Genus Phytala Westwood, [1851] Forest Flash

Total Page:16

File Type:pdf, Size:1020Kb

Genus Phytala Westwood, [1851] Forest Flash AFROTROPICAL BUTTERFLIES. MARK C. WILLIAMS. http://www.lepsocafrica.org/?p=publications&s=atb Updated 30 December 2020 Genus Phytala Westwood, [1851] Forest Flash In: Doubleday & Westwood, [1846-52]. The genera of diurnal Lepidoptera, London: pl. 77 (1: 1-250 pp.; 2: 251-534 pp.). London.). Type-species: Phytala elais Westwood, by monotypy. The genus Phytala belongs to the Family Lycaenidae Leach, 1815; Subfamily Poritiinae Doherty, 1886; Tribe Epitolini Jackson, 1962. The other genera in the Tribe Epitolini in the Afrotropical Region are Iridana, Teratoneura, Epitola, Cerautola, Hewitola, Geritola, Stempfferia, Cephetola, Deloneura, Batelusia, Tumerepedes, Pseudoneaveia, Neaveia, Epitolina, Hypophytala, Neoepitola, Aethiopana, Hewitsonia, and Powellana. Phytala (Forest Flash) is a purely Afrotropical genus containing a single species. *Phytala elais Westwood, [1851] Forest Flash Phytala elais Westwood, [1851]. In: Doubleday & Westwood [1846-52]. The genera of diurnal Lepidoptera, London: pl. 77 [1851], 471 [1852] (1: 1-250 pp.; 2: 251-534 pp.). London. Phytala elais Westwood, 1851. d’Abrera, 2009: 664. Phytala elais elais. Male. Left – upperside; right- underside. Mbalmayo, Cameroon. Jan. to Apr. 2002. Gardiner Collection. Images M.C. Williams ex Gardiner Collection. 1 Phytala elais elais. Female. Left – upperside; right – underside. Camp Kombo, Cameroon. 5 May 2019. J. Dobson. Images M.C.Williams ex Dobson Collection. Type locality: Ghana: “Ashanti”. Distribution: Sierra Leone, Liberia, Ivory Coast, Ghana, Benin, Nigeria, Cameroon, Gabon, Congo, Central African Republic, Democratic Republic of Congo, Sudan, Uganda. Habitat: Forest. Habits: Generally scarce (Larsen, 2005a). Often perches on dry palm fronds, which may have earned it its specific name (the same as the generic name for the oil palm) (Larsen, 2005a). A single male has been noted patrolling 20 m of a forest path at high speed (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published. Note: Larsen (2005a) states that there is a possibility that the two subspecies in West Africa represent two distinct species. Phytala elais elais Westwood, [1851] Phytala elais Westwood, [1851]. In: Doubleday & Westwood [1846-52]. The genera of diurnal Lepidoptera, London: pl. 77 [1851], 471 [1852] (1: 1-250 pp.; 2: 251-534 pp.). London. Phytala elais Westwood, 1851. d’Abrera, 2009: 664. Phytala elais elais. Male. Left – upperside; right- underside. Mbalmayo, Cameroon. Jan. to Apr. 2002. Gardiner Collection. Images M.C. Williams ex Gardiner Collection. Phytala elais elais. Female. Left – upperside; right – underside. Camp Kombo, Cameroon. 5 May 2019. J. Dobson. Images M.C.Williams ex Dobson Collection. 2 Type locality: Ghana: “Ashanti”. Distribution: Ivory Coast (east), Ghana, Benin (south), Nigeria (south and Cross River loop), Cameroon, Gabon, Congo, Central African Republic, Democratic Republic of Congo. Specific localities: Ghana – Kakum (Larsen, 2005a). Benin – see Coache et al., 2017. Gabon – Waka (Vande weghe, 2010). Central African Republic – Dzanga (Noss 1998). Democratic Republic of Congo – Central Forest Block (Ducarme, 2018). Phytala elais catori Bethune-Baker, 1903 Phytala elais catori Bethune-Baker, 1903. Annals and Magazine of Natural History (7) 12: 328 (324-334). Phytala elais catori Bethune-Baker, 1903. d’Abrera, 2009: 664. Type locality: Sierra Leone: “Sierra Leone”. Distribution: Sierra Leone, Liberia, Ivory Coast (west). Specific localities: Sierra Leone – Ohne (Strand, 1920). Liberia – Wologizi (Safian et al., 2020). elaidina Strand, 1920 (as sp. of Phytala). Internationale Entomologische Zeitschrift 19: 148 (146-152). Sierra Leone: “Ohne Lokalitätsangbe”. Phytala elais ugandae Jackson, 1964 Phytala elais ugandae Jackson, 1964. Bulletin of the British Museum (Natural History) (Entomology) 15: 60 (59-80). Phytala elais ugandae Jackson, 1964. d’Abrera, 2009: 664. Type locality: Uganda: “Bwamba Forest, Semliki Valley”. Distribution: Democratic Republic of Congo, Sudan (south), Uganda. Specific localities: Democratic Republic of Congo – Ituri Forest (Ducarme, 2018); Mt Mitumba (Ducarme, 2018); Mt Blue (Ducarme, 2018). Uganda – Bwamba Forest, Semuliki Valley (TL); Semuliki N.P. (S. Forbes & H. Selb, unpublished, 2016). 3.
