Atlas of Yorkshire Coleoptera (VCs 61-65) Part 10 – Cleroidea and Cucujoidea

Introduction

This section of the atlas deals with the Superfamilies Cleroidea and Cucujoidea, a total of 464 species, of which there are 315 recorded in Yorkshire.

Each species in the database is considered and in each case a distribution map representing records on the database (at 1 April 2019) is presented. The number of records on the database for each species is given in the account in the form (a,b,c,d,e) where 'a' to 'e' are the number of records from VC61 to VC65 respectively. These figures include undated records (see comment on undated records in the paragraph below on mapping). As a recorder, I shall continue to use the vice-county recording system, as the county is thereby divided up into manageable, roughly equal, areas for recording purposes. For an explanation of the vice-county recording system, under a system devised in Watson (1883) and subsequently documented by Dandy (1969), Britain was divided into convenient recording areas ("vice-counties"). Thus Yorkshire was divided into vice-counties numbered 61 to 65 inclusive, and notwithstanding fairly recent county boundary reorganisations and changes, the vice-county system remains a constant and convenient one for recording purposes; in the text, reference to “Yorkshire” implies VC61 to VC65 ignoring modern boundary changes. For some species there are many records, and for others only one or two. In cases where there are five species or less full details of the known records are given. Many common species have quite a high proportion of recent records. This is because the older county recorders tended not to record species they regarded as "common" quite as assiduously as we do today. “Recent” records are those considered to be later than 1 January 2000 – this is also the break point used in the production of the distribution maps. (In earlier parts of this Atlas, the break point for ‘recent’ and otherwise was 1 January 1990). Habitat A brief statement on habitat is included where known and relevant. Taxon status Inclusion of a statement on the status of species presents serious problems in that those designations in Recorder 6 are often out of date. The species status designations given in the species accounts below I believe are the current ones when this part of the Atlas was prepared. The situation with species statuses is a fluid one, with Natural England now producing reports on various groups of Coleoptera with statuses according to standard IUCN criteria. But some groups retain earlier defined statuses as listed in Hyman and Parsons (1992 and 1994)

The following statuses are used in this part of the Atlas:

Nationally Notable A – known to occur in <30 10km squares (Hyman and Parsons, 1992) Nationally Notable B – known to occur in 31-100 10km squares (Hyman and Parsons, 1992) Nationally Scarce – known from 16-100 10km squares RDB1 – Endangered (Hyman and Parsons, 1992) RDB3 – Rare (Hyman and Parsons, 1992) RDBI – Indeterminate – endangered/vulnerable/rare but with insufficient information available (Hyman and Parsons, 1992) RDBK – Insufficiently known (Hyman and Parsons, 1992)

Phenology Details are given for the range of months in the year for which there are records, with or without a peak month of occurrence, where the number and detail of records are sufficient to give a meaningful statement. Also bear in mind that the phenology data from the database may be skewed by the fact that coleopterists tend to be out collecting and recording in the better weather of April to October.

Maps Distribution maps have been produced using Dr Alan Morton’s DMAP software (version 7.4b of 2013). It may be worthwhile to note that as DMAP is asked to map pre- and post-2000 records in this application, any undated records will not be mapped. While efforts have been made to establish dates of records, some are difficult to ascertain, and I do not want to make guesses which may be wrong. In addition to this, records flagged in the database as unconfirmed will also

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 1 | P a g e not be mapped. The YNU database possesses a few peat sub-fossil records. These are not considered for inclusion in the Atlas.

Nomenclature The YNU database is presently held in a copy of Recorder 6, which uses the Coleoptera checklist based on Duff (2012) and this new list incorporates a very large number of taxon name changes since the checklist of Pope (1977). As this section of the Atlas was being prepared, a third edition of the British Checklist (Duff, 2018) was published, and has not yet been fully incorporated into Recorder 6. Occasionally I receive records using “noddy names” (see Cooter, (1999) and Key (1999) for an excellent treatment of this subject), e.g. “Snail Hunter”, “Crucifix Beetle” and “Heath-glory ground beetle” (Cychrus caraboides, Panagaeus crux-major and Carabus nitens respectively). Such inventions seem to have originated somewhere within Natural England (previously English Nature) or JNCC. For the purposes of serious science these terms are, in my opinion, either vague, ridiculous, confusing or unhelpful. I do not accept records using such names only, unless accompanied by specimens or other convincing evidence.

For the purposes of this section of the Atlas the Duff (2012) checklist will be used as the basis of the nomenclature and arrangement of species.

Quality of the records A regrettable aspect of many records within the YNU database is the lack of detail accompanying the record, especially of accurate grid references, lack of pertinent comments as to the micro-habitat of the capture, its association with plants/animals, and sometimes, in the case of some modern records, a lack of care in determination, and the recording of the sex of the specimen. Old records, although sound, tend not to give much or indeed any of this detail. Old records on the database so rarely give habitat details but sometimes interpretation of the site name can give clues to the habitat type. Grid references could not be applied before 1930 anyway, and grid references do not figure in the paper archive of the YNU Coleoptera database until the late 1970s. Apparently it was not considered necessary to give that level of accuracy. Entering records into Recorder 6 requires a grid reference, and with older records I have added an accurate grid reference where I believe this has been possible. If the site name is vague I have used a 10km reference, and where a 1km reference is deemed accurate enough I have included that. There can be little excuse with modern records not to include some accurate comment with the record, and at least a six-figure grid reference.

Taxonomic knowledge and expertise have increased greatly since the days of Walsh, Bayford and Stainforth. Much good literature now exists for the taxonomist in the form of dichotomous keys, and when identifying specimens, especially with critical species pairs or groups, too much reliance on the outcome of keys can be placed on a determination, without considering the habitat in which the specimen was taken, or the host plant/animal. Atlases of the national distributions of various orders of insect are now becoming available and should always be consulted. These often give a guide to the likelihood or otherwise of a determination, especially in scarce taxa. Museum collections containing correctly determined and labelled specimens are invaluable in accurate recording; museum collections are there for our use and should be consulted. If a species is ‘new’ to a vice-county or is very significant in some other respect, a second opinion should always be sought or an acknowledged specialist in the group consulted. A voucher should always be retained by the collector if possible. The recorder can only deal with the information he is given, and often has to beware of identifications made on photographic evidence only, and identifications of ‘difficult’ species by non-specialists. It takes many years to become competent in many areas of entomology, and the recorder often has to rely on the expertise of co-workers, especially when records covering large Orders are being documented.

I have adopted a policy in my county recording whereby any received record of a species afforded Notable B or above, especially if a member of a ‘difficult’ beetle group, should be confirmed by a specialist in that group before the record is entered into the database as a correct record. Ultimately, the responsibility for verification of records on the YNU database rests with me. With modern records received from reputable coleopterists there is little problem. Some older records requiring vouchers for confirmation have been retained in the database but flagged as requiring confirmation.

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Contributors (alphabetically by initials as appearing in the text, not alphabetically by surname)

ABD – Tony Drane MB – Max Barclay AG – Andy Godfrey MBa – Maurice Barnes AL – Alan Lazenby MH – Martin Hammond AME – A M Easton MLD – Mike Denton AP – Adam Parker MLT – M L Thompson AS – Arthur Smith MM – Michael Majerus BC – Barry Constantine NH – Norman Heal BE – Brian Eversham NW – Nikki Whitehouse BK – Basil Kitchen PBM – P B Mander CES – Charles Ernest Stott PH – Peter Hodge CJ – Colin Johnson PHa – P Harwood CWC – C W Coombs PK – Peter Kendall DH – David Heaver PS – Dr Peter Skidmore DL – Dick Loxton PW – Paul Whitehead DM – David Maude RAT – R A Taylor (Rev.) DW – Derek Whiteley RBA – Robert Angus EAW – Edward Alexander Waterhouse RGB – Roger Booth ECH – Ernest Charles Horrell RH – R Hawley EFG – E F Gilmour RJM – Bob Marsh EGB – E G Bayford RL – R Lawson EJS – Eric Smith RM – Richard Moore EWA – Ted Aubrook RSK – Roger Key FEK – Frank Kenington RRUK – R R Uhtoff-Kaufmann GBW – George Beckworth Walsh TS – T Stainforth GEW – G E Woodroffe TVW – Thomas Vernon Wollaston HB – Harry Britten (jnr.) TW – Thomas Wilkinson HF – Hilda Flint WAE – Bill Ely HHC – Herbert Henry Corbett WB – William Bunting HM – Howard Mendel WCH – W C Hey JDC – John Coldwell WDH – Walter Douglas Hincks JHF – John Flint WES – W E Sharp JJW – unknown WJF – William John Fordham JWC – J W Carter WJL – W J Lee LA – Len Auckland WJS – W J Sanders LB – Leslie Barringer WRD – Bill Dolling

Superfamily CLEROIDEA Latreille, 1802..

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Family PHLOIOPHILIDAE Kiesenwetter, 1863 FFamily48. Family PHLOIOPHILIDAE Kiesenwetter, 1863 PHLOIOPHILUS Stephens, 1830 edwardsii Stephens, 1830

Phloiophilus edwardsii Stephens, 1830

3 WATSONIAN (1,5,0,1,0 ) Nationally Scarce. Very scarce in Yorkshire with only 7 records 2 YORKSHIRE in the database. Recorded from the 1 fungus Peniphora on old oaks in 0 NY NZ NZ OV SD SE SE TA parklands such as Duncombe Park, 9 Studley Park and Castle Howard, which 8 provide typical habitats. 7 6 5 4 3 2 1

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FFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFFF F Family TROGOSSITIDAE Latreille, 1802

Subfamily PELTINAE Latreille, 1806

OSTOMA Laicharting, 1781 ferrugineum (Linnaeus, 1758)

NO RECORDS

THYMALUS Latreille, 1802

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Thymalus limbatus (Fabricius, 1787)

3 WATSONIAN (0,5,3,6,0) Nationally Scarce. Develops in 2 YORKSHIRE the loose bark and rotted heatwood of, particularly, oak in old parkland 1 situations. Waterhouse (1871) recorded 0 NY NZ NZ OV SD SE SE TA the species in beech. In 1970 a specimen 9 was recorded as a subfossil in a bog oak 8 at SE5615. 7 6 5 4 3 2 1

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Subfamily LOPHOCATERINAE Crowson, 1964

LOPHOCATERES Olliff, 1883 pusillus (Klug, 1832)

Lophocateres pusillus (Klug, 1832)

3 WATSONIAN (0,0,1,0,0) There is a single record of this 2 YORKSHIRE pest of stored products: Doncaster (SE50), 3/10/1978, PS, specimens 1 reported from a town-centre restaurant 0 NY NZ NZ OV SD SE SE TA store room. 9 8 7 6 5 4 3 2 1

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Subfamily TROGOSSITINAE Latreille, 1802

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TENEBROIDES Piller & Mitterpacher, 1783 mauritanicus (Linnaeus, 1758)

Tenebroides mauritanicus (Linnaeus, 1758)

3 WATSONIAN (1,2,4,0,0) A synanthropic introduction 2 YORKSHIRE and a pest of stored products, in mouldy cereals etc. In Yorkshire recorded from 1 bakeries and restaurants. Interestingly 0 NY NZ NZ OV SD SE SE TA HB recorded the species in bamboo 9 canes washed up on shore at Whitby 8 NZ8911 (see Nat 1938:84). 7 6 5 4 3 2 1

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NEMOZOMA Latreille, 1804 elongatum (Linnaeus, 1761)

NO RECORDS Nn50. Family CLERIDAE Latreille, 1802 553 Family CLERIDAE Latreille, 1802

Subfamily THANEROCLERINAE Chapin, 1924

THANEROCLERUS Lefebvre, 1838 buqueti (Lefebvre, 1835)

NO RECORDS

Subfamily TILLINAE Fischer von Waldheim, 1813

TILLUS Olivier, 1790

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elongatus (Linnaeus, 1758)

Tillus elongatus (Linnaeus, 1758)

3 WATSONIAN (0,2,0,0,0) Nationally Scarce. Very rare 2 YORKSHIRE in Yorkshire with only two records in the YNU database: Duncombe Park 1 (SE6082), 26/6/93, MLD,PK, beaten from 0 NY NZ NZ OV SD SE SE TA old oaks; York (SE6254), 20/6/1951, AS, 9 in house. This species is a predator of 8 anobiid larvae in old deciduous trees. 7 6 5 4 3 2 1

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TILLOIDEA Laporte, 1833 unifasciata (Fabricius, 1787)

NO RECORDS

Subfamily CLERINAE Latreille, 1802

OPILO Latreille, 1802

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mollis (Linnaeus, 1758)

Opilo mollis (Linnaeus, 1758)

3 WATSONIAN (0,1,0,0,0) Natonally Scarce. There is a single record of this species in the YNU 2 YORKSHIRE database: Raincliffe Woods (SE9988), 1 1951, GBW, under bark of spruce. The 0 NY NZ NZ OV SD SE SE TA larvae predate upon anobiid larvae in 9 old timber. 8 7 6 5 4 3 2 1

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THANASIMUS Latreille, 1806 femoralis (Zetterstedt, 1838)

NO RECORDS formicarius (Linnaeus, 1758)

Thanasimus formicarius (Linnaeus, 1758)

3 WATSONIAN (10,9,10,3,0) Reported from conifers, 2 YORKSHIRE birch, ash and oak . Not often recorded but widely distributed in a more or less 1 central band in the county. Adults and 0 NY NZ NZ OV SD SE SE TA larvae reported as predatory on 9 scolytids in decaying trees, but also 8 recorded from a garden and inside a 7 house. 6 5 4 3 2 1

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TRICHODES Herbst, 1792 alvearius (Fabricius, 1792)

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NO RECORDS apiarius (Linnaeus, 1758)

NO RECORDS

Subfamily TARSOSTENINAE Jacquelin du Val, 1860

PARATILLUS Gorham, 1878 carus (Newman, 1840)

NO RECORDS

TARSOSTENUS Spinola, 1844 univittatus (Rossi, 1792)

Tarsostenus univ ittatus (Rossi, 1792)

3 WATSONIAN (0,0,1,0,0) There is a single record of this 2 YORKSHIRE species, predatory on Lyctus larvae: Milnsbridge (SE1016), 21/7/1948, EFG, 1 imported(?) (see Nat 1949:151). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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Subfamily KORYNETINAE Laporte, 1836

KORYNETES Herbst, 1792

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caeruleus (De Geer, 1775)

Korynetes caeruleus (De Geer, 1775)

3 WATSONIAN (2,1,2,1,0) Nationally Scarce. There are six records of this species in the YNU 2 YORKSHIRE database. Predatory on anobiids in old 1 timber in and outside buildngs. We have 0 NY NZ NZ OV SD SE SE TA records from inside a house, from 9 hawthorn and in flood refuse. Widely 8 but centrally distrbuted in the county. 7 6 5 4 3 2 1

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NECROBIA Olivier, 1795 ruficollis (Fabricius, 1775)

Necrobia ruficollis (Fabricius, 1775)

3 WATSONIAN (3,3,3,2,0) This necrophilic species is 2 YORKSHIRE rather scarce with only one recent record: Hollym Carrs Reserve (TA3224), 1 22/4/2004, FEK, a specimen in a deer 0 NY NZ NZ OV SD SE SE TA skull. Reported from old bones and 9 carrion. 8 7 6 5 4 3 2 1

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rufipes (De Geer, 1775)

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Necrobia rufipes (De Geer, 1775)

3 WATSONIAN (8,5,11,1,0) Another carrion feeder, there are very few recent records 2 YORKSHIRE probably due to changing farming and 1 public health practices. Recorded in 0 NY NZ NZ OV SD SE SE TA Yorkshire from the carcasses of sheep, 9 birds, hides and Roe Deer. Widely 8 reported except in the north and west. 7 6 5 4 3 2 1

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violacea (Linnaeus, 1758)

Necrobia v iolacea (Linnaeus, 1758)

3 WATSONIAN (15,4,17,5,2) The most widespread and 2 YORKSHIRE the most commonly met-with member of the genus. In carrion generally 1 including that of deer, sheep, badger, 0 NY NZ NZ OV SD SE SE TA rabbit, crow, Muscovy Duck, in a dead 9 porpoise and other old bones on the 8 shoreline, and one anomalous record 7 from the kitchen of a suburban house. The proportion of recent records hints 6 that the species is maintaining healthy 5 populations in the countryside as 4 opposed to the apparent declines in 3 other members of the genus 2 1

SD SE SE TA 0 SJ SK SK TF 9 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 13/8/2018 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 51. Family DASYTIDAE Laporte, 1840 555

Family DASYTIDAE Laporte, 1840

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Subfamily RHADALINAE LeConte, 1861

APLOCNEMUS Stephens, 1830 impressus (Marsham, 1802)

Aplocnemus impressus (Marsham, 1802)

3 WATSONIAN (0,1,1,0,0) Nationally Notable B. There is only one historical record of this 2 YORKSHIRE species in Yorkshire: Newgate Bank 1 (SE5794), 5/1937, GBW, 2 exx from pine 0 NY NZ NZ OV SD SE SE TA (see Nat 1938:96). The Thorne Moor 9 record (SE7117) is a sub-fossil specimen 8 from the peat formation, excavation dated 1993 (NW). The species is a 7 scolytid predator in coniferous and 6 broadleaved trees. 5 4 3 2 1

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nigricornis (Fabricius, 1792)

Aplocnemus nigricornis (Fabricius, 1792)

3 WATSONIAN (0,6,0,1,0) Nationally Notable A. Very 2 YORKSHIRE scarce in Yorkshire. Recorded from under the bark and on the foliage of conifers. 1

NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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Subfamily DASYTINAE Laporte, 1840

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DASYTES Paykull, 1799 aeratus Stephens, 1830

Dasytes aeratus Stephens, 1830

3 WATSONIAN (6,17,11,11,3) Widespread in the 2 YORKSHIRE eastern half of the county and mainly recorded from hawthorn blossom, oak 1 and sycamore. The larvae are 0 NY NZ NZ OV SD SE SE TA carnivorous and saproxylic in live and 9 dead wood, particularly oak (Alexander, 8 2002) 7 6 5 4 3 2 1

SD SE SE TA 0 SJ SK SK TF 9 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 13/8/2018 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 cyaneus (Fabricius, 1775)

NO RECORDS niger (Linnaeus, 1761)

NO RECORDS plumbeus (Müller, O.F., 1776)

Dasytes plumbeus (Müller, O.F., 1776)

3 WATSONIAN (0,2,7,0,0) Nationally Scarce. Very rare 2 YORKSHIRE in the county with no recent records. Usually found by beating shrubs and 1 sweeping low plants. (vi-ix). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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NO RECORDS

PSILOTHRIX Küster, 1850 viridicoeruleus (Geoffroy in Fourcroy, 1785)

Psilothrix v iridicoeruleus (Geoffroy in Fourcroy, 1785)

3 WATSONIAN (0,0,1,0,0) There is a single record of this species in the YNU database: 2 YORKSHIRE Crookesmoor (SK3387), 14/4/2006, EJS, 1 probably an import – found on a bunch 0 NY NZ NZ OV SD SE SE TA of flowers, provenance unknown, in a 9 house. 8 7 6 5 4 3 2 1

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DOLICHOSOMA Stephens, 1830 lineare (Rossi, 1794)

Dolichosoma lineare (Rossi, 1794)

3 WATSONIAN (5,0,0,0,0) Restricted to the Spurn, 2 YORKSHIRE Welwick and Patrington areas and at the northern limit of its range in Britain. A 1 species of saltmarshes, tidal creeks and 0 NY NZ NZ OV SD SE SE TA coastal grassland. The larvae are 9 predatory, adults are usually swept from 8 vegetation. 7 6 5 4 3 2 1

SD SE SE TA 0 SJ SK SK TF 9 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 13/8/2018 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 52. Family MALACHIIDAE Fleming, 1821 557

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COLOTES Erichson, 1840 punctatus (Erichson, 1840)

NO RECORDS

EBAEUS Erichson, 1840 pedicularius (Linnaeus, 1758)

NO RECORDS but see below under “Species removed from Yorkshire list”

HYPEBAEUS Kiesenwetter, 1863 flavipes (Fabricius, 1787)

NO RECORDS

AXINOTARSUS Motschulsky, 1853 marginalis (Laporte, 1840)

NO RECORDS pulicarius (Fabricius, 1777)

NO RECORDS ruficollis (Olivier, 1790)

NO RECORDS

TROGLOPS Erichson, 1840 cephalotes (Olivier, 1790)

NO RECORDS

CORDYLEPHERUS Evers, 1985

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viridis (Fabricius, 1787)

Cordylepherus v iridis (Fabricius, 1787)

3 WATSONIAN (1,0,1,3,0) Scarce in Yorkshire with five records, only one of these recent: 2 YORKSHIRE Escrick (SE6343), 25/5/1917, WJF; 1 Thorne Moor (SE7115), 1983, BE; Selby 0 NY NZ NZ OV SD SE SE TA (SE63), 1884, EGB; Pannal Ash (SE3052), 9 vi/1938, RRUK, one specimen in field; 8 Studley Royal (SE2769), 5/6/2016, AG det MLD. Recorded from grassland, this 7 elusive species develops in the stems of 6 various plants. 5 4 3 2 1

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MALACHIUS Fabricius, 1775 aeneus (Linnaeus, 1758)

NO RECORDS bipustulatus (Linnaeus, 1758)

Malachius bipustulatus (Linnaeus, 1758)

3 WATSONIAN (62,41,274,42,10) Widespread mainly in 2 YORKSHIRE a central band in the county, away from coasts and the high land to the north 1 and west, and very common in 0 NY NZ NZ OV SD SE SE TA grassland, the adults seen feeding on 9 pollen at flowers and by general 8 sweeping. The larvae are said to be 7 predatory on other arthropods under bark. (iv-ix, peak vi). 6 5 4 3 2 1

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CLANOPTILUS Motschulsky, 1854 Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 16 | P a g e

barnevillei (Puton, 1865)

NO RECORDS marginellus (Olivier, 1790)

NO RECORDS strangulatus (Abeille de Perrin, 1885)

NO RECORDS

SPHINGINUS Mulsant & Rey, 1867 lobatus (Olivier, 1790)

NO RECORDS

CERAPHELES Mulsant & Rey, 1867 terminatus (Ménétries, 1832)

NO RECORDS

ANTHOCOMUS Erichson, 1840 fasciatus (Linnaeus, 1758)

NO RECORDS rufus (Herbst, 1784)

Anthocomus rufus (Herbst, 1784)

3 WATSONIAN (11,0,79,1,0) Widespread in the southern half of the county, this fen 2 YORKSHIRE species was very rare in Yorkshire before 1 1990 but has since become much more 0 NY NZ NZ OV SD SE SE TA evident especially post-2010, even in 9 recently-established marshy areas (e.g. 8 areas of mining subsidence). (v-x, peak ix). 7 6 5 4 3 2 1

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Superfamily CUCUJOIDEA Latreille, 1802 Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 17 | P a g e

53. Family SPHINDIDAE Jacquelin du Val, 1860 Family SPHINDIDAE Jacquelin du Val, 1860

SPHINDUS Dejean, 1821 dubius (Gyllenhal, 1808)

Sphindus dubius (Gyllenhal, 1808)

3 WATSONIAN (0,7,1,0,0) Nationally Notable B. Scarce 2 YORKSHIRE in Yorkshire with only eight records from widely separated localities. Recorded 1 from puffball fungi on horse chestnut 0 NY NZ NZ OV SD SE SE TA logs, in unspecified fungi on stumps of 9 felled oaks and in a flight interception 8 trap. 7 6 5 4 3 2 1

