A CATALOG OF THE COLEÓPTERA OF AMERICA NORTH OF MEXICO FAMILY: PSELAPHIDAE

ah52931

/^^ UNITED STATES AGRICULTURE PREPARED BY fiJUl DEPARTMENT OF HANDBOOK AGRICULTURAL "^^7 AGRICULTURE NUMBER 529-31 RESEARCH SERVICE FAMILIES OF COLEóPTERA IN AMERICA NORTH OF MEXICO

Fascicy Family Year issued Fascicle^ Family Year issued Fascicle^ Family Year issued

1 Cupedidae 1979 46 Callirhipidae 102 Biphyllidae 2 Micromalthidae 1982 47 Heteroceiidae 1978 103 Byturidae 1991 3 Carabidae 48 Limnichidae 1986 104 Mycetophagidae 4 Rhysodidae 1985 49 Dryopidae 1983 105 ...... Ciidae 1982 5 Amphizoidae 1984 50 Elmidae 1983 107 Prostomidae 6 Haliplidae 51 Buprestidae 109 Colydiidae 8 Noteridae 52 Cebrionidae 110...... Monommatidae 9 Dytiscidae 53 Elateridae Ill Cephaloidae 10 Gyrinidae 54 Throscidae 112 Zopheridae 13 Sphaeriidae 55...... Cerophytidae 115 ...... Tenebrionidae 14 Hydroscaphidae 56 Perothopidae 116 AUeculidae 15 Hydraenidae 57 Eucnemidae 117 Lagriidae 16 Hydrophilidae 58 Telegeusidae 118 Salpingidae ^ 17 Georyssidae 61 Phengodidae 119...... Mycteridae 18 Sphaeritidae 62 Lampyridae 120 Pyrochroidae 1983 20 Histeridae 63 ]. Cantharidae 121 Othniidae 21 Ptiliidae 64 Lycidae 122...... Inopeplidae 22 Limulodidae.. 65 Derodontidae 1989 123 Oedemeridae 23 Dasyceridae ..... 66 Nosodendridae 124 Melandryidae 24 Micropeplidae 1984 67 Dennestidae 125 Mordellidae 1986 25 .'. Leptinidae 69 Ptinidae 126 Rhipiphoridae 26 Leiodidae 70 Anobiidae 1982 127 ...... Meloidae 27 Scydmaenidae 71 Bostrichidae 128 ...... Anthicidae 28 Silphidae 1993 72 Lyctidae 129 Pedilidae 29 Scaphidiidae 74 Trogositidae 130 Euglenidae 30 Staphylinidae 76 Cleridae 131 Cerambycidae 31 Pselaphidae 1997 78 Melyridae 132 Bruchidae 32 Lucanidae 79 Lymexylidae 133 Chrysomelidae 33 Passalidae 81 Sphindidae 134 Nemonychidae 1994 34 Scarabaeidae 1984 82 Nitidulidae 135 Anthribidae 35 Eucinetidae 83 Rhizophagidae 138 AUocorynidae 1991 36 Helodidae 86 Cucujidae 140 Brentidae 37 Clambidae r..... 90 Cryptophagidae 141 Platypodidae 1979 38 Dascillidae 92 Languriidae 1983 142 Scolytidae 39 Rhipiceridae 93 Erotylidae 143 Curculionidae 1983 40 Byrrhidae 94 Phalacridae 144 Stylopidae 41 Psephenidae 1983 95 Cerylonidae 1982 145 Fossil Coleóptera 42 Brachypsectridae 96 Corylophidae 146 Ithyceridae 1994 43 Artematopidae 97 Coccinellidae 44 Ptilodactylidae 98 Endomychidae 1986 45 Chelonariidae 100 Lathridiidae

^Missing numbers are those assigned ¡n the computer program to families not found in the United States and Canada.

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To file a complaint, write the Secretary of Agriculture, U. S. Department of Agriculture, Washington, DC 20250, or call 1 (800) 245-6340 (voice) or (202) 720-1127 (TDD). USDA is an equal employment opportunity employer. A CATALOG OF THE COLEÓPTERA OF AMERICA NORTH OF MEXICO FAMILY: PSELAPHIDAE

BY DONALD S. CHANDLER DEPARTMENT OF ZOOLOGY UNIVERSITY OF NEW HAMPSHIRE DURHAM, NH 03824

.^'^ss^ I iMiTFn <;TATFS AGRICULTURE PREPARED BY ^^ nFPARTMFNTOF HANDBOOK AGRICULTURAL W ^™™TU^E^ NUMBER 529-31 RESEARCH

February 1997

FOREWORD

Many species of beetles are important pests of agricultural crops, stored food products, forests, wood products and structures, and fabrics. Many other species, in contrast, are beneficial in the biological suppression of pest arthropods and weeds, as well as in the decomposition of plant detritus, animal carcasses, and dung. Part of our national responsibility to American agriculture is to provide correct identification of species of American beetles so that appropriate controls can be applied. Most information about animal species, whether agricultural, biological, or experimental, is filed under the species' scientific names. These names are therefore the keys to retrieval of such information. Because some species have been known by several names, a complete listing of these names for each species is necessary. For the user of scientific names an up-to-date taxonomic catalog providing currently accepted names and pertinent bibliographic and distributional data is an indispensable tool. Although taxo- nomic literature is constantly changing to reflect current work, the traditional published taxonomic catalog remains static, with updating left to the individual user until it is revised. Production of catalogs in the past has been laborious, with long printing delays resulting in data that are obsolete before being published. However, the computer now provides the capability of storing, updating, and retrieving taxonomic data; rapid publication through computer-driven typesetting machinery; and a greater degree of currentness and flexibility. All the fascicles in this catalog of the beetles of America north of Mexico are produced by an original group of computer programs, designed and written during a pilot project by personnel of the Systematic Entomology Laboratory and the Communication and Data Services Division, Agricultural Research Service.

Floyd P. Horn Administrator Agricultural Research Service

111

PREFACE

The Coleóptera, or beetles, are represented in the world by about 220,000 described species, of which about 24,000 occur in the United States and Canada. A comprehensive taxonomic catalog of beetles for this area has not been available except the series of worid-based "Coleopterorum Catalogus" volumes (1909-present, Junk, Berlin). The Leng "Catalogue of the Coleóptera of America North of Mexico" (J.D. Sherman, Jr., Mt. Vemon, NY), which was published in 1920 with supplements to the end of 1947, is a checkUst. However, it has served professional and amateur alike for nearly 60 years as the principal source of scientific names of beetles. Since 1947, many new taxa have been described and many changes in status and nomenclature have appeared in numerous scattered publications, but little effort has been made to summarize these changes. This catalog will supplant the Leng catalog and supply additional essential information. It is produced by an original suite of storage, retrieval, and printing programs written especially for automated taxonomic catalogs. The catalog for each family is published as a separate fascicle with its introductory text, bibliography, and sequence. The publishing of separate fascicles makes data available shortly after they are assembled. Computer tapes for each fascicle are maintained for updating and necessary reprinting. The information on each family is the responsibility of the respective author or authors. The editors modify it only to correct obvious errors and to make it conform to the requirements of the computer programs. No original proposal for a new name, taxon, status, or classification is given, such data having been previously published, but new host and distributional data are often listed. The rules of "The International Code of Zoological Nomenclature" are followed. The geographic scope of this catalog includes the continental United States, Canada, Greenland, and the associated continental islands. Names of taxa found only in other regions are excluded. If the range of a species extends outside these geographic limits, this fact is indicated. On (or inside of) the back cover is a map of the 12 faunal regions based on historical and faunal criteria to simplify distribution recordings. Two-letter Postal Service style abbreviations are used for States and Provinces, and faunal regions are indicated in each distribution record by a diagonal line between groups of abbreviations. It is not the purpose of this catalog to present a complete scheme of higher classification within the order. The familial makeup is somewhat intermediate between that of R.H. Arnett in "The Beetles of the United States" (1960-62, CathoUc University Press, Washington, DC) and that of R.A. Crowson in "The Natural Classification of the Famines of Coleóptera" (1967, Biddies Ltd., Guildford, England). Modifications of these two systems are largely those advocated by J.F. Lawrence based in part on suggestions by taxonomic specialists for certain families. Generic groups and higher categories within the family are arranged phylogenetically as indicated by the author of the particular fascicle, and species group names with their respective synonyms are arranged alphabeti- cally. Names referable to incertae sedis and nomen dubium are listed separately at the end of the nearest appli- cable taxon with notations as to their status. Each available name is followed by its author, date proposed, and page number referring to the complete bibliographic citation containing the original description. Following each generic name are the type-species and method of its designation, necessary explanatory notes, and pertinent references on immature stages, taxonomy, redescription, ecology, and keys. After the specific name entry are the original genus (if different from the present placement), type-locality, geographical distribution by State, Province, and broad extralimital units, explanatory notes, pertinent references to immature stages, taxonomy, redescription, and ecology, depository of type-specimen and its sex, and hosts. In addition to the list under the map (on or inside of the back cover) of faunal regions, the following abbreviations are used in this catalog:

ABBREVIATIONS, GENERAL

Amer. Bor.—America Borealis Mus.—Museum Amer. Sept.—America Septentrionalis N.—North Automat.—Automatic N. Amer.—North America C. Amer.—Central America Orig. des.—Original designation Co.—County pi.—^plate Cosmop.—Cosmopolitan Preocc.—Preoccupied Design.—Designated S.—South E.—East S. Amer.—South America F.—Female ser.—series fig.—figure Sp.—Species Holarc.—Holarctic Subseq. monot.—Subsequent monotypy Isl.—Island Subsp.—Subspecies Km.—Kilometers Taut.—^Tautonymy M.—Male Univ.—University Mex.—Mexico USA—United States of America mi.—mile Var.—Variety Monot.—Monotypy W.—West Mts.—Mountains W. Ind.—West Indies

MUSEUMS IN THE UNITED STATES AND CANADA^

AMNH—American Museum of Natural History, FSCA—Florida State Collection, Gainesville New York HAHC—H. & A. Howden Collection, Ottawa, ANSP—Academy of Natural Sciences, Phila- Canada delphia, PA ICCM—Carnegie Museum, Pittsburgh, PA BPBM—Bernice P. Bishop Museum, Honolulu INHS—^Illinois Natural History Survey, Urbana BYUC—Brigham Young University, Provo, UT JGEC—J. G. Edwards Collection, San Jose, CA CASC—California Academy of Sciences, San KMFC—K. M. Fender Collection, McMinnville, Francisco OR CISC—University of California, Berkely KSUC—Kansas State University, Manhattan CMNH—Carnegie Museum of Natural History LACM—Los Angeles County Museum, CA CNCI—Canadian National Collections, Ottawa LSUC—^Louisiana State University, Baton Rouge CUIC—Cornell University, Ithaca, NY MCZC—Museum of Comparative Zoology, CWOB—C. W. O'Brien Collection, Tallahassee, FL Harvard University, Cambridge, MA DHKC—D. H. Kistner Collection, Chico State MSUC—Michigan State University, East Lansing College, CA NCSM—North Carolina State University, Raleigh ELSC—E. L. Sleeper Collection, Long Beach, CA NYSM—New York State Museum, Albany FMNH—Field Museum of Natural History, Chicago, IL

^Abbreviations for U. S. and Canadian museums abridged from Amett, R. H., Jr., and Samuelson, G. A., 1969, "Directory of Coleóptera Collections of North America (Canada Through Panama)," Cushing-Malloy, Ann Arbor, MI, 123 pp.

VI OSEC—Oklahoma State University, Stillwater SJSC—San Jose State College, CA OSUC—Ohio State University, Columbus SLWC—S. L. Wood Collection, Provo, UT OSUO—Oregon State University, Corvallis SMSH—Stovall Collection, University of Oklahoma, PMNH—Peabody Museum, Yale University, Norman New Haven, CT TAMU—Texas A. & M. University, College Station PSUC—Pennsylvania State Museum, University UANH—Insect Collection, Alabama Museum of Natural History Park UCDC—University of California, Davis PURC—Purdue University, West Lafayette, IN UICM—University of Idaho, Moscow RUIC—Rutgers University, New Brunswick, NJ UMMZ—University of Michigan, Ann Arbor SEMC—Snow Museum, University of Kansas, UMRM—University of Missouri, Columbia Lawrence USNM—U.S. National Museum of Natural History, Washington, DC WSUC—Washington State University, Pullman

MUSEUMS IN FOREIGN COUNTRIES

BMNH—British Museum (Natural History), NMPC—Narodni Museum, Prague, London Czechoslovakia GUHC—Glasgow University, Hunterian SCUT—Spinola College, University of Turin, College, Scotland Italy HMOX—Hope Museum, Oxford, England SMTD—Staatliches Museum für Tierkunde, IPZE—Institut Pflanzenschutzforschung Dresden, East Germany Zweigstelle, Eberswald, East Germany UNAM—Universidad Nacional Autónoma, IRSB—Institut Royal Sciences Belgique, Mexico City Brussels UZMC—University Zoological Museum, MFNB—^Museum für Naturkunde (Humboldt), Copenhagen, Denmark Berlin UZMH—University Zoological Museum, MGFT—Museum G. Prey, Tutzing, Munich, Helsinki, Finland West Germany ZMAS—Zoological Museum, Academy of MHNG—Muséum d'Histoire Naturelle, Geneva Sciences, Leningrad MHNL—Museum d'Histoire Naturelle, Lyon, ZMPA—Zoological Museum, Polish Academy of France Sciences, Warsaw MNHP—Museum National d'Histoire Naturelle, ZMUL—^Zoological Museum, University of Paris Lund, Sweden MNSL—Museum of Natural Sciences, Leipzig, ZMUM—Zoological Museum, University of East Germany Moscow MZBS—Museum Zoología, Barcelona, Spain ZSBS—Zoologische Sammlung Bayerischen NHRS—^Naturhistoriske Riksmuseet, Stockholm Staates, Munich, West Germany

vn ACKNOWLEDGMENTS

We are indebted to many individuals who contributed to the planning and development of this catalog. We are especially grateful to the following specialists who helped to make it as complete and accurate as possible: Richard H. Foote, Systematic Entomology Laboratory (SEL), Agricultural Research Service (ARS), for his sug- gestions, guidance, and encouragement; C. W. Sabrosky, SEL, for valuable counsel on nomenclatural problems; J. F. Lawrence, Division of Entomology, Commonwealth Scientific and Industrial Research Organization, Can- berra, Australia, for his recommendations on higher categories; and more than 50 coleopterists in Canada, the United States, and Mexico for voluntarily contributing information about their specialty groups. We wish to acknowledge the extensive computer-programming and editing-system support provided by members of the former Communications and Data Services Division of ARS during the early years of this project. We thank Roslyn W. Norman for typing the text for this fascicle.

Systematic Entomology Laboratory, Agricultural Research Service U. S. Department of Agriculture, Beltsville, MD

Editorial Board

J. M. Campbell, Biosystematics Research Institute, Agriculture Canada, Ottawa, ON T. L. Erwin, Department of Entomology, Smithsonian Institution, Washington, DC H. F. Howden, Department of Biology, Carleton University, Ottawa, ON J. M. Kingsolver, Florida State Collection of Arthropods, Gainesville, FL P. J. Spangler, Department of Entomology, Smithsonian Institution, Washington, DC T. J. Spilman, Systematic Entomology Laboratory, ARS, Washington, DC R. E. White, Systematic Entomology Laboratory, ARS, Washington, DC

vni CONTENTS

Family Pselaphidae 1 Division Brachyscelia 2 Subfamily Faroninae 2 Tribe Faronini 2 Subfamily Euplectinae 5 Tribe Trogastrini 5 Tribe Euplectini 12 Tribe Jubini 35 Subfamily Bythinoplectinae 36 Tribe Bythinoplectini 36 Tribe Mayetiini 36 Subfamily Batrisinae 38 Tribe Batrisini 38 Tribe Amauropini 49 Subfamily Goniacerinae 52 Tribe Trichonychini 52 Tribe Brachyglutini 53 Tribe Iniocyphini 69 Tribe Bythinini 69 Tribe Valdini 74 Tribe Tychini 75 Tribe Speleobamini 80 Division Macroscelia 81 Subfamily Pselaphinae 81 Tribe Tyrini 81 Tribe Ceophyllini 84 Tribe Tmesiphorini 84 Tribe Ctenistini 84 Tribe Arhytodini 86 Tribe Pselaphini 87 Subfamily Clavigerinae 88 Tribe Clavigerini 88 Bibliography 90 Index 111

IX

Family PSELAPHIDAE Latreille, 1802

Clavigerides Leach, 1815.

By Donald S. Chandler

Taxonomy: Brendel, 1890, 1891; Bowman, 1934; Park, 1951 and 1953d; Amett, 1963; Chandler, 1990b; Newton and Chandler, 1989. Ecology: Park, 1947a and 1964; Park, Auerbach, and Corley, 1950; Park, Auerbach, and Wilson, 1953; Chandler, 1987. Immatures: (eggs) de Marzo, 1986b; Carlton 1989b; (larvae): Böving and Craighead, 1931; de Marzo, 1986a and 1987; Newton, 1991; (pupae): de Marzo, 1988.

At this time there are 712 species in 102 genera known in the family Pselaphidae from America north of Mexico, and over 8,000 species are described for the world. This group has been revised extensively for the Nearctic region, since their striking cuticular projections, foveae, and sulci have long aroused the interest of taxonomists. The family has been considered close to the Staphylinidae, and it has been placed near the staphylinid subfamilies Oxytelinae (Park, 1942), Aleocharinae (Jeannel, 1950), Steninae (Crowson, 1955), and Leptotyphlinae (Naomi, 1985). The family has also been placed in the Omaliinae group (Thayer, 1987) and as a sister-group to the staphylinid subfamily Protopselaphinae (New1;on and Thayer, 1995). Newton and Thayer (1995) placed the family Pselaphidae as a subfamily within the Staphylinidae as was also done in the recent classification of Coleóptera of the world (Lawrence and Newton, 1995). In the first Nearctic monograph of this group, LeConte (1849) added 50 new species to the 10 previously described and characterized many of the distinctive North American genera. In the next monograph (Brendel, 1890-1891), the number of species had tripled. The last work which could be termed a monograph of the Nearctic species is that of Bowman (1934). It is principally a compilation of the previous works; an important part of this monograph is the designation of many of the type- species. The most productive worker has been Casey (1884-1924), who described almost a third of the North American species and genera. However, the most important worker has been Park (1932-65), who not only increased the knowledge of many poorly known groups, such as the cave inhabitants and ant guests, but also reorganized the higher classification of the family for the New World. Grigarick, Schuster, and Marsh (1954-80) revised almost all of the genera of western North America in the last 50 years, making the fauna in this area more thoroughly treated than that in any other area. The origin of the modem classification of the Pselaphidae is in the work of Raffray (1890a), who recognized many of the current tribes. Park (1942, 1951) modified Raffray's system chiefly by adding and redefining the tribes and subtribes of the New World. The last major change to the system was by Jeannel (1959), whose analysis of the world fauna resulted in the family's division into six subfamilies, with Newton and Thayer (1995) adding a seventh. Newton and Chandler (1989) produced an integrated classification of the groups of Jeannel and Park in their catalog of the genera of the world. The biology of the family is not well known; only the larvae of three species have been described for the Nearctic region (Böving and Craighead, 1931; Wagner, 1975). Most biological informafion is confined to the type of habitat in which the adults are collected. The majority of the species are predators of mites, CoUembola, worms, and other small invertebrates (Park, 1947b; Engelmann, 1956; Reichle, 1969). The Clavigerinae and a few species scattered through the other subfamilies are obligate myrmecophiles (Kistner, 1982). They feed on the living or dead larvae of the host ants and may also be fed by trophallaxis (Park, 1932a; Akre and Hill, 1973). Several pselaphid species, which are ordinarily free-living, may also occur in ant colonies. These facultative myrmecophiles tend to increase their frequency in ant nests with an increase in latitude (Park, 1947b). Data on the life cycles and cytology of two species are presented by Mickey and Park (1956). Carlton (1990) discussed the biogeographic patterns of the genera of eastern North America. 2 COLEÓPTERA CATALOG

Species are generally found in moist leaf litter or rotten wood and are usually extracted with a Berlese/Tullgren funnel (Chandler, 1990b). A few species may be collected under rocks or under the bark of dead trees. The myrmecophiles are often found with ant species of the genus Lasius, and less frequently with Aphaenogaster and Formica and other genera. Ultraviolet and white lights attract many species whose habitats are often difficult to identify. Recent ecological interest in this group has focused on the use of species as indicators of old-growth forests (Vit, 1985; Chandler, 1987; Balletto and Cásale, 1991). This manuscript was received November, 1979 and last modified in June, 1996. Division BRACHYSCELIA Raffray, 1890a Subfamily FARONINAE Reitter, 1882 Tribe FARONINI Reitter, 1882 Genus MEGARAFONUS Casey Megarafonus Casey, 1897: 550 (as genus). Type-species: Megarafonus ventralis Casey (monot.). TAXONOMY: Schuster and Marsh, 1958b. REDESCRIPTION: Schuster and Marsh, 1958b.

Subgenus MEGARAFONUS Casey lentus Schuster and Marsh, 1958b; 189, figs. 1, 4, 10. OR: Clatsop Co., Seaside; OR. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From leaf litters of Douglas fir, Sitka spruce, maple, fir, and hemlock. mancus Schuster and Marsh, 1958b: 192, fig. 12. OR: Curry Co., Gold Beach; OR/ CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From redwood leaf litter. ventralis Casey, 1897: 550. OR: Multnomah Co., Portland; AK/ BC WA OR/ CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Schuster and Marsh, 1958b. HOST: From leaf litters of redwood, Douglas fir, maple/alder mixtures, and oak/maple mixtures; from rotten logs.

Subgenus NANORAFONUS Schuster and Marsh

Nanorafonus Schuster and Marsh, 1958b: 192 (as subgenus of Megarafonus). Type-species: Megarafonus parvus Schuster and Marsh (orig. des.). parvus Schuster and Marsh, 1958b: 192, figs. 2, 8, 9. CA: Butte Co., near Stringtown Hill, NE. Oroville; CA TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From litters of pine, oak, madrone, and moss.

Genus SONOMA Casey

Sonoma Casey, 1886: 195 (as genus). Type-species: Sonoma corticina Casey (des. by Lucas, 1920: 597). Faronus (of authors, not North American) (not Faronus Aube, 1844: 157). Sagola (of authors, not North American) (not Sagola Sharp, 1874: 506). REDESCRIPTION: Marsh and Schuster, 1962. KEYS: Marsh and Schuster, 1962. PSELAPHIDAE

Rafonus Casey, 1894: 441 (as genus) (synonymized by Raffray, 1904a). Type species: Faronus tolulae LeConte (monot.). cascadia Chandler, 1986b: 335, fig. 2. OR: Lane Co., H. J. Andrews Experimental Forest, Road 1508; OR. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From litters of old-growth Douglas-fir leaves and logs (Chandler, 1986b). cavifrons Casey, 1887: 481. CA: Mendocino Co.; OR/ CA. Neotype: CA: Santa Clara Co.; apparently no types from Mendocino Co. were found. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. longicollis Casey, 1894: 438 (synonymized by Marsh and Schuster, 1962). CA: Santa Cruz Co., Santa Cruz Mts. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. subsimilis Casey, 1894: 439, pi. I, fig. 4 (synonymized by Marsh and Schuster, 1962). CA: Sonoma Co. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: From leaf litters of hemlock, maple, and alder. conifera Chandler, 1986b: 336, fig. 4. OR: Benton Co., Mary's Peak; OR. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From leaf litters of cedar, hemlock, Douglas fir, and Sitka spruce. corticina Casey, 1887: 480, pi. XVI, fig. 10. CA: Mendocino Co., Soda Springs, Anderson Valley; WA OR/ CA. TYPE DEPOSITORY: USNM. REDESCRIPTION: Marsh and Schuster, 1962. HOST: From redwood leaf litter; under bark.

cuneata Marsh and Schuster, 1962: 43, figs. 13, 23. CA: Del Norte Co., Fort Dick; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From redwood leaf litter.

dilopha Marsh and Schuster, 1962: 47, fig. 17. CA: Humboldt Co., Blue Lake; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From alder and skunk cabbage, and pine leaf litters.

dolabra Marsh and Schuster, 1962: 37, fig. 4. CA: Del Norte Co., 18 mi. S. Klamath; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From alder and fern litter.

grandiceps Casey, 1894: 437. CA: Santa Cruz Co.; CA. TYPE DEPOSITORY: USNM. REDESCRIPTION: Marsh and Schuster, 1962.

héspera Park and Wagner, 1962: 6, pi. I, figs. 4, 5. OR: Polk Co., Boyer; OR BC/ CA. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Marsh and Schuster, 1962. HOST: Primarily from Douglas-fir leaf litter; also from leaf litters of moss, maple, cedar, and canyon live-oak.

humilis Marsh and Schuster, 1962: 45, figs. 15, 22. CA: Mendocino Co., Little River; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From redwood litter. 4 COLEÓPTERA CATALOG isabellae LeConte, 1851: 215 (Faronus), CA: San Diego Co., Santa Ysabel; CA. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. REDESCRIPTION: Marsh and Schuster, 1962. HOST: From leaf litter. margemina Park and Wagner, 1962: 5, pi. I, fig. 3. OR: Yamhill Co., Peavine Ridge near McMinnville; BC WA OR/ AK. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Marsh and Schuster, 1962. HOST: From mixed conifer leaf litter and moss; from alder leaf litter. olycalida Park and Wagner, 1962: 6, pi. I, fig. 6. WA: Clallami Co., Olympic National Park, Olympic Hot Springs; WA OR. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Marsh and Schuster, 1962. HOST: Primarily from Douglas-fir and fir leaf litters; also in alder leaf litter. parviceps Mäklin, 1852: 372 (Euplectus), AK: Sitka; AK/ WA OR ID BC/ CA. TYPE DEPOSITORY: MCZC, UZMH. REDESCRIPTION: Park and Wagner, 1962. HOST: From leaf litters of Douglas fir, red alder, and mixed conifer; from moss. petersi Chandler, 1986b: 333, fig. 1. OR: Benton Co., MacDonald Forest, Oak Creek area; OR. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From oak leaf litter. priocera Marsh and Schuster, 1962: 48, fig. 19. OR: Linn Co., Berlin; OR. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. quercicola Chandler, 1986b: 336, fig. 3. OR: MacDonald Forest, Oak Creek area; OR. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. repanda Marsh and Schuster, 1962: 35, figs. 1, 2. CA: Santa Clara Co., Steven's Creek Reservoir; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From Douglas fir and oak leaf litters. rubida Casey, 1894: 439. CA: Santa Cruz Co., Santa Cruz Mts.; CA/ OR. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Marsh and Schuster, 1962. HOST: From redwood litter. russelli Chandler, 1986b: 337, fig. 5. OR: Benton Co., Mary's Peak; OR. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From leaf litters of alder, Douglas fir, cedar, and hemlock; from rotten wood; from moss. spadica Marsh and Schuster, 1962: 36, fig. 3. CA: Mendocino Co., Mendocino; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From leaf litter. PSELAPHIDAE 5 tolulae LeConte, 1849: 109 {Faronus). GA: "cataractam Georgiae" (northern Georgia); PA VA/ GA NC. TYPE DEPOSITORY: MCZC. REDESCRIPTION: Marsh and Schuster, 1962. HOST: From Rhododendron and oak leaf litters; in well decomposed wood with roots (Schmitt, 1901). triloba Marsh and Schuster, 1962: 43, fig. 12. CA: Mendocino Co., Fort Bragg; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. vanna Marsh and Schuster, 1962: 38, fig. 5. CA: Monterey Co., 7 mi. S. of Big Sur; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From redwood leaf litter. yahiorum Chandler, 1983: 218, fig. 2. CA: Butte Co., Little Chico Creek; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From rotten live-oak logs. Subfamily EUPLECTINAE LeConte, 1861

Grigarick and Schuster, 1980 Tribe TROGASTRINI Jeannel, 1949 Subtribe TROGASTRINA Jeannel, 1949

Genus RHEXmiUS Casey

Rhexidius Casey, 1887: 478 (as genus). Type-species: Rhexidius granulosus Casey (monot.). TAXONOMY: Schuster and Grigarick, 1962 (revision). REDESCRIPTION: Schuster and Grigarick, 1962; Grigarick and Schuster, 1980. KEYS: Schuster and Grigarick, 1962. aggestus Schuster and Grigarick, 1962: 6, figs. 10, 19. CA: Santa Cruz Co., Boulder Creek; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From redwood and madrone leaf litters. asperulus Casey, 1894: 449. CA: Santa Cruz and San Francisco Cos.; CA. TYPE DEPOSITORY: USNM. REDESCRIPTION: Schuster and Grigarick, 1962. HOST: From leaf litters of redwood, laurel, and oak. crenatus Schuster and Grigarick, 1962: 6, figs. 9, 18. CA: Marin Co., 2 mi. S. of Olema; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From laurel and redwood leaf litters.

cuspidatus Schuster and Grigarick, 1962: 10, figs. 14, 23. CA: Mendocino Co., Mendocino; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From redwood leaf litter. glareosus Schuster and Grigarick, 1962: 10, figs. 1-5, 13, 22. CA: Marin Co., Samuel P. Taylor State Park; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1980. HOST: From redwood leaf litter. 6 COLEÓPTERA CATALOG granulosus Casey, 1887: 478, pi. XVI, fig. 8. CA: Alameda Co., Alameda; CA. TYPE DEPOSITORY: USNM. REDESCRIPTION: Schuster and Grigarick, 1962. HOST: From redwood leaf litter. hispidus Schuster and Grigarick, 1962: 11, figs. 15, 24. CA: Mendocino Co., Mendocino; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From redwood leaf litter. impensus Schuster and Grigarick, 1962: 13, figs. 6, 25. CA: Napa Co., Mount George, 7 mi. E. of Napa; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From oak and laurel leaf litters. incomptus Schuster and Grigarick, 1962: 8, figs. 11, 20. CA: Monterey Co., 7 mi. S. of Point Sur; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From leaf litters of mixed oak, laurel, and cedar.

Genus CONOPLECTUS Brendel

Conoplectus Brendel, 1888a: 301 (as genus, not Brendel, 1888b). Type-species: Euplectus canaliculatus LeConte (monot.). Brendel published the same paper twice (1888a and 1888b), with the 1888a paper having priority, published in June 1888, while the 1888b paper was published in November. Prorhexius Raffray, 1890a: 107 (as genus, synonymized by Brendel, 1892a). Ty^Q- spQCiQs: Prorhexius sylvaticus Rdiñmy {s\xhsQÇ\. monot. Raffray, 1890b: 197). TAXONOMY: Carlton, 1983 (revision). REDESCRIPTION: Carlton, 1983. KEYS: Carlton, 1983. Hexirhexius Grigarick and Schuster, 1980: 20 (as genus, synonymized by Carlton, 1983). acornus Carlton, 1983:71, figs. 25-27, 38. FL: Marion Co., 22.6 km. E. of Lynne, Ocala National Forest; FL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From litter of sand pine, oak, and palmetto; at UV light. bicentennialis Carlton, 1983: 73, figs. 33-38. FL: Franklin Co., 4.8 km. NW. of Alligator Point; FLGA. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From litter of sand pine, oak, and palmetto; in rotten tree root. canaliculatus LeConte, 1849: 107 {Euplectus). SC: Richland Co., Columbia; GA: Clarke Co., Athens; WI MI ON/ IL IN OH KY lA MO/ PA NY NJ DE MD WV VA DC/ RI CT MA NH/ MS AL GA SC NC FL LA AR TN/ TX OK. TYPE DEPOSITORY: MCZC. sylvaticus Raffray, 1890b: 197 {Prorhexius) (synonymized by Casey, 1894). "Amérique boréale." TYPE DEPOSITORY: MNHP. SEX OF TYPE: F. REDESCRIPTION: Park, Auerbach, and Corley, 1950; Grigarick and Schuster, 1980; Carlton 1983. ECOLOGY: Reichle, 1966 and 1969. HOST: From various deciduous and coniferous leaf litters and rotten woods (Chandler, 1990a); from sphagnum PSELAPHIDAE 7 excavoides Carlton, 1983: 68, figs. 19-24, 38. FL: Gadsden Co., 1.6 km. W. of Greensboro; FL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From beech and pine litters. susae Carlton, 1983: 72, figs. 28-32. AR: Marion Co., Buffalo Point near Mull; AR OK. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From various leaf and log litters; collected with ants (Chandler, 1990a).

Genus OROPUS Casey

Oropus Casey, 1886: 196 (as genus). Type-species: Oropus convexus Casey (orig. des.). Trichonyx (of authors, not North American) (not Chaudoir, 1845:64) TAXONOMY: Schuster and Grigarick, 1960. REDESCRIPTION: Schuster and Grigarick, 1960; Grigarick and Schuster, 1980. KEYS: Schuster and Grigarick, 1960. abbreviatus Casey, 1886: 200. CA: Sonoma Co.; CA. TYPE DEPOSITORY: A pseudotype is in USNM from Fort Gaston, Hoopa Valley, Humboldt Co., CA. The type is not at USNM (perhaps lost). acriculus Schuster and Grigarick, 1960: 292, fig. 38. CA: Tulare Co., 29 km. NE. of California Hot Springs; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From pine and mixed-conifer leaf litters; leaf litter by stream. acumenis Schuster and Grigarick, 1960: 296, fig. 42. CA: Amador Co., 10 km. N. of lone; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. basalis Casey, 1908: 276. CA: Santa Cruz Co.; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Schuster and Grigarick, 1960. HOST: From leaf litters of redwood, cypress, and Douglas fir. brevipennis Casey, 1908: 275. BC: Metlakatla; BC. This name is probably a synonym of O. keenU fide Schuster and Grigarick, 1960. TYPE DEPOSITORY: USNM.

castaneus Casey, 1908: 276. CA: Marin Co.; OR/ CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. REDESCRIPTION: Schuster and Grigarick, 1960. HOST: From leaf litters of redwood, laurel, Douglas fir, and alder.

cavicauda Casey, 1894: 448. CA: Marin Co.; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Schuster and Grigarick, 1960. HOST: From leaf litters of redwood, live oak, laurel, oak, and maple.

convexus Casey, 1886: 198. CA: Sonoma Co.; CA. TYPE DEPOSITORY: USNM. REDESCRIPTION: Schuster and Grigarick, 1960.

cristatus Schuster and Grigarick, 1960: 285, figs. 15, 30. CA: Napa Co., 11 km. E. of Rutherford; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. 8 COLEÓPTERA CATALOG curtipennis Casey, 1908: 275. CA: Humboldt Co.; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. REDESCRIPTION: Schuster and Grigarick, 1960. HOST: From fir and cedar leaf litters. cyranus Schuster and Grigarick, 1960: 289, figs. 18, 34. CA: Calaveras Co., Mokelumne Hill; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. debilis Casey, 1908: 277. CA: Siskiyou Co.; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Schuster and Grigarick, 1960. HOST: From leaf litters of Douglas fir, oak, redwood, and buckeye. delimatus Schuster and Grigarick, 1960: 295, figs. 22, 40. CA: Napa Co., Oakville; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From redwood and oak leaf litters. fenderi Park and Wagner, 1962: 14, pi. II, figs. 5, 6. OR: Yamhill Co., Peavine Ridge, near McMinnville; OR. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. grigaricki Chandler, 1983: 220, fig. 3. CA: Butte Co., 5 mi. NE. Forest Ranch; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From various leaf litters and rotten woods. helferi Grigarick and Schuster, 1962d: 307, figs. 1, 2. CA: Mendocino Co., Mendocino; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. interruptus Casey, 1886: 199. CA: Sonoma Co., Fisks Mill; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Schuster and Grigarick, 1960. HOST: From redwood and Douglas-fir leaf litters. keeni Casey, 1908: 274. BC: Metlakatla; BC WA OR. TYPE DEPOSITORY: USNM. HOST: From leaf litters of hemlock, Douglas fir, cedar, maple, and alder. macneilli Schuster and Grigarick, 1960: 287, fig. 33. CA: Marin Co., Alpine Lake; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. magnidens Schuster and Grigarick, 1960: 286, figs. 9, 17, 32. OR: Clatsop Co., Seaside; OR. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From leaf litters of hemlock, alder, cedar, big-leaf maple, sugar pine, and Douglas fir. micropthalmus Chandler, 1986b: 338, fig. 7. OR: Lane Co., H. J. Andrews Experimental Forest, Watershed 10; OR. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From old-growth fir and Douglas-fir leaf litters. PSELAPHIDAE 9 minimus Schuster and Grigarick, 1960: 295, fig. 41. CA: Mendocino Co., Ryan Creek; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From redwood and oak leaf litters. montanus Casey, 1887: 479. CA: Placer Co.; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. obtusus Schuster and Grigarick, 1960: 290, figs. 20, 37. CA: Siskiyou Co., 13 km. S. Dunsmuir; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From oak leaf litter; from moss.

orbatus Schuster and Grigarick, 1960: 297, fig. 43. CA: Marin Co., Fort Baker; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M.

pectinis Schuster and Grigarick, 1960: 298, fig. 45. OR: Coos Co., Bridge; OR. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From moss. schusteri Chandler, 1983: 222, fig. 4. CA: Butte Co., 6 mi. N. of Forest Ranch, Ponderosa Way; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: Primarily from oak leaf litters; also from other deciduous leaf litters.

sinifundus Schuster and Grigarick, 1960: 279, fig. 25. CA: Humboldt Co., Freshwater; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From redwood leaf litter.

striatus LeConte, 1874: 49 {Trichonyx). BC: Vancouver Isl.; BC WA OR/ CA/ MT. TYPE DEPOSITORY: MCZC. REDESCRIPTION: Schuster and Grigarick, 1960. HOST: From leaf litters of Sitka spruce, Douglas fir, maple, and alder; from moss.

testaceus Casey, 1908: 274. CA: Marin, Alameda and Santa Cruz Cos.; CA. TYPE DEPOSITORY: USNM. REDESCRIPTION: Schuster and Grigarick, 1960. HOST: From leaf litters of redwood, oak, madrone, laurel, and sword fern.

tuberculatus Schuster and Grigarick, 1960: 289, figs. 19, 35. CA: Mariposa Co., 22 km. E. of Briceburg; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. umbraticus Schuster and Grigarick, 1960: 281, figs. 1, 26. CA: Siskiyou Co., 13 km. S. of Dunsmuir; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From leaf litters of redwood, alder, and Douglas fir; from moss.

vellosus Schuster and Grigarick, 1960: 283, figs. 7, 14, 29. CA: Sonoma Co., Petrified Forest; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From redwood and oak leaf litters. 10 COLEÓPTERA CATALOG

verrucifundus Schuster and Grigarick, 1960: 297, fig. 44. CA: Tulare Co., Ash Mt., Sequoia National Park; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M.

