30 April 1996 PROC. ENTOMOL. SOc. WASH. 98(2), 1996, pp. 173-187

NEOTROPICAL , VI: PROSETOMORPHA FALCATA, A NEW GENUS AND SPECIES OF SETOMORPHINAE (: TINEOIDEA) FROM COLOMBIA ASSOCIATED WITH CURCULIONID GALLERIES IN STEMS OF SOLANUM

DONALD R. DAVIS

Department of Entomology, National Museum of Natural History, MRC-127, Smith­ sonian Institution, Washington, DC 20560 U.S.A.

Abstract.-Adult and immature stages of Prosetomorpha jalcata, a new genus and species of tineid from Colombia, are described and illustrated. Prosetomorpha is remarkable in being the most plesiomorphic setomorphine tineid discovered, as evidenced by the five-segmented maxillary pal pi, moderately well developed haustellum, mandibles vestigial but distinct, female with signum present, and larva with six pairs of stemmata. Synapomorphies shared with other Setomorphinae include similar semi-appressed head vestiture, annulate but completely fused, eighth abdominal tergostemum in the male, mem­ branous eighth abdominal segment without rami in the female, and larvae with unidentate mandibles. The larva is believed to be primarily a scavenger within the bUlTows of a curculionid weevil, Faustinus apicalis (Faust), that bores in the stems of Solanum qui­ toensis var. quitoensis Lamarck and S. quitoensis var. septentrionale R. E. Schultes. A braconid parasitoid, Promicrogaster sp., has been reared from burrows occupied by Pro­ setomorpha. Key Words: Tineidae, Setomorphinae, Prosetomorpha, immatures, scavenger, host rec­ ord, Solanum, Curculionidae, Faustinus, Neotropical