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ASPIDIPHORUS Dejean, 1821 orbiculatus (Gyllenhal, 1808)

Aspidiphorus orbiculatus (Gyllenhal, 1808)

3 WATSONIAN (2,10,18,2,0) Associated with fungi including brackets on Salix species, 2 YORKSHIRE under bark of decaying deciduous wood, 1 and in flight interception traps. Widely 0 NY NZ NZ OV SD SE SE TA distributed but not common. (v-ix). 9 8 7 6 5 4 3 2 1

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Family KATERETIDAE Kirby, 1837

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 18 | P a g e

KATERETES Herbst, 1793 pedicularius (Linnaeus, 1758)

Kateretes pedicularius (Linnaeus, 1758)

3 WATSONIAN (1,3,6,7,0). A wetland species, recorded mainly from Carex spp. in marshes, at 2 YORKSHIRE the margins of reservoirs and by drains 1 in wet woodland. Widely but thinly 0 NY NZ NZ OV SD SE SE TA distributed and rather scarce with only 9 two recent records. (iv-ix). 8 7 6 5 4 3 2 1

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pusillus (Thunberg, 1794)

Kateretes pusillus (Thunberg, 1794)

3 WATSONIAN (18,12,1,6,2) Widely distributed, more common than pedicularius and also on 2 YORKSHIRE Carex spp. in similar habitats. (iii-viii, 1 peak v). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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rufilabris (Latreille, 1807) Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 19 | P a g e

Kateretes rufilabris (Latreille, 1807)

3 WATSONIAN (27,12,37,15,7) The most widely 2 YORKSHIRE distributed and the most frequently recorded of our Kateretes. In fens, 1 around ponds and reservoirs, and in 0 NY NZ NZ OV SD SE SE TA litter under willows and alders in wet 9 places. Recorded from Juncus, Carex and 8 Sparganium spp. 7 6 5 4 3 2 1

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BRACHYPTERUS Kugelann, 1794 glaber (Newman, 1834)

Brachypterus glaber (Stephens, 1832)

3 WATSONIAN (68,34,307,35,13) Widespread and very 2 YORKSHIRE common on Urtica dioica in a wide variety of habitats, particularly disturbed 1 ground, verges and roadsides. 0 NY NZ NZ OV SD SE SE TA (v-x, peak vi). 9 8 7 6 5 4 3 2 1

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urticae (Fabricius, 1792) Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 20 | P a g e

Brachypterus urticae (Fabricius, 1792)

3 WATSONIAN (78,45,386,47,21) Distribution and habitats as the previous species, the two 2 YORKSHIRE often being found together. A little more 1 frequently recorded than glaber. (v-x, 0 NY NZ NZ OV SD SE SE TA peak vi). 9 8 7 6 5 4 3 2 1

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BRACHYPTEROLUS Grouvelle, 1913 antirrhini (Murray, 1864)

NO RECORDS linariae (Stephens, 1830)

Brachypterolus linariae (Stephens, 1830)

3 WATSONIAN (0,0,5,0,0) Rare in Yorkshire, on 2 YORKSHIRE toadflaxes, with only five known records: Thorne (SE61), 3/8/1946, WDH; 1 Denaby Ings (SE4901), 8/5/1983, RJM 0 NY NZ NZ OV SD SE SE TA det CJ; Treeton Dike (SK4386), 9 14/8/1990, WAE det CJ; Walton 8 (SE3618), 11/8/1999, MLD det CJ; 7 Sharlston Common (SE3720), 16/6/2010, MLD. (v-viii). 6 5 4 3 2 1

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pulicarius (Linnaeus, 1758)

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 21 | P a g e

Brachypterolus pulicarius (Linnaeus, 1758)

3 WATSONIAN (4,0,28,2,1) Widespread but mainly in 2 YORKSHIRE the southern half of the county. On Linaria spp. as with B. linariae but 1 relatively much more frequent. Few 0 NY NZ NZ OV SD SE SE TA recent records. (v-viii). 9 8 7 6 5 4 3 2 1

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vestitus (Kiesenwetter, 1850)

Brachypterolus v estitus (Kiesenwetter, 1850)

3 WATSONIAN (0,0,2,0,0) GBES – Established. Rare with only two Yorkshire records: Scabba 2 YORKSHIRE Wood (SE5201), 18/5/2005, RJM, 1 general sweeping disturbed ground at 0 NY NZ NZ OV SD SE SE TA woodland edge; Melton Brand (SE5203), 9 29/6/2008, RJM, on Malva sylvestris on 8 roadside verge adjacent gardens. Usually associated with Antirrhinum. 7 6 5 4 3 2 1

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55. Family NITIDULIDAE Latreille, 1802

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 22 | P a g e

Family NITIDULIDAE Latreille, 1802

Subfamily CARPOPHILINAE Erichson, 1842

Tribe CARPOPHILINAE Erichson, 1842

UROPHORUS Murray, 1864 humeralis (Fabricius, 1798)

NO RECORDS

CARPOPHILUS Stephens, 1829 dimidiatus (Fabricius, 1792)

Carpophilus dimidiatus (Fabricius, 1792)

3 WATSONIAN (1,0,3,0,0) A known pest of stored 2 YORKSHIRE products but may establish in the wild, with four known county records: 1 Marfleet (TA12), c.1915, TS; Doncaster 0 NY NZ NZ OV SD SE SE TA (SE50), 8/5/1915, HHC see Nat . 9 1915:209; Eggborough Power Station 8 (SE5724), 22/6/1983, RJM, a single 7 specimen on window pane in building; Sprotbrough (SE5402), 19/10/1984, 6 RJM, in a garden compost heap. 5 4 3 2 1

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flavipes Murray, 1864

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 23 | P a g e

hemipterus (Linnaeus, 1758)

Carpophilus hemipterus (Linnaeus, 1758)

3 WATSONIAN (2,1,4,2,0) Rarely recorded in the county 2 YORKSHIRE with nine records. In dried fruit and similar stored products, and recorded 1 from shops and restaurants. All our 0 NY NZ NZ OV SD SE SE TA records are certainly imports. 9 8 7 6 5 4 3 2 1

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ligneus Murray, 1864

Carpophilus ligneus Murray, 1864

3 WATSONIAN (0,1,4,1,0) Recorded from dried fruit 2 YORKSHIRE including figs and raisins in stores and other buildings. Our six records are all 1 believed to result from imports of 0 NY NZ NZ OV SD SE SE TA commodities. There are no recent 9 records. 8 7 6 5 4 3 2 1

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maculatus Murray, 1864

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 24 | P a g e

marginellus Motschulsky, 1858

Carpophilus marginellus Motschulsky, 1858

3 WATSONIAN (8,0,29,1,1) Widespread in the southern half of the county. Recorded in the wild 2 YORKSHIRE from compost heaps, in putrefying 1 vegetation in the ground layer in wet 0 NY NZ NZ OV SD SE SE TA places, in piles of old mowings and 9 under hay bales, and probably a fungus 8 feeder. The most commonly met-with Carpophilus in our region. 7 6 5 4 3 2 1

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Carpophilus mutilatus Erichson, 1843

3 WATSONIAN (0,0,2,0,0) There are two records of this species in the YNU database: Doncaster 2 YORKSHIRE (SE50), 1915, HHC, specimens infesting 1 damaged hides at a tannery; Hatfield 0 NY NZ NZ OV SD SE SE TA Moor (SE7006), 9/9/1991, PS, in pitfall 9 traps. 8 7 6 5 4 3 2 1

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nepos Murray, 1864

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 25 | P a g e

obsoletus Erichson, 1843

Carpophilus obsoletus Erichson, 1843

3 WATSONIAN (0,0,2,0,0) Two old records exist in the YNU database: Doncaster (SE50), 1912, 2 YORKSHIRE EGB; Edlington (SK59), 1912, WES. This 1 species is a minor pest of stored cereals 0 NY NZ NZ OV SD SE SE TA but neither record gives habitat details. 9 8 7 6 5 4 3 2 1

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sexpustulatus (Fabricius, 1792)

Carpophilus sexpustulatus (Fabricius, 1792)

3 WATSONIAN (1,2,16,5,0) Widely distributed in the 2 YORKSHIRE southern half of the county. Recorded from fungi, under the bark of oak and 1 beech and in dead birds. NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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truncatus Murray, 1864

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 26 | P a g e

Tribe EPURAEINI Kirejtshuk, 1986

EPURAEA Erichson, 1843

Subgenus EPURAEA Erichson, 1843 aestiva (Linnaeus, 1758)

Epuraea (Epuraea) aestiv a (Linnaeus, 1758)

3 WATSONIAN (42,45,176,31,13) Widespread and 2 YORKSHIRE common throughout most of the county. Recorded mainly from blossoms 1 on trees and shrubs, but also in ground 0 NY NZ NZ OV SD SE SE TA litter and in debris from old bees’ nests. 9 (iv-x, peak vi). 8 7 6 5 4 3 2 1

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angustula Sturm, 1844

Epuraea (Epuraea) angustula Sturm, 1844

3 WATSONIAN (0,2,3,1,0) Nationally Notable B. A rarity in Yorkshire with only six records. A 2 YORKSHIRE known associate of Trypodendron in 1 moribund trees in old woodland. Our 0 NY NZ NZ OV SD SE SE TA records mention birch and holly as 9 habitats. 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 27 | P a g e biguttata (Thunberg, 1784)

Epuraea (Epuraea) biguttata (Thunberg, 1784)

3 WATSONIAN (15,24,100,19,7) Widespread and fairly 2 YORKSHIRE common in the county. Recorded from a wide variety of habitats: sweeping under 1 trees, at sap runs on trees, in compost 0 NY NZ NZ OV SD SE SE TA heaps, in pitfall and flight interception 9 traps, under bark and in fungi. (iv-x, 8 peak v). 7 6 5 4 3 2 1

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binotata Reitter, 1872

NO RECORDS distincta (Grimmer, 1841)

NO RECORDS fuscicollis (Stephens, 1835)

Epuraea (Epuraea) fuscicollis (Stephens, 1832)

3 WATSONIAN (0,1,0,1,0) Nationally Notable B. Very 2 YORKSHIRE rare in Yorkshire with only two records: Harewood (SE3245), 10/1921, BK det 1 JJW; Saltburn (NZ62), 9/1894, MLT, on 0 NY NZ NZ OV SD SE SE TA thistles. Both these records are very old 9 and vouchers would be desirable for 8 confirmation. Nationally the species is 7 very scarce and is recorded from sap runs on trees damaged by Cossus larvae. 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 28 | P a g e guttata (Olivier, 1811)

NO RECORDS longula Erichson, 1845

Epuraea (Epuraea) longula Erichson, 1845

3 WATSONIAN (0,1,0,1,0) Nationally Notable B. There 2 YORKSHIRE are two records on the YNU database: Grassington (SE0064), 9/1909, JWC; 1 Sandsend (NZ8613), 21/7/1934, HB (see 0 NY NZ NZ OV SD SE SE TA Nat. 1934:276). Nationally, recorded 9 from sap runs and swept from various 8 woodland plants, and, as in fuscicollis, 7 noted from trees damaged by Cossus larvae. Recorded from general sweeping 6 in woodland, at sap runand in fungis on 5 trees, in comost bins in gardens 4 3 2 1

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marseuli Reitter, 1872

Epuraea (Epuraea) marseuli Reitter, 1872

3 WATSONIAN (3,5,10,3,0) Widely but very thinly distributed throught except in the 2 YORKSHIRE northwest of Yorkshire. Formally known 1 as E. pusilla this species has been 0 NY NZ NZ OV SD SE SE TA recorded by general sweeping in 9 woodland, at sap runs, in compost bins 8 and in fungi. (v-ix). 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 29 | P a g e melina Erichson, 1843

Epuraea (Epuraea) melina Erichson, 1843

3 WATSONIAN (4,7,12,1,0) Widely but unevenly distributed in the county with records 2 YORKSHIRE from the northeast and southwest. 1 Recorded habitats include general 0 NY NZ NZ OV SD SE SE TA vegetation in woodland rides, in tidal 9 refuse on the seashore, on hawthorn 8 blossom and in debris from the nest of a fieldmouse. The latter record accords 7 with an accepted association with the 6 burrows of small mammals. (v-xii). 5 4 3 2 1

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neglecta (Heer, 1841)

NO RECORDS

pallescens (Stephens, 1835)

Epuraea (Epuraea) pallescens (Stephens, 1832)

3 WATSONIAN (9,5,19,2,0). Widely distributed except in the north and west. Recorded from 2 YORKSHIRE fungi and flowers of hawthorn, cherry 1 and sloe, and a single record from a sap 0 NY NZ NZ OV SD SE SE TA run. (iv-x). 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 30 | P a g e rufomarginata (Stephens, 1830)

Epuraea (Epuraea) rufomarginata (Stephens, 1830)

3 WATSONIAN (0,3,11,0,0) Rare in the county with only 2 YORKSHIRE 14 records. Noted from oak and from leaf litter in mature woodlands, and 1 once in a compost heap in a suburban 0 NY NZ NZ OV SD SE SE TA garden. Nationally, recorded from leaf 9 litter, fungi and under bark of deciduous 8 and coniferous trees. 7 6 5 4 3 2 1

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silacea (Herbst, 1783)

Epuraea (Epuraea) silacea (Herbst, 1783)

3 WATSONIAN (3,13,12,8,1) Widely distributed in 2 YORKSHIRE Yorkshire. Formally known to coleopterists as E. deleta, this species 1 has been recorded in fungi on oak, 0 NY NZ NZ OV SD SE SE TA willow, sycamore and horse chestnut, in 9 the red-rot of old logs and on one 8 accasion in ground litter at a pond 7 margin. In spite of the entry in Hyman and Parsons (1994:63) this species 6 appears to be widely distributed 5 nationally. (iv-ix). 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 31 | P a g e terminalis (Mannerheim, 1843)

Epuraea terminalis Mannerheim, 1843

3 WATSONIAN (0,2,0,0,0) RDBI. Very scarce in Yorkshire 2 YORKSHIRE with only two records: Duncombe Park (SE6082), 10/9/1985, RJM det CJ, 1 around old logs; Castle Hill Wood 0 NY NZ NZ OV SD SE SE TA (SE5783), 5/6/2000, RJM, by sweeping 9 at edge of woodland ride. A species of 8 relict old forest. First recognised in 7 Britain in 1972 (as Epuraea adumbrata) (KNAA, 2002), (Johnson, 1972). 6 5 4 3 2 1

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thoracica Tournier, 1872

Epuraea (Epuraea) thoracica Tournier, 1872

3 WATSONIAN (0,2,0,1,0) Nationally Scarce. There are three records on the YNU database: 2 YORKSHIRE Grassington (SE06), 9/1909, JWC; 1 Ellerburn (SE8484), 9/7/1967, EWA; 0 NY NZ NZ OV SD SE SE TA Mulgrave Woods (NZ8513), c.1930, HB. 9 Another record from Wykeham Forest 8 (SE9588), 2001, (LA) requires confirmation by way of a voucher. 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 32 | P a g e variegata (Herbst, 1793)

Epuraea (Epuraea) v ariegata (Herbst, 1793)

3 WATSONIAN (0,1,0,0,0) RDBK. There is a single old record of this little known species on the 2 YORKSHIRE YNU database: Scarborough district 1 (TA08), c.1900, RL. A voucher specimen NY NZ NZ OV 0 SD SE SE TA would be required to confirm this 9 record. Mainly Scottish, this species is 8 known from bracket fungi on Betula (Alexander, 2002). 7 6 5 4 3 2 1

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Subgenus EPURAEANELLA Crotch, 1874 limbata (Fabricius, 1787)

Epuraea limbata (Fabricius, 1787)

3 WATSONIAN (2,1,0,0,0) Very rarely recorded in the county with three, all old, records: Hull 2 YORKSHIRE (TA02), pre-1940, TS; Market Weighton 1 (SE8742), pre-1950, EGB; Wykeham 0 NY NZ NZ OV SD SE SE TA (SE9684), 22/6/1901, EGB, see Nat. 9 1901:296. Recorded nationally from 8 fungi on trees. 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 33 | P a g e

Subgenus MICRURIA Reitter, 1875 melanocephala (Marsham, 1802)

Epuraea (Micruria) melanocephala (Marsham, 1802)

3 WATSONIAN (19,22,149,18,6). Widespread and fairly 2 YORKSHIRE common, being recorded mostly from blossoms on hawthorn but also from 1 cherry and lime, in leaf litter in 0 NY NZ NZ OV SD SE SE TA woodland, general sweeping in 9 woodland rides, in rotting fungi, in moss 8 and leaf litter in grassland, and once in a 7 flight interception trap. (iii-x, peak vi). 6 5 4 3 2 1

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Subfamily MELIGETHINAE Thomson, C.G., 1859

PRIA Stephens, 1829 dulcamarae (Scopoli, 1763)

Pria dulcamarae (Scopoli, 1763)

3 WATSONIAN (10,1,23,6,0) Widespread in the 2 YORKSHIRE southern half of the county, and often numerous where its foodplant Solanum 1 dulcamara occurs. Usually in wet places. 0 NY NZ NZ OV SD SE SE TA (v-x, peaks vi and ix). 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 34 | P a g e

MELIGETHES Stephens, 1829 aeneus (Fabricius, 1775)

Meligethes aeneus (Fabricius, 1775)

3 WATSONIAN (233,148,1249,127,56) Widespread and extremely common on a wide variety of 2 YORKSHIRE yellow Brassicaceae, especially Oil-seed 1 Rape. With over 1800 records this 0 NY NZ NZ OV SD SE SE TA species is one of our most commonly- 9 recorded beetle species. May be 8 confused with M. viridescens in the field. (iii-x, peaks vi and vii). 7 6 5 4 3 2 1

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atramentarius Förster, 1849

Meligethes atramentarius Förster, 1849

3 WATSONIAN (0,1,0,1,0) Very scarce in Yorkshire with only two records: Studley Royal Park 2 YORKSHIRE (SE2969), 1871, EAW, see EMM 1871:38; 1 Kirkdale (SE672864), 8/8/1998, WRD, 0 NY NZ NZ OV SD SE SE TA swept from Galeopsis. According to Kirk- 9 Spriggs (1996:62) the larval foodplant is 8 Lamiastrum galeobdolon Yellow 7 Archangel. 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 35 | P a g e atratus (Olivier, 1790)

Meligethes atratus (Oliv ier, 1790)

3 WATSONIAN (7,14,78,13,1) Widespread and fairly 2 YORKSHIRE common but unrecorded in the north and west. The species is associated with 1 Rosa spp. particularly R. canina agg. 0 NY NZ NZ OV SD SE SE TA Mainly beaten and swept from this plant 9 but also recorded in ground litter in 8 winter, in flood debris, pitfall traps, in 7 flight interception traps, and by beating various trees and shrubs in summer. 6 (iv-x, peak v). 5 4 3 2 1

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bidens Brisout de Barneville, 1863

Meligethes bidens Brisout de Barnev ille, 1863

3 WATSONIAN (1,0,0,0,0) There is a single record on the 2 YORKSHIRE YNU database: Great Givendale (SE8153), 24/7/2003, WRD. The 1 foodplant is stated to be Clinopodium 0 NY NZ NZ OV SD SE SE TA vulgare Wild Basil (Kirk-Spriggs, 1996). 9 8 7 6 5 4 3 2 1

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bidentatus Brisout de Barneville, 1863

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 36 | P a g e brevis Sturm, 1845

Meligethes brev is Sturm, 1845

3 WATSONIAN (5,4,0,1,0) RDBI. Very scarce in Yorkshire 2 YORKSHIRE with only ten records. Recorded from Helianthemum species, and scarce owing 1 to its habitat requirements. (vi-viii). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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brunnicornis Sturm, 1845

Meligethes brunnicornis Sturm, 1845

3 WATSONIAN (3,7,10,2,0) Widespread but very infrequent in the eastern half of the 2 YORKSHIRE county. Several of our records mention 1 the accepted foodplant Stachys sylvatica 0 NY NZ NZ OV SD SE SE TA Hedge Woundwort. Very infrequently 9 recorded since 2000. (v-x). 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 37 | P a g e carinulatus Förster, 1849

Meligethes carinulatus Förster, 1849

3 WATSONIAN (22,11,53,2,2) Widespread and fairly 2 YORKSHIRE common except in the far north and west. On Lotus corniculatus Bird’s-foot- 1 trefoil. Also recorded in a garden 0 NY NZ NZ OV SD SE SE TA Malaise trap. (iv-ix, peak v-vi). 9 8 7 6 5 4 3 2 1

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coracinus Sturm, 1845

NO RECORDS corvinus Erichson, 1845

NO RECORDS

difficilis (Heer, 1841)

Meligethes difficilis (Heer, 1841)

3 WATSONIAN (15,3,29,1,0) Recorded from the south and east of the county. Widespread and 2 YORKSHIRE fairly common in those areas. On 1 Lamium album White Deadnettle. (iv-xi, 0 NY NZ NZ OV SD SE SE TA peak vi). 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 38 | P a g e erichsonii Brisout de Barneville, 1863

NO RECORDS exilis Sturm, 1845

NO RECORDS flavimanus Stephens, 1830

Meligethes flav imanus Stephens, 1830

3 WATSONIAN (5,10,9,3,2) Widespread on various flowers including the probable main 2 YORKSHIRE foodplant Rosa species. (v-ix). 1

NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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fulvipes Brisout de Barneville, 1863

Meligethes fulv ipes Brisout de Barnev ille, 1863

3 WATSONIAN (5,3,1,1,0) Nationally Scarce. A scarce species in Yorkshire with only ten 2 YORKSHIRE records, none of which are recent. Our 1 records are mainly from marshy areas 0 NY NZ NZ OV SD SE SE TA such as Bubwith and Askham Bog. There 9 is a doubtful record (unmapped) from 8 Spurn Peninsula: see Denton 1995:82. 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 39 | P a g e gagathinus Erichson, 1845

Meligethes gagathinus Erichson, 1845

3 WATSONIAN (0,1,0,0,0) Nationally Scarce. There is a 2 YORKSHIRE single record on the YNU database: Forge Valley (SE9886), 11/8/1948, AME, 1 on Helianthemum. The foodplant given 0 NY NZ NZ OV SD SE SE TA in this record is at odds with the 9 comments in Kirk-Spriggs (1996:73), 8 where the foodplant is said to be 7 Mentha arvensis Corn Mint. Dr Easton reared gagathinus from this foodplant 6 (Kirk-Spriggs, loc.cit). There has been 5 nomenclatural confusion with 4 Meligethes lugubris which is associated 3 with drier habitats than gagathinus, but 2 this does not clarify what species was intended with the Forge Valley record. 1

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haemorrhoidalis Förster, 1849

NO RECORDS

incanus Sturm, 1845

Meligethes incanus Sturm, 1845

3 WATSONIAN (0,0,1,0,0) Nationally Scarce. There is a 2 YORKSHIRE single record on the YNU database: Lindrick Common (SK5482), 9/5/1991, 1 WAE. The host-plant is Nepeta cataria 0 NY NZ NZ OV SD SE SE TA Cat Mint although host-plant details are 9 not given in the record. The record is not 8 included in Kirk-Spriggs (1995:80), who 7 either was not aware of the record or excluded it as unlikely. A voucher would 6 be required to confirm the database 5 entry. 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 40 | P a g e kunzei Erichson, 1845

