APPENDIX 1

FLORISTIC CHECKLIST OF THE NOURAGUES AREA PierreBelbenoit, OdilePoncy, DanielSabatier, Marte- Francoise Prevost, BernardRiera, Patrick Blanc, DenisLarpin and CorinneSarthou

The list includes data gathered at the 'Inselberg site' on an area of about 10 km2 (Charles­ Dominique Chapter I). It was compiled mainly from (a) the 'AUBLET' database of the 'Herbier de Guyane' (CAY) (Hoffet al. 1989); (b) the tree inventory in high forest (see Poncy et al. Chapter 4); (c) the botanical study of the inselberg (rock-savanna, Sarthou Chapter 6; low forests, Larpin Chapter 5). A few additional tree species recorded only from a 6-ha plot studied in the Saut-Parare area are also included (see Poncyet al. Chapter 4). The botanical information was provided by several botanists and ecologists, including S. Barrier, C. Julliot, D. Larpin, O. Poncy, M.-F. Prevost, B. Riera, D. Sabatier, B. Simmen, I-F. ViIliers (tree inventories), and also P. Blanc, M. Born, A. Cockle, G. Cremers, C. Feuillet, S. Gonzalez, J.-J. de Granville, M. Hoff, C. Sarthou, S.Y. Zhang. Nomenclatural and systematic arrangements follow the Checklist ofthe ofthe Guianas (, Surinam, ) (Boggan et al. 1997), except when taxonomic novelties have been published subsequently. Incompletely identified taxa are mentioned only when they are sufficiently well documented to be sure that they are different from the other taxa. When collection references are missing or not reliable (e.g. records based on field observations without a voucher collection, or on a poor collection), names of taxa are placed between brackets. Differences may occur between this list and data presented in certain papers ofthis book. For example, as far as Myrtaceae are concerned, only 23 species out ofthe 33 mentioned by Larpin (Chapter 5) are listed. The 10 excluded taxa were considered insufficiently documented to be included in the checklist. The occurrence of the species in three major habitats is given: high dense forest (H), low forest on the slopes of the inselberg (L), and rock-savanna (RS). 'H' comprises various vegetation types, including mixed transitional forest at the foothill of the inselberg, swamp forest with Euterpe palms, liana forest, and bamboo forest, the two latter eventually being considered (cf Poncy et al. Chapter 4) as having originated from ancient anthropic influence. Information on habitat type may be missing, for instance when not indicated by the collector, or when the record of the species is based on seeds from the seed bank or from faeces, or on a specimen collected in a disturbed area (such as around the camp). The comparative importance of the species in these three major habitats was estimated using the information at our disposal. The symbol (x) indicates that the species can be expected in that habitat, based either on our knowledge

.;ynamicnncl pI_-lRimal interactions in a neotropical rainforest, 301-341. -, er ACldemic Publislrcrs, Printed inthe Netherlands, 302 P. Belbenoit et al. from the Nouragues area (e.g., a species recorded in RS and H is predicted to be present also in L) or from other sites in French Guiana and adjacent areas.

Forest types H = Taxon recorded from the high forest L = Taxon recorded from the low forest RS = Taxon recorded from the rock savanna (x) = Taxon assumed to be present in high and low forest habitats

Lifeforrn Emergent tree = tree > 35 m Tall tree = tree 25-35 m Medium tree = tree 15-25 m Smail tree = tree 10-15 m Tree = single-stemmed woody ~ 5 cm DBH and > 2 m tall Treelet = single-stemmed woody plant < 5 cm DBH and > 2 m tall Shrub = woody plant < 2 m tall Liana = woody c1imber Vine = non-woody climber Epilithic = attached on rocks Epiphyte = growing on other plants, not rooted in the soit Hemi-epiphyte = growing for only a part ofthe life as an epiphyte Saprophyte = plant feeding on dead organic matter F/oristic checkJist o/the Nouragues area 303

FLORISTIC CHECKLIST OF THE NOURAGUES AREA HL RS L1FE FORM

SPERMATOPHYTA Dicotyledones Acanthaceae Ruellia inflata L.e. Rich. x Terrestrial herb Anacardium spruceanum Benth. ex Engl. x TaI! Iree Astronium ulei Mattick Emergent tree Spondias mombin L. X Emergent Iree Tapirira bethanniana J.D. Mitchell X TaU lree Aubl. XXX SmaIl lree Tapirira oblUsa (Benth.) lD. Mitchell X SmaU lree Thyrsodium guianense Sagot ex March. X X TaI! tree Thyrsodium puberulum J.D. Mitchell & Daly X TaU Iree Thyrsodium spruceanum Benth. X Medium tree Annonaceae Anaxagorea dolichocarpa Sprague & Sandw. X Treelet Anaxagoreaprinoit/es (Dunal) A. OC. X Treelel Annona densicoma Mart. X SmaIl tree Annona haematantha MiQ. X Tree Cardiopetalum surinamense R.E. Fries X (X) Treele! Crematosperma brevipes (OC. ex Dunal) R.E.Fries X Treelet Duguetia cadaverica Huber X Treelet Duguetia calycina Benoist X SmaIl tree Duguetia eximia Diels X Treelet Duguetia inconspiCUQ Sagot X Tree/et Duguetia pycnastera Sandw. X SmaIl Iree Duguetia surinamensis R.E. Fries X Mediumtree EpJredranthus guianensis R.E. Fries X Medium lree Fusaea longifolia (Aubl.) Safford X Medium lree Guatterio ouregou (Aubl.) Dunal X Treelel Guatteria punclata (Aubl.) RA Howard X Tree [Gualteria cf. schomburgkiana Mart.]. after Larpin 1993 X Medium lree Gualteria sp. nov. X Oxandra asbeckii (Pulle) R.E. Fries XX X Medium tree Pseudoxandra cuspidata Maas X Mediumlree Rollinia exsucca (DC. ex Dunal) A. OC. X Treelet Unonopsis pe"Olletii (A. OC.) R.E. Fries X Medium lree Unonopsis rujescens (Saill.) R.E. Fries X Medium tree Unonopsis stipitata Diels XX Tree Xylopia nitida Dunal X TaI! tree 304 P. Belbenoit et al.

FLORISTIC CHECK LIST OF THE NOURAGUES AREA HL RS LIFE FORM

Apocynaceae Ambe/ania acida Aubl. x Trcelet Aspidasperma album (Vahl) Benoist ex Pichon X X Tall tree Aspidosperma carapanauba Pichon X Trce Aspidosperma cruentum Woodson X Tree Aspidosperma he/slonei van Donselaar X Tall Iree Aspidosperma marcgravianum Woodson X Tall Iree Aspidosperma ob/ongum A. OC. X Emergent trce Aspidosperma sandwilhianum Markgr. x Medium tree Couma guianellSis Aubl. X Medium lree Geissospermum /aevis (Vell.) Miers X Tall tree Himalanlhus arlicu/OlUS (Vahl) Woodson X Medium Irce Himalanlhus bracleOlus (A. OC.) Woodson X Medium tree Lacmellea acu/ea/a (Ducke) Monachino X Small tree Lacmelleaj10ribunda (Poepp.) Benth. & J.O. Hook. X Small Iree Macoubea guianellSis Aubl. X Tall Iree [Mandevilla hirsula (A. Rich.) K. Schum.l. alter Sarthou 1992 (x) (X) X Liana Mandevilla subspicala (Vahl) Markgr. XX Liana Odonladenia perrollelii (A. OC.) Woodson X Liana Odonladenia punclicu/osa (A. Rich.) Pulle X Parahancornia!ascicu/a/a (Poir.) Benoist ex Pichon X Tall Iree Tabernaemonlana allenuala (Miers) Urb. X Trce Tabernaemontana diSlicha A. OC. XX Trce Tabernaemonlana macroca/yx Muell. Arg. (x) X Tree Aquifoliaceae lIex sp. X Araliaceae Oreopanax capilalus (Jacq.) Decne. & Planch. X Liana SchefJ1era decaphylla (Seem.) Hanns X X Tall trce SchefJ1era mor%loni (Aubl.) Maguire. Steyenn. & Frodin X Talltree Aristolochiaceae Arislo/ochia iquilellSis O.c. Schmidt X Liana Asclepiadaceae Ditassa cf. cOllSanguinea Decne. X Male/ea saslrei Morillo X Vine Asteraceae F/eischmannia microslemon (Cass.) R. King & H. Robinson Terrestria. herb Piplocoma schomburgkii (Schultz Bip.) Pruski X X X Treelel Ro/andra!rulicosa (L.) Kuntze (x) Terreslrial herb Synedrella nodiflora (L.) Gaertn. (x) X Terrestria. herb Unxia camphorala L.f. (x) X Epilithic herb Floristie eheek/ist ofthe Nouragues area 305

FLORISTIC CHECKLIST OF THE NOURAGUES AREA HL RS L1FE FORM

BaIanophoraceae He/osis cayennensis (Swartz) Spreng. X Parasite herb Begoniaceae Begonia g/abra Aubl. x Vine Begonia hirtella Link X Epilithic herb Bignoniaceae Arrabidaea mollis (Vahl.) Bureau ex K. Schum. X Cydista aequinoctialis (L.) Miers x X Liana Distictella e/ongata (Vahl) Urb. XX Liana Distictella magnoliifo/ia (Kunth) Sandw. X Uana Distictis pu/veTU/enta (Sandw.) A. Gentry x Liana Jacaranda copaia (Aubl.) D. Don X Tall tree Mansoa alliacea (Lam.) A. Gentry X Liana [Memora a/ba (Aubl.) Miers], after Larpin 1993 X Liana cf. Memoraflavida (OC.) Bureau & K.Schum. x Liana Memoraflaviflora (Miq.) Pulle X X Liana Parabignonia steyermarkii Sandw. X Liana Schlege/ia paraensis Ducke x Liana Tabebuia capitata (Bureau & K. Schum.) Sandw. x XX Emergenttree Tabebuia serratifolia (Vahl) Nichols. x XX Medium tree Bixaceae Coch/ospermum orinocense (Kunth) Steudel X X Bombacaceae Catostemma sp. X Tall tree Ceibapentandra (L.) Gaertn. x Emergenttree Eriotheca crassa (U ittien) A. Robyns X Tree Eriotheca g/obosa (Aubl.) A. Robyns (X) X Small tree Eriotheca surinamensis (Uittien) A. Robyns XX X Tall tree Pachira aquatica Aubl. X Tree Pachira dolichoca/yx A. Robyns x Tree Quararibea duclœi Huber X Tree Quararibea guianensis Aubl. x Tree Boraginaceae Cordiafu/va LM. Johnston x Tree Cordia goe/diana Huber X Emergent tree Cordia /omat%ba LM. Johnston X Smalltree Cordia nodosa Lam. X (x) Treele1 Cordia sagotii LM. Johnston X Tree Cordia tetrandra Aubl. x Tree Burseraceae Crepidospermum rhoifolium (Benth.) Triana & Planch. X Medium tree 306 P. Be/benoît et al.