Recommended publications
  • Title Lorem Ipsum Dolor Sit Amet, Consectetur Adipiscing Elit
    Volume 26: 102–108 METAMORPHOSIS www.metamorphosis.org.za ISSN 1018–6490 (PRINT) LEPIDOPTERISTS’ SOCIETY OF AFRICA ISSN 2307–5031 (ONLINE) Classification of the Afrotropical butterflies to generic level Published online: 25 December 2015 Mark C. Williams 183 van der Merwe Street, Rietondale, Pretoria, South Africa. E-mail: [email protected] Copyright © Lepidopterists’ Society of Africa Abstract: This paper applies the findings of phylogenetic studies on butterflies (Papilionoidea) in order to present an up to date classification of the Afrotropical butterflies to genus level. The classification for Afrotropical butterflies is placed within a worldwide context to subtribal level. Taxa that still require interrogation are highlighted. Hopefully this classification will provide a stable context for researchers working on Afrotropical butterflies. Key words: Lepidoptera, Papilionoidea, Afrotropical butterflies, classification. Citation: Williams, M.C. (2015). Classification of the Afrotropical butterflies to generic level. Metamorphosis 26: 102–108. INTRODUCTION Suborder Glossata Fabricius, 1775 (6 infraorders) Infraorder Heteroneura Tillyard, 1918 (34 Natural classifications of biological organisms, based superfamilies) on robust phylogenetic hypotheses, are needed before Clade Obtectomera Minet, 1986 (12 superfamilies) meaningful studies can be conducted in regard to their Superfamily Papilionoidea Latreille, 1802 (7 evolution, biogeography, ecology and conservation. families) Classifications, dating from the time of Linnaeus in the Family Papilionidae Latreille, 1802 (32 genera, 570 mid seventeen hundreds, were based on morphology species) for nearly two hundred and fifty years. Classifications Family Hedylidae Guenée, 1858 (1 genus, 36 species) based on phylogenies derived from an interrogation of Family Hesperiidae Latreille, 1809 (570 genera, 4113 the genome of individual organisms began in the late species) 20th century.
    [Show full text]
  • Check-List of the Butterflies of the Kakamega Forest Nature Reserve in Western Kenya (Lepidoptera: Hesperioidea, Papilionoidea)
    Nachr. entomol. Ver. Apollo, N. F. 25 (4): 161–174 (2004) 161 Check-list of the butterflies of the Kakamega Forest Nature Reserve in western Kenya (Lepidoptera: Hesperioidea, Papilionoidea) Lars Kühne, Steve C. Collins and Wanja Kinuthia1 Lars Kühne, Museum für Naturkunde der Humboldt-Universität zu Berlin, Invalidenstraße 43, D-10115 Berlin, Germany; email: [email protected] Steve C. Collins, African Butterfly Research Institute, P.O. Box 14308, Nairobi, Kenya Dr. Wanja Kinuthia, Department of Invertebrate Zoology, National Museums of Kenya, P.O. Box 40658, Nairobi, Kenya Abstract: All species of butterflies recorded from the Kaka- list it was clear that thorough investigation of scientific mega Forest N.R. in western Kenya are listed for the first collections can produce a very sound list of the occur- time. The check-list is based mainly on the collection of ring species in a relatively short time. The information A.B.R.I. (African Butterfly Research Institute, Nairobi). Furthermore records from the collection of the National density is frequently underestimated and collection data Museum of Kenya (Nairobi), the BIOTA-project and from offers a description of species diversity within a local literature were included in this list. In total 491 species or area, in particular with reference to rapid measurement 55 % of approximately 900 Kenyan species could be veri- of biodiversity (Trueman & Cranston 1997, Danks 1998, fied for the area. 31 species were not recorded before from Trojan 2000). Kenyan territory, 9 of them were described as new since the appearance of the book by Larsen (1996). The kind of list being produced here represents an information source for the total species diversity of the Checkliste der Tagfalter des Kakamega-Waldschutzge- Kakamega forest.
