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Fifth International Symposium on Trichoptera Series Entomologica FIFTH INTERNATIONAL SYMPOSIUM ON TRICHOPTERA SERIES ENTOMOLOGICA EDITOR K.A. SPENCER VOLUME 39 PROCEEDINGS OF THE FIFTH INTERNATIONAL SYMPOSIUM ON TRICHOPTERA LYON, FRANCE 21-26 JULY 1986 Edited by MICHEL BOURNAUD and HENRI TACHET 1987 DR W. JUNK PUBLISHERS ~ .. a member of the KLUWER ACADEMIC PUBLISHERS GROUP •• DORDRECHT / BOSTON / LANCASTER .~ Distributors for the United States and Canada: Kluwer Academic Publishers, P.O. Box 358, Accord Station, Hingham, MA 02018-0358, USA for the UK and Ireland: Kluwer Academic Publishers, MTP Press Limited, Falcon House, Queen Square, Lancaster LAI IRN, UK for all other countries: Kluwer Academic Publishers Group, Distribution Center, P.O. Box 322, 3300 AH Dordrecht, The Netherlands Library of Congress Cataloging in Publication Data International Symposium on Trichoptera (5th: 1986 : Lyon, France) Proceedings of the fifth International Symposium on Trichop-tera, Lyon (France) 21-26 July 1986. (Series entomologiea i v. 39) Includes indexes. 1. Caddis-flies--Congresses. I. Bournaud, Michel. II. Tachet, Henri. III. Arens, Marie Fran~oise. IV. Title. V. Series. QL516.157 1986 595.7'45 87-4218 ISBN-13: 978-94-010-8294-5 e-ISBN-13: 978-94-009-4043-7 001: 10.1007/978-94-009-4043-7 Copyright © 1987 by Dr W. Junk Publishers, Dordrecht. Softcover reprint of the hardcover I st edition 1987 All rights reserved. No part of this publication may be reproduced, stored in a retrieval system, or transmitted in any form or by any means, mechanical, photocopying, recording, or otherwise, without the prior written permission of the publishers, Dr W. Junk Publishers, P.O. Box 163, 3300 AD Dordrecht, The Netherlands. PROCE EDINGS OF THE FIFTH INTERNATIONAL SYMPOSIUM ON TRICHOPTE RA LYON (FRANCE) 21 - 26 JULY 1986 Thremm. gallicum Ml:Lachkm Laboratoire de Biologie animale et Ecologie et Unite associee au C.N.R.S. n° 367 "Ecologie des eaux douces" Institut d' Analyse des Systemes Biologiques et Socio-Economiques Universite Claude Bernard, LYON I AAA)\ 1\ 1. B.S. SVENSSON - 2. S.W. HAMIL TON - 3. G. BECKER - 4. G. VAN DER VELDE - 5. R.H. WAGNER - 6. B. HIGLER - 7. K. KUMANSKI - 8. H. TACHET - 9. B. CELLOT - 10. P.W. SCHEFTER - 11. M. BOURNAUD - 12. M.A. GONZALEZ - 13. C. DENIS - 14. L. BOTOSANEANU - 15. S.T. DARLINGTON - 16. T. ITO - 17. B. GULLEFORS - 18. P. CHANTARAMONGKOL - 19. G. GISLASON - 20. M.G. GLAPSKA - 21. R.C. PETERSEN 22. C. TOMASZEWKI - 23. L. PETERSEN - 24. B. STATZNER - 25. P. ZWICK - 26. C. ROUX - 27. M.I. CRICHTON - 28. J. MURILLO I ROYO - 29. C. OTTO - 30. M.A. PUIG - 31. M. GONZALEZ del TANAGO - 32. A. NEBOISS - 33. D. GARCIA del JALON - 34. N.A. ERMAN - 35. M.C. MOLtES - 36. V.H. RESH - 37. M.C. BICCHIERAI - 38. J. MAJECKI - 39. F. CIANFICCONI - 40. G. SPINNELLI SATTA - 41. C. CORALLINI SORCETTI 42. K. TANIDA - 43. A. BOUZIDI - 44. N. WILLIAMS - 45. O. FOSSATI - 46. J.O. SOLEM - 47. F.M. GIBON - 48. L. RECASENS I ALBALADEJO - 49. M.J. WEVERS - 50. T. PITSCH - 51. P. STROOT - 52. P. GREEN - 53. J.S. RICHARDSON - 54. R.N. VINEYARD - 55. F. VAILLANT - 56. A. WELLS - 57. G.P. MORETTI - 58. P. BAGGE - 59. M. BALES - 60. R.M. BADCOCK - 61. S.W.L. da TERRA - 62. S.J. WILKINSON - 63. S. WILLIAMS 64. G. MARLIER - 65. O.S. FLINT - 66. J.A. GORE - 67. R.W. WISSEMAN - 68. M. OEMKE - 69. H. MALICKY - 70. E. PETERSSON - 71. D. DUDGEON 72. A.P. NIMMO - 73. J.C. MORSE - 74. B.J. ARMITAGE - 75. W. WICHARD - 76. T. ANDERSEN - 77. B. FAESSEL - 78. S. SMITH - 79. R. LEUVEN 80. J. WARD - 81. J. WEAVER III - 82. D. WILLIAMS - 83. O. WILLIAMS - 84. P. USSEGLIO-POLATERA - 85. L. COGERINO - 86. J. O'CONNOR. Not in photograph: E. CASTELLA, B. FROCHOT, J. GIUDICELLI, R. HOUP, F. SCHMID, A. TAVARES, L.X. TIAN. Registered but not present: N.H. ANDERSON, W.G. DOWNS, P. GATTAPONI, V.A. NELSON, C. SIEGENTHALER, G. WIGGINS. < VI CONTENTS Preface .•••..•..........••.•.•....••••••••.•.••••...•.••.•.•.•........••••...•....•.••.••.•...••...... VII List of participants ••.••.....••••..•.•.••.•...•••.•••••.••••••••.•.••..••••.......•..........•.•••• IX List of poster displays and oral communications .....•...•...•••..•••••....••.••... XIII Editorial comment .•••••. ••.••••••.••••.••..•....•••.......•..........••..•....•.•.•..••.••••.•.• XVIII List of contributions •.•.••..•.•.••......••..•...••••.••...•••.....••••...••••.•........•.•...... XIX I - Morphology and Anatomy •.•.......••..•.••••.......•••.••.•.•......•••••••.•.•..•....•. 2 - Taxonomy •...•...••••...•...•....•.•.•••..••••.••...•..••••••••••.••••.......•••••.••••••.•••. 23 3 - Fossil forms .••.•••....•..•.......••....•....•..••...•.••.....•••..•••.•••••••••• •••••• •...••. 55 lj. - Biogeography ....•.•••.•.•.•••••.••.•••...••...•••..•...••..........•.•........•.••••.•.....• 71 5 - Phylogeny •• •••.•••••.•.•...••••••••••.••••••••.