DOCSLIB.ORG

A Junior Synonym of Plastocerus Thoracicus, with Nomenclatural Notes on Plastocerus (Coleoptera: Elateridae: Plastocerinae)

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
A Junior Synonym of Plastocerus Thoracicus, with Nomenclatural Notes on Plastocerus (Coleoptera: Elateridae: Plastocerinae) 2020 ACTA ENTOMOLOGICA 60(2): 391–396 MUSEI NATIONALIS PRAGAE doi: 10.37520/aemnp.2020.23 ISSN 1804-6487 (online) – 0374-1036 (print) www.aemnp.eu SHORT COMMUNICATION Binhon atrum Pic – a junior synonym of Plastocerus thoracicus, with nomenclatural notes on Plastocerus (Coleoptera: Elateridae: Plastocerinae) Ji ří HÁJEK1), Michael A. IVIE2) & John F. LAWRENCE3) 1) Department of Entomology, National Museum, Cirkusová 1740, CZ-19300 Praha 9 – Horní Počernice, Czech Republic; e-mail: [email protected] 2) Montana Entomology Collection, Marsh Labs, Room 50, Montana State University, 1911 W. 6 Lincoln Street, Bozeman, MT 59717, USA; e-mail: [email protected] 3) Australian National Insect Collection, CSIRO, GPO 1700, Canberra, ACT and 61 Glenbar Road, The Palms, QLD, Australia; e-mail: [email protected] Accepted: Abstract. Binhon atrum Pic, 1922, an enigmatic genus and species described in the family 9th June 2020 Dascillidae based on a single female from Vietnam, is illustrated, diagnosed and formally Published online: synonymised with Plastocerus thoracicus Fleutiaux, 1918 (Elateridae: Plastocerinae). The 17th June 2020 species is recorded from China (Guangxi, Yunnan) for the fi rst time. The taxonomic history of the genus Plastocerus Schaum, 1852 (not Plastocerus LeConte, 1853) with emphasis on its synonyms, is briefl y summarised. Key words. Coleoptera, Dascillidae, Elateridae, Binhon, Plastocerus, new synonym, new records, China, Vietnam, Oriental Region Zoobank: http://zoobank.org/urn:lsid:zoobank.org:pub:AD67D5CD-E5B0-4CC6-8CA1-D11D2E6B6DDD © 2020 The Authors. This work is licensed under the Creative Commons Attribution-NonCommercial-NoDerivs 3.0 Licence. Introduction While the biggest problem in Pic’s species-group Maurice Pic (1866–1957), an eminent French coleo- taxa are the varieties – largely overlooked by other en- pterist of the turn of 19th and 20th Centuries, described tomologists – but representing available (and in many more than 20,000 species and varieties as well as several case also valid) names (e.g. H 2011, B hundred genera across many families of Coleoptera (V- R 2015), another problem is that his new genera 1958, C 2006). Like several entomologists in sometimes lack explicit family assignment, which has the 19th Century, Pic described taxa from almost all beetle led to misplacement or even omission of many taxa by groups from all parts of the world. Generally speaking, he subsequent entomologists. This problem has only been described anything which did not have a name in his (rather addressed recently, with the study of authentic material chaotically organized) reference collection, without much in Pic’s collection (see, e.g., I 2002, P 2005, eff ort to check for names that already existed elsewhere I P 2012). One of the enigmatic genera (C 2006). Consequently, he produced a great described by Pic is the monotypic Binhon Pic, 1922, with number of synonyms, including repeated descriptions the species Binhon atrum Pic, 1922 from the locality of the same taxon. Among his contemporaries, Pic was Hoa Binh in northern Vietnam (P 1922). The genus already infamous for very short and uninformative descrip- and species were described based on a single female, tions, insuffi cient for recognition of the taxon concerned. and placed in the family Dascillidae, but close to the A large proportion of these descriptions were published Neotropical genus Aploglossa Guérin-Méneville, 1849 in his three private journals: L’Échange, Révue Linnéenne (currently Ptilodactylidae: Aploglossinae). Although P (1885–1956, 543 issues), Mélanges Exotico-Entomolo- (1926) added subsequent information