189 Genus Cephetola Libert

AFROTROPICAL BUTTERFLIES. MARK C. WILLIAMS. http://www.lepsocafrica.org/?p=publications&s=atb Updated 27 June 2021.

Genus Cephetola Libert, 1999 Dark Flashes

In: Libert, 1999. Révision des Epitola (l.s.): 141 (219 pp.). Type-species: Epitola cephena Hewitson, 1873, by original designation.

The genus Cephetola belongs to the Family Lycaenidae Leach, 1815; Subfamily Poritiinae Doherty, 1886; Tribe Epitolini Jackson, 1962. The other genera in the Tribe Epitolini in the Afrotropical Region are Iridana, Teratoneura, Epitola, Cerautola, Hewitola, Geritola, Stempfferia, Deloneura, Batelusia, Tumerepedes, Pseudoneaveia, Neaveia, Epitolina, Hypophytala, Phytala, Neoepitola, Aethiopana, Hewitsonia, and Powellana.

Cephetola (Dark Flashes) is a purely Afrotropical genus containing 69 species. Many, if not all, species are associated with arboreal Crematogaster ants, imagines often being found near ant-infested trees (Safian, 2015). The larvae of a few species have been found in the proximity of Crematogaster ants and a larva of C. collinsi has been observed entering the carton nest of the ants, indicating a strong myrmecophilous relationship (Safian, 2015). Males of several species may be observed displaying on hilltops at fixed times of the day (Libert, 1993, Congdon & Collins, 1998, Safian, 2015). Females are mostly seen flying around ant-trees or larger bushes, which are also visited by ants. Oviposition on branches and twigs of ant-trees by a few species has been noted (Safian, 2015).

Note: Libert (1999) and D’Abrera (2009: 682) list Cephetola lamborni (Bethune-Baker, 1922) [Transactions of the Entomological Society of London 54: 461] as a nomen dubium.

*Cephetola cephena (Hewitson, 1873) Cephena Epitola

Epitola cephena Hewitson, 1873. Entomologist’s Monthly Magazine 10: 151 (149-151). Roche (1954) describes the male but this is actually a redescription of Cephetola mercedes (Suffert, 1904) (Jackson, 1962). Cephetola cephena (Hewitson, 1873). Libert, 1999 comb. nov. Cephetola cephena Hewitson, 1873. d’Abrera, 2009: 680. Cephetola cephena (Hewitson, 1873). Libert, 2020: 83.

Type locality: Gabon: Holotype (female): Gabon (Rogers); in NHM, London. Distribution: Nigeria, Cameroon, Gabon. Specific localities: Nigeria – Uzuakoli (Libert, 2020); Eket (Libert, 2020). Cameroon – Ebogo (Libert, 2020). Habitat: Forest. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola daveyi Libert, 2020

Cephetola daveyi Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 84 (157 pp.; 15 plates). Published by the author.

Type locality: Cameroon: Holotype (female): Lolodorf, VII 2013 (ABRI Leg.); BOLD: MLIB-1495; in 1 ABRI Collection, Nairobi. Allotype (male): Mt Mille, Lolodorf, III 2012 (ABRI Leg.); genitalia Libert 114-102; BOLD: MLIB-1482; in ABRI Collection, Nairobi. Distribution: Cameroon. Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

Cephetola daveyi daveyi Libert, 2020

Cephetola daveyi Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 84 (157 pp.; 15 plates). Published by the author.

Type locality: Cameroon: Holotype (female): Lolodorf, VII 2013 (ABRI Leg.); BOLD: MLIB-1495; in ABRI Collection, Nairobi. Allotype (male): Mt Mille, Lolodorf, III 2012 (ABRI Leg.); genitalia Libert 114-102; BOLD: MLIB-1482; in ABRI Collection, Nairobi. Distribution: Cameroon. Specific localities: Ghana – Lolodorf (TL); Mount Mille (Libert, 2020).

Cephetola daveyi occidentalis Libert, 2020

Cephetola daveyi occidentalis Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 85 (157 pp.; 15 plates). Published by the author.

Type locality: Ghana: Holotype (male): Mate, II 2011 (ABRI Leg.); BOLD: MLIB-1785; in ABRI Collection, Nairobi. Allotype (female): Mate, Volta, IX 2011 (ABRI Leg.); BOLD: MLIB-0923; in ABRI Collection, Nairobi. Distribution: Ghana, Togo, ?Benin, Nigeria. Specific localities: Ghana – Mate (TL); Agona (Libert, 2020); Apemedi (Libert, 2020); Bonkro (Libert, 2020); Tano Ofin (Libert, 2020); Sanko (Libert, 2020); Amedzofe (Libert, 2020); Hohoe (Libert, 2020); Biakpa- Avatime (Safian et al., 2012); Kakum (Safian et al., 2012); Bobiri (Safian et al., 2012). Togo – Kloto (Libert, 2020); Kpalime (Libert, 2020) Nigeria – Gambari (Libert, 2020); Vom (Libert, 2020); Jos Plateau (Libert, 2020).

*Cephetola doleta (Kirby, 1890)

Epitola doleta Kirby, 1890. Annals and Magazine of Natural History (6) 6: 273 (261-274). Synonym of C. cephena (Hewitson, 1873). Libert, 1999. Cephetola doleta (Kirby, 1890). Libert, 2020: 86, stat. rev.

Type locality: Sierra Leone: “Sierra Leone”. Lectotype (male) designated by Libert (2020: 86): Sierra Leone (Preuss); Museum für Naturkunde, Berlin. Distribution: Guinea, Sierra Leone, Liberia, Ivory Coast. Specific localities: Guinea – Fouta Djalon (Libert, 2020); Labe (Libert, 2020). Sierra Leone – Gola Forest (Belcastro & Larsen, 2006: 34); Mabang (Libert, 2020); Moyamba (Libert, 2020). Liberia – Nimba Mountains (Safian, 2014: 53). Ivory Coast – Banco (Libert, 2020); Bayota (Libert, 2020); Mt Peko (Libert, 2020); Mt Sangbe (Libert, 2020); Tiassale (Libert, 2020); Abidjan (Libert, 2020). Habitat: Nothing published. 2 Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

pinodoides Grose-Smith & Kirby, 1893. In Grose-Smith & Kirby, 1892-7 (as sp. of Epitola). Rhopalocera exotica, being illustrations of new, rare and unfigured species of butterflies 2: 85 (261 pp.). London. Sierra Leone: “Mocquerys”. Given as a good species of Epitola by Ackery et al., 1995: 524. Synonymized with Cephetola cephena by Libert, 1999.

leonina Bethune-Baker, 1903 (as sp. of Phytala). Annals and Magazine of Natural History (7) 12: 328 (324-334). Sierra Leone: “Sierra Leone”.

leonensis Bethune-Baker, 1904 (as sp. of Epitola). Annals and Magazine of Natural History (7) 14: 227 (222-233). Sierra Leone: “Sierra Leone”.

*Cephetola entebbeana (Bethune Baker, 1926)

Epitola entebbeana Bethune-Baker, 1926. Annals and Magazine of Natural History (9) 17: 392 (384-402). Cephetola cephena entebbeana (Bethune-Baker, 1926). Libert, 1999 comb. nov. Cephetola cephena entebbeana Bethune-Baker, 1926. d’Abrera, 2009: 680. Cephetola entebbeana (Bethune-Baker, 1926). Libert, 2020: 88, stat. rev.

Cephetola cephena entebbeana. Male. Left – upperside; right – underside. Radio Hill, Mabira Forest, Uganda. 14 June, 2009. J. & C. Dobson. Images M.C.Williams ex Dobson Collection.

Type locality: [Uganda]: “Entebbe”. Distribution: Uganda, Tanzania (north-west). Specific localities: Uganda – Entebbe (TL); Bwamba (Libert, 2020); Mbale (Libert, 2020); Buddo (Libert, 2020); Mpanga Mpigi (Libert, 2020); Radio Hill, Mabira Forest (male illustrated above). Tanzania – Kere Hill, Minziro Forest (common) (Congdon & Collins, 1998). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola cephenoides Libert, 2020

Cephetola cephenoides Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 89 (157 pp.; 15 plates). Published by the author.

Type locality: Democratic Republic of Congo: Holotype (male): Mapimbi, II 2014 (ABRI Leg.); BOLD: MLIB-1806; in ABRI Collection, Nairobi. Allotype (female): Maliva, N. Kivu, I 2015 (ABRI Leg.); BOLD: MLIB-2019; in ABRI Collection, Nairobi. 3 Distribution: Democratic Republic of Congo (north-east), Uganda. Specific localities: Democratic Republic of Congo – Mapimbi (TL); Maliva (Libert, 2020); Mbau (Libert, 2020); Mt Mitumba (Libert, 2020); Kasugho (Libert, 2020). Uganda – Bwamba (Libert, 2020). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola maculata (Hawker-Smith, 1926) Macular Epitola

Epitola maculata Hawker-Smith, 1926. Revue Zoologique Africaine 14: 240 (237-241). Cephetola maculata (Hawker-Smith, 1926). Libert, 1999 comb. nov. Cephetola maculata Hawker-Smith, 1926. d’Abrera, 2009: 684. Cephetola maculata (Hawker-Smith), 1926. Libert, 2020: 92.

Type locality: [Democratic Republic of Congo]: Holotype (male): Elizabetha, near Basoko, (Mme Tinant); MRAC, Tervuren, Belgium. Distribution: Ivory Coast, Cameroon, Congo, Central African Republic, Democratic Republic of Congo, Uganda. Recorded, in error, from Ivory Coast by Larsen (2005a) and Congo (Jackson, 1964). Specific localities: Cameroon – Mekas (Libert, 2020); Dja (Libert, 2020) Democratic Republic of Congo – Elisabehta, near Basoko (TL); Ituri Forest (Ducarme, 2018); Mt Mitumba (Ducarme, 2018); Biakatu (Libert, 2020); Epulu-RFO (Libert, 2020); Kasugho (Libert, 2020); Kasuo (Libert, 2020); Kenge (Libert, 2020); Kithokolo (Libert, 2020); Maliva (Libert, 2020); Mambungu (Libert, 2020); Vhovho (Libert, 2020). Uganda – Bwamba (Libert, 2020). Habits: A rare species in West Africa, with only about a dozen specimens known from Ivory Coast (Larsen, 2005a). Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

Note: There is a possibility that the populations in Ivory Coast represent an undescribed taxon that is an Ivory Coast endemic, should it be genuinely absent from the countries between the Ivory Coast and Congo (Larsen, 2005a).