Meligethes kunzei Erichson, 1845

3 WATSONIAN (0,0,0,1,0) Our single record is from 2 YORKSHIRE VC64: Studley Royal Park (SE26), 1879, EAW. As this record is very old a voucher 1 would decide its authenticity. The 0 NY NZ NZ OV SD SE SE TA foodplant is Lamiastrum galeobdolon 9 Yellow Archangel. Kirk-Spriggs (1996:64) 8 includes the vice-county of Mid-W. York 7 as being one of Dr Easton’s unpublished records – could this refer to our record? 6 5 4 3 2 1

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lugubris Sturm, 1845

Meligethes lugubris Sturm, 1845

3 WATSONIAN (2,0,1,0,0) Nationally Scarce. There are 2 YORKSHIRE three records on the YNU database: Wharram Quarry (SE8665), 9/6/1981, 1 RSK det MLD; Wharram-le-Street 0 NY NZ NZ OV SD SE SE TA (SE8666), 13/8/2002, WRD, in flowers of 9 Campanula; Blackburn Meadows 8 (SK4192), 13/8/2000, AL. Associated 7 with Thymus polytrichus Wild Thyme. 6 5 4 3 2 1

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matronalis Audisio & Spornraft, 1990

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 41 | P a g e morosus Erichson, 1845

Meligethes morosus Erichson, 1845

3 WATSONIAN (11,4,11,1,0) Widespread in the south- eastern half of the county but with few 2 YORKSHIRE recent records. Not a frequently reported 1 species in spite of the nationally 0 NY NZ NZ OV SD SE SE TA accepted status of ‘common’. Associated 9 with Lamium album White Dead-nettle. 8 7 6 5 4 3 2 1

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nanus Erichson, 1845

NO RECORDS

nigrescens Stephens, 1830

Meligethes nigrescens Stephens, 1830

3 WATSONIAN (42,16,89,13,2) Widespread and fairly common, this species is mostly recorded 2 YORKSHIRE from the south and the east of the 1 county, and is associated with Trifolium 0 NY NZ NZ OV SD SE SE TA repens White Clover. (iii-x, peaks v and 9 vii). 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 42 | P a g e obscurus Erichson, 1845

Meligethes obscurus Erichson, 1845

3 WATSONIAN (2,7,9,4,0) Widespresd but infrequent. 2 YORKSHIRE Not recorded from the far north-west. Associated with Teucrium scorodonia 1 Wood Sage. (iv-x, peak vii). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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ochropus Sturm, 1845

Meligethes ochropus Sturm, 1845

3 WATSONIAN (0,0,2,0,0) Rare with only two known 2 YORKSHIRE Yorkshire records: Swinton (SK4498), 16/7/1978, WAE, see Nat 1988:71; 1 Wharncliffe Wood (SK3094), 1979, JDC. 0 NY NZ NZ OV SD SE SE TA Both these records would require 9 vouchers for confirmation. The habitat is 8 damp places in association with Stachys 7 palustris Marsh Woundwort. The two records do not mention habitat or plant. 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 43 | P a g e ovatus Sturm, 1845

Meligethes ov atus Sturm, 1845

3 WATSONIAN (5,2,6,0,0) Scarce and apparently 2 YORKSHIRE confined to the eastern half of the county. Several of our thirteen records 1 mention the foodplant which is 0 NY NZ NZ OV SD SE SE TA Glechoma hederacea Ground Ivy (Kirk- 9 Spriggs, 1996) 8 7 6 5 4 3 2 1

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pedicularius (Gyllenhal, 1808)

Meligethes pedicularius (Gyllenhal, 1808)

3 WATSONIAN (7,2,16,2,2) Widely distributed but local, with few recent records. Associated with 2 YORKSHIRE Galeopsis tetrahit Common Hemp-nettle, 1 but recorded from other plants including 0 NY NZ NZ OV SD SE SE TA Stachys and Senecio species. Also taken 9 in a Malaise trap in a suburban garden. 8 (v-x). 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 44 | P a g e persicus (Faldermann, 1835)

NO RECORDS

planiusculus (Heer, 1841)

Meligethes planiusculus (Heer, 1841)

3 WATSONIAN (0,1,6,0,0) Very scarce in the county with only seven records. Associated with 2 YORKSHIRE Echium vulgare Viper’s-bugloss and 1 recorded from this plant in two of our 0 NY NZ NZ OV SD SE SE TA records. 9 8 7 6 5 4 3 2 1

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rotundicollis Brisout de Barneville, 1863

Meligethes rotundicollis Brisout de Barnev ille, 1863

3 WATSONIAN (0,0,1,0,0) Nationally Scarce. There is a single record on the YNU database: Ulley 2 YORKSHIRE (SE4587), 14/5/1988, WAE. The host 1 plant was not recorded here but the 0 NY NZ NZ OV SD SE SE TA species is associated with Sinapsis 9 arvensis Charlock and Sisymbrium 8 officinale Hedge Mustard (Kirk-Spriggs, 1996). 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 45 | P a g e ruficornis (Marsham, 1802)

Meligethes ruficornis (Marsham, 1802)

3 WATSONIAN (1,0,0,0,0) We possess a single record: 2 YORKSHIRE Spurn (TA4212), 16/8/2006, WRD, swept off Solanum dulcamara, although Kirk- 1 Spriggs (1996) gives Balota nigra Black 0 NY NZ NZ OV SD SE SE TA Horehound as the larval host plant from 9 which Easton reared adults (op.cit.). 8 7 6 5 4 3 2 1

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serripes (Gyllenhal, 1827)

NO RECORDS

solidus (Kugelann, 1794)

Meligethes solidus (Kugelann, 1794)

3 WATSONIAN (0,0,1,3,0) Nationally Scarce. We have four records on the YNU database: 2 YORKSHIRE Hetchell Crags Reserve (SE3842), 8/1959 1 and 8/1961, JHF, on Helianthemum; 0 NY NZ NZ OV SD SE SE TA Ripon Parks (SE3075), 15/6/2002, FEK; 9 Lindrick (SK5482), 11/5/1985, WAE, see 8 Nat 1988:71. Associated with Helianthemum vulgare Common Rock- 7 rose. 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 46 | P a g e subrugosus (Gyllenhal, 1808)

Meligethes subrugosus (Gyllenhal, 1808)

3 WATSONIAN (7,0,0,0,0) Nationally Scarce. All our records are from VC61. Kirk-Spriggs 2 YORKSHIRE (1996) notes Jasione montana Sheep’s- 1 bit as the host-plant, but recorded from 0 NY NZ NZ OV SD SE SE TA other plants so the host-plant situation is 9 unclear. 8 7 6 5 4 3 2 1

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symphyti (Heer, 1841)

NO RECORDS

umbrosus Sturm, 1845

NO RECORDS

viridescens (Fabricius, 1787)

Meligethes v iridescens (Fabricius, 1787)

3 WATSONIAN (4,15,46,6,2) Many records, especially 2 YORKSHIRE from yellow brassicas, though not apparently as frequent as M. aeneus. 1 Some of our aeneus records may be 0 NY NZ NZ OV SD SE SE TA referrable to viridescens as the two 9 species are similar, best distinguished by 8 genitalic characters and form of the mid- 7 femora. (iv-ix). 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 47 | P a g e

Subfamily NITIDULINAE Latreille, 1802

NITIDULA Fabricius, 1775 bipunctata (Linnaeus, 1758)

Nitidula bipunctata (Linnaeus, 1758)

3 WATSONIAN (4,5,10,5,2) An infrequently-recorded 2 YORKSHIRE species of carcasses, usually of sheep and larger birds, although on one occasion in 1 a dead porpoise on the sea shore. (v-vii). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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carnaria (Schaller, 1783)

NO RECORDS

flavomaculata Rossi, 1790

Nitidula flav omaculata Rossi, 1790

3 WATSONIAN (0,1,0,0,0) There is a single record on our database: Scarborough (TA08), 1871, RL, 2 YORKSHIRE in ram’s horn at bone mill. 1

NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 48 | P a g e rufipes (Linnaeus, 1767)

Nitidula rufipes (Linnaeus, 1767)

3 WATSONIAN (1,0,5,3,0) Very scarce with no recent 2 YORKSHIRE records, and unrecorded since 1943. A species associated with carrion and other 1 rotting animal material. NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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OMOSITA Erichson, 1843 colon (Linnaeus, 1758)

Omosita colon (Linnaeus, 1758)

3 WATSONIAN (13,1,15,6,1) Widespread but not 2 YORKSHIRE common, and recorded from the remains of birds, deer, rabbit and fox in advanced 1 state of decay. Mostly in the southern 0 NY NZ NZ OV SD SE SE TA half of the county – there is a single VC65 9 record. (iv-x). 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 49 | P a g e depressa (Linnaeus, 1758)

Omosita depressa (Linnaeus, 1758)

3 WATSONIAN (0,3,12,3,0) Widespread and scarce. This little-recorded species has been noted 2 YORKSHIRE from dead birds and sheep carcasses. 1 Apparently restricted to upland locations 0 NY NZ NZ OV SD SE SE TA in Yorkshire. 9 8 7 6 5 4 3 2 1

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discoidea (Fabricius, 1775)

Omosita discoidea (Fabricius, 1775)

3 WATSONIAN (11,9,36,13,1) This, the commonest species of the genus, is rather centrally 2 YORKSHIRE distributed in the county. Recorded from 1 dead rabbits, sheep, in old bones and in 0 NY NZ NZ OV SD SE SE TA dry bracket fungi. (iv-x). 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 50 | P a g e

SORONIA Erichson, 1843 grisea (Linnaeus, 1758)

Soronia grisea (Linnaeus, 1758)

3 WATSONIAN (7,8,25,9,1) Widespread in the county but not common, this seems to favour 2 YORKSHIRE sap runs on trees, but also recorded from 1 under bark of birch, ash and oak. (iii-ix). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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punctatissima (Illiger, 1794)

Soronia punctatissima (Illiger, 1794)

3 WATSONIAN (0,4,6,6,0) Widely distributed but scarce. 2 YORKSHIRE Recorded from sap runs, in carrion, in rotting fungi under under bark of birch. 1 (iv-xi). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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AMPHOTIS Erichson, 1843 marginata (Fabricius, 1781)

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 51 | P a g e

CYCHRAMUS Kugelann, 1794 luteus (Fabricius, 1787)

Cychramus luteus (Fabricius, 1787)

3 WATSONIAN (1,11,13,2,0) Patchily distributed in the south and east of the county. Adults 2 YORKSHIRE reported from flowers of Crataegus 1 monogyna Hawthorn, Filipendula ulmaria NY NZ NZ OV 0 SD SE SE TA Meadowsweet, Rubus Bramble and 9 Sambucus nigra Elder, also in light traps 8 and fungi. (v-x). 7 6 5 4 3 2 1

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POCADIUS Erichson, 1843 adustus Reitter, 1888

Pocadius adustus Reitter, 1888

3 WATSONIAN (0,4,8,0,0) Scarce in the county with only 2 YORKSHIRE twelve records, this species is recorded from puffballs Lycopendon, but once in 1 rotting bracket fungi on oak Quercus spp. NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 52 | P a g e ferrugineus (Fabricius, 1775)

Pocadius ferrugineus (Fabricius, 1775)

3 WATSONIAN (3,3,13,2,1) Relatively more common and widespread than P. adustus, again in 2 YORKSHIRE Lycoperdon species, and in other 1 unidentified fungi. On one occasion both NY NZ NZ OV 0 SD SE SE TA Pocadius species were recorded in the 9 same Lycoperdon specimen. 8 7 6 5 4 3 2 1

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THALYCRA Erichson, 1843 fervida (Olivier, 1790)

Thalycra ferv ida (Oliv ier, 1790)

3 WATSONIAN (0,0,1,2,0) Nationally Notable B. Very 2 YORKSHIRE rare in the county with only three known records: Hail Mary Hill Wood (SK4386), 1 29/7/1981, WAE det CJ; Cawood 0 NY NZ NZ OV SD SE SE TA (SE5737), 26/8/1990, KP det RJM, in a 9 garden light trap; Brayton Barff (SE5830), 8 2000, AG, at sap runs on tree trunk. This 7 little-known species probably develops in fungi in its larval stage, the adults being 6 found in a wide variety of situations. 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 53 | P a g e

Subfamily CRYPTARCHINAE Thomson, C.G., 1859

CRYPTARCHA Shuckard, 1839 strigata (Fabricius, 1787)

Cryptarcha strigata (Fabricius, 1787)

3 WATSONIAN (0,0,3,2,0) Nationally Notable B. Very 2 YORKSHIRE rare in Yorkshire with only five records known: Wheatley Wood (SE6005), 1910, 1 EGB; Bretton Lakes (SE2812), 9/9/1986, 0 NY NZ NZ OV SD SE SE TA MLD; Lindholme (SE7006), 16/9/2019, 9 PB, in flight trap on oak; Brayton Barff 8 (SE5830), 19/7/2000 and 8/9/2000, AG 7 det MLD, at sap runs on oak. 6 5 4 3 2 1

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undata (Olivier, 1790)

Cryptarcha undata (Oliv ier, 1790)

3 WATSONIAN (0,0,1,3,0) Very rare in Yorkshire with only four known records: Lindholme 2 YORKSHIRE (SE7006), 16/9/2019, PB, in flight trap on 1 oak; Brayton Barff (SE5830), three 0 NY NZ NZ OV SD SE SE TA records - 23/7/1996, AG det PW, at sap 9 runs on oak; - 19/7/2000, AG det MLD, at 8 sap run on oak; - 8/9/2000, AG det MLD, at sap run on oak. 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 54 | P a g e

GLISCHROCHILUS Reitter, 1873

Subgenus GLISCHROCHILUS Reitter, 1873 quadripunctatus (Linnaeus, 1758)

Glischrochilus quadripunctatus (L, 1758)

3 WATSONIAN (4,8,7,3,1) Widespread but not a 2 YORKSHIRE common insect in the county. Recorded from under bark of conifers, and at sap 1 runs on stumps. 0 NY NZ NZ OV SD SE SE TA (iii-x). 9 8 7 6 5 4 3 2 1

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Subgenus LIBRODOR Reitter, 1884 hortensis (Geoffroy in Fourcroy, 1785)

Glischrochilus hortensis (Geoffroy in Fourcroy, 1785)

3 WATSONIAN (43,30,115,43,3) Widespread and common throughout except for upland 2 YORKSHIRE areas. Our records indicate a wide variety 1 of habitats including sap runs, in fungi, in NY NZ NZ OV 0 SD SE SE TA pitfall traps in woodland, in old straw 9 piles on farms, in a dead rabbit, in horse 8 dung, in a dead fox, in sedge litter and in a garden Malaise trap. (iii-xii). 7

6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 55 | P a g e quadriguttatus (Fabricius, 1777)

Glischrochilus (Librodor) quadriguttatus (Fabricius, 1777)

3 WATSONIAN (1,3,6,3,0) Widely distributed but scarce. 2 YORKSHIRE Recorded from sap runs and in fungi on trees. The VC61 record would require a 1 voucher specimen to confirm, but none 0 NY NZ NZ OV SD SE SE TA has been forthcoming. 9 8 7 6 5 4 3 2 1

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PITYOPHAGUS Shuckard, 1839 ferrugineus (Linnaeus, 1761)

NO RECORDS

Subfamily CYBOCEPHALINAE Jacquelin du Val, 1858

CYBOCEPHALUS Erichson, 1844 fodori Endrödy-Younga, 1965

NO RECORDS 56. Family MONOTOMIDAE Laporte, 1840 578

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 56 | P a g e

Subfamily RHIZOPHAGINAE Redtenbacher, 1845

RHIZOPHAGUS Herbst, 1793

Subgenus RHIZOPHAGUS Herbst, 1793 bipustulatus (Fabricius, 1792)

Rhizophagus (Rhizophagus) bipustulatus (Fabricius, 1792)

3 WATSONIAN (10,18,76,25,2) Widely distributed and 2 YORKSHIRE common under bark and in fungi on deciduous trees. (iii-xii). 1

NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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dispar (Paykull, 1800)

Rhizophagus (Rhizophagus) dispar (Paykull, 1800)

3 WATSONIAN (12,28,190,35,2) Distribution, habitat and 2 YORKSHIRE frequency similar to R. bipustulatus. The two species may be easily confused in 1 the field. (iii-xii). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 57 | P a g e fenestralis (Linnaeus, 1758)

Rhizophagus fenestralis (Linnaeus, 1758)

3 WATSONIAN (0,0,3,0,0) RDB3. Known previously as 2 YORKSHIRE Rhizophagus parvulus, we have three records for this species all from the same 1 site: Lindholme Hall (SE7006), July-Sept 0 NY NZ NZ OV SD SE SE TA 2019, PB, in flight traps on oak. Found 9 repeatedly in flight traps durng this 8 period in a very restricted location. 7 Associated with the dead or dying parts of broad-leaved trees, and only 6 otherwise known in Britain from Monks 5 Wood Huntingdonshire and the Scottish 4 Highlands. 3 2 1

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ferrugineus (Paykull, 1800)

Rhizophagus (Rhizophagus) ferrugineus (Paykull, 1800)

3 WATSONIAN (2,6,5,2,0) Widely distibuted cenrally but rather scarce with few recent records. 2 YORKSHIRE Under bark of decaying deciduous trees. 1 (ii-ix). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 58 | P a g e nitidulus (Fabricius, 1798)

Rhizophagus (Rhizophagus) nitidulus (Fabricius, 1798)

3 WATSONIAN (0,3,27,6,0) Nationally Notable B. Records are mainly from the south of the 2 YORKSHIRE county, and usually from old woodland 1 sites. (iii-xii). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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oblongicollis Blatch & Horner, 1892

Rhizophagus oblongicollis Blatch & Horner, 1892 (0,1,0,0,0). RDB1. We have a single 3 WATSONIAN YORKSHIRE record of this species in the database: 2 Langdale End (SE99), 1914, ECH. No 1 voucher specimen has been traced for

NY NZ NZ OV 0 SD SE SE TA this record so there is always the 9 possibility of a misidentification. 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 59 | P a g e parallelocollis Gyllenhal, 1827

Rhizophagus parallelocollis Gyllenhal, 1827

3 WATSONIAN (1,4,5,0,0) Usually associated with the bark layer on deadwood, this species, rarely 2 YORKSHIRE reported in Yorkshire, has been reported 1 from inside a nest of Vespula vulgaris and 0 NY NZ NZ OV SD SE SE TA from a pile of vegetable refuse on a farm. 9 There is only one recent record, from Brook 8 Farm, Elstronwick (TA2332), 2006, WRD, in 7 vegetable refuse. 6 5 4 3 2 1

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perforatus Erichson, 1845

Rhizophagus (Rhizophagus) perforatus Erichson, 1845

3 WATSONIAN (3,4,31,8,0) Widely distributed but not 2 YORKSHIRE common, and recorded from under bark of fallen or standing deciduous trees. 1 (iii-xi). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 60 | P a g e picipes (Olivier, 1790)

Rhizophagus (Rhizophagus) picipes (Oliv ier, 1790)

3 WATSONIAN (1,0,8,3,0) Nationally Notable A. This 2 YORKSHIRE species is distributed in a central area of the county and scarce. Found under bark 1 of coniferous and deciduous trees. (iii-ix). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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Subgenus ANOMOPHAGUS Reitter, 1907 cribratus Gyllenhal, 1827

Rhizophagus (Anomophagus) cribratus Gyllenhal, 1827

3 WATSONIAN (0,0,5,4,0) Very scarce with no recent records. Often associated with tree roots 2 YORKSHIRE (Alexander, 2002), our records mention 1 the species being found in Polyporus and 0 NY NZ NZ OV SD SE SE TA in a pile of decaying grass cuttings. There 9 are no recent records. 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 61 | P a g e

Subgenus EURHIZOPHAGUS Méquignon, 1909 depressus (Fabricius, 1792)

Rhizophagus (Eurhizophagus) depressus (Fabricius, 1792)

3 WATSONIAN (1,3,11,3,0) Scarce in the county, and 2 YORKSHIRE recorded from under bark of unidentified conifers. Only two recent records. 1 (iii-xii). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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grandis Gyllenhal, 1827

NO RECORDS

CYANOSTOLUS Ganglbauer, 1899 aeneus (Richter, 1820)

Cyanostolus aeneus (Richter, 1820)

3 WATSONIAN (0,1,0,0,0) Nationally Notable A. There is 2 YORKSHIRE a single, unconfirmed, record on the database: Scalby Mills (TA0191), 1900, 1 anon, no habitat details. A rare saproxylic 0 NY NZ NZ OV SD SE SE TA and mainly southern Britain, at sap runs 9 and under bark of deciduous and 8 coniferous trees, sometimes recorded 7 with scolytids. A voucher specimen as yet untraced would confirm the identity. 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 62 | P a g e

Subfamily MONOTOMINAE Laporte, 1840

MONOTOMA Herbst, 1793

Subgenus GYROCECIS Thomson, C.G., 1859 angusticollis (Gyllenhal, 1827)

Monotoma angusticollis (Gyllenhal, 1827)

3 WATSONIAN (0,3,0,0,0) RDB3. Rare in Yorkshire, we have three records on the database: York 2 YORKSHIRE (SE65), pre-1900, WCH; Scarborough 1 (TA08), pre-1918, RL, abundant in ants’ 0 NY NZ NZ OV SD SE SE TA nest; Farndale (SE6697), 1920, anon 9 (entry in ISR). 8 7 6 5 4 3 2 1

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conicicollis Guérin-Méneville, 1837

Monotoma (Gyrocecis) conicicollis Guérin-Ménev ille, 1837

3 WATSONIAN (0,4,7,0,0) Very rarely recorded, this species is myrmecophilous in the nests of 2 YORKSHIRE various Formica species. 1

NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 63 | P a g e quadrifoveolata Aubé, 1838

Monotoma (Gyrocecis) quadrifov eolata Aubé, 1838

3 WATSONIAN (0,1,0,0,0) There is a single unconfirmed 2 YORKSHIRE record on the YNU database: Scarborough (TA08), pre-1918, RL, in a 1 flour mill. NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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Subgenus MONOTOMA Herbst, 1793 bicolor Villa & Villa, 1835

Monotoma (Monotoma) bicolor Villa & Villa, 1835

3 WATSONIAN (5,2,32,5,1) A fairly frequent and widespread species, recorded from 2 YORKSHIRE decaying vegetable matter such as straw 1 heaps, compost heaps, piles of grass 0 NY NZ NZ OV SD SE SE TA cuttings and the like. (iii-xi). 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 64 | P a g e brevicollis Aubé, 1838

Monotoma (Monotoma) brev icollis Aubé, 1838

3 WATSONIAN (0,0,1,0,0) There is a single record on our database: Rotherham (SK4493), 2 YORKSHIRE 5/8/1981, LB det WAE. A southern 1 English species of decaying vegetable 0 NY NZ NZ OV SD SE SE TA matter. 9 8 7 6 5 4 3 2 1

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longicollis (Gyllenhal, 1827)

Monotoma (Monotoma) longicollis (Gyllenhal, 1827)

3 WATSONIAN (8,7,13,4,0) Widespread, recorded in Yorkshire from decaying vegetable 2 YORKSHIRE matter, straw heaps, compost piles etc, 1 also in debris in bird nest boxes. 0 NY NZ NZ OV SD SE SE TA (v-xi). 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 65 | P a g e picipes Herbst, 1793

Monotoma (Monotoma) picipes Herbst, 1793

3 WATSONIAN (13,8,37,9,2) A fairly common and widespread species, mainly recorded 2 YORKSHIRE from compost heaps and piles of cut 1 grass, also in piles of dry manure. (ii-xi, 0 NY NZ NZ OV SD SE SE TA peak vii). 9 8 7 6 5 4 3 2 1

