10 © Sahlbergia Vol. 10: 10–27, 2005

Check-list of Finnish flies: families – Microphoridae Jere Kahanpää & Kaj Winqvist

Kahanpää, J. & Winqvist, K. 2005: Check-list of Finnish flies: families Xy- lophagidae – Microphoridae. — Sahlbergia 10: 10–27. Helsinki, Finland, ISSN 1237–3273.

A checklist is presented for the Finnish species of the fly families Xylophagidae, , , , , Tabanidae, , , , , , , Atelesti- dae, , and Microphoridae. Seventeen species previously not recorded from Finland are here added to the Finnish fauna. The following species have been erroneously reported from Finland and are here deleted from the list of Finnish species: Tabanidae: Haematopota italica Meigen, 1804. Stra- tiomyidae: hydroleon (Linnaeus, 1758). Asilidae: Cyrtopogon ma- culipennis (Macquart, 1834), Machimus gonatistes (Zeller, 1840). Bombyliidae: Phthiria canescens Loew, 1846, Anastoechus nitidulus (Fabricius, 1794), Villa fasciata (Meigen, 1804), Villa longicornis Lyneborg, 1965, Villa panisca (Rossi, 1790). Therevidae: Cliorismia ardea (Fabricius, 1794), Cliorismia rustica (Pan- zer, 1804). Empididae: Wiedemannia fallaciosa (Loew, 1873), Empis punctata Meigen, 1804, hirtella Collin, 1927, Hilara pseudochorica Strobl, 1892, Hilara scrobiculata Loew, 1873, Hilara tetragramma Loew, 1873, Rhampho- myia poissoni (Theren, 1966), Rhamphomyia tibialis Meigen, 1822. Hyboti- dae: Crossopalpus abditus Kovalev, 1972. The Finnish records of Rhagionidae: Chrysophilus splendidus (Meigen), Asilidae: Epitriptus arthriticus (Zeller), Epitriptus cingulatus (Fabricius), Choerades fimbriata (Meigen) and Hyboti- dae: Drapetis incompleta Collin, 1926 are considered dubious, and they are not included in the checklist. The total number of species found from Finland in these families is now 483.

Jere Kahanpää, Finnish Environment Institute, P.O. Box 140, FI-00251 Helsinki, Finland. E-mail: kahanpaa@iki.fi

Kaj Winqvist, Zoological Museum, University of Turku, FI-20014 Turku, Finland E-mail: [email protected]

Introduction have been recorded from Finland. In a global About 25 years have elapsed since Hackman comparison Finnish flies are known relatively published his checklist of Finnish Diptera well. Nevertheless, a lot of work is still need- (1980). A new checklist of Finnish flies is ed to cover gaps of faunistic and taxonomical needed already now, since the nomenclature knowledge in this large group. Because has had great changes in many fly families the study of such a big amount of species with and the taxonomical knowledge has consider- the available small resources is slow the new ably advanced. To date, about 3600 fly species checklist will be published in suitable smaller Kahanpää & Winqvist: Check-list of Finnish flies 11 parts. Updated checklists for families Doli- earlier checklists (Frey 1911, 1913, 1941, chopodidae (Kahanpää & Grichanov 2004) Hackman 1980, the Palaearctic Catalogue, nu- and Syrphidae (Haarto & Kerppola 2004) merous individual articles). Each record was have already been published. In this paper we then checked against existing museum mate- present a new checklist of Finnish flies for su- rial. The junior author examined the Diptera perfamilies Xylophagoidea, , Fennica collection in the Zoological Museum, , , and University of Helsinki (MZH), the universi- (excl. ). ty collections in Turku and Oulu, the insect Richard Frey published two papers dea- collections held by natural history and con- ling with the fly families covered here already servation societies in Kuopio and Forssa and in the early 20th century (Frey 1911, 1913). the private