•••.••.•••••••• •.•••••....•.•.••.•.•...•.....•• 131 6 - Biology and Physiology..................... ........... ....................... .... ....... 151 7 - Light traps and flight patterns ••••....•.•••••••••.•• •.••.•••..• .••.••.••..•.•••..... 195 8 - Ecology ••••...•........•.••....••..•.••••...••••.•..•••••.••..••.............•...•••••..•••.•.• 235 Author index •••..••...•.•••••••••.•.....•••••..••. •• ••••••.••••...•.••••••.•.•••••...•••.•••••••.. 381 Geographical index .•••.•••... .••• .•••. ...•...•.•.•. .•.....••.....•. ..•....•••.•..•....•.•.•..•. 383 Subject index ••• •..••.••••.....••... ..••••• ••.. •• •• •••• ••••.•• •••••.• •.•.•...•..••••..••••.•..•... 385 Taxonomic index •• •..•••••...••• •...•••.••••.•.•..•.•...•.•....•......•.......•.•.••••••••...•.. 387 VII PREFACE When we offered to host the 5th International Symposium on Trichoptera in Lyon in July 1986, we knew that for us the great Adventure was about to begin. On that July morning in 1983 in Clemson (South Carolina), we could only dimly imagine what the future held in store. One of the worst moment came, in fact, when we were told by th official scientific authorities that no subsidies would be forthcoming. This refusal meant that we would be unable to give any financial help to those colleagues with only modest means at their disposal. Let us hope that the publication of the Proceedings will convince certain French scientists, and the powers that be, that the study of Trichoptera represents a valid and exciting field of research. The American raid on Tripoli, which did not at the time seem to have any link with Trichoptera, threatened to act as a deterrent as far as the North American participants were concerned. However, the fears expressed by some of our colleagues (and not without reason, when we consider the series of terrorist attacks later carried out in France) were soon allayed, although the organisers of the Congress must confess to having kept a discreet eye open for any suspicious brown paper parcels or unattended luggage. In spite of these difficulties and other minor problems, the 5th Symposium attracted 89 participants from 23 countries. This success was due in part to the impact of the previous Symposia (1974 Lunz am See, Austria ; 1977 Reading, England ; 1980 Perugia, Italy ; 1983 Clemson, U.S.A.), and also to an ever-increasing desire to participate. 20 posters and 57 lectures were presented. Two field trips took place, one to the Massif du Pilat (on the eastern edge of the Massif Central) and the other to the Massif de la Chartreuse (Prealps). These excursions gave participants the opportunity to get to know the landscapes and rivers of the Lyonnais region (as well as its culinary attractions). During the frrst excursion some participants were even lucky enough to collect a few specimens of Thrernrntl gallicum McL., whose larva was the symbol of the Symposium. The guest programme was followed by about forty courageous souls who discovered the charms (hidden, in the case of the traboules) of Lyon and its surroundings with a maximum of good humour and a minimum (we hope) of wear and tear. This symposium brought together many participants from previous Symposia, but also welcomed for the first time several colleagues from the Far East: Dr T. Ito and K. Tanida from Japan, Dr Dudgeon from Hong Kong, Dr P. Chantaramongkol from Thailand, and Dr LX. Tian from the People's Republic of China. Dr Tian's presence was the result of a process which had begun in Clemson ; at the time Dr Morse's efforts had had no immediate results, but the try, try and try again principle proved successful in the end. Dr Tian was able to come to Lyon thanks to a travel grant from the Agricultural University of Nanjing, and the dynamism of Dr Liu Dajun, the President of his University, whom we would like to thank here. The scientific success (lectures, posters, and especially unofficial discussions) of a Symposium like ours depends essentially on the personalities of the participants. The quality of its organisation and cultural content (in the widest sense of the term) depends on the work carried out by the members of the Organising Committee, among whom we would like to thank in particular : Mme Chantal Roux who organised the wine and cheese workshop on Monday July 21st, thereby giving back to the word Symposium its original meaning (Symposium in Greek means banquet). Mr Bernard Cellot who was in charge of the poster session. Professor Franr;ois Vaillant from Grenoble who organised the field trip to the Massif de la Chartreuse. Michele Pariset and Glyn and Philippe Thoiron who worked on translations for the Symposium, and provided linguistic security for us and for the participants while it was in progress, thus ensuring that our beautiful French language did not pose any major
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