about the males of giques (1911–1939, 71 issues), and Opuscula Martialis Binhon, neither the fi ve-line long Latin description, nor (1940–1944, 13 issues). the supplementary information, provide suffi cient infor- Hajek.indd 391 17.06.2020 15:29:14 392 HÁJEK et al.: Binhon atrum Pic – a junior synonym of Plastocerus thoracicus (Coleoptera: Elateridae) mation to allow recognition of the beetle. Therefore, it bed Californian species, Plastocerus schaumii LeConte, is not surprising that Binhon has never been mentioned 1853, which led to a diff erent, broader concept of the in any entomological literature since 1926, either within genus for many years. L (1857) and C- Dascillidae (cf. J et al. 2013), or Ptilodactylidae (cf. (1863) correctly attributed Plastocerus to Schaum, S 1986). Only the fi nding of the type specimen but accepted LeConte’s broad defi nition to include both of Binhon in Pic’s collection of Dascillidae in the Mu- the Palaearctic and Nearctic species. Subsequently, séum national d’Histoire naturelle, Paris, enabled us to apparently considering the Nearctic species as the ‘true’ clarify the identity of this enigmatic beetle and propose Plastocerus, H (1883) proposed a new genus name the following synonymy with Plastocerus Schaum, 1852. Ceroplastus for the Palaearctic taxon; unfortunately, as As the classifi cation of Plastocerus Schaum, 1852 (not he did it again only in a catalogue and without descripti- Plastocerus LeConte, 1853) is rather complicated, and on, several authors considered the name as unavailable. several of its synonymies are missing from the recent These authors (e.g. H 1921) incorrectly attributed literature (cf. B 2007), the taxonomic history of the the name Ceroplastus to S (1888a,b), who gave a genus Plastocerus Schaum is summarised. redescription of the taxon in the generic part of both his ‘Fauna Baltica’ and ‘Fauna Transsylvanica’. The female of Plastocerus angulosus was described Materials and methods by R (1896) as a new genus and species: Pseudo- Habitus photographs were taken using a Canon EOS phyllocerus atricolor Reitter, 1896; but its synonymy was 550D digital camera with an attached Canon MP-E65mm soon recognized by S (1897). S (1902) f/2.8 1–5× macro lens, as numerous separate images at was the fi rst author to explicitly state that the Palaearctic diff erent focal planes and afterwards combined using and Nearctic Plastocerus species were not congeneric. He Helicon Focus 6.3.0 software. attributed the Palaearctic taxon name to C (1863) See B (2010) and B et al. (2018) for and thus incorrectly considered it junior to LeConte’s morphological terminology and detailed redescription of name, so he proposed another replacement name, Clado- Plastocerus. cerus Schwarz, 1902. Schwarz’s name was immediately The material mentioned in this study is deposited in the synonymised by H (1902), who pointed out that his following collections: name Ceroplastus was senior – although he mentioned ANIC Australian National Insect Collection, CSIRO Australian the later ‘Catalogus coleopterorum Europae, Caucasi National Research Collections, Canberra, ACT, Australia; et Armeniae rossicae’ (H et al. 1891), and not the MNHN Muséum national d’Histoire naturelle, Paris, France; original ‘Catalogus coleopterorum Europae et Caucasi’ NHMB Naturhistorisches Museum Basel, Switzerland; (H et al. 1883). In addition, H (1902) noted NMPC National Museum, Prague, Czech Republic. that Cladocerus Schwarz is a junior homonym of Cla- docerus Kirsch, 1865: 68 (Coleoptera: Lycidae) [in fact, Systematics K (1865) described Cladoceras; Cladocerus is its Plastocerus Schaum, 1852 unjustifi ed emendation by G H (1869: 1633)]. For the next 70 years, Plastocerus was considered Plastocerus Schaum, 1852: 49. Type species: Callirhipis angulosus Germar, 1844 (by monotypy). to be an American genus and Ceroplastus was the genus Ceroplastus Heyden, 1883 in H et al. (1883: 111), as an unnecessary occurring in the Old World, until C (1972) cor- replacement name for Plastocerus Schaum, 1852 [not Plastocerus rected the usage: recognizing Plastocerus Schaum, with LeConte, 1853]. Ceroplastus as its junior synonym, for the Mediterranean Pseudophyllocerus Reitter, 1896: 234. Type species: Pseudophyllocerus species, and replaced Plastocerus LeConte with its junior atricolor Reitter, 1896 (by monotypy); synonymized by S (1897: 64). synonym Octinodes Candèze, 1863. Cladocerus Schwarz, 1902: 200, as an unnecessary replacement name S (1852) originally placed Plastocerus in for ‘Plastocerus Cand.’ [= Plastocerus sensu C (1863); i.e. the family Cebrionidae. However, L (1857) Plastocerus Schaum, not Plastocerus LeConte]. included Plastocerus in the family Elateridae, where, Binhon Pic, 1922: 29. Type species: Binhon atrum Pic, 1922 (by mono- with the single sojourn to the Rhipiceridae in H typy); syn. nov. (1921), it remained until C (1972) placed it in Comments on the classifi cation. The genus Plastocerus its own family, Plastoceridae. Quite recently, B et was proposed by S (1852) in his ‘Catalogus Co- al. (2018) and K et al. (2018) revised the taxonomic leopterorum Europae’ for Callirhipis angulosus Germar, placement of Plastocerus using molecular analysis and 1824, an enigmatic elaterid species from the Eastern Me- downgraded its status to the subfamily Plastocerinae, diterranean (Greece, Turkey, Syria and Israel – see P back in the family Elateridae. However, the family group N 2011). Although Schaum’s act fully conforms name Plastoceridae Crowson, 1972 is a junior homonym with the conditions of ICZN (1999: Article 12.2.5), many of Plastocerini LeConte, 1861. Although B subsequent authors considered the genus a nomen nudum et al. (2011) called for a petition to the International and thus formally unavailable. Most importantly of the Commission on Zoological Nomenclature to suppress authors who thought Schaum’s name invalid, LC the LeConte name, no such petition is listed
Load more

Recommended publications
  • The Beetle Fauna of Dominica, Lesser Antilles (Insecta: Coleoptera): Diversity and Distribution
    INSECTA MUNDI, Vol. 20, No. 3-4, September-December, 2006 165 The beetle fauna of Dominica, Lesser Antilles (Insecta: Coleoptera): Diversity and distribution Stewart B. Peck Department of Biology, Carleton University, 1125 Colonel By Drive, Ottawa, Ontario K1S 5B6, Canada stewart_peck@carleton. ca Abstract. The beetle fauna of the island of Dominica is summarized. It is presently known to contain 269 genera, and 361 species (in 42 families), of which 347 are named at a species level. Of these, 62 species are endemic to the island. The other naturally occurring species number 262, and another 23 species are of such wide distribution that they have probably been accidentally introduced and distributed, at least in part, by human activities. Undoubtedly, the actual numbers of species on Dominica are many times higher than now reported. This highlights the poor level of knowledge of the beetles of Dominica and the Lesser Antilles in general. Of the species known to occur elsewhere, the largest numbers are shared with neighboring Guadeloupe (201), and then with South America (126), Puerto Rico (113), Cuba (107), and Mexico-Central America (108). The Antillean island chain probably represents the main avenue of natural overwater dispersal via intermediate stepping-stone islands. The distributional patterns of the species shared with Dominica and elsewhere in the Caribbean suggest stages in a dynamic taxon cycle of species origin, range expansion, distribution contraction, and re-speciation. Introduction windward (eastern) side (with an average of 250 mm of rain annually). Rainfall is heavy and varies season- The islands of the West Indies are increasingly ally, with the dry season from mid-January to mid- recognized as a hotspot for species biodiversity June and the rainy season from mid-June to mid- (Myers et al.