collections of Dr. Antti Haarto and Later, checklists of the whole fauna of Fin- the authors. Richard Frey’s collection (now a nish Diptera were published by Frey (1941) part of MZH Diptera collection) and the in- and Walter Hackman (1980). Ever since 1921 sect collection of the Department of Applied new additions and deletions of species of all Biology (University of Helsinki, now also a Finnish insect groups have been published in part of MZH) were examined by the senior 5-year intervals, first by Hellén 1921-1976, author, who also helped with some empidid then by Silfverberg 1981-2001. Until 1986 genera in MZH. Critical specimens were sent these articles were published in Notulae En- to specialists for identification. New species tomologicae, subsequently in Entomologica for Finland found during the identification Fennica. Also the Catalogue of Palaearctic work were added into the list. Diptera (Soós & Papp 1984-1994) contains For a majority of species at least one cor- faunistic information on Finnish flies. rectly identified Finnish reference specimen This checklist covers the area of the sta- was found in the abovementioned collections. te of Finland by the current definition. After If no such specimen was found, we cleared out the Second World War the borders of Finland the reasons why these species were originally were moved especially in Karelia and Kuusa- reported from Finland. In most such cases it mo region. Old articles published before 1945 was possible to classify all published records include many species which were recorded either as misidentifications or as originating only in the areas now belonging to Russia. In from the current Russian territory. A mere li- Hackman’s checklist this transition of borders terary record was accepted as evidence for a is mainly taken into consideration, but not in species being Finnish only if the identifyer all cases. Also catalogues published by foreign was known and could be considered a specia- researchers may neglect this border problem. list of the species-group or family in question. For example both Hackman’s checklist and If no records were confirmed, the species was Palaearctic catalogue regard two Psilocephala deleted from the checklist. A few borderline (Therevidae) species as Finnish even though cases were tagged dubious and are listed in they have never been recorded in the territory a separate cathegory after the checklist. The of post-war Finland. status of all nontrivial species is presented in detail in Results. Criteria for inclusion The compilation of this checklist was initi- Structure of the checklist (appendix A) ated by preparing a ”proto-checklist”, which The classification of superfamilies follows included all the earlier published records and mainly the Nearctic Manual, Vol. 3 (McAlpi- 12 Kahanpää & Winqvist: Check-list of Finnish flies ne 1989). This choice is purely pragmatic; it Table 1. Number of species (by family) recorded from is used in most recent checklists for European Finland in published checklists (Frey 1911-13, Frey countries. Subtaxa of each superfamily (fa- 1941, Hackman 1980, this paper). Species in older checklists are counted only if they are included in the milies, subfamilies, tribes, genera) are listed current checklist; misidentifications and exclusively alphabetically. Russian species have been excluded. Species are listed in alphabetic order un- der the name. Synonyms are listed here Family Checklist published only if they have been used in major Finnish 1911-13 1941 1980 2004 Xylophagidae 2 4 4 5 papers, especially in the previous