FLORISTfC CHECKLIST OF THE NOURAGUES AREA H L RS LlFE FORM

Dacryodes cuspidata (Cuatrec.) Daly x Tree Dacryodes nitens Cuatrec. X Talltree Protium apicu/atum Swart X Medium Iree Protium aracouchini (Aubl.) March. X Medium lree Protium araguense Cuatrec. X Tree Protium decandrum (Aubl.) March. X Tall Iree Protium cf. demerarense Swart X Tree Protium giganteum Engl. var. crassifo/ium (Engl.) Daly X Tree Protium guianense (Aubl.) March. X Medium Iree Protium heptaphyl/um (Aubl.) March. XX Medium Iree Protium morii Daly X Tree Protium opacum Swart X Tree Protium plagiocarpum Benoist (X) Small tree Protium robustum (Swart) Porter X Tree Protium sagotianum March. X Talltree Protium subserratum (Engl.) Engl. X Trec Protium tenuifo/ium (Engl.) Engl. X Tree Protium trifo/io/atum Engl. X Trec Tetragastris a/tissima (Aubl.) Swart X Tall Iree Tetragastris panamensis (Engl.) Kuntze X Talltree Tral/innickia burserifo/ia Man. X Small tree Tratlinnickia demerarae Sandw. X Tall Iree Tral/innickia rhoifo/ia Willd. X Emergenl Iree Caetaceae Epiphyl/um phyl/anthw (L.) Haw. (X) X EpiphYle Cactaceae sp. X Caesalpiniaceae Batesiaj/oribunda Spruce ex Benth. X Tree Bauhinia cupreonitens Ducke X Liana Bocoa prouacensis Aubl. X Medium Iree Cassia sp. X Chamaecrista dJ!SlIauxii (Collad.) Killip var. saxo/i/is (Amsh.) Irwin & Bameby X Chamaecris/a diphyl/a (L.) Greene (X) (X) X Epililhie herb Crudiasp. X Dia/ium guianense (Aubl.) Sandw. X Tall tree Dicorynia guianensis Amsh. X Tall tree Dimorphandra pul/ei Amsh. X Emergenl Iree Eperuafa/ca/a Aubl. X Tall Iree Eperua grandiflora (Aubl.) Benth. X Tall Iree Eperua rubiginosa Miq. X Medium tree Hymenaea courbari/ L. X Mediumlree F/oristic checkJist o/the Nouragues area 307

FLORISTfC CHECKLIST OF THE NOURAGUES AREA HL RS LIFEFORM

Macr%bium bifolium (Aubl.) Pers. x Tall Iree Pe/logyne panicu/ala Benth. subsp. pubescens (Benth.) M.F. da Silva x Emergent tree Pe/logyne wmosa (Vahl) Benth. x Tall tree Sc/er%bium a/biflorum (Benoist) Zaruch. & Herend. x X Tall tree Swartzia arborescens (Aubl.) Pinier x Medium lree Swartzia grandifolia Bong. el( Benth. x Medium tree Swartzia guianensis (Aubl.) Urb. x Small Iree Swartzia /eioca/ycina Benth. x Swartzia cf. ob/anceo/ata Sandw. x Tr.. Swartzia panacoco (Aubl.) Cowan var. sagolii (Sandw.) Cowan x Tall tree Swartzia pa/yphy//a OC. X Tan Iree Tachiga/i melinonii (Benth.) Zaruch. & Herend. x Tree Tachigali myrmecophi/a (Ducke) Ducke x Tree Tachigali cf. panicu/ata Aubl. x Tree Tachigali paraensis (Benth.) Barneby x Tree Vouacapoua americQI/Q Aubl. x Tall lree Campanulaceae Cenlropogon comutus (L.) Druce x Terrestrial herb Canellaceae Cinnamodendron lenuifo/ium Uinien x Small Ir.. Caricaceae Jacaratia spinosa (Aubl.) A. OC. X Small lree Caryocaraceae Caryocar g/abrum (Aubl.) Pers. x Tan Ir.. Cecropiaceae Cecropia distachya Huber x Tree Cecropia graffl'i//eana C.C. Berg x Small lree Cecropia obtusa Trécul x Small tree Cecropia sciadophylla Mari. X TallIT.. Coussapoa anguslifolia Aubl. x X Epiphyte & lree Coussapoa asperifolia Trécul X Epiphyte & tree Coussapoa /atifo/ia Aubl. x x X Epiphyle & tree Pourouma bic%r Mari. X Small lree Pourouma cf. cecropiaefolia Mari. X Tree Pourouma guianensis Aubl. x Tan Ir.. Pourouma melinonii Benoist x Medium lree Pourouma minor Benoist x Medium lree Pourouma mollis Trécul X Medium tree Pourouma lomen/osa Miq. X Medium lree Pourouma ve/utina Miq. X Small tree Pourouma vil/osa Trécul x TalllTee 308 P. Belbenoit et al.

FLORISTIC CHECKLlST OF THE NOURAGUES AREA HL RS LlFEFORM

Pouroumo sp. x Celastraceae Goupia g/abra Aubl. x Tall lree May/enus myrsinoides Reissek x X Tall tree May/enus ob/onga/a Reissek x Treelet (May/enus pruinosa Reissek], alter Larpin 1993 x Tree Chrysobalanaceae Couepia caryophyl/oides Benoist x Medium tree Couepia guianensis Aubl. subsp. guianensis X Tall tree Couepia obova/a Ducke x Medium tree Couepia paril/o A.P. OC. X Medium tree Hir/el/a araguariensis Prance X Treelet Hir/el/a bicornis Malt. & Zucc. X Medium tree Hir/el/a davisii Sandw. X tree Hir/el/a g/andu/osa Spreng. X Medium tree Hir/e/Ia hispidu/a Miq. X Tree Hir/el/a physophora Malt. & Zucc. X Tree Hir/e/Ia racemosa Lwn. var. hexandra (Willd. ex Roern. & Schult.) X Treelet Hir/el/a racemosa Lwn. var. racemosa X Treelet Hir/el/a silicea Griseb. X X Smalltree Hir/el/a suffu//a Prance X Talltree Hir/el/a /enuifolia Prance X Small tree Licania a/ba (Bernoulli) Cuatrec. X Talltree Licania ope/a/a (E. Mey.) Fritsch X Tree Licania canescens Benoist X Medium tree Licania couda/a Prance X Small tree Licania cya/hodes Benoist X Smalltree Licania densiflora Kleinh. X Talltree Licania granvil/ei Prance X Talltree Licania guianensis (Aubl.) Griseb. X Tree Licania he/eromorpha Benth. X Talltree Licania hypo/euca Benth. XX Talltree Licania irwinii Prance X X Medium tree Licania lcun/hiana J.O. Hook. X Talltree Licania /a/ifo/ia Benth. ex J. O. Hook. X Medium tree Licania /alis/ipu/a Prance X Talltree Licania /axiflora Fritsch X Talltree Licania /ongisty/a (J.O. Hook.) Fritsch X Medium tree Licania majuscula Sagot X Medium tree Licania membranacea Sagot ex Laness. X Tallltee Licania micron/ha Miq. X Talltree F/oristic check/ist ofthe Nouragues area 309

FLORISTIC CHECK LIST OF THE NOURAGUES AREA HL RS L1FEFORM

Licania minutiflora (Sagot) Fritsch X Emergenl lree Parinari exce/sa Sabine x Emergenl lree Parinari parvifo/ia Sandw. X Emergenllree Clusiaceae C/usia grandiflora Splitg. XX X Epiphyte & ueele1 C/usia minor L. XX EpiphYle & tree C/usia nemorosa G. Mey. x x X Epiphyle & tree C/usia panapanari (Aub!.) Choisy X Epiphyte & lreclel C/usia p/atystigma Eyrna X Epiphyte & treele1 C/usia purpurea (Splitg.) Eng!. Epiphyte & tree Moronobea coccinea Aubl. x (x) Emergent tree P/atonia insignis Man. x T.JI tree Rheedia acuminata (Ruiz & Pav.) Planch. & Triana XX Sm.1I uee cf. Rheedia benthamiana Planch. & Triana X T.II uee Rheedia rnacrophy//a (Man.) Planch. & Triana X Tree Symphonia g/obu/ifera L.f. X Tall uee Symphonia sp. X Tovomita cepha/ostigma Vesque X Trce Tovomita choisyana Planch. & Triana X Medium cree Vismia guianensis (Aubl.) Choisy X Small Irce Vismia /atifo/ia (Aubl.) Choisy X Small uee Vismia macrophy//a Kunth X Vismia ramu/iflora Miq. X Tree Vismia sandwithii Ewan X Treelet Cornbretaceae Buchenavia amazonia Alwan & Stace X Tree Buchenavia grandis Ducke X Trce Buchenavia nitidissima (L.C. Rich.) Alwan & Stace X Tree Buchenavia cf. tetraphy//a (Aubl.) R. Howard X Emergent lrce Termina/ia amozonia (J.F. Grnelin) Exell XX Medium tree Termina/ia dichotoma G. Mey. X X Medium tree Termina/ia guyanensis Eichl. X Trec Connaraceae Connarusfascicu/atus (OC.) Planch. X Connarus perrottetii (OC.) Planch. X Liana Rourea sp. X Convolvulaceae Bonamia maripoides H. Hallier X Dicranosty/es cf. guianensis Mennega X Dicranosty/es cf. vi//osus Mennega X Ipomoea angu/ata Man. ex Choisy X Epilithiç herb 310 P. Be/benoit et al.

FLORISnC CHECKLIST OF THE NOURAGUES AREA HL RS LIFEFORM

Maripa cf. g/abra Choisy (X) Liana Maripa scandens Aubl. X Liana Cucurbitaceae Cayaponia cf. ophlluJ/mica R.E. Schultes X Cayapania racemosa (Mill.) Cogn. (X) Vine Cayaponia rigida (Cogn.) Cogn. (X) Gurania bignoniacea (Poepp. & Endl.) C. Jeffrey X Vine Psiguria triphy//a (Miq.) C. Jeffrey X Liana Se/ysia prunifera (Poepp. & Endl.) Cogn. X Liana DichapetaJaceae [DicluJpeta/um peduncu/atum (OC.) Baill.]. after Larpin 1993 X Liana Tapura amazonica Poepp. & Endl. X SmalllrCC Tapura capitu/ifera Baill. X Medium lrcc Tapura singu/aris Ducke X Trcc Dilleniaceae Davi//a sp. X [Do/iocarpus cf. denlalus (Aubl.) Standl.]. after Larpin 1993 (X) X Liana [Do/iocarpus guianensis (Aubl.) Gilg.]. after Larpin 1993 X Liana Do/iocarpus major J.F. Gmelin X Do/iocarpus sp. XX X Ebenaceae Diospyros carbonaria Benoist X trec Diospyros dichroa Sandw. X Talltree Diospyros duckei Sandw. X Medium lrce Diospyros guianensis (Aubl.) Guerke X Trce Diospyros cf. malheriana A.C. Smith X Trec El aeocarpaceae S/oanea garckeana K. Schum. X SmalllrCC S/oanea graci/is Uittien X Trce S/oanea grandiflora J.E. Smith x Trec S/oanea sinemariensis Aubl. X Trec S/oanea lerniflora (Moçino & Sessé ex OC) Standl. X Trcc Ericaceae Cavendishia ca//ista J.D. Smith XX X Li ana Salyria cerander (Dunal) A.C. Smith x x X Liana Sphyrospermum cordifo/ium Benth. (x) X X Woodyepiphyte Erythroxylaceae Erylhroxy/um citrifo/ium St. Hil. (x) XX Trcc & treelcl Erylhroxy/um /iguslrinum OC. X X X Tree & shrub Erylhroxy/um macrophy//um Cav. X SmalllrCC Erylhroxy/um mucronalum Benth, (x) X Trec & IrCClet Floristic checklist ofthe Nouragues area 311

FLORISTIC CHECKLIST OF THE NOURAGUES AREA H L RS LIFEFORM

Erythroxylum squamotum Swanz x x X Treclel Erythroxylum vemicosum O.E. Schulz x X Trec Euphorbillceae Acaiypha sp. X Alchornea trip/inervia (Spreng.) Muell. Arg. X Tree Alchorneopsisjloribunda (Benth.) Muell. Arg. X Medium Iree cf. Amanoa guianensis Aubl. X Conceveiba guianensis Aubl. X SmaUlree Croton hostmannii Miq. XX X Treelet Croton matourensis Aubl. X Croton schiedeanus Schlechlend. X Croton tafelbergicus Croizat XXX Treelet Dalechampia parvibracteata Lanj. X Liana Drypetes cffanshawei Sandw. X Tree Drypetes variabi/is Vittien X TaU tree Glycydendron amazonicum Ducke X TaU tree Hevea guianensis Aubl. X Medium tree Hyeronimo alchorneoides AliemllO var. alchorneoides X Tree Mabea piriri Aubl. X Tree Mabea specioSQ Muel!. Arg. subsp. speciosa X Small tree Mabea subsessi/is PlIX & K. Hoffm. X Trec Maprounea guianensis Aubl. X Medium tree Micrandra brownsbergensis Lanj. X X Medium Iree Omphalea diandra L. X Liana Phyllanthus cf. altenuatus Miq. X X Small tree Plukenetia paiyadenia Muell. Arg. X Liana Pogonophora schomburgkiana Miers ex Benth. XX Medium tree [cf. Sagotia racemoSQ BlÙlI.], lifter Larpin 1993 X Trec Sapium montanum Lanj. X XX Treclet Sapium paucinervium Hemsley X Medium tree Fabllceae Andira coriacea Pulle X Medium tree Andira surinamensis (BondI) Splitg. ex Amshoff X Taii tree Clitoria sagotii Fantz X Liana Da/bergia monetaria L.f. X Small tree Diocka elliptica MlIXwell X Liana Diockasp. X Dip/otropis purpurea (Rich.) Amsh. X Emergent tree Dipteryx odorata (Aubl.) Willd. X Taii tree Dipteryx punctoto (Blake) Amsh. X X Emergent tree

Dussia disc%r (Benth.) Amsh. X Emergentlree 312 P. Belbenoit et al.