    [Show full text]
  • The Volta Region
    WILDLIFE DIVISION (FORESTRY COMMISSION) REPUBLIC OF GHANA Wildlife Division Support Project (WDSP) The Butterflies of Kyabobo National Park, Ghana, and those of the Volta Region by Torben B Larsen (WDSP Report No. 64) March 2006 In collaboration with: Butterflies of Kyabobo and Volta Region. WDSP Report no 64 March 2006 TABLE OF CONTENTS EXECUTIVE SUMMARY …………………………………… 4 ACKNOWLEDGEMENTS ………………………………….. 7 1. INTRODUCTION ………………………………………… 8 2. KYABOBO NATIONAL PARK …………………………. 9 2.1 Location and characteristics of Kyabobo National Park …… 9 2.1.1 Habitat types ………………………………………………… 9 2.2 The butterflies of Kyabobo National Park …………………. 10 2.2.1 Material and methods ……………………………………….. 10 2.2.2 Analysis of the Kyabobo butterflies ……………………….... 11 2.2.3 Conservation value of Kyabobo National Park ……..………. 14 2.3 Ecotourism potential ..……………………………………... 14 3. VOLTA REGION – ECOLOGY AND BIOGEOGRAPHY 17 3.1 The Volta Region setting …………………………………… 17 3.2 History of butterfly collecting in the Volta Region ………… 18 3.3 Review of the Volta Region butterfly fauna ………………… 19 3.3.1 Total butterfly fauna …………………………………………… 19 3.3.2 Endemics of Africa west of the Dahomey Gap ……………….. 21 3.3.3 Eastern species not found west of the Volta River ……………. 23 3.4 Biogeographical summary …………………………………… 25 3.5 Conservation priorities in the Volta Region ………...………. 25 3.6 Ecotourism …………………………………………………... 27 4. CONCLUDING REMARKS ……………………………… 29 REFERENCES ……………………………………………. 30 APPENDICES: Appendix 1 The butterflies of the Volta Region, Kyabobo, Wli Falls, and Kalakpa…………….….…………….. 33 Appendix 2 Butterflies recorded by Karsch (1893) from Adeli Mountains, German Togoland …………… 55 2 Butterflies of Kyabobo and Volta Region. WDSP Report no 64 March 2006 LIST OF TABLES: Table 2.2.2.
    [Show full text]
  • Butterfly Community Assemblages in Relation to Human Disturbance in a Tropical Upland Forest in Ghana, and Implications for Conservation
    Journal of Insect Biodiversity 3(6): 1-18, 2015 http://www.insectbiodiversity.org RESEARCH ARTICLE Butterfly community assemblages in relation to human disturbance in a tropical upland forest in Ghana, and implications for conservation Patrick Addo-Fordjour1* Benjamin A. Osei1 Elizabeth A. Kpontsu1 1 Department of Theoretical and Applied Biology, College of Science, Kwame Nkrumah University of Science and Technology (KNUST), Kumasi, Ghana *Corresponding author e-mail: [email protected] / [email protected] Abstract: The present study determined butterfly diversity, species composition and abundance in different forests of varying human disturbance intensities in the Atewa Range Forest Reserve, Ghana (i.e. non-disturbed, moderately disturbed and heavily disturbed forests). Vegetation characteristics and butterflies were sampled within ten 50 m × 50 m plots in each forest type. The study revealed that butterfly Shannon diversity index was similar in the non-disturbed and moderately disturbed forests although it was significantly lower in the heavily disturbed forest. Butterfly abundance differed significantly among all the forest types. Significant relationships were detected between some vegetation characteristics, and butterfly diversity and abundance (P<0.001). Using Non-metric Multidimensional Scaling (NMDS) and cluster analysis, three main butterfly assemblages were identified on the basis of species composition, with each one in a particular forest type. Furthermore, butterfly species composition differed significantly among the forest types (ANOSIM; P<0.0001). The intermediate form of human disturbance in the moderately disturbed forest maintained butterfly diversity, suggesting that management efforts aimed at butterfly conservation should be geared towards protecting forests from excessive human disturbance; selective logging is recommended.