checklists. Athericidae 1 1 1 1 All names used in the previous checklists are Rhagionidae 14 15 14 16 included. Preoccupied names (preocc.) and Tabanidae 27 30 37 40 emendations (emend.) are labelled according- Stratiomyidae 21 25 27 27 ly. Xylomyidae 1 1 1 1 Misidentifications (misident.) are listed Acroceridae 3 4 5 5 only if the species was reported as new for Asilidae 30 33 34 35 Finland under the given name. In this case the Bombyliidae 16 18 18 18 Mythicomyiidae 0 1 1 1 author name and year refers to the publication Scenopinidae 2 2 2 3 of the Finnish record. Species with notes in Therevidae 10 12 17 17 Results are marked with an asterisk (*) after 0 1 1 2 the species name. Empididae 125 142 151 172 Hybotidae 79 98 129 137 Results Microphoridae 2 2 2 3 The number of species found from Finland Total 333 389 444 483 new % 14% 12% 8% in the families included in this study is 483. Table 1 shows the total number of species for each family, and a comparison with the spe- dae: Cliorismia ardea (Fabricius, 1794), Clio- cies numbers recorded in previous checklists. rismia rustica (Panzer, 1804). Empididae: Only the species that have been found in post- Wiedemannia fallaciosa (Loew, 1873), Empis war Finland were counted in all cases. Seven- punctata Meigen, 1804, Hilara hirtella Col- teen species have previously not been recor- lin, 1927, Hilara pseudochorica Strobl, 1892, ded from the country and are here added to Hilara scrobiculata Loew, 1873, Hilara tetra- the Finnish fauna. The following species have gramma Loew, 1873, Rhamphomyia poissoni erroneously been reported from Finland - but (Theren, 1966), Rhamphomyia tibialis Mei- not necessarily included in previous checklists gen, 1822. Hybotidae: Crossopalpus abditus - and are here deleted from the list of Finnish Kovalev, 1972. The circumstances leading to species: Tabanidae: Haematopota italica Mei- the deletion of each species are presented in gen, 1804. Stratiomyidae: Odontomyia hydro- the notes below. leon (Linnaeus, 1758). Asilidae: Cyrtopogon The Catalogue of Palaearctic Diptera lists maculipennis (Macquart, 1834), Machimus Rhagionidae: Chrysophilus splendidus (Mei- gonatistes (Zeller, 1840). Bombyliidae: Pht- gen), Asilidae: Epitriptus arthriticus (Zeller), hiria canescens Loew, 1846, Anastoechus Epitriptus cingulatus (Fabricius), Choerades nitidulus (Fabricius, 1794), Villa fasciata fimbriata (Meigen) and Hybotidae: Drape- (Meigen, 1804), Villa longicornis Lyneborg, tis incompleta Collin from Finland. We have 1965, Villa panisca (Rossi, 1790). Therevi- found no proof for these records and consider © Sahlbergia Vol. 10: 10–27, 2005 13 them dubious. These species are not included Haematopota italica Meigen: This species in the checklist. was reported as new for Finland by Vuorimies (1984). We have re-examined Notes the specimen from Al: Kökar and it Xylophagus ater Meigen: According to belongs to H. pluvialis Linnaeus. Chandler (1998), the name ater Hybomitra astuta (Osten Sacken): Kauri Meigen, 1804 must be used for the (1976) mentions one specimen (1♀ species commonly called X. compeditus Le: Kilpisjärvi, Malla). There are no Wiedemann in Meigen in Continental specimens from Finland in Finnish Europe. Xylophagus kowarzi (Pleske, museum collections. We accept Kauri’s 1925) is the oldest available name for record as proof for the presence of astuta ‘ater’ of ‘continental’ authors. in Finland. Xylophagus matsumurae Miyatake: 1♂ Tb: (Linnaeus): Rozkošný Saarijärvi, Pyhä-Häkki (697:342), 2.vi.