Genus EUBOARHEXIUS Grigarick aed Schuster

Euboarhexius Grigarick and Schuster, 1966: 31 (as genus). Type-species: Euboarhexius sinus Grigarick and Schuster (orig. des.). TAXONOMY: Carlton and Allen, 1986 (revision). REDESCRIPTION: Grigarick and Schuster, 1980; Carlton and Allen, 1986. KEYS: Carlton and Allen, 1986. dybasi Carlton and Allen, 1986: 294, figs. 5, 9, 11. WA: Pend Oreille Co., Roundtop Mountain, Kaniska National Forest near Metaline Falls; OR WA. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From litter on mossy bog margin. perscitus Fletcher, 1932: 29 (Rhexidius). NC: Buncombe Co., Black Mt.; NC SC GA TN/ VA. TYPE DEPOSITORY: AMNH. SEX OF TYPE: M. REDESCRIPTION: Carlton and Allen, 1986. HOST: From Rhododendron and mixed leaf litters. sinus Grigarick and Schuster, 1966: 31, figs. 1-4. CA: Mendocino Co., Little River; CA/ OR. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1980; Carlton and Allen, 1986. HOST: From Douglas-fir and vine maple leaf litters; from Douglas-fir logs. trogasteroides Brendel, 1892a: 12 (Rhexidius). PA: Westmoreland Co., Youngstown, Chestnut Ridge; PA MD WV VA/ NC GA TN/ KY. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. intermedius Brendel, 1892a: 13 {Rhexidius) (synonymized by Chandler, 1994). VA (PA erroneously listed as type locality in original description, see Chandler, 1994). TYPE DEPOSITORY: ICCM. SEX OF TYPE: F. TAXONOMY: Chandler, 1994. REDESCRIPTION: Brendel, 1893; Carlton and Allen, 1986. HOST: From Rhododendron and oak leaf litters.

Genus BONTOMTES Grigarick and Schuster

Bontomtes Grigarick and Schuster, 1980: 24 (as genus). Type-species: Bontomtes ripariae Grigarick and Schuster (orig. des.). ripariae Grigarick and Schuster, 1980: 26, pi. 27, figs. 1-9. OR: Linn Co., Tombstone Prairie, east of Santiam Pass; OR. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: Under moss on log.

Genus TETRASCAPHA Schuster and Marsh

Tetrascapha Schuster and Marsh, 1957: 149 (as genus). Type-species: Tetrascapha dasycera Schuster and Marsh (orig. des.). REDESCRIPTION: Grigarick and Schuster, 1980. PSELAPHIDAE ^ j dasycerca Schuster and Marsh, 1957: 149, figs. 1-7. CA: Mendocino Co., Mendocino: CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1980. HOST: From alder and redwood leaf litters. dentata Park and Wagner, 1962: 12, pi. II. fig. 1-2. OR: Tillamook Co., Little Nestucca River ORBC. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: In debris from Oxalis ferns; from leaf litters of hemlock, Douglas fir, and alder; from moss on logs. krysiae Chandler, 1983: 223, fig. 6. CA: Butte Co., 2 mi. E. of Jonesville, Scotts John Creek; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From alder leaf litter by springs or in snow melt areas. Genus FOVEOSCAPHA Park and Wagner Foveoscapha Park and Wagner, 1962: 13 (as genus). Type-species: Foveoscapha terracola ?2irk and Wagner (orig. des.). REDESCRIPTION: Grigarick and Schuster, 1980. terracola Park and Wagner, 1962: 13, pi. II, fig. 3. WA: Clallam Co., Olympic National Park, Olympic Hot Springs; WA BC OR. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1980. HOST: From leaf litters of maple, alder, hemlock, and fern. Subtribe RHEXIINA Park, 1951 Genus RHEXIUS LeConte Rhexius LeConte, 1849: 102 (as genus) (other subgenus Neotropical). Type-species: Rhexius insculptus LeConte (monot.). TAXONOMY: Chandler, 1990a (revision). REDESCRIPTION: Grigarick and Schuster, 1980. KEYS: Casey, 1908; Chandler, 1990a.

Subgenus RHEXIUS LeConte ferrugineus Casey, 1908: 280. PA: Westmoreland Co., St. Vincent; PA MD DC VA/ IN OH/ NC GA AL MS FL LA/ OK. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. rwèer Casey, 1908: 279 (synonymized by Chandler, 1990a). DC. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. transversus Csisey, 1908: 281 (synonymized by Chandler, 1990a). IN? TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Chandler, 1990a. HOST: From grass roots; litter at edge cypress marsh; at UV light. insculptus LeConte, 1849: 103. LA: Orleans Parish, New Orleans; IL OH MO/ OK/ LA MS FL GA. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. REDESCRIPTION: Casey, 1908; Chandler, 1990a. ECOLOGY: Reichle, 1969; Park, Auerbach, and Wilson, 1949 and 1953. HOST: From hardwood leaf litter, rotten wood, and sphagnum moss; in flood debris; at UV light. 12 COLEÓPTERA CATALOG ouachita Chandler, 1990a: 510, fig. 8. OK: Latimer Co.; OK. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From forest litter; in flood debris. schmitti Brendel, 1893: 281, pi. IV, fig. 20. PA; OH KY IN MO/ PA MD DC VA WV/ NC GA FL AL MS AR/ OK. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. hirsutus Casey, 1908: 279 (synonymized by Chandler, 1990a). MO: St. Louis Co., St. Louis. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Casey, 1908; Chandler, 1990a. HOST: From rotten wood and leaf litter; at UV light. simplex Motschulsky, 1856: 7. AL: Mobile Co., Mobile; AL. TYPE DEPOSITORY: ZMUM. TAXONOMY: Placed as Incertae sedis by Chandler, 1990a. stephani Chandler, 1990a: 507, fig. 2. OK: Latimer Co.; OK TX/ IL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From grass roots; in flood debris; at UV light. substriatus LeConte, 1878a: 383. FL: Hillsborough Co., Tampa; FL AL MS NC/ VA MD. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. virginicus Casey, 1908: 280 (synonymized by Chandler, 1990a). VA: Norfolk. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Chandler, 1990a. HOST: From leaf litter of sand pine, oak, and mixed hardwood; at UV light. Subtribe TRISIGNINA Park and Schuster, 1955 Genus TRISIGNIS Park and Schuster

Trisignis Park and Schuster, 1955: 1 (as genus). Type-species: Trisignis marshi Park and Schuster (orig. des.). REDESCRIPTION: Grigarick and Schuster, 1980. helferi Park and Schuster, 1955: 5, figs. 7, 8. CA: Mendocino Co., near Caspar; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1980. HOST: From redwood and Lithocarpus densiflora leaf litter. marshi Park and Schuster, 1955: 2, figs. 1-6. CA: Humboldt Co., near Freshwater; CA/ OR. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From redwood and Douglas-fir litter. Tribe EUPLECTINI LeConte, 1861 Subtribe RHINOSCEFSINA Bowman, 1934 Genus RHINOSCEPSIS LeConte Rhinoscepsis LeConte, 1878a: 382 (as genus)(other subgenus Neotropical). Type-species: Rhinoscepsis bistriatus LeConte (monot.). Rhinosceptis (error) Reitter, 1882: 209. PSELAPHIDAE 13

Rhynosceptis (error) Raffray, 1890a: 97. Rhynoscepsis (error) Raffray, 1898b: 223. REDESCRIPTION: Grigarick and Schuster, 1980.

Subgenus RHINOSCEPSIS LeConte bistriatus LeConte, 1878a: 382. FL: Tampa and Enterprise; FL SC GA. TYPE DEPOSITORY: MCZC. REDESCRIPTION: Grigarick and Schuster, 1980. HOST: From litter of sandpine, oak, and palmetto; at UV light.

Genus MORIUS Casey

Morius Casey, 1894: 445 (as genus). Type-species: Morius occidens Casey (monot.). REDESCRIPTION: Grigarick and Schuster, 1980. occidens Casey, 1894: 446, pi. I, fig. 16. CA: Santa Cruz Co.; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. REDESCRIPTION: Schuster, 1959; Grigarick and Schuster, 1980. HOST: From leaf litters of redwood, madrone, and oak. Subtribe PANAPHANTINA Jeannel, 1950 Genus THESIUM Casey Thesium Casey, 1884b: 117 (as genus). Type-species: Euplectus cavifrons LeConte (des. by Bowman, 1934: 144). Apothinus Sharp, 1887: 35 (as genus) (synonymized by Brendel, 1891). Type-species: Apothinus impressifrons Sharp (des. by Lucas, 1920: 113). REDESCRIPTION: Grigarick and Schuster, 1980. cavifrons LeConte, 1863b: 28 (Euplectus). AL: Mobile Co., Mobile; ON/ lA OH/ PA/ ME NH/ LA AL MS FL GA/ OK TX. This name was listed by Motschulsky (1856: 7) without a description and is a nomen nudum; LeConte validated this name when he based his description in part on a specimen sent by Motschulsky, which was collected at Mobile, AL, rather than New Orleans, LA, as stated by LeConte. TYPE DEPOSITORY: MCZC. laticolle Casey, 1884b: 118 (synonymized by Chandler, 1990a). TX: Colorado Co., Columbus. TYPE DEPOSITORY: USNM. REDESCRIPTION: Park, Auerbach and Corley, 1950; Grigarick and Schuster, 1980. HOST: From tree holes and rotten wood, less common in leaf litter; at UV light (Chandler, 1990a). Subtribe BIBLOPORINA Park, 1951 Genus BIBLOPORUS Thomson Bibloporus Thomson, 1859: 53 (as genus). Type-species: Euplectus bicolor Denny (orig. des.). United States species were not assigned to subgenus by Löbl, 1961. Faliscus Casey, 1884b: 120 (as genus) (synonymized by Brendel, 1891: 6). Type- species: Faliscus bicanalis Casey (monot.). REDESCRIPTION: Grigarick and Schuster, 1980.

bicanalis Casey, 1884b: 120 (Faliscus), NY: Oneida Co., Trenton Falls; NY PA WV/ PQ/ ME NH. TYPE DEPOSITORY: USNM. REDESCRIPTION: Grigarick and Schuster. ECOLOGY: Chandler, 1987. HOST: From white pine leaf litter. 14 COLEÓPTERA CATALOG

Genus EUTYPHLUS LeCoute

Eutyphlus LeConte, 1880: 185 (as genus). Type-species: Eutyphlus similis LeConte (monot.). Nicotheus Casey, 1884b: 121 (as genus) (synonymized by Horn, 1885). Type-species: Nicotheus tibialis Casey (monot.) (=similis (LeConte)). Nicothaeus (Qrror) Raffray, 1890a: 102. REDESCRIPTION: Grigarick and Schuster, 1980. KEYS: Park, 1956.

Subgenus EUTYPHLUS LeConte dybasi Park, 1956: 58. TN: Sevier Co., Great Smoky Mountain National Park; TN NC. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From leaf litters of oak, alder, and Rhododendron. prominens Casey, 1894: 460. VA: Lee Co.; MD VA NY WV/ TN NC GA SC. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. REDESCRIPTION: Park, 1956. HOST: From Rhododendron leaf litter. schmitti Raffray, 1904a: 546, fig. 9. PA: Westmoreland Co., St. Vincent; OH/ PA NJ NY/ NH. TYPE DEPOSITORY: MNHP. SEX OF TYPE: M. ECOLOGY: Chandler, 1987. HOST: From birch/beech and Rhododendron leaf litters; in Sphagnum moss. similis LeConte, 1880: 186. DC; OH IL/ NY DC PA MD WV VA/ TN NC GA AL. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. tibialis Casey, 1884b: 121 (Nicotheus) (synonymized by Horn, 1885). DC, VA: Lee Co. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1980. HOST: From various hardwood leaf litters and rotten logs.

Subgenus PLANITYPHLUS Park

Planityphlus Park, 1956: 62 (as subgenus). Type-species: Eutyphlus thoracicus Park (orig. des.). KEYS: Park, 1956. thoracicus Park, 1956: 60. TN: Sevier Co., Smoky Mountain National Park; TN SC NC. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From spruce-fir and Rhododendron leaf litters.

Genus ABDIUNGUIS Park and Wagner

Abdiunguis Park and Wagner, 1962: 10 (as genus). Type-species: Abdiunguis fenderi Park and Wagner (orig. des.). REDESCRIPTION: Grigarick and Schuster, 1980. fenderi Park and Wagner, 1962: 11, pi. I, figs. 9, 10. OR: Yamhill Co., Peavine Ridge, near McMinnville; OR WA. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1980. HOST: From moss in spring areas; in flood debris. PSELAPHIDAE 15

Genus HATCHIA Park and Wagner

Hatchia Park and Wagner, 1962: 9 (as genus). Type-species: Hatchia carinata Park and Wagner (orig. des.). REDESCRIPTION: Grigarick and Schuster, 1980. carinata Park and Wagner, 1962: 9. WA: King Co., Baring; WA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1980. Subtribe EUPLECTINA LeConte, 1861

Genus OROPODES Casey

Oropodes Casey, 1894: 453 (as genus). Type-species: Oropodes orbiceps Casey (monot.). TAXONOMY: Grigarick and Schuster, 1976 (revision). REDESCRIPTION: Grigarick and Schuster, 1976; Grigarick and Schuster, 1980. KEYS: Grigarick and Schuster, 1976. arcaps Grigarick and Schuster, 1976: 103, figs. 18-22. CA: Mendocino Co., Casper; CA/ WA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From redwood and fern leaf litter. dybasi Grigarick and Schuster, 1976: 101, figs. 11-17. OR: Benton Co., Well's Creek, 9 mi. SW. of Philomath; OR. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From Douglas-fir and big-leaf maple leaf litters. ishii Chandler, 1983: 224, fig. 7. CA: Butte Co., 5 mi. NE. of Forest Ranch; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From tanbark oak and other deciduous leaf litters. nuciere Grigarick and Schuster, 1976: 105, figs. 23, 24, 41. CA: Lake Co., Lucerne; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. orbiceps Casey, 1894: 453. CA: Los Angeles Co.; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. ra#ay/Brendel, 1894b; 196, fig. 5 (Euplectus) (synonymized by Raffray, 1904a). CA. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1976; Grigarick and Schuster, 1980. rumseyensis Grigarick and Schuster, 1976: 100, figs. 7-10. CA: Yolo Co., 3 mi. N. of Rumsey; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M.

Genus EUPLECTERGA Park and Wagner

Euplecterga Park and Wagner, 1962: 15 (as subgenus of Oropodes). Type-species: Oropodes impressicollis Park and Wagner (orig. des.). Euplecturga (error) Grigarick and Schuster, 1976: 107. TAXONOMY: Grigarick and Schuster, 1976 (revision). REDESCRIPTION: Grigarick and Schuster, 1976; Grigarick and Schuster, 1980. KEYS: Grigarick and Schuster, 1976. 16 COLEÓPTERA CATALOG fideli Grigarick and Schuster, 1976: 108, figs. 31-34, 40. CA: Santa Cruz Co., Sequel; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. impressicollis Park and Wagner, 1962: 15, pi. II, fig. 7 {Oropodes). OR: Washington Co., Forest Grove; OR. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. norstelcha Grigarick and Schuster, 1976: 109, figs. 35-39. OR: Coos Co., Charleston; OR. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1980. HOST: From Douglas-^- leaf litter.

Genus RAMECIA Casey

Ramecia Casey, 1894: 450 (as genus). Type-species: Euplectus crinitis Brendel (design, by Lucas, 1920: 564). Liniolis Grigarick and Schuster, 1980: 34 (synonymized by Newton and Chandler, 1989). Type species: Euplectus crinitis Brendel (orig. des.). TAXONOMY: Carlton and Chandler, 1994. REDESCRIPTION: Grigarick and Schuster, 1980 (as Liniolis). capitula Casey, 1884b: 112 {Euplectus). FL: Leon Co., Tallahassee: FL. TYPE DEPOSITORY: USNM. TAXONOMY: Carlton and Chandler, 1994. REDESCRIPTION: Park, 1962. crinita Brendel, 1865b: 260 {Euplectus). IL; MI PQ/ NS/ ME NH/ lA IL IN OH /NY PA VA/ NC. TYPE DEPOSITORY: ANSP. TAXONOMY: Moxey, 1962. ECOLOGY: Chandler, 1987. REDESCRIPTION: Park, 1962; Grigarick and Schuster, 1980. HOST: From leaf litter; under bark of maple and oak logs.

Genus PSEUDACTIUM Casey

Pseudactium Casey, 1908: 271 (as genus). Type-species: Pseudactium carolinae Casey (orig. des). Ramecia (of authors, not Casey, 1894). Racemia Newton and Chandler, 1989: 7 (as genus) (synonymized by Newton and Chandler, 1989). Type species: Euplectus arcuatus LeConte (orig. des.). TAXONOMY: Cariton and Chandler, 1994. REDESCRIPTION: Grigarick and Schuster, 1971 (as Ramecia)', Grigarick and Schuster, 1980 (as Ramecia); Carlton and Chandler, 1994. KEYS: Carlton and Chandler, 1994. atexanderi Carlton and Chandler, 1994: 181, figs. 26-29. AL: Winston Co., Natural Bridge Cave; AL TN. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. arcuatum LeConte, 1849: 106 {Euplectus). GA: Clarke Co., Athens; TN AL SC GA FL. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. REDESCRIPTION: Park, 1962; Grigarick and Schuster, 1980; Carlton and Chandler, 1994. HOST: From litter from an oak^eech mixture. PSELAPHIDAE 17 carolinae Casey, 1908: 271. NC; NC FL /TX. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. REDESCRIPTION: Carlton and Chandler, 1994. HOST: From litter of sand pine, oak, and palmetto; from magnolia leaf litter. copelandi Carlton and Chandler, 1994: 182, figs. 30-33. AL: Tuscaloosa Co., Northport, Lurleen State Park; AL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. magazinensis Carlton and Chandler, 1994: 182, figs. 34-39. AR: Logan Co., Mount Magazine, 1.4 km. W. of Greenfield Picnic Area; AR. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From a mixture of beech-maple litter. mendicum Park, 1962: 155, figs. 7, 10 (Ramecia). AL: Jackson Co., Pisgah, near Saltpeter Cave; AL TN FL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Carlton and Chandler, 1994. HOST: From forest litter; from rotten yellow poplar log; from sand pine and palmetto litter. paraboiicum Brendel, 1893: 281, pi. IV, fig. 7 {Trimioplectus), PA: Westmoreland Co. (designated by Carlton and Chandler, 1994); PA VA /KY /TN NC. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. dentiventris Casey, 1894: 452 {Ramecia) (synonymized by Carlton and Chandler, 1994). VA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. discreta CsiSQy, 1894: 451 (Ramecia) (synonymized by Carlton and Chandler, 1994). PA. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. cephalicum Casey, 1908: 273 (synonymized by Carlton and Chandler, 1994). PA: Westmoreland Co. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. REDESCRIPTION: Carlton and Chandler, 1994. HOST: From deciduous forest litter. mellineum Casey, 1908: 272 (synonymized by Carlton and Chandler, 1994). PA: Westmoreland Co. TYPE DEPOSITORY: USNM. SEX OF TYPE: M.

pecki Carlton and Chandler, 1994: 184, figs. 40-43. AL: Jackson Co., 5 mi. N. of Garth; AL. TYPE DEPOSITORY: CNCI. SEX OF TYPE: M.

stannardi Carlton and Chandler, 1994: 187, fig. 49. lA: Woodbury Co., Sioux City, Stone State Park; lA. TYPE DEPOSITORY: INHS. SEX OF TYPE: M. HOST: From Yucca glauca.

steevesi Carlton and Chandler, 1994: 186, figs. 44-48. AL: Blount Co., near Wildcat Cave; AL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. 18 COLEÓPTERA CATALOG

ursum Carlton, 1995: 170, figs. 1-3. AR: Newton Co., Buffalo National River, 3 km. NW. of Erbie Campground; AR. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M.

Genus SAXET Grigarick and Schuster

Saxet Grigarick and Schuster, 1980: 37, pi. 50, figs. 1-9 (as genus). Type-species: Euplectus decoris Casey (orig. des.). aravaipa Chandler, 1985: 119, fig. 6, a-d. AZ: Graham Co., Aravaipa Canyon; AZ. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From oak leaf litter. decora Casey, 1884b: 113 {Euplectus), TX: Colorado Co., Columbus; TX OK/ AL. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. ^REDESCRIPTION: Park, 1962; Grigarick and Schuster, 1980. yavapai Chandler, 1985: 119, fig. 7, a-e. AZ: Yavapai Co., Kirkland; AZ. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: At UV light.

Genus THESIASTES Casey

Thesiastes Casey, 1894: 457 (as genus). Type-species: Euplectus debilis LeConte (design, by Bowman, 1934: 144). TAXONOMY: Chandler, 1990a (revision). REDESCRIPTION: Grigarick and Schuster, 1980. KEYS: Chandler, 1990a. atratus Casey, 1894: 457. RI; RI/ NJ/ OK TX/ SC FL. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: From grass clumps and tree holes; in sphagnum moss; at UV light. debilis LeConte, 1878a: 386 {Euplectus). FL: Hillsborough Co., Tampa; NC FL GA AL MS LA/ KY lA IL IN OH. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. tenuis LeConte, 1878a: 386 {Euplectus) (synonymized by Casey, 1884b). FL: Capron. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. planipennis Brendel, 1889: 195 {Euplectus) (synonymized by Chandler, 1990a). lA: Linn Co. TYPE DEPOSITORY: ANSP. SEX OF TYPE: F. TAXONOMY: Moxey, 1962. REDESCRIPTION: Chandler, 1990a. HOST: At UV light.

fossulatus Brendel, 1891: 59, pi. XII, fig. 124 {Euplectus). IL: Peoria Co., Peoria; lA IL IN OH KY/ NC AL MS FL/ OK TX. TYPE DEPOSITORY: ANSP. SEX OF TYPE: F. TAXONOMY: Moxey, 1962. PSELAPHIDAE 19

rotundicollis Brendel, 1891: 61, pis. XI, XII (Euplectus) (synonymized by Chandler, 1990a). lA: Linn Co. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Moxey, 1962. REDESCRIPTION: Chandler, 1990a. HOST: From rotten stumps and tree holes; at UV light. pumilis LeConte, 1849: 106 (Euplectus). SC: Charleston Co., Charleston; LA MS GA SC/ OK TX. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. REDESCRIPTION: Chandler, 1990a. HOST: From oak tree holes, and rotten wood.

Genus ACOLONIA Casey

Acolonia Casey, 1894: 454 (as genus). Type-species: Euplectus cavicollis LeConte (monot.). TAXONOMY: Wagner, 1975 (revision). REDESCRIPTION: Wagner, 1975; Grigarick and Schuster, 1980. KEYS: Wagner, 1975. cavicollis LeConte, 1878a: 387 {Euplectus). FL: Hillsborough Co., Tampa; FL. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. REDESCRIPTION: Wagner, 1975; Grigarick and Schuster, 1980. HOST: Under pine bark.

Genus EUPLECTUS Leach

Euplectus Leach, 1817: 80. Type-species: Euplectus reichenbachii Leach (= Pselaphus nanus Reichenbach). Only introduced European species have been assigned to subgenera. TAXONOMY: Wagner, 1975 (Nearctic species). REDESCRIPTION: Wagner, 1975; Grigarick and Schuster, 1980. ECOLOGY: Wagner, 1975; Engelmann, 1956. KEYS: Wagner, 1975; Park and Wagner, 1962 (Pacific Northwest spp.). acomanus Casey, 1908: 265. NM: Otero Co., Cloudcroft; IN/ MD PA/ AZ NM OK/ MA/ PQ/ MS. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Wagner, 1975. HOST: Under pine bark; from rotten pine and fir logs (Chandler, 1985); in oak tree holes; in old sawdust. californicus Casey, 1887: 476. CA: Placer Co., Lake Tahoe; CA. TYPE DEPOSITORY: USNM. REDESCRIPTION: Wagner, 1975. HOST: Under bark. confluens LeConte, 1849: 105. MA: Middlesex Co., Cambridge; ID/ WI MI PQ/ NH MA ME/ MD PA DC DE NY WV VA/ lA IL IN OH KY/ NC SC FL AR AL TN GA MS/ AZOK. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. IMMATURE STAGES: Wagner, 1975 (larva); Böving and Craighead, 1931. REDESCRIPTION: Wagner, 1975; Grigarick and Schuster, 1980. ECOLOGY: Wagner, 1975; Park, Auerbach, and Corley, 1950. HOST: From various deciduous rotten woods and material in tree holes; in sawdust; in barn debris; at UV light.

duryi Casey, 1908: 266. OH: Hamilton Co., Cincinnati; ID/ WI PQ ON/ CO/ lA IL OH KY/ BC/ ME CT NH/ NS/ VA PA MD DC/ AZ/ TN GA NC. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. 20 COLEÓPTERA CATALOG

barri Park and Wagner, 1962: 17, pi. Ill, fig. 7 (synonymized by Wagner, 1975). ID: Latah Co., Moscow Mt. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. ECOLOGY: Chandler, 1987. REDESCRIPTION: Wagner, 1975. HOST: Berlesed primarily from rotten wood; also from sawdust and leaf litter. elongatus Brendel, 1893: 281. PA: Westmoreland Co., Youngstown, Chestnut Ridge; IL/ PA MD DC/ CT/ ON. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Moxey, 1962. REDESCRIPTION: Wagner, 1975. HOST: Berlesed from leaf litter and under bark of rotten log. filiformis Casey, 1908: 267 {Leptoplectus), PA: Westmoreland Co., Latrobe, St. Vincent; IN OH/ PA VA/ NC. TYPE DEPOSITORY: USNM. REDESCRIPTION: Wagner, 1975. HOST: Berlesed from leaf litter and sawdust.

¡dahoensis Park and Wagner, 1962: 17, pi. Ill, fig. 9. ID: Latah Co., Moscow Mt.; ID/ MT. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. REDESCRIPTION: Wagner, 1975. HOST: In Salix leaf litter. longicoUis Casey, 1884b: 99. MD: Garrett Co., Deer Park; ON/ IL IN OH KY/ PA MD DC VA/ AL TN se NC. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Wagner, 1975. HOST: Berlesed from rotten wood and material in tree holes. silvicolus Chandler, 1986a: 121, figs. 1-3. NH: Carroll Co., The Bowl, 2.5 mi. NW. of Wonalancet; NH/ NS/ OR. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. ECOLOGY: Chandler, 1987. HOST: From conifer logs in old-growth forests.

Subgenus DIPLECTELLUS Reitter

Diplectellus Reitter, 1909: 207 (as subgenus of Euplectus). Type-species: Euplectus afer Reitter (design, by Jeannel, 1950: 100). karsteni Reichenbach, 1816: 71 {Pselaphus) (introduced into North America). Germany: "Rhodopole," Berlin, and Leipzig; MN WI MI ON PQ/ lA IL/NY PA DC/ ME NH MA/ PE/ MT SK. TYPE DEPOSITORY: Types lost. iowensis Casey, 1894: 456 (synonymized by Besuchet, 1974). lA: Linn Co., Cedar Rapids. The type-locality fixed by Wagner, 1975. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Wagner, 1975. HOST: From compost, material in tree holes, and lawn clippings; at UV light. signatus Reichenbach, 1816: 73 {Pselaphus) (introduced into North America). Germany: Berlin; WI ON PQ/ IL OH/ VT ME/ MB SK. TYPE DEPOSITORY: Types lost. REDESCRIPTION: Wagner, 1975. HOST: From compost and grass cuttings; also in spilled grain. PSELAPHIDAE 21

Unplaced Species disjunctus Casey, 1897: 554. VA: Lee Co.; VA. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. TAXONOMY: Wagner, 1975: 129 placed as nomen dubium.

schmitti Raffray, 1904a: 540. PA; PA. TYPE DEPOSITORY: MNHP. SEX OF TYPE: M. TAXONOMY: Wagner, 1975:129 placed as nomen dubium.

Genus PYCNOPLECTUS Casey

Pycnoplectus Casey, 1897: 552 (as genus). Type-species: Euplectus linearis LeConte (des. by Wagner, 1975: 158). TAXONOMY: Wagner, 1975 (revision). REDESCRIPTION: Wagner, 1975; Grigarick and Schuster, 1980. KEYS: Wagner, 1975.

cediosus Wagner, 1975: 167, figs. 41, 48. OH: Hamilton Co., Cincinnati; OH KY. TYPE DEPOSITORY: CMNH. SEX OF TYPE: M. HOST: From rotten wood.

congener Casey, 1884b: 101 {Euplectus). DC; MI ON/ IL OH/ PA DC/ CT. TYPE DEPOSITORY: USNM. tenellus Casey, 1897: 552 (synonymized by Wagner, 1975). PA: Westmoreland Co., St. Vincent. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Wagner, 1975. HOST: From rotten wood.

difficilis LeConte, 1849: 106 {Euplectus). GA: Clarke Co., Athens; GA SC FL. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. longissimusBxQnau, 1891: 71, pis. XI, XII {Euplectus) (synonymized by Wagner, 1975). GA. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Chandler, 1994. REDESCRIPTION: Wagner, 1975. HOST: Berlesed from material from tree holes; from sand pine and palmetto litter.

falcatus Wagner, 1975: 179, fig. 60. VA: Washington Co., Abingdon; VA. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From rotten wood.

infossus Raffray, 1904a: 541, fig. 6 {Euplectus). PA; KY/ PA WV/ TN GA. TYPE DEPOSITORY: MNHP. SEX OF TYPE: M. excavatus Bowman, 1934: 141 {Euplectus) (synonymized by Wagner, 1975). PA: Franklin CO., Penn-Mar. TYPE DEPOSITORY: ICCM. SEX OF TYPE: M. REDESCRIPTION: Wagner, 1975. HOST: From rotten wood, sawdust, and material from tree holes. 22 COLEÓPTERA CATALOG interruptus LeConte, 1849: 105 {Euplectus) GA: Clarke Co., Athens; MN WI MI ON/ lA MO IL IN OH/ NY PA DC VA WV/ ME/ AL GA FL MS/ OK/ KY. TYPE DEPOSITORY: MCZC. hudsonicus Casey, 1894; 455 {Euplectus) (synonymized by Wagner, 1975). NY: Kings Co., Brooklyn. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Wagner, 1975. HOST: From rotten wood and material from tree holes. linearis LeConte, 1849: 104 {Euplectus). LA; MI ON/ IL IN OH KY MO/ NY PA NJ MD DC/ TX OK/ LA GA FL. TYPE DEPOSITORY: MCZC. impressiceps CdiSQy, 1908: 265 (synonymized by Wagner, 1975). PA: Westmoreland Co. TYPE DEPOSITORY: Lost? Not in USNM? (Wagner 1975: 161). REDESCRIPTION: Wagner, 1975; Grigarick and Schuster, 1980. HOST: From rotten wood, sawdust, material from tree holes, and under bark. longipennis Casey, 1908: 264. PA: Westmoreland Co., St. Vincent; KY/ PA MD DC VA WV. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. REDESCRIPTION: Wagner, 1975. HOST: From rotten pine logs and sawdust. parki Wagner, 1975: 168, figs. 44, 49. IL: Adams Co., Quincy; IL OH/ NC LA/ MD. TYPE DEPOSITORY: INHS. SEX OF TYPE: M. sexualis Casey, 1884b: 103 {Euplectus). SC; MS AL SC FL/ MD. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Wagner, 1975. HOST: From sawdust and tree-hole litter. spinifer Casey, 1884b: 104 {Euplectus). TX: Colorado Co., Columbus; lA IL IN KY/ MD DC/ TX OK/ AR LA MS AL TN FL. TYPE DEPOSITORY: USNM. REDESCRIPTION: Wagner, 1975. HOST: Berlesed from hardwood rotten logs and sawdust; from willow tree-hole debris.

Unplaced Species floridae Casey, 1908: 264. FL; FL. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. TAXONOMY: Wagner, 1975: 129 placed as nomen dubium. imperfectus Casey, 1897: 553. DC; DC. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. TAXONOMY: Wagner, 1975: 129 placed as nomen dubium.

Genus LEPTOPLECTUS Casey

Leptoplectus Casey, 1908: 266 (as genus). Type-species: Euplectus pertenuis Casey (design, by Lucas, 1920: 371). TAXONOMY: Wagner, 1975 (revision). REDESCRIPTION: Wagner, 1975; Grigarick and Schuster, 1980. KEYS: Wagner, 1975. PSELAPHIDAE 23 pertenuis Casey, 1884b: 109 (Euplectus). DC; lA IL IN OH KY/ PA NJ DE MD DC VA/ RI CT/ MS AR LA AL TN GA SC NC FL. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. exilissimus Casey, 1908: 269 (synonymized by Wagner, 1975). OH: Hamilton Co., Cincinnati. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. insolens Casey, 1908: 268 (synonymized by Wagner, 1975). PA: Westmoreland Co., St. Vincent. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Wagner, 1975; Grigarick and Schuster, 1980. HOST: Primarily from rotten wood, also from leaf litter, sawdust, and material from tree holes; from eastern hemlock litter; at UV light.