*Cephetola pulchra (Jackson, 1964)

Phytala pulchra Jackson, 1964. Bulletin of the British Museum (Natural History) (Entomology) 15: 65 (59-80). Cephetola pulchra (Jackson, 1964). Libert, 2020: 93, stat. rev.

Type locality: Congo: Holotype (male): Ketta, Ouesso, XII 1959 (T.H.E. Jackson); in NHM, London. Distribution: Cameroon, Congo, Central African Republic. Specific localities: Cameroon – Ebogo (Libert, 2020); Mintom (Libert, 2020). Congo – Ketta Forest, Ouesso (TL); Etoumbi (Libert, 2020). Central African Republic – Bayanga, Sangha N.P. (Libert, 2020); Botambi (Libert, 2020); Maka (Libert, 2020); Mbata (Libert, 2020); Mombo (Libert, 2020). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published.

4 Larval food: Nothing published.

*Cephetola cameruna Libert, 2020

Cephetola cameruna Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 94 (157 pp.; 15 plates). Published by the author.

Type locality: Cameroon: Holotype (male): West Koutaba, VIII 2013 (ABRI Leg.); BOLD: MLIB- 1465; in ABRI Collection, Nairobi. Allotype (female): West Koutaba, VIII 2013 (ABRI Leg.); BOLD: MLIB-1466; in ABRI Collection, Nairobi. Distribution: Cameroon. Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

Cephetola cameruna cameruna Libert, 2020

Cephetola cameruna Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 94 (157 pp.; 15 plates). Published by the author.

Type locality: Cameroon: Holotype (male): West Koutaba, VIII 2013 (ABRI Leg.); BOLD: MLIB- 1465; in ABRI Collection, Nairobi. Allotype (female): West Koutaba, VIII 2013 (ABRI Leg.); BOLD: MLIB-1466; in ABRI Collection, Nairobi. Distribution: Cameroon (west). Known only from the type locality. Specific localities: Cameroon – West Koutaba (TL).

Cephetola cameruna reducta Libert, 2020

Cephetola cameruna reducta Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 95 (157 pp.; 15 plates). Published by the author.

Type locality: Cameroon: Holotype (male): Maan, XI 2002 (ABRI Leg.); in ABRI Collection, Nairobi. Allotype (female): Maan, VIII 2014 (ABRI Leg.); BOLD: MLIB-1794; in ABRI Collection, Nairobi. Distribution: Cameroon (south). Specific localities: Cameroon – Maan (TL); Dja (Libert, 2020).

*Cephetola belcastroi Libert, 2020

Cephetola belcastroi Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 95 (157 pp.; 15 plates). Published by the author.

Type locality: Ivory Coast: Holotype (male): Tiassale, Abidjan, X 1965 (T.H.E. Jackson); in National Museums of Kenya, Nairobi. Distribution: Ivory Coast. Specific localities: Ivory Coast – Tiassale (TL). Habitat: Nothing published. Habits: Nothing published. 5 Early stages: Nothing published. Larval food: Nothing published.

*Cephetola mariae Libert, 1999

Cephetola mariae Libert, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 148 (219 pp.). Cephetola mariae Libert, 1999. d’Abrera, 2009: 682. Cephetola mariae Libert, 1999. Libert, 2020: 96.

Type locality: Tanzania: “Kere Hill, Minziru for., 1250 m, Bukoba, Tanzanie”. Distribution: Cameroon, Central African Republic, Democratic Republic of Congo (north-east), Kenya, Tanzania (Libert, 1999). Habitat: Forest. Specific localities: Cameroon – Koutaba (Libert, 2020). Democratic Republic of Congo – Kasugho (Libert, 2020); Mamove (Libert, 2020). Kenya – Kakamega Forest (Libert, 2020). Tanzania – Kere Hill, Minziro Forest (TL). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola gerdae (Kielland & Libert, 1998)

Epitola gerdae Kielland & Libert, 1998. In: Congdon & Collins, 1998. Supplement to Kielland’s butterflies of Tanzania : 117 (143 pp.). ABRI and Lambillionea. Cephetola gerdae (Kielland & Libert, 1998). Libert, 1999 comb. nov. Cephetola gerdae Kielland & Libert, 1998. d’Abrera, 2009: 684.

Type locality: Tanzania: “Minziro, Bukoba dist., Tanzanie, 26 I 1993 (I. Kielland)”. Holotype in ABRI Collection, Nairobi. Distribution: Tanzania (Libert, 1999). Specific localities: Tanzania – Kere Hill, Minziro Forest (uncommon) (TL; Congdon and Collins, 1998). Known only from the type locality. Habitat: Forest. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola viridana (Joicey & Talbot, 1921) Vivid Epitola

Males of Cephetola viridana. Kibale Forest, Uganda. October 2014. Images courtesy Raimund Schutte.

Epitola viridana Joicey & Talbot, 1921. Bulletin of the Hill Museum, Witley 1: 84 (40-166). Cephetola viridana (Joicey & Talbot, 1921). Libert, 1999 comb. nov. Cephetola viridana Joicey & Talbot, 1921. d’Abrera, 2009: 680. Cephetola viridana (Joicey & Talbot, 1921). Libert, 2020: 96.

Cephetola viridana. Male. Left – upperside; right – underside. Mabira Forest, Uganda. 8 May 2011. P. Ward. Images M. C. Williams ex P. Ward collection.

Cephetola viridana. Female. Left – upperside; right – underside. Mabira Forest, Uganda. 5 June 2011. P. Ward. Images M. C. Williams ex P. Ward collection.

Type locality: [Democratic Republic of Congo]: “South side Ituri River, five days west of Irumu”. Distribution: Cameroon, Congo, Central African Republic, Democratic Republic of Congo, Uganda, Kenya (west), Tanzania (north-west), Zambia (north-west) (Libert, 1999). Specific localities: Cameroon – Bitje (Bethune-Baker, 1926). Democratic Republic of Congo – Ituri River, five days west of Irumu (TL); Ituri Forest (Ducarme, 2018); Semuliki Valley (Ducarme, 2018); Mt Mitumba (Ducarme, 2018). Uganda – Bwamba (Libert, 2020); Kibale Forest (R. Schutte, unpublished); Mabira Forest (male and female illustrated above).

7 Kenya – Kakamega Forest (Larsen, 1991c). Tanzania – Minziro Forest (Congdon & Collins, 1998); Kikuru Forest (Congdon & Collins, 1998); Munene Forest (very common in all) (Congdon & Collins, 1998). Zambia – Ikelenge (Heath et al., 2002); Chikonke Stream. Habitat: Primary forest. Habits: Nothing published. Early stages:

Congdon et al., 2017 [final instar larva].

Larval food: Cyanobacteria on trees [Congdon et al., 2017; Minziro, Tanzania].

radiata Bethune-Baker, 1926 (as subsp. of Epitola viridana). Annals and Magazine of Natural History (9) 17: 393 (384-402). Cameroon: “Bitje”. Given by Ackery et al., 1995: 525 as a good subspecies of Epitola viridana. Synonymized with Cephetola viridana (Joicey & Talbot, 1921) by Libert (1999: 150). Treated informally, and without explanation, as a subspecies of viridana by d’Abrera (2009: 680).

*Cephetola orientalis (Roche, 1954)

Epitola orientalis Roche, 1954. Bulletin of the British Museum (Natural History) (Entomology) 3: 499 (489-501). Cephetola orientalis (Roche, 1954). Libert, 1999 comb. nov. Cephetola orientalis Roche, 1954. d’Abrera, 2009: 680.

Cephetola orientalis. Male. Left – upperside; right – underside. Mabira Forest, Uganda. 16 January 2010. P. Ward. Images M. C. Williams ex P. Ward collection.

Type locality: Uganda: “Bwamba”. Distribution: Democratic Republic of Congo (north-east), Uganda, Tanzania (north-west). Recorded, in error, from Cameroon, Gabon and Kenya. Specific localities: Democratic Republic of Congo – Ituri Forest (Ducarme, 2018); Mt Mitumba (Ducarme, 2018). Uganda – Bwamba (TL). Tanzania – Kere Hill, Minziro Forest (common) (Congdon & Collins, 1998). Habitat: Primary forest. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola neorientalis Libert, 2020

Cephetola neorientalis Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 99 (157 pp.; 15 plates).

8 Published by the author.

Type locality: Cameroon: Holotype (male): Maan, VIII 2014 (ABRI Leg.); genitalia Libert 117-118; BOLD: MLIB-1842; in ABRI Collection, Nairobi. Distribution: Camerron (south-west). Specific localities: Cameroon – Maan (TL); Mt Febe (Libert, 2020); Ebogo (Libert, 2020); Mt Mille, Lolodorf (Libert, 2020). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola obscura (Hawker-Smith, 1933) Obscure Epitola

Epitola obscura Hawker-Smith, 1933. Stylops 2: 11 (1-12). Cephetola obscura (Hawker-Smith, 1933). Libert, 1999 comb. nov. Cephetola obscura Hawker-Smith, 1933. d’Abrera, 2009: 682.

Type locality: Nigeria: “Lagos”. Distribution: Sierra Leone, Liberia, Ivory Coast, Ghana, Nigeria (west). Specific localities: Liberia – Wologizi (Safian et al., 2020). Ghana – Kakum National Park (Larsen, 2005a); Atewa Range (Larsen, 2005a). Nigeria – Lagos (TL). Habitat: Forest. Habits: A scarce butterfly (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published.