- (1973) records this species from Åland 7.vii.1993, L. Kaila & P. Martikainen Islands and Russian Karelia. We consider leg. 1♂ Tb: Konginkangas, Patrikonmäki, it probable that the record from Åland is a 2.vi.-7.vii.1993, L. Kaila & P. Martikainen result of a geographical mistake, since all leg. These specimens are paratypes of examined museum material comes from X. matsumurae inermis Krivosheina & Russian localities. Krivosheina, 2000. borealis Cole: A specimen from Chrysophilus suomianus (Szilády): The type Ab: Uskela (1♂ mid-19th century, E. J. locality of this species is Enontekiö, Bonsdorff leg.) was identified by W. Finland. According to Szilády’s original Hackman (1970) as this North American description the type specimen should be species. The male genitalia of this in MZH, but no specimens were found specimen were examined by the senior there and the species has never been author and they support Hackman’s entered into the type catalogue. The status identification. The wing venation differs of this species is unresolved until the type slightly from North American forms (as is found. described in Schlinger 1960), but match Ptiolina oculata (Becker): Nartshuk (1995) has the description of the type specimen. synonymised Omphalophora lapponica According to Schlinger it is possible that Frey, 1911 with Ptiolina oculata (Becker, several species are included in the current 1900). The type locality of lapponica is concept of O. borealis. ‘Lapponia fennica’. The type specimen Machimus gonatistes (Zeller): Known from Terijoki in Russia. All alleged gonatistes was collected in 1905 by Bernhard Armas specimens from Finland belong either to Nyman (who later changed his name to Tolmerus or Neoitamus. B. A. Ensiö) somewhere along the rivers Tolmerus pyragra Zeller: New for Finland. Tenojoki and Ounasjoki. 2♂ Sa: Luumäki, Kiurula (57689:35377), Rhagio notatus (Meigen): Missing from 26.vii.2002, J. Kahanpää leg. Hackman (1980), but included in Frey Choerades ursula (Loew): C. ursula was (1941). Two records from Finland: 1♂, synonymized with C. fuliginosa (Panzer) ‘Coll. Nyland.’ and 1♂ Ta: Hattula, year by Lehr (1991). Bosák & Hradsky (2001) 1921, W. Hellén leg. reinstalled ursula as a valid species. 14 © Sahlbergia Vol. 10: 10–27, 2005

Cyrtopogon maculipennis (Macquart): on Hedström (1969) who recorded it There is one specimen in MZH, labelled from Ta: Tammela. He wrote as follows: ‘Tohmajärvi, Åbo, H:fors ?’/’?Fennia’ “The identification is presented with (Nordmann leg). Frey (1911) considered some hesitancy as the specimen is this record suspicious and noted that the unusually small and shows some minor specimen is probably not collected in differences in coloration as compared Finland. We agree with Frey’s opinion with unambiguous halteralis specimens”. and delete C. maculipennis from the Hans Bartch (private comm.) found one checklist. small female in the insect collections Anastoechus nitidulus (Fabricius): There of the Uppsala University (UZIU): 1♀, is one specimen in MZH, bearing the Ta: Jokioinen, 31.vii.1966, A. Väre label ‘Tohmajärvi, Åbo, Helsingfors.’ leg. Bearing in mind the considerable (Nordmann leg). Frey (1911) considered variability of Villa females, this record this record suspicious and noted that the must remain somewhat doubtful. specimen is probably not collected in Villa modesta (Meigen): New for Finland. Finland. We agree with Frey’s opinion The examination of male genitalia and and delete A. nitidulus from the list of external characters revealed that most Finnish species. Finnish specimens identified as V. panisca Villa fasciata (Meigen) and V. longicornis (Rossi) and V. circumdata belong to V. Lyneborg: These