Genus TRIMIOPLECTUS Brendel

Trimioplectus Brendel, 1890: 226 (as genus). Type-species: Trimioplectus obsoletus Brendel (design, by Lucas, 1920: 655). TAXONOMY: Casey, 1894: 459; Chandler, 1990a. REDESCRIPTION: Grigarick and Schuster, 1980. KEYS: Chandler, 1990a.

auerbachi Park, 1949: 319, pi. II, figs. 1, 3. MI: Allegan Co., Glenn; MI/ IL OH/ PA DC. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From sugar maple, white oak, American elm, and silver maple tree holes.

australis Chandler, 1990a: 515, figs. 14-15. OK: Latimer Co.; OK/ AR FL/ KY. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M HOST: From oak tree holes.

obsoletus Brendel, 1891: 50, pis. XI, XII. lA: Linn Co., Cedar Rapids; lA IL OH/ ON PQ/ NH MA/ PA VA/ NC. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Chandler, 1994. ECOLOGY: Chandler, 1987. REDESCRIPTION: Park, 1949; Grigarick and Schuster, 1980. HOST: From rotten wood and tree holes. Subtribe BIBLOPLECTINA Jeannel 1959

Genus BIBLOPLECTUS Reitter

Bibloplectus Reitter, 1881: 529 (as subgenus). Type-species: Pselaphus ambiguus Reichenbach (des. by Lucas, 1920: 137). TAXONOMY: Chandler, 1990a (revision). Jeannel (1949) placed all North American species m the subgenus Bibloplectodes based on Euplectus ruficeps Motschulsky. However, the accompanying figure of this species was oí Bibloplectus choctaw Chandler. Species described by Chandler (1990a) were not placed in a subgenus. REDESCRIPTION: Grigarick and Schuster, 1980. ECOLOGY: Engelmann, 1956. KEYS: Casey, 1894; Chandler, 1990a.

Subgenus BIBLOPLECTODES Jeannel

Bibloplectodes Jeannel, 1949: 49 (as subgenus). Type-species: Euplectus ruficeps LeConte (orig. des.)- 24 COLEÓPTERA CATALOG integer LeConte, 1878a: 386 (Euplectus). MI: Wayne Co., Detroit; MI ON PQ/ IL IN OH/ PA NY NJ/ NB NS/ ME NH VT/ MB. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. REDESCRIPTION: Park, Auerbach, and Wilson, 1953; Chandler, 1990a. ECOLOGY: Park, Auerbach and Wilson, 1949 and 1953; Chandler, 1987. HOST: In conifer and hardwood leaf litters; in sphagnum moss. leviceps Casey, 1884b: 115 {Euplectvs). DC; MD DC NJ VA/ MA/ SC FL AL. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. REDESCRIPTION: Chandler, 1990a. HOST: From hardwood leaf litter; from Sphagnum moss. ruficeps Motschulsky, 1856:7. AL: Mobile Co., Mobile; LA MS AL FL GA SC AR/ NJ PA/ CT/ OH lA/ ON/ OK TX. This species has been attributed to LeConte (1863b: 28); however LeConte received his specimen from Motschulsky and it is a syntype. Motschulsky's description indicates that the types are from Mobile, AL, and not New Orleans, LA, as LeConte stated. TYPE DEPOSITORY: MCZC, ZMUM. SEX OF TYPE: F. REDESCRIPTION: Chandler, 1990a. HOST: From leaf litters, sawdust, grass clumps, old hay, and moss (Chandler, 1990a). sobrinus Casey, 1897: 555. RI: Boston Neck; RI CT. TYPE DEPOSITORY: USNM. HOST: From leaf litter. REDESCRIPTION: Grigarick and Schuster, 1980; Chandler, 1990a.

Unplaced Species

Cherokee Chandler, 1990a: 519, figs. 25-26. OK: Latimer Co; OK. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From flood debris. chickasaw Chandler, 1990a: 521, figs. 35-36. OK: Latimer Co.; OK/ AR/ IL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From grass clumps, and litter from oak/pine mixture. choctaw Chandler, 1990a: 517, figs. 21-22. OK: Latimer Co., 5 mi. W. of Red Oak; OK TX/ MO. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From oak logs; at UV light. creek Chandler, 1990a: 519, figs. 23-24. OK: Latimer Co.; OK/ MO KY. TYPE DEPOSITORY: FMNH SEX OF TYPE: M. HOST: From flood debris; at UV light. exilis Bowman, 1934: 141. PA: Allegheny Co., Imperial; PA. TYPE DEPOSITORY: Not in ICCM (lost?). TAXONOMY: Placed as incertae sedis by Chandler, 1990a. osage Chandler, 1990a: 521, figs. 33-34. OK: Latimer Co.; OK. TYPE DEPOSITORY: FMNH. SEX OF TYPE: F. HOST: From leaf litter and flood debris. PSELAPHIDAE 25

Genus TRIGONOPLECTUS Bowman

Trigonoplectus Bowman, 1934: 37 (as genus). Type-species: Trigonoplectus minutus Bowman (orig. des.). Bowman (1934) designated the type-species for this genus on p. 144. REDESCRIPTION: Grigarick and Schuster, 1980. KEYS: Bowman, 1934. minutus Bowman, 1934: 38. PA: Allegheny Co., near Clinton; PA. TYPE DEPOSITORY: ICCM. SEX OF TYPE: F. REDESCRIPTION: Grigarick and Schuster, 1980. rostratus Bowman, 1934: 38. PA: Allegheny Co., near Clinton: PA. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. Subtribe TRIMIINA Bowman, 1934

Genus ACTIZONA Chandler

Actizona Chandler, 1985: 112 (as genus). Type species: Actizona chuskae Chandler (orig. des.). chuskae Chandler, 1985: 113, fig. 2, a-i. AZ: Apache Co., Chuska Mountains, Wagon Wheel Forest Campground; CO/ AZ. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. REDESCRIPTION: Carlton, 1989a (aedeagal isomerism). trifoveatum Park, 1963: 179, figs. 2, 5-6 (Actium). MI: Delta Co., 13 mi. N. of Nahma Junction; ME NH/ PQ MI/ MB MT AB/ WY/ BC. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. borealis Chandler, 1986a: 123, fig. 4 (synonymized by Chandler, 1993). NH: Coos Co., Jefferson Notch. ECOLOGY: Chandler, 1987. REDESCRIPTION: Grigarick and Schuster, 1971. HOST: From rotten beech or birch logs, or beech tree-hole material (Chandler, 1993).

Genus ACTIUM Casey

Actium Casey, 1886: 201. Type-species: Actium pallidum Casey (design, by Brendel, 1891: 40). Brendel, 1891: 40 designated Actium pallidum Casey as type of genus, which is a junior subjective synonym of Trimium californicum LeConte. Proplectus Raffray, 1890a: 107 (as genus) (synonymized by Casey, 1894). Type-species: Proplectus decipiens Raffray (subseq. monot. Raffray, 1890b: 197) = californicum (LeConte). The type of the genus, Proplectus decipiens, is a junior subjective synonym of Actium californicum (LeConte), type of the genus Actium. Basolum Casey, 1897: 571 (as genus) (synonymized by Chandler, 1994). Type-species: Trimium impunctatum Brendel (orig. des.). TAXONOMY: Grigarick and Schuster, 1971 (revision). REDESCRIPTION: Grigarick and Schuster, 1971; Grigarick and Schuster, 1980. KEYS: Grigarick and Schuster, 1971. aculeatum Grigarick and Schuster, 1971: 23, figs. 126-132. CA: Fresno Co., 10 mi. NW. of Auberry; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. 26 COLEÓPTERA CATALOG arundineum Grigarick and Schuster, 1971: 5, figs. 8-11. CA: Mariposa Co., Wawona; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. barbatum Grigarick and Schuster, 1971: 17, figs. 100, 101. NM: San Miguel Co., Porvenir; NM. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. barri Park and Wagner, 1962: 19, pi. V, figs. 104. ID: Latah Co., Moscow, Moscow Mt.; BC WA OR ID/ AB MT. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: Most frequent in old-growth red fir and silver fir leaf litters; also from alder, cedar/hemlock, maple, and Douglas-fir leaf litters. bicolor Casey, 1908: 269. NM: Otero Co., Cloudcroft; NM AZ. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1971. HOST: Under ponderosa pine bark. bifoveatum Casey, 1897: 559. CA: Siskiyou Co.; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. REDESCRIPTION: Grigarick and Schuster, 1971. brevipenne Casey, 1894: 467. CA: Santa Cruz Co.; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1971. HOST: From redwood and Douglas-fir leaf litters; from rotten wood. calcaris Grigarick and Schuster, 1971: 11, figs. 57-61. CA: Tehama Co., Mineral; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From leaf litters of Douglas fir, big-leaf maple, live oak, and Rhododendron; from rotten logs. californicum LeConte, 1878a: 383 (Trimium). CA; CA. TYPE DEPOSITORY: MCZC. pallidum Casey, 1887: 473, pi. XVI, fig. 7 (synonymized by Casey, 1894). CA: Monterey Co. TYPE DEPOSITORY: USNM. decipiens Raffray, 1890b: 197 {Proplectus) (synonymized by Casey, 1894). CA: Napa Co., St. Helena. TYPE DEPOSITORY: MNHP. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1971; Grigarick and Schuster, 1980. HOST: Under bark of Quercus agrifolia; from poplar and grape litter near intermittent streams; from rotting vegetation along streams. candidum Casey, 1894: 465. CA: Santa Cruz Co.; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1971. chiloquinensis Grigarick and Schuster, 1971: 15, figs. 95-97. OR: Klamath Co., 1 mi. SW. of Chiloquin; OR. TYPE DEPOSITORY: OSUC. SEX OF TYPE: M. HOST: From manzanita leaf litter; from Douglas-fir logs. PSELAPHIDAE 27 cochísei Chandler, 1985: 111, fig. 1, a-d. AZ: Cochise Co., Cochise Stronghold, Dragoon Mountains; AZ. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: At UV light. falcatum Grigarick and Schuster, 1971: 20, figs. 110, 111. CA: Tuolumne Co., Leland Meadows; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: Under pine or fir bark; from mixed conifer litter. fastigiiim Grigarick and Schuster, 1971: 23, figs. 133-139. CA: Calaveras Co., 1.5 mi. N. of Murphys; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From oak and pine leaf litters. fastosum Grigarick and Schuster, 1971: 21, figs. 112-114. CA: Mariposa Co., Miami; OR/ CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: Under conifer bark; from white fir leaf litter. femineum Grigarick and Schuster, 1971: 25, figs. 142-148. CA: Trinity Co., 11 mi. E. of Douglas City; CA/ OR. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From leaf litters of oak, maple, Douglas fir, and sugar pine; under ponderosa pine bark; from moss on conifer log. formosum Grigarick and Schuster, 1971: 6, figs. 12-17. CA: Tulare Co., Balch Park; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From leaf litter of oak/pine mixture. functum Grigarick and Schuster, 1971: 12, figs. 62-68. CA: Mendocino Co., Mendocino; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. hazeltinei Chandler, 1983: 224, fig. 8. CA: Butte Co., 2 mi. E. of Jonesville, Scotts John Creek; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From leaf litters of alder, willow, and grass in snow melt areas. helferi Grigarick and Schuster, 1971: 8, figs. 38-42. OR: Klamath Co., Klamath Falls; OR. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. REDESCRIPTION: From mixed-conifer leaf litter. impunctatum Brendel, 1865b: 259 {Trimium). VA; OH/ PA MD/ NC/ VA WV MD VA. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. angustum Casey, 1897: 559 (synonymized by Chandler, 1994). PA: Westmoreland Co., St. Vincent. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. TAXONOMY: Chandler, 1994. REDESCRIPTION: Grigarick and Schuster, 1971 (as angustum). HOST: From rotten logs and material in tree holes; from leaf litters of white pine, Rhododendron, hemlock, oak, and maple. 28 COLEÓPTERA CATALOG marinicum Casey, 1894: 465. CA: Marin Co.; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 197L microphthalmum Park and Wagner, 1962: 19, pi. IV, fig. 5. OR: Yamhill Co., Sourgrass Creek; OR. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 197L HOST: Most common in leaf litter from mature Douglas-fir trees; also from rotten wood. ojaiensis Grigarick and Schuster, 1971: 22, figs. 119-125. CA: Ventura Co., Ojai; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. pacificum Casey, 1894: 466. CA: Siskiyou Co.; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1971. HOST: From forest litter. pandum Grigarick and Schuster, 1971: 13, figs. 75-80. CA: Madera Co., Sugar Pine; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From ponderosa pine litter. politum Casey, 1887: 473. CA: Mendocino Co.; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. REDESCRIPTION: Grigarick and Schuster, 1971. retractum Casey, 1908: 270. BC: Queen Charlotte Isls.; AK/ BC WA OR/ CA/ MT. TYPE DEPOSITORY: USNM. hatchi Park and Wagner, 1962: 20, pi. IV, fig. 9 (synonymized by Grigarick and Schuster, 1972). WA: Kittitas Co., Snoqualmie Pass. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1971. HOST: From leaf litters of redwood, Sitka spruce, and hemlock; from streamside moss. robustulum Casey, 1887: 474. CA: Mendocino Co., Anderson Valley; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1971. HOST: From pine leaf litter. rothi Grigarick and Schuster, 1971: 9, figs. 47-51. OR: Coos Co., Charleston; OR. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From leaf litters of Douglas fir, hemlock, maple, and alder; in moss. tempus Grigarick and Schuster, 1971: 14, figs. 86-89. CA: Eldorado Co., Fred's Place: CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. tenellum Grigarick and Schuster, 1971: 13, figs. 81-85. CA: Calaveras Co. 5.6 mi. E. of Dorrington; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From pine and fir litters. PSELAPHIDAE 29 tentum Grigarick and Schuster, 1971: 12, figs. 69-74. CA: Mendocino Co., 24 mi. SE. of Covelo; CA/ OR. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From leaf litters of Douglas fir, cedar, maple, Averts, and oak, particularly near streams or in wet areas. testaceum Casey, 1887: 475. CA: Mendocino Co., Anderson Valley; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1971. HOST: From redwood leaf litter. wawonaensis Grigarick and Schuster, 1971: 7, figs. 21-30. CA: Mariposa Co., Wawona; OR BC/ CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From leaf litters of Douglas fir, ponderosa pine, cedar, white fir, maple, alder, and redwood.

Genus ACTIASTES Casey

Actiastes Casey, 1897: 561 (as genus). Type-species: Trimium foveicolle LeConte (orig. des.). TAXONOMY: Grigarick and Schuster, 1971 (revision). REDESCRIPTION: Grigarick and Schuster, 1971; Grigarick and Schuster, 1980. KEYS: Grigarick and Schuster, 1971.

desertorum Grigarick and Schuster, 1971: 30, figs. 169-171. CA: Riverside Co., Deep Canyon; CA NV/ UT; Mex. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From manzanita leaf litter; in horse dung; at UV light.

foveicoUis LeConte, 1878a: 384 {Trimium). MA: Middlesex Co., Cambridge; BC/ PA NJ DE MD VA WV NY/ MA NH RI CT/ TN/ ON PQ. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. costale Brendel, 1892b: 166 {Trimium) (synonymized by Grigarick and Schuster, 1971). PA: Franklin Co. Grigarick and Schuster (1971) treated a specimen in ICCM as the holotype; however. Chandler (1994) placed the holotype at ANSP. TYPE DEPOSITORY: ANSP. SEX OF TYPE: F. pennsylvanicum Bowman, 1934: 141 {Actium) (synonymized by Grigarick and Schuster, 1971). PA: Franklin Co., Penn-Mar. TYPE DEPOSITORY: ICCM. SEX OF TYPE: M. ECOLOGY: Chandler, 1987. REDESCRIPTION: Grigarick and Schuster, 1971; Grigarick and Schuster, 1980. HOST: From hardwood leaf litters, often near water; from rotten wood.

fovicinus Grigarick and Schuster, 1971: 32, figs. 181-183. GA: Dade Co., Cloudland Canyon State Park; AL GA. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From forest leaf litter. fundatum Grigarick and Schuster, 1971: 33, figs. 195-197. TN: Blount Co., Smoky Mountain National Park, Sinks Canyon; TN NC. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From sycamore tree holes; from hardwood litter near logs. 30 COLEÓPTERA CATALOG globiferum LeConte, 1849: 107 {Euplectus). GA: White Co., Nacoochee (Nakutshi); PA MD VA DC/KY OH/ PQ/ MA/ NC SC GA AL MS TN/ OK. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. blandum Casey, 1908: 271 {Actium){synonyrmzQá by Grigarick and Schuster, 1971). PA: Westmoreland Co. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 197L HOST: Under pine bark; from pine and oak litter; in tree holes. spatium Grigarick and Schuster, 1971: 29, figs. 156-162. MO; MO. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. suteri Park, 1963: 183, figs. 3, 4, 8 (Actium). TN: Blount Co., Sinks Canyon: TN GA NC. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1971. HOST: From Rhododendron, and leaf litter from pine/oak mixture. wagneri Park, 1963: 182, figs. 4, 7, 9 (Actium). GA: Rabun Co., Mountain City; AL GA NC/ VA WV. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1971. HOST: From Rhododendron leaf litter.

Genus SIMPLONA Casey

Simplona Casey, 1897: 561 (as genus). Type-species: Simplona arizonica Casey (orig. des.). REDESCRIPTION: Grigarick and Schuster, 1980. arizonica Casey, 1897: 562. AZ: Coconino Co., Williams; AZ. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. REDESCRIPTION: Grigarick and Schuster, 1980. HOST: From rotting pine and Douglas-fir logs (Chandler, 1985); from aspen leaf litter. dybasi Chandler, 1985: 121, fig. 8, a-c. AZ: Cochise Co., Chiricahua Mountains, Onion Saddle; AZ. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From rotten conifer logs; under bark of conifer logs.

Genus PYGMACTIUM Schuster and Grigarick

Pygmactium Schuster and Grigarick, 1968: 112 (as genus). Type-species: Pygmactium steevesi Schuster and Grigarick (orig. des.). TAXONOMY: Chandler, 1990a (revision). REDESCRIPTION: Grigarick and Schuster, 1980. KEYS: Chandler, 1990a. mollyae Park, 1956: 56, fig. 1 {Actium). NC: Polk Co., near Tryon; AL TN NC SC/ TX. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Schuster and Grigarick, 1968. HOST: From rotten wood, and rotten yellow pine stump; from pine leaf and log litter. PSELAPHIDAE 31 quercavum Chandler, 1990a: 526, figs. 43-44. OK: Latimer Co., 5 mi. W. of Red Oak; OK/ AR. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From oak tree holes. steevesi Schuster and Grigarick, 1968: 113, figs. 1-7, 10-12, 14-15. FL: Highlands Co., Highlands Hammocks State Park; FL MS GA SC NC TN/ VA NJ DE NY. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1980. HOST: From rotten wood, palmetto leaf litter, sawdust, and mixture of sandpine and oak litter.

Genus PILACTIUM Grigarick and Schuster

Pilactium Grigarick and Schuster, 1970: 37 (as genus). Type-species: Pilactium summersi Grigarick and Schuster (orig. des.). REDESCRIPTION: Grigarick and Schuster, 1980. benedictae Grigarick and Schuster, 1978: 9, figs. 1-3. OR: Jackson Co., Soda Mountain Road, SE. Ashland; OR/ CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From oak leaf litter.

summersi Grigarick and Schuster, 1970: 37, figs. 1-4. CA: Plumas Co., Quincy; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1980. HOST: From pine forest soil.

Genus ALLOBROX Fletcher

Allobrox Fletcher, 1928: 205, pi. XII, figs. 20-23 (as genus). Type-species: Allobrox dampfi Fletcher (monot.). Cutrimia Park, 1945: 336 (as subgenus of Cupild) (synonymized by Grigarick and Schuster, 1980). Type-Species: Cupila mexicana Park. REDESCRIPTION: Grigarick and Schuster, 1977; Grigarick and Schuster, 1980.

stephani Grigarick and Schuster, 1977: 219, figs. 1-10. AZ: Pima Co., Santa Catalina Mts.; AZ. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From oak and sycamore leaf litter; from rotten pine and fir logs (Chandler, 1985).

Genus CUPILA Casey

Cupila Casey, 1897: 561 (as genus). Type-species: Trimium clavicorne Mäklin (orig. des.). TAXONOMY: Grigarick and Schuster, 1968 (revision). REDESCRIPTION: Grigarick and Schuster, 1968; Grigarick and Schuster, 1980. KEYS: Grigarick and Schuster, 1968.

clavicornis Mäklin, 1852: 371 {Trimium). AK: Sitka; AK/ BC WA OR. TYPE DEPOSITORY: UZMH. REDESCRIPTION: Grigarick and Schuster, 1968; Grigarick and Schuster, 1980. HOST: From leaf litter of Labrador tea and leaf litter of spruce/hemlock mixture; from moss.

excavata Park and Wagner, 1962: 18, pi. IV, figs. 2, 3. OR: Coos Co., Charleston; OR. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1968. HOST: From leaf litters of Douglas fir, Abies, cedar, and mixture of maple/alder; from moss. 32 COLEÓPTERA CATALOG multífossa Grigarick and Schuster, 1968: 43, figs. 1, 2, 5. CA: Humboldt Co., Freshwater: CA/ OR. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From leaf litters of redwood, oak, and fir.

Genus LEMELBA Park

Lemelba Park, 1953c: 1 (as genus). Type-species: Lemelba davisi Park (orig. des.). REDESCRIPTION: Grigarick and Schuster, 1980. davisi Park, 1953c: 2, fig. 1. FL: Dade Co., Everglades near Miami; FL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1980. HOST: From hardwood leaf and log litters; from sawtooth palm litter.

Genus MELBA Casey

Melba Casey, 1897: 565 (as genus) (other subgenera Neotropical). Type-species: Trimium thoracicum Brendel (design, by Bowman, 1934: 144). Trimium (of authors, not North American) (not Aube, 1833: 44). TAXONOMY: Chandler, 1990a (revision). REDESCRIPTION: Park, 1942; Grigarick and Schuster, 1980. KEYS: Chandler, 1990a.

Subgenus MELBA Casey maja Brendel, 1892b: 167 {Trimium). lA; Linn Co., Cedar Rapids; lA. TYPE DEPOSITORY: ANSP. SEX OF TYPE: F. TAXONOMY: Chandler, 1994. párvula LeConte, 1849: 108 {Euplectus). SC: Charleston Co., Charleston; lA IL IN OH NE KS/ PA NY NJ VA DC MD/ SC NC AL GA FL MS TN LA/ TX OK/ MA NH RI CT/ MI ON. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. durum Brendel, 1892b: 166 {Trimium) (synonymized by Chandler, 1990a). lA: Linn Co., Cedar Rapids. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. gracile Brendel, 1892b: 167 {Trimium) (synonymized by Chandler, 1990a). lA: Linn Co., Cedar Rapids. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. laticolle Brendel, 1892b: 167 {Trimium) (synonymized by Chandler, 1990a). lA: Linn Co., Cedar Rapids. TYPE DEPOSITORY: ANSP. SEX OF TYPE: F. TAXONOMY: Moxey, 1962. texana Casey, 1897: 566 (synonymized by Chandler, 1990a). TX: Colorado Co., Columbus. TYPE DEPOSITORY: USNM. uniformis Casey, 1897: 569 (synonymized by Chandler, 1990a). RI: Boston Neck. TYPE DEPOSITORY: USNM. REDESCRIPTION: Brendel, 1892b; Chandler, 1990a. HOST: From various rotten woods and litter from tree holes; less common in leaf litters (Chandler, 1990a). PSELAPHIDAE 33 quercae Chandler, 1985: 117, fig. 5, a-e. AZ: Graham Co., Pinaleño Mountains, Noon Creek; AZ. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From oak and sycamore leaf litters. sulcatula Casey, 1897: 567. PA; Westmoreland Co.; MI/ OH lA MO/ OK/ PA MY MD/ LA AR MS. TYPE DEPOSITORY: USNM. fossiger CñSQy, 1897: 568 (synonymized by Chandler, 1990a). NY. TYPE DEPOSITORY: USNM. dentipes Raffray, 1904a: 536 (synonymized by Chandler, 1990a). PA: Westmoreland Co., St. Vincent. TYPE DEPOSITORY: MNHP. SEX OF TYPE: M. REDESCRIPTION: Park, Auerbach, and Corley, 1950; Chandler, 1990a. HOST: From oak stumps and oak leaf litter; at UV light.

thoracica Brendel, 1889: 196 {Trimium). lA; lA IL MO/ GA NC SC FL AL MS AR LA/ TX OK/ PA DC VA. TYPE DEPOSITORY: ANS?. SEX OF TYPE: F. TAXONOMY: Moxey, 1962. REDESCRIPTION: Grigarick and Schuster, 1980; Chandler, 1990a. HOST: From mixed hardwood litter and rotten wood.

Unplaced Species

simplex LeConte, 1878a: 384 {Trimium). FL: Hillsborough Co., Tampa; FL. Probably a Pygmactium (Chandler, 1990a). TYPE DEPOSITORY: MCZC; type lost.

Genus ALLOTRIMIUM Park

AUotrimium Park, 1943: 187 (as genus). Type-species: Allotrimium michoacanensis Park (orig. des.). REDESCRIPTION: Grigarick and Schuster, 1980.

excavatnm Chandler, 1985: 114, fig. 3, a-d. AZ: Graham Co., Pinaleño Mountains, Hospital Flat; AZ, TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From various rotten woods.

tuberculatum Chandler, 1985: 115, fig. 4, a-e. AZ: Pima Co., Santa Catalina Mountains, Mount Lemmon Highway; AZ/ CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From oak and cottonwood leaf litters.

Genus ZONAIRA Grigarick and Schuster

Zonaira Grigarick and Schuster, 1980: 43 (as genus). Type-species: Zonaira trilinea Grigarick and Schuster (orig. des.).

puncticoUis LeConte, 1878a: 384 {Trimium). AZ; AZ. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. HOST: With ants. 34 COLEÓPTERA CATALOG

trilinea Grigarick and Schuster, 1980: 43, pi. 72, figs. 1-11. AZ: Pima Co., foothills of Santa Catalina Mountains; AZ; Mex. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From rotting saguaro cactus; from rotting sotol (Chandler, 1985).

Genus TRIMIOMELBA Casey

Trimiomelba Casey, 1897: 563 (as genus). Type-species: Euplectus dubius LeConte (design, by Lucas, 1920: 655). REDESCRIPTION: Grigarick and Schuster, 1980. KEYS: Casey, 1897.

americana Motschulsky, 1856: 7 (Trimium). AL: Mobile Co., Mobile; AL. This species was attributed to LeConte (1863b: 28) by previous authors. LeConte received his specimen from Motshulsky, and it is a syntype. TYPE DEPOSITORY: MCZC, ZMUM.

convexula LeConte, 1878a: 383 (Trimium). FL: Hillsborough Co., Tampa; FL. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. REDESCRIPTION: Grigarick and Schuster, 1980.

dubia LeConte, 1849: 108 (Euplectus). Type locality not given; MO IL IN OH/ MA RI CT/ PA NJ DC MD VA DE NY/ GA NC SC AL MS TN KY AR LA FL/ TX OK. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. REDESCRIPTION: Park, 1953d: pi. I. HOST: Primarily from various leaf litters; also from rotten logs; at UV light.

laevis Casey, 1897: 564. DC; DC. TYPE DEPOSITORY: USNM. SEX OF TYPE: F.

Genus DALMOSELLA Casey

Dalmosella Casey, 1897: 570 (as genus). Type-species: Dalmosella tenuis Casey (orig. des.). REDESCRIPTION: Park, 1942; Grigarick and Schuster, 1980. tennis Casey, 1897: 570. PA: Westmoreland Co; OH/ PA NJ DC/ ME NH MA/ NC GA FL AL TN MS KY LA/ OK. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. REDESCRIPTION: Grigarick and Schuster, 1980. HOST: From various rotten woods and material in tree holes (Chandler, 1990a).

Genns TRIMIUM Aube

Trimium Aube, 1833: 44 (as genus) (other subgenus Palearctic). Type species: Pselaphus brevicornis Reichenbach (monot.).

Subgenus TRIMIUM Aube brevicorne Reichenbach, 1816: 47 (Pselaphus). Germany: Berlin; LA. TYPE DEPOSITORY: Types lost. discolor hQConXQ, 1878a: 384 (Trimium) (synonymized by Chandler, 1990a; introduced into North America, but not established?). LA. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. PSELAPHIDAE 35

Genus TOMOPLECTUS Raffray

Tomoplectus Raffray, 1898b: 266 (as genus). Type-species: Tomoplectus cordicollis Raffray (monot.). REDESCRIPTION: Grigarick and Schuster, 1980. apache Chandler, 1985: 122, fig. 9, a-e. AZ: Cochise Co., Dragoon Mountains, Cochise Stronghold; AZ. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: At UV light.

Genus TRIMIOARCUS Park

Trimioarcus Park, 1952: 130 (as genus). Type-species: Trimioarcus incisurus Park (orig. des.). REDESCRIPTION: Grigarick and Schuster, 1980.

musamator Chandler, 1992b: 251, figs. 24-26. TX: Travis Co., Twin Dig Pit; TX. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From leaf litter in pit.

pajarito Chandler, 1985: 123, fig. 10, a-d. AZ: Santa Cruz Co., Pajarito Mountains, Peña Blanca; AZ. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M.

Genus PSEUDOTRIMIUM Raffray

Pseudotrimium Raffray, 1898b: 230 (as genus, not North American). Type-species: Pseudotrimium microcephalum Raffray (monot.).

microcephalum (not North American) Raffray, 1898b: 230. Mex.: Tabasco, Teapa. Raffray cited the type-locality as "New Orleans" in the description. This is probably an error. TYPE DEPOSITORY: MNHP. SEX OF TYPE: F. Tribe JUBINI Raffray, 1904a Jubinini Raffray, 1908; Auxenocerini Jeannel, 1962; Park, 1942; Park, 1952. Genus SEBAGA Raffray Sebaga Raffray, 1891: 300 (as genus). Type-species: Sebaga centralis Raffray (monot.). KEYS: Park, 1942; Park, 1952a.

ocampi Park, 1945: 333. Mexico: Tamaulipas, El Monte; and Tabasco, Emiliano; OK TX; Mex. TYPE DEPOSITORY: FMNH. REDESCRIPTION: Chandler, 1990a. HOST: From leaf litter. Genus CANTHODERUS Motschulsky

Canthoderus Motschulsky, 1855:15 (as genus, not North American). Type-species: Canthoderus villosus Motschulsky (monot.) Stratus L. W. Schaufiiss, 1872: 246 (as genus; not North American) (synonymized by Reitter, 1882). Type-species: Stratus ursinus L. W. Schaufiiss (subseq. monot., L. W. Schaufuss, 1877: 452). 36 COLEÓPTERA CATALOG

ursinus L. W. Schaufiíss, 1877: 452 {Stratus). Mex.: Tabasco, Teapa. New Orleans, LA, record is most likely an error. TYPE DEPOSITORY: MNHP.

Subfamily BYTHINOPLECTINAE L. W. Schaufuss, 1890

Tribe BYTHINOPLECTINI L. W. Schaufuss, 1890

Pyxidicerini Raffray, 1904a.

Subtribe BYTHINOPLECTINA L. W. Schaufuss, 1890

Zethopsina Jeannel, 1952; Comellini, 1985; Coulon, 1989.

Genus BYTHINOPLECTUS Reitter

Bythinoplectus Reitter, 1882: 209 (as genus). Type-species: Bythinoplectus foveatus Reitter (sub. monot. Reitter, 1883a: 37). TAXONOMY: Comellini, 1985. REDESCRIPTION: Comellini, 1985; Coulon, 1989. gloydi Park, 1949: 316, pi. I. AZ: Pinal Co., 4 mi. W. of Superior; AZ; Mex. TYPE DEPOSITORY: FMNH. SEX OF TYPE: F. REDESCRIPTION: Comellini, 1985. HOST: At UV light; under rock with ant, Paratrechina melanderi (Coulon 1989: 154). Tribe MAYETIINI Winkler, 1925

Jeannel and Coffait, 1955.

Genus MAYETIA Mulsant and Rey

Mayetia Mulsant and Rey, 1875: 87 (as genus). Type-species: Mayetia sphaerifera Mulsant and Rey (monot.). TAXONOMY: Schuster, Marsh, and Park, 1959 (revision of U.S. spp., excluding CA) and 1960 (revision of CA spp.). KEYS: Schuster, Marsh, and Park, 1959 (key to U.S. spp., excluding CA). bicorona Schuster, et al, 1959: 124, figs. 7, 13, 18, 19. GA: Upson Co., west edge of Thomasville; GA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: From Cecil sandy loam. bowmani Schuster, et al, 1959: 120, figs. 5, 11, 16. NC: Buncombe Co., Andropogon Bald, Shut-in-Ridge; NC. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. bulla Schuster, et al, 1959: 125, figs. 2, 10, 22. NC: Durham Co., Durham, Duke Forest; NC. TYPE DEPOSITORY: INHS. SEX OF TYPE: M. HOST: From clay soil beneath pines. PSELAPHIDAE 37 domestica Schuster, et al, 1959: 125, figs. 9, 15, 21. MO: Stoddard Co., Dexter; MO/ AR/ OK. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: From grass clumps in sandy soil; sandy soil beneath pine litter and moss (Chandler, 1990a). fistula Schuster, et al, 1960: 22, figs. 15, 16. CA: Eldorado Co., near Nashville; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From clay loam. grayae Schuster, 1961: 23, fig. 2. CA: Marin Co., Fort Ross, Cazadero Township; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: Under sod. judsoni Schuster, 1961: 26, fig. 3. CA: Marin Co., 1 mi. W. of Bolinas; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From litter and soil beneath cypress. langei Schuster, et al, 1960: 21, fig. 14. CA: Placer Co., 4 mi. W. of Newcastle; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: In soil under oak. mendocinoensis Schuster, et al, 1960: 15, figs. 1-5, 9, 10. CA: Mendocino Co., Mendocino; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: In soil. pearsei Schuster, et al, 1959: 124, figs. 1, 3, 4, 8, 14, 20. NC: Durham Co., Durham, Duke Forest; NC GA AR FL LA MS SC/ IN. TYPE DEPOSITORY: INHS. SEX OF TYPE: M. HOST: From clay soil under oaks; soil and litter in beech-maple forest (Waltz, 1981). pravitas Schuster, et al, 1960: 22, fig. 17. CA: Eldorado Co., near Nashville; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From clay loam. putahensis Schuster, 1961: 23, fig. 1. CA: Yolo Co., Putah Creek, 5.4 mi. SW. of Winters; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. raneyi Schuster, et al, 1960: 20, fig. 12. CA: Napa Co., 9 mi. S. of Monticello; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From soil under oaks.

scobina Schuster, et al, 1960: 20, fig. 13. CA: Placer Co., 4 mi. W. of Newcastle; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From soil under oak and grass. skeeteri Royce and Kistner, 1970: 84, figs. A-C. CA: Butte Co., 1 mi. E. of Chico, Bidwell Park; CA. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From grass sod in clay soil (Chandler, 1983). 38 COLEÓPTERA CATALOG smithi Schuster, 1961: 26, fig. 4. OR: Douglas Co., 2.7 mi. N. of Loon Lake; OR. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From Douglas-fir litter in forests. turneri Schuster, et al, 1959: 122, figs. 6, 12, 17. NC: Moore Co.; NC. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. walkeri Schuster, et al, 1960: 18, figs. 6-8, 11. CA: Del Norte Co., 6.4 mi. S. of Klamath; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: In redwood litter and soil. Subfamily BATRISINAE Reitter, 1882

Tribe BATRISINI Reitter, 1882 Subtribe BATRISINA Reitter, 1882

Genus BATRISODES Reitter

Batrisodes Reitter, 1882: 205. Type-species: Batrisus delaporti Aube (design, by Lucas, 1920: 135). Newton and Chandler, 1989 discuss the problems of type designations for this genus, Batrisus Aube, 1833 and Alytus Hampe, 1863. Batrisus (oí authors, not North American) (not Aube, 1833: 45). TAXONOMY: Grigarick and Schuster, 1962c (revision western spp.); Park, 1947b (revision eastern spp.), 1958 (revision cavernicolous spp.). KEYS: Park, 1947b (eastern U.S. spp.); Park, 1958 (cavernicolous spp.); Park and Wagner, 1962 (Pacific Northwest spp.); Grigarick and Schuster, 1962c (western U.S. spp.); Chandler, 1992b (Texas cave spp.).