*Cephetola obscuralis Libert, 2020

Cephetola obscuralis Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 99 (157 pp.; 15 plates). Published by the author.

Cephetola obscuralis. Male. Left – upperside; right – underside. Mabira Forest, Uganda. 11 January 2009. P. Ward. Images M. C. Williams ex P. Ward collection.

Type locality: Uganda: Holotype (male): Mabira Forest (00° 23,3'N, 33° 00,7'E), 9 VII 2011 (P.R.F. Ward); BOLD: MLIB-0891; in ABRI Collection, Nairobi. Distribution: Cameroon (undescribed ssp. (Libert, 2020: 101)), Democratic Republic of Congo, Uganda (east). 9 Specific localities: Cameroon – Biwon-Bane, near Ebolowa (Libert, 2020). Democratic Republic of Congo – Mt Mitumba (Libert, 2020). Uganda – Mabira (TL; male imaged above). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola praecox Sáfián, 2021

Cephetola praecox Sáfián, 2021. Zootaxa 4981 (3): 568 (554-576).

Type locality: Liberia: Holotype (male): Grand Gedeh County, Mount Jideh Ridge, Putu Range 18.XII.2012 Leg.: Sáfián, Sz., Tropek, R. Coordinates: 5°38’23.32»N, 8°11’7.60»W, altitude: 700-760 m asl. ANHRT unique ID number: ANHRTUK00056662. Gen. prep.: SAFI00384. Deposited in ANHRT. The female is unknown. Distribution: Liberia. Specific localities: Liberia – Mt Jideh Ridge, Putu Range (TL). Known only from the type locality. Habitat: Submontane forest (Safian et al., 2021). Habits: Males display early in the morning, flying rapidly up and down tree trunks, from three to ten metres above the ground (Safian et al., 2021). Early stages: Nothing published. Larval food: Nothing published.

*Cephetola eliasis (Kielland & Congdon, 1998)

Epitola eliasis Kielland & Congdon, 1998. In: Congdon & Collins, 1998. Supplement to Kielland’s butterflies of Tanzania : 116 (143 pp.). ABRI and Lambillionea. Cephetola eliasis (Kielland & Congdon, 1998). Libert, 1999 comb. nov. Cephetola eliasis Kielland & Congdon, 1998. d’Abrera, 2009: 684. Cephetola eliasis (Kielland & Congdon, 1998). Libert, 2020: 102.

Type locality: Tanzania: “Minziro (1200m), Bukoba dist., Tanzanie, 16 II 1991 (I. Kielland)”. In ABRI, Nairobi. Distribution: Democratic Republic of Congo (north-east), Tanzania (north-west). Specific localities: Tanzania – Kere Hill, Minziro Forest (TL; Congdon & Collins, 1998). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola angustata Libert & Collins, 1999

Cephetola eliasis angustata Libert & Collins, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 153 (219 pp.). Cephetola eliasis angustata Libert & Collins, 1999. d’Abrera, 2009: 684. Cephetola angustata Libert & Collins, 1999. Libert, 2020: 102, stat. rev.

Type locality: Cameroon: “Ebogo, Nyong, Cameroun”. Distribution: Cameroon, Central African Republic. Specific localities:

10 Cameroon – Ebogo, Nyong (TL). Central African Republic – Bayanga, Sagha N.P. (Libert, 2020); Ituri Forest (Ducarme, 2018); Mt Mitumba (Ducarme, 2018). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola ouesso (Jackson, 1962)

Epitola ouesso Jackson, 1962. Bulletin of the British Museum (Natural History) (Entomology) 12: 147 (125-163). Cephetola ouessa (Jackson, 1962). Libert, 1999 comb. nov. Cephetola ouesso Jackson, 1962. d’Abrera, 2009: 682.

Type locality: Congo: “Ouesso, Ketta Forest”. Distribution: Cameroon, Congo, Central African Republic (Libert, 1999). Specific localities: Congo – Ketta Forest, Ouesso (TL). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola barnsi Libert, 1999

Cephetola barnsi Libert, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 154 (219 pp.). Cephetola barnsi Libert, 1999. d’Abrera, 2009: 682.

Type locality: [Democratic Republic of Congo]: “Middle Low Valley, nr. Walikale, 3000-4000 ft, forest”. Distribution: Democratic Republic of Congo (Libert, 1999). Specific localities: Democratic Republic of Congo – Middle Low Valley, near Walikale (TL); Ituri Forest (Ducarme, 2018). Habitat: Forest. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola mercedes (Suffert, 1904)

Epitola mercedes Suffert, 1904. Deutsche Entomologische Zeitschrift, Iris 17: 53 (12-107). Cephetola mercedes (Suffert, 1904). Libert, 1999 comb. nov. Cephetola mercedes Suffert, 1904. d’Abrera, 2009: 680. Cephetola mercedes (Suffert, 1904). Libert, 2020: 102.

Type locality: Cameroon: “Johann Albrechthöhe” [Kumba]. Distribution: Ivory Coast, Ghana, Nigeria, Cameroon, Gabon, Central African Republic, Democratic Republic of Congo. Habitat: Forest. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

Cephetola mercedes mercedes (Suffert, 1904)

Type locality: Cameroon: “Johann Albrechthöhe” [Kumbo]. Distribution: Nigeria (south and Cross River loop), Cameroon (west), Gabon, Central African Republic, Democratic Republic of Congo (south-west). Specific localities: Nigeria – Lagos area (Larsen, 2005a). Cameroon – Kumbo (TL); Ndoupe (TL of dejeani); Mt Kupe (Libert, 2020). Gabon – Mondah (Libert, 2020). Central African Republic – Bayanga (Libert, 2020). Democratic Republic of Congo – Mbata Siala (Libert, 2020).

dejeani Libert, 1999 (as ssp. of Cephetola mercedes (Suffert, 1904). In: Libert, 1999. Révision des Epitola (l.s.): 157 (219 pp.). Type locality: Cameroon: “Ndoupé, ex larva, Cameroun”. Larsen (2005a) avers that this subspecies may be a distinct species. D’Abrera (2009: 680), however, suggests that it may be a synonym of Cephetola mercedes (Suffert, 1904). D’Abrera does not make any formal taxonomic changes and includes it in his book as a valid subspecies. Libert (2020: 102) synonymized dejeani with mercedes.

Cephetola mercedes ivoriensis (Jackson, 1967)

Epitola ivoriensis Jackson, 1967. Journal of Natural History 1: 333 (327-336). Epitola ivoiriensis Jackson, 1967. Ackery et al., 1995: 522. [misspelling of species name]. Cephetola mercedes ivoiriensis (Jackson, 1967). Libert, 1999 stat. rev., comb. nov. [misspelling of species name]. Cephetola mercedes ivoriensis (Jackson, 1967). Larsen, 2005a. Cephetola mercedes ivoirensis Jackson, 1967. d’Abrera, 2009: 680. [misspelling of species name; should be ivoriensis]. Cephetola mercedes ivoriensis (Jackson, 1967). Libert, 2020: 102.

Type locality: Ivory Coast: “Abidgan, Tiassalé”. Distribution: Ivory Coast, Ghana. Specific localities: Ivory Coast – Abidjan, Tiassale (TL).

Note: D’Abrera (2009: 680) suggests that Cephetola ivoriensis (Jackson, 1967) is a synonym of Cephetola mercedes (Suffert, 1904) but makes no taxonomic changes and includes it in his book.

*Cephetola quentini Bouyer & Libert, 1999

Cephetola quentini Bouyer & Libert, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 157 (219 pp.). Cephetola quentini Bouyer & Libert, 1999. d’Abrera, 2009: 680.

Type locality: Cameroon: “Pinda, Cameroun”. Distribution: Cameroon (Libert, 1999). Specific localities: Cameroon – Pinda (TL). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published.

12 Larval food: Nothing published.

*Cephetola catuna (Kirby, 1890)

Epitola catuna Kirby, 1890. Annals and Magazine of Natural History (6) 6: 273 (261-274). Cephetola catuna (Kirby, 1890). Libert, 1999 comb. nov. Cephetola catuna Kirby, 1890. d’Abrera, 2009: 680.

Type locality: Cameroon: “Cameroons”. Distribution: Cameroon, Gabon, Congo, Central African Republic, Gabon, Democratic Republic of Congo, Uganda (Libert, 1999). Habitat: Forest. Habits: Females oviposit on lichens on the bark of twigs and tree trunks; eggs sometimes laid on dead bare twigs. Ants in attendance (Van Someren, 1974). Early stages: Nothing published. Larval food: Nothing published.

Cephetola catuna catuna (Kirby, 1890)

Epitola catuna Kirby, 1890. Annals and Magazine of Natural History (6) 6: 273 (261-274). Cephetola catuna (Kirby, 1890). Libert, 1999 comb. nov. Cephetola catuna Kirby, 1890. d’Abrera, 2009: 680.

Type locality: Cameroon: “Cameroons”. Distribution: Cameroon, Gabon, Congo, Central African Republic, Gabon, Democratic Republic of Congo, Uganda (Libert, 1999). Specific localities: Cameroon – Barombi Station (Suffert, 1904). Gabon – Waka (Vande weghe, 2010); Ndougou (Vande weghe, 2010); Lope N.P. (zone SEGC and Mikongo) (Vande weghe, 2010). Democratic Republic of Congo – Ituri Forest (Ducarme, 2018); Semuliki Valley (Ducarme, 2018); Central Forest Block (Ducarme, 2018); Mt Blue (Ducarme, 2018).

mus Suffert, 1904 (as sp. of Epitola). Deutsche Entomologische Zeitschrift, Iris 17: 53 (12-107). Cameroon: “Barombi-Station”.