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spinicollis Aubé, 1838

Monotoma (Monotoma) spinicollis Aubé, 1838

3 WATSONIAN (3,0,4,3,0) An infrequent species with no recent records. Noted from compost 2 YORKSHIRE heaps and an old potato dump. 1

NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 66 | P a g e testacea Motschulsky, 1845

Monotoma testacea Motschulsky, 1845

3 WATSONIAN (0,0,6,2,0) Widely distributed but scarce 2 YORKSHIRE in Yorkshire with no recent records. In grass cuttings and other decaying 1 vegetable matter. NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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57. Family SILVANIDAE Kirby, 1837

Subfamily BRONTINAE Blanchard, 1845

ULEIOTA Latreille, 1796 planatus (Linnaeus, 1761)

Uleiota planatus (Linnaeus, 1761)

3 WATSONIAN (6,0,0,0,0) Nationally Notable A. Found repeatedly at a single site: Cali Heath 2 YORKSHIRE (SE7549) from 2007 to 2010, RJM, PK, 1 FEK, WRD, under bark of oak in log pile. 0 NY NZ NZ OV SD SE SE TA The Nationally Notable A designation 9 may need reconsideration as this species 8 has expanded its national range considerably since the 1980s (James, 7 2018:294). 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 67 | P a g e

DENDROPHAGUS Schönherr, 1809 crenatus (Paykull, 1799)

NO RECORDS

CRYPTAMORPHA Wollaston, 1854 desjardinsii (Guérin-Méneville, 1844)

NO RECORDS

PSAMMOECUS Latreille, 1829 bipunctatus (Fabricius, 1792)

Psammoecus bipunctatus (Fabricius, 1792)

3 WATSONIAN (5,0,11,0,0) Very scarce in Yorkshire. Until 1996 only known recently from Hornsea 2 YORKSHIRE Mere (TA1947), with sub-fossil records 1 from Thorne Moor and Conisbrough

NY NZ NZ OV 0 SD SE SE TA (SK5198). There is also an old record for 9 Askern: Rushy Moor (SE5714), 1911, HHC. The species then appeared as follows: 8 Hatfield Moor (SE6947), 25/8/2005, AG det 7 MLD; Hexthorpe Ings (SE5502), 2016-2019, 6 RJM, swept in huge numbers on a few days in May in each year; Sprotbrough (SE5503), 5 8/5/2018, RJM, in garden Malaise trap. This 4 last record demonstrates the flight ability of 3 this species as the specimen can only have 2 flown from the Hexthorpe site about a kilometre away. The species develops in the 1 stems of sedges and reeds in swamps and 0 SD SE SE TA SJ SK SK TF also in the ground layer debris in reed beds, 9 and occurs extensively in the southern half 8 post 31/12/1999 of England. Map produced for Yorkshire Naturalists' Union by DMAP 14/8/2018 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4

Subfamily SILVANINAE Kirby, 1837

CATHARTUS Reiche, 1854 quadricollis (Guérin-Méneville, 1844)

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 68 | P a g e

NAUSIBIUS Lentz, 1857 clavicornis (Kugelann, 1794)

Nausibius clav icornis (Kugelann, 1794)

3 WATSONIAN (0,0,1,0,0) There is a single record of this 2 YORKSHIRE stored products silvanid in Yorkshire: Barnsley (SE30), 1912, EGB (with no 1 other details given). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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AHASVERUS des Gozis, 1881 advena (Waltl, 1834)

Ahasv erus adv ena (Waltl, 1834)

3 WATSONIAN (8,2,19,3,0) Widespread in Yorkshire and 2 YORKSHIRE recorded from old straw bales, grass cuttings, compost heaps and once from 1 Laetiporus sulphureus on old oak, this 0 NY NZ NZ OV SD SE SE TA species is a stored-products pest that has 9 established in the wild. (ii-x). 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 69 | P a g e

ORYZAEPHILUS Ganglbauer, 1899 mercator (Fauvel, 1889)

Oryzaephilus mercator (Fauv el, 1889)

3 WATSONIAN (1,0,1,0,0) Very rarely recorded in 2 YORKSHIRE Yorkshire with two records: Raleigh Lodge (TA1432, 15/7/2006, WRD, a 1 breeding population in a domestic flour 0 NY NZ NZ OV SD SE SE TA bin; Storthes Hall (SE1712), 19/8/1984, 9 DM det MLD. A beetle of stored 8 products. 7 6 5 4 3 2 1

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surinamensis (Linnaeus, 1758)

Oryzaephilus surinamensis (Linnaeus, 1758)

3 WATSONIAN (2,0,4,0,0) This cosmopolitan pest of stored products is recorded from 2 YORKSHIRE locations in the south of the county, from 1 commercial food premises and grain 0 NY NZ NZ OV SD SE SE TA stores. 9 8 7 6 5 4 3 2 1

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SILVANUS Latreille, 1804 bidentatus (Fabricius, 1792)

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 70 | P a g e unidentatus (Olivier, 1790)

Silv anus unidentatus (Oliv ier, 1790)

3 WATSONIAN (0,2,2,0,0) There are four records of this 2 YORKSHIRE saproxylic species on the YNU database: Castle Howard (SE7170), 5/9/1951, WDH, 1 one specimen under bark; Langdale End 0 NY NZ NZ OV SD SE SE TA (SE9492), 1980, RSK; Sandall Beat 9 (SE6103), 1912, HHC; Roche Abbey 8 (SK5489), pre-1970, PS. Develops under 7 the sappy bark of Quercus Oak and Fagus Beech (Alexander, 2002). 6 5 4 3 2 1

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SILVANOPRUS Reitter, 1911 fagi (Guérin-Méneville, 1844)

NO RECORDS 58. Family CUCUJIDAE Latreille, 1802

PEDIACUS Shuckard, 1839 depressus (Herbst, 1797)

Pediacus depressus (Herbst, 1797)

3 WATSONIAN (0,0,1,0,0) Nationally Notable A. There is a single record of this rare saproxylic 2 YORKSHIRE species for Yorkshire: Barnsley area 1 (SE30), 1910, EGB. Nationally, the species 0 NY NZ NZ OV SD SE SE TA is recorded from freshly cut or broken 9 stumps (Alexander, 2002). 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 71 | P a g e dermestoides (Fabricius, 1792)

Pediacus dermestoides (Fabricius, 1792)

3 WATSONIAN (0,10,5,2,1) Very scarce with a central distribution. Under bark of Quercus Oak 2 YORKSHIRE and Fagus Beech, particularly in old 1 parkland sites. NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

SD SE SE TA 0 SJ SK SK TF 9 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 14/8/2018 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 59. Family LAEMOPHLOEIDAE Ganglbauer, 1899 LAEMOPHLOEUS Dejean, 1835 monilis (Fabricius, 1787)

NO RECORDS

CRYPTOLESTES Ganglbauer, 1899 capensis (Waltl, 1834)

NO RECORDS duplicatus (Waltl, 1839)

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 72 | P a g e ferrugineus (Stephens, 1831)

Cryptolestes ferrugineus (Stephens, 1831)

3 WATSONIAN (3,5,17,3,0) Widely distributed but 2 YORKSHIRE infrequent. Under bark of Quercus Oak, Fagus Beech and Betula Birch, once in a 1 flour mill and once in a tannery amongst 0 NY NZ NZ OV SD SE SE TA old hides. (i-xii). 9 8 7 6 5 4 3 2 1

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pusilloides (Steel & Howe, 1952)

NO RECORDS

pusillus (Schönherr, 1817)

Cryptolestes pusillus (Schönherr, 1817)

3 WATSONIAN (0,0,2,0,0) There are only two very old 2 YORKSHIRE records of this rare species on the YNU database: Sheffield (SK38), 1870, YNU 1 report; Great Houghton (SE4207), 0 NY NZ NZ OV SD SE SE TA 29/6/1911, EGB. 9 8 7 6 5 4 3 2 1

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spartii (Curtis, 1834)

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 73 | P a g e turcicus (Grouvelle, 1876)

NO RECORDS

NOTOLAEMUS Lefkovitch, 1959 unifasciatus (Latreille, 1804)

NO RECORDS

LEPTOPHLOEUS Casey, 1916 clematidis (Erichson, 1846)

NO RECORDS janeti (Grouvelle, 1899)

NO RECORDS 60. Family PHALACRIDAE Leach, 1815

PHALACRUS Paykull, 1800 caricis Sturm, 1807

Phalacrus caricis Sturm, 1807

3 WATSONIAN (6,4,9,1,0) Restricted to the south- 2 YORKSHIRE eastern quarter of the county. Infrequently recorded from smutted 1 flowers of wetland sedges. 0 NY NZ NZ OV SD SE SE TA (iv-x). 9 8 7 6 5 4 3 2 1

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championi Guillebeau, 1892

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 74 | P a g e corruscus (Panzer, 1796)

Phalacrus corruscus (Panzer, 1796)

3 WATSONIAN (1,0,3,0,0) Very rare in the county with 2 YORKSHIRE only three instances recorded: Welwick (TA3421), 9/6/1946, WDH; Thorne Moor 1 (SE71), 3/8/1946; Sykehouse (SE6316), 0 NY NZ NZ OV SD SE SE TA 5/7/1977, PS. Feeds on smutted grasses 9 rather than sedges (c.f. P. caricis above). 8 7 6 5 4 3 2 1

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fimetarius (Fabricius, 1775)

Phalacrus fimetarius (Fabricius, 1775)

3 WATSONIAN (27,2,19,3,0) Like P. caricis, apparently 2 YORKSHIRE restricted to the south-eastern quarter of the county, and is associated with smuts 1 on various grasses. The commonest 0 NY NZ NZ OV SD SE SE TA member of the genus in Yorkshire, it is 9 widspread and common within its known 8 range. (v-x, peaks vi and ix). 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 75 | P a g e substriatus Gyllenhal, 1813

Phalacrus substriatus Gyllenhal, 1813

3 WATSONIAN (0,1,2,0,0) Very rare in the county with 2 YORKSHIRE only three records: Scarborough (TA08), pre-1918, RL; Langold Holt (SK5685), 1 14/7/1984, EJS det CJ; Blackmoorfoot 0 NY NZ NZ OV SD SE SE TA (SE0912), 22/2/1987, MLD. On smutted 9 Carex Sedge species. There are no recent 8 records. 7 6 5 4 3 2 1

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OLIBRUS Erichson, 1845 aeneus (Fabricius, 1792)

Olibrus aeneus (Fabricius, 1792)

3 WATSONIAN (41,6,96,10,1) Common and widespread but mainly in the south-eastern quarter. 2 YORKSHIRE Usually found by sweeping in grassland. 1 Some of our records mention Tanacetum 0 NY NZ NZ OV SD SE SE TA Tansies and Matricaria Mayweeds as 9 host plants. 8 (iii-xii, peak ix). 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 76 | P a g e affinis (Sturm, 1807)

Olibrus affinis (Sturm, 1807)

3 WATSONIAN (3,0,1,0,0) We have only four records and all are recent: Hull-Marfleet old rail track 2 YORKSHIRE (TA1329), 21/5/2002, WRD, a single 1 female on yellow Composite; Beacon 0 NY NZ NZ OV SD SE SE TA Lagoons (TA4117), 27/6/2013, a single 9 swept from Hypochaeris radicata Cat’s- 8 ear; Treeton (SK4386), 6/9/2006, FEK; Bridlington (TA1968), 30/6/2012, DW det 7 EJS. Nationally once considered coastal 6 but now producing inland records. 5 4 3 2 1

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corticalis (Panzer, 1796)

Olibrus corticalis (Panzer, 1796)

3 WATSONIAN (9,0,7,0,0) Widespread in the south and east but rather scarce. Recorded from 2 YORKSHIRE various Compositae, this species may be 1 under-recorded in the field in favour of 0 NY NZ NZ OV SD SE SE TA Olibrus aeneus. 9 8 7 6 5 4 3 2 1

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flavicornis (Sturm, 1807)

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 77 | P a g e liquidus Erichson, 1845

Olibrus liquidus Erichson, 1845

3 WATSONIAN (4,0,9,1,0) Restricted to the south of the county and recorded from yellow 2 YORKSHIRE Compositae, this species may be under- 1 recorded in the field in favour of Olibrus 0 NY NZ NZ OV SD SE SE TA aeneus. 9 8 7 6 5 4 3 2 1

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millefolii (Paykull, 1800)

NO RECORDS

pygmaeus (Sturm, 1807)

Olibrus pygmaeus (Sturm, 1807)

3 WATSONIAN (1,0,0,0,0) Nationally Notable B. There is a single record of this scarce species on 2 YORKSHIRE the YNU database: Bubwith (SE73), 1914, 1 WJF. Nationally, this species is recorded 0 NY NZ NZ OV SD SE SE TA from Leontodon and Filago. Ideally, a 9 voucher specimen would confirm this 8 record. 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 78 | P a g e

STILBUS Seidlitz, 1872 atomarius (Linnaeus, 1767)

NO RECORDS

oblongus (Erichson, 1845)

Stilbus oblongus (Erichson, 1845)

3 WATSONIAN (5,0,13,0,0) Scarce and apparently 2 YORKSHIRE restricted to the south and east of the county, this species is recorded from 1 Typha latifolia Bulrush and reedbed 0 NY NZ NZ OV SD SE SE TA ground litter in wetlands and on peat 9 moors. (i-x). 8 7 6 5 4 3 2 1

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testaceus (Panzer, 1796)

Stilbus testaceus (Panzer, 1796)

3 WATSONIAN (122,44,186,28,8) By far the most common member of the genus with 2 YORKSHIRE many records. Usually a dry grassland 1 species, but also recorded from old hay 0 NY NZ NZ OV SD SE SE TA bales, in gardens and in a Malaise trap. 9 (i-xii, peaks v and ix). 8 7 6 5 4 3 2 1

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61. Family CRYPTOPHAGIDAE Kirby, 1826 583

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 79 | P a g e

Subfamily CRYPTOPHAGINAE Kirby, 1826

Tribe CRYPTOPHAGINI Kirby, 1826

TELMATOPHILUS Heer, 1841 brevicollis Aubé, 1862

NO RECORDS caricis (Olivier, 1790)

Telmatophilus caricis (Oliv ier, 1790)

3 WATSONIAN (6,7,48,7,0) Widespread and reasonably 2 YORKSHIRE common on Carex Sedges species at margins of lakes and ponds, in fens, and 1 recorded from stands of Typha latifolia 0 NY NZ NZ OV SD SE SE TA Bulrush. (iv-x, peak vii). 9 8 7 6 5 4 3 2 1

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schonherrii (Gyllenhal, 1808)

Telmatophilus schoenherrii (Gyllenhal, 1808)

3 WATSONIAN (0,0,3,0,0) RDBK. Very rarely recorded in the county with only three records: 2 YORKSHIRE Rushy Moor (SE5712), 14/7/1982, PS, 1 swept in open fen; Rockley Dike 0 NY NZ NZ OV SD SE SE TA (SE3402), 14/7/1985, WAE; Worsborough 9 Reservoir (SE3403), 1985, JDC. Nationally 8 this species is usually recorded from 7 Typha angustifolia Lesser Bulrush. [Note: this species is synonymised with typhae 6 Fallen, 1802 in Duff, 2018.] 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 80 | P a g e sparganii (Ahrens, 1812)

NO RECORDS

typhae (Fallén, 1802)

Telmatophilus typhae (Fallén, 1802)

3 WATSONIAN (23,6,91,11,3) Common and widespread 2 YORKSHIRE on Typha latifolia Bulrush in reed beds, this is the most frequently encountered 1 member of the genus. NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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PARAMECOSOMA Curtis, 1833 melanocephalum (Herbst, 1793)

Paramecosoma melanocephalum (Herbst, 1793)

3 WATSONIAN (0,6,8,9,2) Usually found by sieving 2 YORKSHIRE ground litter in wet places and in flood debris, but also recorded from under 1 bark of wet logs and by sweeping 0 NY NZ NZ OV SD SE SE TA riverside herbage. Surprisingly, 9 unrecorded from Hatfield/Thorne Moors 8 and the numerous wetland sites in the 7 south of VC63. (v-xii). 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 81 | P a g e

HENOTICUS Thomson, C.G., 1868

californicus (Mannerheim, 1843)

NO RECORDS

serratus (Gyllenhal, 1808)

Henoticus serratus (Gyllenhal, 1808)

3 WATSONIAN (2,7,13,3,0) Widespread but rather 2 YORKSHIRE scarce with few recent records. A saproxylic species occurring under bark, 1 with adults sometimes recorded from 0 NY NZ NZ OV SD SE SE TA flowers, particularly of Crataegus 9 hawthorn. 8 (iv-xi). 7 6 5 4 3 2 1

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CRYPTOPHAGUS Herbst, 1792 acutangulus Gyllenhal, 1827

Cryptophagus acutangulus Gyllenhal, 1827

3 WATSONIAN (7,5,18,3,0) Widespread but not common. Recorded from compost 2 YORKSHIRE heaps, rotting grass piles, hay bales, 1 where there is some warmth from 0 NY NZ NZ OV SD SE SE TA putrefaction. 9 (ii-xii). 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 82 | P a g e badius Sturm, 1845

Cryptophagus badius Sturm, 1845

3 WATSONIAN (0,1,0,4,0) Very rarely recorded in Yorkshire with only five records: Little 2 YORKSHIRE Beck Wood (NZ8804) 1936, anon [record 1 requires voucher for confirmation]; 0 NY NZ NZ OV SD SE SE TA Pannal Ash (SE3052), -1939, RRUK, in 9 carrion trap; Leeds (SE33), 23/5/1951, 8 WJS, in a carrion trap; Malham Tarn (SD8866), 1956, JHF; Malham Tarn 7 (SD8866), 1957, JHF. 6 5 4 3 2 1

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Cryptophagus cellaris (Scopoli, 1763)

3 WATSONIAN (1,4,6,1,0) Very scarce with twelve records from a wide variety of habitats: 2 YORKSHIRE recorded from a hen house, a puffball 1 fungus, in stored goods in a warehouse, 0 NY NZ NZ OV SD SE SE TA the only recent records being from pitfall 9 traps on moorland. 8 7 6 5 4 3 2 1

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NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 83 | P a g e corticinus Thomson, C.G., 1863

Cryptophagus corticinus Thomson, C.G., 1863

3 WATSONIAN (0,0,1,0,0) RDBI. There is a single record 2 YORKSHIRE on the YNU database: Thorne Moor (SE7116), 13/5/1972, PS, under bark of 1 burnt birch after moorland fire. NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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dentatus (Herbst, 1793)

Cryptophagus dentatus (Herbst, 1793)

3 WATSONIAN (10,10,62,12,4) A widespread and common species. Some of our records 2 YORKSHIRE may be referrable to C. denticulatus Heer 1 (= pseudodentatus Heer) owing to 0 NY NZ NZ OV SD SE SE TA confusion in identification of these two 9 species. Under bark, Daldinia, and 8 Polyporus species, also beaten from flowers of Crataegus hawthorn. (ii-xii). 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 84 | P a g e denticulatus Heer, 1841

Cryptophagus denticulatus Heer, 1841

3 WATSONIAN (4,3,22,9,3) A widespread species 2 YORKSHIRE (previously known as C. pseudodentatus) easily confused with dentatus. But the 1 habitats in which this species is found 0 NY NZ NZ OV SD SE SE TA tend to be wetter than those in which 9 dentatus is recorded, such as reservoir 8 and lake margins, wet meadows, and 7 carrs. Found in fungi, under bark and in ground litter in such situations. (ii-xi) 6 5 4 3 2 1

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dilutus Reitter, 1874

NO RECORDS

distinguendus Sturm, 1845

Cryptophagus distinguendus Sturm, 1845

3 WATSONIAN (9,1,19,7,1) Widespread and fairly frequent in compost heaps carrion traps, 2 YORKSHIRE nests of small mammals, piles of grass 1 cuttings, farmyard debris and hay bales. 0 NY NZ NZ OV SD SE SE TA (ii-xi). 9 8 7 6 5 4 3 2 1

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falcozi Roubal, 1927

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 85 | P a g e fallax Balfour-Browne, 1953

NO RECORDS fuscicornis Sturm, 1845

NO RECORDS insulicola Roubal, 1919

NO RECORDS intermedius Bruce, 1934

NO RECORDS

labilis Erichson, 1846

Cryptophagus labilis Erichson, 1846

3 WATSONIAN (0,0,1,0,0) RDB1. There is a single record on our database of this extremely rare 2 YORKSHIRE species: Sprotbrough (SE5402), 1 17/4/1984, RJM det CJ, under bark of elm 0 NY NZ NZ OV SD SE SE TA log. 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 86 | P a g e lapponicus Gyllenhal, 1827

Cryptophagus lapponicus Gyllenhal, 1827

3 WATSONIAN (0,1,0,0,0) RDBI. A single record exists on the database of this little-known species: 2 YORKSHIRE Scarborough (TA08), -1918, RL. This 1 record should be taken as requiring 0 NY NZ NZ OV SD SE SE TA confirmation in the absence of a voucher, 9 as this species is apparently confined to 8 Scotland (Johnson, 1988). The NBN Atlas has a single record for the Shropshire 7 area but does not indicate any Scottish 6 records. 5 4 3 2 1

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laticollis Lucas, 1846

Cryptophagus laticollis Lucas, 1846

3 WATSONIAN (7,6,21,2,4) Widespread and fairly 2 YORKSHIRE frequent in fungi, carrion traps, piles of rotting grass cuttings, old haystacks, and 1 farmyard debris. (iii-xi). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 87 | P a g e lycoperdi (Scopoli, 1763)

Cryptophagus lycoperdi (Scopoli, 1763)

3 WATSONIAN (5,1,32,5,0) Widespread in the southern half of the county and usually recorded 2 YORKSHIRE from Scleroderma fungi but also reported 1 from unidentified puffballs. A relatively 0 NY NZ NZ OV SD SE SE TA large, dark and distinctive species, the 9 adults are autumnal as are the host 8 fungal fruiting bodies. (x-xi). 7 6 5 4 3 2 1

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micaceus Rey, 1889

NO RECORDS

obsoletus Reitter, 1879

Cryptophagus obsoletus Reitter, 1879

3 WATSONIAN (1,0,0,0,0) We have a single record for 2 YORKSHIRE Yorkshire on the YNU database: Selby (SE6232), 1951, CWC/GEW, in a flour mill 1 – see EMM 88(1952):260. NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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parallelus Brisout de Barneville, 1863

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 88 | P a g e populi Paykull, 1800

Cryptophagus populi Paykull, 1800

3 WATSONIAN (0,1,4,1,0) Nationally Scarce. Very scarce 2 YORKSHIRE in Yorkshire - a very distinctive species in form and coloration which should not be 1 mistaken for any other. A little-known 0 NY NZ NZ OV SD SE SE TA species, it has been associated variously 9 with bees, fruit trees and warehouses 8 (Coombs and Woodroffe, 1955). 7 6 5 4 3 2 1

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pubescens Sturm, 1845

Cryptophagus pubescens Sturm, 1845

3 WATSONIAN (3,5,13,3,2) Widely but thinly distributed throughout. Apparently this species has 2 YORKSHIRE an affinity with nests of bees and wasps, 1 but also recorded from light traps and by 0 NY NZ NZ OV SD SE SE TA general sweeping. (ii-xi). 9 8 7 6 5 4 3 2 1

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NO RECORDS punctipennis Brisout de Barneville, 1863

NO RECORDS reflexus Rey, 1889

NO RECORDS Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 89 | P a g e ruficornis Stephens, 1830