    [Show full text]
  • 184 Genus Epitola Westwood
    AFROTROPICAL BUTTERFLIES. MARK C. WILLIAMS. http://www.lepsocafrica.org/?p=publications&s=atb Updated 7 March 2021. Genus Epitola Westwood, [1851] Pointed Flashes In: Doubleday & Westwood, [1846-52]. The genera of diurnal Lepidoptera, London: pl.68 (1: 1-250 pp.; 2: 251-534 pp.). London. Type-species: Epitola elion Westwood, by monotypy. The genus Epitola belongs to the Family Lycaenidae Leach, 1815; Subfamily Poritiinae Doherty, 1886; Tribe Epitolini Jackson, 1962. The other genera in the Tribe Epitolini in the Afrotropical Region are Iridana, Teratoneura, Cerautola, Hewitola, Geritola, Stempfferia, Cephetola, Deloneura, Batelusia, Tumerepedes, Pseudoneaveia, Neaveia, Epitolina, Hypophytala, Phytala, Neoepitola, Aethiopana, Hewitsonia, and Powellana. Epitola (Pointed Flashes) is a purely Afrotropical genus containing five species. Revised by Libert, M. 1999 (Révision des genres Epitola Westwood, Hypophytala Clench et Stempfferia Jackson, et description de trois nouveaux genres (Lepidotera Lycaenidae). A.B.R.I. and Lambillonea, February, 1999, 219 pp.). Revised again by Libert in 2020. Note: Epitola lamborni Bethune-Baker, 1922 and Epitola pulverulentula Dufrane, 1953 (as pulverulenta in Ackery et al., 1995: 524) are regarded by Libert (1999) as nomen dubium. D’Abrera (2009: 684) informally lists pulverulentula (as pulverulenta) as a valid species, ignoring Libert’s taxonomic treatment. Classification of the Genus Epitola (Libert, 2020) E. posthumus clade E. posthumus E. uranoides E. occidentalis E. urania clade E. urania E. larseni *Epitola posthumus (Fabricius, 1793) Common Giant Epitola Papilio posthumus Fabricius, 1793. Entomologia Systematica emendata et aucta 3 (1): 149 (488pp.). Epitola posthumus Fabricius, 1793. d’Abrera, 2009: 664. 1 Epitola posthumus. Male. Left – upperside; right – underside. Mbalmayo, Cameroon. Jan.-April 2002.
    [Show full text]
  • Butterflies As an Indicator Group for the Conservation Value of the Gola Forests in Sierra Leone
    BUTTERFLIES AS AN INDICATOR GROUP FOR THE CONSERVATION VALUE OF THE GOLA FORESTS IN SIERRA LEONE Claudio Belcastro* & Torben B. Larsen** * Lungotevere di Pietro Papa 21 00146 Roma, Italia [email protected] ** 358 Coldharbour Lane London SW9 8PL, UK [email protected] EXECUTIVE SUMMARY Less than 5% of Sierra Leone’s original forest cover still exists, though some of that hardly merits the term forest. Besides the remaining forest on the Freetown Peninsula, and the important Loma and Tingi Mountains, with their submontane elements, Gola Forest is the most significant forest in the country. During late April, 2006, a one week field-trip was made to study the butterflies of the Gola Forests by two separate teams, headed by one of the authors of this report. Belcastro also returned to Gola North for three days in early May. In all, 370 species were positively recorded. The estimated total for the area is about 600, accounting for about 80% of the 750 or so known Sierra Leone butterflies. Many rare and interesting butterflies occur and, in general, the Gola Forests are now the westernmost outpost of the West African forest fauna. Many species endemic to Africa west of the Dahomey Gap and to its Liberia subregion were found in Gola. The fact that so many rare and interesting species were collected in, sometimes quite heavily, logged areas of Gola is a strong indicator that the forests have the capacity to return to a state that resembles the original over the next 25 years. In Gola (South), and especially in Gola (North), there appear to be areas of undisturbed forest that act as reservoirs of biodiversity that help to re-populate the regenerating parts of the forest.