Cephetola catuna carpenteri (Bethune-Baker, 1922)

Epitola carpenteri Bethune-Baker, 1922. Transactions of the Entomological Society of London 54: 462 (461-464). Cephetola catuna carpenteri (Bethune-Baker, 1922). Libert, 1999 comb. nov. Cephetola catuna carpenteri Bethune-Baker, 1922. d’Abrera, 2009: 680.

Type locality: [Uganda]: “Damba Island, N.W. Victoria Nyanza, about 20 miles S.E. of Entebbe”. Distribution: Uganda (Libert, 1999). Specific localities: Uganda – Damba Island, about 30 km south-east of Entebbe (TL).

*Cephetola oubanguensis Libert & Collins, 1999

Cephetola oubanguensis Libert & Collins, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 160 (219 pp.). Cephetola oubanguensis Libert & Collins, 1999. d’Abrera, 2009: 680.

Type locality: Central African Republic: “Yakoli, R.C.A.”. Distribution: Central African Republic (Libert, 1999). Specific localities: Central African Republic – Yakoli (TL). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola epitolina Libert & Collins, 1999

Cephetola epitolina Libert & Collins, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 160 (219 pp.). Cephetola epitolina Libert & Collins, 1999. d’Abrera, 2009: 680. Cephetola epitolina Libert & Collins, 1999. Libert, 2020: 104.

Type locality: Central African Republic: “Yakoli, R.C.A.”. Distribution: Central African Republic, Democratic Republic of Congo (Libert, 1999). Specific localities: Central African Republic – Yakoli (TL). Democratic Republic of Congo – Ituri Forest (Ducarme, 2018). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola nigra (Bethune-Baker, 1903) Bethune-Baker’s Epitola

Epitola nigra Bethune-Baker, 1903. Annals and Magazine of Natural History (7) 12: 331 (324-334). Cephetola nigra (Bethune-Baker, 1903). Libert, 1999 comb. nov. Cephetola nigra Bethune-Baker, 1903. d’Abrera, 2009: 680.

Cephetola nigra. Female. Left – upperside; right – underside. Bobiri, Ghana. 20 May 2014. A. Gardiner. Images M. Williams ex Gardiner Collection.

Type locality: Sierra Leone: “Sierra Leone”. Distribution: Sierra Leone, Ivory Coast, Ghana (Larsen et al., 2007), Nigeria (Cross River loop), Cameroon, Democratic Republic of Congo, Uganda. Specific localities: Democratic Republic of Congo – Ituri Forest (Ducarme, 2018). Habitat: Forest.

14 Habits: A very scarce species (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published.

*Cephetola ghesquierei (Roche, 1954)

Epitola ghesquierei Roche, 1954. Bulletin of the British Museum (Natural History) (Entomology) 3: 498 (489-501). Cephetola ghesquierei (Roche, 1954). Libert, 1999 comb. nov. Cephetola ghesquierei Roche, 1954. d’Abrera, 2009: 680.

Type locality: [Democratic Republic of Congo]: “Eala”. Distribution: Cameroon, Gabon, Congo, Central African Republic, Democratic Republic of Congo (Libert, 1999). Specific localities: Gabon – Camp Nouna (Vande weghe, 2010). Democratic Republic of Congo – Eala (TL); Ituri Forest (Ducarme, 2018); Central Forest Block (Ducarme, 2018). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola nigeriae (Jackson, 1962) Nigerian Epitola

Epitola nigeriae Jackson, 1962. Bulletin of the British Museum (Natural History) (Entomology) 12: 145 (125-163). Synonym of Epitola ghesquierei Roche, 1954. Jackson. Cephetola nigeriae (Jackson, 1962). Libert, 1999 stat. rev., comb. nov. Cephetola nigeriae Jackson, 1962. d’Abrera, 2009: 680.

Type locality: Nigeria: “Ogoja Prov., Ikom. X.1955. T.H.E. Jackson.” Distribution: Nigeria (Cross River loop), Cameroon, Gabon, Congo. Specific localities: Nigeria – Ikom, Ogaja Province (TL). Habitat: Forest. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola kamengensis (Jackson, 1962)

Epitola kamengensis Jackson, 1962. Bulletin of the British Museum (Natural History) (Entomology) 12: 150 (125-163). Cephetola kamengensis (Jackson, 1962). Libert, 1999 comb. nov. Cephetola kamengensis Jackson, 1962. d’Abrera, 2009: 682. Cephetola kamengensis (Jackson, 1962). Libert, 2000: 105, stat. rev.

Type locality: Uganda: “Mawakota, Kamengo”. Distribution: Cameroon, Central African Republic, Democratic Republic of Congo, Uganda, Kenya, Tanzania (Libert, 2020). Specific localities: Uganda – Mawakota, Kamengo (TL). Kenya – Nandi (Larsen, 1991c); Kakamega Forest (Congdon & Collins, 1998). Tanzania – Kere Hill, Minziro Forest (single record) (Congdon & Collins, 1998); Munene Forest (single record) (Congdon & Collins, 1998). 15 Habitat: On forest edges and in gallery forest, from 900 to 1 200 metres (Kielland, 1990d). Habits: Males spend considerable periods whirling around tall bushes, often several specimens together. Active from before sunrise (Kielland, 1990d). Females oviposit on lichens on the bark of twigs and tree trunks; eggs sometimes laid on dead bare twigs. Ants in attendance (Van Someren, 1974). Early stages: Nothing published. Larval food: Nothing published.

Cephetola kamengensis kamengensis (Jackson, 1962)

Type locality: Uganda: “Mawakota, Kamengo”. Distribution: Uganda, Kenya (west), Tanzania (north-west) (Libert, 2020). Specific localities: Uganda – Mawakota, Kamengo (TL). Kenya – Nandi (Larsen, 1991c); Kakamega Forest (Congdon & Collins, 1998). Tanzania – Kere Hill, Minziro Forest (single record) (Congdon & Collins, 1998); Munene Forest (single record) (Congdon & Collins, 1998).

Cephetola kamengensis subgriseata (Jackson, 1964)

Epitola subgriseata Jackson, 1964. Bulletin of the British Museum (Natural History) (Entomology) 15: 72 (59-80). Cephetola subgriseata (Jackson, 1964). Libert, 1999 comb. nov. Cephetola subgriseata Jackson, 1964. d’Abrera, 2009: 684. Cephetola kamengensis subgriaeata (Jackson, 1964). Libert, 2020: 106, stat. rev.

Type locality: Uganda: “Bwamba, Mongiro”. Distribution: Cameroon, Central African Republic, Democratic Republic of Congo, Uganda, Tanzania (Libert, 2020). Specific localities: Cameroon – Koutaba (Libert, 2020). Democratic Republic of Congo – Ituri Forest (Ducarme, 2018); Kinshasa (Libert, 2020). Uganda – Mongiro Forest, Bwamba (TL). Tanzania – Kigoma; Sitebi Mountain in Mpanda (single female) (Congdon & Collins, 1998).

*Cephetola martini (Libert, 1998)

Epitola martini Libert, 1998. In: Congdon & Collins, 1998. Supplement to Kielland’s butterflies of Tanzania : 110 (143 pp.). ABRI and Lambillionea. Cephetola martini (Libert, 1998). Libert, 1999 comb. nov. Cephetola martini Libert, 1998. d’Abrera, 2009: 682.

Cephetola martini. Male. Left – upperside; right – underside. Mabira Forest, Uganda. 11 July 2010. P. Ward. Images M. C. Williams ex P. Ward collection.

Cephetola martini. Female. Left – upperside; right – underside. Mabira Forest, Uganda. 10 March 2011. P. Ward. Images M. C. Williams ex P. Ward collection.

Type locality: Uganda: “Katera, Sango Bay, Masaka, Ouganda, V 1956 (T.H.E. Jackson)”. N.H.M., London. Distribution: Uganda, Tanzania (north-west) (Libert, 1999). Recorded, in error, from Cameroon (Bouyer, vide Congdon & Collins, 1998). Specific localities: Uganda – Katera, Sango Bay, Masaka (TL); Damba Island (Bethune-Baker, 1922); Mabira (male and female illustrated above). Tanzania – Minziro Forest (Congdon & Collins, 1998); Kigoma (Congdon & Collins, 1998). Habitat: Forest. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

carpenteri Bethune-Baker, 1922 (as sp. of Epitola). Transactions of the Entomological Society of London 54: 462 (461-464). Uganda: “Damba Island, N.W. Victoria Nyanza, about 20 miles S.E. of Entebbe”. The female is given as a synonym of Epitola azurea Jackson and the male as Epitola catuna carpenteri Bethune-Baker by Ackery et al., 1995: 520 and 521, respectively.

azurea Jackson, 1962 (as sp. of Epitola). Bulletin of the British Museum (Natural History) (Entomology) 12: 148 (125-163). Uganda: “Masaka, Sango Bay, Katera”. Given as a good species of Epitola by Ackery et al., 1995: 520 but regarded as a synonym of Epitola martini by Libert [In: Congdon & Collins, 1998. Supplement to Kielland’s butterflies of Tanzania: 110 (143 pp.). ABRI and Lambillionea)].

*Cephetola maani Libert, 2020

Cephetola maani Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 107 (157 pp.; 15 plates). Published by the author.

Type locality: Cameroon: Holotype (male): Maan, XI 2000 (ABRI Leg.); in ABRI Collection, Nairobi. Distribution: Cameroon (south). Allotype (female): Maan, VII 1999 (S.C. Collins); in ABRI Collection, Nairobi. Distribution: Cameroon (south). Specific localities: Cameroon – Maan (TL); Dja (Libert, 2020); Ebogo (Libert, 2020). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola mpangensis (Jackson, 1962)

Epitola mpangensis Jackson, 1962. Bulletin of the British Museum (Natural History) (Entomology) 12: 149 (125-163). Cephetola mpangensis (Jackson, 1962). Libert, 1999 comb. nov. Cephetola mpangensis Jackson, 1962. d’Abrera, 2009: 682. Cephetola mpangensis (Jackson, 1962). Libert, 2020: 108.