    [Show full text]
  • The Evolution and Genomic Basis of Beetle Diversity
    The evolution and genomic basis of beetle diversity Duane D. McKennaa,b,1,2, Seunggwan Shina,b,2, Dirk Ahrensc, Michael Balked, Cristian Beza-Bezaa,b, Dave J. Clarkea,b, Alexander Donathe, Hermes E. Escalonae,f,g, Frank Friedrichh, Harald Letschi, Shanlin Liuj, David Maddisonk, Christoph Mayere, Bernhard Misofe, Peyton J. Murina, Oliver Niehuisg, Ralph S. Petersc, Lars Podsiadlowskie, l m l,n o f l Hans Pohl , Erin D. Scully , Evgeny V. Yan , Xin Zhou , Adam Slipinski , and Rolf G. Beutel aDepartment of Biological Sciences, University of Memphis, Memphis, TN 38152; bCenter for Biodiversity Research, University of Memphis, Memphis, TN 38152; cCenter for Taxonomy and Evolutionary Research, Arthropoda Department, Zoologisches Forschungsmuseum Alexander Koenig, 53113 Bonn, Germany; dBavarian State Collection of Zoology, Bavarian Natural History Collections, 81247 Munich, Germany; eCenter for Molecular Biodiversity Research, Zoological Research Museum Alexander Koenig, 53113 Bonn, Germany; fAustralian National Insect Collection, Commonwealth Scientific and Industrial Research Organisation, Canberra, ACT 2601, Australia; gDepartment of Evolutionary Biology and Ecology, Institute for Biology I (Zoology), University of Freiburg, 79104 Freiburg, Germany; hInstitute of Zoology, University of Hamburg, D-20146 Hamburg, Germany; iDepartment of Botany and Biodiversity Research, University of Wien, Wien 1030, Austria; jChina National GeneBank, BGI-Shenzhen, 518083 Guangdong, People’s Republic of China; kDepartment of Integrative Biology, Oregon State
    [Show full text]
  • P a Rt 2: N O Tes 4 1
    R ec o rd s o f th e H a w a ii B io lo g ic a l S u rvey fo r 20 0 4 Ð 20 0 5 — P a rt 2: N o tes 4 1 Measurements: F em a le: B L 1 0 .0 8 Ð 1 4 .7 8 m m , B B 2.8 6 Ð 4 .20 . M a le: B L 9 .4 1 Ð 1 3 .9 4 m m ; B 2.6 8 3 .6 1 Ð 3 .7 8 m m . T y p es. H o lo ty p e / (B P B M 1 6 ,4 0 1 ): HAWAI‘I: K o n a F o res t U n it, H a k a la u F o res t N W R [N a tio n a l W ild life R es erve], 6 7 5 m [5 5 0 0 ft], 25 A u g 20 0 0 , rea red ex P itto sp o rum, W . H a in es c o l- lec to r; s a m e d a ta bu t 1 A u g 20 0 0 (a llo ty p e ?, p a ra ty p es : 1 ?, 2 /); s a m e d a ta bu t 5 S ep 20 0 0 (/ p a ra ty p e); s a m e d a ta bu t 9 N o v 20 0 0 (/ p a ra ty p e); s a m e d a ta bu t 1 9 N o v 20 0 0 , H V N P 0 0 3 8 5 4 (? p a ra ty p e, H A V O ); s a m e d a ta bu t 1 A u g 20 0 0 , H V N P 0 0 3 8 4 3 (/ p a ra ty p e, H A V O ) N o te: 2 / p a ra ty p es p res erved in a lc o h o l.
    [Show full text]
  • Banisteria, Number 20, 2002 © 2002 by the Virginia Natural History Society
    Banisteria, Number 20, 2002 © 2002 by the Virginia Natural History Society Thirteen Additions to the Known Beetle Fauna of Virginia (Coleoptera: Scirtidae, Bothrideridae, Cleridae, Tenebrionidae, Melyridae, Callirhipidae, Cerambycidae, Chrysomelidae) Richard L. Hoffman Virginia Museum of Natural History Martinsville, Virginia 24112 Steven M. Roble Virginia Department of Conservation and Recreation Division of Natural Heritage 217 Governor Street Richmond, Virginia 23219 Warren E. Steiner, Jr. Department of Systematic Biology - Entomology National Museum of Natural History, NHB-187 Smithsonian Institution Washington, D. C. 20560 INTRODUCTION SCIRTIDAE Ongoing inventories of the arthropod faunas of Ora troberti (Guérin) Virginia by our respective agencies continue to augment New northernmost and state records the lists of known resident species. Often these newly Members of this genus and the related Scirtes documented species are small and rarely collected, and resemble the chrysomelid “flea beetles” in having the hind their discovery in Virginia represents dramatic extensions femora greatly enlarged for jumping. Ora troberti is in their established areas of distribution, sometimes by listed for FL and TX (Peck & Thomas, 1998; Young, hundreds of miles. 2002a) and AL (Löding, 1945). There is a specimen We take this occasion to formally register thirteen labeled “La.” in the Casey Collection (USNM). White beetle species, most of which fall into the category just (1983) apparently used a specimen of O. troberti for his described, as native to Virginia. Eight of these species are field guide illustration of the genus. Its occurrence along resident in the southeastern Coastal Plain of the state and the Atlantic coast northward from Florida is not reported, three inhabit the Appalachians.