species were previously modesta. This species has been found in jointly known as Villa circumdata the Finnish biogeographical provinces Al, (Meigen) and appeared in previous Ab, N, Ta, Sa and Kl. checklists under that name, now a Villa panisca (Rossi): Finnish specimens junior synonym of fasciata. According identified as panisca belong to V. cingulata to Evenhuis & Greathead (1999, 2003) (Meigen) or V. modesta (Meigen). There both species are found in Finland. In are some V. panisca specimens from the description of longicornis Lyneborg Russian Karelia (Sortavala) in MZH. (1965) writes “The total distribution Phthiria canescens Loew: Finnish according of V. circumdata longicornis cannot be to Zaitzev (1989) and Evenhuis & given, but, no doubt, this subspecies will Greathead (1999, 2003). The only represent circumdata in Scandinavia specimen in Finnish collections is from and Finland”. All Finnish specimens of Tytärsaari (Bolshoj Tuters), which is now ‘circumdata’ were re-examined and they in Russian territory. all belong either to V. hottentotta (L.) or Scenopinus sp.: This species was misidentified V. modesta (Meigen). There is one male as Scenopinus vitripennis Meigen by of V. fasciata labelled only ‘96.’ in the Haarto (2000). Its true identity is still Diptera Fennica collection (MZH). This unknown. Several additional specimens kind of label is otherwise found only in are now known from Finland. Central European material in coll. Frey. It Cliorismia ardea (Fabricius) & C. rustica seems that this specimen had been moved (Panzer): All Eastern Fennoscandian from coll. Frey to the Fennica collection records of these species are from Terijoki by mistake. (Zelenogorsk), Russia. Villa halteralis (Kowarz): The presence of Thereva circumscripta Loew: The real T. this species in Hackman (1980) is based circumscripta Loew has not been found © Sahlbergia Vol. 10: 10–27, 2005 15

from northern Europe and all the old the material from MZH (M. Chvàla, pers. Finnish specimens under that name belong comm.). to T. valida Loew. Hilara coracina Oldenberg & H. quadrifaria Meghyperus sudeticus Loew: Not listed in Strobl: According to Chvàla (2002b) Hackman (1980). Chvàla (1983) found all the Finnish material identified as H. one Finnish specimen in MZH (1♀ Sa: quadrifaria belongs to H. coracina. Joutseno, 18.vi.1960, E. Thuneberg leg.). Hilara eviana Straka: New for Finland. Clinocera nigra Meigen: Reported as new for 1♀ Sb: Tuovilanlahti (=Maaninka), Finland by Frey (1925) who based it on an Lundström leg., previously misidentified old record made by Sahlberg. However, as H. gallica. there are no specimens left in Finnish Hilara hirta Strobl & H. hirtella Collin: All collections and the Palaearctic Catalogue Finnish material identified as H. hirtella (Chvála & Kovalev 1989) does not regard belongs to H. hirta (Chvàla 2002a). the species as Finnish. The old record was Hilara hybrida Collin: New for Finland. in all probability a misidentification. Over one hundred specimens have been Wiedemannia fallaciosa (Loew) and W. collected from provinces Al, Sa, Tb, ObS, simplex (Loew): W. simplex is a Holarctic ObN, LkW, Le and Li (in MZH, M. Chvàla species, previously known in Europe as det.). W. impudica Mik (Sinclair 1997). The Hilara hyposeta Straka: New for Finland. 