FLORISTIC CHECKLIST OF THE NOURAGUES AREAHL RS L1FEFORM

Hymenolobiumflavum Kleinh. x Tall tree Hymenolobium pelraeum Ducke x Emergenl Iree Lonchocarpus aff. heplaphyllus (Pair.) OC. X Medium tree cf. Monopleryx sp. X Ormosia coccil'N!a (Aubl.) Jackson x Tree Ormosia paraensis Ducke x Tree P1atymiscium u/ei Harms x Poecilanthe hoslmannii (Benth.) Amsh. x Medium lree Plerocarpus officinalis Jacq. x Tall Iree Sty/osanthes guial'N!nsis (Aubl.) Swartz (X) (X) X Epililhic herb Valairea erylhrocarpa Ducke X Valairea paraensis Ducke X Emergent tree Valaireopsis surinamensis Lima X Emergent Iree FI acourtiaceae Banara guial'N!nsis Aubl. X Smalllree Carpolroche crispidenlala Ducke X Trce & treelet Casearia acuminala OC. X Tree & treelet Casearia commersoniana Cambo X Treelel Caseariajavilensis Kunth X Small tree Casearia pilumba Sieumer X Tree & Ireelet [Homalium guianense (Aubl.) Oken). after Larpin 1993 X Medium tree Laelia procera (Poepp.) Eichl. X Emergenllree Ryania speciosa Vahl X X Treelet Gentianaceae /r/bachia a/ala (Aubl.) Maas subsp. a/ala X Epilithic herb /r/bachia a/ala (Aubl.) Maas subsp./ongisty/a Persoon & Maas X Epilithic herb /r/bachia purpurascens (Aubl.) Maas (X) X X Terrestrial herb Voyria aphylla (Jacq.) Pers. XX Temstrial hem Voyria auranliaca Splitg. X Saprophyte Voyria caeru/ea Aubl. X Saprophyte Voyria clavala Sp 1itg. X X Saprophyte Voyria corymbosa Splitg. X X Saprophyte Voyria rosea Aubl. X X Saprophyte Voyria lel'N!/Ia Hook. X Saprophyte Voyria lenuiflora Griseb. X Saprophyle Voyriella parviflora (Miq.) Miq. X X X Temstrial herb Gesneriaceae Besleria cf. /axiflora Benth. X Epilithic Ireele! CGdonanlhe ca/carala (Miq.) HansI. X Epiphyte herb CGdonanlhe crassifolia (Focke) C. Morton X X X Epiphyte herb CGdonanlhopsis dissimu/ala (H.E. Moore) Wiehler X Epiphyte herb Floris/ie eheek/is/ of/he Nouragues area 313

FLORISTIC CHECKLIST OF THE NOURAGUES AREA H L RS L1FEFORM

Columnea calotricha J.O. Smith x Epiphyte herb Columnea sanguinea (Pers.) Hanst. x Epiphyte herb Drymonia coccinea (Aubl.) Wiehler X Liana & epiphyle Drymonia serrulata (Jacq.) Malt. X Vine Episcia sphalera Leeuwenberg XX Epilithic herb Episcia mntha Leeuwenberg X Epilithic herb Lembocarpus amoenus Leeuwenberg X X Epilithic herb Napeanthus jelskii Fritsch X Epilithic herb Nautilocalyx mimuloides (Benth.) C. Morton XX Terrestrial herb Nauliloca/yx pictus (Hook.) Sprague X Terrestrial herb Paradrymonia camposlyla (Leeuwenberg) Wiehler XX Vine Paradrymonia densa (C.H. Wright) Wiehler XX Epililhic herb Paradrymonia macu/ata (J.O. Hook.) Wiehler X Vine Hemandiaceae Hernandia guianensis Aubl. X Hippocrateaceae Cheiloclinium cognatum (Miers) A.C. Smith XX Liana Elachypterafloribunda (Benth.) A.c. Smith X Hippocratea volubilis L. X Liana Salacia cf. cordala (Miers) A.M. Mennega X Liana [Salacia impressifolia (Miers) A.C. Smith], alter Larpin 1993 XX Liana Salacia insignis A.C. Smith X Liana Salacia multijlora (Lam.) OC. X Lianalet Salacia sp. X Tontelea cylindrocarpa (A.C. Smith) A.C. Smith X Liana Tontelea laxijlora (Benth.) A.C. Smith XXX Liana Hugoniaceae Hebepelalum humiriifolium (Planch.) Benth. X Medium tree Humiriaceae Humiria balsamifera (Aubl.) St. Hil. var. balsamifera X XX Trcelet & tree Sacoglottis cydonioides Cuatrec. X TaI! tree Sacoglottis guianensis Benth. XX TaU tree [Schistostemon dichotomum (Urb.) Cuatrec.], alter Larpin, 1993 X Medium tree Vantaneaparvijlora Lam. X Emergent tree 1cacinaceae Dendrobangia boliviana Rusby X TaI! tree Poraqueiba guianensis Aubl. X Medium tree Lacistemataceae Lacistema aggregatum (Bergius) Rusby X Treelet Lacistema grandifolium Schnizlein X Medium tree 314 P. Belbenoit et al.

FLORISTIC CHECKLIST OF THE NOURAGUES AREA HL RS LlFEFORM

Lauraceae Aiouea guianensis Aubl. x Tall !ree Aiouea sp. x Aniba guianensis Aubl. x Tree Aniba panurensis (Meissn.) Mez X Tree Aniba rosaeodora Ducke x Tall tree Aniba terminalis Ducke X Tree Cassytha filiformis L. X (x) Parasite hem Endlicheria punctulata (Mez) C.K. Allen X Treelet Endlicheria pyriformis (Nees) Mez X Tree Endlicheria sericea Nees X Tree Kubitzkia mezii (Kosterm.) Van der Werff X Tree Licaria cannelia (Meissn.) Kosterm. X X Talltree Licaria chrysophylla (Meissn.) Kosterm. X Tall tree Licaria guianensis Aubl. XX Tall tree Licaria vernicosa. (Mez) Kosterm X TreeJet cf Mezilaurus sp. X Nectandra globosa (Aubl.) Mez X Tree Nectandra maguireana C.K. Allen X Medium tree Oco/ea aciphylla (Nees) Mez X Medium tree Oco/ea amazonica (Meissn.) Mez X Tree Oco/ea argyrophylla Ducke X Tree Oco/ea ceanothifolia (Nees) Mez X Tree cymbarum Kunth X Tree Oco/ea discrepens C.K. Allen X Medium tree Oco/eafendleri (Meissn.) Rohwer X Tree Oco/eafloribunda (Swartz) Mez X Tall tree Ocotea guianensis Aubl. X Tree Ocotea indirec/inervÎa C.K. Allen X Tree Ocotea leuco;,;ylon (Swartz) de Lanessan X Tree Ocotea nigra Benoist X Tree Ocotea oblonga (Meissn.) Mez X Tree Oco/ea rubra Mez X Tall tree Ocotea schomburgkiana (Nees) Mez X Tree Ocolea splendens (Meissn.) Mez X Tree Ocotea tomentella Sandw. X Small tree Rhodostemonodaphne grandis (Mez) Rohwer X Tree Rhodos/emonodaphne kunthiana (Nees) Rohwer X Lecythidaceae Cory/hophora rimosa W. Rodrigues X Tall tree Coura/ari guianensis Aubl. emend. P,anee X Emergent tree F/oristic checlc/ist ofthe Nouragues area 315

FLORISTIC CHECKLIST OF THE NOURAGUES AREA H L RS LlFEFORM

Coura/ari mu//iflora (J.E. Smith) Eyma x Tan Iree Coura/ari ob/OIIgifolia Ducke & Knuth x Troo Eschwei/era apicu/ala (Miers) AC. Smith x Emergent Iroo Eschwei/era cf. charlaceifo/ia Mori x Tree Eschwei/era collina Eyma x Tan Iroo Eschwei/era conges/iflora (Benoist) Eyma x Medium Iree Eschweilera coriacea (AP. OC.) Mori x Tan tree Eschweilera ckc%rans Sandw. x Tan tree Eschwei/era grandiflora (Aubl.) Sandw. x Medium tree Eschwei/era /aevicarpa Mori x Tree Eschwei/era micran/ha (Berg) Miers x Tan tree Eschwei/era parviflora (Aubl.) Miers x Tan tree Eschweilera pedicellala (Rich.) Mori x Medium tree Eschweilera sago/iana Miers x Tan tree Eschweilera simiorum (Benoist) Eyma x Medium lree Eschweilera squamala Mori x Tree Guslavia augusla L. x Medium Iree Gustavia hexapeta/a (Aubl.) J.E. Smith x Medium tree Lecy/his charlacea Berg X Tree Lecylhis confer/iflora (AC. Smith) Mori x Tan Iroo Lecy/his corrugala Poit. x TaU Iroo Lecy/his ho/cogyne (Sandw.) Mori x TaU Iree Lecylhis ida/imon Aubl. x X Medium tree Lecy/his persis/ens Sagol subsp. auranliaca Mori x Tan Iroo Lecy/his persistens Sagol subsp. persislens X Tan Iroo Lecy/his poileaui Berg x Tan lree Lecylhis zabucajo Aubl. x Emergent Iroo Lentibulariaceae Virieu/aria amel/ryslina Salzm. ex St. Hil. & Girard x Epilithic herb Virieu/aria choriSlo/heca P. Taylor x Epilithic hem Virieu/aria hispida Lam. x Epilithic hem V/rieulariajamesoniana Oliver x Epiphyte hem Virieu/aria subulata L. x Epililhic herb Virieu/aria tenuissima Tulin x Terrestrial hem Loganiaceae Anlonia ovala Pohl X Medium lree POlalia amara Aubl. X Shrub or lreelet Strychnos erichsonii Rich. Schomb. ex Progel X Liana Strychnos g/abra Sagol ex Progel x Liana Strychnos guianensis (Aubl.) Mart. X Liana Sirychnos oiapocensis Froes x x Liana 316 P. Belbenoit et al.

FLORISTIC CHECKLIST OF THE NOURAGUES AREA HL RS UFEFORM

Slrychnos toxijéra Rob. Schomb. ex Benth. x Liana Loranthaceae Phthirusa pyrifolia (Kunth) Eichl. x x x Woody parasite Phthirusa stelis (L.) Kuijt x x X Woody parasite Psillacanthus redaclus Rizzini x Woody parasite Lythraceae Cuphea b/ackii Lourteig X Epilithie herb Malpighiaceae Byrsonima aerugo Sagot X Talltrec Byrsonima densa (Poir.) OC. X Tall Irec Byrsonima /aevigata (Poir.) OC. X Tree Byrsonima stipu/acea Adr. Juss. X Talliree

Heteropterys sp. XX X Malvaceae Pavonia caslanaejolia St. Hil. & Naud. X Terrestrial herb Pavonia schiedeana Steud. X Terrestrial herb Marcgraviaceae Marcgravia coriacea Vahl X Liana Norantea guianensis Aubl. X XX Liana

Souroubea guianensis Aubl. X x X Liana Melastomataceae Aciotis faxa (OC.) Cogn. var./axa x Treelet Aciotis purpurascens (Aubl.) Triana x Shrub Appendicu/aria thymifolia (Bonpl.) DC. x Epilithic herb Bellucia gros!>U/arioides (L.) Triana x Tree C/idemia cong/omerata OC. X Terrestrial herb Clidemia dentata D. Don x Treelet Clidemia hirta (L.) D. Don var. e/egans (Aubl.) Griseb. x Clidemia invo/ucrata OC. x Clidemia septulinervia Cogn. x Clidemia venosa (Gleason) Wurdack x Treelet Ernestia confértiflora Wurdack x Epilithie subshrub Ernestia g/andu/osa Gleason x Trcelet Ernestia granvillei Wurdack (x) x X Treelet Ernestia pullei G1eason x Shrub & treelet Henriettea maroniensis Sagot (x) X Tree Henriellea succosa (Aubl.) DC. x X Trec Henriellella caudata Gleason x Ireclel Henriellellaflavescens (Aubl.) Triana x Treelel Leandra divaricata (Naud.) Cogn. x x Leandra pa/eacea Wurdack x Treelet Floristic checldist ofthe Nouragues area 317