    [Show full text]
  • B a N I S T E R I A
    B A N I S T E R I A A JOURNAL DEVOTED TO THE NATURAL HISTORY OF VIRGINIA ISSN 1066-0712 Published by the Virginia Natural History Society The Virginia Natural History Society (VNHS) is a nonprofit organization dedicated to the dissemination of scientific information on all aspects of natural history in the Commonwealth of Virginia, including botany, zoology, ecology, archaeology, anthropology, paleontology, geology, geography, and climatology. The society’s periodical Banisteria is a peer-reviewed, open access, online-only journal. Submitted manuscripts are published individually immediately after acceptance. A single volume is compiled at the end of each year and published online. The Editor will consider manuscripts on any aspect of natural history in Virginia or neighboring states if the information concerns a species native to Virginia or if the topic is directly related to regional natural history (as defined above). Biographies and historical accounts of relevance to natural history in Virginia also are suitable for publication in Banisteria. Membership dues and inquiries about back issues should be directed to the Co-Treasurers, and correspondence regarding Banisteria to the Editor. For additional information regarding the VNHS, including other membership categories, annual meetings, field events, pdf copies of papers from past issues of Banisteria, and instructions for prospective authors visit http://virginianaturalhistorysociety.com/ Editorial Staff: Banisteria Editor Todd Fredericksen, Ferrum College 215 Ferrum Mountain Road Ferrum, Virginia 24088 Associate Editors Philip Coulling, Nature Camp Incorporated Clyde Kessler, Virginia Tech Nancy Moncrief, Virginia Museum of Natural History Karen Powers, Radford University Stephen Powers, Roanoke College C. L. Staines, Smithsonian Environmental Research Center Copy Editor Kal Ivanov, Virginia Museum of Natural History Copyright held by the author(s).
    [Show full text]
  • Key to Some of the Families of Beetles (Coleoptera)
    © 2003 Identification of Beetles (Coleoptera) Notes: There have been many name changes within the Order Coleoptera. Numerous families have been split, and some have been combined. Familys names listed here are those used in your textbook. When a textbook revision is published in 2004 the new family names will be used. Until then, please use the names as they are presented in your text. Dichotomous Keys to Some Families of Florida Coleoptera Introduction to the Identification of Beetles (Coleoptera) P. M. Choate, 2003 The order Coleoptera probably contains the larg- Suborder Adephaga est number of described species of any insect order. Beetles are found in almost every habitat, and range Carabidae - ground beetles, includes Rhysodidae in size from 1 – 100mm. in length. The heaviest and Cicindelidae known insect is a scarab beetle. Cicindelidae - tiger beetles now treated as Cara- The diversity of this order contributes to the com- bidae, tribe Cicindelini plexity of writing classification keys that encompass Rhysodidae - now placed in Carabidae all exceptions. Frequently generic keys that are writ- Haliplidae - aquatic ten to cover all species become so complex and Noteridae - aquatic lengthy that even experts get bogged down in de- Dytiscidae - aquatic tails. In order to introduce you to this order without Gyrinidae - aquatic, surface inhabitants only adding to the difficulty of identification I have con- stucted a key to 50+ families of beetles that you Suborder Polyphaga should be able to identify (Florida). Keep in mind that there are more than 100 families of beetles, and Staphylinidae - rove beetles, elytra reduced, expos- that this key is only intended to introduce you to the ing several abdominal tergites terminology and morphology that you will use in oth- Silphidae - carrion beetles er, more comprehensive keys.