1♂ Finnish specimens were misidentified as Ta: Sysmä, Herneniemi, 5.viii.1912, W. W. fallaciosa. Hellén leg., M. Chvàla det. (in MZH); Empis punctata Meigen: This species is 1♂ Sa: Ristiina, 20.viii.1935, Linnaniemi reported from South Karelia in Fauna leg., M. Chvàla det.; 8♂ 1♀, Kb: Joensuu, Entomologica Scandinavica (Chvàla, Ylämylly, 8.viii.1982, O. Martin leg. (in 1994). There are no verifiable records ZMUC). from Finland; all South Karelian material Hilara implicata Collin: New for Finland. in Finnish Museums comes from localities 2♂ Al: Finström, R. Frey leg.; 1♂ N: now in Russia (Leningrad district). Helsinki, R. Frey leg.; 1♂ Ab: Karjalohja, Hilara albitarsis von Roser: New for Finland. year 1934, Krogerus leg., 1♀, St: Yläne, 2♀ Ab: Kiikala, 18.vi.1931, R. Krogerus Sahlberg leg. (all M. Chvàla det.) leg., M. Chvàla det. (in MZH); 1♀ Ta: Hilara lapponica Chvàla: Described partly Karkkila, Moksi, Moksjärvi 1.-7.vii.1976 from Finnish material (Chvàla 2002b). O. Martin leg. (Zoological Museum, Univ. Hilara longifurca Strobl: New for Finland. of Copenhagen = ZMUC), M. Chvàla det.; Chvála found this species from Finland 1♀ Sa: Luumäki, Pajari (67551:5145), while checking the MZH museum 30.v.2003, J. Kahanpää leg. material in 1980. He found altogether Hilara biseta Collin: New for Finland. 1♀ Sb: twelve specimens of it from the provinces Tuovilanlahti (=Maaninka), Lundström leg.; 1♂ N: Kirkkonummi, Bobäck, 23.- Ab, N, Ka, Om and Sb. (in MZH, M. 24.viii.1919, R. Frey leg., M. Chvàla det. Chvála det.) (in MZH). Hilara pseudochorica Strobl: According to M. Hilara campinosensis Niesiolowski: New for Chvàla (pers. comm.) this species does not Finland. Milan Chvàla found this species occur in Finland, and all previous reports from the provinces Al, Ab, St and Kb in were based on misidentifications. 16 © Sahlbergia Vol. 10: 10–27, 2005

Hilara scrobiculata Loew: According Rhamphomyia (Pararhamphomyia) caudata to Chvàla (pers. comm.) all Finnish Zetterstedt: All specimens hitherto specimens identified as H. scrobiculata determined as R. caudata in Finnish belong to other species. collections and in Finnish entomological Hilara sturmii Wiedemann in Meigen: The literature represent R. aethiops Zetterstedt. species was earlier known in Finland The real R. caudata Zetterstedt, a northern as H. cingulata Dahlbom. Actually, H. species, was described by Frey as R. cingulata is a junior synonym of H.flavipes longestylata. (Meigen), recorded in Southern Sweden Rhamphomyia (P.) helleni Frey: New for but not in Finland (Chvála 1997). Finland. 1♀ LkW: Muonio, Jerisjoki, Hilara tetragramma Loew: Hedström’s old 10.vii.1943, R. Frey leg. (in MZH). literary record (1964) is the only piece of Rhamphomyia (Rhamphomyia) cinerascens evidence indicating the possible presence (Meigen): Listed in Frey (1941) but not of this species in Finland. The specimen in Hackman (1980). In MZH there are has, however, got lost already in the 1960’s several specimens from Russian Karelia, and thus could not be located. Chvála has and one from Finland (1♂ Sb: Kuopio, in his later revision work (2002ab) found 19th century, Lundström leg.). this species only from Central Europe and Rhamphomyia (R.) laevipes (Fallén): Listed in the species complex being difficult, it is Frey (1941) but, probably by mistake, not in unlikely that Hedström’s specimens were Hackman (1980). There are ten specimens correctly identified. from the biogeographical provinces Sa Hilara woodiella Chvàla: New for Finland. and Sb in Finnish collections. 1♀ Ab: Turku, 17.vii.1917, R. Frey leg.; Rhamphomyia (R.) tibialis Meigen: All 1 ex (pin only) N: Espoo, B. Poppius leg., specimens in Finnish museum material M. Chvàla det. (in MZH). determined as this species are either R. Rhamphomyia (Aclonempis) galactoptera sulcata (Meigen) or R. sulcatina Collin. Strobl: New for Finland. 