Water heated by the sun Guinea by the sun Pool UpwellingUpwelling South America Cool lower water Thermocline Cool lower water W.S.Kessler, NOAA/PMEL 1 Meridional circulation and processes z y TIW x Green = Wind Brown = Fluxes Red = Solar Blue = Velocity = Mixing = Isotherms EUC Upwelling is O(m/day) vertical mixing must be very strong Consider the modeling challenge ... 2 MARCH 2001 JOHNSON ET AL. 845 How well can we describe this circulation? Observed v in the east-central Pacific: Two limitations MARCH 2001 JOHNSON ET AL. 845 Near-surface Ekman divergence drives upwelling, but ... Limitations: 1) Ship ADCP does not see upper 25m 2) TIW fluctuations Need near-surface Red=Northward currents to Blue=Southward describe the structure of the (Johnson et al. 2001) divergence. (Scatterometer winds also need referencing over Mean meridional current in the east-central Pacific FIG.5.Vertical–meridionalsectionsbasedoncentered28 lat linear fitsstrong to data currents.) taken from 1708 to 958W, regardless of longitude (see 22 21 22 21 text). (a) MeridionalShipboard velocity, ADCPy (10 overms 170°W-95°W); CI 2, positive (northward) shaded.