Subgenus BABNORMODES Park

Babnormodes Park, 1951: 12 (as subgenus). Type-species: Pselaphus riparius Say (orig. des.). TAXONOMY: Park, 1956 (males). KEYS: Park, 1956 (all species) and 1960 (cavernicolous species). antennatus Schaeffer, 1906: 262. NC: Buncombe Co., Black Mountain; PA VA/ NC TN. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Park, 1947b. HOST: From litter by logs. appalachianus Casey, 1908: 262. PA: Westmoreland Co.; PA MD. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Park, 1947b. barri Park, 1958: 40. TN: Dickson Co., 1.5 mi. S. of Dickson, Dickson Cave; TN. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Cavernicole. cavicornis Casey, 1897: 579 {Batrisus), PA: Westmoreland Co.; IL OH KY IN/ PA WV. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Park, 1947b. HOST: From rotten wood and leaf litter. PSELAPHIDAE 39 clypeospecus Park, 1960: 71. TN: Fentress Co., Reed Creek Cave; TN. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Cavernicole. ferulifer Park, 1960: 69. TN: Maury Co., Benderman Cave, near Southport; TN. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Cavernicole. foveicornis Casey, 1887: 462 (Batrisus). TN; lA IN OH KY/ NY/ TN LA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Park, 1947b. HOST: With ant, Lasius umbratus (Wickham, 1896). gemmoides Park, 1960: 68. TN: Dickson Co., Columbia Caverns; TN. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Cavernicole. gemmus Park, 1956: 71. TN: Dickson Co., 12 mi. NW. of Dickson, Jewel Cave; TN. TYPE DEPOSITORY: UANH. SEX OF TYPE: M. HOST: Cavernicole. hairstoni Park, 1947b: 101, pi. VI, fig. 5. IN: Clarke Co., Clark Co. State Park, near Uno; IN KY/ PA. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Under rocks; in stomach of salamander, Plethodon cinereus cinereus (Park, 1947b). henroti Park, 1956: 70, fig. 10. KY: Barren Co., Vance Cave; KY. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Park, 1958. HOST: Cavernicole.

hubrichti Park, 1958: 41, fig. 1. KY: Barren Co., 0.4 mi. NW. of Beckton, Beckton Cave; KY. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Cavernicole. jocuvestus Park, 1960: 70. AL: Madison Co., Aladdin Cave, near Maysville; AL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Cavernicole.

jonesi Park, 1951: 27, figs. 5, 6. AL: Colbert Co., Tuscumbia, Little Bear Cave; AL. TYPE DEPOSITORY: UANH. SEX OF TYPE: M. HOST: Cavernicole.

krekeleri Park, 1960: 74. IN: Clarke Co., 2.5 mi. SSW. of New Washington; IN. TYPE DEPOSITORY: FMNH. SEX OF TYPE: F. HOST: Cavernicole.

pannosus Park, 1960: 72 TN: Putnam Co., 2 mi. N. of Cookeville, Pedigo Cave; TN. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Cavernicole. 40 COLEÓPTERA CATALOG

profundas Park, 1956: 68, figs. 7, 8. AL: Conecuh Co., near Brooklyn, Turk's Cave- AL TYPE DEPOSITORY: UANH. ' SEX OF TYPE: M. HOST: Cavernicole.

punctifrons Casey, 1887: 463 (Batrisus). PA; OH/ PA NY WV MD/ NH MA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. ECOLOGY: Chandler, 1987. REDESCRIPTION: Park, 1947b. HOST: From leaf litters of beeclVbirch mixture, maple, oak, and conifer; in rotting conifer and hardwood logs.

riparius Say, 1824: 98 {Pselaphus). MO; MO IL IN OH/ OK/ NY PA NJ DC WV/ GA NC KY MS/ PQ ON MI. TYPE DEPOSITORY: Types lost. REDESCRIPTION: Park, 1947b. HOST: Under bark; in tree holes; with ant, Aphaenogaster tennesseenis (Park, 1935b); from leaf litter by fallen trees.

rossi Park, 1947b: 100, pi. VI, fig. 2. IL: Pope Co., Herod; IL. TYPE DEPOSITORY: INHS. SEX OF TYPE: M.

schaefferi Park, 1947b: 104, pi. VI, fig. 6. NC; VA/ GA NC SC. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From rotten stumps; from pine and hardwood leaf litters.

specus Park, 1951: 35, fig. 9. AL: Madison Co., Sharp's Cave, Hutton Cave; AL TN/ OH TYPE DEPOSITORY: UANH. SEX OF TYPE: M. HOST: Cavernicole. spretus LeConte, 1849: 100 {Batrisus). GA: White Co., Nacoochee (Nakutshi); IL IN OH KY lA MO/ NY WV PA DC VA/ VT ME/ NC TN GA. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. luculentus Casey, 1887: 460 (Batrisus) (synonymized by Brendel, 1891) DC TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Park, 1947b. HOST: With termite, Reticulitermes flavipes (Blatchley, 1910); from hemlock leaf litter. subterraneus Park, 1951: 30, fig. 7. AL: Marshall Co., Guntersville, Griffith Cave; AL TYPE DEPOSITORY: UANH. SEX OF TYPE: M. HOST: Cavernicole. tumoris Park, 1960: 73. AL: Colbert Co., McClusky Cave; AL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: In caves, beneath rocks. uncicornis Casey, 1897: 576 {Batrisus). NY: New York City; NY PA NJ VA/ MA RI/ MS AL TN NC FL GA LA/ OK TX. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. schneiderensis Park, 1960: 75 (synonymized by Chandler, 1992b). TX: Kendall Co., 12 mi. NNE. of Boerne, Schneider Cave. TYPE DEPOSITORY: FMNH. SEX OF TYPE: F. REDESCRIPTION: Park 1956, 1947b; Chandler, 1992b. HOST: From sawdust; under pine bark; under stones; troglophile (Chandler, 1992b); at UV light. PSELAPHIDAE 41 valentinei Park, 1951: 32, fig. 8. AL: Madison Co., Keel Mt., Goat Cave; AL TN. TYPE DEPOSITORY: UANH. SEX OF TYPE: M. REDESCRIPTION: Park, 1958. ECOLOGY: Park, 1956. HOST: Cavernicole. Subgenus DECLIVODES Park

Declivodes Park, 1951: 13 (as subgenus). Type-species: Batrisus bistriatus LeConte (orig. des.). bistriatus LeConte, 1849: 101 {Batrisus). PA; PA NJ MD DC/ CT. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. REDESCRIPTION: Park, 1947b. HOST: With ants, Formica exsectoides (Park, 1947b), and Lasius claviger? (Schwarz, 1890; Hamilton, 1886). cartwrighti Sanderson, 1940: 169. SC: Oconee Co., Clemson College; SC. TYPE DEPOSITORY: INHS. SEX OF TYPE: M. REDESCRIPTION: Park, 1947b. declivis Casey, 1908: 262. lA: Johnson Co., Iowa City; lA IL. TYPE* DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Park, 1947b. fossicauda Casey, 1897: 575 (Batrisus). PA: Westmoreland Co.; lA/ PA NY/ CT/ LA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Park, 1947b. HOST: With ants, Formica exsectoides and F. subsericea (Wickham, 1900a). lineaticoUis Aube, 1833: 50, pi. 90, fig. 3 (Batrisus). "America septentrionali"; OH/ NY PA NJ/ GA. TYPE DEPOSITORY: MNHP. REDESCRIPTION: Park, 1947b. HOST: With ants, Formica subsericea (Wickham, 1894), and F. exsectoides. Subgenus ELYTRODES Park

Elytrodes Park, 1951: 13 (as subgenus). Type-species: Batrisus ionae LeConte (orig. des.). caseyi Blatchley, 1910: 326. IN: Posey Co.; IN. TYPE DEPOSITORY: CMNH. HOST: Beneath a chunk of wood in open sandy woods.

ioîiae LeConte, 1849: 94 (Batrisus). GA; IN OH KY MO KS/ GA MS/ PA NJ DC VA MD/ OK. TYPE DEPOSITORY: MCZC. REDESCRIPTION: Park, 1947b. HOST: From hardwood leaf litter; with ant, Lasius alienus (Schwarz, 1890).

Subgenus EMPINODES Park

Empinodes Park, 1953d: 315 (as subgenus). Type-species: Batrisus albionicus Aube (orig. des.). TAXONOMY: Grigarick and Schuster, 1962c. KEYS: Grigarick and Schuster, 1962c.

albionicus Aube, 1833: 49, pi. 90, fig. 2 (Batrisus). BC; AK/ BC WA OR/ CA. TYPE DEPOSITORY: MNHP. REDESCRIPTION: Grigarick and Schuster, 1962c. HOST: From leaf litters of redwood, alder, fern, maple, fir, Jeffrey and Oregon pines, madrone, oak, incense cedar, and hemlock. 42 COLEÓPTERA CATALOG

aphenogastri Fall, 1912: 11. ID: Latah Co., Kendrick; ID. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. REDESCRIPTION: Park and Wagner, 1962. HOST: In nest of ant, Aphaenogaster subterránea var. occidentalis.

cicatricosus Brendel, 1891: 15, pi. X, fig. 83 {Batrisus) (not Batrisus cicatricosus Raffray 1897- 248 (of Sumatra)). CA: Placer Co.; CA. ^' • TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. pygidialis Casey, 1894: 470 {Batrisus) (synonymized by Grigarick and Schuster, 1962c).

TYPE DEPOSITORY: USNM. REDESCRIPTION: Grigarick and Schuster, 1962c. HOST: Under bark of dead pines or firs; in rotten conifer logs. denticauda Casey, 1894: 471 {Batrisus), CA: Siskiyou Co.; WA OR BC/ CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1962c. HOST: Under bark of dead conifers; in rotten conifer logs (Chandler, 1983). indistinctus Grigarick and Schuster, 1962c: 201, fig. 11. CA: Shasta Co., 7 mi. N. of Lakehead;

TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: Under stones. lustrans Casey, 1908: 260. CA: Lake Tahoe; CA. TYPE DEPOSITORY: USNM. REDESCRIPTION: Grigarick and Schuster, 1962c. martini Grigarick and Schuster, 1962c: 201, figs. 6, 10, 12. CA: Marin Co., Cypress Ridge- CA TYPE DEPOSITORY: CASC. ^ ' SEX OF TYPE: M. mendocino Casey, 1886: 174 {Batrisus). CA: Mendocino Co., Anderson Valley; CA TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1962c. HOST: From leaf litters of redwood, oak, Douglas fir, alder, and skunk cabbage. montícola Casey, 1886: 177 {Batrisus) (probably a synonym of Batrisodes lustrans Casey, Grigarick and Schuster, 1962c). CA: Eldorado Co.; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. nebulosus Grigarick and Schuster, 1962c: 202, figs. 5, 13. CA: Placer Co., 4 mi. W. of Newcastle; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From Quercus wislizenii litter. obscuras Grigarick and Schuster, 1962c: 206, figs. 4, 14. CA: Placer Co., 4 mi. W. of Newcastle;

TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From Adenostema and Quercus litter. occiduus Casey, 1886: 178 {Batrisus). CA: Humboldt Co; CA. TYPE DEPOSITORY: USNM. PSELAPHIDAE 43 opacus Grigarick and Schuster, 1962c: 210, fig. 19. CA: San Mateo Co., 6 mi. SE. of Half Moon Bay; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From leaf litter of redwood and laurel. oro Chandler, 1983: 230, fig. 15. CA: Butte Co., Mountain House; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From moist leaf litter near streams and springs. speculum Casey, 1886: 176 {Batrisus). CA: Alameda Co.; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. REDESCRIPTION: Grigarick and Schuster, 1962c. tulareanus Casey, 1908: 261. CA: Tulare Co.; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1962c. zephyrinus Casey, 1886: 175 {Batrisus), NV: Washoe Co, Reno; CA NV. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1962c. HOST: With ant, Formica rufa (Mann, 1911); from rotten wood and litter along streams (Chandler, 1983).

Subgenus EXCAVODES Park

Excavodes Park, 1951: 12 (as subgenus). Type-species: Batrisus frontalis LeConte (orig. des.). auerbachi Park, 1956: 80, figs. 11, 12. TN: Sevier Co., Great Smoky Mountains National Park; TN NC GA. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From leaf litter. beyeri Schaeffer, 1906: 261. NC: Buncombe Co., Black Mountain; NC. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Park, 1947b.

cavernosus Park, 1951: 25, fig. 4. AL: Butler Co., Greenville, Hinson Cave; AL. TYPE DEPOSITORY: UANH. SEX OF TYPE: M. HOST: Cavernicole.

clypeonotus Brendel, 1893: 280, pi. IV, fig. 4 {Batrisus). LA: Tangipahoa Co., Ponchatoula; LA AL AR/ MO/ OK TX. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. tridens Casey, 1908: 263 (synonymized by Chandler, 1992b). MO: St Louis Co., St. Louis; MO. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Park, 1956; Chandler, 1992b. HOST: From tree-hole debris; with ant, Camponotus americanus (Chandler, 1992b).

festinatus Park, 1956: 82, fig. 14. MS: George Co., near Lucedale; MS. TYPE DEPOSITORY: CUIC. SEX OF TYPE: M. 44 COLEÓPTERA CATALOG frontalis LeConte, 1849: 96 (Batrisus). PA; MB AB/ MN WI PQ ON/ CO/ KS lA MO IL OH/ PA DC/ ME/ LA. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. REDESCRIPTION: Park, 1947b. HOST: Under bark with ants, Lasius claviger (Wickham, 1898), L americanus (Wickham, 1900b), and L. umbratus (Park, 1947b). furcatus Brendel, 1891: 82 {Batrisus). Southern PA; MN MI PQ/ IL IN lA/ ME NH RJ/ PA DE NY VA/ MA/ AL TN NC KY GA/ OK. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. REDESCRIPTION: Park, 1947b. ECOLOGY: Reichle, 1966 and 1969. HOST: From rotten pine logs, leaf litter, under bark, and sphagnum moss. globosos LeConte, 1849: 100 {Batrisus). PA, GA; MN WI MI ON PQ/ CO/ KS lA MO IL IN OH KY/ NY PA NJ MD VA DC WV/ ME VT NH MA RI CT/ NB NS/ WA/ AB/ TX OK/ MS AL FL AR LA TN GA NC. TYPE DEPOSITORY: MCZC. REDESCRIPTION: Park, 1947b; Chandler, 1992b. ECOLOGY: Reichle, 1966; Engelmann, 1956; Park, 1932b, 1935a, 1935b, and 1947b; Park et al, 1953; Chandler 1987. HOST: Primarily in rotten wood; often with ants, Lasius niger americanus, L claviger, L. umbratus, Camponotus caryae subbarbatus, C. herculeanus, C. pennsylvanicus, C. noveboracensis. Formica cinérea neocinerea, F. ulkei (Schwarz, 1890; Wickham, 1898; Park, 1947b); in tree holes; under bark; in sphagnum moss; in caves (Chandler, 1992b). grubbsi Chandler, 1992b: 245, figs. 5-6. TX: Hays Co., Grapevine Cave; TX. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Cavernicole. kahli Bow^man, 1934: 141. TN:^ TN. TYPE DEPOSITORY: ICCM. SEX OF TYPE: M. mississippiensis Park, 1956: 81, fig. 13. MS: George Co., Lucedale; MS. TYPE DEPOSITORY: CUIC. SEX OF TYPE: M. reyesi Chandler, 1992b: 246, figs. 7-8. JX: Burnet Co., Fenceline Sink; TX. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Cavernicole. scabriceps LeConte, 1849: 98 {Batrisus). LA: Orleans Parish, New Orleans; WI MI ON PQ/ lA MO IL OH IN/ NY PA DC MD WV/ CT/ AR LA MS NC TN/ OK TX. TYPE DEPOSITORY: MCZC. harringtoni Casey, 1897: 578 {Batrisus) (synonymized by Park, 1947b). ON: Ottawa; WI: Bayfield; PA: Westmoreland Co. TYPE DEPOSITORY: USNM. REDESCRIPTION: Park, 1947b. HOST: With ants, Lasius umbratus (Park, 1935b), Formica exsectoides, Aphaenogaster tennesseensis (Park, 1935a), F. subsericea (Wickham, 1896), A. fulva (Park, 1947b); under oak bark. sinuatifrons Brendel, 1893: 280, pi. IV, fig. 5 {Batrisus). TN: Shelby Co., Memphis; AR LA TN MS/ ON. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Moxey, 1962. REDESCRIPTION: Park, 1947b. HOST: From beech and maple litters; at UV light. PSELAPHIDAE 45 temporalis Casey, 1897: 573 (Batrisus). PA: Westmoreland Co.; PA MD/ FL. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Park, 1947b. HOST: From beech logs. texanus Chandler, 1992b: 247, figs. 13-14. TX: Williamson Co., Inner Space Caverns; TX. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Cavernicole (now on the federal list of threatened and endangered species (Beacham, 1994)). venyivi Chandler, 1992b: 247, figs. 15-16. TX: Bexar Co., Helotes Hilltop Cave; TX. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Cavernicole. virginiae Casey, 1884b: 90 (Batrisus), VA: Lee Co., Stone Creek; VA WV/ NC TN. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: From hardwood litter; under bark; with ant, Aphaenogaster tennesseensis (Park, 1935b).

Subgenus PUBIMODES Park

Pubimodes Park, 1951: 12 (as subgenus). Type-species: Batrisus nigricans LeConte (orig. des.).

denticollis Casey, 1884b: 89 {Batrisus), DC; lA MO XL OH/ NY PA NJ MD DC VA/ GA NC FL SC TN KY AR/ OK. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Park, 1947b. HOST: From Rhododendron and other deciduous leaf litters by fallen trees and along streams.

nigricans LeConte, 1849: 99 (Batrisus). SC: Richland Co., Columbia; GA SC. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. HOST: From oak leaf litter.

schmitti Casey, 1897: 581 (Batrisus). PA: Westmoreland Co.; IN KY OH/ PA VA WV/ NC SC AL MS TN. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Park, 1947b. HOST: From hardwood litter, and from litter along streams.

spinifer Brendel, 1887: 205, figs., 1-3 (Batrisus). NY: Long Isl.; IN OH/ CT/ NY NJ DC MD/ CO. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Chandler, 1994. triangulifer BrrndQl 1891: 28, pi. X, figs. 91, 93b (Batrisus) (unjustified replacement name for B. spinifer). REDESCRIPTION: Park, 1947b. ECOLOGY: Reichle, 1969. HOST: From sphagnum moss in white cedar bog; at UV light.

striatus striatus LeConte, 1849: 99 (Batrisus). PA; MN WI MI ON PQ/ NE KS lA MO IL IN/ NY PA NJ DC/ GA/ MA RI CT NH/ NC KY. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. 46 COLEÓPTERA CATALOG

simplex LeConte, 1878b: 598 {Batrisus) (synonymized by Casey, 1894: 472) (not Batrisus simplex Raffray, 1898a: 440). MI: Wayne Co., Detroit. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. REDESCRIPTION: Park, 1947b. ECOLOGY: Reichle, 1966 and 1969. HOST: From sphagnum moss; from Typha drift on beach. aterrimus Casey, 1884b: 91 {Batrisus) (synonymized by Casey, 1894: 472). MA: Middlesex Co., Cambridge. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. cephalotes Casey, 1887: 459 {Batrisus) (synonymized by Casey, 1894: 472). NY. TYPE DEPOSITORY: USNM. SEX OF TYPE: M.

striatus psotai Park, 1947b: 105 (described as variety). IL: Lake Co., Antioch; IL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M.

Subgenus SPIFEMODES Park

Spifemodes Park, 1953d: 315 (as subgenus). Type-species: Batrisus schaumi Auhé (orig. des.).

schaumi Aube, 1844: 84 {Batrisus). "Amérique du Nord"; MI/ MO IL IN OH/ NY PA NJ VA/ MS GA NC SC FL TN LA/ KY. TYPE DEPOSITORY: ZSBS. punctatus LeConte, 1849: 97 {Batrisus) (synonymized by LeConte, 1863a). GA: White Co. TYPE DEPOSITORY: MCZC. REDESCRIPTION: Park, 1956. HOST: With ants, Lasius umbratus, Aphaenogaster tennesseenis (Park, 1935b); from leaf litter along streams; under bark.

Not assigned to subgenus juvencus Brendel, 1865b: 258 {Batrisus) (not assigned to subgenus). Northern IL; IL. TYPE DEPOSITORY: Not at ANSP, ICCM, or MCZC; apparently lost (Chandler, 1994). SEX OF TYPE: F.

NOMEN NUDUM aculeatus LeConte, 1863a: 21. The name aculeatus was placed in a catalog by LeConte and is a nomen nudum. However, it has been treated as a synonym of Batrisodes albionicus by C. Schaufiiss (1888) and Leng (1920). LeConte (1866a: 78) noted that this name was to be deleted in a list of corrections to the catalog.

Genus TEXAMAUROPS Barr and Steeves

Texamaurops Barr and Steeves, 1963: 117 (as genus). Type-species: Texamaurops reddelli Barr and Steeves (orig. des.). This genus was initially placed in Amauropsini; later placed in Batrisini by Barr, 1974. TAXONOMY: Barr, 1974; Chandler, 1992b. REDESCRIPTION: Barr, 1974. reddelli Barr and Steeves, 1963: 118, fig. 1. TX; Travis Co., 15 mi. NW. of Austin, Kretschmarr Cave; TX. TYPE DEPOSITORY: FMNH. SEX OF TYPE: F. REDESCRIPTION: Barr, 1974; Chandler, 1992b. HOST: Cavernicole (now on the federal list of threatened and endangered species (Lowe et al., 1990)). PSELAPHIDAE 47

Genus BATRIASYMMODES Park Batriasymmodes Park, 1951: 11, figs. 1-3 (as subgenus oí Batrisodes). Type-species: Batrisus monstrosus LeConte (orig. des.). Batrisus (of authors, not North American) (not Aube, 1833: 45). TAXONOMY: Park, 1965. REDESCRIPTION: Park, 1965. KEYS: Park, 1965. Subgenus BATRIAS ^^MMODES Park armiger LeConte, 1849: 94 {Batrisus). PA; PA/ NC SC FL. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. REDESCRIPTION: Park, 1947b. HOST: With ants; from mixed hardwood litter. beameri Park, 1956: 66 {Batrisodes). FL: Highlands Co., Sebring; FL NC AL. TYPE DEPOSITORY: SEMC. SEX OF TYPE: M. HOST: From sand pine, oak, and palmetto litter; at UV light. carolinae Casey, 1894: 468 {Batrisus). NC: Buncombe Co., Asheville; NC. TYPE DEPOSITORY: USNM. REDESCRIPTION: Park, 1947b. cavicrus Casey, 1894: 468 {Batrisus). NC: Buncombe Co., Asheville; IN OH/ PA/ TN GA SC NC. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Park, 1947b. HOST: From litters of pine, oak, and hardwood; from rotten wood; at UV light.

confmis LeConte, 1849: 96 {Batrisus). GA: Clarke Co., Athens; MO IL IN OH/ PA MD/ AL GA SC NC FL. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. curvatus Sanderson, 1940: 170 {Batrisodes) (synonymized by Park, 1947b). SC: Oconee Co., Clemson College. TYPE DEPOSITORY: INHS. SEX OF TYPE: M. REDESCRIPTION: Park, 1947b. HOST: From sand pine and palmetto litter; from mixed hardwood litter.

greeveri Park, 1965: 189, fig. 4. TN: Sullivan Co., Potters Cave; TN. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Cavernicole.

monstrosus LeConte, 1849: 95 {Batrisus). GA: Clarke Co., Athens; IL IN OH/ NY PA NJ DE MD DC VA/ GA NC FL. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. TAXONOMY: Barr, 1987. cristatus LeConte, 1849: 96 {Batrisus) (synonymized with ferox by Brendel, 1865b). PA. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. ferox LeConte, 1849: 95 {Batrisus) (synonymized by Brendel, 1866). PA, OH. TYPE DEPOSITORY: MCZC. REDESCRIPTTON: Park, 1947b. IMMATURE STAGES: Böving and Craighead, 1931. HOST: With ants, Lasius umbratus (Park, 1935b), L claviger, L interjectus, Amblyopone serrata (Schwarz, 1890, ^s ferox); facultative myrmecophile; at UV light; under bark of logs; from leaf litter; from material from tree holes. 48 COLEÓPTERA CATALOG parki Barr, 1987: 22, fig. 1. WV: Mercer Co., Neely Farm Cave, near Athens; WV. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. - HOST: Cavernicole,. quisnamus Park, 1951: 20 {Batrisodes), TN: Picket Co., Byrdstown, Bunkum Cave; KY/ TN. TYPE DEPOSITORY: UANH. SEX OF TYPE: F. jeanneli Park, 1951: 21^ fig. 2 {Batrisodes) (synonymized by Park, 1960). TN: Dekalb CO., Doweltown, Lindsay Williams Cave. TYPE DEPOSITORY: UANH. SEX OF TYPE: M. reduncus Park, 1956: 65 {BaMsodes) (synonymized by Park, 1960). TN: Sumner Co., Sam Lackey Cave. TYPE DEPOSITORY: FMNH. SEX OF TYPE: F. REDESCRIPTION: Park, 1965. HOST: Cavernicole. sandersoni Park, 1947b: 97, pi. VI, fig. 3 {Batrisodes). IL: Pope Co., Herod; IL. TYPE DEPOSITORY: INHS. SEX OF TYPE: F. spelaeus Park, 1951: 16, fig. 1 {Batrisodes). AL; Blount Co., Bangor, Bangor Cave; AL TN. TYPE DEPOSITORY: UANH. SEX OF TYPE: M. REDESCRIPTION: Park, 1965. HOST: Cavernicole. suteri Park, 1965: 187, fig. 2. TN: Blount Co., Great Smoky Mountains National Park; TN. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From leaf litter; under bark of logs; beneath rocks. Subgenus EXTOLLODES Park Extollodes Park, 1965: 185 (as subgenus). TypQ-s^QC\ts\ Batriasymmodes wagneriYdixk (orig. des). frosti Park, 1965: 191, fig. 8. FL: Highlands Co., Archbold Biological Station; FL, AL. TYPE DEPOSITORY: PSUC. SEX OF TYPE: M. HOST: From rhizome forest floor debris; at UV light. wagneri Park, 1965: 191, fig. 7. AL: Mobile Co., Theodore; MS AL FA LA. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From pine leaf litter and decomposing stumps; from beech litter; at UV light. Subgenus SPELEODES Park Speleodes Park, 1965: 186 (as subgenus). Type-species: Batrisodes troglodytes Park (orig. des.). troglodytes Park, 1951: 23 {Batrisodes). AL: Butler Co., Greenville, Rock Cave; AL. TYPE DEPOSITORY: UANH. SEX OF TYPE: M. HOST: Cavernicole. Genus ARTHMIUS LeConte Arthmius LeConte, 1849: 91 (as genus). Type-species: Arthmius globicollis LeConte (monot.). REDESCRIPTION: Park, 1942. KEYS: Casey, 1894. PSELAPHIDAE 49 bulbifer Casey, 1894: 474, pi. I, fig. 7. TX; TX OK/ AR/ KS. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: From debris on lake edge; in grass clumps; from hackberry leaf litter; at UV light. globicoUis LeConte, 1849: 91. GA, PA; OH XL KY/ PA VA NJ MD DC/ TN GA NC FL AL MS LA/ TX OK. TYPE DEPOSITORY: MCZC. HOST: From hardwood and pine leaf litters; at UV light. gracilior Casey, 1884b: 88. FL: Hillsborough Co., Tampa; FL. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: From leaf litter along the banks of springs and streams; at UV light. involutus Casey, 1894: 474. NC: Buncombe Co., Asheville; NC SC GA FL AL LA TN/ NJ VA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: From pine and hardwood litter; from swamp litter; at UV light. morsus Fletcher, 1932: 33. AZ: Coconino Co., Oak Creek Canyon; AZ. TYPE DEPOSITORY: SEMC. SEX OF TYPE: M. Genus EUPHALEPSUS Reitter Euphalepsus Reitter, 1883b: 378 (as genus, not North American). Type-species: Euphalepsus globipennis Reitter (design, by Park, 1942: 258). TAXONOMY: Park, 1942. REDESCRIPTION: Park, 1942. KEYS: Park, 1942. dentipes Raffray, 1904b: 63. LA?; LA. This species was described with a question mark as to whether it is from Louisiana; it probably is not from Louisiana. TYPE DEPOSITORY: MNHP. SEX OF TYPE: M. Tribe AMAUROPINI Jeannel, 1948 (Amauropsini Jeannel, 1948, incorrect original spelling). Genus ARIANOPS Brendel Arianops Brendel. 1893: 278, pi. IV, fig. 3 (as genus). Type-species: Anops amplyoponica Brendel (monot). ^ .«.^/^ Anops Brendel, 1891b; 80 (as genus, preoccupied; not Anops Oken, 1815 (Crustacea) or others). Type-species: Anops amplyoponica Brendel (monot.). Eusanops Brendel, 1893: 278 (published in synonymy with Arianops; unavailable, never validated). Arispeleops Park, 1951: 41 (as subgenus) (synonymized by Barr, 1974). Type-species: Arianops cavernensis Park (orig. des). TAXONOMY: Ban-, 1974 (revision). REDESCRIPTION: Barr, 1974. KEYS: Barr, 1974. allatoona Barr, 1974: 33, figs. 17A, 17B. GA: Cherokee Co., 5 mi. NE. of Acworth; GA. TYPE DEPOSITORY: AMNH. SEX OF TYPE: M. HOST: Under rocks. alticola Barr, 1974: 22, figs. 8A, 8B. NC: Macon Co., 4.5 mi. N. of Highlands, Buck Creek Rd.; NC. TYPE DEPOSITORY: AMNH. SEX OF TYPE: M. HOST: Under stones. 50 COLEÓPTERA CATALOG ampiyoponica Brendel, 1891: 80 (Anops), PA: Westmoreland Co., Latrobe; PA. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Chandler, 1994. amblyoponica (of authors). {Anops). The species was named after ant genus Amblyopone, and was originally misspelled by Brendel as ampiyoponica. Subsequent spellings have been amblyoponica (except Barr, 1974). REDESCRIPTION: Barr, 1974. HOST: With ants, Amblyopone pallipes and A. serrata (Schwarz, 1895). barbata Barr, 1974: 23, figs. 9A, 9B. NC: Swain and Macon Co., Leatherman Gap; NC. TYPE DEPOSITORY: AMNH. SEX OF TYPE: M. HOST: Under rocks. cavernensis Park, 1951: 41. AL: Marshall Co., Salpêtre Cave; AL. TYPE DEPOSITORY: UANH. SEX OF TYPE: F. REDESCRIPTION: Barr, 1974. HOST: Cavernicole, under rocks. copelandi Carlton, 1990: 366, fig. 1. AR: Pulaski Co., Pinnacle Mountain State Park, northeast slope of Pinnacle Mt. near summit; AR. coweeta Barr, 1974: 31, figs. 14A-C. NC: Macon Co., Coweeta Hydrologie Laboratory; NC. TYPE DEPOSITORY: AMNH. SEX OF TYPE: M. digitata Barr, 1974: 27, figs. 12A, 12B. TN: Sevier Co., Great Smoky Mountains National Park; TN. TYPE DEPOSITORY: AMNH. SEX OF TYPE: M. HOST: Under rocks. extera Barr, 1974: 39, figs. 20A, 20B. AL: Jackson Co., Horseshoe Cave; AL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Under stones. fovealis Barr, 1974: 46. NC: Macon and Swain Co., Leatherman Gap; NC. TYPE DEPOSITORY: AMNH. SEX OF TYPE: F. gigantea Barr, 1974: 41, figs. 23A, 23B. NC: Macon and Swain Co., Leatherman Gap; NC. TYPE DEPOSITORY: AMNH. SEX OF TYPE: M. henroti Park, 1956: 84. GA: Rabun Co., Rabun Bald, near Rabun Gap; GA. TYPE DEPOSITORY: MNHP. SEX OF TYPE: M. HOST: From ground cover and under rocks. jeanneli Park, 1956: 85. VA: Lee Co., Pennington Gap, Gilly's Cave; VA. TYPE DEPOSITORY: MNHP. SEX OF TYPE: F. REDESCRIPTION: Barr, 1974. kingi Barr, 1974: 40, figs. 22A, 22B. AL: St. Clair Co., 2.5 mi. SW. of Oneonta; AL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. PSELAPHIDAE 51 laminata Barr, 1974: 17, figs. 5A, 5B. NC: Buncombe Co., Round Mt.; NC. TYPE DEPOSITORY: AMNH. SEX OF TYPE: M. HOST: Under rocks. nantahalae Joanna Barr, 1974: 25. NC: Cherokee and Graham Cos., Teyahalee Bald; NC. TYPE DEPOSITORY: AMNH. SEX OF TYPE: M. nantahalae nantahalae Barr, 1974: 25, figs. lOA, lOB. NC: Macon Co., Winespring Bald; NC GA. TYPE DEPOSITORY: AMNH. SEX OF TYPE: M. HOST: Under rocks; from leaf litter. neglecta Barr, 1974: 29, figs. 1, 13A, 13B. GA: Rabun Co., near Rabun Bald; GA NC. TYPE DEPOSITORY: AMNH. SEX OF TYPE: M. HOST: Under stones and with Amblyopone pallipes (Haldeman). nodosa Barr, 1974: 20, figs. 7A, 7B. NC: 1.2 mi. E. of Busick, Buck Creek Gap; NC. TYPE DEPOSITORY: AMNH. SEX OF TYPE: M. norithe Barr, 1974: 46, figs. 26A, 26B. NC: Graham Co., Cheoah Bald; NC. TYPE DEPOSITORY: AMNH. SEX OF TYPE: M. HOST: Beneath moss carpet. obliqua Barr, 1974: 47, figs. 27A, 27B. GA: Towns and White Cos., Unicoi Gap; GA. TYPE DEPOSITORY: AMNH. SEX OF TYPE: M. HOST: Under rocks.

parki Barr, 1974: 31, figs. 15A, 15B. NC: Macon and Swain Cos., Leatherman Gap; NC. TYPE DEPOSITORY: AMNH. SEX OF TYPE: M. HOST: Under stones.

pecki Barr, 1974: 36. TN: Cannon Co., Henpeck Mill Cave; TN. TYPE DEPOSITORY: FMNH. SEX OF TYPE: F. HOST: Cavernicole, under stones.

plectrops Casey, 1897: 582. NC: McDowell Co., Round Knob; NC. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Barr, 1974. HOST: From forest litter; with ant, Amblyopone serrata (Schwarz, 1895, as Arianops amblyoponica in part).

sandersoni Barr, 1974: 21. AR: Logan Co., Magazine Mt.; AR. TYPE DEPOSITORY: INHS. SEX OF TYPE: F. REDESCRIPTION: Carlton and Allen, 1989. HOST: From forest debris beneath logs, stumps, and large rocks (Carlton and Allen, 1989).

sewanee Barr, 1974: 39, figs. 21A, 21B. TN: Franklin Co., Sewanee; TN. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Under rocks. 52 COLEÓPTERA CATALOG spinícollis Barr, 1974: 18, figs. 6A, 6B. NC: Buncombe Co., Mount Pisgah; NC. TYPE DEPOSITORY: AMNH. SEX OF TYPE: M. HOST: Under rocks. steevesi Barr, 1974: 38, figs. 19A, 19B. AL: Jackson Co., 7.5 mi. N. of Princeton, Horseshoe Cave; AL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Cavernicole; under rocks. stephani Carlton, in Carlton and Allen, 1989: 62, figs. 4, 7-9. OK: Latimer Co., Robber's Cave State Park. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Under rocks in upland forest. stygica Park, 1960: 67. TN: Warren Co., Cumberland Caverns; TN. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Barr, 1974. HOST: Cavernicole; under rocks. teyahalee Barr, 1974: 47, figs. 28A, 28B. NC: Graham Co., Teyahalee Bald; NC. TYPE DEPOSITORY: AMNH. SEX OF TYPE: M. HOST: Under stones. thornei Barr, 1974: 45, figs. 25A, 25B. NC: Macon Co., Coweeta Hydrologie Laboratory; NC. TYPE DEPOSITORY: AMNH. SEX OF TYPE: M. HOST: Under stones. truncata Barr, 1974: 33, figs. 16A, 16B. GA: Rabun Co., 10 mi. W. of Clayton; GA. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. unicoi Barr, 1974: 27, figs. 11 A, IIB. NC: Graham Co., 0.5 mi. S. of Stratton Meadows; NC. TYPE DEPOSITORY: AMNH. SEX OF TYPE: M. Subfamily GONIACERINAE Reitter, 1882

Bryaxinae LeConte, 1861. Tribe TRICHONYCHINI Reitter, 1882

Trichonyni Raffray, 1890a.