In the course of their investigations on Morphological compruisons of the larva, the pests of Solanum quitoensis La­ pupa, and adult reveal that Prosetomorpha marck, known locally as "Iulo" or "nar­ is a member of the Setomorphinae, a small, anj illo," entomologists at the Instituto Col­ mostly neotropical subfamily, previously ombiano Agropecuario (lCA) of Bogota, represented by two genera (Setomorpha, Colombia, have reared a previously un­ Lindera) and approximately eight species known, stem-boring setomorphine moth, (Robinson and Nielsen 1993). Prosetomor­ Prosetomorpha Jalcata. The larva was pha is of phylogenetic interest in being the found frequently in association with larvae most plesiomorphic member of the subfam­ of the stem-boring weevil, Faustinus api­ ily, as evidenced by the five-segmented calis (Faust). Considering what is known of maxillary palpi (Figs. 13, 14), relatively the larval biology of other Setomorphinae (Hinton 1956, Gozmany and Vari 1973, well developed haustellum, mandibular ves­ Zimmerman 1978, Robinson and Nielsen tiges distinct, female with signum present 1993), the new species probably feeds pri­ (Figs. 24, 25), and larva with six pairs of marily as a scavenger within the tunnels of stemmata (Figs. 35, 55). The discovery of the weevil. Observations indicate, however, Prosetomorpha consequently broadens our that living plant tissue is also consumed. definition of the subfamily, which previ­ 174 PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON VOLUME 98 , ously had been characterized in part by of maxillary pal pus. Maxillary palpus 5­ with more reduced mouthparts and segmented, with ratios from base = 0.2: by larvae lacking stemmata (Hinton 1956, 0.18: 0.18: I: 0.33. Labial palpus 3-seg­ Zimmerman 1978, Robinson and Nielsen mented; ratios from base = 0.43: I: 0.88; 1993). Synapomorphies shared with other segment II smooth dorsally and laterally, Setomorphinae include si mi lar semi-ap­ with rough, triangular tuft ve nL raliy; bris Lles pressed head vestiture (Figs. 3, 4), slender absent; segment III entirel y smooth, de­ annulate eighth abdominal sclerite with pressed. stout projections directed anteriorly in the Thorax: Metafurcasternum (Fi gs. 16, 17) male (Fig. 19), membranous eighth abdom­ with anteromedial process (APM) elongate, inal segment without rami in the female slender, with a pair of fu rcal ap physes (Fig. 24), and larva with unidentate man­ (FA) directed sharply anlerio rl y; furcal dibles (Fig. 56). In addition to retaining the ridge continues caudall y and terminates as plesiomorphies listed above, Prosetomor­ a pair of rounded lobes arising inside lam­ pha differs most notably from Lindera and ina of secondary furcal arms (LSF). Fore­ Setomorpha by the presence of I and Y2 an­ wing (Fig. 15) falcate, abruptly indented nuli of flagellar scales (a basal, dorsally ab­ between apex and M I ; R 5-branched; R4 breviated row and an entire distal annulus and 5 stalked half th ir I ngth; R5 termi­ (Figs. 5, 6), falcate forewings, and a com­ nating before apex; all 3 branches of M sep­ pletely fused, eighth abdominal tergosternal arate; euA I and 2 arising approximate; A I ring in the male. In other Setomorphinae and 2 fused except for fork at basal third; the adults possess a single annulus of fla­ accessory cell present; base of media usu­ gellar scales, the forewings terminate in a ally not evident within discal cell; retinac­ simple rounded apex, and the eighth ab­ ulum of male a tightly curled flap (Figs. 9, dominal sclerite of the male is interrupted 10). Hindwi.lg with all veins arising sepa­ dorsally. rate; M 3-branched : base of media divided within cel!; male with a single stout frenu­ Prosetomorpha Davis, NEW GENUS lum; female with 2 smaller frenular br istl es. Type species.- Prosetomorpha falcata, Foreleg (Fig. I 1, 12, 18) with epiphysi. ca. new species. 0.5 the length of shorten d ti bia, the latter Adult.-Moderately large moths with equal in length to tarsomere I. Ti bial purs falcate forewings; length of forewing = = 0-2-4. Apices of a ll ti biae and tarso­ 8.5-13 mm. meres with 3 stout, spiniform setae (Fig. Head (Figs. 3, 4, 13, 14): Vertex mod­ 11) . erately rough with lateral tufts converging Abdomen (Fig. 19): Sternal apophyses mesally over occiput and projecting anteri­ slender, ca. half the length of S2. Eighth orly between antennae over frons; scales segment of male with a slender, scleroliz d slender with bidentate apices. Frons rela­ ring completely encircling segment; a pair tively smooth, scales directed mesally. An­ of slender projections arising laterally from tenna simple in both sexes; length ca. 113 of ring; T7 reduced, triangu lar. A pair of non­ forewing; scape cylinrical, without pecten; retractible coremata arising ventrally from Figs. 1- 6. mm. J, Head flagellum (Figs. 5, 6) with two rows of slen­ S8 of male. Corethrogyne of female con­ showing two der scales per segment dorsally, with only sisting of a dense ring of elongate, straight row of scal c ~ the distal row forming a complete ring. Pil­ setae. scale for all ~ ifers well developed, densely setose. Man­ Male genitalia: Tegumen reduced to a dibles (Fig. J3) present but vestigial, narrow dorsal ring. Vinculum similar to and equalling apical segment of maxillary pal­ contiuous with tegumen as a narrow ventral pus in length. Haustellum moderately de­ ring; saccus short, ca. 0.14 the length of veloped (Fig. 14), slightly exceeding length valva, compressed. Valva long, ca. 1.3 X the : WAS HI NGTO VOLUME 98, NUMB R 2 175 ary palpus 5­ 1 base = 0.2: palpus 3-seg­ 0.43: I: 0.88; and lat rail y, nll'ally ; bristles f smooth, de­

~ (Figs. 16. 17) lPM) elongate, 'cal apophyses !riorly ; fu rcal I terminates as 2 ng inside lam­ s (LSF). Fore­ uptly indented -branched; R4 glh; RS termi­ Ichc s of M sep­ lproxim ate; A l at basal third: of media usu­ II cell; reti nac­ d flap (Figs. 9, S ilrising sepa­ media divided ~ Ie stout fren u­ 'cnular bristles. h epiphysis ca. libia. the latter I. Tibial spurs liac and tarso­ nll setae (Fig.