FLORISTIC CHECKLIST OF THE NOURAGUES AREA HL RS LJFEFORM

Leandra purpurea Gleason x Leandra cf rufescens (De.) Cogn. x Treelet Leandra cf verticillata (OC.) Cogn. x x Loreya mespiloides Miq. x Smal! tree Macrocentrum cristamm (OC.) Triana var. cristamm x x x Subshrub Maieta guianensis Aubl. x Shrub & treelet Miconia acuminata (Sleud.) Naud. x Medium lree Miconia affinis OC. x Smalllree Miconia argyrophylla De. x Treeler Miconia bracteata (OC.) Triana x Treelel Miconia cacatin (Aubl.) Renner x Treelet Miconia ceramicarpa (De.) Cogn. var. candolleana Cogn. x Shrub Miconia ceramicarpa (OC.) Cogn. var. ceramicarpa x x Shrub Miconia ciliata (Rich.) OC. x x x Shrub & lreelel Miconia diaphanea G1eason x Treelet Miconia diSpar Benth. (x) x Treelel Miconia h%sericea (L.) De. x x x Treelet Miconia cf. /epidota OC. x Miconia mirabilis (Aubl.) L.O. Williams x Smalltree Miconia p/ukenetii Naud. x Tree Miconia prasina (Swartz) OC. x x Tree Miconia trimera Wurdack x Shrub Mouriri crassifolia Sagol X Talllree Mouriri huberi Cogn. X Tall tree Mouriri sagotiana Triana X Epiphyle & terrestrial Topobea parasitica Aubl. (X) (X) X Epiphyte & tree Meliaceae Carapa guianensis Aubl. X Tall tree Carapa procera A. OC. X Tall \Jee Cedre/a odorata L. X Emergent tree Ouarea gomma Pulle X Tree Ouarea grandifo/ia A. De. X Tree Ouarea guidonia (L.) Sieumer x Medium tree Guarea kunthiana Adr. Juss. X Medium tree [Ouarea cf. macrophylla Vahl], after Larpin 1993 X Medium tree Ouarea micM/·moddei Pennington & Mori X Tree Ouarea pubescens (L.e. Rich.) Adr. Juss. X X Smalllree Ouarea si/vatica e. OC. X Medium lree Trichilia cipo (Adr. Juss.) e. OC. X Trichilia /epidota Man. X Medium tree Trichilia micrantha Benlh. X Medium tree 318 P. Belbenoit et al.

FLORISTIC CHECKLIST OF THE NOURAGUES AREA HL RS L1FEfORM

Trichilia pa/lida Swartz X Trce Trichilia quadrijuga Kunth x Medium trce Trichilia schomburgkii C. OC. X Medium tree Trichilia .seplentrionalis C. OC. X Tall trce Trichilia surinamensis (Miq.) C. OC. XX Tall tn:e Mendonciaceae Mendoncia hoffmannseggiana Nees X (x) Vine Menispermaceae Abula bul/ala Moldenke XX Liana Abula grandifolia (MlIrt.) Sandw. X Tree Abula cf. obovala Diels (X) Liana Abula cf. rufescens AubL X Liana Anomospermum cf. ch/oranlhum Diels X Liana E/ephanlomene cf. eburnea Barneby & Krukotf X Liana Mimosaceae Abarema barbouriana (Standl.) Bameby & Grimes X X Trec Abarema curvicarpa (Irwin) Bameby & Grimes X Trec Abarema gal/orum Bameby & Grimes X X Trec Abaremajupunba (Willd.) Britton & Killip var.jupunba XX Abarema malaybifolia (Sandw.) Bameby & Grimes X Tree Acacia lenuifolia (L.) Willd. X Balizia pedicel/aris (OC.) Bameby & Grimes XXX Medium tree Cal/iandra atf. surinamensis Benth. (X) XX Trcelet & !rce Enler%bium o/demanii Bameby & Grimes X Tree Enler%bium schomburgkii (Benth.) Benth. X Tall !rce Inga acreana Harms X Tree Inga acrocepha/a Steud. X Medium tree Inga a/ala Benoist X Trec Inga a/ba (Swartz) Willd. X Trec Inga a/bicoria O. Poncy X Trec Inga aurislel/ae Harms X Small tn:e Inga bourgoni (AubL) OC. X Tall tree Inga brachyslachys Ducke X Trcelet Inga bracleosa Benth. X Small tree Inga capi/ala Desv. X Medium !rce Inga cayennensis Sagot ex Benth. XXX Trec Inga edulis (VeiL) MlIrt. X Tall tree Ingafanchoniana O. Poncy X Trec Ingafastuosa (N.l. Jacq.) Willd. X Treclet Ingaflagel/iformis MlIrt. X Trec Inga graciliflora Benth. X Tree FJoristic checklist ofthe Nouragues area 319

FLORIsnc CHECKLIST OF THE NOURAGUES AREA H L RS LIFEFORM

Inga graci/ifo/ia Ducke X Small tree Inga grandiflora Ducke X Tree Inga huberi Ducke X Medium tree /ngajenmanii Sandw. X Medium tree Inga /ateriflora Miq. XX Mediumlree Inga /eioca/ycina Benth. X Medium lree Inga /omatophy/la (Benth.) Pitt. X Trec Inga /ongipeduneu/ata Ducke X Tree Inga marginata Willd. X Tree Inga me/inonis Sagot X Trec Inga nouragensis O. Poncy X Tree Inga paraensis Ducke X Tree Inga pezizifera Benth. X Tall tree Inga poeppigiana Benth. X Tree Inga retinocarpa. O. Poncy X Trec Inga rubiginosa (L.C. Rich.) OC. X Talltree Inga stipu/aris OC. X Medium tree Inga thibaudiana OC. X X Medium lree Inga tubaejormis Benoist XX Trec Inga umbe//ifera (Vahl) Steud. ex OC. XXX Tree Inga virgu/tosa (Vahl) Desv. XX Small tree Mimosa annu/aris Benth. var. xinguensis (Ducke) Bameby (X) Liana [Mimosa gui/andinae (OC.) Bameby var. gui/andinae). after Julliot 1992 (x) Liana Mimosa myriadena Benth. X Mimosa po/ydacty/a Humb. & Bonp!. ex Willd. X Mimosa pudica L. X Parkia ni/ida Miq. X Emergent lree Parkia pendula (Willd.) Benth. ex Walpers X Emergenllree Parkia retieu/ata Ducke X Trec Parkia u/ei (Hanns) Kuhlmann X Emergenllree Parkia ve/utina Benoist X Emergenl tree Pseudopiptadenia psi/ostachya (OC.) Lewis & M.P. Lima X Tal1lree Pseudopiptadenia suaveo/ens (Miq.) Grimes X Talllree Stryphnodendron guianense (Aubl.) Benth. X Tree Stryphnodendron moric%r Bameby & Grimes X Tree Stryphnodendron po/ystachyum (Miq.) Kleinh. X Emergenl tree Zygia racemosa (Dueke) Barneby & Grimes X Talllree Zygia sabatieri Bameby & Grimes X Trec Zygia tetragona Bameby & Grimes XX Trec Monimiaceae Siparuna cristata (Poepp. & Endl.) A. OC. X Tree 320 P. Belbenoit et al.

FLORISTIC CHECKLIST OF THE NOURAGUES AREA H L RS L1FEFORM

Sipanma cuspidala (Tul.) A. OC. x Small trce Siparuna decipiens (Tul.) A. OC. x SmaU tree Sipanma guionensis Aubl. X SmaU tree Siparuna pachyanlha A.C. Smith X SmaU teee Sipanma sprucei A. OC. X X SmaU tree guianensis Aubl. X Em ergent tree Balocarpus amazonicus (Ducke) Fosb. X Medium Iree aculifo/ium Huber subsp. aculifo/ium X TaU tree Brosimum guianense (Aubl.) Huber XX Tait tree Brosimum parinorioides Ducke X Em ergenltree Brosimum rubescens Taub. X Emergenllree Brosimum ulile (Kunth) Pitt X Emergent tree ilicifolia (Spreng.) Lanj. & Rossberg X Tree amazonica (Miq.) Miq. XX Hemi...piphYlic trce Ficus broadwayi Urb. X Hemi...piphYlic tree Ficus gomelleira Kunth & Bouché X Medium lree Ficus guianensis Desv. ex Hamilton XXX TaU tree Ficus insipida Willd. X TaU teee Ficus cf. leiophylla C.C. Berg X Medium trce Ficus malhewsii (Miq.) Miq. X TaU teee Ficus rrymphaeifolia Mill. X TaUlree Ficus pertusa L.f. X Medium lree Ficus Irigono L.f. X Hemi...piphYlic tree He/icOSlylis pedunculala Benoist X Medium lree He/icosly/is lomenlosa (Poepp. & Endl.) Rusby X TaU tree calophylla (Poepp. & Endl.) C.C. Berg X Tree Maquira guianensis Aubl. X Tait tree glabra Spruce ex Pitt. X Medium lrce Naucleopsis guianensis (Mildbr.) C.C. Berg X Medium \ree Perebea guianensis Aubl. X Medium lree Perebea mollis (Poepp. & Endl.) Huber X TaUtree laevis (Ruiz & Pav.) Macbr. X Em ergenl trce Trymalococcus o/igandrus (Benoist) Lanj. X Medium lree Trymalococcus paraensis Ducke X TaUlree Myristicaceae Iryanthera hoslmannii (Benth.) Warb. X Medium lree Iryanthera sagoliana (Benth.) Warb. X TaUlree OSleophloeumplalyspermum (Spruce ex A. OC.) Warb. X TaU \ree Virola /ewalae Sabatier X TaU tree Virola michelii Heckel X TaU Iree Floristic checklist ofthe Nouragues area 321

FLORISTfC CHECKLIST OF THE NOURAGUES AREA H L RS L1FEFORM

Virola mul/icos/a/a Ducke X Tree Virola sebifera Aubl. X Trce Virola surinamensis (Rolander) Warb. X Emergent Irce Myrsinaceae Cybian/hus guianensis (A.OC.) Miq. subsp. pseudoicacoreus (A. OC.) XXX Agostini Cybian/hus venezuelanus Mez X Treelet Myrtaceae [Ca/ycorec/es berg;; Sandw.], alter Larpin 1993 X Trcelet Calycorec/es grandifolius O. Berg X Shrub Ca/yp/ran/hes amshoffae McVaugh XX Medium trce Calyp/ran/hes lepida McVaugh XXX Tree Eugenia albicans (O.C. Berg) Urban XX Treelet Eugenia biflora (L.) OC. X Trcelet Eugenia coffeifolia OC. X Tree Eugenia cucullalo Amsh. X Talllree Eugenia cupula/a Amsh. X Trce Eugeniajlorida OC. X XX Trec Eugenia la/ifolia Aubl. X Treelet Eugenia macrocalyx (Rusby) McVaugh X Smalllrce Eugenia marowynensis Miq. XXX Smalltree Eugenia mon/icola (Swartz) OC. X X Tree Eugenia pa/ris;; Vahl X Smalltree Eugenia pseudopsidium Jacq. X Treelet Eugenia ramiflora Desv. XXX Treelet Eugenia le/ramera (McVaugh) M.L. Kawasaki & B. Holst X Tree Marliereaferruginea (Poir.) McVaugh X Myrcia amazonica OC. X Trcelet Myrcia citrifolia (Aubl.) Urb. XXX Treelet MyrciafaJlax (L.C. Rich.) OC. XXX Treelet [Myrcia cf. graciliflora Sagot], after Larpin 1993 X Treelet Myrcia guianensis (Aubl.) OC. X X X Treelet Myrcia cf. paivae O. Berg X Treelet Myrcia cf platyclada OC. X Treelet Myrcia quilorensis (Bentham) Sagat XXX Treelet Myrcia saxalilis (Amsh.) McVaugh X X Treelet Myrcia sylva/ica (G. Mey.) OC. XX X Treelet Myrciariajloribunda (West ex Willd.) O. Berg X XX Smalllree cf Psidium aculongulum Mari. ex OC. X Nyctaginaceae Guapira eggersiana (Heimerl) Lundell X X X Neeajloribunda Poepp. & Endl. X Smalltree 322 P. Belbenoit et al.