Genus TRICHONYX Chaudoir

Trichonyx Chaudoir, 1845: 164 (as genus). Type-species: Pselaphus sulcicollis Reichenbach (monot.). sulcicollis Reichenbach, 1816: 62 {Pselaphus) (introduced into North America). Germany (Berlin), central Sweden, Austria; NY. TYPE DEPOSITORY: Lost. HOST: Under bark with ants (Park, 1953b); with ants, Ponera pennsyhanicus and Lasius brunneus (Cooper, 1961). PSELAPHIDAE 53

Tribe BRACHYGLUTINI Raffray, 1904b Subtribe BRACHYGLUTINA Raffray, 1904b

Genus RYBAXIS Saulcy

Rybaxis Saulcy, 1876: 96. Type-species: Bryaxis sanguínea Leach (design, by Bowman, 1934: 144) (=longicornis Leach); for treatment of this complicated problem, see Besuchet, 1969, and Newton and Chandler, 1989. Bryaxis Leach, 1817: 85 (as genus, [Rejected Name No. 1953, ICZN, 1969: 133], preoccupied by Bryaxis Kugelann, 1794: 580). TAXONOMY: Fall, 1927 (Nearctic species). KEYS: Fall, 1927 (Nearctic species). appressicornis Park, 1956: 86, figs. 16, 17. TN; TN FL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: At UV light. arkansana Fall, 1927: 226. AR: Lonoke Co., Carlisle; MO/ AR. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. bifalxa Park, 1956: 87, figs. 18-21. AL: Mobile Co., Mount Vernon; AL. TYPE DEPOSITORY: CUIC. SEX OF TYPE: M. HOST: From cypress leaf litter. clavata Brendel, 1865b: 256 (Bryaxis). PA, IL; MN WI MI ON PQ/ IL IN/ NB/ NY PA NJ MD/ VT RI CT. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Moxey, 1962. brendeli LtContQ, 1880: 181 (Bryaxis) (unjustified replacement name). REDESCRIPTION: Fall, 1927. ECOLOGY: Reichle, 1966, 1967, and 1969. HOST: From sphagnum moss; from alder litter on bog edge. conjuncta LeConte, 1849: 85 (Bryaxis). "provinciis orientalibus"; ID/ WI MI ON PQ/ lA IL/ NY NJ MD DC/ ID/ WA/ NS NB/ ME NH VT MA RI CT. TYPE DEPOSITORY: MCZC. REDESCRIPTION: Fall, 1927. ECOLOGY: Reichle, 1966. HOST: From litter on swamp margins and along streams; from sphagnum moss; at UV light. geminata Fall, 1927: 225. PA: Westmoreland Co., St. Vincent; PA. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. mystica Casey, 1894: 476. RI; MI/ IL/ WI ON PQ/ NB NS/ MB SK AB/ NH ME MA RI/ NJ. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Fall, 1927. HOST: From alder and birch leaf litters on bog and marsh margins; from sphagnum moss; at UV light. obliquedens Fall, 1927: 222. PA; lA/ PA VA/ NC. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. 54 COLEÓPTERA CATALOG phantasma Park, 1958: 53, fig. 15. AL: Limestone Co., near Greenbrier; AL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: At light trap. transversa Fall, 1927: 223. MA: Hampden Co., Springfield; AK/ AB SK MB ND/ NT/ OH/ ME NH VT MA CT RI/ NY NJ/ ON PQ MN WI/ NF NB. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. ECOLOGY: Reichle, 1966 and 1969. HOST: From Rhododendron swamp litter, and alder/birch leaf litter on bog margins; from sphagnum moss; at UV light. truncaticornis Brendel, 1890: 274 {Bryaxis) (as variety). lA; lA IL/ AR. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. REDESCRIPTION: Fall, 1927. TAXONOMY: Moxey, 1962. ECOLOGY: Park, Auerbach, and Wilson, 1953. HOST: From hardwood leaf litter; in leaves over nest oí Lasius americanus (Wickham, 1900a). valida Brendel, 1890: 273 {Bryaxis). IL; IL IN / NY NJ/ MA/ NC MS. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Moxey, 1962. sanguínea (of authors). LeConte, 1880: 181 {Bryaxis). This was a misidentification by LeConte, 1880 using the name of a European species. REDESCRIPTION: Fall, 1927. HOST: From hardwood leaf litters; from rotten logs; at UV light. varicornis Brendel, 1890: 274 {Bryaxis) (as variety). Canada; WI MI PQ ON/ NS/ IL/ NY NJ PA VA/ MA RI CT NH ME/ ND. TYPE DEPOSITORY: ANSP. SEX OF TYPE: F. TAXONOMY: Moxey, 1962. REDESCRIPTION: Fall, 1927. ECOLOGY: Reichle, 1966 and 1969. HOST: From leaf litters on bog margins; in grass roots; in sphagnum moss.

Genus BRACHYGLUTA Thomson

Brachygluta Thomson, 1859: 54 (as genus). Type-species: Pselaphus fossulatusKQ\QhQnh?iQh (orig. design.). U.S. species have not been assigned to subgenera in this Holarctic genus. Bryaxis (of authors, not North American) (not Leach, 1817 nor Kugelann, 1794). Nisa CasQy, 1886: 182 (synonymized by Raffray, 1904b). Type-species: Bryaxis cavicornis Brendel (design, by Newton and Chandler, 1989). abdominalis Aube, 1833: 27, pi. 82, fig. 2 {Bryaxis). "Atlantic States": NY PA NJ DC MD VA/ GA/ ME NH MA CT. TYPE DEPOSITORY: MNHP. SEX OF TYPE: M. REDESCRIPTION: Brendel, 1890. HOST: From grass and deciduous leaf litters at margins of salt marshes; under beach drift; at UV light. arguta Casey, 1897: 585 {Bryaxis). MI, PA: Westmoreland Co.; MI ON/ lA OH IL IN/ PA WV. TYPE DEPOSITORY: USNM. HOST: From river drift and pond and lake shore debris; from leaf litter; at UV light. arizonae Casey, 1887: 465 {Bryaxis). AZ: Pima Co., Tucson; CA NV/ AZ. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: At UV light. PSELAPHIDAE 55 belfragei LeConte, 1880: 181 (Bryaxis). TX; MO/ TX OK/ LA AR/ KY/ DC. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. REDESCRIPTION: Brendel, 1890. HOST: On gravel bars in streams and in flood debris; at UV light. cavicornis Brendel, 1865a: 30 (Bryaxis). VA; IN/ DC VA MD NY/ NC. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. TAXONOMY: Chandler, 1994. REDESCRIPTION: Brendel, 1890. HOST: With ant, Myrmica sabuleti, in salt marsh; at UV light. corniventris Motschulsky, 1856: 7 {Bryaxis). AL: Mobile Co., Mobile; AL. TYPE DEPOSITORY: ZMUM. SEX OF TYPE: M. curvicera Motschulsky, 1854: 4 {Bryaxis). NY: New York City; NY TYPE DEPOSITORY: ZMUM. SEX OF TYPE: M. HOST: Under stones on seashore. dentata Say, 1824: 99 {Pselaphus). "United States"; NY NJ DC MD/ MA/ FL ND SC. TYPE DEPOSITORY: Lost. REDESCRIPTION: Brendel, 1890. HOST: Under washed-up log; at UV light. elegans Brendel, 1890: 276, pi. VIII, fig. 48 {Bryaxis). AZ: Coconino Co., Williams; AZ. TYPE DEPOSITORY: ANSP. SEX OF TYPE: F. TAXONOMY: Chandler, 1994. floridana Brendel, 1865b: 257 {Bryaxis). FL: St. John River; MD VA NY/ FL NC SC. TYPE DEPOSITORY: ANSP. SEX OF TYPE: F. TAXONOMY: Moxey, 1962. REDESCRIPTION: Brendel, 1890. HOST: Under washed-up logs and boards on beaches; at UV light. foveata LeConte, 1851: 215 {Bryaxis). CA: mouth of Gila River; CA/ UT/ NV. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. REDESCRIPTION: Brendel, 1893. HOST: At UV light. illinoiensis Brendel, 1865b: 257 {Bryaxis). IL; WI MI/ lA IL IN OH KY MO/ OK/ PA NJ MD/ SC AL MS TN AR. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Moxey, 1962. REDESCRIPTION: Brendel, 1890. TAXONOMY: Chandler, 1994. HOST: Under boards; at UV light. infinita Casey, 1886: 184 {Bryaxis). TX: Travis Co., Austin; TX. TYPE DEPOSITORY: USNM. HOST: At UV light. intermedia Brendel, 1866: 191 {Bryaxis) (as variety). DC; DC NY/ TN NC. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Chandler, 1994. REDESCRIPTION: Brendel, 1890. HOST: At UV light. 56 COLEÓPTERA CATALOG intricata Casey, 1894: 479 {Bryaxis). NY; OH/NY. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: From leaf litter along streams; at UV light. jacobina Casey, 1908: 259. CA: San Diego Co., San Diego; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: Under rocks on grassy hills; in bat guano; at UV light. labyrinthea Casey, 1894: 477 {Bryaxis). NY; NY. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. loripes Casey, 1894: 480 {Bryaxis). Southern CA; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. luniger LeConte, 1849: 87 {Bryaxis). MA: Middlesex Co., Cambridge; NY NJ MD VA DE DC/ FL SC GA/ MA. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. REDESCRIPTION: Brendel, 1890. HOST: In salt marshes; at UV light. mormon Bowman, 1934: 142. UT: Salt Lake Co., Salt Lake City; UT. TYPE DEPOSITORY: ICCM. perpunctata Brendel, 1890: 275 {Bryaxis). GA; GA. TYPE DEPOSITORY: ANSP. SEX OF TYPE: F. terebrata Casey, 1894: 481 {Bryaxis) (replacement name for Bryaxis perforata Brendel). NY; NY CT RI WV MD/ WI ON PQ/ NH MA/ SK/ IL OH. perforata BvQXiáQX, 1866: 192 {Bryaxis) (not Bryaxis perforata Awhe, 1833: 27). NY. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Chandler, 1994. REDESCRIPTION: Brendel, 1890. HOST: From alder and sphagnum litter in marshes; in beaver houses and muskrat nests; in beach debris; at UV light. texana Casey, 1886: 183 {Bryaxis). TX: El Paso Co., El Paso; NM TX. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Brendel, 1890. ulkei Brendel, 1866: 193 {Bryaxis). DC; MD DC VA. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Chandler, 1994. REDESCRIPTION: Brendel, 1890.

Genus REICHENBACHIA Leach

Reichenbachia Leach, 1826: 451. Type-species: Bryaxis juncorum Leach (design, by Bowman, 1934: 144). Bryaxis (of authors, not North American) (not Bryaxis Leach, 1817: 85, or Bryaxis Kugelann, 1794: 580). Reichenbachius Casey, 1906: 359 (unjustified replacement name). TAXONOMY: Grigarick and Schuster, 1967 (revision, western species); Chandler, 1990b (eastern species). KEYS: Grigarick and Schuster, 1967 (western spp.); Casey, 1897; Bowman, 1934 (U. S. spp.); Park and Wagner, 1962 (Pacific Northwest spp.). PSELAPHIDAE 57 albionica Motschulsky, 1845: 43, pi. Ill, fig. 2 (Bryaxis). Sitka and northern CA; AK/ WA OR ID/ CA. TYPE DEPOSITORY: ANSP, ZMUM. compar LeConte, 1851: 215 {Bryaxis) (synonymized by Grigarick and Schuster, 1971). CA: Santa Clara Co., San Jose. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. franciscana Casey, 1886: 193 (synonymized with compar by Brendel, 1893). CA: San Mateo Co. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. fundata Casey, 1886: 192 (synonymized by Brendel, 1894b). CA: Sonoma Co. TYPE DEPOSITORY: USNM. americana Kdifxdiy, 1904b: 241 (unjustified replacement name). REDESCRIPTION: Grigarick and Schuster, 1967. HOST: Collected around cattails; at UV light; from cottonwood and willow leaf litter; from leaf litter along slough; sweeping Bermuda grass. appendiculata appendiculata Raffray, 1904b: 174, fig. 70 (other subspecies from Panama (Park, 1942)). Mex.; TX; Mex., Central America. TYPE DEPOSITORY: MNHP. SEX OF TYPE: M. REDESCRIPTION: Park, 1942, 1945. HOST: At UV light. arcifer Casey, 1897: 604. KS; KS/ PQ/ OK. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Chandler, 1989 (aedeagus). HOST: In beaver houses; at UV light. arthritica Brendel, 1894a: 159, pi. V, fig. 7 (Bryaxis). CA: San Luis Obispo Co., San Luis Obispo; CA. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1967.

atlántica Brendel, 1866: 193 (Bryaxis). LA; LA SC FL/ DC. TYPE DEPOSITORY: ANSP. SEX OF TYPE: F. TAXONOMY: Chandler, 1994. REDESCRIPTION: Chandler, 1989 (aedeagus). HOST: From mixed hardwood litter; at UV light.

bicolor Brendel, 1890: 294 (Bryaxis). lA: Linn Co., Cedar Rapids; lA NE IL/ MN/ MA/ NY MD/NCGA. TYPE DEPOSITORY: ANSP. SEX OF TYPE: F. TAXONOMY: Moxey, 1962. subsimilis Casey, 1897: 594 (synonymized by Chandler, 1989). lA: Johnson Co., Iowa City. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Chandler, 1989 (aedeagus). ECOLOGY: Park, Auerbach, and Wilson, 1949 and 1953, and Mickey and Park, 1956 (as subsimilis). HOST: Under stones.

binodifer Casey, 1897: 608, fig. B, 2. BC: Vancouver Isl.; BC OR WA/ AK /AB MT/ CA NV. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1967. HOST: From grass litter and rotten wood by spring (Chandler, 1983); from boxelder leaf litter in grass; from Labrador tea leaf litter; in sphagnum moss in marshes and bogs; at UV light. 58 COLEÓPTERA CATALOG borealis Casey, 1897: 601. ON: Ottawa; MN WI MI ON PQ/ MB AB/ IL IN NE/ PE NB NS/ RI ME NH MA CT/ PA DC VA NY NJ/ NC. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. distans Casey, 1897: 602 (synonymized by Chandler, 1989). ON: Ottawa. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Chandler, 1989 (aedeagus). ECOLOGY: Reichle, 1966, 1967, and 1969. HOST: Primarily from mosses on logs and bog margins; at UV light. caseyi Raffray, 1904b: 240 (replacement name for Reichenbachia articularis Casey); WY UT CO. articular is C2LSQy, 1897: 608, fig. B, 3; C, 1 (preoccupied Raffray, 1893: 494). Southern CO. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1967. HOST: Under pine bark, in moss, and in ant nests. chiricahuensis Raffray, 1904b: 172. AZ: Cochise Co., Chiricahua Mts.; AZ NM. TYPE DEPOSITORY: MNHP. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1967. HOST: In pine leaf litter; at UV light. clamer Grigarick and Schuster, 1967: 10, figs. 50-53. CA: Mendocino Co., Mendocino; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: In shore debris and under logs on beaches; at UV light. complectens LeConte, 1880: 183 (Bryaxis). TX; TX; Mex. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. dilatipes Schaeffer, 1906: 262 (synonymized by Chandler, 1989). TX: Comal Co., New Braunfels. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. latipes Fletcher, 1928: 212, pi. XII, figs. 9, 29 (synonymized by Chandler, 1989). Mex.: Veracruz. TYPE DEPOSITORY: CUIC. SEX OF TYPE: M. REDESCRIPTION: Park, 1945; Chandler, 1989 (aedeagus). HOST: At UV light. congener Brendel, 1865b: 257 {Bryaxis). NY: Long Island; lA IL/ PQ/ NY PA NJ/ MA RI/ FL MS/ OK. TYPE DEPOSITORY: ANSP. SEX OF TYPE: F. TAXONOMY: Moxey, 1962. REDESCRIPTION: Chandler, 1989 (aedeagus). HOST: From maple stumps; in flood debris; at UV light. corporalis Casey, 1897: 600. MI, ON: Ottawa; MI ON WI PQ/ ME MA CT/ NJ NY/ TN. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Chandler, 1989 (aedeagus). ECOLOGY: Reichle, 1969. HOST: From sphagnum moss; from leaf litter of alde^^irch mixture on marsh edge; at UV light. PSELAPHIDAE 59 cylindrarta Brendel, 1893: 279, pi. IV, fig. 18 (Bryaxis). North IL; IL OH/ ON/ DC/ DC MS/ OK. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Moxey, 1962. REDESCRIPTION: Chandler, 1989 (aedeagus). HOST: From oak leaf litter by water; from swamp litter; in rotten logs. deformata LeConte, 1880: 183 (Bryaxis). CA: Los Angeles Co., Los Angeles; CA. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1967. demissa Casey, 1894: 485. DC; DC/ FL. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. REDESCRIPTION: Chandler, 1989 (aedeagus). HOST: From litter of sand pine, oak, and palmetto. depressifrons Brendel, 1893: 279, pi. IV, fig. 17 (Bryaxis). CA: Alameda Co.; CA. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1967. HOST: At UV light.

divergens LeConte, 1880: 182 (Bryaxis). MA; ON MI/ lA IN IL OH/ MA RI NH/ NJ DC VA NY PA/ VA NC. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. cribricollis BYQndQl, 1890: 291 (Bryaxis) (synonymized by Chandler, 1989). IL, lA. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Moxey, 1962 (designated lectotype; type locality not restricted; lectotype lacks locality label). furtiva Casey, 1894: 483 (synonymized by Chandler, 1989). PA. TYPE DEPOSITORY: USNM. SEX OF-TYPE: M. fusciventris Casey, 1897: 599 (synonymized by Chandler, 1989). NY. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Chandler, 1989 (aedeagus). HOST: From grass litter in salt marsh; leaf litter around pond margin.

dorothyae Park and Wagner, 1962: 26, pi. VI, fig. 1-2. OR: Bear Springs; OR/ CA. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1967. HOST: From alder leaf litter.

ectofacilis Park, 1956: 91, figs. 25, 26. AL: Mobile Co., Mount Vernon; AL FL LA MD/ TX. TYPE DEPOSITORY: CUIC. SEX OF TYPE: M. HOST: From river drift; at UV light.

emoyteis Grigarick and Schuster, 1967: 13, figs. 71-74. CA: Mariposa Co., Happy Isls., Yosemite National Park; OR/ CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: Beating pine in meadow. 60 COLEÓPTERA CATALOG

facilis Casey, 1884b: 87 {Bryaxis). NY: Long Island, Willets Point; NY NJ/ ON/ lA IL 7 MA CT RI/ GA MS. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Park, 1956. HOST: From leaf litters of oak, beech, and maple; from mixture of alder and birch leaf litter at marsh edges* from river drift; at UV light. '

falli Casey, 1897: 606, fig. A, 2. CA: Los Angeles Co., Santa Monica; OR/ CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1967. HOST: Under log on beach.

fusticornis Casey, 1897: 608, figs. B, 5; C, 3. CA: northern coast region; OR WA BC/ MT AB/ CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1967. HOST: From bunch grass leaf litter in spring area (Chandler, 1983); also from alder litter, and rotten wood.

gemmifer LeConte, 1880: 182 {Bryaxis). MI; MI ON/ lA IN IL; NY DC VA/ MS LA AR. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. canadensis Brendel, 1887: 206 {Bryaxis) (synonymized by Chandler, 1989). "Canada." TYPE DEPOSITORY: ANSP. SEX OF TYPE: F. TAXONOMY: Moxey, 1962. REDESCRIPTION: Brendel, 1887; Chandler, 1989 (aedeagus).

gentilis Fletcher, 1932: 33. CA; CA NV/ UT. TYPE DEPOSITORY: ANSP. SEX OF TYPE: F. utahensis Tanner, 1934: 43, fig. 1 (synonymized by Grigarick and Schuster, 1971). UT: Washington Co., Zion National Park. TYPE DEPOSITORY: BYUC. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1967. gracilicornis Casey, 1886: 190. TX: Travis Co., Austin; TX OK/ MS. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Chandler, 1989 (aedeagus). HOST: From rotten wood;4n flood debris; at UV light. gracilis Casey, 1884a: 32 {Bryaxis). ML Wayne Co., Detroit; MI/ IN OH MO KS/ LA. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. ursina Casey, 1897: 596 (synonymized by Chandler, 1989). Western MO. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Chandler, 1989 (aedeagus). HOST: From leaf litter. hardyi Park, 1956: 89, fig. 22. FL: Palm Beach Co., W. Palm Beach, Morrison Field; FL SC GA AR LA/ TX OK/ PA DC VA. TYPE DEPOSITORY: SEMC. SEX OF TYPE: M. HOST: From sand pine and oak litter; from maple stumps; at UV light. PSELAPHIDAE 61 howardi Park, 1958: 46, fig. 4. LA: Lake Pontchartrain, near New Orleans; LA FL GA KY TN / IL/ NJ/ TX. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Beneath littoral debris; from vegetation in marsh; at UV light. ignobilis Fletcher, 1932: 32. AZ: Cochise Co., San Bernardino Ranch; AZ; Mex. TYPE DEPOSITORY: SEMC. SEX OF TYPE: M. sonorensis ??irk, 1945: 371, pi. VII, figs. 4-5 (synonymized by Park, 1958). Mex.: Sonora, Yaqui Valley. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1967. HOST: At UV light. inepta Casey, 1894: 484. NJ; NJ/ GA/ OK. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. REDESCRIPTION: Chandler, 1989 (aedeagus). HOST: From pine litter; at UV light. informis Casey, 1886: 189. CA: Mendocino Co.; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1967. inopia Casey, 1884a: 33 (Bryaxis). FL: Sand Point; FL. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. REDESCRIPTION: Park, 1958. insólita Casey, 1897: 597. RI: Boston Neck; RI. TYPE DEPOSITORY: USNM. REDESCRIPTION: Chandler, 1989 (aedeagus). HOST: From swamp litter.

liansana Casey, 1897: 595. KS; KS/ MS. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Chandler, 1989 (aedeagus). HOST: From deciduous leaf litter.

kapones Grigarick and Schuster, 1967: 11, figs. 59-62. WA: Spokane Co., Mount Spokane; WA/ MT AB/ BC. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M.

Utoralis Brendel, 1890: 296 (Brycais). DC; DC VA/ TX OK/ FL AL TN LA MS/ IL. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Chandler, 1994. . turba Park, 1958: 52, fig. 14 (synonymized by Chandler, 1989). AL: Limestone Co., near Greenbrier. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From sand pine and oak litter; at UV light.

nevadensis Casey, 1886: 191. NV: Washoe Co., Reno; NV. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. 62 COLEÓPTERA CATALOG

nodesa Grigarick and Schuster, 1967: 14, figs. 82-87. AZ: Coconino Co., Sedona; AZ. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: At UV light

peregrinator Casey, 1897: 599. lA: Iowa City; ND SD MB/ WI MI/ NE lA IL OH/ PA/ SC. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. shenefelti Park, 1958: 50, fig. 13 (synonymized by Chandler, 1989). WI: Grant Co., Boscobel, State Nursery. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. ECOLOGY: Park, Auerbach, and Wilson, 1953; Mickey and Park, 1956. HOST: From river drift; at UV light.

polita Brendel, 1890: 297 {Bryaxis). NY; NY PA NJ/ lA IN MO/ NH MA/ FL. TYPE DEPOSITORY: ANSP. SEX OF TYPE: F. TAXONOMY: Moxey, 1962. procera C?iSQy, 1897: 603 (synonymized by Chandler, 1989). lA?. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. pygidialis Raffray, 1904b: 160 (synonymized by Chandler, 1989). MO. TYPE DEPOSITORY: MNHP. SEX OF TYPE: F. REDESCRIPTION: Park, 1958 {^s procera Casey); Chandler, 1989 (aedeagus). HOST: In sphagnum moss. propinqua LeConte, 1849: 88 {Bryaxis). "Lake Superior"; MI ON PQ/ NS/ NH MA CT/ PA MD/ MB ND. TYPE DEPOSITORY: MCZC. REDESCRIPTION: Chandler, 1989 (aedeagus). HOST: From cedar leaf litter near lake; from litter at marsh edges; from moss. puncticoUis LeConte, 1849: 87 {Bryaxis). LA: Orleans Parish, New Orleans; LA FL NC SC GA MS/ TX/ PA. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. howdeni Park, 1958: 47, fig. 6 (synonymized by Chandler, 1989). NC: Hyde Co., Swanquarter. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. ECOLOGY: Reichle, 1966 and 1969. HOST: From sphagnum moss; at UV light. radians LeConte, 1880: 182 {Bryaxis). IL; IL IN MO/ MI/ NH/ OK. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. trigona LeConte, 1880: 183 {Bryaxis) (synonymized by Chandler, 1989). MO. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. trigma (error) Brendel, 1890: 294 {Bryaxis). pluridentata Park, 1949: 322, pi. Ill (aedeagus figured in Park 1942: pi. I, figs. 2-3, but without accompanying written description) (synonymized by Chandler, 1989). IL: Champaign Co., Urbana, Brownfield Woods. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Park, 1958. HOST: Beneath piece of rotten wood. PSELAPHIDAE 63 sagax LeConte, 1880: 183 {Bryaxis). CA: Los Angeles Co., Los Angeles; CA. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1967. HOST: From hillside seepage area. scabra Brendel, 1865a: 29 {Bryaxis). NY: Long Island; MN WI MI/ ON/ CT RI/ IL IN OH/ NY NJ. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Moxey, 1962. REDESCRIPTION: Chandler, 1989 (aedeagus). ECOLOGY: Reichle, 1966 and 1969. HOST: From mixture of alder and birch leaf litters at marsh and bog margins; in sphagnum moss.

semirugosa Brendel, 1895: 183. PA: Westmoreland Co., St. Vincent; PA. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. TAXONOMY: Chandler, 1994.

snowi Fletcher, 1932: 31. AZ: Coconino Co., Oak Creek Canyon; AZ. TYPE DEPOSITORY: SEMC. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1967. HOST: From leaf litter in dense grass; from leaf litter along streams.

sodalis Casey, 1897: 598. AR: Pulaski Co., Little Rock; AR MS GA SC. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Chandler, 1989 (aedeagus). HOST: From hardwood leaf litter.

spatulifer Casey, 1897: 609, fig. B, 4. ON: Ottawa; WA BC/ CA/ AB SK MB/ YT/ MN WI MI ON PQ/ lA IL/ NF NB/ ME. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1967. ECOLOGY: Reichle, 1966 and 1969. HOST: Primarily from sphagnum mosses; also from alder litter on bog margins.

subtilis LeConte, 1851: 215 {Bryaxis). CA: mouth of Gila River; CA. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. REDESCRIPTION: Grigarick and Schuster, 1967.

taphrocera Casey, 1897: 605. CA: Monterey Co.; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M.

túmida LeConte, 1880: 183 {Bryaxis). TX; TX; Mex. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. pattersoniY2iXk, 1958: 49, figs. 9, 10, 11 (synonymized by Chandler, 1989). TX: Presidio CO., Porvenir. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: At UV light.

tumidicornis Casey, 1886: 187. CA: Santa Cruz and Santa Clara Cos.; CA/ OR. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1967. HOST: Sweeping Bermuda grass; in flood debris along river; in moss. 64 COLEÓPTERA CATALOG

tumorosa Casey, 1886: 186. CA: Sonoma Co.; CA. TYPE DEPOSITORY: USNM. REDESCRIPTION: Grigarick and Schuster, 1967. HOST: From Salix, Rubus, and alder litter near streams; from grass litter in bogs; at UV light. turgidicornis Casey, 1897: 607, fig. A, 1. CA: Los Angeles Co., Santa Monica; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1967. HOST: At UV light. visda Grigarick and Schuster, 1967: 9, figs. 38, 39. CA: Yolo Co., Davis; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: At UV light. wickhami Brendel, 1893: 280, pi. IV, fig. 19 {Bryaxis). CO: Fremont Co., Canyon City; CO/ AZ NM/ KS. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Moxey, 1962. REDESCRIPTION: Grigarick and Schuster, 1967. HOST: At UV light.

Unplaced Species rubricunda Aube, 1844: 116 {Bryaxis). PA; IN/ NY PA/ CT/ GA FL. rubicunda (error), of authors. TYPE DEPOSITORY: Lost? Not in MNHP or ZSBS (Chandler, 1989).

Genus BRIARAXIS Brendel

Briaraxis Brendel, 1894a: 158, pi. V, fig. 6 (as genus). Type-species: Briaraxis depressaBxQnáéi (monot). Briaraxys Raffray, 1908: 219 (error). TAXONOMY: Chandler, 1992a. REDESCRIPTION: Chandler, 1992a. depressa Brendel, 1894a: 159, pi. V, fig. 6. FL: Monroe Co., Dry Tortugas; FL, Caribbean. TYPE DEPOSITORY: ANSP. SEX OF TYPE: F. REDESCRIPTION: Chandler, 1992a. HOST: Under beach debris; at UV light.

Genus NISAXIS Casey Nisaxis Casey, 1886: 183 (as genus). Type-species: Bryaxis tomentosa Aube (monot). Nisaxys {QYVOX) Raffray, 1890a: 124. caudata Schaeffer, 1905: 124. TX: Cameron Co., Brownsville, Esperanza Ranch; OK TX; Mex. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: At UV light. marítima Casey, 1887: 468. TX: Galveston Co., Galveston; TX/ LA MS. TYPE DEPOSITORY: USNM. REDESCRIPTION: Park, 1958. HOST: Beneath seashore beach drift; at UV light in salt marshes. parviceps Casey, 1897: 610. NE; NE. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. PSELAPHIDAE 65 tomentosa Aube, 1833: 33, pi. 84, fig. 4 (Bryaxis). "America septentrionali"; NE IL/ NY NJ DE DC MD/ CT/ TX OK/ LA AL MS NC FL; Caribbean. TYPE DEPOSITORY: MNHP? minuta Brendel, 1865^ 30 (Bryaxis) (synonymized by Casey, 1897). NY. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. TAXONOMY: Chandler, 1994. cincinnata Casey, 1887: 466, pi. XVI, fig. 1 (as subspecies) (synonymized by Park, 1958). TX: Galveston Co., Galveston. TYPE DEPOSITORY: USNM. REDESCRIPTION: Park, 1958. HOST: Under seashore beach drift; at UV light. Subtribe PSELAPTINA Park, 1976

Genus EUTRICHITES LeConte

Eutrichites LeConte, 1880: 184 (as genus). Type-species: Eutrichites zimmermanni LQCOíúQ (monot.) (=zonatus Brendel). TAXONOMY: Cariton, 1989b (revision). REDESCRIPTION: Cariton, 1989b. KEYS: Cariton, 1989b.

arizonensis Cariton, 1989b: 113, figs. 5, 10, 18, 20. AZ: Yavapai Co., Verde River, Cottonwood; AZ/ CA; Mex. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: At UV light.

zonatus Brendel, 1865a: 28 {Bythinus), LA; WI MN ON PQ/ MO IL IN/ DC VA DE MD WV/ FL AL TN AR SC NC GA MS LA/ TX OK/ NH. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. zimmermanni LQCOïUQ, 1880: 184 (synonymized by Brendel, 1890). TX. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. REDESCRIPTION: Cariton, 1989b. HOST: From grass debris; in sawdust; from old river drift; at UV light.

Genus ANCHYLARTHRON Brendel

Anchylarthron Brendel, 1887: 208 (as genus). Type-species: Decarthron cornutum Brendel (monot.). Verticinotus Brendel, 1888a: 300 (as genus; not Brendel, 1888b). Type-species: Decarthron cornutum Brendel (monot.). Brendel published the same paper tw^ice with 1888a published in June 1888 and 1888b published in November 1888.

caviceps Casey, 1897: 612. FL: Palm Beach Co., Lake Worth; FL GA MS AL. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: From leaf litter; at UV light.

cornutum Brendel, 1865a: 31 {Decarthron). IL; KS lA IL/ TX/ SC. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. TAXONOMY: Chandler, 1994. HOST: From leaf litter; at UV light.

curtipenne Casey, 1897: 611. lA; lA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. 66 COLEÓPTERA CATALOG

inornatum Brendel, 1865b: 258 {Bryaxis). SC. Moxey (1962) lists type-locality as KS, but Brendel mentions only SC in description. TYPE DEPOSITORY: Lost? (Chandler, 1994). SEX OF TYPE: F.

Genus SCALENARTHRUS LeConte

Scalenarthrus LeConte, 1880: 185 (as genus). Type-species: Scalenarthrus horni LQCOTUQ (monot.). Cylindrembolus L. W. Schaufiiss, 1887: 103 (as genus) (synonymized by Raffray, 1904b). Type-species: Cylindrembolus marginalis L. W. Schaufiiss (monot.). Abryxis Raffray, 1890a: 124 (as genus; synonymized by Raffray, 1904b). Type-species: Bryaxis denticornis L. W. Schaufuss (design, by Newton and Chandler, 1980: 46). horni LeConte, 1880: 185. AZ; CA/AZ; Mex. TYPE DEPOSITORY: MCZC. HOST: With ant, Solenopsis aurea; from oak and sycamore leaf litters; under low plants in corn fields.

Genus PSELAPTUS LeConte

Pselaptus LeConte, 1880: 184 (as genus). Type-species: Pselaptus belfragei LeConte (monot.).

belfragei LeConte, 1880: 185. TX; CA/AZ TX. TYPE DEPOSITORY: MCZC. HOST: At UV light.

Subtribe DECARTHRINA Park, 1951

(Decarthonina Park, 1951, incorrect original spelling).

Genus DECARTHRON Brendel

Decarthron Brendel, 1865a: 30 (as genus, other subgenus Neotropical). Type-species: Bryaxis formiceti LeConte (design, by Bowman, 1934: 144). TAXONOMY: Park, 1958. REDESCRIPTION: Park, 1958. KEYS: Park, 1958.

