The Distribution of Protected Areas in Relation to the Needs of Biotic Community Conservation in Eastern Africa
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The Distribution of Protected Areas in Relation to the Needs of Biotic Community Conservation in Eastern Africa A Paper presented at the Regional Meeting on Coordinated System of National Parks and Reserves in Eastern Africa Seronera Lodge, Serengeti National Park, Tanzania 14-19 October 1974 by Hugh F. Lamprey IUCN Consultant IUCN Occasional Paper No. 16 International Union for Conservation of Nature and Natural Resources Morges, Switzerland, 1975 THE DISTRIBUTION OF PROTECTED AREAS IN RELATION TO THE NEEDS OF BIOTIC COMMUNITY CONSERVATION IN EASTERN AFRICA A Paper presented at the Regional Meeting on Coordinated System of National Parks and Reserves in Eastern Africa Seronera Lodge, Serengeti National Park, Tanzania 14-19 October 1974 by Hugh F. Lamprey IUCN Consultant IUCN Occasional Paper No. 16 International Union for Conservation of Nature and Natural Resources Morges, Switzerland, 1975 © 1975 International Union for Conservation of Nature and Natural Resources FOREWORD IUCN has launched a programme aimed at ensuring that representative samples of the various types of natural biomes and ecosystems are conserved in a coordinated system of national parks and related protected areas. Such action was called for in Recommendation 1 of the Second World Conference on National Parks (Grand Teton National Park, U.S.A., September 1972). As part of this programme Dr. Hugh Lamprey, IUCN/WWF consultant, undertook a survey of areas in eastern Africa within the framework of UNEP Project No. 0603-73-001. The results of his work are set down in the present paper. The survey was carried out during most of 1974 and covered nine countries: Botswana, Ethiopia, Kenya, Malawi, Somalia, Sudan, Tanzania, Uganda, and Zambia. The report on the survey and related recommendations for action were considered by the Regional Meeting on Coordinated System of National Parks and Reserves in Eastern Africa held at Seronera Lodge, Serengeti National Park, Tanzania, 14-19 October 1974. The version presented here incorporates suggestions made at that meeting. Whilst it will be included in the Proceedings of the Seronera meeting which will appear in the IUCN Supplementary Paper series, the report is being issued separately as IUCN Occasional Paper No. 16 because of its broad interest as a source document. Grateful acknowledgement is made to the Fund of UNEP and to WWF for financial support of Dr. Lamprey's work. 3 CONTENTS FOREWORD 3 I. INTRODUCTION 7 II. CLASSIFICATION OF BIOTIC COMMUNITIES 7 III. THE PARKS AND RESERVES 9 IV. CONSERVATION OF BIOTIC COMMUNITIES IN EASTERN AND CENTRAL AFRICA - CHECKLIST 12 V. CONCLUSIONS AND RECOMMENDATIONS 48 VI. BIBLIOGRAPHY A. Africa General 54 B. East Africa - regional 55 C. Central Africa - regional 56 D. Southern Africa - regional 56 E. Botswana 57 F. Ethiopia 58 G. Kenya 59 H. Malawi 59 I. Somalia 60 J. Sudan 61 K. Tanzania 63 L. Uganda 64 M. Zambia 65 N. General 66 5 MAPS Botswana . 69 Ethiopia 71 Kenya 73 Malawi 75 Somalia 77 Sudan 79 Tanzania 81 Uganda . 83 Zambia 85 I. INTRODUCTION This paper aims to review the extent to which the biomes and their important biotic communities in nine eastern and central African countries are protected by their inclusion in national parks and game and forest reserves. It consists largely of tabulated and abbreviated information and it enumerates the countries and their parks and reserves in which the different communities are represented. Short notes provide additional information on the special importance of the conservation areas and on their security. In cases where a conspicuous community is not represented or is inadequately represented in a protected area, this is indicated. Maps of the vegetation of each country are combined with maps showing the distribution of parks and reserves. Owing to the small scale used this can give only an approximate representation of the inclusion of the major communities within the protected areas. Similarly, the protection of natural ecosystems in an African region can only be reviewed superficially in a paper of this length. Nevertheless it is hoped that the most important gaps in the conservation coverage of the region can be identified. Above all there is a need to draw attention to biotic communities which are vulnerable to irreversible change and to possible extinction. Experience has shown that progressive ecological changes are usually degrading and lead to a reduction in species diversity or biological productivity or both. Such changes should not be confused with successional processes which are cyclical or with the relatively rare instances of recovery towards more diverse and productive associations. The objectives of biological conservation have been discussed else- where (e.g. Lamprey 1972) and the subject will not be pursued in this paper. It is assumed that the maintenance of the highest possible degree of endemic diversity is the main objective and this paper reviews the extent to which this is achieved by the