'nal apophyses of S2. E ig hth der, sclerotized egment; a pair ~ latl:rally from A pair of non­ ve ntrally from Figs. 1-6. Proselomorpha jlilcaia. I, Adult male, forewing length 10.6 mm. 2, Female, forew ing length 14 mm. 3, Head, lateral view (0.75 mm). 4, Anterior view of Fig. 3. 5, Lateral view of antenna at Jistal third Dr female con­ showing two dor,,~1 rows of scales (annuli; 100 fJ.m). 6, Ventral view of Fig. 5 showing continuation of distal ngate, straight row of scales and position of sensilla coeloconicum (see arrow; 100 fJ.m ). (Scale lengths in parentheses; bar scale for a ll photographs = ("ig. 3.) 1 reduced to a 1simi lar to and narrow v ntral the length of Ig. ca. 1.3 X the 176 PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON VOLUME

length ( ately bl tremely a small subrect,1 ca.2 X pIe, slig ly curvl absent. Fema te!lescop length ( physes broad; a ite (lam! mediatel vaginali: extendin broad, I elongate terior el corpus enlargec spinose Etym. rived fr combine plUi. ft .

P.

Male 10.1 ml buff to buff. AI paler ve ered wi t:: ked veIl of short sensilla pus cre; cream tc Figs. 7-12. ProselOmorpha j'afcalo, adult. 7 , Sensilla coeloconicum shown in Fig. 6 (5 fLm) , 8, Ventral [J irrom sensilla of antenna showing details of sensi lla chaetica (aITow) and sensilla auricillica (asterisk) (IS fLm). 9, brown, Male retinac ulum, ventral view, (120 fLm). 10, Distal view o f Fig. 9 (l00 fLm ), II, Foretibia with epiphysis and mostly apical spines (176 fLm ). 12, Detail of epiphysis s howing flat spines (comb; 6 fLm). (Scale lengths in parentheses; bar scale for all photographs = Fig. 7.) Forewitl Iy SpOll. cous SC Y OF WASHINGTON VOLUME 98, NUMBER 2 177

length of genital capsule; sacculus moder­ light to dark brown scales with suffusion of ately broad, gradually narrowing to ex­ cream around torn us. Hindwing uniformly tremely slender cucullus (Fig. 22), bealing fuscous except for streak of white along a small recurved spine at apex. Juxta a flat, costa, broadest at wing base and narrowing subrectangular, elongate plate, with length toward apex; fringe generally lighter in col­ ca. 2X width. Aedoeagus a relatively sim­ or, gray along hind margin becoming more ple, slightly depressed, elongate, and slight­ white toward apex with streaks of fuscous. ly curved (viewed laterally) tube; cornuti Legs mostly dark fuscous dorsally, pale absent. buff ventrally, with buff banding on tibia Female genitalia: Ovipositor elongate, and more faint at apices of tibia and tarso­ telescoping. Posterior apophyses 1.7X the meres. Hindleg much paler, lighter brown. length of anterior pair. Ventral pseudapo­ Abdomen: Dark fuscous dorsally and lat­ physes absent. Ostium bursae indistinct, erally, cream ventraHy with long, cream, broad; a small, sclerotized, triangular scler­ piliform scales over AS largely covering ite (lamella postvaginalis; Fig. 24, LP) im­ genitalia. mediately caudad to ostium; lamella post­ Female (Fig. 2).-Length of forewing vaginalis with a slender keel-like process 10.5-13.0 mm. Generally paler than male. extending anteriorly; ductus bursae mostly Head: Light brown to white with light broad, membranous except for a dense, brown scaling laterally on labial palpus. elongate patch of minute spinules from an­ Forewing light brown with fuscous spots terior end that extends to signum within more evident. Hindwing gray with whitish corpus bursae; corpus bursae moderately costal streak as in male. Legs with similar enlarged (Fig. 24), with a single densely markings to male but generally lighter in spinose signum (Fig. 25). color. Etymology.-The generic name is de­ Male genitalia (Figs. 20-23).-As de­ rived from the Greek prefix pro (before) scribed for genus. combined with the generic name Setomor­ Female genitalia (Figs. 24, 25).-As de­ pha. It is considered feminine in gender. scribed for genus. Larva (Figs. 30-42, 49-58).- Length of largest larva 20 mm, maximum diameter Prosetomorpha falcata Davis, 2.9 mm. Body generally white with distinct NEW SPECIES dark brown plates and pinacula (Figs. 49, Male (Fig. l).-Length of forewing 8.5­ 50). 10.1 mm. Head (Figs. 3,4, 13, 14): Vertex Head: Maximum width 2.7 mm. Color buff to dark brown; frons paler, cream to reddish brown, becoming darker over frons buff. Antenna with scape mostly fuscous, and along anterior margins. Chaetotaxy as paler ventrally; Aagellum light brown, cov­ illustrated (Figs. 30-32, 34, 52, 53, 55); AF ered with narrow scales dorsally, mostly na­ I absent. Stemmata (Figs. 35, 55) consist­ ked ventrally except for dense pubescence ing of 6 pairs, with 3-5 contiguous near of short sensi lla chaetica and dense, slender lateral rim of antennal socket, and 6 more sensilla auricillica (Fig. 8). Maxillary pal­ remote and ventral. Mandible unidentate pus cream to white. Labial pal pus mostly (Fig. 56). Spinneret (Fig. 33) slender and cream to white with external sides of I and elongate with a minute orifice. Labial pal­ g. 6 (5 fLm) . 8, Ventral II irrorated with fuscous. Thorax: Dorsum pus 2-segmented, slender and elongate, ca. a (asterisk) (15 fLm). 9, brown, irrorated with fuscous; tegulae O.S the length of spinneret; apical segment :tibia with epiphysis and mostly fuscous. Venter pale buff to cream. less than l!J the length of basal segment; api­ c lengths in parentheses; Forewing variably light to dark brown, fine­ cal seta 1.5 X the length of second segment. ly spotted with small clusters of dark fus­ Apex of mentum with a pair of minute sec­ cous scales, fringe similarly mottled with ondary labial setae (Fig. 33). 178 PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON VOLUME 9R .