Subgenus DECARTHRON Brendel abnorme LeConte, 1849: 89 (Bryaxis). "provinciis mediis Australibus et Borealibus"; MN WI MI ON PQ/ lA IL/ FL/ ME NH MA RI CT/ NY PA NJ/ NT/ AB MB SK/ WA BC. TYPE DEPOSITORY: MCZC. REDESCRIPTION: Park, 1958 (aedeagus). ECOLOGY: Reichle, 1966 and 1969. HOST: From leaf and moss litters on margins of streams, ponds, and marshes; from sphagnum moss; at UV light. anneae Park, 1958: 57, fig. 21. NC: Moore Co., Southern Pines; OH/ VA/ NC AL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: In moss; from swamp litter. brendeli Casey, 1887: 464. TX: Galveston Co., Galveston; IN/ TX OK/ MS AL. TYPE DEPOSITORY: USNM. REDESCRIPTION: Park, 1958 (aedeagus). HOST: At UV light. PSELAPHIDAE 67 defectum Park, 1958: 64, fig. 31. WI: Grant Co., Boscobel, State Nursery; MN WI MI ON/ IL/ NY PA. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. ECOLOGY: Reichle, 1966, 1967, and 1969. HOST: Floating vegetation layer of tamarack bog; at UV light; from sphagnum moss. discolor Brendel, 1890: 265. NM: Bemalillo Co., Albuquerque; NM AZ. TYPE DEPOSITORY: ANSP. SEX OF TYPE: F. TAXONOMY: Moxey, 1962. HOST: From leaf litter; at UV light. diversum Park, 1958: 63, fig. 29. FL: Palm Beach Co., W. Palm Beach, Morrison Field; LA FL MS. TYPE DEPOSITORY: SEMC. SEX OF TYPE: M. HOST: In marshes; at UV light. exiguum Notman, 1920: 187. NY: Broome Co., Windsor; NY. TYPE DEPOSITORY: Staten Isl. Museum. SEX OF TYPE: F. exsectum Brendel, 1865a: 32. NY; NY PA NJ/ FL. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Moxey, 1962. REDESCRIPTION: Park, 1958 (aedeagus). formiceti LeConte, 1849: 90 (Bryaxis). PA; PA NY WV DC/ CT NH MA/ TX/ LA AL FL GA MS/ ON. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. velutinum LeConte, 1849: 86 (Bryaxis) (synonymized by Brendel, 1865b). LA: Orleans Parish, New Orleans. TYPE DEPOSITORY: MCZC. HOST: With ant, Formica pennsylvanica; from sand pine leaf litter; at UV light. howdeni Park, 1956: 93, figs. 27-29. NC: Moore Co., Southern Pines; NJ/ MS GA NC FL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: In sphagnum moss; at UV light. inusitatum Park, 1958: 61, figs. 26, 27, 28. AL: Mobile Co., Mount Vemon; MS AL NC SC TN/ ON PQ/ KY IL lA/ OK/ NY/ MA. TYPE DEPOSITORY: CUIC. SEX OF TYPE: M. HOST: From cypress litter at lake margin; from hardwood litter in ravine; at UV light. justum Park, 1958: 59, fig. 23. VA: Fairfax Co.; VA NJ DC/ FL NC. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From leaf litters of cypress and of sand pine/oak mixtures; at UV light.

laurenticum Casey, 1897: 584. MA, ON: Ottawa; ON/ MA NH/ PA NY NJ. TYPE DEPOSITORY: USNM. REDESCRIPTION: Park, 1958 (aedeagus). HOST: From moss and grass in bogs.

longulum LeConte, 1849: 89 (Bryaxis). NY, "Lake Superior"; WI ON PQ/ NY NJ DC/ MA CT. TYPE DEPOSITORY: MCZC. REDESCRIPTION: Park, 1958 (aedeagus). HOST: From moss; at UV light. 68 COLEÓPTERA CATALOG

marinum Brendel, 1893: 278. MS: Gulfview; LA MS. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Moxey, 1962.

mimicum Park, 1958: 63, fig. 30. GA: Tifton Co., Tifton; KY/ GA FL LA. TYPE DEPOSITORY: SEMC. SEX OF TYPE: M. HOST: From litter along stream margins; at UV light. nigrocavum Park, 1958: 56, fig. 18. NC: Hyde Co., Swanquarter; LA NC/ IL/ OK. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Under log on lake shore; in straw pile; in flood debris; at UV light. rayi Park, 1949: 325, pi. IV. IL: Pope Co., Eddyville; IL/ GA AL LA/ VA NY. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Park, 1958 (aedeagus). HOST: From rotten logs; under logs on lake shore; at UV light. reciprocum Park, 1958: 60, fig. 24. MS: George Co., Lucedale; MS SC FL LA. TYPE DEPOSITORY: CUIC. SEX OF TYPE: M. HOST: From hardwood leaf litter; at UV light. rostratrum Park, 1958: 57, fig. 20. FL: Pinellas Co., Tarpon Springs; FL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From maple logs; at UV light. scarificatum Brendel, 1893: 278. lA: Linn Co., Cedar Rapids; lA. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. seriepunctatum Brendel, 1893: 278. TN; TN. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. setosum Bowman, 1934: 142. PA: Franklin Co., Penn-Mar; PA. TYPE DEPOSITORY: ICCM. SEX OF TYPE: F. snowi Park, 1958: 58, fig. 22. AL: Limestone Co., near Greenbrier; AL TN. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: At UV light. stigmosum Brendel, 1865a: 31. NY; IL lA/ ON/ NY PA NJ/ VT MA CT NH/ FL. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. TAXONOMY: Chandler, 1994. strenuum Brendel, 1865a: 32 (synonymized by Chandler, 1994). PA. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. REDESCRIPTION: Park, 1958. HOST: With ants, Aphaenogaster fiiha, A. treati (Schwarz, 1890); with Formica fusca; at UV light. varicum Park, 1958: 61, fig. 25. MS: George Co., Lucedale; MS FL AL/ OK. TYPE DEPOSITORY: CUIC. SEX OF TYPE: M. HOST: From mixtures of leaf litters of sand pine and palmetto, and of leaf litters of beech and magnolia- at UV light. PSELAPHIDAE 69

Subtribe EUPSENIINA Park, 1951

Genus EUPSENIUS LeConte

Eupsenius LeConte, 1849: 90 (as genus). Type-species: Eupsenius glaber LeConte (monot.). Euplenius (error) Motschulsky, 1855: 16. Eupsinus (error) Motschulsky, 1856: 7. Euprenius (error) Raffray, 1897: 262. KEYS: Park et al., 1976 (Southeast U.S. and Caribbean). dilatatus Motschulsky, 1856: 7. AL: Mobile Co., Mobile; AL. TYPE DEPOSITORY: ZMUM. glaber LeConte, 1849: 90. SC: Richland Co., Columbia; IL/ LA AL MS GA SC FL NC/ TX, Bahamas. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. REDESCRIPTION: Brendel, 1893. HOST: From hardwood litter; at UV light. rufus LeConte, 1863b: 28. AL: Mobile Co., Mobile; AL FL. Motschulsky (1856: 7) originally listed this name without a description, and is a nomen nudum. LeConte validated this name (1863b: 28), basing his description on a specimen sent by Motschulsky. TYPE DEPOSITORY: MCZC. Tribe INIOCYPHINI Park, 1951 Tanypleurini Jeannel, 1949. Subtribe NATYPLEURINA Newton and Thayer, 1992 Genus BYTHINOPHYSIS Raffray

Bythinophysis Raffray, 1908: 266. Type-species: Bythinophysis punctipennis Raffray (monot.). Dalmodes (of authors, not North American) (not Reitter, 1882: 210). REDESCRIPTION: Park, 1942. KEYS: Park, 1942. schaufussi Raffray, 1897: 267 {Dalmodes). Mex.: Tabasco, Teapa. New Orleans, LA, record is probably an error. TYPE DEPOSITORY: MNHP. SEX OF TYPE: M. Tribe BYTHININI Raffray, 1890a

Subtribe BYTHININA Raffray, 1890a

Genus MACHAERODES Brendel

Machaerodes Brendel, 1890: 252 (as genus). Type-species: Bythinus carinatus Brendel (design, by Lucas, 1920: 386). Bythinus (of authors, not North American) (not Bythinus Leach, 1817: 82).

carinatus Brendel, 1865a: 29 {Bythinus). PA: ON/ OH/ PA VA/ TN NC WV GA. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Chandler, 1994. HOST: From leaf litters of pine, oak, Rhododendron, and beech. 70 COLEÓPTERA CATALOG

Subtribe XENOBYTHINA Jeannel, 1950 Genus PSELAPTRICHUS Brendel

Pselaptrichus Brendel, 1889: 194 (as genus). Type-species: Pselaptrichus tuberculipalpus Brendel (monot.). TAXONOMY: Schuster and Marsh, 1956 (revision). REDESCRIPTION: Marsh and Schuster, 1954. KEYS: Schuster and Marsh, 1956; Park and Wagner, 1962 (Pacific Northwest).

Subgenus PSELAPTRICHUS Brendel

auctumnus Chandler, 1983: 229, fig. 14. CA: Butte Co., 3 mi. NE. of Forest Ranch; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From forest floor litter.

burdicki Schuster and Marsh, 1956: 144, fig. 70. CA: Santa Clara Co., Mount Madonna; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M.

carinatus Marsh and Schuster, 1954: 16, figs. 4, 10, 21. CA: Del Norte Co., 18 mi. S of Klamath; CA/ OR. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From leaf litters of redwood, pine, and black oak.

cavatus Marsh and Schuster, 1954: 21, figs. 7, 11, 17, 24. CA: Humboldt Co., Freshwater; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From redwood leaf litter; from alder logs.

chandleri Schuster and Marsh, 1956: 130, figs. 12, 50. CA: Shasta Co., McArthur Bumey State Park; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M.

cornus Chandler, 1983: 228, fig. 13. CA: Butte Co., Mountain House; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From dogwood leaf litter; from leaf litter by streams; from ponderosa pine stump.

cuspidatus Schuster and Marsh, 1956: 145, fig. 73. CA: Monterey Co., 7 mi. S. of Big Sur; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From redwood litter.

frigidus Schuster and Marsh, 1956: 123, figs. 5, 43. CA: Mariposa Co., Yosemite National Park; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. gibbosus Marsh and Schuster, 1954: 23, figs. 8, 14, 25. CA: Humboldt Co., 14 mi. E. of Blue Lake; OR/ CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From redwood and Sitka-spruce leaf litters. helferi Schuster and Marsh, 1956: 137, figs. 20, 35, 55. CA: Mendocino Co., Mendocino; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From swamp litter. PSELAPHIDAE 71 hocus Schuster and Marsh, 1956: 144, fig. 71. CA: Monterey Co., Bixby Creek, 8 mi. N. of Big Sur; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. incognitus Schuster and Marsh, 1956: 124, figs. 6, 27, 44. CA: Eldorado Co., Riverton; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From mixed conifer litter. intimus Schuster and Marsh, 1956: 127, figs. 9, 30, 47. OR: Big Meadows, near Santiam Pass; OR. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From Douglas fir, silver fir, and alder leaf litters; in moss mixed with conifer leaf litter.

levinei Schuster and Marsh, 1956: 138, figs. 21, 36, 56. CA: Eldorado Co., Riverton; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From Douglas-fir leaf litter.

magaliae Chandler, 1983: 228, fig. 12. CA: Butte Co., 2 mi. N. of Hurleton, Stringtown Road; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From moist leaf litters by streams.

marshi Chandler, 1983: 227, fig. 11. CA: Butte Co., Mountain House; CA. TYPE DEPOSITORY: CA. SEX OF TYPE: M. HOST: From wet leaf litter by springs.

minimus Schuster and Marsh, 1956: 140, figs. 23, 38, 58. CA: Lake Co., Clear Lake, Sulfiir Bank Point; CA. TYPE DEPOSITORY: CASC- SEX OF TYPE: M. HOST: From toyon, laurel, and knobcone pine leaf litters.

oculatus Marsh and Schuster, 1954: 18, figs. 5, 12, 22. CA: Mariposa Co., Wawona; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From manzanita and ponderosa pine leaf litters.

ornatus Marsh and Schuster, 1954: 20, figs. 6, 13, 18, 23. CA: Mendocino Co., Hartsook Grove; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. / , / , HOST: From leaf litters of redwood, Douglas fir, big-leaf maple, alder, and mixtures of dogwood/oak/maple.

parki Schuster and Marsh, 1956: 139, figs. 22, 37, 57. CA: Mariposa Co., 14 mi. E. of Briceburg; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From oak, moss, and fern litter.

pennatus Schuster and Marsh, 1956: 133, figs. 16, 52, 62, 63, 64. CA: Marin Co., near southern entrance of Samuel P. Taylor State Park; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From redwood, laurel, and fern leaf litters. 72 COLEÓPTERA CATALOG

perditus Schuster and Marsh, 1956: 128, figs. 11, 31, 49. OR: Linn Co., Detroit; OR. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From hemlock leaf litter and from moss with mixed conifer litter.

perfidus Schuster and Marsh, 1956: 126, figs. 8, 29, 46. OR: Clatsop Co., Seaside; WA OR. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From mixed conifer litters; in moss.

plusculus Park and Wagner, 1962: 28, pi. VII, figs. 9-10. OR: Coos Co., Charleston; OR. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M.

propinquus Schuster and Marsh, 1956: 131, figs. 14, 65. CA: Trinity Co., 11 mi. E. of Douglas City; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From leaf litters of pine, black oak, and Douglas fir.

proprius Schuster and Marsh, 1956: 126, figs. 7, 28, 45. CA: Del Norte Co., 2 mi. N. of Fort Dick; OR/ CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From leaf litters of Douglas fir, redwood, and Eurhynchium.

rectus Marsh and Schuster, 1954: 13, fig. 2. CA: Marin Co., Alpine Lake; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From leaf litters of redwood, Douglas fir, and laurel.

rothi Park, 1953a: 256, pi. 1, B; 2, A, F; 3, C. OR: Benton Co., 8 mi. N. of Corvallis, MacDonald Forest; WA OR/ CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From leaf litters of moss, cedar, hemlock, maple, oak, and Douglas fir.

shastensis Schuster and Marsh, 1956: 130, figs. 13, 32, 51. CA: Siskiyou Co., 8 mi. S. of Dunsmuir; CA/ OR. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From leaf litters of maple, Douglas fir, oak, Jeffrey and Oregon pine, and willow.

silvanus Schuster and Marsh, 1956: 141, fig. 66. CA: Marin Co., Inverness; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From laurel and Lithocarpus leaf litters.

similis Schuster and Marsh, 1956: 145, fig. 72. CA: Monterey Co., near Point Sur; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From redwood litter. spinosus Marsh and Schuster, 1954: 15, figs. 3, 9, 20. CA: Santa Cruz Co., Ben Lomond; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From leaf litters of redwood, oak, Douglas fir, and sword fern. tenuis Schuster and Marsh, 1956: 132. CA: Mendocino Co., Mendocino; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From redwood and alder leaf litters. PSELAPHIDAE 73

íuberculipalpus Brendel, 1889: 194. CA: Alameda Co.; CA. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. REDESCRIPTION: Marsh and Schuster, 1954. HOST: From redwood leaf litter. vanus Schuster and Marsh, 1956: 127, figs. 10, 48. OR: Linn Co., Berlin; OR. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From Douglas fir leaf litter. venustrus Schuster and Marsh, 1956: 138, figs. 17, 33, 53. CA: Sonoma Co., Mark West Springs; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From oak leaf litters. Subgenus VESTITRICHUS Park Vestitrichus Park, 1953a: 259 (as subgenus). Type-species: Pselaptrichus curiosus Park (orig. des.). curiosus Park, 1953a: 258, pi. 1, C; 2, B. ID: Shoshone Co., near Avery; ID. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From fir and cedar leaf litters. Subtribe MACHAERITINA Jeannel, 1950 Genus SPELEOCHUS Park Speleochus Park, 1951: 46 (as subgenus). Type-species: Machaerites stygicus Park (orig. des.). Machaerites (of authors, not North American) (not Miller, 1855: 509). TAXONOMY: Park, 1960. REDESCRIPTION: Park, 1960. KEYS: Park, 1960. croceus Park, 1960: 78, fig. 7. AL: Madison Co., Lott Cave, near Brownsboro; AL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Cavernicole.

Stygicus Park, 1951: 47, figs. 11, 12. AL: Madison Co., Monte Sano; AL. TYPE DEPOSITORY: UANH. SEX OF TYPE: M. HOST: Cavernicole. synstygicus Park, 1956: 64, fig. 6. AL: Madison Co., King Mt., Barclay Cave; AL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Cavernicole. Genus SUBTERROCHUS Park Subterrochus Park, 1960: 91 (as subgenus). Type-species: Machaerites ferus Park (orig. des.). Machaerites (of authors, not North American) (not Miller, 1855: 509). TAXONOMY: Park, 1960. KEYS: Park, 1960.

eurous Park, 1960: 79, fig. 8 {Speleochus). AL: Jackson Co., Jess Elliot Cave; AL. TYPE DEPOSITORY: UANH. SEX OF TYPE: F. HOST: Cavernicole. 74 COLEÓPTERA CATALOG

férus Park, 1951: 49, figs. 13, 14 {Speleochus). AL: Madison Co., Aladdin Cave, Sharp's Cove; AL. TYPE DEPOSITORY: UANH. SEX OF TYPE: M. HOST: Cavernicole.

steevesi Park, 1960: 80, figs. 4, 6, 9 {Speleochus). AL: Marshall Co., Guffey Cave; AL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Cavernicole.

Genus TYCHOBYTHINUS Ganglbauer

Tychobythinus Ganglbauer, 1896: 170 (as subgenus oí Bythinus). Type-species: Bythinus ottonis Ganglbauer (design, by Besuchet, 1962: 348). Bythinus (of authors, not North American) (not Leach, 1817: 82). Tychus (of authors, not North American) (not Leach, 1817: 84). Bythinopsis Raffray, 1908: 282 (as genus) (synonymized by Besuchet, 1962). Type- species: Bythinus myrmido Reitter (design, by Bowman, 1934: 144). TAXONOMY: Besuchet, 1982a (Nearctic species).

bythinioides Brendel, 1865b: 259 (Tychus). NY: New York City; MN WI MI/ IL IN OH/ NY PA NJ/ ME VT NH MA RI CT/ NS/ ON PQ. TYPE DEPOSITORY: ANSP. SEX OF TYPE: F. bythinoides (error) Brendel, 1890: 253 {Tychus). tychoides Brendel. 1890: 253, pi. VII, fig. 35 {Machaerodes). This was an unexplained change of name for the valid Bythinopsis bythinioides; invalid replacement name. ECOLOGY: Reichle, 1966, 1967, and 1969. HOST: From sphagnum moss, swamp debris, and material in tree holes; from mixture of alder and birch litter at marsh edge. carolinae Casey, 1897: 614 {Bythinus). NC: Buncombe Co., Asheville; NC/ VA WV. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: From Rhododendron leaf litter. hubrichti Park, 1960: 76, fig. 1 {Bythinopsis). KY: Hart Co., 2 mi. E. of Millerstown, Copelin Cave; KY. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: Cavernicole. jonesi Park, 1951: 43, fig. 10 {Bythinopsis). AL: Colbert Co., Wolf Den Cave, near Maud; AL. TYPE DEPOSITORY: UANH. SEX OF TYPE: F. HOST: Cavernicole. strinatii Besuchet, 1982a: 50, figs. 1-3. TN: Warren Co., Cumberland Caverns; TN. TYPE DEPOSITORY: MHNG. SEX OF TYPE: M. HOST: Cavernicole.

Tribe VALDINI Park, 1953a

Genus VALDA Casey

Valda Casey, 1894: 493 (as genus). Type-species: Valda frontalis Casey (monot.). REDESCRIPTION: Park, 1953a. PSELAPHIDAE 75 frontalis Casey, 1894: 493, pi. I, fig. 17. CA: Siskiyou Co.; CA. TYPE DEPOSITORY: USNM. HOST: Under oak, incense cedar, and pine bark. Tribe TYCHINI Raffray, 1904b

Genus CYLINDRARCTUS L. W. Schaufuss

Cylindrarctus L. W. Schaufuss, 1887: 91 (as genus). Type-species: Cylindrarctus americanus L. W. Schaufiiss (monot.). Tychus (of authors, not North American) (not Leach, 1817: 84). TAXONOMY: Chandler, 1988a (revision). REDESCRIPTION: Chandler, 1988a, 1988b. KEYS: Chandler, 1988a. americanos L. W. Schaufuss, 1887: 92. IL; IL lA MO KS/ AR/ MI. TYPE DEPOSITORY: MNHP. SEX OF TYPE: M. REDESCRIPTION: Park, 1942; pi. I; Chandler, 1988a. HOST: From hardwood litter. bicornis Chandler, 1988a: 135, fig. 5. TX; TX. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Chandler, 1992b. HOST: From washed-in litter in cave.

comes Casey, 1894: 489, pi. I, fig. 20b. FL; FL MS. TYPE DEPOSITORY: USNM, missing. SEX OF TYPE: M. REDESCRIPTION: Chandler, 1988a. HOST: From grass and leaf litter along marshes; at UV light.

crinifer Casey, 1894: 489, pi. I, fig. 20a. IN, lA; lA NE IL IN MO OH KS KY/ TX OK/ NC FL MS TN AR LA/ WV. TYPE DEPOSITORY: USNM. REDESCRIPTION: Chandler, 1988a. HOST: From swamp litter; debris in oak woods; piles of grass; lake beach debris; at UV light.

fluvialis Chandler, 1988b: 138, figs. 12-14. FL: Suwanee Co., 7 mi. SE. of Branford, Santa Fe River; FL/ TX. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From flood drift; at UV light.

longipalpis LeConte, 1849: 82 {Tychus), "Carolina"; FL NC SC GA AL MS LA/ VA. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. REDESCRIPTION: Park and Wagner, 1962; Chandler, 1988a. HOST: From wet leaf litter in the forest; in sawdust; at UV light.

ludovicianus Brendel, 1893: 277 {Tychus). LA; LA MS NC GA FL AL. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Moxey, 1962. REDESCRIPTION: Chandler, 1988a. HOST: From hardwood forest litter; from rotten logs; at UV light. 76 COLEÓPTERA CATALOG

obrieni Chandler, 1988a: 134, figs. 3-4. FL: Franklin Co., 3 mi. NW. of Alligator Point; FL AL. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From sand pine, oak, and palmetto litter. onaga Chandler, 1988b: 135, fig. 6. KS: Pottawotamie Co., Onaga; KS. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M.

Orientalis Chandler, 1988a: 138, fig. 11. NJ: Ocean Co., Lakehurst; NY NJ DC VA. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M.

Genus LUCIFOTYCHUS Park and Wagner

Lucifotychus Park and Wagner, 1962: 22 (as genus). Type-species: Lucifotychus impellus Park and Wagner (orig. design.). Tychus (of authors, not North American) (not Leach, 1817: 84). Hylotychus Grigarick and Schuster, 1962b: 170 (as genus) (synonymized by Chandler, 1988b). Type-species: Hylotychus dentatus Grigarick and Schuster (orig. design.). REDESCRIPTION: Chandler, 1988b. KEYS: Chandler, 1991 (eastern species); Park and Wagner, 1962 (Pacific Northwest species). agomphius Grigarick and Schuster, 1962b: 169, fig. 1. OR; Coos Co., near Bridge Camp; OR. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. bipuncticeps Casey, 1887: 470 {Tychus), CA: Lake Tahoe. TYPE DEPOSITORY: USNM. REDESCRIPTION: Grigarick and Schuster, 1962b. cognatus LeConte, 1874: 50 {Tychus), BC: Vancouver, Stanley Park. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. REDESCRIPTION: Grigarick and Schuster, 1962b. HOST: From leaf litters of Douglas fir, alder, Labrador tea, spruce, and hemlock; from moss. confusus Chandler, 1983: 226, fig. \Q {Hylotychus). CA: Butte Co., 3 mi. NE. of Forest Ranch; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From forest floor litter. cornus Grigarick and Schuster, 1962b: 176, fig. 10 {Hylotychus). CA: Mendocino Co., Mendocino. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From leaf litters of alder, redwood, and fern. dentatus Grigarick and Schuster, 1962b: 170, fig. 2 {Hylotychus), CA: Del Norte Co., 19 mi. NE. of Gasquet. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From leaf litters of alder, maple, fern, and grass near streams. hirsutus Chandler, 1991: 50, figs. 5-7. NH: Coos Co., Norton Pool, 3 mi. NE. of East Inlet Dam; NH/ NS NF/ PQ/ MB SD. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From conifer leaf and log litters; in mosses. PSELAPHIDAE 77 impellus Park and Wagner, 1962: 23, pi. V (7); pi. X (3). OR; Coos Co., Charleston; WA OR. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From leaf litters of hemlock, alder, and Douglas fir; from moss. inornatus Grigarick and Schuster, 1964: 201, fig. 1. OR: Curry Co., Myrtle Grove, Loeb State Park; OR. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From forest floor litter. intellectus Grigarick and Schuster, 1962b: 171, fig. 3 {Hylotychus). OR: Lane Co., Triangle Lake. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. newelli Park and Wagner, 1962: 23, pi. V, fig. 9. OR: Coos Co., Coos Head. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. plumas Chandler, 1983: 226, fig. 9 {Hylotychus). CA: Butte Co., 2 mi. N. of Merrimac; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From maple and wet litter by streams. puberulus LeConte, 1851: 214 (Tychus). CA: Santa Clara Co., San Jose; CA. TYPE DEPOSITORY: MCZC. quirsfeldi Chandler, 1991: 52, fig. 8. NC: Yancey Co., Mount Mitchell; NC. TYPE DEPOSITORY: CNCI. SEX OF TYPE: M. remipennis Grigarick and Schuster, 1962b: 172, fig. 5 (Hylotychus). CA: Siskiyou Co., 8 mi. S. of Dunsmuir. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From Douglas-fir and white-fir leaf litters; from litter along springs. schuhi Chandler, 1986b: 338, fig. 6 (Hylotychus). OR: Klamath Co., above Geary Ranch; OR. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From leaf litters of pine, willow, and aspen.

simplicis Grigarick and Schuster, 1962b: 176, fig. 8 (Hylotychus). OR: Coos Co., Coos Bay, Shore Acres State Park. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From oak litter.

sonomae Casey, 1887: 469, pi. XVI, fig. 6 (Tychus) (nomen dubium, Grigarick and Schuster, 1962b: 177). CA: Mendocino Co. TYPE DEPOSITORY: USNM. SEX OF TYPE: F.

stellatus Grigarick and Schuster, 1962b: 172, fig. 4 (Hylotychus). CA: Humboldt Co., Freshwater. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From leaf litters of oak, Douglas fir, willow, redwood, and alder and skunk cabbage.

testaceus Casey, 1884a: 31 (Tychus). MI: Ann Arbor; MN WI MI ON PQ/ lA IL IN OH / PA NY NJ WV VA/ RI CT ME NH MA/ NB. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. 78 COLEÓPTERA CATALOG

minor (of authors, not LeConte, 1849: 82). REDESCRIPTION: Chandler, 199L ECOLOGY: Reichle, 1966 and 1969; Chandler, 1987 (all as minor). HOST: From deciduous leaf litters, often near bogs and swamps; in mosses; at UV light.

Genus NEARCTITYCHUS Chandler

Nearctitychus Chandler, 1988b: 155 (as genus). Type species: Tychus sternalis Rsiifrsiy (orig. des.). Tychus (of authors, not North American) (not Leach, 1817: 84).

Sternalis Raffray, 1904b: 268 {Tychus). NC; NJ DE MD VA/ NC SC GA AL MS/ KY. TYPE DEPOSITORY: MNHP. SEX OF TYPE: M. eisenachi Park, 1956: 95, figs. 30, 31 (Tychus) (synonymized by Chandler, 1988b). NC: Polk Co., Tryon. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Chandler, 1988b. HOST: From pine and sweetgum leaf litters; in tree-hole debris.

Genus CUSTOTYCHUS Park and Wagner

Custotychus Park and Wagner, 1962: 23 (as subgenus). Type-species: Tychus daggyi Park (orig. des.). Tychus (of authors, not North American) (not Leach, 1817: 84). REDESCRIPTION: Chandler, 1988b. daggyi Park, 1949: 327, pi. V {Tychus). NC: Cabarrus Co., Egg Rock Woods; GA NC AL TN/ MD VA WV. TYPE DEPOSITORY: FMNH. SEX OF TYPE: F. REDESCRIPTION: Park, 1956. HOST: From mixed oak and beech leaf litter; at UV light.

minor LeConte, 1849: 82 {Tychus). GA: White Co., Nacoochee (Nakutshi) Valley; WV/ NC SC GA AL TN KY LA. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. HOST: From deciduous leaf litters; at UV light. pocahontas Casey, 1897: 615 {Tychus). VA: Norfolk; MD VA WV/ NC SC GA FL. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. HOST: From pine leaf litter and mixed magnolia^eech leaf litter; from rotten logs; in moss on logs. spiculifer Casey, 1894: 491 {Tychus). PA. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. verticalis Casey, 1894: 492 {Tychus). DC; IL/ NY NJ DC VA/ NC GA. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. HOST: From silver maple leaf litter; at UV light.

Genus HESPEROTYCHUS Schuster and Marsh

Hesperotychus Schuster and Marsh, 1958a: 126 (as genus). Type-species: Hesperotychus adustus Schuster and Marsh (orig. des.). PSELAPHIDAE 79

Tychus (of authors, not North American) (not Leach, 1817: 84). TAXONOMY: Schuster and Marsh, 1958a (revision). REDESCRIPTION: Chandler, 1988b. aculeatus Schuster and Marsh, 1958a: 132, fig. 11. CA: Yolo Co., Davis; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. adustus Schuster and Marsh, 1958a: 126, figs. 1-3, 15, 19, 20. CA: San Diego Co., Palm Canyon, Borrego State Park; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From Washingtonia fllifera debris. aspersus Schuster and Marsh, 1958a: 129, figs. 6, 18, 22, 23. CA: Santa Clara Co., 2 mi. E. of Morgan Hill; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. claudus Schuster and Marsh, 1958a: 128, fig. 4. CA: Tulare Co., Ash Mountain Road, Sequoia National Park; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. fenyesi Schuster and Marsh, 1958a: 130, figs. 9, 16, 24, 25. CA: Los Angeles Co., Pasadena; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. hexagonus Casey, 1897: 616 (Tychus). CA: Ventura Co., Ojai; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Schuster and Marsh, 1958a.

macclayi Schuster and Marsh, 1958a: 131, figs. 10, 17, 26, 27. CA: Sonoma Co., Sebastopol; OR/CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From leaf litters of boxelder, ash, oak, willow, juniper, and Chtysothamnus; at UV light.

micropthalmus Brendel, 1893: 277 {Tychus). CO: Fremont Co., Canon City; CO. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Chandler, 1994.

moratus Schuster and Marsh, 1958a: 129, fig. 7. CA: Mendocino Co., Mendocino; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From redwood leaf litter.

nanus Schuster and Marsh, 1958a: 128, figs. 5, 21. CA: Yolo Co., Davis; CA/ OR. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From leaf litters of oak, manzanita, sugar pine, and grape.

ramus Park and Wagner, 1962: 21, pi. V, fig. 5. OR: Lake Co., Summer Lake; OR. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M.

stangei Grigarick and Schuster, 1962a: 99, figs. 1-3. CA: Yolo Co., Davis; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: At UV light. 80 COLEÓPTERA CATALOG tantillus Schuster and Marsh, 1958a: 136, figs. 12, 14, 28, 29. CA: Los Angeles Co., Pasadena; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. teneUus LeConte, 1851: 214 (Tychus). CA: Santa Clara Co., San Jose; CA. TYPE DEPOSITORY: MCZC.

Genus OUACHITYCHUS Chandler

Ouachitychus Chandler, 1988b: 160, figs. 20-23 (as genus). Type species: Ouachitychus parvoculus Chandler (orig. des.). parvoculus Chandler, 1988b: 160, figs. 20-23. AR: Logan Co., Mount Magazine, 0.5 mi. S. of Greenfield. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From mixed hardwood and pine litters. Tribe SPELEOBAMINI Park, 1951

Genus SPELEOBAMA Park

Speleobama Park, 1951: 52 (as genus). Type-species: Speleobama vana Park (orig. des.). vana Park, 1951: 53, fig. 15. AL: Jefferson Co., near Clay; AL. TYPE DEPOSITORY: UANH. SEX OF TYPE: M.

Genus PRESPELEA Park

Prespelea Park, 1953a: 251 (as genus). Type-species: Prespelea quirsfeldi Park (orig. des.). KEYS: Park, 1956.

Subgenus PRESPELEA Park quirsfeldi Park, 1953a: 251, pi. J; 2, E; 3, B. NC: Haywood Co., 30 mi. W. of Asheville; TN NC GA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: From Rhododendron leaf litter.

Subgenus FUSJUGAMA Park

Fusjugama Park, 1956: 55 (as subgenus). Type-species: Prespelea copelandi (orig. des.). copelandi Park, 1956: 55. TN: Blount Co., Cades Cove; TN NC. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. HOST: From hardwood leaf litter by logs. PSELAPHIDAE gj

Division MACROSCELIA Raffray, 1890a

Subfamily PSELAPHINAE Latreille, 1802 Tribe TYRINI Reitter, 1882

Subtribe TYRINA Reitter, 1882

Genus TYRUS Aube

Tyrus Aube, 1833: 15 (as genus). Type-species: Pselaphus mucronatus Panzer (monot). Pytna Casey, 1887: 457 (as genus) (synonymized by Brendel, 1887). Type-species: Pytna corticinus Casey (monot.). KEYS: Casey, 1897. carinifer Casey, 1897: 628. Southwestern UT; UT. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. consimilis Casey, 1897: 629. KY, IN; IN KY IL OH KS MO/ NJ MD DC VA WV/ NC SC GA FL AL MS TN AR LA/ TX OK TYPE DEPOSITORY: USNM. HOST: Primarily from under pine bark; also from boxelder tree holes, in rotten pine log; at UV light. corticinus Casey, 1887: 458 {Pytna), CA: Lake Tahoe; BC WA OR ID/ CA/ AZ NM/ WY. TYPE DEPOSITORY: USNM. REDESCRIPTION: Park and Wagner, 1962. HOST: Primarily from under bark of pines and Douglas fir; under bark of Betula occidentalis\ with ant, Formica sanguínea (Schv^aiz, 1895). humeralis Aube, 1844: 93 (Hamotus). "Caroline du Nord"; MB/ WI MI ON PQ/ MO IN OH IL/ NY PA NJ/ NB/ NT/ ME NH MA/ NC GA. TYPE DEPOSITORY: ZSBS. SEX OF TYPE: F. compar LeConte, 1849: 80 (synonymized by LeConte, 1863a). GA, Carolina, and Missouri Territory. TYPE DEPOSITORY: MCZC. HOST: Primarily from under bark of white pine; also from under bark of spruce and balsam fir; with ant, Aphaenogaster tennesseensis (Schwarz, 1890); at UV light. semiruber Casey, 1897: 629. "Lake Superior." TYPE DEPOSITORY: USNM. SEX OF TYPE: F.

Genus CEDIUS LeConte

Cedius LeConte, 1849: 74 (as genus). Type-species: Cedius spinosus LeConte (design, by Lucas, 1920: 169). TAXONOMY: Park, 1949. REDESCRIPTION: Park, 1949. KEYS: Park, 1949.

Subgenus CEDIUS LeConte

spinosus LeConte, 1849; 75. SC: Richland Co., Columbia; MO IL IN lA OH KY/ DE NY PA NJ MD WV/ NC SC GA FL AL TN/ TX. TYPE DEPOSITORY: MCZC. 82 COLEÓPTERA CATALOG

obsoletus Park, 1949: 340 (described as variety). NY; Massapequa, NJ: Jamesburg, IL; Willow Springs. Park's use of this name for an infrasubspecific group renders this name a junior synonym [ICZN, 1985, article 45g]. TYPE DEPOSITORY: FMNH. REDESCRIPTION: Park, 1949. ECOLOGY: Engelmann, 1956: Park, 1964. HOST: With ants, Aphaenogaster tennesseensis, Lasius umbratus (Park, 1935b); from under oak and pine bark; at UV light.

Subgenus SINISTROCEDIUS Newton and Chandler

Sinistrocedius Newton and Chandler, 1989: 59 (as subgenus). Type-species: Cedius ziegleri LeConte (orig. des.). Sinistrocedius Park, 1949: 336 (as subgenus) (nomen nudum). cruralis Park, 1949; 338, pi. VI; VII, 1, 2, 9. IN: Laporte Co., Davis Woods, near Smith; MI/ IL IN/ TN MS AL LA. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. ECOLOGY: Park, 1964. HOST: With ant, Aphaenogaster tennesseensis (Park, 1964); from tree holes; at UV light. robustos Casey, 1897: 626. DC; DC. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Park, 1949. ziegleri LeConte, 1849: 74. PA; ON/ lA IL MO IN OH/ PA NJ DC MD VA/ MA CT/ NC GA TN AL. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. REDESCRIPTION: Park, 1949. HOST: Under bark; with ants Formica rufa, F. integra, F. exsectoides (Schwarz, 1890), Camponotus pennsylvanicus (Schmitt, 1901); in oak tree holes.

Genus MIPSELTYRUS Park

Mipseltyrus Park, 1953a: 262 (as genus). Type-species: Mipseltyrus nicolayi Park (orig. des.). levini Chandler, 1978: 324, fig. 1. CA: Solano Co., Lake Solano; CA. TYPE DEPOSITORY: UCDC. SEX OF TYPE: M. HOST: From leaf litters of digger pine, maple, grape, madrone, and oak (Chandler, 1983). mirus Schuster, 1956: 84, fig. 3. CA: Eldorado Co., Riverton; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: From leaf litters of Douglas fir, madrone, maple, and live oak. nicolayi Park, 1953a: 263, pi. 4, D; 5, A-D. TN: Sevier Co., near Gatlinburg, The Chimneys; TN NC. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: From leaf litters of Rhododendron, pine, and oak. parki Schuster, 1956: 83, figs. 1, 2, 4. CA: Mariposa Co., Briceburg; CA. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. PSELAPHIDAE 83

Genus ANITRA Casey

Anitra Casey, 1894: 499 (as genus). Type-species: Anitra glaberula Casey (monot.). TAXONOMY: Chandler, 1974b. REDESCRIPTION: Chandler, 1974b. glaberula Casey, 1894: 500, pi. I, fig. 15. AZ; AZ. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: With ant, Paratrechina melanderi (Chandler, 1974b). Subtribe HAMOTINA Park, 1951

Genus HAMOTUS Aube

Hamotus Aube, 1844: 91 (nominate subgenus Neotropical). Type-species: Hamotus lateritius Aube (design, by Lucas, 1920: 314).