FLORISTIC CHECKLIST OF THE NOURAGUES AREA HL RS LIFEFORM

Neea ova/ifo/ia Spruce ex Schmidt XXX SmaU tree Nyctaginaceae sp. XX Ochnaceae E/vasia macroslipu/aris Sastre & Lescure X Trec OUl'alea candollei (Planch.) Tiegh. X XX Trec OUl'alea decogyna Maguirc X Medium Irce Ouralea guianensis Aubl. X Treelet Owalea impressa (Tiegh.) Lemée X Shrub & Iree Ouralea /eb/ondii (Tiegh.) Lemée XXX Trec Ouralea macrocorpa Sastre X Ouralea melinonii (Tiegh.) Lemée X Trcelel SaUllagesia aliciae Sastre X XX Shrub SaUllagesia erecla L. X Epililhic herb SaU\IQgesia lafe/bergensis Sastre X Terreslrial herb Olacaceae Cathedra acuminala (Benth.) Miers XX Tree Chaunochilon kapp/eri (Sagot ex Engl.) Ducke X Du/acia guianensis (Engl.) Kuntze X X TaU tree Heisleria cau/iflora I.E. Smith X Shrub & Ireelel Heisleria densifrons Engl. X Treelel Minquarlia guianensis Aubl. X TaUlJce Plychopela/um o/acoides Benth. X Medium Iree Opiliaceae [Agonandra silvalica DuckeJ. after Larpin 1993 X Talltree Passifloraceae Dilkea relUsa Mast. X Passiflora amoena Escobar X Liana Passiflora auricu/ala Kunth X Liana Passiflora candida (Poepp. & Endl.) Masters (X) Liana Passiflora coccinea Aubl. X Liana Passiflora crenala Feuillet & Cremers X Liana Passiflora exura Feuillet (X) Liana Passiflorafanchonae Feuillet XX Liana Passiflorafuchsiiflora Hemsley XX Liana Passiflora garckei Masters XX Liana Passiflora g/andu/osa Cav. X Liana Passiflora aff. vario/ala Poepp. & Endl. X Liana Passiflora vesperlilio L. (x) Liana Passiflora spp. Phytolaccaceae Peliveria cf. alliacea L. X FJoristic check/ist ofthe Nouragues area 323

FLORISTIC CHECKLIST OF THE NOURAGUES AREA H L RS L1FEFORM

Piperaceae Peperomia glabella (Swartz) A. Dictr. X Epiphyte herb Peperomia macrostachya (Vahl) A. Dietr. XX Epiphyte herb Peperomia obtusifolia (L.) A. Dietr. X Epiphyte herb Peperomia ouabianae C. OC X Peperomia pernambucensis Miq. X Epiphyte herb Peperomia rotundifolia (L.) Kunth X X Epiphyte herb Peperomia serpens (Swartz) J.e. Loudon X Epiphyte herb Peperomia sp. X Piper adenandrum (Miq.) e. OC. X Treelet Piper aequale Vahl X Tn:elet Piper angustifolium Lam. X Subshrub Piper anonifolium (Kunth) e. OC. X Treelet Piper arboreum Aubl. X Tree Piper augustum Rudge X Treelel Piper avellanum (Miq.) e. OC. X Treelel Piper brownsbergense Yunck. X Treelel Piper consanguineum Kunth X Subshrub Piper demeraranum (Miq.) e. OC. X Tn:elet Piper divaricatum G. Mey. X Piper hispidum Swartz X Treelet Piper hostmonnianum (Miq.) e. OC. X Liana Piper humistratum GOrts & K.U. Kramer X Tem:slrial herb Piper nemotanthera (Miq.) e. De. X Liana Piper obliquum Ruiz & Pav. X Piper cf. oblongifolium (Klotzsch) C. OC. X Tn:elet Piper paramaribense C. OC. X Subshrub Piper cf. pulleanum Yunck. X Tn:elet Piper tectoniifolium Kunth X Tn:elet Piper trichoneuron (Miq.) e. OC. X Piper wachenheimii Tre!. X Tn:elet PolygaIaceae Moutabea guianensis Aubl. X Liana Moutabea cf. longifolia Poepp. & End!. X Liana Polygala adenophora OC. (X) (X) X Epilithic herb Polygala spectabi/is OC. XX X Shrub & Ireelet Securidaca uniflora Oort X X X Liana Securidoca sp. X Polygonaceae Coccoloba marginata Benth. X Liana Coccoloba lucid1lla Benth. X Liana 324 P. Belbenoit et al.

FLORISTIC CHECKLIST OF THE NOURAGUES AREA HL RS LIFEFORM

Cocc%ba ovala Benth. x Liana Cocc%ba parimensis Benth. x Liana Proteaceae Eup/assa pinnata (Lam.) Johnston x Medium tree Roupa/a montana Aubl. x XX Medium tree Quiinaceae Lacunaria crenata (Tul.) A.C. Smith X Smalltre<: Lacunariajenmanii (Oliver) Dueke X Small tre<: Quiina cf. guianensis Aubl. X Quiina obovata Tul. x Small tree Quiina oiapocensis Pires X Talltree Tourou/ia guianensis Aubl. x Small tree Rhabdodendraceae Rhabdodendron amazonicum (Spruce ex Benth.) Huber x Small tree Rhamnaceae Ampe/ozizyphus amazonicus Dueke x Tree Gouania b/anchetiana Miq. Llana Ziziphus cinnamomum Triana & Planeh. x Tree Rhizophoraeeae Cassipourea guianensis Aubl. x X Trec Rubiaceae Alibertia surinamensis (Bremek.) Steyerm. X Treclet Aliberlia myrciifolia K. Schumann x x x Amaioua corymbosa Kunth x Amaioua guianensis Aubl. x Borreria a/ata (Aubl.) OC. (x) x Epilithic herb Borreria capi/ata (Ruiz & Pav.) OC. var. tenel/a (Kunth) Steyerm. x Epilithic herb Capirona decorticans Spruce X Tall Irec Chimarrhis turbinala OC. X Talltree Chomelia sp. x Coccocypse/um guianense (Aubl.) K. Sehum. x X TelTestrial herb Coussarea racemosa A. Rich. x Small tree Ouroia aquatica (Aubl.) Bremek. x Small tree Ouroia eriopi/a L.f. x Smail tree Ouroia genipoides J.D. Hook. ex K. Sehum. x Ouroia /ongij/ora Dueke x Medium tree Ouroia micrantha Ladbr. Zaraeehi & Kirkbride X Faramea corymbosa Aubl. X Treelet &r tree

Faramea guianensis (Aubl.) Bremek. X Shrub &treelet Ferdinandusa paraensis Ducke x Medium tre<: F/oristic check/ist ofthe Nouragues area 325

FLüRISTIC CHECKLIST OF THE NOURAGUES AREA H L RS L1FEFüRM

Geophi/a cordifo/ia Miq. x x Epilithic herb Geophi/a sp. x Guettardo acreana Krause x Small tree Guettarda aroma/ica Poell. & Endl. x Hil/ia iI/lU/ris (Vell.) K. Schum. x Epiphyte & treelet /ser/ia spiciformis DC. X X Treelet lxora graci/iflora Benth. X X Tree lxora pires;; Steyenn. Treelet lxora u/ei Krause X Ma/anea macrophyl/a Bartling ex Griseb. X X Liana MafU!ttia a/ba (Aubl.) Wernham (X) X Liana Morinda brachyca/yx (Bremek.) Steyenn. X X Treelet Morinda sp. XX O/den/andia cf. corymbosa L. X Palicourea ca/ophyl/a OC. X Shrub Pa/icourea expe/ens Standl. X Tree Pa/icourea guianensis Aubl. X [Pa/icourea quadrifo/ia (Rudge) OC.]. after Larpin 1993 X Tree Pa/icourea /riphy//a OC. X Treelet Perama hirsula Aubl. X Epilithic herb Posoqueria /a/ifo/ia (Rudge) Roem. & Schult. X Small tree Psycho/ria anceps Kunth X Tree Psycho/ria apoda Steyerm. X Treelet Psycho/ria cf. bahiensis OC. X Treelet Psycholria b/ephorophyl/a (Standl.) Steyenn. X Shrub Psycholria brac/eocardia (OC.) Muel1. Arg. (x) X Treelet Psycho/ria capila/a Ruiz & Pav. X X Tree Psycholria c/enophora Steyerm. X XX Shrub & treelet Psycho/ria cupu/aris (Muel\. Arg.) Stand\. X Trec Psycho/ria deflexa OC. X Treelet Psycho/ria erec/a (Aubl.) Stand\. & Steyenn. X X Shrub & treelet Psycho/ria ga/baoensis Steyerm. X Tree Psycho/ria hoffmannseggiana (Willd. ex Roem. & Schult.) Muell. Arg. X XX Shrub & treelet Psycho/ria cf. kapp/eri (Miq.) Muel\. Arg. ex Benoist (x) Shrub & treelet Psycho/ria /igu/aris (Rudge) Steyenn. var. carapichea (Poir.) Steyerm. X Treelet Psycho/ria mapourioides OC. X Treelet Psycho/ria microbo/rys Ruiz ex Standley X Shrub & treelet Psycholria moroidea Steyerm. XX Shrub & treelet Psycho/ria poeppigiano Muell. Arg. X Shrub Psycho/ria racemosa (Aubl.) Raeusch. X X Shrub & treelet Psycho/ria surinamensis Bremek. X 326 P. Belbenoit et al.

FLORISTIC CHECKLIST OF THE NOURAGUES AREA H L RS LlFEFORM

Psycho/ria ulviformis Steyerm. x Psycho/ria variegata Steyerm. X TelTestrial herb Randia arma/a (Swartz) OC. X Tree Rudgea crassiloba (Benth.) Robinson XX X Rudgea tanaosepala Sandw. X Treelet Sipanea pratensis Aubl. (x) (x) X Epililhic herb Sipanea wilson-brownei Cowan X TelTestrial herb cf. Tocoyena guianensis K. Schum. X Uncaria guianensis (Aubl.) J.F. Gmelin X Liana Rutaceae Ertela /rifolia (L.) Kuntze (X) X Esenbeckia cowanii Kaastra (X) X Small tree Esenbeckia grandiflora Man. (x) X Small tree Galipea /rifoliata Aubl. (x) X Tree Zanthoxylumpentandrum (Aubl.) R. Howard X Tree Sapindaceae Cupania diphylla Vahl XX Small lree Cupania hirsu/a Radlk. X Medium lree Cupania scrobiculata L.e. Rich. X Smail tree Ma/ayba arborescens (Aubl.) Radlk. X Li ana Ma/ayba guianensis Aubl. X Small tree Ma/ayba inelegans Spruce ex Radlk. X Tree Paullinia anodonta Radlk. X Liana Paullinia capreolata (Aubl.) Radlk. X Liana Paullinia cf. plagiop/era Radlk. X Lians Paullinia rufescens Rich. ex Juss. X Paullinia cf. tricornis Radlk. X Liana Paullinia cf. venosa Radlk. X Tree Paullinia sp. X Porocys/is /oulicioides Radlk. X Pseudimafru/escens (Aubl.) Radlk. X Medium tree Talisia carina/a Radlk. X Small tree Talisia c/a/hra/a Radlk. X Tree Talisia hexaphylla Vahl X Tree [Talisia longifolia (Benth.) Radlk.]. after Larpin 1993 X Small tree Talisia micran/ha Radlk. X Treelet Talisia mollis Kunth ex Cambo X Talisia pedicellaris Sagot ex Radlk. X Tree Talisia praealta (Sagot) Radlk. X Mediumtree Talisia simaboides K.U. Krarner X Tall tree Vouarana guianensis Aubl. X Smalltree Floristie eheeklist ofthe Nouragues area 327

FLORISTIC CHECKLIST OF THE NOURAGUES AREA H L RS LIFEFORM

Sapotaceae Chrysophy/lum argenteum Jacq. subsp. auralum (Miq.) Pennington x Trec Chrysophy/lum argenteum Jacq. subsp. nilidum (G. Mey.) Pennington x Medium lree Chrysophy/lum cuneifo/ium (Rudge) A. OC. X Mediumlree Chrysophy/lum eximium Ducke X Medium lrec Chrysophy/lum /ucentifo/ium Cronq. X Emergent Iree Chrysophy/lum pomiferum (Eyma) Pennington X Tall tn:e Chrysophy/lum prieurii A. OC. X Tall tn:e Chrysophy/lum sanguino/entum (Pierre) Baehni X Medium lree Chrysophy/lum venezue/anense (Pierre) Pennington X Treele! Dip/olJn cuspic/alum (Hoehne) Cronquist X Ecc/inusa guiQJliinsis Eyma X Tall tree Ecc/inusa /anceo/ala (Malt. & Eichl.) Pierre X Smalllree Ecc/inusa ramiflora Malt. X Tree E/aeo/uma nuda (Baehni) Aubr. X Mani/kara bidentala (A. OC.) Chevalier subsp. bidentala X X Tall tree Mani/kara huberi (Ducke) Chevalier x X Tall tree Manilkara zapota (L.) van Royen [not a native species] x Micropholis cayennensis Pennington X Tall tree Micropho/is egensis (A. OC.) Pierre x Medium lree Micropho/is guyanensis (A. OC.) Pierre subsp. guyanensis X Tall tree Micropho/is /ongipedice//ala Aubr. x Tree Micropho/is me/inoniana Pierre x Tall tree Micropho/is mensa/is (Baehni) Aubr. x X Tall tree Micropho/is obscura Pennington x Tall tree Micropho/is porphyrocarpa (Baehni) Monachino x Tree Micropho/is sanclae-rosae (Baehni) Pennington x Tree Micropho/is venu/osa (Malt. & Eichl.) Pierre x Tall tree Pouleria ambe/aniifo/ia (Sandw.) Pennington x Talllree Pouleria benai (Aubr. & Pellegr.) Pennington x Tall tree Pouleria afr. bi/ocu/aris (Winkler) Baehni x Talllree Pouleria caimilo (Ruiz & Pav.) Radlk. x Tree [Pouleria af[ cicalricala Pennington], after Larpin 1993 x Tree Pouleria c/adanlha Sandw. x Tree Pouleria afr. congeslifo/ia Pilz x Trec Pouleria coriacea (Pierre) Pierre x Tree Pouleria cuspidala (A. OC.) Baehni subsp. dura (Eyma) Pennington x Trec Pouleria decorlicans Pennington x Trec Pouleria de/iciosa Pennington x Tree Pouleria egregia Sandw. x Talllree Pouleria eugeniifo/ia (Pierre) Baehni x Tall tree 328 P. Belbenoil el al.