Cephetola mpangensis mpangensis. Male. Left – upperside; right – underside. Mabira Forest, Uganda. 6 March 2010. P. Ward. Images M. C. Williams ex P. Ward collection.

Cephetola mpangensis mpangensis. Female. Left – upperside; right – underside. Mubenda, Uganda. Em. 17 May 2010. P. Ward. Images M. C. Williams ex P. Ward collection.

Type locality: Uganda: “Bwamba”. Distribution: Cameroon, Democratic Republic of Congo, Uganda, Tanzania. Recorded, in error, from Kenya by Larsen (1991c). Habitat: Primary forest. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

Cephetola mpangenis mpangensis (Jackson, 1962)

Cephetola mpangensis mpangensis. Male. Left – upperside; right – underside. Mabira Forest, Uganda. 6 March 2010. P. Ward. Images M. C. Williams ex P. Ward collection.

Cephetola mpangensis mpangensis. Female. Left – upperside; right – underside. Mubenda, Uganda. Em. 17 May 2010. P. Ward. Images M. C. Williams ex P. Ward collection.

Type locality: Uganda: “Bwamba”. Distribution: Democratic Republic of Congo, Uganda, Tanzania (north-west). Recorded, in error, from Kenya by Larsen (1991c). Specific localities: Democratic Republic of Congo – Semuliki Valley (Ducarme, 2018); Mt Mitumba (Ducarme, 2018). Uganda – Bwamba (TL); Mabira (male images above); Mubenda (female images above). Tanzania – Kere Hill, Minziro Forest (not uncommon) (Congdon & Collins, 1998); MII in Minziro Forest (Congdon & Collins, 1998).

Cephetola mpangenis christinae Libert, 2020

Cephetola mpangensis christinae Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 109 (157 pp.; 15 plates). Published by the author.

Cameroon: Holotype (male): Maan, VI 2001 (ABRI Coll.); in ABRI Collection, Nairobi. Allotype (female): Maan, XI 2003 (ABRI Coll.); in ABRI Collection, Nairobi.

19 Distribution: Cameroon (south). Specific localities: Cameroon – Maan (TL); Lolodorf (Libert, 2020); Dja (Libert, 2020).

*Cephetola kiellandi (Libert & Congdon, 1998)

Epitola kiellandi Libert & Congdon, 1998. In: Congdon & Collins, 1998. Supplement to Kielland’s butterflies of Tanzania : 113 (143 pp.). ABRI and Lambillionea. Cephetola kiellandi (Libert & Congdon, 1998). Libert, 1999 comb. nov. Cephetola kiellandi Libert & Congdon, 1998. d’Abrera, 2009: 682.

Type locality: Tanzania: “Minziro, 1200 m, Bukoba, Tanzanie, 1 I 1995 (I. Kielland)”. Holotype in ABRI, Nairobi. Distribution: Central African Republic, Democratic Republic of Congo, Uganda, Tanzania (north- west) (Libert, 1999). Specific localities: Democratic Republic of Congo – Ituri Forest (Ducarme, 2018); Mt Mitumba (Ducarme, 2018). Tanzania – Kere Hill, Minziro Forest (common) (TL; Congdon & Collins, 1998). Habitat: Forest. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola mitumbensis Libert, 2020

Cephetola mitumbensis Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 111 (157 pp.; 15 plates). Published by the author.

Type locality: Democratic Republic of Congo: Holotype (male): Mambungu, Kivu, I 2014 (ABRI Leg.); BOLD: MLIB-1941; in ABRI Collection, Nairobi. Allotype (female): Mambungu, Kivu, II 2015 (ABRI Leg.); BOLD: MLIB-2084; in ABRI Collection, Nairobi. Distribution: Democratic Republic of Congo (north-east). Specific localities: Democratic Republic of Congo – Mambungu (TL). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola soror Libert, 2020

Cephetola soror Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 112 (157 pp.; 15 plates). Published by the author.

Type locality: Cameroon: Holotype (female): Bikoman, V 2012 (ABRI Leg.); BOLD: MLIB-2035; in ABRI Collection, Nairobi. Distribution: Cameroon. Specific localities: Cameroon – Bikoman (TL); Dja (Libert, 2020). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola marginalis Libert, 2020

Cephetola marginalis Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 113 (157 pp.; 15 plates). Published by the author.

Type locality: Democratic Republic of Congo: Holotype (male): Lubango, Kivu, I 2016 (ABRI Leg.); BOLD: MLIB-1938; in ABRI Collection, Nairobi. Distribution: Democratic Republic of Congo, Uganda. Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

Cephetola marginalis marginalis Libert, 2020

Cephetola marginalis Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 113 (157 pp.; 15 plates). Published by the author.

Type locality: Democratic Republic of Congo: Holotype (male): Lubango, Kivu, I 2016 (ABRI Leg.); BOLD: MLIB-1938; in ABRI Collection, Nairobi. Distribution: Democratic Republic of Congo. Specific localities: Democratic Republic of Congo – Lubango (TL).

Cephetola marginalis ambigua Libert, 2020

Cephetola marginalis ambigua Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 114 (157 pp.; 15 plates). Published by the author.

Type locality: Uganda: Mabira Forest, 6 III 2011 (P.R.F. Ward); genitalia Libert 112-133; BOLD: MLIB-0907; in ABRI Collection, Nairobi. Distribution: Uganda. Specific localities: Uganda – Mabira (TL).

*Cephetola marci Collins & Libert, 1999

Cephetola marci Collins & Libert, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 168 (219 pp.). Cephetola marci Collins & Libert, 1999. d’Abrera, 2009: 680.

Type locality: Cameroon: “Ebogo, Nyong, Cameroun”. Distribution: Cameroon (Libert, 1999). Specific localities: Cameroon – Ebogo, Nyong (TL). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola karinae Bouyer & Libert, 1999

Cephetola karinae Bouyer & Libert, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 169 (219 pp.). Cephetola karinae Bouyer & Libert, 1999. d’Abrera, 2009: 680.

Type locality: Central African Republic: “Ngbo, R.C.A.”. Distribution: Central African Republic (Libert, 1999). Specific localities: Central African Republic – Ngbo (TL). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola katerae (Jackson, 1962)

Epitola katerae Jackson, 1962. Bulletin of the British Museum (Natural History) (Entomology) 12: 149 (125-163). Cephetola katerae (Jackson, 1962). Libert, 1999 comb. nov. Cephetola katerae Jackson, 1962. d’Abrera, 2009: 682.

Cephetola katerae. Male. Left – upperside; right – underside. Mabira Forest, Uganda. 15 May 2011. P. Ward. Images M. C. Williams ex P. Ward collection.

Cephetola katerae. Female. Left – upperside; right – underside. Zambezi Rapids, Ikelenge, Zambia. 28 March 2007. A. Gardiner. Images M. Williams ex Gardiner Collection.

Type locality: Uganda: “Masaka, Sango Bay, Katera”. Distribution: Cameroon (south-west), Gabon (east-central), Democratic Republic of Congo, Uganda, Kenya (south-west), Tanzania (west), Zambia (Libert, 1999). Specific localities: Gabon – Bateke Plateau (Kebiri Forest and camp PPG) (Vande weghe, 2010). 22 Democratic Republic of Congo – Ituri Forest (Ducarme, 2018); Semuliki Valley (Ducarme, 2018). Uganda – Masaka, Sango Bay, Katera (TL). Tanzania – Ntakatta Forest in Mpanda (Kielland, 1990d); Kemfu Forest in Kigoma (Kielland, 1990d); Kasye Forest in Kigoma (Kielland, 1990d); Kere Hill (not uncommon) and Bulembe, Minziro Forest (Congdon & Collins, 1998); Kikuru Forest (Congdon & Collins, 1998); Kampisa River near Sitebi Mountain (Bampton and Congdon). Zambia – Mwinilunga district (female illustrated above); Zambezi Rapids, Ikelenge (male illustrated above). Habitat: Forest. At altitudes between 900 and 1 500 metres in Tanzania (Kielland, 1990d). Also in riparian forest (Congdon & Collins, 1998). Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola kisusui Libert, 2020

Cephetola kisusui Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 117 (157 pp.; 15 plates). Published by the author.

Type locality: Democratic Republic of Congo: Holotype (male): Lubango, Kivu, I 2016 (ABRI Leg.); in ABRI Collection, Nairobi. Allotype (female): Kasuo, N. Kivu, V 2012 (ABRI Leg.); BOLD: MLIB- 1468; in ABRI Collection, Nairobi. Distribution: Democratic Republic of Congo (north-east). Specific localities: Democratic Republic of Congo – Lubango (TL); Kasuo (Libert, 2020). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola bwamba (Jackson, 1964)

Epitola bwamba Jackson, 1964. Bulletin of the British Museum (Natural History) (Entomology) 15: 73 (59-80). Cephetola bwamba (Jackson, 1964). Libert, 1999 comb. nov. Cephetola bwamba Jackson, 1964. d’Abrera, 2009: 682. Cephetola bwamba (Jackson, 1964). Libert, 2020: 118.

Type locality: Uganda: “Bwamba, Mongiro”. Distribution: Democratic Republic of Congo (north-east), Uganda (Libert, 2020). Specific localities: Uganda – Mongiro Forest, Bwamba (TL); Semuliki N.P. (Forbes, 2018). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola hartorum Libert, 2020

Cephetola hartorum Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 119 (157 pp.; 15 plates). Published by the author.

Type locality: Democratic Republic of Congo: Holotype (male): Kasuo, 2000 m, N Kivu, V 2012 (ABRI Leg.); BOLD: MLIB-1469; in ABRI Collection, Nairobi. Allotype (female): Lubango, V 2016

23 (ABRI Leg.); in ABRI Collection, Nairobi. Distribution: Democratic Republic of Congo Specific localities: Democratic Republic of Congo – Kasuo (TL); Lubango (Libert, 2020); Kasugho (Libert, 2020); Maliva (Libert, 2020). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola wardi Libert, 2020

Cephetola wardi Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 24 (157 pp.; 15 plates). Published by the author.