I~~ ~ / I I ( I I I I I I

I t I I,'I I ~ 1\ 1\1 I'\)! \/\' ~ ' ~I "

~!/ 20

19

y I 26 18 1 T 1

Figs. 20-2. Mesal view 0­ Figs. 13-19. ProselOmorpha falcara, adult. ]3, Head, anterior view; mandible (see arrow; 0.5 mm). 14, gi nal is (0.5 111 Detail of maxilla (0.2 mm). 15, Wing venation. 16, Metafurcasternum, lateral view (0.5 mm). 17 , Caudal view vicw. (Scale I of Fig. 16. 18, Leg struc ture (I mm). 19, Ventral view of maJe abdominal segments 7-8; note complete, scJerotized ring on A8 (0.5 mm). (Scale lengths in parentheses.) VOLUME 98, N MBER 2 179 'y OF WASHINGTON

14

1 1

23 21 22

25 24

19

T

I 26 1 27 1 ) 1 . ~

CJ

Figs. 20- 27. Proselomorpha fa/cola. adult. 20. Male genitalia, ventral view (0.5 mm). 21, Lateral view. 22, Mesal view of valva. 23, Ventral view of aedoeagus. 24, Female genitalia, ventral view; LP = lamella postva­ sec arrow; 0.5 mm). 14, ginalis (OS mm). 25, Enlarged detail of signum shown in Fig. 24. 26, Pupa, dorsal view (2 mm). 27, Ventral Ij 111m). 17. Caudal view view. (Scale lengths in parentheses.) WIS 7- 8; note complete, 180 PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON VOLUME 98.

Thorax: dark reddist and togetheJ crescent sha diately ante! and metatho pinacula fro of similar lei sal. claw mo enlarged bas arated, beari Abdomell: segments A. SV trisetose gether on s; plate with 5 seta XDI (F on A3-6 anc uniseria'l, anc of approxim band of smal apex of plar consisting of 20-22 hook~ (Fig. 42). Pupa (Fi§ Length 9-1 Color usuall ing darker .. for a minu spine projeci head (Fig. 4 long. LabrUi (Fig. 59). Fo dal margin c to middle of row of spine spines; tabu) anterior = ::; (Fig. 60), a 67-75; AS, 70-77; A6, 63-71; A7, Figs. 28-33. Prosetomorphafalca ta. 28-29, Adult: 28. Pretarsus of hindleg, ventral view (38 J.l.m). 29, Detail 24-28; A8 = of unguitractor plate in Fig. 28 (12 J.l.m). 30-33, Last instar larva: 30, Head, dorsal view (0.43 mm). 31, Head. circled by 8 ventral view (0.43 mm). 32, Head. anterior view (0.43 mm). 33. ventral view of maxilla and labium (100 J.l.m). 61-64) as f (Scale lengths in parentheses; bar scale for all photographs = Fig. 28.) pairs bearinl! separated fllJ Holotype.­ 'y OF WASHINGTON VOLUME 98, NUMBER 2 181