FLûRlsnc CHECKLIST OF THE NOURAGUES AREA HL RS L1FEFûRM

Pouleria engleri Eyma x Tree Pouleriafilipes Eyma x Tall tree Pouleria glomerala (Miq.) Radlk. subsp. slylosa (Pierre) Pennington x Tree Pouleria gonggrijpii Eyma x Medium trce Pouleria grandis Eyma x Trec Pouleria guianensis Aubl. x Talltree Pouleria hispida Eyma X Small tree Pouleriajariensis Pires & Pennington x Trec Pouleria laevigaJa (Mart.) Radlk. x Tall tree Pouleria macrophylla (Lam.) Eyma x Tree Pouleria melanopoda Eyma x Medium tree Pouleria oblanceolala Pires X Tree Pouleria reliculala (Engl.) Eyma X Trec Pouleria relinervis Pennington X Small tree Pouleria sagoliana (Baill.) Eyma X Small tree PouJeria singularis Pennington X Trce Pouleria lorla (Mart.) Radlk. subsp. glabra Pennington X Smail tree Pouleria lorla (Mart.) Radlk. subsp. luberculala (Sleumer) Penninglon X Tree Pouleria venosa (Mart.) Baehni X Tall trce Pouleria virescens Baehni X Tree Pouleria williamii (Aubr. & Pellegr.) Pennington X tree Pradosia huberi (Ducke) Ducke X Troo Pradosia plychandra (Eyma) Penninglon X Medium Irce [Pradosia verlicillala Duckel. after Larpin 1993 X Tree Sarcaulus brasiliensis (A. OC.) Eyma X Scrophulariaceae Scrophulariaceae sp. X Simaroubaceae Picramnia guianensis (Aubl.) Jansen·Jacobs X X Treelet Simaba guianensis Aubl. X X Smalltroo Simaba orinocensis Kunth X Treelel Simaba polyphylla (Cavalcante) W. Thomas X Trcelel Simarouba amara Aubl. X Talllree Solanaceae Brunfrlsia guianensis Benth. X (X) X Treelel Cyphomandra endopogon Biller X Liana Cyphomandra harlWegii (Miers) Walp. X Treelel Lycianlhes pauciflora (Vahl) Bitter X Morlœa camponoli Ducke x Marlœa coccinea L.C. Rich. x Liana Marlœaformicarum Dammer x Liana Floristie eheeklist ofthe Nouragues area 329

FLORIsnc CHECKLIST OF THE NOURAGUES AREA H L RS LIFEFORM

Morua sessifif/ora Ducke (x) x Sofanum anceps Ruiz & Pav. x TerreSirial herb Sofanum asperum L.C. Rich. x Trcele! Sofanum coriaceum Dunal x Liana Solanum fepidotum Dunal x Woody Sofanum schfechlendalianum M. Walpers x Trcele! Solanum vefulinum Dunal x Treele! Sterculiaceae Byttneria morii L. Sarnen & DOIT X Trce Siercufia excefsa Malt. X Tall Iree Siercufiojrondosa A. Rich. x Tall trce Siercufio pruriens (Aubl.) K. Schum. x Tall tree Siercufio sp. x Theobroma subincanum Malt. X Medium tree Theobroma lIefulinum Benoist x Smalliree Styracaceae Styrax sp. x Symplocaceae Sympfocos martinicensis Jacq. x Medium tree Theaceae Ternslroemio den/ala (Aubl.) Swartz x x X Tree Theophrastaceae Cfallija lam:ifolia Desf. x Thymeleaceae Thymefeaceae sp. XX x Tiliaceae Apeiba glabra Aubl. x Tall Iree Apeiba peloumo Aubl. X TaI( Iree Luehea cf speciosa Willd. x Lueheopsis rugosa (Pulle) Burret x Tall trce Tumeraceae Tumera rupeslris Aubl. x Treelet Urticaceae Pilea sp. X Verbenaceae Aegiphifa lIiffosa (Aubl.) J.F. Gmelin X Tree Pelrea bracleala Steud. (X) (x) X Liana Vi/ex guianensis Moldenke X Medium tree Vilex Irif/ora Vahl XX Tree Violaceae Amphirrhox fongifolia (St. Hil.) Spreng. X 330 P. Be/benoit et al.

FLORlSTlC CHECKLIST OF THE NOURAGUES AREA H L RS LIFEFORM

Leonia glycycarpa Ruiz & Pav. x Medium tree Noisel/ia orehidiflora (Rudge) Gingins X Terreslrial herb Paypayrola guianensis Aubl. X Small tree Rinorea amapensis Hekking X Tree Rinoreajlaveseens (Aubl.) Kuntze X Rinorea negleela Sandw. X Rinorea riana Kuntze X Tree Viscaceae Phoradendron piperoides (Kunth) Trel. XX Woody parasite Vochysiaceae Erismajloribundum Rudge X TaU tree Erisma uncinalum Warm. X TaU tree Qualea polyehroma Stail. X Qualea rosea Aubl. X TaU tree Ruiz/erania albiflora (Warm.) Marcano-Berti X Emergent tree [Voehysia guianensis Aubl.], after Larpin 1993 X X TaUtree Vochysia neyra/ii Nonnand X Tal\tree Vochysia surinamensis Stail. X TaUtree SPERMATOPHYTA Monocotyledones Araceae Anaphyllopsis amerieanum (Engl.) Hay X Terrestrial herb An/hurium clavigerum Poepp. & Endl. X Vine Anthu/ium afr. erassinervium (Jacq.) Scholl X (X) An/hurium gracile (Rudge) Lind!. X X Epiphyte herb An/huriumjenmanii Engl. X x x Epilithic herb An/hurium cf. lanjouwii Jonker & Jonker X Epiphyte herb Anthurium rubrinervium (Link) G. Don X Epilithic herb An/hurium seandens (Aubl.) Engl. X Epiphyte herb An/hurium sinualUm Benth. ex Scholl X Vine Anthurium /hrifIQX Madison X Epilithic herb Anthurium /rinerve Miq. X An/hurium sp. Caladium bieolor (Aiton) Ventenal XX Epilithic herb Die.ffenbaehia duidae (Steyenn.) Bunting X Terrestrial herb Dieffenbaehia paludieola N.E. Brown ex Gleason x Aquatic herb Dieffenbaehia seguine (Jacq.) Scholl X Terrestrial herb Draeontium polyphyllum 1.. x X Epilithic herb He/eropsisjlexuosa (Kunth) Bunting x Vine Homalomena pielUrala (Linden & André) Regel X Terrestrial herb Mons/era adansonii Scholl XX Vine Mons/era dubia (Kunth) Engl. & Krause X Vine Floristic checklist o/the Nouragues area 331

FLORlsnc CHECKLlST OF THE NOURAGUES AREA HL RS LlFEFORM

Monstera obliqua Miq. X Vine Monstera spruceana (Scholl) Engl. X Vine Philodendron bi/lietiae Croal X X X Epiphyte herb Philodendron ca/losum Krause X Epiphyte herb Philodendron deflexum Poepp. ex SchoU X X Epiphyte herb Philodendron duclœi Croal & Grayum X Philodendronfragrantissimum (Hook.) Kunth XX X Epiphyte herb Philodendron goeldii Barroso X Epiphyte herb Philodendron grandifolium (Jacq.) Schou XX Vine Philodendron guianense Croat & Grayum X Philodendron insigne SchOll X X X Epiphyte herb Philodendron linnaei Kunth XXX Epiphyte herb

Philodendron melinonii Brongn. ex Regel X X Epiphyte herb Philodendron ornatum SchOll XX Epilithic herb

Philodendron pedatum (Hook.) Kunth X X Vine Philodendron placidum SchoU X Philodendron rudgeanum Schott X Vine Philodendron scandens K. Koch & Sello X Vine Philodendron solimoesence A.C. Smith X Epilithic herb Philodendron sphalerum SchOll X Vine Philodendron squamiferum Poepp. XX Epiphyte herb Philodendron cf surinamense (Scholl) Engler X Philodendron wittianum Engl. X Rhodospatha cf. oblongata Poepp. X Rhodospatha venosa G1eason X Spathiphyllum humboldtii SchOll X Aquatic herb Syngonium podophyllum Scholl X Xanthosoma conspurcatum Scholl XX Epilithic herb Arecaceae Astrocaryum gynacanthum Mart. X Palm Astrocaryum paramaca Mart. X Palm Astrocaryum sciophilum (Miq.) Pulle X Palm Bactris acanthocarpoides Barb. Rodr. X Palm Baclris aubletiana Trail X Palm Bactris constanciae Barb. Rodr. X Palm Bactrisfloccosa Spruce X Palm Bactris gastoniana Barb. Rodr. X Palm Bactris hirta Man. (x) X Palm Bac/ris oligocarpa Barb. Rodr. & Trail ex Barb. Rodr. X (x) Palm Bactris rhaphidacantha Wessels Boer X Palm

Bactris simplicifrons Mart. X Palm 332 P. Belbenoit et al.

