10 © Sahlbergia Vol. 10: 10–27, 2005 Check-list of Finnish flies: families Xylophagidae – Microphoridae Jere Kahanpää & Kaj Winqvist Kahanpää, J. & Winqvist, K. 2005: Check-list of Finnish flies: families Xy- lophagidae – Microphoridae. — Sahlbergia 10: 10–27. Helsinki, Finland, ISSN 1237–3273. A checklist is presented for the Finnish species of the fly families Xylophagidae, Stratiomyidae, Xylomyidae, Athericidae, Rhagionidae, Tabanidae, Acroceridae, Asilidae, Bombyliidae, Mythicomyiidae, Therevidae, Scenopinidae, Atelesti- dae, Empididae, Hybotidae and Microphoridae. Seventeen species previously not recorded from Finland are here added to the Finnish fauna. The following species have been erroneously reported from Finland and are here deleted from the list of Finnish species: Tabanidae: Haematopota italica Meigen, 1804. Stra- tiomyidae: Odontomyia hydroleon (Linnaeus, 1758). Asilidae: Cyrtopogon ma- culipennis (Macquart, 1834), Machimus gonatistes (Zeller, 1840). Bombyliidae: Phthiria canescens Loew, 1846, Anastoechus nitidulus (Fabricius, 1794), Villa fasciata (Meigen, 1804), Villa longicornis Lyneborg, 1965, Villa panisca (Rossi, 1790). Therevidae: Cliorismia ardea (Fabricius, 1794), Cliorismia rustica (Pan- zer, 1804). Empididae: Wiedemannia fallaciosa (Loew, 1873), Empis punctata Meigen, 1804, Hilara hirtella Collin, 1927, Hilara pseudochorica Strobl, 1892, Hilara scrobiculata Loew, 1873, Hilara tetragramma Loew, 1873, Rhampho- myia poissoni (Theren, 1966), Rhamphomyia tibialis Meigen, 1822. Hyboti- dae: Crossopalpus abditus Kovalev, 1972. The Finnish records of Rhagionidae: Chrysophilus splendidus (Meigen), Asilidae: Epitriptus arthriticus (Zeller), Epitriptus cingulatus (Fabricius), Choerades fimbriata (Meigen) and Hyboti- dae: Drapetis incompleta Collin, 1926 are considered dubious, and they are not included in the checklist. The total number of species found from Finland in these families is now 483. Jere Kahanpää, Finnish Environment Institute, P.O. Box 140, FI-00251 Helsinki, Finland. E-mail: kahanpaa@iki.fi Kaj Winqvist, Zoological Museum, University of Turku, FI-20014 Turku, Finland E-mail: [email protected] Introduction have been recorded from Finland. In a global About 25 years have elapsed since Hackman comparison Finnish flies are known relatively published his checklist of Finnish Diptera well. Nevertheless, a lot of work is still need- (1980). A new checklist of Finnish flies is ed to cover gaps of faunistic and taxonomical needed already now, since the nomenclature knowledge in this large insect group. Because has had great changes in many fly families the study of such a big amount of species with and the taxonomical knowledge has consider- the available small resources is slow the new ably advanced. To date, about 3600 fly species checklist will be published in suitable smaller Kahanpää & Winqvist: Check-list of Finnish flies 11 parts. Updated checklists for families Doli- earlier checklists (Frey 1911, 1913, 1941, chopodidae (Kahanpää & Grichanov 2004) Hackman 1980, the Palaearctic Catalogue, nu- and Syrphidae (Haarto & Kerppola 2004) merous individual articles). Each record was have already been published. In this paper we then checked against existing museum mate- present a new checklist of Finnish flies for su- rial. The junior author examined the Diptera perfamilies Xylophagoidea, Stratiomyoidea, Fennica collection in the Zoological Museum, Tabanoidea, Nemestrinoidea, Asiloidea and University of Helsinki (MZH), the universi- Empidoidea (excl. Dolichopodidae). ty collections in Turku and Oulu, the insect Richard Frey published two papers dea- collections held by natural history and con- ling with the fly families covered here already servation societies in Kuopio and Forssa and in the early 20th century (Frey 1911, 1913). the private collections of Dr. Antti Haarto and Later, checklists of the whole fauna of Fin- the authors. Richard Frey’s collection (now a nish Diptera were published by Frey (1941) part of MZH Diptera collection) and the in- and Walter Hackman (1980). Ever since 1921 sect collection of the Department of Applied new additions and deletions of species of all Biology (University of Helsinki, now also a Finnish insect groups have been published in part of MZH) were examined by the senior 5-year intervals, first by Hellén 1921-1976, author, who also helped with some empidid then by Silfverberg 1981-2001. Until 1986 genera in MZH. Critical specimens were sent these articles were published in Notulae En- to specialists for identification. New species tomologicae, subsequently in Entomologica for Finland found during the identification Fennica. Also the Catalogue of Palaearctic work were added into the list. Diptera (Soós & Papp 1984-1994) contains For a majority of species at least one