Cephetola wardi. Male. Left – upperside; right – underside. Mabira Forest, Uganda. 11 July 2010. P. Ward. Images M. C. Williams ex P. Ward collection.

Type locality: Type locality: Uganda: Holotype (male): Mabira Forest, 29 XII 2009 (P.R.F. Ward); genitalia Libert 117-046; BOLD: MLIB-0905; in ABRI Collection, Nairobi. Distribution: Uganda. Specific localities: Uganda – Bobiri (TL). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola izidori (Kielland & Congdon, 1998)

Epitola izidori Kielland & Congdon, 1998. In: Congdon & Collins, 1998. Supplement to Kielland’s butterflies of Tanzania : 115 (143 pp.). ABRI and Lambillionea. Cephetola izidori (Kielland & Congdon, 1998). Libert, 1999 comb. nov. Cephetola izidori Kielland & Congdon, 1998. d’Abrera, 2009: 684. Cephetola izidori (Kielland & Congdon, 1998). Libert, 2020: 120.

Cephetola izidori izidori. Male. Left – upperside; right – underside. Mabira Forest, Uganda. 1 January 2010. P. Ward. Images M. C. Williams ex P. Ward collection.

Cephetola izidori izidori. Female. Left – upperside; right – underside. Mabira Forest, Uganda. 20 December 2009. P. Ward. Images M. C. Williams ex P. Ward collection.

Type locality: Tanzania: “Minziro, Bukoba dist., Tanzanie, 12 XII 1994 (I. Kielland)”. Holotype male in ABRI, Nairobi. Distribution: Cameroon, Central African Republic, Democratic Republic of Congo, Uganda, Kenya, Tanzania, Zambia (Libert, 2020). Habitat: Forest. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

Cephetola izidori izidori (Kielland & Congdon, 1998)

Cephetola izidori izidori. Male. Left – upperside; right – underside. Mabira Forest, Uganda. 1 January 2010. P. Ward. Images M. C. Williams ex P. Ward collection.

Cephetola izidori izidori. Female. Left – upperside; right – underside. Mabira Forest, Uganda. 20 December 2009. P. Ward. Images M. C. Williams ex P. Ward collection.

Type locality: Tanzania: “Minziro, Bukoba dist., Tanzanie, 12 XII 1994 (I. Kielland)”. Holotype male in ABRI, Nairobi. Distribution: Cameroon, Central African Republic, Democratic Republic of Congo (north-east), Uganda, Kenya (west), Tanzania (north-west) (Libert, 2020). Specific localities: Cameroon – Dja (Libert, 2020); Maan (Libert, 2020). Central African Republic – Bangui (single male) (Congdon & Collins, 1998). Democratic Republic of Congo – Biakatu (Libert, 2020); Lume (Libert, 2020) Uganda – Mabira Forest (mal and femal images above). Kenya – Kakamega Forest (Congdon & Collins, 1998). Tanzania – Kere Hill, Minziro Forest (TL; Congdon & Collins, 1998).

Cephetola izidori zambeziae Libert & Collins, 1999

Cephetola izidori zambeziae Libert & Collins, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 172 (219 pp.). Cephetola izidori zambeziae Libert & Collins, 1999. Libert, 2020: 120.

Cephetola izidori zambeziae. Male. Left – upperside; right – underside. Zambezi Rapids, Ikelenge, Zambia. 29 March 2007. A. Gardiner. Images M. Williams ex Gardiner Collection.

Cephetola izidori zambeziae. Female. Left – upperside; right – underside. Hillwood Sakeji, Ikelenge, Zambia. 22 March 2007. A. Gardiner. Images M. Williams ex Gardiner Collection.

Type locality: Zambia: “Zambezi R., Hillwood, Ikelenge, Zambie”. Distribution: Zambia (north-west) (Libert, 2020). Specific localities: Zambia – Ikelenge (Heath et al., 2002); Hillwood Farm, Ikelenge (male and female illustrated above).

Note: D’Abrera (2009) does not list Cephetola izidori zambeziae Libert & Collins, 1999.

*Cephetola aureliae Libert, 1999

Cephetola aureliae Libert, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 174 (219 pp.). Cephetola aureliae Libert, 1999. d’Abrera, 2009: 680.

Type locality: [Democratic Republic of Congo]: “Lume, Zaïre”. Distribution: Democratic Republic of Congo. Specific localities: Democratic Republic of Congo – Lume (TL); Semuliki Valley (Ducarme, 2018); Mt Mitumba (Ducarme, 2018). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola wologizi Sáfián, 2021

Cephetola wologizi Sáfián, 2021. Zootaxa 4981 (3): 571 (554-576).

Type locality: Liberia: Holotype (male): Lofa County, Wologizi Mountains, Belegizi Ridge and summit Mountains general collecting, 8°7’27.24»N, 9°56’10.91»W, 1000-1086 m, 4–9.XI.2018 Sáfián, Sz., Simonics, G. Leg. ANHRT unique ID number: ANHRTUK00058086. Gen. prep. SAFI00385. Deposited in ANHRT. The female is unknown. Distribution: Liberia. Specific localities: Liberia – Belegizi Ridge, Wologizi Mountains (TL). Habitat: Submontane forest (Safian et al., 2021). Habits: Males display in the subcanopy of forest on hilltops at about 1000 m.a.s.l., between 09h30 and 10h00 (Safian et al., 2021). Early stages: Nothing published. Larval food: Nothing published.

*Cephetola wingae Safian, 2015

Cephetola wingae Safian, 2015. Metamorphosis 26: 16 (12-19).

Type locality: Liberia: “Mount Beeton, Nimba Mountains, Western Range, Nimba County, Liberia. Leg.: Safian, Sz. & Horvath, A. 10–16.xii.2013. Coordinates: 07°31'52"N 08°39'22"W. Gen. prep: SAFI00068”. Holotype (male) deposited in the African Butterfly Research Institute collection, Nairobi, Kenya. Distribution: Liberia. Specific localities: Liberia: Mount Beeton, Nimba Mountains (TL); Mount Bele (Blei), Blei Coomunity Forest, Nimba Mountains (Safian, 2015); Wologizi Mtns (Safian et al., 2020). Habitat: Upland forest (Safian, 2020). Habits: Males hilltop between 08h30 and 09h00. They fly in small clearings, from 5 to 6 m above the ground, rarely settling (Safian, 2015). Early stages: Nothing published. Larval food: Nothing published.

*Cephetola peteri (Kielland & Congdon, 1998)

Epitola peteri Kielland & Congdon, 1998. In: Congdon & Collins, 1998. Supplement to Kielland’s butterflies of Tanzania : 116 (143 pp.). ABRI and Lambillionea. Cephetola peteri (Kielland & Congdon, 1998). Libert, 1999 comb. nov. Cephetola peteri Kielland & Congdon, 1998. d’Abrera, 2009: 684. Cephetola peteri (Kielland & Congdon, 1998). Libert, 2020: 122.

Type locality: Tanzania: “Minziro, Bukoba dist., Tanzanie, 20 I 1995 (I. [sic] Kielland)”. Holotype in ABRI, Nairobi. Distribution: ? Uganda, Kenya, Tanzania (Libert, 2020). Habitat: Forest. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

Cephetola peteri peteri (Kielland & Congdon, 1998)

Type locality: Tanzania: “Minziro, Bukoba dist., Tanzanie, 20 I 1995 (I. [sic] Kielland)”. Holotype in ABRI, Nairobi. Distribution: ? Uganda, Kenya, Tanzania (north-west) (Libert, 2020). Specific localities: ?Uganda - ?Mabira (Libert, 2020). Kenya – Kakamega Forest (Congdon & Collins, 1998). Tanzania – Kere Hill, Minziro Forest (very common) (TL; Congdon & Collins, 1998).

Cephetola peteri uniformis Libert, 2020

28 Cephetola peteri uniformis Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 122 (157 pp.; 15 plates). Published by the author.

Type locality: Democratic Republic of Congo: Holotype (male): Maliva, E Kivu, II 2014 (ABRI Leg.); genitalia Libert 117-126; in ABRI Collection, Nairobi. Allotype (female): Maliva, E Kivu, IV 2014 (ABRI Leg.); in ABRI Collection, Nairobi. Distribution: Democratic Republic of Congo (north-east). Specific localities: Democratic Republic of Congo – Maliva (TL); Kanyatsi (Libert, 2020); Kasugho (Libert, 2020); Kibale (Libert, 2020); Kithokolo (Libert, 2020); Mambungu (Libert, 2020).

*Cephetola godarti Libert & Collins, 1999

Cephetola godarti Libert & Collins, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 173 (219 pp.). Cephetola godarti Libert & Collins, 1999. d’Abrera, 2009: 680.

Type locality: Central African Republic: “Botambi, R.C.A.”. Distribution: Central African Republic (Libert, 1999). Specific localities: Central African Republic – Botambi (TL). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola australis Libert, 1999

Cephetola australis Libert, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 174 (219 pp.). Cephetola australis Libert, 1999. d’Abrera, 2009: 680.

Type locality: Mozambique: “Delagoa Bay, Mozambique”. Distribution: Mozambique (Libert, 1999). According to Congdon et al., 2010 this is biogeographically improbable. Specific localities: Mozambique – Delagoa Bay [=Maputo] (TL). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola kakamegae Libert & Collins, 1999

Cephetola kakamegae Libert & Collins, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 187 (219 pp.). Cephetola kakamegae Libert & Collins, 1999. d’Abrera, 2009: 682. Cephetola kakamegae Libert & Collins, 1999. Libert, 2020: 123.