Thorax: Pronotum and spiracular plates l560 m, Cundinamarica Prov.; Oct 1988, I. dark reddish brown. Lateral setae trisetose Zenner, ex stem of Solanum quitoensis [var. and together with spiracle on same plate; a septentrionalis R. E. Schultes], USNM. crescent shaped, hyaline separation imme­ Paratypes.-Same data as holotype: 6 0", diately anterior to spiracle (Fig. 51). Meso­ 9

Figs. 34- 39. ProselOmorpha faicala, last instar larva. 34, Head, lateral view (0.46 mm). 35, Detail of Figs. 40­ stemmatal area in Fig. 34 (l50 fJ-m) . 36, Maxilla (43 fJ-m). 37, Detail of sensilla at apex of maxillary pal pus Caudal vic\1iiI (8.6 fJ-m). 38, Pretarsus of metathoracic leg (43 fJ-m). 39, Right proleg, A4 (arrow = anterior; 158 fJ-m) . (Scale mill). 44. L lengths in parentheses; bar scale for all photographs = Fig. 34.) 75 f.1m). (Sc ( OF WASHI NGTON VOL ME 98, UMBER 2 183

J46 mm). 35, Detail of Figs, 40- 45. Proselom.orph.a fa/caw. 40-42, last instar larva : 40, Late ral view of A9-1 0 (0,5 mm). 41 , ~pcx ()f maxillary pal pu s Caud al view of A J0 (0.43 mm). 42, Right proJeg of A 10 (150 ""m). 43-45, Pupa: 43, Head, ventral view (0.6 anterior: 158 f1m). (Scale mm). 44, L a tl~ ra l view of hg. 43 (0.6 mm). 45, Detail of frontal spine (with broken tip) in Fi g . 44 (see arrow; 75 ""m ). (Scale le ngths in parentheses; bar scale for a ll photographs = Fig. 40.)

r 184 PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON VOL U ~ "Ii;'01 -I~ -

U 51

r~~ I I ! I k-L 52 "­'-.

cc 55

Figs. 5 abdomina 10, dorsa mm). (Sp

plant dE sion of : hole" i never u the last pupates

cocoon~ 6 mm i inside t Figs. 46--50. ProselOm.orph.a falcaw. 46, Solanum quilaensis da maged by stem-boring larvae of weevil, FolJowi Faustina apicalis, and tineid, Proselomorph.a falcala. Arrow points to inrested site indicated by swollen stem. vium c 47, Closeup o f swo lle n stem with protruding frass of Proselomorpha (arrows). 48, Cocoon (23 mm). 49, Last in star larva, lateral view (20 mm). 50, Dorsal view. (Specimen lengths in parentheses.) hole. Becal ~TY OF WASH[NGTON VOLUME 98, NUMBER 2 185

\ XDl --...... 0 1 --.---. . I ..... '-. I I D2/ ~.../ ~ . 0 r~-, J D2 r:" SD1 ::::'-!" ~~l ~ ~~) p i jzIT'- ---/-'U ~

51 T-1 T-2 A-1 A-2 A-6 A-7 A-6 A-9

5 3 48 AI A3

e ~:: I \ o . , ./S2~ . ~ ~ rz V ...... V " 't1 58 ss ~~SS I 1~~rH-r-Jt--Y\ 55 r~ 56 , 57

Figs. 51-58. Prosetomorpha!alcata, morphology of last instar larva. 51, Chaetotaxy of pro- and mesothorax, abdominal segments 1,2,6-[0.52, Head, dorsal view (0.5 mm). 53, Ventral view. 54, Abdominal segments 8­ 10, dorsal vjew. 55, Head, lateral view. 56, Mandible (0.3 mm). 57, Labrum, dorsal view. 58, ventral view (0,3 mm). (Specimen lengths in parentheses.)