1♂ Sa: Joutseno, According to Bartàk (1982) it is likely that 4.vi.1953, E. Thuneberg leg., M. Bartàk R. tibialis is a Central European species det. (in MZH). missing from Northern Europe. Rhampomyia (Holoclera) trigemina Chelifera astigma Collin: Hellén adds Oldenberg: New for Finland. 1♀ Ab: astigma Collin to the Finnish fauna in his Parainen, Lofsdal, 24.vii.1917, R. Frey list of additions (1952) referring to Frey’s leg., M. Bartàk det. (in MZH). article (1950). That article does not, Rhamphomyia (Megacyttarus) anomala however, give faunistic information of the species. There are no museum specimens Oldenberg: All Finnish specimens in Finland and the Palaearctic catalogue identified as R. anomala belong to R. (Chvála & Wagner 1989) does not regard anomalipennis Meigen. C. astigma as Finnish. Rhamphomyia (M.) poissoni (Trehen): In MZH Hemerodromia adulatoria Collin: New for there is a single specimen determined as Finland. 1♂ Ks: Kuusamo, Kitkajoki, R. poissoni (Trehen) (under the name R. 16.vii.1926, R. Frey leg. (in MZH). tephraea Frey, 1953 nec. Meigen, 1822). Iteaphila furcata (Zetterstedt). There are four Misidentified by Frey, it is a female R. exx. from Ks: Kuusamo (R. Frey leg.) maculipennis Zetterstedt. in MZH. They were all taken near Lake © Sahlbergia Vol. 10: 10–27, 2005 17

Paanajärvi, which is now in Russian Russian Isthmus Karelia are mentioned. territory. According to Tuomikoski (1958) Many of Krogerus’s identifications have this species has also been collected from since been proved erroneous. As no Ks: Salla by himself, but no specimens or samples were found, we could not confirm more detailed records exist. It is probable his records of this species. that also Tuomikoski’s observation is from the Russian side. Acknowledgements Crossopalpus abditus Kovalev: Chvála (1975) We would like to thank everyone who helped reports one specimen from Paanajärvi, us by eximining critical specimens or com- Finland. The locality is in Russia and menting on draft species lists: Dr. Miroslav the species is therefore deleted from the Bartak (Empididae), Dr. Milan Chvála (Hy- Finnish checklist. botidae, Acroceridae), Dr. Neal Evenhuis Platypalpus alpinus Chvála: Chvála (1975) (Bombyliidae), Dr. Kevin Holston (Therevi- mentions this species from Finland but dae), Dr. Milan Hradsky (Asilidae), Mr. Sa- later doubts this record in his monograph muli Lehikoinen (Tabanidae) and Dr. Norman on Platypalpus (1989). We found two E. Woodley (Stratiomyidae). candidate specimens of this species in MZH (provinces N and Ta, collected References by Frey and Hellén respectively), both Barták, M. 1982: The Czechoslovak species of in poor condition. Chvála has recently Rhamphomyia (Diptera, Empididae), with re-examined the specimen from Ta and description of a new species from Central considers it to be alpinus. Europe. — Acta Univ. Carol. (Biol.) 1980: 381–461. Platypalpus annulipes (Meigen): Recently Bosák, J. & Hradský, M. 2001: Some remarks on reported as new for Finland (Kahanpää & the distribution of robber flies (Diptera: Asi- Winqvist 2003). lidae) in Turkey. — Journal of the Ent. Res. Platypalpus fuscicornis (Zetterstedt): New Soc. 3(3):1–28. for Finland. 1♂ N: Espoo, Suomenoja Chandler, P. J. 1998: The identity of Xylopha- (66736:33730), 18.vi.2003, J. Kahanpää gus ater Meigen (Diptera, Xylophagidae). leg. — Dipt. Digest (2nd series) 5: 1–5. Platypalpus scandinavicus Chvála, 1972: Chvála, M. 1975: The Tachydromiinae (Dip. New for Finland. 1♀ Oa: Ilmajoki, Empididae) of Fennoscandia and Denmark. Kivistönmäki, 2♂ (window trap) and 1♀ Scandinavian Science Press (Klampenborg, (pitfall trap), 18.vi.