Type locality: Kenya: “Kakamega for., W. Kenya”. Distribution: Democratic Republic of Congo, Uganda, Kenya (Libert, 2020). Specific localities: Democratic Republic of Congo – Ituri Forest (Ducarme, 2018); Mt Mitumba (Ducarme, 2018); Manzumbu (Libert, 2020); Mapimbi (Libert, 2020). Uganda – Mabira (Libert, 2020). Kenya – Kakamega Forest (TL).

29 Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola formosa Collins & Libert, 2020

Cephetola formosa Collins & Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 124 (157 pp.; 15 plates). Published by the author.

Type locality: Democratic Republic of Congo: Holotype (male): Kanyambia, 10 km south-west of Lubero, Kivu, VII 2011 (ABRI Leg.); in ABRI Collection, Nairobi. Allotype (female): Kanyambia, 10 km south-west of Lubero, Kivu, X 2011 (ABRI Leg.); in ABRI Collection, Nairobi. Distribution: Democratic Republic of Congo (north-east). Specific localities: Democratic Republic of Congo – Kanyambia (TL); Bikara (Libert, 2020); Kanyatsi (Libert, 2020); Kasuo (Libert, 2020); Kasugho (Libert, 2020); Kibale (Libert, 2020); Lubango (Libert, 2020). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola ducarmei Libert, 1999

Cephetola ducarmei Libert, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 188 (219 pp.). Cephetola ducarmei Libert, 1999. d’Abrera, 2009: 682. Cephetola ducarmei Libert, 1999. Libert, 2020: 125.

Type locality: [Democratic Republic of Congo]: “Mapimbi (1150 m), Zaïre”. Neallotype (female): Mamove, Kivu, IX 2016 (ABRI Leg.); BOLD: MLIB-2221; in ABRI Collection, Nairobi. Distribution: Democratic Republic of Congo (Libert, 2020). Specific localities: Democratic Republic of Congo – Mapimbi [Mapembe] (TL); Ituri Forest (Ducarme, 2018); Mt Mitumba (Ducarme, 2018); Biakatu (Libert, 2020); Cantine (Libert, 2020); Epulu (Libert, 2020); Mamove (Libert, 2020). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola frater Libert, 2020

Cephetola fraterLibert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 126 (157 pp.; 15 plates). Published by the author.

Type locality: Democratic Republic of Congo: Holotype (female): Maliva, N Kivu, II 2015 (ABRI Leg.); BOLD: MLIB-1936; in ABRI Collection, Nairobi. Allotype (male): Biakatu, Orientale Province, II 2011 (R. Ducarme); genitalia Libert 117-021; BOLD: MLIB-2174; in collection of R. Ducarme. Distribution: Democratic Republic of Congo (north-east). Specific localities: Democratic Republic of Congo – Maliva (TL); Biakatu (Libert, 2020); Mamove (Libert, 2020). Habitat: Nothing published. Habits: Nothing published.

30 Early stages: Nothing published. Larval food: Nothing published.

*Cephetola rileyi (Audeoud, 1936)

Epitola rileyi Audeoud, 1936. Bulletin de la Société Lépidoptérologique de Genève 7: 186 (184-187). Cephetola rileyi (Audeoud, 1936). Libert, 1999 comb. nov. Cephetola rileyi Audeoud, 1936. d’Abrera, 2009: 682.

Type locality: Cameroon: “Lolodorf”. Distribution: Cameroon (Libert, 1999). Specific localities: Cameroon – Lolodorf (TL). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola pinodes (Druce, 1890) Druce’s Epitola

Epitola pinodes Druce, 1890. Annals and Magazine of Natural History (6) 5: 24 (24-31). Cephetola pinodes (Druce, 1890). Libert, 1999 comb. nov. Cephetola pinodes Druce, 1890. d’Abrera, 2009: 680.

Cephetola pinodes budduana. Male. Left – upperside; right – underside. Radio Hill, Mabira Forest, Uganda. 14 June, 2009. J. & C. Dobson. Images M.C.Williams ex Dobson Collection.

Type locality: Nigeria: “Lagos”. Distribution: Sierra Leone, Liberia, Ivory Coast, Ghana, Nigeria, Cameroon, Gabon, Congo, Central African Republic, Democratic Republic of Congo, Uganda, Tanzania. Habitat: Nothing published. Habits: This is a localized species but numbers can be found around individual ant-trees (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published.

Cephetola pinodes pinodes (Druce, 1890)

Epitola pinodes Druce, 1890. Annals and Magazine of Natural History (6) 5: 24 (24-31). Cephetola pinodes (Druce, 1890). Libert, 1999 comb. nov. Cephetola pinodes Druce, 1890. d’Abrera, 2009: 680. 31

Type locality: Nigeria: “Lagos”. Distribution: Sierra Leone, Liberia, Ivory Coast, Ghana, Nigeria. Specific localities: Sierra Leone – Mocquerys (Grose-Smith & Kirby, 1893); Freetown (Libert, 2020); Bumbuna and Farangbaya forests near Mt Loma (Libert, 2020). Liberia – Wologizi (Safian et al., 2020). Ivory Coast – Issia (Jackson’s collectors, vide Larsen 2005a). Nigeria – Lagos (TL).

conjuncta Grose-Smith & Kirby, 1893 (as sp. of Epitola). In: Grose-Smith & Kirby, 1892-7 (Rhopalocera exotica, being illustrations of new, rare and unfigured species of butterflies 2: 86 (261 pp.). London.) Sierra Leone: “Mocquerys”. Given by Ackery et al., 1995: 521 as a good species but regarded as a synonym of Epitola pinodes Druce, 1890 by Congdon & Collins, 1998 ( Supplement to Kielland’s butterflies of Tanzania: 78 (143 pp.). ABRI and Lambillionea).

Cephetola pinodes budduana (Talbot, 1937)

Epitola conjuncta budduana Talbot, 1937. Transactions of the Royal Entomological Society of London 86: 62 (59-72). Epitola pinodes budduana Talbot, 1937. Congdon & Collins, 1998: 78. Cephetola pinodes budduana (Talbot, 1937). Libert, 1999 comb. nov. Cephetola pinodes budduana Talbot, 1937. d’Abrera, 2009: 680.

Cephetola pinodes budduana. Male. Left – upperside; right – underside. Radio Hill, Mabira Forest, Uganda. 14 June, 2009. J. & C. Dobson. Images M.C.Williams ex Dobson Collection.

Type locality: [Uganda]: “Katera”. Distribution: Democratic Republic of Congo, Uganda, Tanzania (north-west) (Libert, 2020). Specific localities: Gabon – Waka (Vande weghe, 2010). Democratic Republic of Congo – Ituri Forest (Ducarme, 2018); Central Forest Block (Ducarme, 2018); Mt Mitumba (Ducarme, 2018); Katako-Kombe (Libert, 2020); Kisangani (Libert, 2020). Uganda – Katera (TL); Semuliki N.P. (H. Selb, unpublished, 2016); Buddu (Libert, 2020). Tanzania – Minziro Forest (Congdon & Collins, 1998); Kikuru Forest (Congdon & Collins, 1998); Munene Forest (uncommon) (Congdon & Collins, 1998).

*Cephetola intricata Libert, 2020

Cephetola intricata Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 128 (157 pp.; 15 plates). Published by the author.

Type locality: Cameroon: Holotype (male): Nyong River, Ebogo, X 199? (S. Collins); in ABRI Collection, Nairobi.

32 Distribution: Cameroon, Gabon, Congo, Central African Republic, Democratic Republic of Congo (west). Specific localities: Cameroon – Nyong Rivier, Ebogo (TL); Edea (Libert, 2020); Ngambe Tikar (Libert, 2020); Bertoua (Libert, 2020); Maan (Libert, 2020); Dja (Libert, 2020); Lolodorf (Libert, 2020); Mbega (Libert, 2020); Meyo Messi (Libert, 2020); Obout (Libert, 2020). Gabon – Oyem (Libert, 2020); Waka (Libert, 2020). Congo – Sembe (Libert, 2020); Tchissanga (Libert, 2020). Central African Republic – Boukoko (Libert, 2020); Yakoli (Libert, 2020). Democratic Republic of Congo – Mbandaca (Libert, 2020); Kinshasa (Libert, 2020). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola dolorosa (Roche, 1954)

Epitola dolorosa Roche, 1954. Bulletin of the British Museum (Natural History) (Entomology) 3: 498 (489-501). Cephetola dolorosa (Roche, 1954). Libert, 1999 comb. nov. Cephetola dolorosa Roche, 1954. d’Abrera, 2009: 684. Cephetola dolorosa (Roche, 1954). Libert, 2020: 127.

Type locality: Uganda: “Bwamba Forest, Semliki Valley”. Distribution: Sudan, Democratic Republic of Congo, Uganda (Libert, 2020). Recorded, apparently in error, from Tanzania by Kielland (1990d). Specific localities: Democratic Republic of Congo – Masau (Libert, 2020). Uganda – Bwamba Forest, Semliki Valley (TL). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola overlaeti Libert, 1999

Cephetola overlaeti Libert, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 179 (219 pp.). Cephetola overlaeti Libert, 1999. d’Abrera, 2009: 680. Cephetola overlaeti Libert, 1999. Libert, 2020: 127.

Type locality: [Democratic Republic of Congo]: “R. Mokweji, Kafakumba (Katanga), Zaïre”. Distribution: Democratic Republic of Congo (Libert, 1999). Specific localities: Democratic Republic of Congo – Mokweji River, Kafakumba, Katanga (TL). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola sublustris (Bethune-Baker, 1904) Greasy Epitola

Epitola sublustris Bethune-Baker, 1904. Annals and Magazine of Natural History (7) 14: 228 (222-233). Cephetola sublustris (Bethune-Baker, 1904). Libert, 1999 comb. nov. Cephetola sublustris Bethune-Baker, 1904. d’Abrera, 2009: 682. 33 Cephetola sublustris (Bethune-Baker, 1904). Libert, 2020: 130.