FLORlsnc CHECKLIST OF THE NOURAGUES AREA HL RS L1FEFORM

Desmoncus sp. x Lianapalm EUlerpe o/eracea Mart. X Palm Geonoma deversa (Poi!.) Kunth x Palm Geonoma slricla (Poi!.) Kunth x Palm Hyospalhe e/egans Mart. x (x) Palm Jessenia balaua (Mart.) Burret x X Palm Maximiliano maripa (Corrêa) Drude x Palm Oenocarpw bacaba Mart. X Palm Orbignya sp. x Palm Socralea exorrhiza (Mart.) H. Wend1. x Palm Syagrw slralinco/a Wessels Boer x Palm Bromeliaceae Aechmea aqui/ega (Salisb.) Griseb. x x X Epiphyte & epilithic Aechmea brome/iifolia (Rudge) Baker x x X Epiphyte & epilithic Aechmea melinonii Hook. XXX Epiphyte & epilithic Ananas anonassoides (Baker) L.B. Smith (x) XX Epilithic Araeococcus micranlhus Brongn. (x) X Epiphyte Billbergia vio/acea Beer XX Epiphyte & epilithic Brome/ia agavifo/ia Brongn. ex Houllett XX Terrestrial herb Calopsis berleroniana (Schult.f.) Mez (x) (x) X Epiphyte Disleganlhus basilaleralis Lem. X X Epilithic Disleganlhus /aleralis (L.B. Smith) Gouda XX Guzmania lingu/ata (L.) Mez X XX Epiphyte & epilithic Guzmania me/inonis Regel X X Epiphyte Pilcairnia geyskesii (L.B. Smith) Varadarajan & Gilmartin X Epilithic Pilcairnia saslrei L.B. Smith & R.W. Read XX Epilithic Racinae spicu/osa (Griseb.( Spencer &L.B. Smith (x) X X Epiphyte Slreploca/yx /ongifolius (Rudge) Baker (x) X Epiphyte Tillandsia anceps Lodd. XX Epiphyte Tillandsia bu/bosa Hook. (X) X X Tillandsiaflexuosa Swartz X Epiphyte Tillandsia monade/pha (E. Morren) Baker XX Epiphyte Tillandsia lenuifo/ia L. (X) X Epiphyte Vriesea p/eioslicha (Griseb.) Gouda XXX Epiphyte & epilithic Vriesea splendens (Brongn.) Lem. var. sp/endens XXX Epiphyte & terreslrial Weraulhia g/adio/iflora (Wendl.) J.R. Grant XX Epilithic Burmanniaceae Apleria aphylla (Nut!.) Barnhart ex Small XXX Saprophyte Burmannia capitala (Walter ex J.F. Gmel.) Mart. X (X) X Epilithic Campy/osiphon purpurascens Benth. (x) Saprophyte Dictyoslega orobanchoides (Hook.) Miers XX Tem:strial herb FJoristic checklist ofthe Nouragues area 333

FLORISTIC CHECKLIST OF THE NOURAGUES AREA HL RS LIFE FORM

Gymnosiphon breviflorus G1eason x x Terrestrial herb Gymnosiphon minulus Snelders & Maas (x) Saprophyle Hexapterel/a gentianoides Urb. (X) Saprophyte Commelinaceae Buforrestia candol/eana C.B. Clarke x Terreslrial herb Comme/ina nif1pes Seub. var. glabrata (DR Hunt) Faden & D.R. Hunt x X Epililhic Costaceae Costus c/aviger Benoist X Terreslrial herb Costus congestif/orus L.C. Rich. ex Gagnep. X Terreslrial herb Costus curcumoides Maas X Terreslrial herb Costus erythrothyrsus Loes. XX Epililhic herb Cyclanthaceae Asp/undia brachyphyl/a Harling X Terreslrial herb Asp/undia g/andu/osa (G1eason) Harling X Terrestrial herb Asp/undia heteranthera Harling X Vine Asp/undia sp. X Cyclanthus bipartitus Poit. X Terrestrial herb Evodianthusfunifer (Poit.) Lindm. subsp.funifer XX Vine Ludovia /ancifo/ia Brongn. X X X Steksty/is surinamensis Harling XX Terrestrial herb Thoracocarpus bissectus (Vell.) Harling X Liana Cyperaceae Becquere/ia cymosa Srongn. X (X) Terreslrial herb Bisboecke/era /ongifo/ia (Rudge) Kuntze X Bisboecke/era microcepha/a (Boeck.) T. Koyama X Terresuial her" Calyptrocarya bic%r (H. PfeilT.) T. Koyama X Ca/yptrocarya g/omerulata (Brongn.) Urb. X Terrestrial herb Cyperus /uzu/ae (L.) Roltb. ex Retz. X Terresuial herb Cyperus odoralus L. X Terreslrial herb Cyperus pa/ystachyos Roltb. X Fimbristy/is dichotoma (L.) Vahl X Terrestrial herb Fimbristy/is mi/iacea (L.) Vahl [not a native species] X Terrestrial herb Hypo/ytrumjenmanii C.B. Clarke subsp.jenmanii X Terrestrial herb Hypolytrum /ongifo/ium (L.C. Rich.) Nees X Terreslrial herb Kyl/inga brevifo/ia Roltb. X Terreslrial herb Kyllinga odorata Vahl X Terrestrial herb Maponia sy/vatica Aubl. X Terreslrial herb Rhynchospora barbata (Vahl) Kunth (x) (X) X Terrestrial herb Rhynchospora capitata (Homboldt & Kunth) Roem. & Schult. X Terrestrial herb Rhynchospora cepha/otes (L.) Vahl (x) XX Terrestrial herb Rhynchosporafi/iformis Vahl X 334 P. Belbenoit et al.

FLORISTIC CHECKLI ST OF THE NOURAGUES AREA HL RS L1FEFORM

Rhynchospora globosa (Kunth) Roem. & Schult. x Terrestrial herb Rhynchasporajunciformis (Kunth) Boeck. x Terrestrial herb Rhynchospora /enuis Willd. ex Link x (X) X Terrestrial herb Scleria cyperina Willd. ex Kunth (x) (x) X Terrestrial herb Scleria disions Poir. x Epilithie Scleria secam (L.) Urb. x x X Vine Dioscoreaceae Dioscorea sp. x Eriocaulaceae Paepalanthus lamarckii Kunth x Epilithie Syngonan/hus caulescens (Poir.) Ruhland x Terrestrial herb Heliconiaceae He/iconia acumina/a L.e. Rich. x Terreslrial herb He/iconia bihai (L.) L. x He/iconia hirsu/a L. f. x Terrestrial herb He/iconia pendula Wawra X Terrestrial herb He/iconia richardiana Miq. X (X) Terrestrial herb Marantaceae Cala/hea cyc/ophora Baker X Terrestrial herb Cala/hea cy/indrica (Roscoe) K. Schum. X Terrestrial herb Cala/hea grandis Petersen X Terrestrial herb Calatheapropinqua (Poepp. & Endl.) KOm. XX Terrestrial herb Cala/hea squarrosa L. Andersson & H. Kennedy XX X Epilithie herb Ischnosiphon arouma (Aubl.) KOm. X Terrestrial herb Ischnosiphon graci/is (Rudge) KOm. X Terrestrial herb fschnosljJhon marlianus Eichl. ex Petersen X (x) Terrestrial herb fschnosljJhon pe/iolatus (Rudge) L. Andersson X Terreslrial herb Monophyl/an/he o/igophylla K. Schum. X Terrestrial herb

Mono/agma spicatum (Aubl.) Macbr. X Terrestrial herb Orchidaceae Aganisia pulchella Lindl. (X) XX Epiphytie herb Ba/emannia colleyi Lindl. (X) X Epiphytie herb Brassia lanceana Lindl. (X) X Epiphytie herb Bulbophyllum roraimense Rolfe X Epiphytie herb Campylocenlrum micran/hum (Lindl.) Rolfe (X) XX Epiphytie herb Cheiradenia cuspida/a Lindl. XX Epiphytie & epilithie Cleis/es rosea Lindl. (X) (X) X Terrestrial herb Cleis/es /enuis (Rchb.f. ex Griseb.) Schlechter X Terrestrial herb Cranichis sp. X Cyr/opodium andersonii (Lamb.) R. Brown (x) XX Dichaea pendula (Aubl.) Cogn. XX Epiphytie herb Floristie eheekJist ofthe Nouragues area 335

FLORISTIC CHECKLIST OF THE NOURAGUES AREA H L RS L1FEFORM

Dichaea splilgerberi Rchb.f. (X) X Epiphytie hem Dichaea /rul/a Rchb.f. (X) X Epiphytie hem EI/ean/hus brasiliensis (Lind!.) RchbJ. XX Epiphytie herb EI/ean/hus carava/a (Aub!.) Rchb.f. (X) XX Epiphytie herb Ellean/hw graminifolius (Barb. Rodr.) Loijtnant (X) X X Epiphytie hem Encyclia ionosma (Lindl.) Schlechter (X) X Epilithie herb Epidendrum anceps Jacq. X Epiphytie hem Epidendrum noclUmum Jacq. (X) X Epiphytie herb Epidendrum purpurascens Focke (X) XX Epiphytie hem Ery/hrodes sp. X Habenaria rodriquezii Cogn. X Terrestrial herb Jacquiniel/a globosa (Jacq.) Schlechter (X) XX Epiphytie hem Koel/ens/einia carraoënsis Garay & Dunsterv. X Terrestrial herb Koel/ensteinia graminea (Lind!.) Rchb.f. (X) X Epiphytie herb Koel/ens/einia sp. X Epiphytie herb Masdeval/ia minuta Lind!. (X) XX Epiphytie hem Maxil/aria brunnea Linden & Rchb.f. (x) X Epiphytie herb Maxil/aria camaridii Rchb.f. (X) X Epiphytie herb Maxil/aria crassifolia (Lind!.) Rchb.f. XX Epiphytie herb Maxillaria desvauxiana Rchb.f. X Epiphytie herb Maxillaria unca/a Lind!. XXX Epiphytie hem Maxillaria vil/osa (Barb. Rodr.) Cogn. (X) XX Epiphytie herb Maxil/aria violaceopunc/a/a Rchb.f. (X) XX Epiphytie hem Maxil/aria sp. XX Epiphytie hem NOlylia sagillifera (Kunth) Linle, Klotzsch, & Otto X Epiphytie herb NOlylia yauaperiensis Barb. ROOr. X Epiphytie hem Oc/omeria sar/houae Luer X Epiphytie herb Oc/omeria sp. X Ornithocephalus kruegeri Rchb.f. X Epiphytie Palmorchis pabslii Veyret X Terrestrial Palmorchis praspec/orum Veyret X Terrestrial Paphinia sp. X Pelexia cal/ifera (C. Schweinf.) Garay X Terrestrial Phragmipedium lindJeyanum (Schomb.) Rotfe X X Epilithie hem Pleuro/ha/lis arisla/a W.J. Hook. (X) X Epiphytie hem Pleuro/hallis pic/a Lind!. (X) X Epiphytie hem Pleuro/hallis spiculifera Lind!. (X) XX Epiphytie herb Polys/achyaj/avescens (Blume) U. Smith X Epiphytie herb Polys/achyafoliosa (Hook.) Rchb.f. (X) XX Epiphytie herb Prescollia sp. X Rodriguezia candida (Lind!.) Rchb.f. X Epiphytie herb 336 P. Belbenoit et al.

FLORISTIC CHECKLIST OF THE NOURAGUES AREA H L RS LIFEFORM

Rodrigueria lanceola/a Ruiz & Pav. (x) x x Epiphytie hem Scaphyglollis bifida (Rchb.f.) Schweinf. X Epiphytie herb Scaphyglollisfusiformis (Griseb.) Schultes X Epiphytie herb Scaphyglottis prolifera Cogn. (X) X x Epiphytie hem Scaphyglo//is sickii Pabst X Epiphytie hem &aphyglo//is s/ellala Lodd. ex Lindl. XX Scaphyglollis violacea Lind!. X Epiphytie herb Seu/icaria s/eelei (Hook.) Lindl. X Epiphytie herb Sigma/os/alu: huebneri Mansf. X Epiphytie hem S/elis argen/ala Lindl. (x) X Epiphytie herb Trichosalpinx egleri (Pabst) Luer X Epiphytie herb Trichosalpinx orbieularis (Lind!.) Luer (X) XX Epiphytie hem Vanilla ova/a Rolfe (x) X Vine Wullschlaegelia calcara/a Benth. X Saprophyte Zygosepalum labiosum (L.e. Rich.) Schweinf. (X) X Epiphytie herb Zygosepalum Iindeniae (Rolfe) Garay & Dunsterv. (X) X Epiphytie Poaceae Axonopus compressus (Swartz) P. Beauv. X Terrestrial hem Axonopus ramosus Swallen X Terreslrial herb Cenchros echinalUs L. X Terrestrial herb Digilaria ciliaris (Retz.) Koeler X Terreslrial herb Guadua Ia/ifolia (Humb. & Bonpl.) Kunth X Terrestrial herb Ichnan/hus nemoralis (Schrad.) Hitchc. & Chase (x) X X Terrestrial herb Ichnan/hus panicoides P. Beauv. X lchnamhus /enuis (J. Presl) Hitchc. & Chase (x) X Terreslrial herb Olyra obliquifolia Steud. X X Terreslrial hem Panieum miliaceum L. [not a native species] X Terreslrial herb Panicum rivale Swallen x Temslrial herb Pariana campes/ris Aubl. XXX Terreslrial herb Pariana radiciflora Sagot ex Doell X Paspalum conjugalUm Berg. X Terreslrial hem Paspalum deeumbens Swartz x Terreslrial herb Paspalum marilimum Trin. X Temslrial hem Paspalum mullinervium A.G. Burman (x) (x) X Terreslrial hem Pharos lalifolius L. X Terreslrial herb Pharos parvifolius Nash X Temslrial herb Pharus virescens Doell X Terreslrial herb Selaria ilalica (L.) P. Beauv. [not a native species) x Terresrrial hem Sorghum halepense (L.) Pers. [not a native species) X Tcrreslrial hem Thrasya scandens (Tutin) Soderstrom ex A.G. Burm. x Terreslrial herb Floristie eheeklist ofthe Nouragues area 337