plant damage can be detected by the extru­ their concentrated feeding often weakens sion of silk-entwined frass from a "cleaning the stems sufficiently to facilitate breakage hole" in the stem (Fig. 47). This hole is on windy days. In this manner, the larval never used as an exit by the moth; instead, feeding of P. falcata can be instrumental in the last instar larva chews another hole and the death of damage stems and, at times, pupates within a dark silken, frass covered even the entire plant. cocoon, measuring up to 23 mm long and The full complement of six pairs of stem­ 6 mm in diameter (Fig. 48), that is formed mata easily distinguishes larvae of P. fal­ inside the stem near the future exit hole. cata from that of Setomorpha and Lindera lem-boring larvae of weevil, Following adult emergence, the pupal exu­ (both without stemmata). The chaetotaxy of Ie indicated by swollen stem. vium can be found protuding from this aJi three genera is very similar, including g, Cocoon (23 mm). 49, Last hole. the shared presence of five pairs of setae on cscs.) Because the larvae tend to be gregarious, the anal plate (Fig. 54). Prosetomorpha dif­ 186 PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON VOLUM 98.

fe rs in havi and in POSS( on AI O (Fig

Most of a Zenner-PolLIi Agropccuari, lecling the s is based, as

biological 11( debtcd to Yo nton1ology and EI is:1bctl ings. Susanr SEM lab an sonian Phot( photographic (retired), Sy tory, USDA,

Figs. 59-64. ProselOmorpha fa/COla, pupa. 59, Frontal region of head, ventral view (2 50 j..l.m). 60, Double rows of dorsal spines on abdominal segment 5 and portions of A4 and 6 (0. 3 mm). 61 , Dorsal view of A9- JO (0.38 mm). 62, Ventral view of A9 - 10 (0.38 mm). 63, Lateral view of A9-10 (0.38 mm). 64, Caudal view of AlO. (Scale lengths in pare ntheses; bar scale for all photographs = Fig. 59.) ( OF WASHINGTON VOLUME 98. NUMBER 2 187

rers in having lost the first adrronlal seta wi sh to thank Gaden Robinson of The Nat­ and in possessing an additional seta (PP I ) ural History Museum and two anonymous on A I U (Fig. 5 I ). reviewers for their comments on the manu­ scri pt. ACKNOWLEDG MENTS LITERATURE [TED Most of all I wi sh to thank Dr. Jngeborg Da vi s, D . R. and 1. E. Pena. 1990. Biology and mor­ Zenner-Polania or the Instituto Colombiana phology of the banana moth, OpogonG sacchari Agro pecuario. Bogota. Colombia fo r col­ (Boje rl. and its introduction into Fl ori da (Lepi­ do ptera: T incidac). P roceed ings of the E ntomolog­ lecting th e specimens on which this report ical Soc iety of Washington 92(4): 593-618. is based. a wel l as for providing pertinent G ozmany, L. A . and L. Vari. 1973. The Tineidae of biological notes and photographs. I am in­ the Ethopian Region. Memoir Transvaal Museum debted to Young Sohn of the Department of No. 18: i-vi i 1- 238. Entomology and volunteers Rachel Collins Hinto n, H. E . 1956. The larvae of the species o f T i­ ncidae o f economic impo rtance. Bulletin o f En­ and Elisabeth Ni c holson for th e line draw­ tomological Research 47: 251-346. ings. Susann Braden of the Smithsonian Robinson, G . S. and E. S. Nielsen. 1993. Tineid gen­ SEM lab and Vi ctor Kranz of the Smith­ era of Australia. Monographs o n Australian Lep­ sonian Photographic Laboratory provided idoptera 2: i-xvi + 1-344. photographic assistance. and Paul Marsh Zimmerman, E . C. 1978. M icrolepidoptera. Part I. Mo notrys ia, Tineoidea, Gracillarioidea, Ypono­ (retired), Systematic Entomology Labora­ mcutoidea, and Aluc ito idea. o f Hawai i 9: tory, USDA. identified the parasitoid. I also i- xvi ii , 1-88 1, pi s. 1- 8.

-- .-, ., -, ---­ ' 62

I (250 Ilm ). 60, Double Dorsal view of A9-10 m). 64, Caudal view of