Cryptophagus ruficornis Stephens, 1830

3 WATSONIAN (5,0,12,1,0) Nationally Scarce. This 2 YORKSHIRE species is restricted to the southern half of the county, rather scarce and recorded 1 from Daldinia, on Fraxinus Ash and 0 NY NZ NZ OV SD SE SE TA Betula Birch. 9 (iii-xi). 8 7 6 5 4 3 2 1

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saginatus Sturm, 1845

Cryptophagus saginatus Sturm, 1845

3 WATSONIAN (2,1,4,0,0) Very scarce in Yorkshire with 2 YORKSHIRE only seven county records. Recorded from farmyard debris including old oats, 1 straw and compost heaps. Also in piles of 0 NY NZ NZ OV SD SE SE TA roadside grass mowings. 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 90 | P a g e scanicus (Linnaeus, 1758)

Cryptophagus scanicus (Linnaeus, 1758)

3 WATSONIAN (3,12,23,8,2) Widely but thinly distributed throughout the county. 2 YORKSHIRE Found in synanthropic situations and 1 outdoors, in compost heaps, farmyard 0 NY NZ NZ OV SD SE SE TA debris and in piles of grass cuttings. 9 (iii-x). 8 7 6 5 4 3 2 1

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schmidtii Sturm, 1845

Cryptophagus schmidtii Leunis in Sturm, 1845

3 WATSONIAN (0,2,3,0,0). RDBK. Rare in Yorkshire with 2 YORKSHIRE only five records: Newburgh Priory (SE5675), 16/7/2003, ABD, on Quercus 1 Oak; Scabba Wood (SE5201), 9/2004 and 0 NY NZ NZ OV SD SE SE TA 10/2004, coll and det RJM teste CJ, in 9 piles of pheasant feed and in pitfall traps 8 in woodland rides; Yorkshire Museum 7 (SE6052), 10/2017, AP det RJM, 1 ex. in sticky trap. 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 91 | P a g e scutellatus Newman, 1834

Cryptophagus scutellatus Newman, 1834

3 WATSONIAN (5,3,22,3,0) Widely distributed but mainly in the southern half of the county. 2 YORKSHIRE Recorded from various habitats including 1 compost heaps, haystacks, heaps of grass 0 NY NZ NZ OV SD SE SE TA cuttings and old fungi on logs. (iii-xii). 9 8 7 6 5 4 3 2 1

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Cryptophagus setulosus Sturm, 1845

3 WATSONIAN (8,15,17,5,4) Widely distributed throughout the county. Recorded from 2 YORKSHIRE bees’ nests, sweeping around woodland 1 edges, in gill fungi on the woodland floor, 0 NY NZ NZ OV SD SE SE TA straw debris in farmyards, in pitfall traps 9 on peat moors, under bark and in grass 8 piles, attracted to substrates where there is fungal decay. (iii-xi). 7 6 5 4 3 2 1

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simplex Miller, 1858

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 92 | P a g e subdepressus Gyllenhal, 1827

Cryptophagus subdepressus Gyllenhal, 1827

3 WATSONIAN (0,1,0,0,0) There is a single county record 2 YORKSHIRE on the YNU database: Whitby (NZ81), 10/11/1935, HB. There is an association 1 with coniferous trees (Coombs and 0 NY NZ NZ OV SD SE SE TA Woodroffe, 1955). 9 8 7 6 5 4 3 2 1

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subfumatus Kraatz, 1856

NO RECORDS

uncinatus Stephens, 1830

Cryptophagus uncinatus Stephens, 1830

3 WATSONIAN (1,3,2,2,0). Rare with only eight records 2 YORKSHIRE on the YNU database. C. uncinatus Stephens, previously known as 1 postpositus Sahlberg, J., (see Coombs and 0 NY NZ NZ OV SD SE SE TA Woodroffe, 1955) and immixtus Rey 9 (Duff, 2012) has been confused in the 8 past with C. badius Sturm. Also, this 7 species along with badius Sturm, ‘pallidus’ of Coombs and Woodroffe (op. 6 cit.) and scanicus (Linnaeus) can be very 5 difficult to separate (see Coombs and 4 Woodroffe, 1955:260) and are the 3 subject of long-standing nomenclatural 2 confusion. Recorded from grass piles and carrion traps. 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 93 | P a g e

MICRAMBE Thomson, C.G., 1863 abietis (Paykull, 1798)

Micrambe abietis (Paykull, 1798)

3 WATSONIAN (1,0,0,0,0) There is a single county record 2 YORKSHIRE known to us: Spurn – Warren Cottage (TA4115), 7/1953, det WDH teste GEW, 1 one specimen on wing near cottage. 0 NY NZ NZ OV SD SE SE TA There is an association of this species 9 with conifers as the specific name 8 suggests (Coombs and Woodroffe, 7 1955:263). 6 5 4 3 2 1

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bimaculata (Panzer, 1798)

NO RECORDS

pilosula (Erichson, 1846)

Micrambe pilosula (Erichson, 1846)

3 WATSONIAN (7,0,1,0,0) Previously known as 2 YORKSHIRE lindbergorum Bruce, there are eight records from five sites in the south of the 1 county. In the Spurn area this species 0 NY NZ NZ OV SD SE SE TA may be found by sweeping or shaking 9 dead thistle stems but also from drift 8 refuse on the beach, while the Thorne 7 locality yielded its record by sweeping ruderal vegetation around the levelled 6 colliery site. A Bedale (SE28) record with 5 no date or recorder has been discounted 4 and deleted from the database. 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 94 | P a g e ulicis (Stephens, 1830)

Micrambe ulicis (Stephens, 1830)

3 WATSONIAN (50,21,105,5,16) Previously widely known to coleopterists as M. vini (Panzer) this 2 YORKSHIRE species is widely distributed and 1 common wherever Ulex Gorse or Cytisus 0 NY NZ NZ OV SD SE SE TA Broom occur – adults are often found in 9 large numbers in the flowers. 8 (iii-xii, peaks v and viii). 7 6 5 4 3 2 1

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woodroffei Johnson, 2007

Micrambe woodroffei Johnson, 2007

3 WATSONIAN (16,5,10,0,0) Previously known to British 2 YORKSHIRE coleopterists as M. villosus Heer, this species has often been recorded on 1 Cirsium Thistles and Cytisus Broom 0 NY NZ NZ OV SD SE SE TA species. Widely distributed in the eastern 9 half of the county. (vi-ix). 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 95 | P a g e

ANTHEROPHAGUS Dejean, 1821 pallens (Linnaeus, 1758)

Antherophagus pallens (Linnaeus, 1758)

3 WATSONIAN (4,7,36,4,1) Previously known to British 2 YORKSHIRE coleopterists as A. nigricornis, this species is widely distributed but with few 1 recent records and may be declining in 0 NY NZ NZ OV SD SE SE TA the Yorkshire region. Adults visiting 9 flowers are carried into the nests of 8 Bombus species where the larvae 7 develop. Little else is known of the life cycle. 6 The name ‘pallens’ has been applied to 5 both species of this genus in the past so 4 some of our records may apply to similis 3 (see below). 2 (vi-x). 1

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silaceus (Herbst, 1792)

NO RECORDS

similis Curtis, 1835

Antherophagus similis Curtis, 1835

3 WATSONIAN (3,6,34,8,1) Previously known to British coleopterists as A. pallens. Widely 2 YORKSHIRE distributed with a healthy proportion of 1 recent records and with a life cycle as in 0 NY NZ NZ OV SD SE SE TA the previous species. (vi-ix). 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 96 | P a g e

Tribe CAENOSCELINI Casey, 1900

CAENOSCELIS Thomson, C.G., 1863 ferruginea (Sahlberg, C.R., 1820)

Caenoscelis ferruginea (Sahlberg, C.R., 1820)

3 WATSONIAN (0,4,4,0,0) Very scarce in the county with a single recent record. May be declining. 2 YORKSHIRE Recorded from ground litter by water, in 1 dead leaves in woodland and in a water 0 NY NZ NZ OV SD SE SE TA trap on a riverbank. A species which 9 inhabits substrates with fungal decay, 8 including woodland leaf litter and 7 compost heaps. 6 5 4 3 2 1

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sibirica Reitter, 1889

Caenoscelis sibirica Reitter, 1889

3 WATSONIAN (0,0,1,0,0) A single record exists on the 2 YORKSHIRE YNU database: TP Wood, Huddersfield (SE1317), 7/2/1981, MLD det CJ, 1 recorded from woodland litter, with the 0 NY NZ NZ OV SD SE SE TA specimen in Manchester Museum. 9 Currently the sole British record. 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 97 | P a g e subdeplanata Brisout de Barneville, 1882

Caenoscelis subdeplanata Brisout de Barnev ille, 1882

3 WATSONIAN (2,0,19,1,0). This species was once fairly 2 YORKSHIRE commonly recorded, mainly in VC63, but with only a single recent record, so 1 probably declining for reasons unknown. 0 NY NZ NZ OV SD SE SE TA Recorded variously by beating oak 9 foliage, in shoreline refuse, in ground 8 debris on post-industrial (steelworks) 7 site, and at light in a house. 6 5 4 3 2 1

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Subfamily ATOMARIINAE LeConte, 1861

Tribe HYPOCOPRINI Reitter, 1879

HYPOCOPRUS Motschulsky, 1839 latridioides Motschulsky, 1839

NO RECORDS

Tribe ATOMARIINI LeConte, 1861

ATOMARIA Stephens, 1829

Subgenus ATOMARIA Stephens, 1829 badia Erichson, 1846

NO RECORDS bella Reitter, 1875

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 98 | P a g e diluta Erichson, 1846

Atomaria (Atomaria) diluta Erichson, 1846

3 WATSONIAN (0,0,2,5,0) RDBK. A very scarce Atomaria from upland and inland sites in the 2 YORKSHIRE western half of the county. Recorded 1 particularly from fungi around old logs by 0 NY NZ NZ OV SD SE SE TA riverbanks and in other damp places. 9 One of our records, Roundhay Park 8 (SE3337), dated June 1910, is unsubstantiated (CJ pers comm to RJM). 7 6 5 4 3 2 1

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fimetarii (Fabricius, 1792)

Atomaria fimetarii (Fabricius, 1792)

3 WATSONIAN (2,1,6,2,0) Nationally Scarce. Very scarce with only a small number of records in 2 YORKSHIRE the south of the county. Recorded from 1 Coprinus comatus, swept on raodside 0 NY NZ NZ OV SD SE SE TA verges and at a lake margin, and in a 9 straw heap. Mainly a grassland species. 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 99 | P a g e impressa Erichson, 1846

Atomaria (Atomaria) impressa Erichson, 1846

3 WATSONIAN (0,1,0,1,1) Nationally Scarce. Very rare in 2 YORKSHIRE the county with only three records, all confirmed by CJ from voucher 1 specimens: Scarborough (TA08), pre- 0 NY NZ NZ OV SD SE SE TA 1918, RL; Langton Bridge (SE2996), 1984, 9 HM; Studley Royal Park (SE2969), 8 19/6/1989, RJM, in debris under old logs. 7 A species associated with broadleaved trees in river valleys (Johnson, 1993). 6 5 4 3 2 1

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linearis Stephens, 1830

Atomaria (Atomaria) linearis Stephens, 1830

3 WATSONIAN (23,16,58,13,7) A widespread and common species of ground litter, 2 YORKSHIRE compost heaps, piles of grass cuttings 1 and by general sweeping of ruderal 0 NY NZ NZ OV SD SE SE TA vegetation, in carrion traps in reed litter 9 and in fungi on logs and under bark. 8 (iii-xii, peak vi). 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 100 | P a g e lohsei Johnson & Strand, A., 1968

Atomaria lohsei Johnson & Strand, A., 1968

3 WATSONIAN (0,1,2,0,0) RDBK. A rare species associated with coniferous trees and with 2 YORKSHIRE only three county records: Haw Park 1 (SE3615), 15/8/1992, RJM det CJ, under 0 NY NZ NZ OV SD SE SE TA pine bark; May Moss (SE8795), 9 27/7/1996, LA det CJ, in flight 8 interception trap adjacent Lodgepole Pine; Swinden Plantation (SE1900), 7 9/3/1998, EJS det CJ. 6 5 4 3 2 1

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longicornis Thomson, C.G., 1863

NO RECORDS

nigrirostris Stephens 1830

Atomaria (Atomaria) nigrirostris Stephens 1830

3 WATSONIAN (9,7,62,15,3) A fairly common species of compost heaps, and in ground litter, old 2 YORKSHIRE fungi, piles of grass cuttings particularly 1 in damp places. (iii-xii, peak vi). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 101 | P a g e nigriventris Stephens, 1830

Atomaria nigriv entris Stephens, 1830

3 WATSONIAN (0,2,3,1,0) Nationally Scarce. Very scarce 2 YORKSHIRE with only six known records. The Leven Bridge (NZ4412) record (pre-1900, RL) is 1 unsubstantiated. The others have been 0 NY NZ NZ OV SD SE SE TA verified from specimens by CJ. A 9 declining grassland species recorded 8 from ground litter and garden refuse. 7 6 5 4 3 2 1

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pulchra Erichson, 1846

Atomaria pulchra Erichson, 1846

3 WATSONIAN (1,0,1,2,0) Rarely recorded in the county with only four known records with none 2 YORKSHIRE recent: Stanks Pond (SE3735), -1916, 1 EAW, det CJ; Wheldrake (SE6845), 0 NY NZ NZ OV SD SE SE TA 10/3/1991, RJM det CJ, in flood refuse; 9 Thorne Moor (SE7215), 26/6/1994, NH 8 det CJ; Staveley YWT Reserve (SE3663), 13/11/1994, MLD det CJ, in ground litter 7 by pond in woodland. (Previously known 6 to coleopterists as barani Brisout de 5 Barneville, 1863). 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 102 | P a g e puncticollis Thomson, C.G., 1868

Atomaria (Atomaria) puncticollis Thomson, C.G., 1868

3 WATSONIAN (1,0,0,0,0) RDBI. A single record exists on 2 YORKSHIRE the YNU database: Manor Farm, Eddlethorpe (SE7766), 2001, DL det CJ. 1

NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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punctithorax Reitter, 1888

Atomaria (Atomaria) punctithorax Reitter, 1887

3 WATSONIAN (2,0,1,1,0) Nationally Scarce. Rare in 2 YORKSHIRE Yorkshire with only four records: Stocksmoor YWT Reserve (SE2714), 1 16/7/1965, EWA; Tockwith (SE4652), 0 NY NZ NZ OV SD SE SE TA 26/7/1969, EWA det CJ, in manure heap, 9 specimen in Manchester Museum; 8 Blacktoft Sands (SE8524), 17/8/1982, 7 collector unknown, det CJ, specimen in Manchester Museum; Filey Old Quay 6 Rocks (TA1281), 21/5/2005, WRD, sieved 5 from beach drift. 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 103 | P a g e strandi Johnson, 1967

Atomaria strandi Johnson, 1967

3 WATSONIAN (0,1,3,0,0) Nationally Scarce. Another rarity with only four known records: 2 YORKSHIRE Scarborough (TA08), c1860, TW det CJ, 1 determined from specimen in 0 NY NZ NZ OV SD SE SE TA ?Manchester Museum; Howell Wood 9 (SE4309), 9/7/1977, PS det CJ; Pot 8 Ridings Wood (SE5200), 24/8/1989, WAE det CJ; Hatfield Moor (SE7006), 7 9/9/1991, PS det CJ. A woodland species 6 (coniferous and deciduous) in litter 5 samples and under bark. 4 3 2 1

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umbrina (Gyllenhal, 1827)

Atomaria (Atomaria) umbrina (Gyllenhal, 1827)

3 WATSONIAN (0,1,0,1,0) Nationally Scarce. Rare in 2 YORKSHIRE Yorkshire with only two records on the YNU database: Scarborough (TA08), - 1 1918, RL, record unsubstantiated and 0 NY NZ NZ OV SD SE SE TA requires a voucher for confirmation; 9 Ellington Banks (SE2773), 2/9/1989, WAE 8 det CJ, 1 specimen by sweeping. Usually 7 associated with fungi in woodland (Johnson, 1993). 6 5 4 3 2 1

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vespertina Mäklin, 1853

NO RECORDS wollastoni Sharp, 1867

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 104 | P a g e

Subgenus ANCHICERA Thomson, C.G., 1863

apicalis Erichson, 1846

Atomaria (Anchicera) apicalis Erichson, 1846

3 WATSONIAN (11,6,31,3,0) Widespread and fairly 2 YORKSHIRE common throughout. In grassland, ground litter, compost heaps, piles of 1 putrefying grass cuttings and similar 0 NY NZ NZ OV SD SE SE TA media. (iii-xi). 9 8 7 6 5 4 3 2 1

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atra (Herbst, 1793)

Atomaria (Anchicera) atra (Herbst, 1793)

3 WATSONIAN (0,1,0,0,0) Nationally Scarce. There is a 2 YORKSHIRE single unsubstantiated record for the county: Whitby (NZ81), 20/3/1936, HB, in 1 Sphagnum. A voucher would be required 0 NY NZ NZ OV SD SE SE TA for confirmation, although this is a 9 species of marshes and fenland. 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 105 | P a g e atricapilla Stephens, 1830

Atomaria (Anchicera) atricapilla Stephens, 1830

3 WATSONIAN (29,21,136,16,6) A widespread and 2 YORKSHIRE common grassland species but little recorded from the high ground in the 1 north and west. In mowings, garden 0 NY NZ NZ OV SD SE SE TA compost heaps, flood refuse, woodland 9 pitfall traps and in fungal growths under 8 bark. (iii-xii). 7 6 5 4 3 2 1

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basalis Erichson, 1846

Atomaria (Anchicera) basalis Erichson, 1846

3 WATSONIAN (27,1,6,1,0) A species of wetlands, 2 YORKSHIRE marshes and fens. Recorded from very few sites in Yorkshire with the majority of 1 records resulting from extensive surveys 0 NY NZ NZ OV SD SE SE TA of Blacktoft Sands grazing marshes 9 (SE8323) in 2013, MLD, RJM, MH. In 8 ground litter and in pitfall traps. 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 106 | P a g e clavigera Ganglbauer, 1899

Atomaria clav igera Ganglbauer, 1899

3 WATSONIAN (1,0,0,0,1) RDBK. Very rare with only two records for the county: Bubwith (SE73), 2 YORKSHIRE 1919, Anon det CJ, specimen in 1 Manchester Museum; Masham (SE28), 0 NY NZ NZ OV SD SE SE TA 1919, WJF det CJ, specimen in 9 Manchester Museum. This species is 8 associated with grassland and has been 7 recorded from margins of ponds, riverbanks and carr land. 6 5 4 3 2 1

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fuscata (Schönherr, 1808)

Atomaria (Anchicera) fuscata (Schönherr, 1808)

3 WATSONIAN (67,31,159,35,6) A very common and 2 YORKSHIRE widespread species in a variety of habitats including ground litter in reed 1 beds, in compost heaps, flood refuse, 0 NY NZ NZ OV SD SE SE TA grass cuttings, horse dung and in fungi. 9 Probably our commonest Atomaria. 8 (i-xi, peak vi). 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 107 | P a g e fuscipes (Gyllenhal, 1808)

Atomaria fuscipes (Gyllenhal, 1808)

3 WATSONIAN (4,5,2,2,1) Very scarce with no recent records. A little-known species recorded 2 YORKSHIRE from old straw and by sieving ground 1 litter in woodland and arable margins. 0 NY NZ NZ OV SD SE SE TA (iv-ix). 9 8 7 6 5 4 3 2 1

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gutta Newman, 1834

NO RECORDS

hislopi Wollaston, 1857

Atomaria (Anchicera) hislopi Wollaston, 1857

3 WATSONIAN (0,1,0,0,0) RDBI. There is a single record 2 YORKSHIRE on the YNU database: Scarborough (TA08), Pre-1918, RL det CJ. This boreal 1 species is associated with microhabitats 0 NY NZ NZ OV SD SE SE TA (e.g. deer, sheep and grouse dung) in or 9 near coniferous woodland. (Johnson, 8 1993) 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 108 | P a g e lewisi Reitter, 1877

Atomaria (Anchicera) lewisi Reitter, 1877

3 WATSONIAN (74,30,301,46,9) Very common and 2 YORKSHIRE widespread and one of our most frequent Atomaria. A relative newcomer 1 to Britain (Allen, 1938) and described on 0 NY NZ NZ OV SD SE SE TA specimens from Japan, so presumably 9 introduced from the Far East. Recorded 8 from compost heaps, leaf litter, fungi and 7 piles of grass cuttings. (i-xi, peak vi). 6 5 4 3 2 1

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mesomela (Herbst, 1792)

Atomaria (Anchicera) mesomela (Herbst, 1792)

3 WATSONIAN (17,2,38,11,3) Widespread and recorded 2 YORKSHIRE mainly in the southern half of the county, this species inhabits wetlands, marshes, 1 fens, reedbeds, and margins of ponds 0 NY NZ NZ OV SD SE SE TA and lakes. (v-x). 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 109 | P a g e morio Kolenati, 1846

Atomaria (Anchicera) morio Kolenati, 1846

3 WATSONIAN (2,0,0,0,0) RDBI. Very rare in Yorkshire with two records from a single site: 2 YORKSHIRE Hornsea Mere (TA1947), 4/1980, PS det 1 CJ, one in a jackdaw nest; and 6/1980, RH 0 NY NZ NZ OV SD SE SE TA det CJ, 2 specimens in barn owl pellet. 9 Associated with bird nests in woodland, 8 it has been found in nest boxes in open situations. There are very few records 7 nationally. 6 5 4 3 2 1

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munda Erichson, 1846

Atomaria munda Erichson, 1846

3 WATSONIAN (1,1,8,3,0) Thinly distributed in the 2 YORKSHIRE southern half of the county, this grassland species is rather rare with no 1 recent records. Recorded from piles of 0 NY NZ NZ OV SD SE SE TA mouldy straw, old grass cuttings, and in 9 farmyard debris. 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 110 | P a g e nigripennis (Kugelann, 1794)

Atomaria (Anchicera) nigripennis (Kugelann, 1794)

3 WATSONIAN (0,0,2,0,0) Nationally Scarce. There are only two records on the YNU database, 2 YORKSHIRE both from the same site: Blackmoorfoot 1 – Edge Moor (SE1012), 6/8/2009 and 0 NY NZ NZ OV SD SE SE TA 8/8/2010, MLD. A grassland species 9 associated with heaps of decaying straw 8 and grass cuttings. 7 6 5 4 3 2 1

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nitidula (Marsham, 1802)

Atomaria (Anchicera) nitidula (Marsham, 1802)

3 WATSONIAN (84,33,331,39,13) Widespread and very 2 YORKSHIRE common – one of our most frequently encountered Atomaria. Recorded from a 1 wide range of habitats, from reed beds to 0 NY NZ NZ OV SD SE SE TA dry grassland, in compost heaps, old 9 straw piles and in putrefying grass 8 cuttings. 7 (iii-xi). 6 5 4 3 2 1

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ornata Heer, 1841

NO RECORDS peltata Kraatz, 1853

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 111 | P a g e pseudatra Reitter, 1888

NO RECORDS

pusilla (Paykull, 1798)

Atomaria (Anchicera) pusilla (Paykull, 1798)

3 WATSONIAN (3,4,7,7,4,2) A grassland species usually occurring in man-made habitats such as 2 YORKSHIRE piles of grass cuttings, compost heaps 1 and around farms. Our records are 0 NY NZ NZ OV SD SE SE TA distributed thinly and centrally in the 9 county; a tiny species which may be 8 easily overlooked in the field. (vi-x). 7 6 5 4 3 2 1