FLORISTIC CHECKLIST Of THE NOURAGUES AREA H L RS L1FE FORM

Rapateaceae Ropatea paludosa Aubl. x Terrestrial herb Spathanthus uni/aleralis (Rudge) Desv. x Aqualic herb Smïlacaceae Smilax spp. [5 laxa 1 Thumiaceae Thurnia sphaerocephala (Rudge) J.O. Hook. X Terreslrial Triuridaceae Sciaphila a/bescens Benth. X Saprophyte Triuris hyalina Miers Saprophyte Xyridaceae Abo/boda americana (Aubl.) Lanj. X Epilithic herb Xyris savanensis Miq. X Epilithic herb Zingiberaceae Renealmia guianensis Maas X Terrestrial herb Renea/mia monosperma Miq. X TerreSlrial hem Renea/mia orinocensis Rusby X Terrestrial herb SPERMATOPHYTA Gnelales Gnelaceae Gnetum nodiflorum Brongn. X Woody epipbyle Gnetum panicu/atum Spruce ex Benlh. X Liana HEPATOPHYTA Herbertaceae Herbertus divergens (Sleph.) Herz. X Lejeuneaceae Cerato/ejeunea sp. x Cystolejeunea /ineata (Lehm. & Lindenb.) Evans X BRYOPHITA Calymperaceae Calymperes erosum C. MOII. x Ca/ymperes /onchophyllum Schwaegr. x Ca/ymperes nicOl'aguense Ren. & Cardo x Ca/ymperes venezue/anum (Mitt.) Broth. ex Pitt. X Sy"hopodon hornschuchii Mart. X Sy"hopodon /eprieurii Mont. x Syrrhopodon pro/ifer Schwaegr. x (x) x Oicranaceae Campylopus richardii Brid. x Campy/opus savannarum (C. MolI.) Mill. x uuc%ma cruegerianum (C. MolI.) Jaeg. & Sauerb. x uucolomo serrulatum Brid. x 338 P. Belbenoit et al.

FLORlsnc CHECKLIST OF THE NOURAGUES AREA HL RS LIFEFORM

Hookeriaceae Callicostella pallida (Homsch.) Angstr. x Hypnella pallescens (Hook.) Jaeg. x Hypnaceae Chrysohypnum diminulivum (Hampe) Buck X Isoplerygium lenerum (Swartz) Mitt. X Rhocopilopsis Irinilensis (C. Moll.) Britt. & Dix. x Leucobryaceae Leucobryum marlianum (Homsch.) C. MOII. x x x OCloblepharum cocuiense Mitt. XX OCloblepharum ereclifolium Min. ex Williams x OCloblepharum slramineum Min. x Meteoriaceae Squamidium leucolrichum (Tay!.) Broth. x Onhotrichaceae Macromilrium cirrosum (Hedw.) Brid. x Macromilrium pellucidum Min. x Phyllodrepaniaceae Phyllodrepaniumfalcifolium (Schwaegr.) Crosby x Pterobryaceae Renauldia paradoxica Allen x Sematophyllaceae Acroporium pungens (Hedw.) Broth. x Semalophyllum subsimplex (Hedw.) Mitt. x x x Taxilheliumplanum (Brid.) Min. x x Trichosleleum papillosum (Homsch.) Jaeg. x x Stereophyllaceae Pilosium chlorophyllum (Homsch.) C. Moll. x Thuidiaceae Cyrlohypnum involvens (Hedw.) Buck & Crum x LYCOPHYTA Lycopodiaceae Huperzia linifolia (L.) Trevis. (x) (x) x Epiphyte Selaginellaceae Selaginella epirrhizos Spring x TeITCstrial Selaginellaj1agellala Spring x TeITCstrial Selaginella producla Baker x x x Selaginella radiala (Aub!.) Spring x X Terrestrial Selaginella revoluta Baker x x TeITCstrial Floristic checklist ofthe Nouragues area 339

FLORlsnc CHECKLIST OF THE NOURAGUES AREA HL RS LlFE FORM

PTEROPHYTA Adiantaceae Adianlopsis radiata (L.) Fée X Terrestrial fem Adiantum adiantoides (J.E. Smith) C. Chr. x Terrestrial fem Adiantum cajennense Willd. ex Klotzsch x Terrestrial fem Adiantum sp. nov. aff. cordatum Maxon x TerreSlrial fem Adiantumfructuosum Poepp. ex Spreng. x Terrestrial fem Adianlumfu/iginosum Fée x TerreSlrial fem AdianlUm glauœscens Klotzsch x Terrestrial fem Adiantum Iatifo/ium Lam. x Terrestrial fem Adiantum leprieurii Hook. x Terrestrial fem Adiantum oyapokense Jenm. x Terrestrial fem Adiantum tomentosum Klotzsch x Terrestrial fem Pityrogramma calomelanos (L.) Link Terrestrial fem Aspleniaceae Asplenium angustum Swartz (x) X Epiphytie fem

Asplenium auritum Swartz X Epiphytie fem

Asp/eniumjuglandifolium Lam. X X Epiphytie & terrestrial fem Asplenium laetum Swartz X Epiphytie & epilithie Asplenium serratum L. X X Epiphytie fem Blechnaceae Salpichlaena volubilis (Kaulf.) J. Smith X Terrestrial fem Cyatheaceae Cyathea andina (H. Karst.) Domin X Treefem Cyathea cyatheoides (Desv.) K.U. Kramer X Treefem Cyathea pungens (Willd.) Domin X Treefem Cyathea surinomensis (Miq.) Domin X Treefem Dennstaedtiaceae Dennstaedtia obtusifo/la (Willd.) T. Moore X Terrestrial hem Hypo/epis repens (L.) K. Presl x Terrestrial hem Lindsaea dubia Spreng. x X Terrestriai herb Lindsaea guianensis (Aubl.) Dl)'ander x Terrestrial hem Lindsaea lancea (L.) Bedd. var.falcata(Dl)'ander) Rosenstock x Terrestrial herb Lindsaea lanœa (L.) Bedd. var. lancea x X Terrestrial hem Lindsaea pa//ida Klotzsch x x Terrestrial hem Lindsaea reniformis Dl)'ander x Terrestrial herb Saccoloma inaequale (Kunze) MeU. X Terrestrial herb Dl)'opteridaceae Cyclodium guianense (Klotzsch) L.D. Gomez x X Terrestrial fem Cyclodium inerme (Fée) A.R. Smith x Terrestrial fem Cyclodium meniscioides (Willd.) K. Presl x X Terrestrial & epiphytie 340 P. Belbenoit et al.

FLORISTIC CHECKLIST OF THE NOURAGUES AREA HL RS LIFEFORM

Didymoch/aena /runcaJu/a (Swartz) J. Smith x TenestriaJ fem Olfersia cervina (L.) Kunze x Hemi-epipbytie fern Po/ybo/rya caudata Kunze x Hemi-epipbytie fern Grammitidaceae Cochlidium.fùrcatum (Hook. & GreY.) C. Chr. x X Epipbyte Coch/idium /inearifo/ium (DcsY.) Maxon ex C. Chr. x x X Epipbyte Cochlidium serru/atum (Swartz) L.E. Bishop (x) x X Epipbyte Grammi/is b/anchetii (C. Chr.) A.R. Smilh x Epipbyte Grammi/is suspensa (L.) Proctor x x x Epipnyte Hymenophyllaceae Hymenophyllum btl!vifrons Kunze x Epipbytie Cern Hymenophyllum decurrens (Jacq.) SWarlZ (x) x Epipbytie fern Hymenophyllum hirsutum (L.) Swartz x Epipbytie fem Hymenophyllum po/yan/hos (Swartz) Swartz x X x Epipbytie fem Trichomanes ankersii Parker ex Hook. & Grey. x Epipbytie fem Trichomanes arbuscu/a Desy. x x Epipnytie fem Trichomanes crispum L. (x) x X Terrestrial fem Trichomanes cris/atum Kaulf. (x) X Terrestrial & epipbytie fem Trichomanes diversifrons (Bory) MeIl. ex Sadebeck x X Terrestrial fem Trichomanes e/egans L.C. Rich. x Terrestrial fem Trichomanes kapp/erianum J.W. Sturm x Epipbytie fem Trichomanes maci/entum Bosch x Fern Trichomanes pedicella/um OesY. x Epipbytie fem Trichomanes pinnatum Hedw. x Terrestrial fem Trichomanes radicans Swartz X Epipnytie & epilitbie Trichomanes trollii Bergd. x Terrestrial & epiphytie Lomariopsidaceae Bo/bitis nico/ianifolia (Swartz) Aislon x Terrestrial fem Bo/bitis semipinnatifida (Fée) Alston x Terrestrial fern E/aphog/ossumflaccidum (Fée) T. Moore x Epipbytie fem Elaphog/ossum g/abellum 1. Smith x x x Epipbytie fem E/aphog/ossum herminieri (Bory & Fée ex Fée) T. Moore x .x x Epipbytie fem E/aphog/ossum /aminarioides (Bory ex Fée) T. Moore (x) X Epipbytie fem E/aphog/ossum /uridum (Fée) Christ x x Epipnytie fem E/aphog/ossum p/eropus C. Chr. (x) x Epiphytie fern E/aphog/ossum raywaense (Jenm.) Alston X Epiphytie fern Lomagramma gu/anens/s (Aubl.) Ching X Epiphytie fern Lomariopsisjapurensis (Mart.) J. Smith x Terrestrial & epilitbie fern Marattiaceae Danaea nodosa (L.) J.E. Smith x Terrestria! fem Danaea simp/icifo/ia Rudge x Terrestrial F/oristic check/ist ofthe Nouragues area 341

FLORlSTlC CHECKLIST OF THE NOURAGUES AREA H L RS LlFEFORM

Metaxyaeeae Metaxya rostrata (Kunth) K. Presl x Oleandraeeae Nephrolepis biserra/a (Swartz) Sehou x Terrestrial & epiphytie rem Nephrolepis rivularis (Vahl) Mett. ex Krug (x) x Epiphytie rem Oleandra ar/icula/a (Swartz) K. Presl (x) X Epiphytie rem Polypodiaeeae Campyloneurum phylli/idis (L.) K. Pres1 x x Epiphytie rem Dieranoglossum desvauxii (Klotzseh) Proetor X Epiphytie rem Mierogrammafuseopunelala (Hoak.) Varesehi x Epiphytie & epilithie rem Mierogramma /ycopodioides (L.) Copeland x x x Epiphytie & epilithie rem Mierogramma reptans (Cav.) A.R. Smith (x) x Epiphytie & epilithie rem Niphidium erassifolium (L.) Lellinger x X Terrestrial & epiphytie rem Peclllma pee/ina/a (L.) Priee x Epiphytie rem PoJypodillm dulce Poir. x X Fern PoJypodillm /riseriale Swartz (X) X x Epiphytie rem Pteridaeeae P/erispungens Willd. x Terrestrial rem Schizaeaceae Aelinos/aehys pennilia (Swartz) Hook. x Terrestrial rem Aelinos/aehys sub/rijllga (Mart.) K. Presl x Terrestrial rem Sehizaea incurva/a Sehkuhr x Terreslrial rem Teetariaceae Tee/aria Incisa Cav. x Terreslrial rem Tee/aria plantaginea (Jacq.) Maxon x X Terrestrial rem Tee/aria trifolia/a (L.) Cav. X Terrestrial rem Triplophyllumfunestum (Kunze) Holttum x Terrestrial rem Thelypteridaceae The/yp/eris leprieurii (Hoak.) R. Tryon x Terrestrial rem Thelyp/eris penna/a (Poir.) C. Morton x Terrestrial rem Vittariaceae Anelium eilrifolium (L.) Splitg. x Epiphytie rem Antrophyum eajenense (Desv.) Spreng. X Epiphytie rem Antrophyum guayanense Hieron. x x Epiphytie rem

Nouragues Dynamics and Plant-Animal Interactions in a Neotropical Rainforest

edited by

FRANS BONGERS

PIERRE CHARLES-DOMINIQUE

PIERRE-MICHEL FORGET

MARCTHERY

KLUWER ACADEMIC PUBLISHERS DORDRECHT I BOSTON I LONDON A C.I.P. Catalogue record for this book is available from the Library of Congress.

ISBN 1-4020-0123-1

Published by Kluwer Academie Publishers, P.O. Box 17,3300 AA Dordrecht, The Netherlands.

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