    [Show full text]
  • Kwame Nkrumah Univesity of Science and Technology Kumasi- Ghana College of Science Department of Environmental Science Effects O
    KWAME NKRUMAH UNIVESITY OF SCIENCE AND TECHNOLOGY KUMASI- GHANA COLLEGE OF SCIENCE DEPARTMENT OF ENVIRONMENTAL SCIENCE EFFECTS OF HUMAN DISTURBANCE ON BUTTERFLY DIVERSITY AND ABUNDANCE IN ATEWA RANGE FOREST RANGE FOREST RESERVE, GHANA BY OSEI BENJAMIN ADJEI (B.Sc. ENVIRONMENTAL SCIENCE) THIS THESIS IS PRESENTED TO THE DEPARTMENT OF ENVIRONMENTAL SCIENCE IN PARTIAL FULFILMENT OF THE REQUIREMENT FOR M.Sc. OF ENVIRONMENTAL SCIENCE DECEMBER 2012 i ACKNOWLEDGEMENT First of all, I will give thanks to the Almighty God for helping go through such a study successfully. Secondly, my sincere gratitude should go to my supervisor Mr. Addo-Fordjour Patrick for his assistance and guidance given me throughout the study. God richly bless you. I will also thank my mother Agnes Dankwah Adjei, my siblings and friends. The rest goes to the staff of Bobiri Forest Reserve especially Mr. Zap. As well as the managers of FORIG Eastern region and Ashantiregion. ii DECLARATION I hereby declare that the thesis entitled: The effect of human disturbance on Butterfly diversity and abundance in Atewa Range Forest Reserve was done by me with the guidance and supervision by Mr. Addo-Fordjour Patrick. Exceptions are the sections where previous knowledge is stated in which references have been duly provided. ………………………….. Date…………………… Osei Benjamin Adjei (Student) …………………………… Date…………………… Mr. Addo- Fordjour Patrick (Supervisor) ……………………………. Date……………………. Name………………………………………………. (Head of Department) iii ABSTRACT In the face of increasing human disturbance in tropical forests, it is important to understand how disturbance influence species assemblages. Though butterflies respond dramatically to disturbance, only a limited number of studies have examined the effects of varying levels of disturbance intensities on butterfly diversity and abundance.
    [Show full text]
  • The Biodiversity of Atewa Forest
    The Biodiversity of Atewa Forest Research Report The Biodiversity of Atewa Forest Research Report January 2019 Authors: Jeremy Lindsell1, Ransford Agyei2, Daryl Bosu2, Jan Decher3, William Hawthorne4, Cicely Marshall5, Caleb Ofori-Boateng6 & Mark-Oliver Rödel7 1 A Rocha International, David Attenborough Building, Pembroke St, Cambridge CB2 3QZ, UK 2 A Rocha Ghana, P.O. Box KN 3480, Kaneshie, Accra, Ghana 3 Zoologisches Forschungsmuseum A. Koenig (ZFMK), Adenauerallee 160, D-53113 Bonn, Germany 4 Department of Plant Sciences, University of Oxford, South Parks Road, Oxford OX1 3RB, UK 5 Department ofPlant Sciences, University ofCambridge,Cambridge, CB2 3EA, UK 6 CSIR-Forestry Research Institute of Ghana, Kumasi, Ghana and Herp Conservation Ghana, Ghana 7 Museum für Naturkunde, Berlin, Leibniz Institute for Evolution and Biodiversity Science, Invalidenstr. 43, 10115 Berlin, Germany Cover images: Atewa Forest tree with epiphytes by Jeremy Lindsell and Blue-moustached Bee-eater Merops mentalis by David Monticelli. Contents Summary...................................................................................................................................................................... 3 Introduction.................................................................................................................................................................. 5 Recent history of Atewa Forest................................................................................................................................... 9 Current threats
    [Show full text]