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rhenonum Kraatz, 1853

Atomaria rhenonum Kraatz, 1853

3 WATSONIAN (12,0,0,0,0) Nationally Scarce. A species 2 YORKSHIRE of saline environments and thus coastal in distribution. All Yorkshire records have 1 been noted from Spurn and the Humber 0 NY NZ NZ OV SD SE SE TA estuary where the species inhabits 9 shoreline reedbeds and saltmarshes. 8 (i-xi). 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 112 | P a g e rubella Heer, 1841

Atomaria (Anchicera) rubella Heer, 1841

3 WATSONIAN (21,7,52,10,1) A common and widely distributed species recorded from a wide 2 YORKSHIRE range of habitats including wetlands, 1 reedbeds, compost heaps, woodland 0 NY NZ NZ OV SD SE SE TA ground litter especially in damp 9 situations, and in piles of cut grass and 8 decaying straw. (iii-xi). 7 6 5 4 3 2 1

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rubida Reitter, 1875

NO RECORDS

rubricollis Brisout de Barneville, 1863

NO RECORDS

scutellaris Motschulsky, 1849

Atomaria (Anchicera) scutellaris Motschulsky, 1849

3 WATSONIAN (0,0,2,0,0) RDBK. We have two records from a single site: Ponderosa (SE2122), 2 YORKSHIRE 8/6/2009 and 18/6/2009, MLD teste CJ. 1 This species may be found in a range of 0 NY NZ NZ OV SD SE SE TA habitats including grassland, marshland 9 and in woodland pitfall traps and is 8 mainly confined to the south of England. 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 113 | P a g e testacea Stephens, 1830

Atomaria (Anchicera) testacea Stephens, 1830

3 WATSONIAN (87,22,216,38,19) Very common and widespread in grassland, compost heaps, 2 YORKSHIRE woodland floor litter, heaps of cut grass, 1 decaying straw and in fungi under bark, 0 NY NZ NZ OV SD SE SE TA often occurring in large numbers. (ii-xii). 9 8 7 6 5 4 3 2 1

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turgida Erichson, 1846

NO RECORDS

zetterstedti (Zetterstedt, 1838)

NO RECORDS

OOTYPUS Ganglbauer, 1899 globosus (Waltl, 1838)

Ootypus globosus (Waltl, 1838)

3 WATSONIAN (5,5,9,30) Widespread but always infrequent, a grassland species usually 2 YORKSHIRE occurring in the dung of large herbivores 1 especially horse dung. Sometimes 0 NY NZ NZ OV SD SE SE TA confused with Ephistemus globulus in the 9 field and in collections but the habitat is 8 quite different (see below). (iv-x). 7 6 5 4 3 2 1

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EPHISTEMUS Stephens, 1829 globulus (Paykull, 1798)

Ephistemus globulus (Paykull, 1798)

3 WATSONIAN (42,13,153,20,7) A common and very widespread grassland species often 2 YORKSHIRE found in man-made habitats such as 1 compost heaps, straw heaps and piles of 0 NY NZ NZ OV SD SE SE TA cut grass, especially in gardens and 9 around farms. May be confused in 8 appearance with Ootypus globosus in the 7 field and in collections (see note above). (iii-xi). 6 5 4 3 2 1

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reitteri Casey, 1900

NO RECORDS 62. Family EROTYLIDAE Latreille, 1802

Subfamily CRYPTOPHILINAE Casey, 1900

CRYPTOPHILUS Reitter, 1874 integer (Heer, 1841)

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 115 | P a g e

Subfamily EROTYLINAE Latreille, 1802

Tribe DACNINI Gistel, 1848

DACNE Latreille, 1796 bipustulata (Thunberg, 1781)

Dacne bipustulata (Thunberg, 1781)

3 WATSONIAN (11,14,47,24,2) A common species in fungi, usually brackets on deciduous 2 YORKSHIRE trees. A very widespread species from all 1 parts of the county. Various fungi are 0 NY NZ NZ OV SD SE SE TA mentioned in our records including 9 Pleurotus, Polyporus, Trametes and 8 Auricularia. (i-xii). 7 6 5 4 3 2 1

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rufifrons (Fabricius, 1775)

Dacne rufifrons (Fabricius, 1775)

3 WATSONIAN (4,9,9,8,2) Very similar to, but much less 2 YORKSHIRE frequent than, D. bipustulata and easily confused with that species, and found in 1 the same range of habitats. (vi-x). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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Tribe TRITOMINI Curtis, 1834

TRIPLAX Herbst, 1793 aenea (Schaller, 1783)

Triplax aenea (Schaller, 1783)

3 WATSONIAN (2,13,77,18,7) A fungus-dependent species with a broad central distribution, 2 YORKSHIRE it is usually associated with Pleurotus 1 species particularly in old woodland in 0 NY NZ NZ OV SD SE SE TA and pastures with isolated old trees. 9 (i-xii). 8 7 6 5 4 3 2 1

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lacordairii Crotch, 1870

NO RECORDS

russica (Linnaeus, 1758)

Triplax russica (Linnaeus, 1758)

3 WATSONIAN (2,0,20,0,0) Nationally Notable B. Restricted to bracket fungi on mature 2 YORKSHIRE trees in woodland. Some of our records 1 mention the fungus Fomes in association 0 NY NZ NZ OV SD SE SE TA with Betula Birch. (iv-x). 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 117 | P a g e scutellaris Charpentier, 1825

Triplax scutellaris Charpentier, 1825

3 WATSONIAN (0,2,0,0,0) RDB3. A rare species of the 2 YORKSHIRE northern half of Britain and associated with Pleurotus. We have two records on 1 our database: Ancliffe Wood (NZ4601), 0 NY NZ NZ OV SD SE SE TA 17/9/1970, EWA, on Pleurotus; Raincliffe 9 Woods (SE9988), 1986, MLD. 8 7 6 5 4 3 2 1

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TRITOMA Fabricius, 1775 bipustulata Fabricius, 1775

Tritoma bipustulata Fabricius, 1775

3 WATSONIAN (0,2,4,0,0) Nationally Notable A. Very 2 YORKSHIRE scarce in Yorkshire, the only recent record being: Potteric Carr, (SE5900), 1 29/4/2003, RJM, 1 ex swept in vicinity of 0 NY NZ NZ OV SD SE SE TA old wood lying on the ground. 9 8 7 6 5 4 3 2 1

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63. Family BYTURIDAE Gistel, 1848

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BYTURUS Latreille, 1796 ochraceus (Scriba, 1790)

Byturus ochraceus (Scriba, 1790)

3 WATSONIAN (10,9,52,9,2) Widely distributed in the county but few records from the high 2 YORKSHIRE ground of the north and west. The larvae 1 may be associated with Geum urbanum 0 NY NZ NZ OV SD SE SE TA Wood Avens in particular (Hammond et 9 al., 2019), the adults at flowers of Rubus 8 Brambles, Crataegus Hawthorn and others. (v-viii, peak vi). 7 6 5 4 3 2 1

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tomentosus (De Geer, 1774)

Byturus tomentosus (De Geer, 1774)

3 WATSONIAN (38,67,359,33,13) More widespread and frequent than B. ochraceus, developing 2 YORKSHIRE in the fruits of Rubus Brambles. Adults 1 found in flowers of other plants such as 0 NY NZ NZ OV SD SE SE TA Crataegus monogyna Hawthorn. (v-ix, 9 peak vi). 8 7 6 5 4 3 2 1

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64. Family BIPHYLLIDAE LeConte, 1861

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BIPHYLLUS Dejean, 1821 lunatus (Fabricius, 1787)

Biphyllus lunatus (Fabricius, 1787)

3 WATSONIAN (13,8,25,8,1) Widespread with a mainly 2 YORKSHIRE central distribution in Yorkshire. Recorded from Daldinia, mainly on 1 Fraxinus Ash, in woodland and parkland. 0 NY NZ NZ OV SD SE SE TA (iii-ix). 9 8 7 6 5 4 3 2 1

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DIPLOCOELUS Guérin-Méneville, 1844 fagi (Chevrolat in Guérin-Méneville, 1837)

Diplocoelus fagi (Chev rolat in Guérin-Ménev ille, 1837)

3 WATSONIAN (0,0,0,1,0) Nationally Notable B. There is 2 YORKSHIRE a single record on the YNU database: Studley Royal (SE2769), 8/6/2016, AG det 1 MLD. NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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Subfamily TEREDINAE Seidlitz, 1888

TEREDUS Dejean, 1835 cylindricus (Olivier, 1790)

NO RECORDS

OXYLAEMUS Erichson, 1845 cylindricus (Creutzer in Panzer, 1796)

NO RECORDS variolosus (Dufour, 1843)

NO RECORDS

Subfamily ANOMMATINAE Ganglbauer, 1899

ANOMMATUS Wesmael, 1835 diecki Reitter, 1875

NO RECORDS

duodecimstriatus (Müller, P.W.J., 1821)

Anommatus duodecimstriatus (Müller, P.W.J., 1821)

3 WATSONIAN (3,1,5,0,0) Nationally Notable A. This 2 YORKSHIRE little-known species, of cryptic and mainly subterranean habits, is recorded 1 from the south of the county. Found in 0 NY NZ NZ OV SD SE SE TA pitfall traps in woodland, in a garden 9 compost heap, in a haystack bottom and 8 in a window in a house. Subterranean 7 pitfall trapping is said to yield this species more frequently than the usual collecting 6 methods. 5 4 3 2 1

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Subfamily MURMIDIINAE Jacquelin du Val, 1858

MURMIDIUS Leach, 1821 ovalis (Beck, 1817)

NO RECORDS segregatus Waterhouse, C.O., 1876

NO RECORDS

Subfamily CERYLONINAE Billberg, 1820

CERYLON Latreille, 1802 fagi Brisout de Barneville, 1867

Cerylon fagi Brisout de Barnev ille, 1867

3 WATSONIAN (1,0,0,1,0) Nationally Notable B. There are only two (old) records for this species 2 YORKSHIRE in Yorkshire: Bubwith (SE73), 1917, GBW, 1 under bark of rotten ash in garden; 0 NY NZ NZ OV SD SE SE TA Harewood (SE3245), 7/12/1941, JHF, 9 under bark of felled beech. An ancient 8 woodland and wood pasture insect 7 feeding on decay fungi. 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 122 | P a g e ferrugineum Stephens, 1830

Cerylon ferrugineum Stephens, 1830

3 WATSONIAN (1,18,83,21,4) Widespread and fairly 2 YORKSHIRE commonly encountered in mature woodland and wood pasture, under the 1 bark of decaying deciduous trees, feeding 0 NY NZ NZ OV SD SE SE TA on fungi. (ii-xii). 9 8 7 6 5 4 3 2 1

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histeroides (Fabricius, 1792) Cerylon histeroides (Fabricius, 1792)

3 WATSONIAN (4,3,32,15,4) Less frequent than C. 2 YORKSHIRE ferrugineum and in the same types of habitat, but also recorded from conifers. 1 (i-xii). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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67. Family ALEXIIDAE Imhoff, 1856

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SPHAEROSOMA Stephens, 1832 pilosum (Panzer, 1793)

Sphaerosoma pilosum (Panzer, 1793)

3 WATSONIAN (0,0,1,0,0) Very rare nationally. This very tiny and cryptic beetle is recorded from 2 YORKSHIRE ground litter and in mosses and probably 1 feeds on fungal mycelia. We have a single 0 NY NZ NZ OV SD SE SE TA record on the YNU database: Wharncliffe 9 Wood (SK39), 6/1924, EGB. 8 7 6 5 4 3 2 1

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68. Family ENDOMYCHIDAE Leach, 1815 595

Subfamily ENDOMYCHINAE Leach, 1815

ENDOMYCHUS Panzer, 1795 coccineus (Linnaeus, 1758)

Endomychus coccineus (Linnaeus, 1758)

3 WATSONIAN (4,13,29,11,1) This very distinctive species inhabits fungal growths under 2 YORKSHIRE bark, and on cut ends of logs of Fagus 1 Beech, Malus Apples, Salix Willows and 0 NY NZ NZ OV SD SE SE TA Aesculus Horse-chestnuts. Widely but 9 rather centrally distributed. 8 (ii-xii, peaks vi and ix). 7 6 5 4 3 2 1

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Subfamily ANAMORPHINAE Strohecker, 1953

SYMBIOTES Redtenbacher, 1849 latus Redtenbacher, 1849

Symbiotes latus Redtenbacher, 1849

3 WATSONIAN (0,0,1,0,0) Nationally Notable B. There is a single record on the YNU database: 2 YORKSHIRE Whiteley Wood (SK3185), 1/9/1992, EJS. 1 Associated with dead wood fungi, 0 NY NZ NZ OV SD SE SE TA particularly on Ulmus Elms, Fraxinus Ash 9 and Populus Poplars. 8 7 6 5 4 3 2 1

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Subfamily MEROPHYSIINAE Seidlitz, 1872

HOLOPARAMECUS Curtis, 1833 caularum (Aubé, 1843)

NO RECORDS depressus Curtis, 1833

NO RECORDS singularis (Beck, 1817)

NO RECORDS

Subfamily LYCOPERDININAE Bromhead, 1838

LYCOPERDINA Latreille, 1807 bovistae (Fabricius, 1792)

NO RECORDS succincta (Linnaeus, 1767)

NO RECORDS

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Subfamily MYCETAEINAE Jacquelin du Val, 1857

MYCETAEA Stephens, 1829 subterranea (Fabricius, 1801)

Mycetaea subterranea (Fabricius, 1801)

3 WATSONIAN (28,5,42,3,1) Widely disrtibuted and fairly frequent in compost heaps, piles of 2 YORKSHIRE grass cuttings and around farmyards 1 where larvae and adults feed on fungi. 0 NY NZ NZ OV SD SE SE TA (iii-xii). 9 8 7 6 5 4 3 2 1

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69. Family COCCINELLIDAE Latreille, 1807

Subfamily COCCIDULINAE Mulsant, 1846

COCCIDULA Kugelann in Illiger, 1798

rufa (Herbst, 1783) Coccidula rufa (Herbst, 1783)

3 WATSONIAN (131,56,406,58,16) Very common in 2 YORKSHIRE wetland situations including reedbeds, marshes, pond margins and riverbanks 1 where aphids are the food source. (i-xii, 0 NY NZ NZ OV SD SE SE TA peak vi). 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 126 | P a g e scutellata (Herbst, 1783)

Coccidula scutellata (Herbst, 1783)

3 WATSONIAN (12,2,41,2,0) Much less frequent than C. rufa, but with apparently similar habitat 2 YORKSHIRE preferences and aphid food sources. 1 (i-xii, peak vi). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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RHYZOBIUS Stephens, 1829 chrysomeloides (Herbst, 1792)

NO RECORDS

litura (Fabricius, 1787)

Rhyzobius litura (Fabricius, 1787)

3 WATSONIAN (90,54,279,32,14) A grassland species 2 YORKSHIRE feeding on aphids. Usually collected by sweeping grasses and low plants, and 1 often very numerous in the summer 0 NY NZ NZ OV SD SE SE TA months. An aphid feeder, this species will 9 overwinter in favoured sites as adults, 8 and is generally distributed except for 7 the far west on the higher ground. (i-xi, peaks vi and ix). 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 127 | P a g e lophanthae (Blaisdell, 1892)

Rhyzobius lophanthae (Blaisdell, 1892)

3 WATSONIAN (1,0,5,0,0) A recent newcomer to our 2 YORKSHIRE fauna, native to Australia and used in southern Europe since 1908 to control 1 scale insects on crops. First recorded in 0 NY NZ NZ OV SD SE SE TA Britain in 1999 in Surrey, we have five 9 county records on the database: Tickton 8 (TA0641), 10/8/2017, WRD, one settled 7 on washing hung out to dry; Hatfield Moor (SE7004), 1/10/2015, AG det MLD, 6 in pitfall traps on peat moors; 5 Sprotbrough (SE5503), repeatedly in 4 Malaise trap in garden, 2017, 2018 and 3 2019, RJM teste MB. 2 1

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RODOLIA Mulsant, 1850 cardinalis (Mulsant, 1850)

NO RECORDS

STETHORUS Weise, 1885 punctillum (Weise, 1891)

Stethorus punctillum (Weise, 1891)

3 WATSONIAN (1,1,10,0,0) Our smallest ladybird, this species is recorded from gardens, 2 YORKSHIRE grassland, woodlands and orchards, 1 feeding on spider mites and aphids. We 0 NY NZ NZ OV SD SE SE TA have few records in Yorkshire and it may 9 be under-recorded owing to its small 8 size. It appears regularly in a garden Malaise trap in Sprotbrough (SE5503). 7 (v-x). 6 5 4 3 2 1

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CLITOSTETHUS Weise, 1885 arcuatus (Rossi, 1794) 599

Clitostethus arcuatus (Rossi, 1794)

3 WATSONIAN (0,0,4,0,0) RDB1. Very rarely recorded in Yorkshire with only four records from 2 YORKSHIRE two sites: Sprotbrough (SE559034), 1 9/2003, 4/2004, 9/2004, RJM, numerous 0 NY NZ NZ OV SD SE SE TA by beating Hedera helix Ivy on Fraxinus 9 excelsior Ash in a suburban garden; 8 Sprotbrough (SE553032), 14/7/2019, RJM, 1 ex in garden Malaise trap. 7 Possibly under-recorded owing to its 6 small size, this species feeds on whitefly. 5 4 3 2 1

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SCYMNUS Kugelann, 1794

Subgenus SCYMNUS Kugelann, 1794 femoralis (Gyllenhal, 1827)

Scymnus (Scymnus) femoralis (Gyllenhal, 1827)

3 WATSONIAN (3,2,4,0,0) Nationally Notable B. Very 2 YORKSHIRE infrequently encountered, this is an aphid feeder and in Yorkshire is probably 1 at the northern edge of its range in 0 NY NZ NZ OV SD SE SE TA Britain, although there is a single (pre- 9 1990) record for NE Scotland (Roy et al., 8 2011). Collected from low-growing 7 vegetation in fairly dry habitats. (iv-x). 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 129 | P a g e frontalis (Fabricius, 1787)

Scymnus (Scymnus) frontalis (Fabricius, 1787)

3 WATSONIAN (2,1,5,0,0) A mainly southern species with very few records on our database, 2 YORKSHIRE most of these being recent. This may 1 indicate that the species has been 0 NY NZ NZ OV SD SE SE TA establishing further north in recent years. 9 Repeatedly found in a garden Malaise 8 trap in Sprotbrough (SE5503) during 7 2017-2018. An aphid feeder of dry habitats this species is mainly located by 6 sweeping low-growing vegetation. (vi-ix). 5 4 3 2 1

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interruptus (Goeze, 1777)

NO RECORDS

jakowlewi Weise, 1892

NO RECORDS

nigrinus Kugelann, 1794

Scymnus (Scymnus) nigrinus Kugelann, 1794

3 WATSONIAN (0,2,1,2,0) Very scarce in Yorkshire with 2 YORKSHIRE only five records, excluding a Spurn (TA41) record dated 1948, WDH, which is 1 believed to be a mis-identification: 0 NY NZ NZ OV SD SE SE TA Osmotherley (SE4597), 1/8/1908, MLT, 9 on lower branches of pine; Bishopwood 8 (SE5633), pre-1920, WJF; Aberford 7 (SE43), 1939, PHa; Gundale Wood (SE7987), 30/6/1956, HF, on Larix Larch; 6 Haw Park (SE3615), 7/9/1968, JHF. This 5 species is an aphid feeder and associated 4 with coniferous woodland, and is also 3 recorded from gorse. 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 130 | P a g e rubromaculatus (Goeze, 1777)

NO RECORDS schmidti Fürsch, 1958

Scymnus (Scymnus) schmidti Fürsch, 1958

3 WATSONIAN (3,0,0,0,0) Nationally Notable B. There are three records on the YNU database: 2 YORKSHIRE Spurn – Wire Dump (TA4012), 1 19/7/1948, WDH det CJ; Spurn (TA4212), 0 NY NZ NZ OV SD SE SE TA 6/8/2006, WRD, one male in sand; 9 Haverfield Quarry (TA3220), 15/5/1999, 8 WRD det RJM, swept in tall grasses. A species of dry habitats feeding on aphids. 7 6 5 4 3 2 1

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Subgenus PULLUS Mulsant, 1846 auritus Thunberg, 1795

Scymnus (Pullus) auritus Thunberg, 1795

3 WATSONIAN (8,4,19,4,0) Widely distributed mainly in 2 YORKSHIRE the south of the county, this species is a specialist feeder on Phylloxeridae, a 1 group of aphid-like insects (Roy et al., 0 NY NZ NZ OV SD SE SE TA 2011) and our records indicate that oak is 9 the prime habitat. We have few recent 8 records. (iv-x) 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 131 | P a g e suturalis Thunberg, 1795

Scymnus (Pullus) suturalis Thunberg, 1795

3 WATSONIAN (13,20,22,3,2) Widespread and fairly frequent, an aphid feeder on conifers, 2 YORKSHIRE with a few adventitious records away 1 from conifers. (iii-xii). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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Subgenus NEOPULLUS Sasaji, 1971 haemorrhoidalis Herbst, 1797

Scymnus (Neopullus) haemorrhoidalis Herbst, 1797

3 WATSONIAN (3,3,16,2,0) A rather local species with a 2 YORKSHIRE central distribution in Yorkshire. Recorded from damp meadows, pond 1 and river margins, and grassland on 0 NY NZ NZ OV SD SE SE TA heavy soils. (ii-xi). 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 132 | P a g e limbatus Stephens, 1832

Scymnus (Neopullus) limbatus Stephens, 1832

3 WATSONIAN (0,0,5,1,0) Nationally Notable B. Very 2 YORKSHIRE local and scarce in Yorkshire with six records. Noted from deciduous trees in 1 wet situations, once under bark of fallen 0 NY NZ NZ OV SD SE SE TA rotting poplar. This species is a coccid 9 and aphid feeder. 8 7 6 5 4 3 2 1

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NEPHUS Mulsant, 1846 bisignatus (Boheman, 1850)

NO RECORDS quadrimaculatus (Herbst, 1783)

NO RECORDS redtenbacheri (Mulsant, 1846)

Nephus redtenbacheri (Mulsant, 1846)

3 WATSONIAN (15,10,23,7,1) A ladybird of varied 2 YORKSHIRE habitats including wetlands, dunes, heathland and grassland. Widely 1 distributed in the county. Taxonomic 0 NY NZ NZ OV SD SE SE TA confusion with limbatus may mean that 9 some of our wetland redtenbacheri 8 records may be referrable to limbatus. 7 Without vouchers we will never know. (iii-xi). 6 5 4 3 2 1

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CRYPTOLAEMUS Mulsant, 1853 montrouzieri Mulsant, 1853

Cryptolaemus montrouzieri Mulsant, 1853

3 WATSONIAN (1,0,0,0,0) We have a single record of this vagrant species: Skipsea (TA1755), 2 YORKSHIRE 7/1991, BC det MM, 2 specimens on car 1 bonnet in very hot weather (EMM NY NZ NZ OV 0 SD SE SE TA 130(1994): 45-46). An Australian species 9 introduced to control mealybug and 8 requiring hothouse conditions, so presumably an escape from such a 7 premises nearby. 6 5 4 3 2 1

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HYPERASPIS Dejean, 1836 pseudopustulata Mulsant, 1853

Hyperaspis pseudopustulata Mulsant, 1853

3 WATSONIAN (2,4,6,0,0) Nationally Notable B. This very local ladybird is thinly distributed in 2 YORKSHIRE the south and northeast of the county. 1 There are no recent records, the latest NY NZ NZ OV 0 SD SE SE TA being 1984. An aphid feeder in coastal 9 and wet habitats inland. 8 7 6 5 4 3 2 1