cor- faunistic information on Finnish flies. rectly identified Finnish reference specimen This checklist covers the area of the sta- was found in the abovementioned collections. te of Finland by the current definition. After If no such specimen was found, we cleared out the Second World War the borders of Finland the reasons why these species were originally were moved especially in Karelia and Kuusa- reported from Finland. In most such cases it mo region. Old articles published before 1945 was possible to classify all published records include many species which were recorded either as misidentifications or as originating only in the areas now belonging to Russia. In from the current Russian territory. A mere li- Hackman’s checklist this transition of borders terary record was accepted as evidence for a is mainly taken into consideration, but not in species being Finnish only if the identifyer all cases. Also catalogues published by foreign was known and could be considered a specia- researchers may neglect this border problem. list of the species-group or family in question. For example both Hackman’s checklist and If no records were confirmed, the species was Palaearctic catalogue regard two Psilocephala deleted from the checklist. A few borderline (Therevidae) species as Finnish even though cases were tagged dubious and are listed in they have never been recorded in the territory a separate cathegory after the checklist. The of post-war Finland. status of all nontrivial species is presented in detail in Results. Criteria for inclusion The compilation of this checklist was initi- Structure of the checklist (appendix A) ated by preparing a ”proto-checklist”, which The classification of superfamilies follows included all the earlier published records and mainly the Nearctic Manual, Vol. 3 (McAlpi- 12 Kahanpää & Winqvist: Check-list of Finnish flies ne 1989). This choice is purely pragmatic; it Table 1. Number of species (by family) recorded from is used in most recent checklists for European Finland in published checklists (Frey 1911-13, Frey countries. Subtaxa of each superfamily (fa- 1941, Hackman 1980, this paper). Species in older checklists are counted only if they are included in the milies, subfamilies, tribes, genera) are listed current checklist; misidentifications and exclusively alphabetically. Russian species have been excluded. Species are listed in alphabetic order un- der the genus name. Synonyms are listed here Family Checklist published only if they have been used in major Finnish 1911-13 1941 1980 2004 Xylophagidae 2 4 4 5 papers, especially in the previous checklists. Athericidae 1 1 1 1 All names used in the previous checklists are Rhagionidae 14 15 14 16 included. Preoccupied names (preocc.) and Tabanidae 27 30 37 40 emendations (emend.) are labelled according- Stratiomyidae 21 25 27 27 ly. Xylomyidae 1 1 1 1 Misidentifications (misident.) are listed Acroceridae 3 4 5 5 only if the species was reported as new for Asilidae 30 33 34 35 Finland under the given name. In this case the Bombyliidae 16 18 18 18 Mythicomyiidae 0 1 1 1 author name and year refers to the publication Scenopinidae 2 2 2 3 of the Finnish record. Species with notes in Therevidae 10 12 17 17 Results are marked with an asterisk (*) after Atelestidae 0 1 1 2 the species name. Empididae 125 142 151 172 Hybotidae 79 98 129 137 Results Microphoridae 2 2 2 3 The number of species found from Finland Total 333 389 444 483 new % 14% 12% 8% in the families included in this study is 483. Table 1 shows the total number of species for each family, and a comparison with the spe- dae: Cliorismia ardea (Fabricius, 1794), Clio- cies numbers recorded in previous checklists. rismia rustica (Panzer, 1804). Empididae: Only the species that have been found in post- Wiedemannia fallaciosa (Loew, 1873), Empis war Finland were counted in all cases. Seven- punctata Meigen, 1804, Hilara hirtella Col- teen species have previously not been recor- lin, 1927, Hilara pseudochorica Strobl, 1892, ded from the country and are here added to Hilara scrobiculata Loew, 1873, Hilara tetra- the Finnish fauna. The following species have gramma Loew, 1873, Rhamphomyia poissoni erroneously been reported from Finland - but (Theren, 1966), Rhamphomyia tibialis Mei- not necessarily included in previous checklists gen, 1822. Hybotidae: Crossopalpus abditus - and are here deleted from the list of Finnish Kovalev, 1972. The circumstances leading to species: Tabanidae: Haematopota italica Mei- the deletion of each species are presented in gen, 1804. Stratiomyidae: Odontomyia hydro- the notes below. leon (Linnaeus, 1758). Asilidae: Cyrtopogon The Catalogue of Palaearctic Diptera lists maculipennis (Macquart, 1834), Machimus Rhagionidae: Chrysophilus splendidus (Mei- gonatistes (Zeller,