    [Show full text]
  • In My Thesis I Have Focused on the Alpha-Taxonomy of Two Poorly
    In my thesis I have focused on the alpha-taxonomyof two poorly known beetle families Eulichadidae and Callirhipidae occurring predominantlyin forests of tropical areas.Both families areclassified as incertaesedis within the seriesElateriformia. The smďl elateriformfamily Eulichadidaecomprises of two extant genera:the monoýpic Califomian genusStenocolus LeConte, 1853,and the predominatelyOriental genusEulichas Jacobson,l9l3, with 42 describedspecies. The larvae ofboth geneÍaare aquatic,while the adultslive on vegetationneaÍ wateÍ, and especiallythe genusEulichas is often attractedat light. Altogether182 taxaat the specieslevel belongingto six generaare cunently classiftedin the family Callirhipidae.Members of the family occur mainly in tropics of Oriental,Australian and Neotropicalzoogeographical regions. Larvae feed on rottenwood, adultsare collectedon deadtree trunks, but mostofall they are attractedat light. REVISION OF THE GENUS EULICIIAS JACOBSON, 1913 (COLEOPTERA: EULICHADIDAE) I. INTRODUCTION, MORPHOLOGY OF ADULTS, KEY TO SUBGENERA AND SPECIES GROUPS. AND TAXONOMY OF E. FUNEBRIS SPECIES GROUP [ - seelist of publications] First part of the revision summarizesour knowledgeabout the genusEulichas. It containsa detailedmorphology of adults,and keys to identificationof subgeneraand speciesgroups of the genus.The taxonomyofthe E funebris speciesgroup is revised in detail. The group is characterisedby the long and slender phallobase,which is distinctly longer than the purÍirmeÍes'and by the long basď parameralapophysis. The group contains 16 species, including following nine newly describedspecies: E. birmanica Hájek, sp. nov' (Myanmar: Tenasserim),E.hauch Hájek,sp.nov.(Thailand),E.jaechi Hájek,sp.nov.(Malaysia), .ó.. janbezdeki Hájelq sp' nov. (Laos); E. kubani Hájek, sp. nov. (Laos, Vietnam)' E. meghalayensisHájek, sp. nov. (India:Meghálaya)' E, minutaHájek' sp. nov' (Sumatr4Nias, Siberut),E. strbaiHáje|rysp. nov. (Malaysia)' andE. tanahrataHájek,sp. nov.
    [Show full text]
  • A Catalog of the Coleóptera of America North of Mexico Family: Pselaphidae
    A CATALOG OF THE COLEÓPTERA OF AMERICA NORTH OF MEXICO FAMILY: PSELAPHIDAE ah52931 /^^ UNITED STATES AGRICULTURE PREPARED BY fiJUl DEPARTMENT OF HANDBOOK AGRICULTURAL "^^7 AGRICULTURE NUMBER 529-31 RESEARCH SERVICE FAMILIES OF COLEóPTERA IN AMERICA NORTH OF MEXICO Fascicy Family Year issued Fascicle^ Family Year issued Fascicle^ Family Year issued 1 Cupedidae 1979 46 Callirhipidae 102 Biphyllidae 2 Micromalthidae 1982 47 Heteroceiidae 1978 103 Byturidae 1991 3 Carabidae 48 Limnichidae 1986 104 Mycetophagidae 4 Rhysodidae 1985 49 Dryopidae 1983 105 ....... Ciidae 1982 5 Amphizoidae 1984 50 Elmidae 1983 107 Prostomidae 6 Haliplidae 51 Buprestidae 109 Colydiidae 8 Noteridae 52 Cebrionidae 110....... Monommatidae 9 Dytiscidae 53 Elateridae Ill Cephaloidae 10 Gyrinidae 54 Throscidae 112 Zopheridae 13 Sphaeriidae 55....... Cerophytidae 115 ....... Tenebrionidae 14 Hydroscaphidae 56 Perothopidae 116 AUeculidae 15 Hydraenidae 57 Eucnemidae 117 Lagriidae 16 Hydrophilidae 58 Telegeusidae 118 Salpingidae ^ 17 Georyssidae 61 Phengodidae 119....... Mycteridae 18 Sphaeritidae 62 Lampyridae 120 Pyrochroidae 1983 20 Histeridae 63 ]. Cantharidae 121 Othniidae 21 Ptiliidae 64 Lycidae 122....... Inopeplidae 22 Limulodidae.. 65 Derodontidae 1989 123 Oedemeridae 23 Dasyceridae ..... 66 Nosodendridae 124 Melandryidae 24 Micropeplidae 1984 67 Dennestidae 125 Mordellidae 1986 25 .'. Leptinidae 69 Ptinidae 126 Rhipiphoridae 26 Leiodidae 70 Anobiidae 1982 127 ....... Meloidae 27 Scydmaenidae 71 Bostrichidae 128 ....... Anthicidae 28 Silphidae 1993 72 Lyctidae 129 Pedilidae 29 Scaphidiidae 74 Trogositidae 130 Euglenidae 30 Staphylinidae 76 Cleridae 131 Cerambycidae 31 Pselaphidae 1997 78 Melyridae 132 Bruchidae 32 Lucanidae 79 Lymexylidae 133 Chrysomelidae 33 Passalidae 81 Sphindidae 134 Nemonychidae 1994 34 Scarabaeidae 1984 82 Nitidulidae 135 Anthribidae 35 Eucinetidae 83 Rhizophagidae 138 AUocorynidae 1991 36 Helodidae 86 Cucujidae 140 Brentidae 37 Clambidae r....