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Subfamily CHILOCORINAE Mulsant, 1846

PLATYNASPIS Redtenbacher, 1843 luteorubra (Goeze, 1777)

NO RECORDS

CHILOCORUS Leach, 1815 bipustulatus (Linnaeus, 1758)

Chilocorus bipustulatus (Linnaeus, 1758)

3 WATSONIAN (5,23,37,1,0) Widely distributed on heathy habitats, areas of dunes and in 2 YORKSHIRE scrub, wet areas (e.g. Fen Bog SE89), and 1 wet woodland with sallows and willows, 0 NY NZ NZ OV SD SE SE TA this species is a coccid feeder. (iii-xi). 9 8 7 6 5 4 3 2 1

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renipustulatus (Scriba, 1791)

Chilocorus renipustulatus (Scriba, 1791)

3 WATSONIAN (29,29,96,23,3) Widely distributed 2 YORKSHIRE throughout and fairly common in deciduous woodland, scrub and 1 grasslands, this coccid feeder has been 0 NY NZ NZ OV SD SE SE TA noted from various trees including Salix 9 Willow, Betula Birch and Alnus Alder, but 8 also from ruderal vegetation on verges 7 and by riverbanks. (iii-xi). 6 5 4 3 2 1

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EXOCHOMUS Redtenbacher, 1843 quadripustulatus (Linnaeus, 1758)

Exochomus quadripustulatus (Linnaeus, 1758)

3 WATSONIAN (24,16,171,17,2) Known as the Pine 2 YORKSHIRE Ladybird this name is somewhat of a misnomer because the species has been 1 recorded from a wide range of habitat 0 NY NZ NZ OV SD SE SE TA types including mixed woodland, 9 grassland, sea cliffs, marshland and 8 heaths. Also recorded regularly in a 7 Malaise trap in a suburban garden. (iii-xi). 6 5 4 3 2 1

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Subfamily COCCINELLINAE Latreille, 1807

COCCINULA Dobzhansky, 1925 quattuordecimpustulata (Linnaeus, 1758)

NO RECORDS

ANISOSTICTA Dejean, 1836 novemdecimpunctata (Linnaeus, 1758)

Anisosticta nov emdecimpunctata (Linnaeus, 1758)

3 WATSONIAN (59,10,302,41,7) An insect of marshy 2 YORKSHIRE grassland and reedbeds, the main distribution is in the southern half of the 1 county. Nationally there is a sharp cut-off 0 NY NZ NZ OV SD SE SE TA in distribution north of a Tyne-Solway 9 line, so Yorkshire is toward the northern 8 end of its range. The species develops in 7 a wide range of tall emergent water plants including Typha Bulrushes and 6 Glyceria Sweet-grasses. It is also a colour 5 changeling, with overwintering adults 4 becoming pale yellow and the spring- 3 emerging adults developing a darker red 2 hue, feeding on aphids. (i-xii). 1

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TYTTHASPIS Crotch, 1874 sedecimpunctata (Linnaeus, 1761)

Tytthaspis sedecimpunctata (Linnaeus, 1761)

3 WATSONIAN (13,1,38,2,0) Until 2000 very scarce in 2 YORKSHIRE Yorkshire and at its northern limit here. Since that date quite frequent in 1 grassland and scrub, being best located 0 NY NZ NZ OV SD SE SE TA by sweeping various low-growing plants, 9 and in ground litter towards the end of 8 the year. Feeds on nectar and pollen. 7 (v-x). 6 5 4 3 2 1

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MYZIA Mulsant, 1846 oblongoguttata (Linnaeus, 1758)

Myzia oblongoguttata (Linnaeus, 1758)

3 WATSONIAN (4,15,12,8,2) Widely and thinly 2 YORKSHIRE distributed throughout the county but with very few recent records. Noted from 1 Pinus sylvestris Scots Pine and Larix Larch 0 NY NZ NZ OV SD SE SE TA upon which this ladybird feeds on aphids. 9 (iii-x). 8 7 6 5 4 3 2 1

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MYRRHA Mulsant, 1846 octodecimguttata (Linnaeus, 1758)

Myrrha octodecimguttata (Linnaeus, 1758)

3 WATSONIAN (13,13,18,10,0) A conifer (mainly Scots pine) specialist with few recent records, 2 YORKSHIRE this species is widely distributed but with 1 no records from the far north and west 0 NY NZ NZ OV SD SE SE TA of the county. Aphids are the food 9 source. (iv-x). 8 7 6 5 4 3 2 1

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PROPYLEA Mulsant, 1846 quattuordecimpunctata (Linnaeus, 1758)

Propylea quattuordecimpunctata (Linnaeus, 1758)

3 WATSONIAN (164,126,1302,103,40) Very common, 2 YORKSHIRE widespread and ubiquitous in a wide range of habitat types including 1 woodland, grassland, saltmarsh and 0 NY NZ NZ OV SD SE SE TA scrub, and also in gardens. This species 9 may be collected by sweeping nettles, 8 hogweed, willowherbs and thistles. It is 7 also recorded from wetlands in the ground litter and by sweeping. An aphid 6 feeder. (iii-x, peak v/vi). 5 4 3 2 1

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CALVIA Mulsant, 1846 quattuordecimguttata (Linnaeus, 1758)

Calv ia quattuordecimguttata (Linnaeus, 1758)

3 WATSONIAN (42,65,346,67,31). Very widespread and fairly frequent on the foliage of 2 YORKSHIRE deciduous trees, mainly in woodland, but 1 also on isolated trees in grassland. We 0 NY NZ NZ OV SD SE SE TA also have records from wetlands such as 9 Askham Bog (SE5748), Denaby Ings 8 (SE5000) and Breary Marsh (SE2541). Food sources are aphids and psyllids. 7 (iv-x, peak v). 6 5 4 3 2 1

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VIBIDIA Mulsant, 1846 duodecimguttata (Poda, 1761)

NO RECORDS

HALYZIA Mulsant, 1846 sedecimguttata (Linnaeus, 1758)

Halyzia sedecimguttata (Linnaeus, 1758)

3 WATSONIAN (14,14,95,25,25) Widespread and 2 YORKSHIRE frequent in woodland and in gardens and hedgerows where there are trees. Before 1 2000 this species was a rarity in the 0 NY NZ NZ OV SD SE SE TA county with only three records, all pre- 9 1935, and since 2000 becoming suddenly 8 noticeable everywhere. The food source 7 is mildews on the foliage of Fraxinus Ash, Quercus Oak, Betula Birch and Crataegus 6 Hawthorn. (iii-xi, peak vi). 5 4 3 2 1

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PSYLLOBORA Dejean, 1836 vigintiduopunctata (Linnaeus, 1758)

Psyllobora v igintiduopunctata (Linnaeus, 1758)

3 WATSONIAN (87,33,465,37,8) Widespread and fairly 2 YORKSHIRE common everywhere except in the far west. This mildew feeder may be swept 1 from various low-growing plants and 0 NY NZ NZ OV SD SE SE TA beaten from the foliage of a wide range 9 of deciduous trees, and occurs regularly 8 in a garden Malaise trap. Said to be 7 decreasing nationally (Roy et al., 2011), this is not the experience in Yorkshire 6 where we receive a steady supply of 5 modern records. (iii-xii, peak viii). 4 3 2 1

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ANATIS Mulsant, 1846 ocellata (Linnaeus, 1758)

Anatis ocellata (Linnaeus, 1758)

3 WATSONIAN (6,28,73,14,5) A conifer specialist and an 2 YORKSHIRE aphid feeder, this species is widely recorded in Yorkshire except for the 1 southeast and the far west. There are 0 NY NZ NZ OV SD SE SE TA rather few recent records so may be 9 declining here. (iii-x, peak v). 8 7 6 5 4 3 2 1

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APHIDECTA Weise, 1899 obliterata (Linnaeus, 1758)

Aphidecta obliterata (Linnaeus, 1758)

3 WATSONIAN (11,41,52,15,16) A conifer specialist 2 YORKSHIRE recorded mainly from Larix Larch, but we also have a record from Pinus sylvestris 1 Scots Pine. A feeder on coccids, this 0 NY NZ NZ OV SD SE SE TA species is widespread and fairly frequent 9 in suitable habitat. (iv-xi). 8 7 6 5 4 3 2 1

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HIPPODAMIA Dejean, 1836

Subgenus HIPPODAMIA Dejean, 1836 tredecimpunctata (Linnaeus, 1758)

Hippodamia tredecimpunctata (Linnaeus, 1758)

3 WATSONIAN (1,2,3,0,0) RDBK. There are six records of 2 YORKSHIRE this elusive and rare species on the YNU database. This species occurs in 1 wetlands, riverbanks and reedbeds, and 0 NY NZ NZ OV SD SE SE TA is an aphid feeder, with no recent 9 records. 8 7 6 5 4 3 2 1

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Subgenus ADONIA Mulsant, 1846 variegata (Goeze, 1777) Hippodamia (Adonia) v ariegata (Goeze, 1777)

3 WATSONIAN (31,7,127,13,4) Nationally Notable B. 2 YORKSHIRE This aphid feeder which is becoming more frequent in Yorkshire may be found 1 in fairly dry habitats such as waste 0 NY NZ NZ OV SD SE SE TA ground, peat moors, gravel pits and 9 arable margins. It has also been taken 8 regularly in flight interception traps in 7 suburban gardens. There are few records for the higher ground in the north and 6 west of the county. The Notable B 5 designation requires revision in the light 4 of the number of records, especially 3 since 2000. (iv-x). 2 1

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COCCINELLA Linnaeus, 1758 hieroglyphica Linnaeus, 1758

Coccinella hieroglyphica Linnaeus, 1758

3 WATSONIAN (6,20,64,4,2) Associated with heather Calluna, recent records are from the 2 YORKSHIRE North York Moors, Thorne/Hatfield 1 Moors and Strensall Common. The food 0 NY NZ NZ OV SD SE SE TA sources include the heather aphid Aphis 9 callunae, and the eggs and larvae of the 8 heather leaf beetle Lochmaea suturalis and other chrysomelids in the genera 7 Galerucella and Altica (Roy et al., 2011). 6 None of our records of which there are 5 approximately 100 specifies a food 4 source or feeding observation. (iv-x). 3 2 1

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magnifica Redtenbacher, 1843

Coccinella magnifica Redtenbacher, 1843

3 WATSONIAN (0,0,4,0,0) Nationally Notable A. We have four records of this elusive ladybird: 2 YORKSHIRE Derwent Reservoir (SK1791), 5/9/1971, 1 PBM, specimen in Sheffield Museum; 0 NY NZ NZ OV SD SE SE TA Unsliven Bridge (SK2599), 20/6/1977, 9 WJL; Wharncliffe Heath (SK2997), 8 1/5/1997, DW; Sprotbrough (SE5503), 17/7/2005, RJM, sweep netting along 7 woodland edge. This species is 6 myrmecophilus and associated with 5 Formica rufa, and may be under- 4 recorded owing to its similarity to the 3 very common Coccinella septempunctata. The adults and larvae 2 feed on aphids in and around the ant 1 nest. SD SE SE TA 0 SJ SK SK TF 9 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 1/9/2018 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4

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NO RECORDS septempunctata Linnaeus, 1758

Coccinella septempunctata Linnaeus, 1758

3 WATSONIAN (334,144,1818,126,69) The very common and ubiquitous 7-spot ladybird, and an 2 YORKSHIRE aphid feeder found in almost any habitat 1 type. (ii-xii). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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undecimpunctata Linnaeus, 1758

Coccinella undecimpunctata Linnaeus, 1758

3 WATSONIAN (51,10,159,7,6) Widely distributed but most records concentrated in the south 2 YORKSHIRE of VC63. Seems to be declining with few 1 recent records, and most recent records 0 NY NZ NZ OV SD SE SE TA are coastal. An aphid feeder. (iii-x). 9 8 7 6 5 4 3 2 1

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ADALIA Mulsant, 1846 bipunctata (Linnaeus, 1758)

Adalia bipunctata (Linnaeus, 1758)

3 WATSONIAN (84,55,858,68,28) A widespread and very common aphid-feeding species. 2 YORKSHIRE Recorded in a wide range of habitats and 1 may be seen in all months of the year. 0 NY NZ NZ OV SD SE SE TA (ii-xii, peak vi). 9 8 7 6 5 4 3 2 1

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decempunctata (Linnaeus, 1758)

Adalia decempunctata (Linnaeus, 1758)

3 WATSONIAN (57,56,413,53,23) Widely distributed and common in hedgerows and on the foliage 2 YORKSHIRE of deciduous trees. This species feeds on 1 aphids. (ii-xii, peak v). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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HARMONIA Mulsant, 1846 axyridis (Pallas, 1773)

Harmonia axyridis (Pallas, 1773)

3 WATSONIAN (83,14,316,35,44). This species was first 2 YORKSHIRE recorded in Britain in 2003 and has since spread rapidly and is now (2019) 1 widespread and common in Yorkshire. A 0 NY NZ NZ OV SD SE SE TA naturalised Asian species and a feeder on 9 aphids and coccids, it also feeds on other 8 coccinellids, which has given the beetle a 7 bad press. However, observations in Yorkshire indicate that other ladybird 6 populations have not declined markedly 5 (except possibly Coccinella 4 undecimpunctata) as a result of the 3 spread of H. axyridis, and in fact numbers 2 of Coccinella septempunctata seem to have increased in the period 2015-2019. 1 Adults of axyridis appear to be noticeable 0 SD SE SE TA SJ SK SK TF especially in ix-x. (RJM pers. obs). 9 (i-xii, peak ix) 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 1/9/2018 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4

quadripunctata (Pontoppidan, 1763)

Harmonia quadripunctata (Pontoppidan, 1763)

3 WATSONIAN (4,1,38,2,0) Widespread in the southern half of the county but always rather 2 YORKSHIRE scarce, with few recent reports. Feeding 1 on aphids this species is recorded from 0 NY NZ NZ OV SD SE SE TA conifers, heathland and scrub. (iv-x). 9 8 7 6 5 4 3 2 1

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Subfamily EPILACHNINAE Mulsant, 1846

HENOSEPILACHNA Li, 1961 argus (Geoffory in Fourcroy, 1762)

NO RECORDS

SUBCOCCINELLA Guérin-Méneville, 1844 vigintiquattuorpunctata (Linnaeus, 1758)

Subcoccinella v igintiquattuorpunctata (Linnaeus, 1758)

3 WATSONIAN (57,10,36,1,0) Our Yorkshire records indicate a distribution in the eastern half 2 YORKSHIRE of the county although the map shown in 1 Roy et al., 2011: 70 depicts the presence 0 NY NZ NZ OV SD SE SE TA of records from VC64 that we do not 9 possess. This species is phytophagous on 8 Silene Campions and Arrhenatherum elatius False oat-grass. (iii-x). 7 6 5 4 3 2 1

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Subfamily CORYLOPHINAE LeConte, 1852

SERICODERUS Stephens, 1829 brevicornis Matthews, A., 1890

Sericoderus brev icornis Matthews, A., 1890

3 WATSONIAN (0,1,1,0,0) There are two records on the YNU database: Robin Hood’s Bay 2 YORKSHIRE (NZ9506), 17/8/2006, RGB, sieved from a 1 hay bale; Fenwick (SE5815), 3/8/2016, 0 NY NZ NZ OV SD SE SE TA RJM, in compost heap. This species has 9 only recently been recognised as British 8 and distinct from S. lateralis. 7 6 5 4 3 2 1

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lateralis (Gyllenhal, 1827)

Sericoderus lateralis (Gyllenhal, 1827)

3 WATSONIAN (11,1,29,2,2) Widespread in the southern 2 YORKSHIRE half of the county. Recorded from compost heaps and under piles of rotting 1 straw where there is mould and warmth. 0 NY NZ NZ OV SD SE SE TA Refer to the note under S. brevicornis – 9 our records for lateralis now have a 8 doubtful element owing to the bringing 7 forward of brevicornis to the British list. (v-x). 6 5 4 3 2 1

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CORYLOPHUS Stephens, 1833 cassidoides (Marsham, 1802)

Corylophus cassidoides (Marsham, 1802)

3 WATSONIAN (3,0,2,0,0) A rarely recorded species in 2 YORKSHIRE Yorkshire, there are five records on the YNU database: Hornsea Mere (TA1947), 1 30/5/1915, TS, see Nat. 1915:403, and 0 NY NZ NZ OV SD SE SE TA 26/10/1996, RJM, in Phragmites litter; 9 Pulfin Bog (TA0544), 28/4/2011, RJM, in 8 ground litter at edge of drain and in 7 Glyceria beds; Hatfield Moor (SE6904), 28/5/2005, AG det MLD, in ground litter. 6 There are subfossil records from Cowick 5 Moat (SE6520) and Thorne Moor (SE71) 4 of Bronze Age to medieval date. This tiny 3 species may be easily overlooked in litter 2 samples in reedbeds and similar situations and seems to occur in long- 1 established and unpolluted wetland 0 SD SE SE TA SJ SK SK TF situations. 9 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 25/10/2019 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4

sublaevipennis Jacquelin du Val, 1859

Corylophus sublaev ipennis Jacquelin du Val, 1859

3 WATSONIAN (6,2,0,0,0) Our eight records are all 2 YORKSHIRE strictly coastal, except for a specimen from Hollym Carrs NR (TA3224). Usually 1 brachypterous except for occasional 0 NY NZ NZ OV SD SE SE TA fully-winged specimens which may 9 account for inland records (Bowestead, 8 1999:83). Recorded from ground litter 7 and tussocks usually in dry situations. 6 5 4 3 2 1

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Subfamily ORTHOPERINAE Jacquelin du Val, 1857

ORTHOPERUS Stephens, 1829 aequalis Sharp, 1885

Orthoperus aequalis Sharp, 1885

3 WATSONIAN (0,0,1,0,0) RDBK. We have a single record on the YNU database: Wentworth 2 YORKSHIRE Park (SK3996), 11/5/2008, EJS det CJ, 2 1 exx under peeling bark of poplar log. 0 NY NZ NZ OV SD SE SE TA Associated with moulds under bark of 9 hardwoods (Bowestead, 1999:116). 8 Owing to the minute size of these beetles, specimens may be overlooked 7 very easily in samples of many mouldy 6 substrates. 5 4 3 2 1

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atomarius (Heer, 1841)

NO RECORDS atomus (Gyllenhal, 1808)

Orthoperus atomus (Gyllenhal, 1808)

3 WATSONIAN (0,0,3,1,0) There are four records on the 2 YORKSHIRE YNU database: Healaugh (SE5147), 3/1968, JHF, bred from old ivy with 1 Ochina ptinoides; Rother Valley Country 0 NY NZ NZ OV SD SE SE TA Park (SK4584), 27/6/2004, WAE; Stainton 9 Churchyard (SK5593), 4/10/2000, RJM 8 det CJ, sieved from mouldy grass 7 cuttings; Blackmoorfoot (SE1012), 18/4/2010, MLD det CJ. At the time of 6 the 1968 record the identification of 5 species within this genus was poorly 4 defined owing to the lack of present-day 3 knowledge and literature, so this record 2 may not be correct, and no voucher has been traced. 1

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Orthoperus brunnipes (Gyllenhal, 1808)

3 WATSONIAN (9,2,0,0,0) RDB3. Rarely reported in 2 YORKSHIRE Yorkshire with eleven records. A species of fens or other types of wetland habitat 1 and recorded in accumulations of mouldy 0 NY NZ NZ OV SD SE SE TA vegetation on wet ground. 9 8 7 6 5 4 3 2 1

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corticalis (Redtenbacher, 1849)

Orthoperus corticalis (Redtenbacher, 1849)

3 WATSONIAN (1,1,3,0,0) With five records on the YNU 2 YORKSHIRE this is a species of old woodland associated with moulds and fungi: 1 Skipwith Common (SE6537), 7/9/2000, 0 NY NZ NZ OV SD SE SE TA MLD det CJ; Strensall (SE6559), 9 31/7/2002, WRD det CJ, in Polyporus 8 betulinus with blue mould, in woodland; 7 Langsett (SK2199), 19/11/2006, EJS det CJ, in small bracket fungus on Salix 6 Willows; Pot Ridings (SE5200), 5 29/5/2007, AG det CJ, bred from fungi 4 collected in woodland; Wentworth Park 3 (SK3996), 11/5/2008, EJS det CJ, under 2 loose bark of log of unidentified tree species. 1

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Orthoperus nigrescens Stephens, 1829

3 WATSONIAN (1,0,1,0,0) Nationally Notable B. We 2 YORKSHIRE have two records on the YNU database: Skipwith Common (SE6537), 1984, RGB; 1 Langsett (SK2199), 19/11/2006, EJS, from 0 NY NZ NZ OV SD SE SE TA decaying bracket fungus on sallows. A 9 species of deciduous woodland and 8 found in fungi and in mouldy situations in 7 the ground layer. (Bowestead, 1999:104) 6 5 4 3 2 1

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Subfamily RYPOBIINAE Paulian, 1950

RYPOBIUS LeConte, 1852 praetermissus Bowestead, 1999

NO RECORDS 71. Family LATRIDIIDAE Erichson, 1842 612

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 152 | P a g e

Subfamily LATRIDIINAE Erichson, 1842

LITHOSTYGNUS Broun, 1886 serripennis Broun, 1914

Lithostygnus serripennis Broun, 1914

3 WATSONIAN (14,0,13,1,0) As with many of our tiny 2 YORKSHIRE Coleoptera, this species is probably under-recorded. Noted from compost, 1 old straw, mouldy grass cuttings, in old 0 NY NZ NZ OV SD SE SE TA fungi, in piles of faggots, in bat roost 9 debris and at light in houses. There are 8 few recent records. 7 6 5 4 3 2 1

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STEPHOSTETHUS LeConte, 1878 alternans (Mannerheim, 1844)

NO RECORDS angusticollis (Gyllenhal, 1827)

Stephostethus angusticollis (Gyllenhal, 1827)

3 WATSONIAN (1,0,2,0,0) There are three records on the 2 YORKSHIRE YNU database: Wath-on-Dearne (SE4300), - 1920, EGB; Honley Wood 1 (SE1211), 21/7/1985, JRB det MLD; Spurn 0 NY NZ NZ OV SD SE SE TA – Clubley’s Field (TA4115), 18/6/1990, 9 MLD, collected by evening sweeping. A 8 species of mouldy substrates. 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 153 | P a g e caucasicus (Mannerheim, 1844)

NO RECORDS

lardarius (De Geer, 1775)

Stephostethus lardarius (De Geer, 1775)

3 WATSONIAN (75,40,186,40,5) Widely distributed and common in compost heaps, piles of grass 2 YORKSHIRE cuttings and straw, in old fungi in 1 woodland and grassland and under bark 0 NY NZ NZ OV SD SE SE TA of deciduous trees. 9 8 7 6 5 4 3 2 1

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CARTODERE Thomson, C.G., 1859

Subgenus CARTODERE Thomson, C.G., 1859 constricta (Gyllenhal, 1827)

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 154 | P a g e

Subgenus ARIDIUS Motschulsky, 1866 bifasciata (Reitter, 1877)

Cartodere (Aridius) bifasciata (Reitter, 1877)

3 WATSONIAN (129,47,478,50,10) Widespread and 2 YORKSHIRE extremely common in ground litter, in flight, in old straw heaps, compost piles 1 and in mouldy grass cuttings. A recent 0 NY NZ NZ OV SD SE SE TA newcomer to Britain, first observed in 9 1951 and a species described from 8 Australia (Allen, 1951), this beetle is now 7 ubiquitous. Occurs in all kinds of putrefying mouldy material and is often 6 collected by sweeping and also in 5 Malaise traps. 4 3 2 1