Subgenus HAMOTOIDES Schaufuss

Hamotoides L. W. Schaufuss, 1888: 298 (as subgenus). Type-species: Hamotus tritomus Reitter (design, by Jeannel, 1962: 473). TAXONOMY: Chandler, 1974a. KEYS: Chandler, 1974a (western species). electrae Park, 1942: 327. Mexico: Veracruz, San Juan; TX; Mex. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Chandler, 1992b. HOST: In caves; at UV light. elongatus Brendel, 1890: 239, pi. VI, fig. 21 {Tyrus). AZ: Coconino Co., Williams; AZ NM. TYPE DEPOSITORY: ANSP. SEX OF TWE: M. TAXONOMY: Chandler, 1994. REDESCRIPTION: Chandler, 1974a. HOST: Under pine bark. nodicollis Raffray, 1883: 240. Mex.; AZ; Mex. TYPE DEPOSITORY: MNHP. SEX OF TYPE: M. REDESCRIPTION: Chandler, 1974a. HOST: In rotting saguaro and cardon cacti; at UV light; under bark. opimus Fletcher, 1932: 34. FL: Palm Beach Co., Palm Beach; FL MS/ OK. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. HOST: Sweeping herbage; from hardwood leaf litter; at UV light. populus Chandler, 1974a: 379, figs. 1, 5. AZ: Cochise Co., Cochise Stronghold, Dragoon Mts.; AZ; Mex. TYPE DEPOSITORY: CASC. SEX OF TYPE: M. HOST: Under cottonwood bark; at UV light.

Genus CIRCOCERUS Motschulsky

Circocerus Motschulsky, 1855: 16 (as genus). Type-species: Circocerusperplexus Motschulsky (monot.). Cercocerus {QWOX) LeConte, 1861: 57. 84 COLEÓPTERA CATALOG

Upoluna L. W. Schaufiíss, 1886: 283 (as genus) (synonymized by Raffray, 1908). Type- species: Upoluna flavitarsis L. W. Schaufuss (= Circocerus batrisioides Motschulsky). batrisioides Motschulsky, 1856: 4. LA: Orleans Parish, New Orleans; KS MO IL IN OH KY/ PA NY WV/ AR LA FL TN/ TX OK. TYPE DEPOSITORY: MCZC, ZMUM. batrisoides (error) LeConte, 1863: 27. flavitarsis L. W. Schaufuss, 1886: 283 (Upoluna) (synonymized by Raffray, 1908). Upolu Islands (Samoa). Raffray initially (1896: 135) believed that these two taxa represented separate species but later decided (1908: 397) that the type-locality of flavitarsis was incorrect and the species were the same. TYPE DEPOSITORY: MNHP. SEX OF TYPE: M. HOST: With ant, Aphaenogaster tennesseensis (Park, 1935b); in boxelder and oak tree iioles; from mixed hardwood litter; at UV light. Tribe CEOPHYLLINI Park, 1951 Genus CEOPHYLLUS LeConte Ceophyllus LeConte, 1849: 73 (as genus). Type-species: Ceophyllus monilis LeConte (monot.). monilis LeConte, 1849: 73. MI: St. Clair Co., St. Clair River; WI MI ON/ MB/ MO IL IN OH KY/ PA NJ MD DE WV VA/ MA/ NC TN AL. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. HOST: With ants, Lasius umbratus, L. niger americanus, L flavus marcticus (Park, 1932a and 1935b; Wickham, 1894); with Lasius claviger? (Schwarz, 1890; Hamilton, 1886). Tribe TMESIPHORINI Jeannel, 1949 Genus TMESIPHORUS LeConte Tmesiphorus LeConte, 1849: 75 (as genus). Type-species: Pselaphus carinatus Say (design, by Lucas, 1920: 643). carinatus Say, 1824: 97 (Pselaphus). "United States"; KS MO IL IN OH KY lA/ PA NJ MD VA/ SC MS AL GA FL AR LA/ TX OK. TYPE DEPOSITORY: Lost? HOST: Under pine bark; from sand pine and oak litter; with ants, Aphaenogaster tennesseensis (Park, 1935b); with termite, Reticulitermes sp.; at UV light. costalis LeConte, 1849: 77. PA; KS MO IL IN OH KY lA/ PA NJ VA MD/ AR MS AL TN GA NC FL SC/ TX OK/ ON. TYPE DEPOSITORY: MCZC. rostratus L. W. Schaufuss, 1888: 296 (as var.). IN. Initially proposed as a variety by Schaufuss and maintained by all authors as variety. Last citation is by Leng (1920: 132). The continued infraspecific use of this name renders it a junior synonym [ICZN, 1985, article 45g]. TYPE DEPOSITORY: MFNB. REDESCRIPTION: Chandler, 1992b. ECOLOGY: Englemann, 1956. HOST: Under pine bark; in caves (Chandler, 1992b); from mixed leaf litters of beech/magnolia and sand pine/oak; from rotten wood; with ants, Aphaenogaster fiilva (Park, 1933) and Formica rufa (Schwarz, 1890); at UV light. Tribe CTENISTINI Blanchard, 1845 Genus CTENISIS Raffray Ctenisis Raffray, 1890a: 143 (as genus). Type-species: Desimia dispar Sharp (design, by Raffray, 1897: 271). PSELAPHIDAE 85 raffrayi Casey, 1894: 503, pi. I, fig. 13. AZ: Pima Co., Tucson; CA/ AZ TX; Mex., C. Amer. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: With ant, Veromessor juliana (Chandler, 1976); at UV light.

Genus PILOPIUS Casey

Pilopius Casey, 1897: 617 (as genus). Type-species: Pilopius laeustris Casey (design, by Bowman, 1934: 144). Ctenistes (of authors, not North American) (not Reichenbach, 1816: 75). Sognorus (of authors, not North American) (not Reitter, 1882: 202). KEYS: Casey, 1897. abruptus Casey, 1894: 502, pi. I, fig. 14a {Sognorus). AZ; AZ. TYPE DEPOSITORY: USNM. REDESCRIPTION: Park and Wagner, 1962 (aedeagus).

Cinderella Casey, 1897: 624. TX: Cameron Co., Brownsville; TX OK. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: At UV light. consobrinus LeConte, 1849: 79 {Ctenistes). GA: Clarke Co., Athens; NJ VA/ GA. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. floridanus Casey, 1897: 623. FL; LA FL. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: At UV light. georgianus Casey, 1897: 621. GA; GA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. granicoUis Casey, 1897: 622. NY: Long Island, Willets Point; NY. TYPE DEPOSITORY: USNM. impressipennis Casey, 1897: 624. TX; TX. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. HOST: Under bark; in logs with ant, Cyphomyrmex hmosus; at UV light. iowensis Casey, 1897: 622. lA: Lee Co., Keokuk; KS lA MO/ TN. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: From oak leaf litter; with ants; at UV light. laeustris Casey, 1897: 619. MI, IN, northern IL, lA, Iowa City; MI ON/ lA IL IN OH. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. ECOLOGY: Park, Auerbach, and Wilson, 1949 and 1953; Mickey and Park, 1956; Park, 1964. HOST: At UV light. ocularis Casey, 1894: 502 {Sognorus). AZ; AZ/ CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: With ants, Novomessor albisetosus (Chandler, 1976), Ischnomyrmex cockerelli (Mann, 1924); at UV light.

piceus LeConte, 1849: 78 {Ctenistes). VT; OH/ PA NJ NY/ VT CT ME NH/ MI ON PQ. TYPE DEPOSITORY: MCZC. HOST: From leaf litter along the edges of streams and marshes; from moss; from rotten wood; at UV light. 86 COLEÓPTERA CATALOG pulvereus LeConte, 1851: 214 {Ctenistes). CA: Santa Clara Co., San Jose; CA/ AZ. TYPE DEPOSITORY: MCZC. REDESCRIPTION: Park and Wagner, 1962 (aedeagus). HOST: With ants, Formica parva, F. Schaufussi (Wickham, 1892); at UV light. saginatus Casey, 1897: 620. ON: Ottawa, RI: Boston Neck; ON/ MA RI. TYPE DEPOSITORY: USNM. zimmermani LeConte, 1849: 79 {Ctenistes). GA, LA; KS MO IN lA/ AR LA GA FL; Mex. TwE DEPOSITORY: MCZC. HOST: At UV light. Genus BIOTUS Casey Biotus Casey, 1887: 456 (as genus). Type-species: Biotus formicarius Casey (monot.). formicarius Casey, 1887: 456, pi. XVI, fig. 2. CA: Los Angeles Co., Los Angeles; CA/ AZ. TYPE DEPOSITORY: USNM. HOST: With ants; at UV light. Genus ATINUS Horn Atinus Horn, 1868: 127 (as genus). Type-species: Ctenistes monilicornis Brendel (monot.). Ctenistes (of authors, not North American) (not Reichenbach, 1816: 75). KEYS: Casey, 1894.

brevicornis Casey, 1894: 498. TX; AZ TX. TYPE DEPOSITORY: USNM. HOST: With ants; at UV light.

monilicornis Brendel, 1866: 190 {Ctenistes). TN: Rhea Co., near Washington City; MO IL/ OK/ VA DE WV/ NC FL AL MS TN GA. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Chandler, 1994. REDESCRIPTION: Brendel, 1893. HOST: From leaf litter and rotten stumps; with ants, Prenolepis párvula (Schwarz, 1890) and Pheidole vinlandica (Wickham, 1900); at UV light. Tribe ARHYTODINI Raffray, 1890a Holozodini Raffray, 1900. Genus CACCOPLECTUS Sharp

Caccoplectus Sharp, 1887: 22 (as genus). Type-species: Caccoplectus celatus Sharp (monot.). TAXONOMY: Chandler, 1976; Chandler and Wolda, 1986. REDESCRIPTION: Chandler, 1976; Chandler and Wolda, 1986. KEYS: Chandler, 1976; Chandler and Wolda, 1986. ECOLOGY: Chandler and Wolda, 1986.

nuttingi Chandler, 1976: 305, figs. 4-10. AZ: Pima Co., Santa Rita Range Reserve, IBP site; AZ, Mex. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Chandler and Wolda, 1986. HOST: At UV light.

pectinatus Chandler, 1976: 313, figs. 34-40. AZ: Pima Co., Tucson; AZ. TYPE DEPOSITORY: CNCI. SEX OF TYPE: M. REDESCRIPTION: Chandler and Wolda, 1986. HOST: At UV light. PSELAPHIDAE 87 sentis Chandler, 1976: 309, figs. 11-17. AZ: Cochise Co., Cochise Stronghold, Dragoon Mts.; AZ TX. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Chandler and Wolda, 1986. HOST: At UV light. spinipes Schaeffer, 1906: 263. TX; TX; C. Amer. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Chandler, 1976; Chandler and Wolda, 1986. HOST: At UV light. Tribe PSELAPHINI Latreille, 1802

Genus PSELAPHUS Herbst

Pselaphus Herbst, 1792: 106 (as genus) (other subgenus not from N. Amer. (Jeannel, 1951)). Type-species: Pselaphus heisei Herbst (design, by Shuckard, 1839: 147). KEYS: Casey, 1894.

Subgenus PSELAPHUS Herbst bellax Casey, 1894: 496, pi. I, fig. 18. MA: Bristol Co., Taunton; MI MN WI ON PQ/ IL lA IN/ NB NS/ ME NH/ MB SK AB ND SD/ WA ID BC/ NY NJ PA. TYPE DEPOSITORY: USNM. REDESCRIPTION: Park and Wagner, 1962. ECOLOGY: Reichle, 1966 and 1969. HOST: Most common in leaf litters on marsh edges and in mosses; in beaver houses. erichsoni LeConte, 1840: 81. NY; NY PA NJ/ CT MA ME/ WI ON/ IL IN KS/ FL. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. HOST: Under stones in grassy areas; under mullein leaves; from Andropogon leaf litter. fustifer Casey, 1894: 495, pi. I, fig. 18a. NY; WI MI ON PQ/ MB AB/ NE IL/ NY NJ WV/ ME RIMA/NSNB. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. ECOLOGY: Reichle, 1966; Park, Auerbach, and Wilson, 1949 and 1953; Mickey and Park, 1956. HOST: Primarily from mosses. longiclavus LeConte, 1849: 81. "provinciis australibus"; lA/ LA FL GA MS/ TX/ MA. TYPE DEPOSITORY: MCZC. HOST: From Spartina cynosuroides and Juncus roemerianus litter; from swamp litter; at UV light. ulkei Bowman, 1934: 142. SD; SD. TYPE DEPOSITORY: ICCM. REDESCRIPTION: Park and Wagner, 1962. ECOLOGY: Reichle, 1966, 1967, and 1969. HOST: From sphagnum moss. Genus NEOPSELAPHUS Jeannel

Neopselaphus Jeannel, 1951: 8 (as genus). Type-species: Pselaphus bizonatus L. W. Schaufiiss (orig. design.). TAXONOMY: Besuchet, 1982b (revision). REDESCRIPTION: Besuchet, 1982b. KEYS: Besuchet, 1982b. 88 COLEÓPTERA CATALOG mexicanus Park, 1945: 411, pi. 1, figs. 5-6 {Pselaphellus), Mexico: Veracruz, Vista Hermosa, Rio Tonto; TX; Mex., C. Amer. TYPE DEPOSITORY: FMNH. SEX OF TYPE: M. REDESCRIPTION: Besuchet, 1982b. HOST: At UV light. Subfamily CLAVIGERINAE Leach, 1815

Besuchet, 1991. Tribe CLAVIGERINI Leach, 1815

Clavigerodini L. W. Schaufuss, 1882; Fustigerini Jeannel, 1949; Adranites Chenu and Desmarest, 1857.

Genus FUSTIGER LeConte

Fustiger LeConte, 1866b: 109 (as genus). Type-species: Articerus braziliensis Westwood (design, by Newton and Chandler, 1989). Articerus (of authors, not North American) (not Articerus Dalman, 1826: 394). fuchsii Brendel, 1866: 190. TN: CA/ MO IL IN/ AZ/ MS AL TN. TYPE DEPOSITORY: ANSP. SEX OF TYPE: F. TAXONOMY: Chandler, 1994. californicus Brendel, 1889: 197 {Articerus) (placed by Brendel, 1890 as a variety of fuchsii and thus in synonymy). CA: Los Angeles Co., Los Angeles. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. HOST: With ants, Crematogaster leviuscula (Park, 1935b) and C lineolata (Wickham, 1892). knausii Schaeffer, 1906: 264. NM: Otero Co., Cloudcroft; CO/ AZ NM. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. ECOLOGY: Leschen, 1991. HOST: With ant, Crematogaster cerasi (Leschen, 1991).

Genus ADRANES LeConte

Adranes LeConte, 1849: 83 (as genus). Type-species: Adranes coecus LeConte (monot.). KEYS: Wickham, 1901. angustus Casey, 1924: 150. MO: St. Louis Co., St. Louis; MO. TYPE DEPOSITORY: USNM. coecus LeConte, 1849: 83. GA: Montis Jonae; MO IL OH/ NY PA NJ VA/ GA NC AL MS. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. HOST: With ants, Lasius niger americanus, L niger neoniger, Aphaenogaster fiilva aquia (Park, 1935b). L alienus (Schwarz, 1890). dietzi Schaeffer, 1906: 265. CA: Tulare Co.; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. PSELAPHIDAE 89 lecontei Brendel, 1865b: 255. IL; MT MB SK AB/ MI ON/ MO IL IN OH KY lA/ PA NY/ MA/ MSNC. TYPE DEPOSITORY: ANSP. SEX OF TYPE: M. TAXONOMY: Moxey, 1962. REDESCRIPTION: Wickham, 1901. ECOLOGY: Park, 1964; Kistner, 1982. HOST: With ants, Lasius umbratus, Aphaenogaster fulva (Park, 1932a and 1935b; Wickham, 1894). pacificus Wickham, 1901: 26, fig. 2. CA: Sisson; CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. HOST: With ant, Lasius niger (Wickham, 1901).

taylori Wickham, 1901: 27, fig. 3. OR: Lincoln Co., Newport; BC WA OR ID/ SK/ CA. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. REDESCRIPTION: Park and Wagner, 1962; Akre and Hill, 1973. ECOLOGY: Akre and Hill, 1973; Hill et ai, 1976; Kistner, 1982. HOST: With ants, Lasius niger, L sitkaensis (Akre and Hill, 1973), L pallitarsis (Hill et al, 1976). 90

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Park, O. 1932a The myrmecocoles of Lasius umbratus mixtus aphidicola Walsh. Annals of the Entomological Society of America, vol. 25, pp. 77-88.

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Park O. 1949 New species of Nearctic pselaphid beetles and a revision of the genus Cedius. Bulletin of the Chicago Academy of Sciences, vol. 8, pp. 315-343, pis. 1-8, illus.

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Park, O. 1952b A revisional study of Neotropical pselaphid beetles. Part two. Tribe Euplectini sensu latiore. Chicago Academy of Sciences, Special Publication No. 9 (2), pp. 53-150. Park, O. 1953a New or little known pselaphid beetles of the United States, with observations on taxonomy and evolution of the family Pselaphidae. Bulletin of the Chicago Academy of Sciences, vol. 9, pp. 249-283, illus. Park, O. 1953b An European pselaphid beetle collected in New York. Natural History Miscellanea. The Chicago Academy of Sciences, Chicago, Illinois, no. 117, pp. 1-3. Park, O. 1953c A new genus of pselaphid beetles from the Everglades. Natural History Miscellanea. The Chicago Academy of Sciences, Chicago, Illinois, no. 121, pp. 1-4, illus. Park, O. 1953d Discrimination of genera of pselaphid beetles of the United States. Bulletin of the Chicago Academy of Sciences, vol. 9, pp. 299-331, illus. Park, O. 1956 New or little known species of pselaphid beetles from Southeastern United States. Journal of the Tennessee Academy of Science, vol. 31, pp. 54-100, illus. Park, O. 1958 New or little known species of pselaphid beetles chiefly from Southeastern United States. Journal of the Tennessee Academy of Science, vol. 33, pp. 39-74, illus. Park, O. 1960 Cavemicolous pselaphid beetles of the United States. The American Midland Naturalist, vol. 64, pp. 66-104, illus.

Park, O. 1962 A review of the genus Ramecia (Coleóptera: Pselaphidae). The Ohio Journal of Science vol. 62, pp. 150-156, illus.

Park, O. 1963 Observations on the genus Actium (Coleóptera: Pselaphidae). Transactions of the American Microscopical Society, vol. 82, pp. 178-184, illus. Park, O. 1964 Observations upon the behavior of myrmecophilous pselaphid beetles. Pedobiolosia vol. 4, pp. 129-137, illus. '

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Park, O,, S Auerbach, and M. Wilson 1949 Pselaphid beetles of an Illinois prairie: the fauna, and its relation to the Prairie Peninsula Hypothesis. Bulletin of the Chicago Academy of Sciences, vol. 8, pp. 267- 277, illus.

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Raffray, A, 1883 Psélaphides nouveaux ou peu connus, deuxième mémoire. Revue d'Entomologie, vol. 2, pp. 229-251, pis. 4-5, illus.

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Raffray, A. 1891 (1890) Voyage de M. E. Simon au Venezuela (Décembre 1887-Avril 1888). Annales de la Société Entomologique de France, vol. 10, no. 6, pp. 297-330, pi. 6.

Raffray, A. i • j T- 1893 Révision des Psélaphides de Sumatra. Annales de la Société Entomologique de France, (1892) vol. 61, pp. 463-504, pi. 10, illus.

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Raffray, A. i i i o • ' ' 1897 Nouvelles études sur les Psélaphides et les Clavigérides. Annales de la Société Entomologique de France, (1896) vol. 65, pp. 227-284, illus. 106 COLEÓPTERA CATALOG

Raffray, A. 1898a Révision des Batrisus et genres voisins de l'Amérique centrale et méridionale. Annales de la Société Entomologique de France, (1897) vol. 66, pp. 431-517, pi. 12, illus. Raffray, A. 1898b Notes sur les Psélaphides. Révision générique de la tribu des Euplectini. Descriptions d'espèces nouvelles. Revue d'Entomologie, vol. 17, pp. 198-273. Raffray, A. 1900 Psélaphides et Clavigérides de Madagascar. Annales de la Société Entomologique de France, vol. 68, pp. 516-525.

Raffray, A. 1904a Genera et catalogue des Psélaphides. Annales de la Société Entomologique de France (1903) vol. 72, pp. 484-604, illus.

Raffray, A, 1904b Genera et catalogue des Psélaphides. Annales de la Société Entomologique de France vol. 73, pp. 1-400, illus.

Raffray, A. 1905 Genera et catalogue des Psélaphides. Annales de la Société Entomologique de France (1904) vol. 73, pp. 401-476, pis. 1-3, illus.

Raffray, A. 1908 Coleóptera. Fam. Pselaphidae. Genera Insectorum. P. Wytsmann, ed., Rome, fase 64 487 pp., 9 pis., illus.

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Reichle, D. E. 1966 Some pselaphid beetles with boreal affinities and their distribution along the postglacial fringe. Systematic Zoology, vol. 15, pp. 330-344, illus.

Reichle, D. E. 1967 The temperature and humidity relations of some bog pselaphid beetles Ecology vol 48,_pp. 208-215, illus.

Reichle, D. E. 1969 Distribution and abundance of bog-inhabiting pselaphid beetles. Transactions of the Illinois Academy of Science, vol. 62, pp. 233-264, illus. Reitter, E. 1881 Bestimmungs-Tabellen der europäischen Coleóptera. V. Enthaltend die Familien: Paussidae, Clavigeridae, Pselaphidae und Scydmaenidae. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, vol 31 pp 443-592, pl. 19, illus. ' ' PSELAPHIDAE 107

Reitter, E. 1882 Versuch einer systematischen Eintheilung der Clavigeriden und Pselaphiden. Verhandlungen des naturforschenden Vereines in Brunn, (1881) vol. 20, pp. 177- 211.

Reitter, E. 1883a Beitrag zur Kenntniss der Clavigeriden, Pselaphiden und Scydmaeniden von Westindien. Deutsche Entomologische Zeitschrift, vol. 27, pp. 33-46, illus.

Reitter, E. 1883b Neue Pselaphiden und Scydmaeniden aus Central-und Südamerika. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, (1882) vol. 32, pp. 371-386.

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Sanderson, M. W. 1940 Two new species of Batrisodes from South Carolina (Coleóptera: Pselaphidae). Entomological News, vol. 51, pp. 169-172.

Saulcy, F. H. C. de 1876 Species des Paussides, Clavigérides, Psélaphides et Scydménides de l'Europe et des pays circonvoisins. Bulletin de la Société d'Histoire Naturelle de Metz, vol. 14, pp. 25-100.

Say, T. 1824 Descriptions of coleopterous insects collected in the late expedition to the Rocky Mountains, performed by order of Mr. Calhoun, Secretary of War, under the command of Major Long. Journal of the Academy of Natural Sciences of Philadelphia, vol. 4, pp. 83-99.

Schaeffer, C. F. A. 1905 Additions to the Coleóptera of the United States with notes on some known species. Brooklyn Institute Museum of Arts and Science Bulletin, vol. 1, pp. 123-140.

Schaeffer, C. F. A. 1906 Six new Pselaphidae. Transactions of the American Entomological Society, vol. 32, pp. 261-266.

Schaufuss, C. 1888 Catalogus synonymicus Pselaphidarum adhuc descriptarum. Tijdschrift voor Entomologie, vol. 31, pp. 1-104.

Schaufuss, L. W. 1872 Tabellen-Entwurf zur Bestimmung der Pselaphiden-Gattungen. Nunquam Otiosus, vol. 2, pp. 243-248. 108 COLEÓPTERA CATALOG

Schaufuss, L. W. 1877 Ueber Pselaphidengattungen. Nunquam Otiosus, vol. 2, pp. 450-460.

Schaufuss, L. W. 1882 Pselaphidarum Monographiae. Annali del Museo Civico di Storia Naturale di Genova vol. 18, pp. 349-399.

Schaufuss, L. W. 1886 Beschreibung neuer Pselaphiden aus der Sammlung des Museum Ludwig Salvator. Ein Beitrag zur Fauna Brasiliens, der Kgl. Niedeirlaendischen Besitzungen in Indien und Neuhollands. Tijdschrift voor Entomologie, vol. 29, pp. 241-296, pis. 10-11, illus. Schaufuss, L. W. 1887 Beschreibung neuer Pselaphiden aus der Sammlung des Museum Ludwig Salvator. Ein Beitrag zur Fauna Brasiliens, der Kgl. Niederländischen Besitzungen in Indien und Neuhollands (Forsetzung). Tijdschrift voor Entomologie, vol. 30, pp. 91-165, pis. 7-9, illus.

Schaufuss, L. W. 1888 Ueber Pselaphiden und Scydmaeniden des Königl. zoologischen Museums zu Berlin und verwandte Arten. Berliner Entomologische Zeitschrift, (1887) vol. 31, pp. 287- 320.

Schaufuss, L. W. 1890 System-Schema der Pselaphiden, ein Blick in die Vorzeit, in die Gegenwart und in die Zukunft. Tijdschrift voor Entomologie, vol. 33, pp. 101-162, pis. 2-6 and foldout, illus.

Schmitt, J. 1901 Pselaphidae, pp. 280-285. In H. G. Klages, Supplement to Dr. John Hamilton's list of the Coleóptera of southwestern Pennsylvania. Annals of the Carnegie Museum, vol. I, pp. 265-294.

Schuster, R. O. 1956 Two new species of Mipseltyrus from California. The Pan-Pacific Entomologist, vol. 32, pp. 83-86, illus.

Schuster, R. O. 1959 Notes on Morius occidens Casey with a description of the male (Coleóptera: Pselaphidae). The Pan-Pacific Entomologist, vol. 35, pp. 95-97, illus. Schuster, R. O. 1961 Four new species of Mayetia from western North America (Coleóptera: Pselaphidae). The Pan-Pacific Entomologist, vol. 37, pp. 23-27, illus.

Schuster, R. O. and A. A. Grigarick 1960 A revision of the genus Oropus Casey (Coleóptera: Pselaphidae). Pacific Insects, vol. 2, pp. 269-299, illus.

Schuster, R. O. and A. A. Grigarick 1962 A revision of the genus Rhexidius Casey (Coleóptera: Pselaphidae). The Pan-Pacific Entomologist, vol. 38, pp. 1-14, illus. PSELAPHIDAE 109

Schuster, R. O. and A. A. Grigarick 1968 A new genus of pselaphid beetles from Southeast United States (Coleóptera: Pselaphidae). The Pan-Pacific Entomologist, vol. 44, pp. 112-118, illus.

Schuster, R. O. and G. A. Marsh 1956 A revision of the genus Pselaptrichus Brendel (Coleóptera: Pselaphidae). University of California Publications in Entomology, vol. 11, pp. 117-158, illus.

Schuster, R. O. and G. A. Marsh 1957 A new genus of Euplectini from California. The Pan-Pacific Entomologist, vol. 33, pp. 149-152, illus.

Schuster, R. O. and G. A. Marsh 1958a A new genus of Tychini from California (Coleóptera: Pselaphidae). The Pan-Pacific Entomologist, vol. 34, pp. 125-137.

Schuster, R. O. and G. A. Marsh 1958b A study of the North American genus Megarafonus Casey (Coleóptera: Pselaphidae). The Pan-Pacific Entomologist, vol. 34, pp. 187-194, illus.

Schuster, R. O., G. A. Marsh, and O. Park 1959 Present status of the tribe Mayetini in the United States-Part I (excluding California). The Pan-Pacific Entomologist, vol. 35, pp. 117-127, illus.

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Schwarz, E. A. 1890 Myrmecophilous Coleóptera found in temperate North America. Proceedings of Entomological Society of Washington, (1889) vol. 1, pp. 237-247.

Schwarz, E. A. 1895 Additions to the lists of North American termitophilous and myrmecophilous Coleóptera. Proceedings of the Entomological Society of Washington, vol. 3, pp. 73- 78.

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Names are indexed as follows: arkansana Fall, Rybaxis 53 CAPITALS: All names for taxa above the generic level; armiger LeConte, Batriasymmodes 47 Boldface: Valid generic and subgeneric names; Arthmius LeConte 48 Roman: Valid specific and subspecific names; arthritica Brendel, Reichenbachia 57 Italics: All invalid names such as synonyms, nomina nuda, Articerus (of authors) 88 and extra-limital taxa even though valid. articularis Casey, Reichenbachia 58 arundineum Grigarick and Schuster, Actium 26 The generic name following the author's name indicates the aspersus Schuster and Marsh, Hesperotychus 79 present placement of the species. Synonyms of species-group asperulus Casey, Rhexidius 5 names are listed with the original spelling. aterrimus Casey, Batrisodes 46 Atinus Horn 86 abbreviatus Casey, Oropus 7 atlántica Brendel, Reichenbachia 57 Abdiunguis Park and Wagner 14 atratus Casey, Thesiastes 18 abdominalis Aube, Brachygluta 54 auctumnus Chandler, Pselaptrichus 70 abnorme LeConte, Decarthron 66 auerbachi Park, Batrisodes 43 abruptus Casey, Pilopius 85 auerbachi Park, Trimioplectus 23 Abryxis Raffray 66 austral is Chandler, Trimioplectus 23 Acolonia Casey 19 Babnormodes Park 38 acomanus Casey, Euplectus 19 barbata Barr, Arianops 50 acornus Carlton, Conoplectus 6 barbatum Grigarick and Schuster, Actium 26 acriculus Schuster and Grigarick, Oropus 7 barri Park and Wagner, Actium 26 Actiastes Casey 29 barri Park and Wagner, Euplectus 20 Actium Casey 25 barri Park, Batrisodes 38 Actizona Chandler 25 basalis Casey, Oropus 7 aculeatum Grigarick and Schuster, Actium 25 Basolum Casey 25 aculeatus LeConte, Batrisodes 46 Batriasymmodes Park 47 aculeatus Schuster and Marsh, Hesperotychus 79 BATRISINA 38 acumenis Schuster and Grigarick, Oropus 7 BATRISINAE 38 Adranes LeConte 88 BATRISINI 38 adustus Schuster and Marsh, Hesperotychus 79 batrisioides Motschulsky, Circocerus 84 aggestus Schuster and Grigarick, Rhexidius 5 Batrisodes Reitter 38 agomphius Grigarick and Schuster, Lucifotychus 76 batrisoides (error) LeConte, Circocerus 84 albionica Motschulsky, Reichenbachia 57 Batrisus (of authors) 38, 47 albionicus Aube, Batrisodes 41 beameri Park, Batriasymmodes 47 alexanderi Carlton and Chandler, Pseudactium 16 belfragei LeConte, Brachygluta 55 allatoona Barr, Arianops 49 belfragei LeConte, Pselaptus 66 Allobrox Fletcher 31 bellax Casey, Pselaphus 87 Allotrimium Park 33 benedictae Grigarick and Schuster, Pilactium 31 alticola Barr, Arianops 49 beyeri Schaeffer, Batrisodes 43 AMAUROPINI 49 BIBLOPLECTINA 23 amblyoponica (of authors), Arianops 50 Bibloplectodes Jeannel 23 americana Motschulsky, Trimiomelba 34 Bibloplectus Reitter 23 americana Raffray, Reichenbachia 57 BIBLOPORINA 13 americanus L. W. Schaufuss, Cylindrarctus 75 Bibloporus Thomson 13 amplyoponica Brendel, Arianops 50 bicanalis Casey, Bibloporus 13 Anchylarthron Brendel 65 bicentennialis Carlton, Conoplectus 6 angustum Casey, Actium 27 bicolor Brendel, Reichenbachia 57 angustus Casey, Adranes 88 bicolor Casey, Actium 26 Anitra Casey 83 bicornis Chandler, Cylindrarctus 75 anneae Park, Decarthron 66 bicorona Schuster, Mayetia 36 Anops Brendel 49 bifalxa Park, Rybaxis 53 antennatus Schaeffer, Batrisodes 38 bifoveatum Casey, Actium 26 apache Chandler, Tomoplectus 35 binodifer Casey, Reichenbachia 57 aphenogastri Fall, Batrisodes 42 Biotus Casey 86 Apothinus Sharp 13 bipuncticeps Casey, Lucifotychus 76 appalachianus Casey, Batrisodes 38 bistriatus LeConte, Batrisodes 41 appendiculata appendiculata Raffray, Reichenbachia 57 bistriatus LeConte, Rhinoscepsis 13 appressicornis Park, Rybaxis 53 blandum Casey, Actiastes * 30 aravaipa Chandler, Saxet 18 Bontomtes Grigarick and Schuster 10 arcaps Grigarick and Schuster, Oropodes 15 borealis Casey, Reichenbachia 58 arcifer Casey, Reichenbachia 57 borealis Chandler, Actizona 25 arcuatum LeConte, Pseudactium 16 bowmani Schuster, Mayetia 36 arguta Casey, Brachygluta 54 Brachygluta Thomson 54 ARHYTODINI 86 BRACHYGLUTINA 53 Arianops Brendel 49 BRACHYGLUTINI 53 Arispeleops Park 49 BRACHYSCELIA 2 arizonae Casey, Brachygluta 54 brendeli Casey, Decarthron 66 arizonensis Carlton, Eutrichites 65 brendeli LeConte, Rybaxis 53 arizonica Casey, Simplona 30 brevicorne Reichenbach, Trimium 34 112 COLEÓPTERA CATALOG brevicornis Casey, Atinus 86 cincinnata Casey, Nisaxis 65 brevipenne Casey, Actium 26 Cinderella Casey, Pilopius 85 brevipennis Casey, Oropus 7 Circocerus Motschulsky 83 Briaraxis Brendel 64 clamer Grigarick and Schuster, Reichenbachia 58 Briaraxys Raffray 64 claudus Schuster and Marsh, Hesperotychus 79 Bryaxis Leach 53 clavata Brendel, Rybaxis 53 Bryaxis (of authors) 54, 56 clavicornis Mäklin, Cupila 31 bulbifer Casey, Arthmius 49 CLAVIGERINAE 88 bulla Schuster, Mayetia 36 CLAVIGERINI 88 burdicki Schuster and Marsh, Pselaptrichus . 70 clypeonotus Brendel, Batrisodes 43 BYTHININA 69 clypeospecus Park, Batrisodes 39 BYTHININI 69 cochisei Chandler, Actium 27 bythinioides Brendel, Tychobythinus 74 coecus LeConte, Adranes 88 bythinoides (error) Brendel, Tychobythinus 74 cognatus LeConte, Lucifotychus 76 Bythinophysis Raffray 69 comes Casey, Cylindrarctus 75 BYTHINOPLECTINA 36 compar LeConte, Reichenbachia 57 BYTHINOPLECTINAE 36 compar LeConte, Tyrus 81 BYTHINOPLECTINI 36 complectens LeConte, Reichenbachia 58 Bythinoplectus Reitter 36 confinis LeConte, Batriasymmodes 47 Bythinopsis Raffray 74 confluens LeConte, Euplectus 19 Bythinus (of authors) 69, 74 confusus Chandler, Lucifotychus 76 Caccoplectus Sharp 86 congener Brendel, Reichenbachia 58 calcaris Grigarick and Schuster, Actium 26 congener Casey, Pycnoplectus 21 californicum LeConte, Actium 26 conifera Chandler, Sonoma 3 caUfornicus Brendel, Fustiger 88 conjuncta LeConte, Rybaxis 53 californicus Casey, Euplectus 19 Conoplectus Brendel 6 canadensis Brendel, Reichenbachia 60 consimilis Casey, Tyrus 81 canal iculatus LeConte, Conoplectus 6 consobrinus LeConte, Pilopius 85 candidum Casey, Actium 26 convexula LeConte, Trimiomelba 34 Canthoderus Motschulsky 35 convexus Casey, Oropus 7 capitula Casey, Ramecia 16 copelandi Carlton, Arianops 50 carinata Park and Wagner, Hatchia 15 copelandi Carlton and Chandler, Pseudactium 17 carinatus Brendel, Machaerodes 69 copelandi Park, Prespelea 80 carinatus Marsh and Schuster, Pselaptrichus 70 corniventris Motschulsky, Brachygluta 55 carinatus Say, Tmesiphorus 84 cornus Chandler, Pselaptrichus 70 carinifer Casey, Tyrus 81 cornus Grigarick and Schuster, Lucifotychus 76 carolinae Casey, Batriasymmodes 47 cornutum Brendel, Anchylarthron 65 carolinae Casey, Pseudactium 17 corporalis Casey, Reichenbachia 58 carolinae Casey, Tychobythinus 74 corticina Casey, Sonoma 3 cartwrighti Sanderson, Batrisodes 41 corticinus Casey, Tyrus 81 cascadia Chandler, Sonoma 3 costale Brendel, Actiastes 29 caseyi Blatchley, Batrisodes 41 costal is LeConte, Tmesiphorus 84 caseyi Raffray, Reichenbachia 58 coweeta Barr, Arianops 50 castaneus Casey, Oropus 7 creek Chandler, Bibloplectus 24 caudata Schaeffer, Nisaxis 64 crenatus Schuster and Grigarick, Rhexidius 5 cavatus Marsh and Schuster, Pselaptrichus 70 cribricollis Brendel, Reichenbachia 59 cavernensis Park, Arianops 50 crinifer Casey, Cylindrarctus 75 cavernosus Park, Batrisodes 43 crinita Brendel, Ramecia 16 cavicauda Casey, Oropus 7 cristatus LeConte, Batriasymmodes 47 caviceps Casey, Anchylarthron 65 cristatus Schuster and Grigarick, Oropus 7 cavicollis LeConte, Acolonia 19 croceus Park, Speleochus 73 cavicornis Brendel, Brachygluta 55 cruralis Park, Cedius 82 cavicornis Casey, Batrisodes 38 Ctenisis Raffray 84 cavicrus Casey, Batriasymmodes 47 Ctenistes (of authors) 85, 86 cavifrons Casey, Sonoma 3 CTENISTINI 84 cavifrons LeConte, Thesium 13 cuneata Marsh and Schuster, Sonoma 3 cediosus Wagner, Pycnoplectus 21 Cupila Casey 31 Cedius LeConte 81 curiosus Park, Pselaptrichus 73 CEOPHYLLINI 84 curtipenne Casey, Anchylarthron 65 Ceophyllus LeConte 84 curtipennis Casey, Oropus 8 cephalicum Casey, Pseudactium 17 curvatus Sanderson, Batriasymmodes 47 cephalotes Casey, Batrisodes 46 curvicera Motschulsky, Brachygluta 55 Cercocerus (error) LeConte 83 cuspidatus Schuster and Grigarick, Rhexidius 5 chandleri Schuster and Marsh, Pselaptrichus 70 cuspidatus Schuster and Marsh, Pselaptrichus 70 Cherokee Chandler, Bibloplectus 24 Custotychus Park and Wagner 78 chickasaw Chandler, Bibloplectus 24 Cutrimia Park 31 chiloquinensis Grigarick and Schuster, Actium 26 Cylindrarctus L. W. Schaufuss 75 chiricahuensis Raffray, Reichenbachia 58 cylindrarta Brendel, Reichenbachia 59 choctaw Chandler, Bibloplectus 24 Cylindrembolus L. W. Schaufuss 66 chuskae Chandler, Actizona 25 cyranus Schuster and Grigarick, Oropus 8 cicatricosus Brendel, Batrisodes 42 daggyi Park, Custotychus 78 PSELAPHIDAE 113