    [Show full text]
  • Mckenna2009chap34.Pdf
    Beetles (Coleoptera) Duane D. McKenna* and Brian D. Farrell and Polyphaga (~315,000 species; checkered beetles, Department of Organismic and Evolutionary Biology, 26 Oxford click beetles, A reP ies, ladybird beetles, leaf beetles, long- Street, Harvard University, Cambridge, MA 02138, USA horn beetles, metallic wood-boring beetles, rove beetles, *To whom correspondence should be addressed scarabs, soldier beetles, weevils, and others) (2, 3). 7 e ([email protected]) most recent higher-level classiA cation for living beetles recognizes 16 superfamilies and 168 families (4, 5). Abstract Members of the Suborder Adephaga are largely preda- tors, Archostemata feed on decaying wood (larvae) and Beetles are placed in the insect Order Coleoptera (~350,000 pollen (adults), and Myxophaga are aquatic or semi- described species). Recent molecular phylogenetic stud- aquatic and feed on green and/or blue-green algae ( 6). ies defi ne two major groups: (i) the Suborders Myxophaga Polyphaga exhibit a diversity of habits, but most spe- and Archostemata, and (ii) the Suborders Adephaga and cies feed on plants or dead and decaying plant parts Polyphaga. The time of divergence of these groups has (1–3). 7 e earliest known fossil Archostemata are from been estimated with molecular clocks as ~285–266 million the late Permian (7), and the earliest unequivocal fossil years ago (Ma), with the Adephaga–Polyphaga split at ~277– Adephaga and Polyphaga are from the early Triassic (1). 266 Ma. A majority of the more than 160 beetle families Myxophaga are not known from the fossil record, but are estimated to have originated in the Jurassic (200–146 extinct possible relatives are known from the Permian Ma).
    [Show full text]
  • Fossil Perspectives on the Evolution of Insect Diversity
    FOSSIL PERSPECTIVES ON THE EVOLUTION OF INSECT DIVERSITY Thesis submitted by David B Nicholson For examination for the degree of PhD University of York Department of Biology November 2012 1 Abstract A key contribution of palaeontology has been the elucidation of macroevolutionary patterns and processes through deep time, with fossils providing the only direct temporal evidence of how life has responded to a variety of forces. Thus, palaeontology may provide important information on the extinction crisis facing the biosphere today, and its likely consequences. Hexapods (insects and close relatives) comprise over 50% of described species. Explaining why this group dominates terrestrial biodiversity is a major challenge. In this thesis, I present a new dataset of hexapod fossil family ranges compiled from published literature up to the end of 2009. Between four and five hundred families have been added to the hexapod fossil record since previous compilations were published in the early 1990s. Despite this, the broad pattern of described richness through time depicted remains similar, with described richness increasing steadily through geological history and a shift in dominant taxa after the Palaeozoic. However, after detrending, described richness is not well correlated with the earlier datasets, indicating significant changes in shorter term patterns. Corrections for rock record and sampling effort change some of the patterns seen. The time series produced identify several features of the fossil record of insects as likely artefacts, such as high Carboniferous richness, a Cretaceous plateau, and a late Eocene jump in richness. Other features seem more robust, such as a Permian rise and peak, high turnover at the end of the Permian, and a late-Jurassic rise.