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nodifer (Westwood, 1839)

Cartodere (Aridius) nodifer (Westwood, 1839)

3 WATSONIAN (81,41,378,48,10) A widespread and 2 YORKSHIRE common species sometimes occurring with the closely-related C. bifasciatus, 1 and in the same types of habitat. NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 155 | P a g e norvegica (Strand, 1940)

Cartodere (Aridius) norv egica (Strand, 1940)

3 WATSONIAN (0,0,1,0,0) We have a single record on the YNU database: Thorne Waterside 2 YORKSHIRE (SE6714), 14/8/1982, PS, in ground litter. 1

NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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LATRIDIUS Herbst, 1793 assimilis Mannerheim, 1844

Latridius assimilis Mannerheim, 1844

3 WATSONIAN (3,2,16,3,1) Infrequent and centrally 2 YORKSHIRE distributed in the county, this species, known in earlier checklists as 1 pseudominutus, is recorded from fungi, 0 NY NZ NZ OV SD SE SE TA ground litter in reedbeds, straw heaps, 9 compost heaps and piles of grass 8 cuttings. (iii-xi). 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 156 | P a g e consimilis Mannerheim, 1844

Latridius consimilis Mannerheim, 1844

3 WATSONIAN (1,0,1,3,0) Nationally Scarce. Very scarce 2 YORKSHIRE in the county with only five records known to us: Skipwith Common 1 (SE6537), 17/5/1948, EWA; Askham Bog 0 NY NZ NZ OV SD SE SE TA (SE5748), 4/4/1980, PH, on Salix Willows; 9 Blackmoorfoot (SE1012), 23/8/1980, 8 MLD; Studley Park (SE2969), 19/6/1989, 7 RJM det CJ, in debris in old logs; Hackfall Wood (SE2377), 15/6/1996, RJM, in fungi 6 on stumps by R. Ure; 5 4 3 2 1

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minutus (Linnaeus, 1767)

Latridius minutus (Linnaeus, 1767)

3 WATSONIAN (1,4,17,1,1) Widely but thinly distributed 2 YORKSHIRE throughout. Recorded from Daldinia on Fraxinus Ash, in hay bales, in decaying 1 Polyporus on Acer pseudoplatanus 0 NY NZ NZ OV SD SE SE TA Sycamore, in farmyard debris and in 9 grass mowings. (v-xii). 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 157 | P a g e porcatus Herbst, 1793

Latridius porcatus Herbst, 1793

3 WATSONIAN (17,1,10,2,1) Widely distributed but 2 YORKSHIRE infrequent, and recorded from compost heaps, old straw bales where mould and 1 some warmth occur. (iii-x). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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THES Semenov, 1910 bergrothi (Reitter, 1880)

Thes bergrothi (Reitter, 1880)

3 WATSONIAN (0,1,2,2,0) A very rare species in the 2 YORKSHIRE county of which there are five records on the YNU database, all very old: 1 Doncaster (SE50), 1907, HHC; Bradford 0 NY NZ NZ OV SD SE SE TA (SE13), 1910, JWC; Scarborough (TA08), 9 1914, RAT det ECH, in cellar; Leeds 8 (SE33), 5/1917, SM, in cellar; Leeds 7 (SE33), 1935, CES. A synanthropic species in stored products. 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 158 | P a g e

ENICMUS Thomson, C.G., 1859 brevicornis (Mannerheim, 1844)

Enicmus brev icornis (Mannerheim, 1844)

3 WATSONIAN (3,0,4,0,1) Nationally Scarce. Very scarce 2 YORKSHIRE with few records reported – noted from ground litter at the edge of a reservoir, in 1 piles of garden refuse, in a compost heap 0 NY NZ NZ OV SD SE SE TA and by general sweeping in woodland. 9 8 7 6 5 4 3 2 1

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fungicola Thomson, C.G., 1868

Enicmus fungicola Thomson, C.G., 1868

3 WATSONIAN (1,0,9,2,0) Very scarce with no recent 2 YORKSHIRE records. Recorded from piles of cut vegetation and in ground litter in damp 1 situations where the substrate is mouldy. 0 NY NZ NZ OV SD SE SE TA (v-xii). 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 159 | P a g e histrio Joy & Tomlin, 1910

Enicmus histrio Joy & Tomlin, 1910

3 WATSONIAN (9,8,12,5,2) Widespread in the county 2 YORKSHIRE but not a common species. May be overlooked in favour of E. transversus, 1 and dissection of males is recommended. 0 NY NZ NZ OV SD SE SE TA In piles of cut vegetation and in compost 9 heaps where moulds are present. (v-x). 8 7 6 5 4 3 2 1

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rugosus (Herbst, 1793)

Enicmus rugosus (Herbst, 1793)

3 WATSONIAN (0,4,0,1,0) Nationally Scarce. This species 2 YORKSHIRE is only known from two sites in Yorkshire, with five records: Avenue Wood Temple 1 Newsam (SE3731), 9/7/1977, PS; 0 NY NZ NZ OV SD SE SE TA Wykeham Forest (SE9588), 7/2001 – 9 9/2001, LA. This latter record would 8 require vouchers to confirm. No habitat 7 details are given with our records but the species is associated with mouldy 6 substrates in woodland. 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 160 | P a g e testaceus (Stephens, 1830)

Enicmus testaceus (Stephens, 1830)

3 WATSONIAN (2,1,7,2,0) A very scarce species, known 2 YORKSHIRE to be associated with slime moulds in woodland. One of our records mentions 1 “Reticularia” as a host. NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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transversus (Olivier, 1790)

Enicmus transv ersus (Oliv ier, 1790)

3 WATSONIAN (78,26,211,36,13) By far the commonest 2 YORKSHIRE member of the genus, this species is widespread throughout the county with 1 over 360 records. Often abundant in leaf 0 NY NZ NZ OV SD SE SE TA litter, old fungi, compost heaps, and in 9 mouldy substrates in piles of cut grass 8 and hay. (ii-xi, peak vi). 7 6 5 4 3 2 1

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DIENERELLA Reitter, 1911

Subgenus CARTODEREMA Reitter, 1911 clathrata (Mannerheim, 1844)

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 161 | P a g e ruficollis (Marsham, 1802)

Dienerella ruficollis (Marsham, 1802)

3 WATSONIAN (9,1,6,1,0) A species of compost heaps, piles of old straw and in similar situations 2 YORKSHIRE where mould and warmth are present. 1 Also noted from rotting seaweed on the NY NZ NZ OV 0 SD SE SE TA shoreline. Very infrequently reported 9 probably owing to its small size and 8 sluggish movements. (iv-xi). 7 6 5 4 3 2 1

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vincenti Johnson, 2007

Dienerella elongata (Curtis, 1830)

3 WATSONIAN (0,1,6,1,0) Very scarce in the county, and as with all these tiny beetles probably 2 YORKSHIRE grossly under-recorded. Noted from 1 fungi on Betula Birch and Taxus baccata 0 NY NZ NZ OV SD SE SE TA Yew, in a celler and in old straw debris 9 feeding on moulds. This taxon, D. 8 elongata (Curtis, 1830) is now known as vincenti Johnson, 2007 in the latest 3rd 7 edition checklist (Duff, 2018) 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 162 | P a g e

Subgenus DIENERELLA Reitter, 1911 argus (Reitter, 1884)

NO RECORDS

elegans (Aubé, 1850)

NO RECORDS

filiformis (Gyllenhal, 1827)

Dienerella filiformis (Gyllenhal, 1827)

3 WATSONIAN (0,1,2,0,0) Very rarely recorded with only three Yorkshire records known to us: 2 YORKSHIRE Scarborough (TA08), pre-1918, RL; 1 Huddersfield (SE1416), 11/1970, EWA, in 0 NY NZ NZ OV SD SE SE TA fungus Merulius lacrymans in a cellar; 9 Crosland Moor (SE1114), 27/10/1984, 8 MLD, in a house. The Scarborough record would require a voucher for 7 confirmation. Although not part of our 6 remit, there is a Derbyshire record from 5 Holbrook Marsh (SK4481), 1/7/1993, DW 4 det JDC, just on the border with VC63. 3 2 1

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filum (Aubé, 1850)

Dienerella filum (Aubé, 1850)

3 WATSONIAN (1,0,3,0,0) Mainly synanthropic with four 2 YORKSHIRE known records: Hull (TA02), 1915, TS; Huddersfield (SE1416), 18/3/1970, EWA, 1 in a house; Crosland Moor (SE1114), 0 NY NZ NZ OV SD SE SE TA 27/7/1983 and 28/8/1997, MLD, in 9 house. 8 7 6 5 4 3 2 1

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NO RECORDS

ADISTEMIA Fall, 1899 watsoni (Wollaston, 1871)

Adistemia watsoni (Wollaston, 1871)

3 WATSONIAN (0,10,0,0,0) All of our ten records are 2 YORKSHIRE from within buildings in the York area, once in a house in central York (SE6052), 1 2011, RM, and then repeatedly in sticky 0 NY NZ NZ OV SD SE SE TA traps in storage areas at the Yorkshire 9 Museum, 2016-2018, AP det RJM. 8 Probably strictly synanthropic. 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 164 | P a g e

Subfamily CORTICARIINAE Curtis, 1829

CORTICARIA Marsham, 1802 alleni Johnson, 1974

NO RECORDS

crenulata (Gyllenhal, 1827)

Corticaria crenulata (Gyllenhal, 1827)

3 WATSONIAN (19,2,10,0,0) Scarce but widely distributed. Early records were mainly 2 YORKSHIRE coastal but now identified from inland 1 sites. This species appears to like 0 NY NZ NZ OV SD SE SE TA wetlands, peatmoors in wetter areas, 9 damp pastures, saltmarshes and dunes 8 on the shore. (v-ix). 7 6 5 4 3 2 1

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dubia Dajoz, 1970

Corticaria dubia Daj oz, 1970

3 WATSONIAN (0,0,1,0,0) We have a single record on our database: Thorne Moor (SE7215), 2 YORKSHIRE 20/6/1990, DH, in pitfalls on peat moor. 1 Associated with slime moulds on trees, a 0 NY NZ NZ OV SD SE SE TA voucher specimen has not been traced 9 for this record, so must remain as 8 requiring confirmation. 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 165 | P a g e elongata (Gyllenhal, 1827)

Corticaria elongata (Gyllenhal, 1827)

3 WATSONIAN (12,10,32,5,0) Widespread and fairly frequent. Recorded from various habitats 2 YORKSHIRE including sweeping in grassland, in dead 1 leaves, in old straw, in reedbeds, in water 0 NY NZ NZ OV SD SE SE TA traps in woodland and at light indoors. 9 (ii-xii). 8 7 6 5 4 3 2 1

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fagi Wollaston, 1854

Corticaria fagi Wollaston, 1854

3 WATSONIAN (0,0,1,0,0) RDBI. We have a single record 2 YORKSHIRE of this species: Holmfirth (SE1508), 26/6/1990, RBA det CJ. No habitat details 1 are provided with this record, but the 0 NY NZ NZ OV SD SE SE TA species is associated with dead wood 9 (Alexander, 2002) 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 166 | P a g e ferruginea Marsham, 1802

Corticaria ferruginea Marsham, 1802

3 WATSONIAN (1,1,3,0,0) A county rarity with five 2 YORKSHIRE records: Scarborough (TA08), 1925, GBW; Haw Park (SE3615), 22/7/1965, 1 EWA; Park Wood Elland (SE1122), 0 NY NZ NZ OV SD SE SE TA 22/9/1985, MLD; Spurn Warren Cottage 9 (TA4115), 22/6/1990, MLD, in pile of 8 grass cuttings; Fosterhouses (SE6714), 7 30/7/2011, RJM, by general sweeping in uncut hay meadow. 6 5 4 3 2 1

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fulva (Comolli, 1837)

Corticaria fulv a (Comolli, 1837)

3 WATSONIAN (1,1,5,1,0) Rare in the county with eight 2 YORKSHIRE records. Most of these records indicate synanthropic situations, houses, cellars 1 etc., except for a single record from 0 NY NZ NZ OV SD SE SE TA woodland requiring confirmation with a 9 voucher specimen. There are no recent 8 records. 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 167 | P a g e impressa (Olivier, 1790)

Corticaria impressa (Oliv ier, 1790)

3 WATSONIAN (84,11,144,14,0) Widespread and fairly 2 YORKSHIRE common especially in the southern half of the county. (i-xii, peak vi). Recorded by 1 general sweeping low vegetaion, in pitfall 0 NY NZ NZ OV SD SE SE TA traps in arable and woodland situations, 9 piles of grass cuttings, compost heaps 8 and in saltmarsh, presumably a fungus 7 feeder. 6 5 4 3 2 1

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inconspicua Wollaston, 1860

Corticaria inconspicua Wollaston, 1860

3 WATSONIAN (0,0,2,0,0) There are two records on the 2 YORKSHIRE YNU database: Ravensknowle Park (SE1616), 11/1964, EWA, in fungi; 1 Crosland Moor (SE1114), 29/6/1981, 0 NY NZ NZ OV SD SE SE TA MLD det CJ. 9 8 7 6 5 4 3 2 1

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longicollis (Zetterstedt, 1838)

NO RECORDS longicornis (Herbst, 1783)

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 168 | P a g e polypori Sahlberg, J., 1900

NO RECORDS

punctulata Marsham, 1802

Corticaria punctulata Marsham, 1802

3 WATSONIAN (13,2,30,3,0) Widespread in the southern 2 YORKSHIRE half of the county, predominantly in wet locations such as fens, drain sides and 1 reedbeds. Recorded from piles of cut 0 NY NZ NZ OV SD SE SE TA reeds, in grass tussocks, but also in 9 manure heaps. (iii-xii). 8 7 6 5 4 3 2 1

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rubripes Mannerheim, 1844

Corticaria rubripes Mannerheim, 1844

3 WATSONIAN (0,1,1,0,0) We have only one firm and another, unsubstantiated, record of this 2 YORKSHIRE elusive species in Yorkshire: Dalby Forest 1 (SE8888), 28/6/1969, EWA, between 0 NY NZ NZ OV SD SE SE TA freshly-cut pine poles; Thorne Moor 9 (SE71), 1982, unknown collector and 8 determiner – record unsubstantiated. 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 169 | P a g e serrata (Paykull, 1798)

Corticaria serrata (Paykull, 1798)

3 WATSONIAN (1,1,0,0,0) Very rare in Yorkshire with only two records: Saltburn (NZ62), 1900, 2 YORKSHIRE MLT; Filey (TA18), 1914, ECH. Both these 1 records are very old and the 0 NY NZ NZ OV SD SE SE TA determinations may not be correct in the 9 light of modern knowledge and 8 identification literature. 7 6 5 4 3 2 1

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umbilicata (Beck, 1817)

Corticaria umbilicata (Beck, 1817)

3 WATSONIAN (1,6,1,1,0) Very rarely reported with nine 2 YORKSHIRE records from five sites. Apparently a hygrophilous species, and recorded 1 variously in a Juncus Rushes flush in wet 0 NY NZ NZ OV SD SE SE TA woodland, at Deschampsia Hair-grass 9 roots and in ground litter at a drain edge. 8 7 6 5 4 3 2 1

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CORTICARINA Reitter, 1880 curta (Wollaston, 1854)

NO RECORDS

Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 170 | P a g e lambiana (Sharp, 1910)

NO RECORDS

latipennis (Sahlberg, J., 1871)

NO RECORDS

minuta (Fabricius, 1792)

Corticarina minuta (Fabricius, 1792)

3 WATSONIAN (87,66,289,61,17) Known from earlier checklists as C. fuscula, this is a 2 YORKSHIRE widespread and very common species of 1 a wide range of habitat types including 0 NY NZ NZ OV SD SE SE TA compost heaps, piles of cut vegetation, 9 on tree foliage, in reed beds, sweeping in 8 grassland, taken in Malaise traps, in straw bales and on blossoms of 7 Crataegus monogyna Hawthorn. 6 (iii-xii, peak v). 5 4 3 2 1

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similata (Gyllenhal, 1827)

Corticarina similata (Gyllenhal, 1827)

3 WATSONIAN (0,4,1,1,0) Very rarely reported with only 2 YORKSHIRE six records. Habitats recorded include coniferous and deciduous woodland, 1 otherwise little is known about this 0 NY NZ NZ OV SD SE SE TA species. 9 8 7 6 5 4 3 2 1

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Atlas of Yorkshire Coleoptera - part 10 – Cleroidea and Cucujoidea 171 | P a g e truncatella (Mannerheim, 1844)

NO RECORDS

CORTINICARA Johnson, 1975 gibbosa (Herbst, 1793)

Cortinicara gibbosa (Herbst, 1793)

3 WATSONIAN (114,77,410,41,8) Widespread and very common particularly on flowers of 2 YORKSHIRE Crataegus monogyna Hawthorn, in leaf 1 litter and by general sweeping in 0 NY NZ NZ OV SD SE SE TA grassland, on roadside verges and in 9 gardens. (i-xii, peaks vi and ix). 8 7 6 5 4 3 2 1

SD SE SE TA 0 SJ SK SK TF 9 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 28/9/2018 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4

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MELANOPHTHALMA Motschulsky, 1866 distinguenda (Comolli, 1837)

NO RECORDS suturalis (Mannerheim, 1844)

NO RECORDS transversalis (Gyllenhal, 1827)

Melanophthalma transv ersalis (Gyllenhal, 1827)

3 WATSONIAN (5,0,0,0,0) There are five records 2 YORKSHIRE on the YNU database, all from 1 Spurn: Caravan Field (TA4115), NY NZ NZ OV 0 SD SE SE TA 18/7/1948, WDH; Kilnsea Warren 9 (TA4115), 21/6/1990, MLD, 8 collected in refuse on Humber 7 shore; Beacon Lagoons (TA4116),

6 27/8/2003, WRD; Spurn (TA4212),

5 16/8/2006, WRD, on sand in

4 Marram tussock; Shore near Blue

3 Bell (TA4115), 12/9/2016, WRD,

2 one under Phragmites australis 1 Common Reed among dunes. 0 SD SE SE TA SJ SK SK TF 9 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 8/11/2019 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4

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MIGNEAUXIA Jacquelin du Val, 1859 lederi Reitter, 1875

NO RECORDS

Species removed from the Yorkshire list

Ebaeus pedicularius (Linnaeus, 1758) – a record exists in the YNU database, dated 1979, recorder Leslie Barringer, location SK445935. This must surely be an error (see Hodge and Jones 1995:85 and Hyman and Parsons 1992:359). The record is maintained for reference but flagged as Incorrect until a voucher specimen is traced. There are three specimens in the Doncaster Museum’s British Coleoptera collection but devoid of data.

Dasytes virens (Marsham, 1802) – there are two records of this species on the YNU database both from the Doncaster area. Specimens in the Doncaster Museum with the same data appear to be Dasytes plumbeus, so therefore the records for virens are considered incorrect.

Acknowledgements

My thanks are due to Colin Johnson, lately of the Manchester Museum where he was Head of Entomology. Colin’s detailed and long-term research into the Cryptophagidae and help and advice provided to me and other entomological colleagues by Colin since the 1980s have been invaluable to our study and understanding of this fascinating but often difficult group of beetles. Thanks are also due particularly to Mike Denton (additionally for proof-reading), to Bill Dolling and to other members of the YNU Entomological Section for advice, assistance and/or the provision of records, and to Dr. Paul Buckland for his important work on the cataloguing of the Doncaster Museum British Coleoptera collection. Also deserving of thanks is Mike Weideli of Littlefield Consultancy who has helped enormously with the software that has allowed me to import data from Recorder 6 into DMAP and to permit analysis of records into Vice-County distributions for this and for previous parts of the Atlas.

References

Alexander, K.N.A., 2002. English Nature Reports 467. The invertebrates of living & decaying timber in Britain and Ireland – a provisional annotated checklist.

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Allen, A.A., 1938. Two British species of Atomaria (Col., Cryptophagidae) new to the European list. Entomologist’s Mon. Mag. 74(1938):82-83.

Allen, A.A., 1951. Lathridius bifasciatus Reitt. (Col., Lathridiidae) an Australian beetle found wild in Britain. Entomologist’s Mon. Mag. 87(1951):114.

Bowestead, S., 1999. A revision of the Corylophidae (Coleoptera) of the West Palaearctic Region. Museum d’histoire naturelle, Geneve.

Buckland P 2018 A catalogue of British Coleoptera in the Doncaster Museum collections: https://www.heritagedoncaster.org.uk/collections/

Coombs, C.W. and Woodroffe, G.E., 1955. A revision of the British species of Cryptophagus (Herbst) (Coleoptera:Cryptophagidae). Trans. R. Ent. Soc. Lond. 106 (6):237-282.

Cooter, J., 1999. On ‘English’ names for British Coleoptera. Coleopterist 8 (1): 43-45.

Dandy J.E. 1969. Watsonian Vice-counties of Great Britain. The Ray Society, London

Denton, M.L., 1995. The Beetles of Spurn Peninsula. Yorkshire Museum/NYCC, York.

Duff, A.G., (ed.), 2012 Checklist of Beetles of the British Isles, 2nd edition. Pemberley Books.

Duff, A.G., (ed.), 2018 Checklist of Beetles of the British Isles, 3rd edition. Pemberley Books (Publishing).

Hodge, P.J., and Jones, R.A. 1995. New British Beetles – Species not in Joy’s practical handbook. British Entomological and Natural History Society.

Hyman, P.S., and Parsons, M.S., 1992. A review of the scarce and threatened Coleoptera of Great Britain, part 1. JNCC, Peterborough.

Hyman, P.S. and Parsons, M.S., 1994. A review of the scarce and threatened Coleoptera of Great Britain, part 2. JNCC, Peterborough.

James, T.J., 2018. Beetles of Hertfordshire. Hertfordshire Natural History Society.

Johnson, C., 1972. Entomologist 125:126-129 Epuraea udumbrata Mannerheim and E. biguttata (Thunberg) (Col., Nitidulidae) New to Btitain.[description figures and key introducing E terminalis to the British list (as adumbrata)]

Johnson, C., 1988. Notes on some British Cryptophagus Herbst (Coleoptera: Cryptophagidae), including confusus Bruce new to Britain. Entomologist’s Gazette 39(1988):329-335.

Johnson, C., 1993. Provisional atlas of the Cryptophagidae-Atomariinae (Coleoptera) of Britain and Ireland. Biological Records Centre, Monks Wood, Huntingdon.

Key, R.S., 1999. More on ‘noddy names’ for British beetles. Coleopterist 8 (2): 94.

Kirk-Spriggs, A.H., 1996. Pollen Beetles – Coleoptera: Kateretidae and Nitidulidae (Meligethinae). Handbk. Ident. Br. Insects Vol 5 part 6a. Royal Entomological Society, London.

Pope, R.D., 1977. Kloet & Hincks. A Check List of British Insects. Part 3: Coleoptera and Strepsiptera. Second revised edition. Handbooks Ident. Brit. Insects volume 11(3), pp. xiv+105. Royal Entomological Society, London.

Roy, H.E., Brown, P.M.J., Frost, R. & Poland, R.L., 2011. The Ladybirds (Coccinellidae) of Britain and Ireland. Biological Records Centre, NERC-CEH, Wallingford, Oxon.

Watson, H.C., 1883. 2nd. ed. Topographical Botany. Quaritch, London.

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Errors and omissions The author would be pleased to receive feedback regarding any errors or omissions noted by users of this Atlas.

R J Marsh FRES YNU Coleoptera recorder 20 November 2019

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