  • Nationally Threatened Species for Uganda
    Nationally Threatened Species for Uganda National Red List for Uganda for the following Taxa: Mammals, Birds, Reptiles, Amphibians, Butterflies, Dragonflies and Vascular Plants JANUARY 2016 1 ACKNOWLEDGEMENTS The research team and authors of the Uganda Redlist comprised of Sarah Prinsloo, Dr AJ Plumptre and Sam Ayebare of the Wildlife Conservation Society, together with the taxonomic specialists Dr Robert Kityo, Dr Mathias Behangana, Dr Perpetra Akite, Hamlet Mugabe, and Ben Kirunda and Dr Viola Clausnitzer. The Uganda Redlist has been a collaboration beween many individuals and institutions and these have been detailed in the relevant sections, or within the three workshop reports attached in the annexes. We would like to thank all these contributors, especially the Government of Uganda through its officers from Ugandan Wildlife Authority and National Environment Management Authority who have assisted the process. The Wildlife Conservation Society would like to make a special acknowledgement of Tullow Uganda Oil Pty, who in the face of limited biodiversity knowledge in the country, and specifically in their area of operation in the Albertine Graben, agreed to fund the research and production of the Uganda Redlist and this report on the Nationally Threatened Species of Uganda. 2 TABLE OF CONTENTS PREAMBLE .......................................................................................................................................... 4 BACKGROUND ....................................................................................................................................
    [Show full text]
  • The Ghana Butterfly Fauna and Its Contribution to the Objectives of the Protected Areas System
    WILDLIFE DIVISION (FORESTRY COMMISSION) REPUBLIC OF GHANA Wildlife Division Support Project (WDSP) The Ghana Butterfly Fauna and its Contribution to the Objectives of the Protected Areas System by Torben B Larsen (WDSP Report No. 63) March 2006 In collaboration with: TABLE OF CONTENTS EXECUTIVE SUMMARY ………………………………..….. 8 ACKNOWLEDGEMENTS …………………………………… 12 1. INTRODUCTION ………………………………………… 13 1.1 Background to the present mission …………………………. 13 1.2 Objectives of the present mission …………………………… 14 2. BUTTERFLIES AS INDICATOR SPECIES ……………. 16 2.1 Butterflies as indicators for overall biodiversity …………… 16 2.2 Butterflies as indicators of comparative diversity ………….. 17 2.3 Butterflies as ecological indicator species …………………. 18 2.4 Butterflies as biogeographical indicator species …………… 19 2.5 Butterflies as conservation indicators ……………………… 19 2.6 Butterflies as indicators of extinction ……………………… 20 2.7 Discussion …………………………………………………. 22 3. BUTTERFLIES OF THE PROTECTED AREAS ………. 23 3.1 Material and methods ……………………………………… 23 3.2 Ghana butterflies and the main ecological zones ………….. 24 3.3 Coverage of butterflies by the protected areas system …….. 26 3.3.1 Overall butterfly coverage by the protected areas system …… 26 3.3.2 Coverage of the endemic species west of Dahomey Gap .…… 28 3.3.3 Specific comments on the Ghana subregion endemics ….…… 31 3.3.4 Coverage of butterflies by the protected areas system in different ecological zones ……………………….………… 33 3.3.4.a Forest zone (excluding Volta Region) ……………… 33 3.3.4.b Volta Region …………………….………………….. 36 3.3.4.c Savannah zone ……………………….……………… 38 i) Forest-savannah transition ……………………… 39 ii) Guinea Savannah …………..…………………… 41 iii) Sudan Savannah ………………………………... 43 3.3.4.d Ubiquitous species …………………………………… 47 3.4 Recapitulation of coverage by the protected areas system ….