    [Show full text]
  • Les Arthropodes Continentaux De Guadeloupe (Petites Antilles)
    Société d’Histoire Naturelle L’Herminier Les Arthropodes continentaux de Guadeloupe (Petites Antilles) : Synthèse bibliographique pour un état des lieux des connaissances. Date Rédaction : François Meurgey 1 Les Arthropodes continentaux de Guadeloupe (Antilles françaises) : Synthèse bibliographique pour un état des lieux des connaissances. Version 1.1 François Meurgey Cette étude a été réalisée sous l’égide de la Société d’Histoire Naturelle L’HERMINIER et a bénéficié d’un financement par le Parc National de Guadeloupe. Ce rapport doit être référencé comme suit : SHNLH (Meurgey, F.), 2011. Les Arthropodes continentaux de Guadeloupe : Synthèse bibliographique pour un état des lieux des connaissances. Rapport SHNLH pour le Parc National de Guadeloupe. 184 pages. Photos page de couverture : Polites tricolor et Thomisidae (en haut), Enallagma coecum , mâle. Clichés Pierre et Claudine Guezennec. 2 AAVERTTISSSSEEMEENTT Ce travail est uniquement basé sur l’analyse et le dépouillement de la bibliographie relative aux Arthropodes de Guadeloupe. Les listes d’espèces proposées dans ce premier état des lieux sont préliminaires et doivent être corrigées et améliorées, mais également régulièrement mises à jour par les spécialistes, au gré des nouvelles données transmises et des compilations bibliographiques. Nous souhaitons prévenir le lecteur (surtout le spécialiste) qu’il est inévitable que des erreurs se soient glissées dans cette étude. Des espèces manquent très certainement, d’autres n’existent pas ou plus en Guadeloupe et un très grand nombre d’entre elles devraient voir leur statut révisé. Nous sommes bien entendu ouverts à toutes critiques, pourvu qu’elles servent à améliorer ce travail. 3 SOOMMMAIIREE INTRODUCTION ET REMERCIEMENTS .................................................................................... 5 PREMIERE PARTIE : OBJECTIFS ET DEMARCHE ......................................................................
    [Show full text]
  • INSECTA MUNDI a Journal of World Insect Systematics
    INSECTA MUNDI A Journal of World Insect Systematics 0144 The beetles of the island of St. Vincent, Lesser Antilles (Insecta: Coleoptera); diversity and distributions Stewart B. Peck Department Biology, Carleton University, 1125 Colonel By Drive Ottawa, Ontario K1S 5B6, CANADA Date of Issue: October 15, 2010 CENTER FOR SYSTEMATIC ENTOMOLOGY, INC., Gainesville, FL Stewart B. Peck The beetles of the island of St. Vincent, Lesser Antilles (Insecta: Coleoptera); diversity and distributions Insecta Mundi 0144: 1-77 Published in 2010 by Center for Systematic Entomology, Inc. P. O. Box 141874 Gainesville, FL 32614-1874 U. S. A. http://www.centerforsystematicentomology.org/ Insecta Mundi is a journal primarily devoted to insect systematics, but articles can be published on any non-marine arthropod taxon. Manuscripts considered for publication include, but are not limited to, systematic or taxonomic studies, revisions, nomenclatural changes, faunal studies, phylogenetic analy- ses, biological or behavioral studies, etc. Insecta Mundi is widely distributed, and referenced or ab- stracted by several sources including the Zoological Record, CAB Abstracts, etc. As of 2007, Insecta Mundi is published irregularly throughout the year, not as quarterly issues. As manuscripts are completed they are published and given an individual number. Manuscripts must be peer reviewed prior to submission, after which they are again reviewed by the editorial board to insure quality. One author of each submitted manuscript must be a current member of the Center for System- atic Entomology. Managing editor: Paul E. Skelley, e-mail: [email protected] Production editor: Michael C. Thomas, e-mail: [email protected] Editorial board: J. H. Frank, M.
    [Show full text]