existing system of parks and reserves. The conservation of biomes and ecosystems in as natural a state as possible has become recognized as a virtual imperative and is incorporated in the O.A.U. African Convention on the Conservation of Nature and Natural Resources. II. CLASSIFICATION OF BIOTIC COMMUNITIES A review of the state of conservation of the biotic communities of a region necessitates the adoption of a practical scheme of classifi- cation. As yet none exists which portrays adequately the full range of diversity of ecosystems, and the construction of a comprehensive scheme presents almost insuperable difficulties. For practical purposes our classification has to be based on major vegetation types characterized by a combination of physiognomic and floristic features (mainly dominant tree and shrub species and genera), However it is important to recognize the limitations of this system as a means of identifying all biotic communities. 7 8 The information available on which to base an assessment of the protection of existing communities is inadequate and unevenly distributed. There are floristic lists, descriptions and maps for most countries or regions, based mainly on the woody plants. In the descriptions and maps the authors have tended to use. their own class- ification and terminology. Nevertheless, present knowledge on vegetation has been used to construct a vegetation map of Africa (AETFAT: Keay 1958) on a scale of 1:10,000,000, in which most (but not all) of these individual approaches have been reconciled. A second map of African vegetation on a larger scale (AETFAT: White 1974) is in press. These maps can only show the distribution of the larger blocks of vegetation with a relatively low degree of resolution but much of their merit lies in the largely successful attempt at a unified classification across the African continent. There are excellent works on the birds and mammals of several countries and regions and more locally on the reptiles and amphibians. A small number of ecological surveys of varying degrees of detail have been carried out, incorporating integrated accounts of plant and animal communities but for the most part biogeographical information tends to be sharply divided into the botanical and the zoological. It is only in recent years that the ecosystem approach has prevailed with its aim of describing the whole range of organisms and their functional relation- ships with each other and their environment. Information on invertebrate fauna and micro-organisms is particularly sparse. Our ability to identify biotic communities is limited by the extent and detail of the ecological exploration of the area concerned. Although we can recognize and define the broader categories of biotic community on the basis of a relatively small number of conspicuous plants and animals, at a higher level of resolution we will probably fail to identify numerous small and local ecosystems. A related problem is the recognition and documentation of variation in the botanical composition within biomes and also in the relative dominance of the component species, particularly in those biomes which have wide geographical ranges. Such variation may be due to topography, soil, drainage, local climate, fire distribution and frequency, or the direct and indirect effects of man. There will also be genetic variation within species across the geographic range of an extensive biome. The process of evolutionary divergence resulting from partial or total genetic isolation may lead to speciation, particularly where a biome is represented by several island ecosystems as in the case of the montane and Afro-alpine communities of eastern Africa. Each community tends to possess its own complement of endemic species and races as well as many that are common to other areas or to the biome as a whole. The mountain massifs of eastern Africa are particularly good examples of localized endemism in plants and animals. The conclusion that must be drawn from our knowledge of the ecological and genetic variation within biomes is that no two sample areas are exactly alike. The implications this has for the selection of areas for the protection of biotic communities, are discussed below. The system of classification of biomes and their sub-types used in this review is based as closely as possible on that of IUCN (1973) but with modifications which are intended to adapt it to the vegetation of 9 Africa and to the terminology usually applied to it. Some difficulty has been experienced in fitting certain localized biomes into the classification system and there is the inevitable uncertainty about equating a vegetation zone with its ecological equivalent in the other hemisphere. However, provided that the system is applied with the full knowledge of its limitations, it can serve the purpose of identifying the main biomes and communities and in plotting their distributions. III. THE PARKS AND RESERVES In the past, the survival of indigenous ecosystems has most commonly occurred in areas which have remained undeveloped through historical accident or owing to the existence of serious obstacles to development.