    [Show full text]
  • Diversity and Habitat Preferences of Anurans in The
    University of Ghana http://ugspace.ug.edu.gh DIVERSITY AND HABITAT PREFERENCES OF ANURANS IN THE ATEWA RANGE FOREST RESERVE, EASTERN REGION, GHANA BY HAMDIA MAHAMA (10395800) THIS THESIS IS SUBMITTED TO UNIVERSITY OF GHANA, LEGON IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE AWARD OF MPHIL ZOOLOGY DEGREE JULY, 2014 University of Ghana http://ugspace.ug.edu.gh DECLARATION I declare that this thesis submitted for the award of Master Philosophy in Zoology, is the result of research undertaken by me. References to other people‟s work have been duly acknowledged. ………………………………………………… HAMDIA MAHAMA (Student) Date………………………………… ………………………………………………… PROFESSOR DANIEL K. ATTUQUAYEFIO (Principal Supervisor) Date……………………………….. …………………………………………………….. DR. JAMES K. ADOMAKO (Supervisor) Date……………………………….. i University of Ghana http://ugspace.ug.edu.gh ACKNOWLEDGEMENTS I thank God for my life and for giving me strength to complete this work successfully. I would like to thank my Supervisors, Prof. Daniel K. Attuquayefio and Dr. James K. Adomako for their patience, guidance and immense support during the research. I am also grateful to my sister, Ruby and husband, Emmanuel J. Wumbeidow for their continual prayers and motivation. I am extremely grateful to Dr. Caleb Ofori-Boateng and Mr. Gilbert Adum for their time, dedication, useful criticisms, technical support and encouragement in amphibian studies. Philip Amankwaa and Jojo Ibrahim, trained specialists in amphibian identification and field techniques, were tremendously hardworking and supported me throughout the field work. I am indebted to Mr. Charles Gyekye, of the Begoro District Forestry Office, Forestry Commission, for assisting with the identification of trees, support and also providing a forest guard during the entire research period.
    [Show full text]
  • Los Muscoidea De La Región Etiópica
    UNIVERSIDAD COMPLUTENSE DE MADRID FACULTAD DE CIENCIAS Sección de Biológicas. TESIS DOCTORAL Los muscoidea de la región etiópica MEMORIA PARA OPTAR AL GRADO DE DOCTOR PRESENTADA POR Pilar Ortega Nuñez Madrid, 2015 © Pilar Ortega Nuñez, 1971 5 9 5 . r R T UNIVEI - ! ' ü COMPLUTENSE DE MADRID FACULTAD DE CIENCIAS LOS MUSCOIDEA DE LA REGION ETIOPICA MEMORIA PRESENTADA PARA ASPIRAR AL GRADO DE DOCTOR EN CIENCIAS Por Pilar Ortega Nunez PRIMERA PARTE P. 25.0 1 3 Dirigida por el Prof. Dr. S.V. PERIS TORRES ÜNIVERSIDAD COMPLUTENSE FACULTAD DE CIENCIAS DE MADRID LOS MUSCOIDEA DE LA REGION ETIOPICA MEMORIA PRESENTADA PARA ASPIRAR AL GRADO DE DOCTOR EN CIENCIAS Por Pilar Ortega Nûnez Dirigida por el Prof. Dr. S.V.PERIS TORRES Mi sincere agradecimiento: A1 Profesor Dr. S.V. PERIS TORRES, Catedràtico de Zoologla (Artrdpodos) de la Facultad de Ciencias de la Universidad Com- plutense, bajo cuya direccidn ha side efectuado este trabajo, que puso a mi disposicidn una coleccidn de 3500 ejemplares de Muscoidea etiôpicos del Museo del Africa Central de Tervuren (Bélgica), en depdsito en su câtedra, y cuyos consejos, obser- vaciones y directrices han hecho posible su realizacidn. Al Profesor P. MAYOR ZARAGOZA, Jefe del Departamento de Bio- qulmica y Biologia Molecular de la Universidad Autdnoma de Ma­ drid, que en todo memento me ha otorgado las mâximas facilida- des para que el presente trabajo pudiera llevarse a cabo. A la Dra. R. MANSO MARTINEZ, Profesor Agregado del mismo depar- tamento, siempre dispuesta a facilitarme mi trabajo. A1 Dr. P. BASILEWSKY, Jefe del Institute de Entomologia del Museo del Africa Central de Tervuren (Bélgica), por su excelen- te acogida durante mi asistencia a dicho museo, y por haber pues- to a mi disposicidn todo el material relacionado con mi traba­ jo que habfa en el mencionado centre, y que compléta el deposi- tado en la Càtedra de Artrdpodos.
    [Show full text]