Dalmodes (of authors) 69 EUPSENIINA 69 Dalmosella Casey 34 Eupsenius LeConte 69 dasycerca Schuster and Marsh, Tetrascapha 11 Eupsinus (error) Motschulsky 69 davisi Park, Lemelba 32 eurous Park, Subterrochus 73 debilis Casey, Oropus 8 Eusanops Brendel 49 debilis LeConte, Thesiastes 18 Eutrichites LeConte 65 DECARTHRINA 66 Eutyphlus LeConte 14 Decarthron Brendel 66 excavata Park and Wagner, Cupila 31 decipiens Raffray, Actium 26 excavatum Chandler, Allotrimium 33 declivis Casey, Batrisodes 41 excavatus Bowman, Pycnoplectus 21 Dieclivodes Park 41 Excavodes Park 43 decora Casey, Saxet 18 excavoides Carlton, Conoplectus 7 defectum Park, Decarthron 67 exiguum Notman, Decarthron 67 deformata LeConte, Reichenbachia 59 exilis Bowman, Bibloplectus 24 delimatus Schuster and Grigarick, Oropus 8 exilissimus Casey, Leptoplectus 23 demissa Casey, Reichenbachia 59 exsectum Brendel, Decarthron 67 dentata Park and Wagner, Tetrascapha 11 extera Barr, Arianops 50 dentata Say, Brachygluta 55 Extollodes Park 48 dentatus Grigarick and Schuster, Lucifotychus 76 facilis Casey, Reichenbachia 60 denticauda Casey, Batrisodes 42 falcatum Grigarick and Schuster, Actium 27 denticollis Casey, Batrisodes 45 falcatus Wagner, Pycnoplectus 21 dentipes Raffray, Euphalepsus 49 Faliscus Casey 13 dentipes Raffray, Melba 33 falli Casey, Reichenbachia 60 dentiventris Casey, Pseudactium 17 FARONINAE 2 depressa Brendel, Briaraxis 64 FARONINI 2 depressifrons Brendel, Reichenbachia 59 Faronus (of authors) 2 desertorum Grigarick and Schuster, Actiastes 29 fastigium Grigarick and Schuster, Actium 27 dietzi Schaeffer, Adranes 88 fastosum Grigarick and Schuster, Actium 27 difficilis LeConte, Pycnoplectus 21 femineum Grigarick and Schuster, Actium 27 digitata Barr, Arianops 50 fenderi Park and Wagner, Abdiunguis . 14 dilatatus Motschulsky, Eupsenius 69 fenderi Park and Wagner, Oropus 8 dilatipes Schaeffer, Reichenbachia 58 fenyesi Schuster and Marsh, Hesperotychus 79 dilopha Marsh and Schuster, Sonoma 3 ferox LeConte, Batriasymmodes 47 Diplectellus Reitter 20 ferrugineus Casey, Rhexius 11 discolor Brendel, Decarthron 67 ferulifer Park, Batrisodes 39 discolor LeConte, Trimium 34 ferus Park, Subterrochus 74 discreta Casey, Pseudactium 17 festinatus Park, Batrisodes 43 disjunctus Casey, Euplectus 21 fideli Grigarick and Schuster, Euplecterga 16 distans Casey, Reichenbachia 58 filiformis Casey, Euplectus 20 divergens LeConte, Reichenbachia 59 fistula Schuster, Mayetia 37 diversum Park, Decarthron 67 flavitarsis L. W. Schaufuss, Circocerus 84 dolabra Marsh and Schuster, Sonoma 3 floridae Casey, Pycnoplectus 22 domestica Schuster, Mayetia 37 floridana Brendel, Brachygluta 55 dorothyae Park and Wagner, Reichenbachia 59 floridanus Casey, Pilopius 85 dubia LeConte, Trimomelba 34 fluvialis Chandler, Cylindrarctus 75 durum Brendel, Melba 32 formicarius Casey, Biotus 86 duryi Casey, Euplectus 19 formiceti LeConte, Decarthron 67 dybasi Carlton and Allen, Euboarhexius 10 formosum Grigarick and Schuster, Actium 27 dybasi Chandler, Simplona 30 fossicauda Casey, Batrisodes 41 dybasi Grigarick and Schuster, Oropodes 15 fossiger Casey, Melba 33 dybasi Park, Eutyphlus 14 fossulatus Brendel, Thesiastes 18 ectofacilis Park, Reichenbachia 59 fovealis Barr, Arianops 50 eisenachi Park, Nearctitychus 78 foveata LeConte, Brachygluta 55 electrae Park, Hamotus 83 foveicollis LeConte, Actiastes 29 elegans Brendel, Brachygluta 55 foveicornis Casey, Batrisodes 39 elongatus Brendel, Euplectus 20 Foveoscapha Park and Wagner 11 elongatus Brendel, Hamotus 83 fovicinus Grigarick and Schuster, Actiastes 29 Elytrodes Park 41 franciscana Casey, Reichenbachia 57 emoyteis Grigarick and Schuster, Reichenbachia 59 frigidus Schuster and Marsh, Pselaptrichus 70 Empinodes Park 41 frontalis Casey, Valda 75 erichsoni LeConte, Pselaphus 87 frontalis LeConte, Batrisodes 44 Euboarhexius Grigarick and Schuster 10 frosti Park, Batriasymmodes 48 Euphalepsus Reitter 49 fuchsii Brendel, Fustiger 88 Euplecterga Park and Wagner 15 functum Grigarick and Schuster, Actium 27 EUPLECTINA 15 fiindata Casey, Reichenbachia 57 EUPLECTINAE 5 fundatum Grigarick and Schuster, Actiastes 29 EUPLECTINI . 12 furcatus Brendel, Batrisodes 44 Euplecturga (error) Grigarick and Schuster 15 furtiva Casey, Reichenbachia 59 Euplectus Leach 19 fusciventris Casey, Reichenbachia 59 Euplenius (error) Motschulsky 69 Fusjugama Park 80 Euprenius (error) Raffray 69 fusticornis Casey, Reichenbachia 60 114 COLEÓPTERA CATALOG fustifer Casey, Pselaphus 87 impunctatum Brendel, Actium 27 Fustiger LeConte 88 incognitus Schuster and Marsh, Pselaptrichus 71 geminata Fall, Rybaxis 53 incomptus Schuster and Grigarick, Rhexidius 6 gemmifer LeConte, Reichenbachia 60 indistinctus Grigarick and Schuster, Batrisodes 42 gemmoides Park, Batrisodes 39 inepta Casey, Reichenbachia 61 gemmus Park, Batrisodes 39 infinita Casey, Brachygluta 55 gentilis Fletcher, Reichenbachia 60 informis Casey, Reichenbachia 61 georgianus Casey, Pilopius 85 infossus Raffray, Pycnoplectus 21 gibbosus Marsh and Schuster, Pselaptrichus 70 INIOCYPHINI 69 gigantea Barr, Arianops 50 inopia Casey, Reichenbachia 61 glaber LeConte, Eupsenius 69 inornatum Brendel, Anchylarthron 66 glaberula Casey, Anitra 83 inornatus Grigarick and Schuster, Lucifotychus 77 glareosus Schuster and Grigarick, Rhexidius 5 insculptus LeConte, Rhexius 11 globicollis LeConte, Arthmius 49 insolens Casey, Leptoplectus 23 globiferum LeConte, Actiastes 30 insólita Casey, Reichenbachia 61 globosus LeConte, Batrisodes 44 integer LeConte, Bibloplectus 24 gloydi Park, Bythinoplectus 36 intellectus Grigarick and Schuster, Lucifotychus 77 GONIACERINAE 52 intermedia Brendel, Brachygluta 55 gracile Brendel, Melba 32 intermedius Brendel, Euboarhexius 10 gracilicornis Casey, Reichenbachia 60 interruptus Casey, Oropus 8 gracilior Casey, Arthmius 49 interruptus LeConte, Pycnoplectus 22 gracilis Casey, Reichenbachia 60 intimus Schuster and Marsh, Pselaptrichus 71 grandiceps Casey, Sonoma 3 intricata Casey, Brachygluta 56 granicollis Casey, Pilopius . . . . 85 inusitatum Park, Decarthron 67 granulosus Casey, Rhexidius 6 involutus Casey, Arthmius 49 grayae Schuster, Mayetia 37 ionae LeConte, Batrisodes 41 greeveri Park, Batriasymmodes 47 iowensis Casey, Euplectus 20 grigaricki Chandler, Oropus 8 iowensis Casey, Pilopius 85 grubbsi Chandler, Batrisodes 44 isabellae LeConte, Sonoma 4 hairstoni Park, Batrisodes 39 ishii Chandler, Oropodes 15 HAMOTINA 83 jacobina Casey, Brachygluta 56 Hamotoides L. W. Schaufuss 83 jeanneli Park, Arianops 50 Hamotus Aube 83 jeanneli Park, Batriasymmodes 48 hardyi Park, Reichenbachia 60 jocuvestus Park, Batrisodes 39 harringtoni Casey, Batrisodes 44 jonesi Park, Batrisodes 39 hatchi Park and Wagner, Actium 28 jonesi Park, Tychobythinus 74 Hatchia Park and Wagner 15 JUBINl 35 hazeltinei Chandler, Actium 27 judsoni Schuster, Mayetia 37 helferi Grigarick and Schuster, Actium 27 justum Park, Decarthron 67 helferi Grigarick and Schuster, Oropus 8 juvencus Brendel, Batrisodes 46 helferi Park and Schuster, Trisignis 12 kahli Bowman, Batrisodes 44 helferi Schuster and Marsh, Pselaptrichus 70 kansana Casey, Reichenbachia 61 henroti Park, Arianops 50 kapones Grigarick and Schuster, Reichenbachia 61 henroti Park, Batrisodes 39 karsteni Reichenbach, Euplectus 20 héspera Park and Wagner, Sonoma 3 keeni Casey, Oropus 8 Hesperotychus Schuster and Marsh . 78 kingi Barr, Arianops 50 hexagonus Casey, Hesperotychus 79 knausii Schaeffer, Fustiger 88 Hexirhexius Grigarick and Schuster 6 krekeleri Park, Batrisodes 39 hirsutus Casey, Rhexius 12 krysiae Chandler, Tetrascapha 11 hirsutus Chandler, Lucifotychus 76 labyrinthea Casey, Brachygluta 56 hispidus Schuster and Grigarick, Rhexidius 6 lacustris Casey, Pilopius 85 hocus Schuster and Marsh, Pselaptrichus 71 laevis Casey, Trimiomelba 34 horni LeConte, Scalenarthrus 66 laminata Barr, Arianops 51 howardi Park, Reichenbachia 61 langei Schuster, Mayetia 37 howdeni Park, Decarthron 67 laticolle Brendel, Melba 32 howdeni Park, Reichenbachia 62 laticolle Casey, Thesium 13 hubrichti Park, Batrisodes 39 latipes Fletcher, Reichenbachia 58 hubrichti Park, Tychobythinus 74 laurenticum Casey, Decarthron 67 hudsonicus Casey, Pycnoplectus 22 lecontei Brendel, Adranes 89 humeralis Aube, Tyrus 81 Lemelba Park 32 humilis Marsh and Schuster, Sonoma 3 lentus Schuster and Marsh, Megarafonus 2 Hylotychus Grigarick and Schuster 76 Leptoplectus Casey 22 idahoensis Park and Wagner, Euplectus 20 leviceps Casey, Bibloplectus 24 ignobilis Fletcher, Reichenbachia 61 levinei Schuster and Marsh, Pselaptrichus 71 illinoiensis Brendel, Brachygluta 55 levini Chandler, Mipseltyrus 82 impellus Park and Wagner, Lucifotychus 77 linearis LeConte, Pycnoplectus 22 impensus Schuster and Grigarick, Rhexidius 6 lineaticollis Aube, Batrisodes 41 imperfectus Casey, Pycnoplectus 22 Liniolis Grigarick and Schuster 16 impressiceps Casey, Pycnoplectus 22 litoralis Brendel, Reichenbachia 61 impressicoUis Park and Wagner, Euplecterga 16 longiclavus LeConte, Pselaphus 87 impressipennis Casey, Pilopius 85 longicollis Casey, Euplectus 20 PSELAPHIDAE 115 longicollis Casey, Sonoma 3 nebulosus Grigarick and Schuster, Batrisodes 42 longipalpis LeConte, Cylindrarctus 75 neglecta Barr, Arianops 51 longipennis Casey, Pycnoplectus 22 Neopselaphus Jeannel 87 longissimus Brendel, Pycnoplectus 21 nevadensis Casey, Reichenbachia 61 longulum LeConte, Decarthron 67 newelli Park and Wagner, Lucifotychus 77 loripes Casey, Brachygluta 56 nicolayi Park, Mipseltyrus 82 Lucifotychus Park and Wagner 76 Nicothaeus (error) Raffray 14 luculentus Casey, Batrisodes 40 Nicotheus Casey 14 ludovicianus Brendel, Cylindrarctus 75 nigricans LeConte, Batrisodes 45 luniger LeConte, Brachygluta 56 nigrocavum Park, Decarthron 68 lustrans Casey, Batrisodes 42 Nisa Casey 54 macclayi Schuster and Marsh, Hesperotychus 79 Nisaxis Casey 64 Machaerites (of authors) 73 nodesa Grigarick and Schuster, Reichenbachia 62 MACHAERITINA 73 nodicollis Raffray, Hamotus 83 Machaerodes Brendel 69 nodosa Barr, Arianops 51 macneilli Schuster and Grigarick, Oropus 8 norithe Barr, Arianops 51 MACROSCELIA 81 norstelcha Grigarick and Schuster, Euplecterga 16 magaliae Chandler, Pselaptrichus 71 nuciere Grigarick and Schuster, Oropodes 15 magazinensis Carlton and Chandler, Pseudactium 17 nuttingi Chandler, Caccoplectus 86 magnidens Schuster and Grigarick, Oropus 8 obliqua Barr, Arianops 51 maja Brendel, Melba 32 obliquedens Fall, Rybaxis 53 mancus Schuster and Marsh, Megarafonus 2 obrieni Chandler, Cylindrarctus 76 margemina Park and Wagner, Sonoma 4 obscurus Grigarick and Schuster, Batrisodes 42 marinicum Casey, Actium 28 obsoletus Brendel, Trimioplectus 23 marinum Brendel, Decarthron 68 obsoletus Park, Cedius 82 marítima Casey, Nisaxis 64 obtusus Schuster and Grigarick, Oropus 9 marshi Chandler, Pselaptrichus 71 ocampi Park, Sebaga 35 marshi Park and Schuster, Trisignis 12 occidens Casey, Morius 13 martini Grigarick and Schuster, Batrisodes 42 occiduus Casey, Batrisodes 42 Mayetia Mulsant and Rey 36 ocularis Casey, Pilopius 85 MAYETIINI 36 oculatus Marsh and Schuster, Pselaptrichus 71 Megarafonus Casey 2 ojaiensis Grigarick and Schuster, Actium 28 Melba Casey 32 olycalida Park and Wagner, Sonoma 4 mellineum Casey, Pseudactium 17 onaga Chandler, Cylindrarctus 76 mendicum Park, Pseudactium 17 opacus Grigarick and Schuster, Batrisodes 43 mendocino Casey, Batrisodes 42 opimus Fletcher, Hamotus 83 mendocinoensis Schuster, Mayetia 37 orbatus Schuster and Grigarick, Oropus 9 mexicanus Park, Neopselaphus 88 orbiceps Casey, Oropodes 15 microcephalum Raffray, Pseudotrimium 35 oriental is Chandler, Cylindrarctus 76 microphthalmum Park and Wagner, Actium 28 ornatus Marsh and Schuster, Pselaptrichus 71 micropthalmus Brendel, Hesperotychus 79 oro Chandler, Batrisodes 43 micropthalmus Chandler, Oropus 8 Oropodes Casey 15 mimicum Park, Decarthron 68 Oropus Casey 7 minimus Schuster and Grigarick, Oropus 9 osage Chandler, Bibloplectus 24 minimus Schuster and Marsh, Pselaptrichus 71 ouachita Chandler, Rhexius 12 minor (of authors) 78 Ouachitychus Chandler 80 minor LeConte, Custotychus 78 pacificum Casey, Actium 28 minuta Brendel, Nisaxis 65 pacificus Wickham, Adranes 89 minutus Bowman, Trigonoplectus 25 pajarito Chandler, Trimioarcus 35 Mipseltyrus Park 82 pallidum Casey, Actium 26 mirus Schuster, Mipseltyrus 82 PANAPHANTINA 13 mississippiensis Park, Batrisodes 44 pandum Grigarick and Schuster, Actium 28 mollyae Park, Pygmactium 30 pannosus Park, Batrisodes 39 monilicornis Brendel, Atinus 86 parabolicum Brendel, Pseudactium 17 monilis LeConte, Ceophyllus 84 parki Barr, Arianops 51 monstrosus LeConte, Batriasymmodes 47 parki Barr, Batriasymmodes 48 montanus Casey, Oropus 9 parki Schuster and Marsh, Pselaptrichus 71 montícola Casey, Batrisodes 42 parki Schuster, Mipseltyrus 82 moratus Schuster and Marsh, Hesperotychus 79 parki Wagner, Pycnoplectus 22 Morius Casey 13 parviceps Casey, Nisaxis 64 mormon Bowman, Brachygluta 56 parviceps Mäklin, Sonoma 4 morsus Fletcher, Arthmius 49 parvoculus Chandler, Ouachitychus ^0 multifossa Grigarick and Schuster, Cupila 32 párvula LeConte, Melba 32 musamator Chandler, Trimioarcus 35 parvus Schuster and Marsh, Megarafonus 2 mystica Casey, Rybaxis 53 pattersoni Park, Reichenbachia 63 Nanorafonus Schuster and Marsh 2 pearsei Schuster, Mayetia 37 nantahalae Joanna Barr, Arianops 51 pecki Barr, Arianops 51 nantahalae nantahalae Barr, Arianops 51 ■ pecki Carlton and Chandler, Pseudactium 17 nanus Schuster and Marsh, Hesperotychus 79 pectinatus Chandler, Caccoplectus 86 NATYPLEURINA 69 pectinis Schuster and Grigarick, Oropus 9 Nearctitychus Chandler 78 pennatus Schuster and Marsh, Pselaptrichus 71 116 COLEÓPTERA CATALOG pennsylvanicum Bowman, Actiastes 29 raneyi Schuster, Mayetia 37 perditus Schuster and Marsh, Pselaptrichus 72 rayi Park, Decarthron 68 peregrinator Casey, Reichenbachia 62 reciprocum Park, Decarthron 68 perfidus Schuster and Marsh, Pselaptrichus 72 rectus Marsh and Schuster, Pselaptrichus 72 perfórala Brendel, Brachygluta 56 reddelli Barr and Steeves, Texamaurops 46 perpunctata Brendel, Brachygluta 56 reduncus Park, Batriasymmodes 48 perscitus Fletcher, Euboarhexius 10 Reichenbachia Leach . . 56 pertenuis Casey, Leptoplectus 23 Reichenbachius Casey 56 petersi Chandler, Sonoma 4 remipennis Grigarick and Schuster, Lucifotychus 77 phantasma Park, Rybaxis 54 repanda Marsh and Schuster, Sonoma 4 piceus LeConte, Pilopius 85 retractum Casey, Actium 28 Pilactium Grigarick and Schuster 31 reyesi Chandler, Batrisodes 44 Pilopius Casey 85 Rhexidius Casey 5 planipennis Brendel, Thesiastes 18 RHEXIINA 11 Planityphlus Park 14 Rhexius LeConte 11 plectrops Casey, Arianops 51 RHINOSCEPSINA 12 plumas Chandler, Lucifotychus 77 Rhinoscepsis LeConte 12 pluridentata Park, Reichenbachia 62 Rhinosceptis (error) Reitter 12 plusculus Park and Wagner, Pselaptrichus 72 Rhynoscepsis (error) Raffray 13 pocahontas Casey, Custotychus 78 Rhynosceptis (error) Raffray 13 polita Brendel, Reichenbachia 62 ripariae Grigarick and Schuster, Bontomtes 10 politum Casey, Actium 28 riparius Say, Batrisodes 40 populus Chandler, Hamotus 83 robustulum Casey, Actium 28 pravitas Schuster, Mayetia 37 robustus Casey, Cedius 82 Prespelea Park 80 rossi Park, Batrisodes 40 priocera Marsh and Schuster, Sonoma 4 rostratrum Park, Decarthron 68 procera Casey, Reichenbachia 62 rostratus Bowman, Trigonoplectus 25 profundus Park, Batrisodes 40 rostratus L. W. Schaufuss, Tmesiphorus 84 prominens Casey, Eutyphlus 14 rothi Grigarick and Schuster, Actium 28 propinqua LeConte, Reichenbachia 62 rothi Park, Pselaptrichus 72 propinquus Schuster and Marsh, Pselaptrichus 72 rotundicollis Brendel, Thesiastes 19 Proplectus Raffray 25 ruber Casey, Rliexius 11 proprius Schuster and Marsh, Pselaptrichus 72 rubicunda (error), Reichenbachia 64 Prorhexius Raffray 6 rubida Casey, Sonoma 4 PSELAPHIDAE 1 rubricunda Aube, Reichenbachia 64 PSELAPHINAE 51 ruficeps Motschulsky, Bibloplectus 24 PSELAPHINI 87 rufus LeConte, Eupsenius 69 Pselaphus Herbst 87 rumseyensis Grigarick and Schuster, Oropodes 15 PSELAPTINA 65 russelli Chandler, Sonoma 4 Pselaptrichus Brendel 70 Rybaxis Saulcy 53 Pselaptus LeConte 66 sagax LeConte, Reichenbachia 63 Pseudactium Casey 16 saginatus Casey, Pilopius 86 Pseudotrimiuiti Raffray 35 Sagola (of authors) 2 puberulus LeConte, Lycifotychus 77 sandersoni Barr, Arianops 51 Pubimodes Park 45 sandersoni Park, Batriasymmodes 48 pulvereus LeConte, Pilopius 86 sanguínea (of authors), Rybaxis 54 pumilis LeConte, Thesiastes 19 Saxet Grigarick and Schuster 18 punctatus LeConte, Batrisodes 46 scabra Brendel, Reichenbachia 63 puncticollis LeConte, Reichenbachia 62 scabriceps LeConte, Batrisodes 44 puncticollis LeConte, Zonaira 33 Scalenarthrus LeConte 66 punctifrons Casey, Batrisodes 40 scarificatum Brendel, Decarthron 68 putahensis Schuster, Mayetia 37 schaefferi Park, Batrisodes 40 Pycnoplectus Casey 21 schaufussi Raffray, Bythinophysis 69 pygidialis Casey, Batrisodes . . 42 schaumi Aube, Batrisodes 46 pygidialis Raffray, Reichenbachia 62 schmitti Brendel, Rhexius 12 Pygmactium Schuster and Grigarick 30 schmitti Casey, Batrisodes 45 Pytna Casey 81 schmitti Raffray, Euplectus 21 quercae Chandler, Melba 33 schmitti Raffray, Eutyphlus 14 quercavum Chandler, Pygmactium 31 schneiderensis Park, Batrisodes 40 quercicola Chandler, Sonoma 4 schuhi Chandler, Lucifotychus 77 quirsfeldi Chandler, Lucifotychus 77 schusteri Chandler, Oropus 9 quirsfeldi Park, Prespelea 80 scobina Schuster, Mayetia 37 quisnamus Park, Batriasymmodes 48 Sebaga Raffray 35 Racemia Newton and Chandler . r 16 semiruber Casey, Tyrus 81 radians LeConte, Reichenbachia 62 semirugosa Brendel, Reichenbachia 63 raffrayi Brendel, Oropodes 15 sentis Chandler, Caccoplectus 87 raffrayi Casey, Ctenisis . 85 seriepunctatum Brendel, Decarthron 68 Rafonus Casey 3 setosum Bowman, Decarthron 68 Ramecia (of authors) 16 sewanee Barr, Arianops 51 Ramecia Casey 16 sexualis Casey, Pycnoplectus 22 ramus Park and Wagner, Hesperotychus 79 shastensis Schuster and Marsh, Pselaptrichus 72 PSELAPHIDAE 117

shenefelti Park, Reichenbachia 62 sulcatula Casey, Melba 33 signatus Reichenbach, Euplectus 20 sulcicollis Reichenbach, Trichonyx 52 silvanus Schuster and Marsh, Pselaptrichus 72 summersi Grigarick and Schuster, Pilactium 31 silvicolus Chandler, Euplectus 20 susae Carlton, Conoplectus 7 similis LeConte, Eutyphlus 14 suteri Park, Actiastes 30 similis Schuster and Marsh, Pselaptrichus 72 suteri Park, Batriasymmodes 48 simplex LeConte, Batrisodes 46 sylvaticus Raffray, Conoplectus 6 simplex LeConte, Melba 33 synstygicus Park, Speleochus 73 simplex Motschulsky, Rhexius 12 tantillus Schuster and Marsh, Hesperotychus 80 simplicis Grigarick and Schuster, Lucifotychus 77 taphrocera Casey, Reichenbachia 63 Simplona Casey 30 taylori Wickham, Adranes 89 sinifiindus Schuster and Grigarick, Oropus 9 temporalis Casey, Batrisodes 45 Sinistrocedius Newton and Chandler 82 tempus Grigarick and Schuster, Actium 28 Sinistrocedius Park 82 tenellum Grigarick and Schuster, Actium 28 sinuatifrons Brendel, Batrisodes 44 tenellus Casey, Pycnoplectus 21 sinus Grigarick and Schuster, Euboarhexius 10 tenellus LeConte, Hesperotychus 80 skeeteri Royce and Kistner, Mayetia 37 tentum Grigarick and Schuster, Actium 29 smithi Schuster, Mayetia 38 tenuis Casey, Dalmosella 34 snowi Fletcher, Reichenbachia 63 tenuis LeConte, Thesiastes 18 snowi Park, Decarthron 68 tenuis Schuster and Marsh, Pselaptrichus 72 sobrinus Casey, Bibloplectus 24 terebrata Casey, Brachygluta 56 sodalis Casey, Reichenbachia 63 terracola Park and Wagner, Foveoscapha 11 Sognorus (of authors) 85 testaceum Casey, Actium 29 Sonoma Casey 2 testaceus Casey, Lucifotychus 77 sonomae Casey, Lucifotychus 77 testaceus Casey, Oropus 9 sonorensis Park, Reichenbachia 61 Tetrascapha Schuster and Marsh 10 spadica Marsh and Schuster, Sonoma 4 Texamaurops Barr and Steeves 46 spatium Grigarick and Schuster, Actiastes 30 texana Casey, Brachygluta 56 spatulifer Casey, Reichenbachia 63 texana Casey, Melba 32 speculum Casey, Batrisodes 43 texanus Chandler, Batrisodes 45 specus Park, Batrisodes 40 teyahalee Barr, Arianops 52 spelaeus Park, Batriasymmodes 48 Thesiastes Casey 18 Speleobama Park 80 Thesium Casey 13 SPELEOBAMINI 80 thoracica Brendel, Melba 33 Speleochus Park 73 thoracicus Park, Eutyphlus 14 Speleodes Park 48 thornei Barr, Arianops 52 spiculifer Casey, Custotychus 78 tibialis Casey, Eutyphlus 14 Spifemodes Park 46 TMESIPHORINI 84 spinicollis Barr, Arianops 52 Tmesiphorus LeConte 84 spinifer Brendel, Batrisodes 45 tolulae LeConte, Sonoma 5 spinifer Casey, Pycnoplectus 22 tomentosa Aube, Nisaxis 65 spinipes Schaeffer, Caccoplectus 87 Tomoplectus Raffray 35 spinosus LeConte, Cedius 81 transversa Fall, Rybaxis 54 spinosus Marsh and Schuster, Pselaptrichus 72 transversus Casey, Rhexius 11 spretus LeConte, Batrisodes 40 triangulifer Brendel, Batrisodes 45 stangei Grigarick and Schuster, Hesperotychus 79 TRICHONYCHINI 52 stannardi Carlton and Chandler, Pseudactium 17 Trichonyx (of authors) 7 steevesi Barr, Arianops 52 Trichonyx Chaudoir 52 steevesi Carlton and Chandler, Pseudactium 17 tridens Casey, Batrisodes 43 steevesi Park, Subterrochus 74 trifoveatum Park, Actizona 25 steevesi Schuster and Grigarick, Pygmactium 31 trigma (error) Brendel, Reichenbachia 62 stellatus Grigarick and Schuster, Lucifotychus 77 trigona LeConte, Reichenbachia 62 stephani Carlton, Arianops 52 Trigonoplectus Bowman 25 stephani Chandler, Rhexius 12 trilinea Grigarick and Schuster, Zonaira 34 stephani Grigarick and Schuster, Allobrox 31 triloba Marsh and Schuster, Sonoma 5 Sternalis Raffray, Nearctitychus 78 TRIMIINA 25 stigmosum Brendel, Decarthron 68 Trimioarcus Park 35 Stratus L. W. Schaufuss 35 Trimiomelba Casey 34 strenuum Brendel, Decarthron 68 Trimioplectus Brendel 23 striatus LeConte, Oropus 9 Trimium (of authors) 32 striatus psotai Park, Batrisodes 46 Trimium Aube 34 striatus striatus LeConte, Batrisodes 45 TRISIGNINA 12 strinatii Besuchet, Tychobythinus 74 Trisignis Park and Schuster 12 stygica Park, Arianops 52 trogasteroides Brendel, Euboarhexius 10 stygicus Park, Speleochus 73 TROGASTRINA 5 subsimilis Casey, Reichenbachia 57 TROGASTRINI 5 subsimilis Casey, Sonoma 3 troglodytes Park, Batriasymmodes 48 substriatus LeConte, Rhexius 12 trúncala Barr, Arianops 52 subterraneus Park, Batrisodes 40 truncaticornis Brendel, Rybaxis 54 Subterrochus Park 73 tuberculatum Chandler, Allotrimium 33 subtilis LeConte, Reichenbachia 63 tuberculatus Schuster and Grigarick, Oropus 9 118 COLEÓPTERA CATALOG tuberculipalpus Brendel, Pselaptrichus 73 vana Park, Speleobama 80 tulareanus Casey, Batrisodes 43 vanna Marsh and Schuster, Sonoma 5 túmida LeConte, Reichenbachia 63 vanus Schuster and Marsh, Pselaptrichus 73 tumidicornis Casey, Reichenbachia 63 varicornis Brendel, Rybaxis 54 tumoris Park, Batrisodes 40 varicum Park, Decarthron 68 tumorosa Casey, Reichenbachia 64 vellosus Schuster and Grigarick, Oropus 9 turba Park, Reichenbachia 61 velutinum LeConte, Decarthron '67 turgidicomis Casey, Reichenbachia 64 ventralis Casey, Megarafonus 2 turneri Schuster, Mayetia 38 venustrus Schuster and Marsh, Pselaptrichus 73 TYCHINI 75 venyivi Chandler, Batrisodes 45 Tychobythinus Ganglbauer 74 verrucifundus Schuster and Grigarick, Oropus 10 tychoides Brendel 74 verticalis Casey, Custotychus 78 Tychus (of authors) 74, 75, 76, 78, 79 Verticinotus Brendel 65 TYRINA 81 Vestitrichus Park 73 TYRINI 81 virginiae Casey, Batrisodes 45 Tyrus Aube 81 virginicus Casey, Rhexius 12 ulkei Bowman, Pselaphus -. 87 visda Grigarick and Schuster, Reichenbachia 64 ulkei Brendel, Brachygluta 56 wagneri Park, Actiastes 30 umbraticus Schuster and Grigarick, Oropus 9 wagneri Park, Batriasymmodes 48 uncicornis Casey, Batrisodes 40 walker] Schuster, Mayetia 38 unicoi Barr, Arianops 52 wawonaensis Grigarick and Schuster, Actium 29 uniformis Casey, Melba 32 wickhami Brendel, Reichenbachia 64 Upoluna L. W. Schaufuss 84 XENOBYTHINA 70 ursina Casey, Reichenbachia 60 yahiorum Chandler, Sonoma 5 ursinus L. W. Schaufuss, Canthoderus 36 yavapai Chandler, Saxet 18 ursum Carlton, Pseudactium 18 zephyrinus Casey, Batrisodes 43 utahensis Tanner, Reichenbachia 60 ziegleri LeConte, Cedius 82 Valda Casey 74 zimmermani LeConte, Pilopius 86 VALDINI 74 zimmermanni LeConte, Eutrichites 65 valentinei Park, Batrisodes 41 Zonaira Grigarick and Schuster 33 valida Brendel, Rybaxis 54 zonatus Brendel, Eutrichites 65

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