Type locality: Sierra Leone: “Sierra Leone [Mabang (Larsen, 2005a)]”. Distribution: Senegal, Sierra Leone, Liberia, Ivory Coast, Ghana, Nigeria, Cameroon, Equatorila Guinea (Bioko), Congo, Central African Republic, Democratic Republic of Congo, Uganda, Tanzania (north-west). Misattributed to the Senegal fauna by Condamin (Larsen, 2005a). Specific localities: Sierra Leone – Mabang (TL). Liberia – Wologizi (Safian et al., 2020). Ivory Coast – Banco (Warren-Gash; single male; vide Larsen, 2005a). Nigeria – Lagos area (Larsen, 2005a); Umuhaia (Larsen, 2005a). Democratic Republic of Congo – Beni (Libert, 1999); Ituri Forest (Ducarme, 2018). Uganda – Kagera (Libert, 1999). Tanzania – Minziro (Libert, 1999). Habitat: Forest. Habits: Fairly common but tied to individual ant-trees (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published.

*Cephetola collinsi Libert & Larsen, 1999

Cephetola collinsi Libert & Larsen, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 181 (219 pp.). = Cephetola sublustris (Bethune-Baker, 1904). d’Abrera, 2009: 682. syn. nov. Cephetola collinsi Libert & Larsen, 1999. Collins et al., 2013: 52. stat. rev. Cephetola collinsi Libert & Larsen, 1999. Libert, 2020: 131.

Type locality: Ghana: “Aburi Botanical Gardens, Ghana”. Distribution: Liberia, Ivory Coast, Ghana, Togo, Benin (south), Nigeria. Specific localities: Ivory Coast – Niangon (Libert, 1999); Tiassale (Libert, 1999). Ghana – Aburi Botanical Gardens (TL); Kpandu (Libert, 1999); Ashanti (Libert, 1999). Togo – Badou (Libert, 1999). Benin – see Coache et al., 2017. Nigeria – Lagos (Libert, 1999); Oni (Libert, 1999); Umuhaia (Libert, 1999). Habitat: Habits: Specimens of collinsi were noted patrolling the lawns at the type locality [Aburi Botanical Gardens, Ghana] between 10h00 and 11h00 (Larsen, 2005a). This behaviour was preceded by similar behaviour, on the same lawns, by Cephetola subcoerulea (08h30 to 09h30) and was followed by Hypophytala hyettoides (11h00 to 13h00). Early stages: Nothing published. Larval food: Nothing published.

*Cephetola maesseni Libert, 1999 Maessen’s Greasy Epitola

Cephetola maesseni Libert, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 182 (219 pp.). Cephetola maesseni Libert, 1999. d’Abrera, 2009: 680. Cephetola maesseni Libert, 1999. Libert, 2020: 136.

Type locality: Ghana: “Lipke [Likpe], Ghana”. Distribution: Ghana (Volta Region), Cameroon, Gabon, Congo, Central African Republic (Libert, 2020). Specific localities: Ghana – Likpe (TL).

34 Habitat: Forest. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

Note: Libert (1999) avers that a specimen figured in Berger (1981) may be referable to this species but conspecificity has not been substantiated.

*Cephetola vinalli (Talbot, 1935)

Epitola vinalli Talbot, 1935. Entomologist’s Monthly Magazine 71: 75 (69-78, 115-127, 147-153). Cephetola vinalli (Talbot, 1935). Libert, 1999 comb. nov. Cephetola vinalli Talbot, 1935. d’Abrera, 2009: 682.

Type locality: [Democratic Republic of Congo]: “Basankusu, 40 miles west of Bongandanga, Baolongo”. Distribution: Congo, Central African Republic, Democratic Republic of Congo, Uganda, Tanzania (north-west). Specific localities: Democratic Republic of Congo – Basankusu (TL); Ituri Forest (Ducarme, 2018); Mt Mitumba (Ducarme, 2018). Tanzania – Kere Hill, Minziro Forest (rare) (Congdon & Collins, 1998). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

pseudoconjuncta Jackson, 1962 ( as sp. of Epitola). Bulletin of the British Museum (Natural History) (Entomology) 12: 153 (125-163). Nigeria: “Ogaja Prov., Ikom”. Given as a good species of Epitola by Ackery et al., 1995: 524 but synonymized with Cephatola vinalli (Talbot, 1935) by Libert, 1999. (In: Libert, 1999. Révision des Epitola (l.s.): 186 (219 pp.)). Since pseudoconjuncta was described from Nigeria and vinalli does not occur this far west some mistake is involved here [MCW].

*Cephetola subcoerulea (Roche, 1954) Roche’s Epitola

Epitola subcoerulea Roche, 1954. Bulletin of the British Museum (Natural History) (Entomology) 3: 498 (489-501). Synonym of Epitola vinalli Talbot, 1935. Ackery et al., 1995: 525. Cephetola subcoerulea (Roche, 1954). Libert 1999: 184 stat. rev., comb. nov. Cephetola subcoerulea Roche, 1954. d’Abrera, 2009: 682. Cephetola subcoerulea (Roche, 1954). Libert, 2020: 134.

Type locality: Sierra Leone: “Sierra Leone”. Distribution: Senegal (Basse Casamance), Sierra Leone, Liberia, Ivory Coast, Ghana, Nigeria (east), Cameroon, Equatorial Guinea (Bioko). Recorded, in error, from Gambia (Jon Baker, pers. comm., May 2020). Specific localities: Liberia – Wologizi (Safian et al., 2020). Ghana – Aburi Botanical Gardens (Larsen, 2005a). Habitat: Forest. Habits: The species is scarce and is closely associated with individual ant-trees (Larsen, 2005a). At Aburi Larsen (2005a) noted that specimens patrolled the lawns between 08h30 and 09h30 (see Cephetola collinsi for further notes). Early stages: Nothing published. Larval food: Nothing published.

*Cephetola chari Libert & Collins, 1999

Cephetola chari Libert & Collins, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 184 (219 pp.). Cephetola chari Libert & Collins, 1999. d’Abrera, 2009: 682. Cephetola chari Libert & Collins, 1999. Libert, 2020: 133.

Type locality: Central African Republic: “Botambi, R.C.A.”. Distribution: Cameroon, Congo, Central African Republic (Libert, 2020). Specific localities: Cameroon – Koutaba (Libert, 2020). Congo – Etoumbi (Libert, 2020). Central African Republic – Botambi (TL). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

*Cephetola mengoensis (Bethune-Baker, 1906)

Epitola mengoensis Bethune-Baker, 1906. Annals and Magazine of Natural History (7) 17: 105 (104-110). Cephetola mengoensis (Bethune-Baker, 1906). Libert, 1999 comb. nov. Cephetola mengoensis Bethune-Baker, 1906. d’Abrera, 2009: 682. Cephetola mengoensis (Bethune-Baker, 1906). Libert, 2020: 136.

Type locality: Uganda: “Mengo”. Distribution: Uganda, Kenya (Libert, 1999). Recorded, apparently in error, from Tanzania by Kielland (1990d). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

Cephetola mengoensis mengoensis (Bethune-Baker, 1906)

Type locality: Uganda: “Mengo”. Distribution: Uganda (Libert, 1999). Specific localities: Uganda – Mengo (TL).

Cephetola mengoensis kenya Libert & Collins, 1999

Cephetola mengoensis kenya Libert & Collins, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 186 (219 pp.). Cephetola mengoensis kenya Libert & Collins, 1999. Libert, 2020: 136.

Type locality: Kenya: “Kakamega, W. Kenya”. Distribution: Kenya (Libert, 1999). Specific localities:

36 Kenya – Kakamega Forest (TL).

Note: This taxon (Cephetola mengoensis kenya Libert & Collins, 1999) is not included in d’Abrera (2009).

*Cephetola tanzaniensis Libert, 1999

Cephetola tanzaniensis Libert, 1999. In: Libert, 1999. Révision des Epitola (l.s.): 186 (219 pp.). Cephetola tanzaniensis Libert, 1999. d’Abrera, 2009: 682. Cephetola tanzaniensis Libert, 1999. Libert, 2020: 134.

Type locality: Tanzania: “Lubalizi, Kigoma, Tanzanie”. Distribution: Democratic Republic of Congo, Tanzania (Libert, 2020). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.

Cephetola tanzaniensis tanzaniensis Libert, 1999

Type locality: Tanzania: “Lubalizi, Kigoma, Tanzanie”. Distribution: Democratic Republic of Congo, Tanzania (Libert, 2020). Specific localities: Democratic Republic of Congo – Ituri Forest (Ducarme, 2018). Tanzania – Lubalizi, Kigoma (TL).

Cephetola tanzaniensis parva Libert, 2020

Cephetola tanzaniensis parva Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 135 (157 pp.; 15 plates). Published by the author.

Type locality: Democratic Republic of Congo: Holotype (male): Baliko-Basayo, 30 km east of Kisangani, Orientale Province, I 2014 (ABRI Leg.); BOLD: MLIB-1867; in ABRI Collection, Nairobi. Distribution: Democratic Republic of Congo (north-east). Specific localities: Democratic Republic of Congo – Baliko-Basayo (TL); Ituri Forest (Ducarme, 2018); Biakatu (Libert, 2020); Mambasa (Libert, 2020); Mamove (Libert, 2020); Mapimbi (Libert, 2020); Manzumbu (Libert, 2020).

*Cephetola yedim Libert, 2020

Cephetola yedim Libert, 2020. Epitola l.s.: mise à jour de la Révision (Lepidoptera, Lycaenidae); 136 (157 pp.; 15 plates). Published by the author.

Type locality: Democratic Republic of Congo: Holotype (male): Yedi-Mongbwalu, Orientale Province, V 2013 (ABRI Leg.); BOLD: MLIB-1491; in ABRI Collection, Nairobi. Distribution: Democratic Republic of Congo (north-east). Specific localities:

37 Democratic Republic of Congo – Yedi-Mongbwalu (TL); Mapimbi (Libert, 2020). Habitat: Nothing published. Habits: Nothing published. Early stages: Nothing published. Larval food: Nothing published.