-3.viii.2002, V.-M. Denmark). 336 p. Chvála, M. 1983: The Empidoidea of Fennoscan- Mukkala leg. dia and Denmark II. General Part. Families Microphor anomalus (Meigen): Included in Hybotidae, Atelestidae and Microphoridae. all previous checklists on the basis of Scandinavian Science Press (Klampenborg, misidentified specimens. One male has Denmark). 279 p. since been found among unidentified Chvála, M. 1989: Monograph of northern and cent- material (1♂ Al: Mariehamn, 19.vii.1922, ral European species of Platypalpus (Diptera, R. Frey leg., in MZH). Hybotidae) with data on the occurrence in Czechoslovakia. — Acta Univ. Carol. (Biol.) Microphorella praecox (Loew): The only 32: 209–376. Finnish record of this species is in Chvála, M. 1994: The Empidoidea of Fennos- Krogerus (1932), where specimens from candia and Denmark III. Genus Hilara. E. J. the geographical province ObS and from Brill, Leiden. 187 p. 18 © Sahlbergia Vol. 10: 10–27, 2005

Chvála, M. 1997: A revision of the European spe- Hackman, W. 1970: Mötesreferat – Kokousselos- cies of the Hilara flavipes-group, with new tuksia (20.3.1970). — Notulae Entomol. 50: synonymies and description of a new species. 135. — Studia dipt. 4: 463–472. Hackman, W. 1980: A check list of the Finnish Chvála, M. 2002a: Review of European species Diptera I. and (s. of the Hilara ‘quadrivittata’ group. — Acta str). — Notulae Entomol. 60: 17–48. Univ. Carol. (Biol.) 46: 229–276. Hedström, L. 1964: Lounais-Hämeen kärpäsiä. Chvála, M. 2002b: The Hilara species described — Lounais-Hämeen Luonto 15: 7 7–89. by L. Oldenberg, with description of a new Hedström, L. 1969: Diptera Orthorrapha from species from Lapland. — Entomol. Fenn. 13: South-West Häme, Part 1. — Lounais-Hä- 65–78. meen Luonto 32: 1–6. Chvála, M. & Kovalev, V. G. 1989: Family Hybo- Hellén, W. 1952: Verzeichnis der in den tidae. In Soós & Papp (q.v.), Vol. 6. p. 174- Jahren 1946-1950 für die Fauna Finnlands 227. neuhinzugekommenen Insektenarten. — Not- Chvála, M. & Wagner, R. 1989: Family Empidi- ulae Entomol. 31: 59–80. dae. In Soós & Papp (q.v.), Vol. 6. p. 228- Kahanpää, J. & Grichanov, I. Ya. 2004: A check- 336. list of Finnish long-legged flies (Diptera: Do- Evenhuis, N. L. & Greathead, D. J. 1999: World lichopodidae). — Int. J. Dipterol. Res. 15(1): catalog of bee flies (Diptera: Bombylii- 57–62. dae). Backhuys Publishers, Leiden. xlviii + Kahanpää, J. & Winqvist, K. 2003: Rare flies re- 756 p. corded in Finland during the year 2002 (Dip- Evenhuis, N. L. & Greathead, D. J. 2003: World tera). — Sahlbergia 8(2): 92–95. catalog of bee flies (Diptera: Bombyliidae) Kauri, H. 1976: The Scandinavian populations of [http://hbs.bishopmuseum.org/bombcat/]. Hybomitra polaris (Frey) (Diptera: Tabani- Frey, R. 1911: Zur Kenntnis der Dipterenfauna dae). With redescription of the male and desc- Finlands. — Acta Soc. pro Fauna & Flora ription of H. polaris viddensis n. ssp. — Ent. Fenn. 34(6): 1–57. Scand. 7: 145–149. Frey, R. 1913: Zur Kenntnis der Dipterenfauna Krogerus, R. 1932: Über die Ökologie und Ver- Finlands II. Empididae. — Acta Soc. pro Fau- breitung der Arthropoden der Triebsandge- na & Flora Fenn. 37(3): 1–88. biete an den Küsten Finnlands. — Acta Zool. Frey, R. 1925: Föreningsmeddelanden. Tiedon- Fenn. 12: 1–308. antoja yhdistyksistä. — Notulae Entomol. 5: Lehr, P. A. 1991: Revision of Robber flies of the 62. genus Choerades Walker, 1851, and notes Frey, R. 1941: Diptera Brachycera (excl. Musci- on the structure of the subfamily Laphriinae (Diptera, Asilidae). — Ent. Obozr. 70(3): dae, ). In Enumeratio Insectorum 694–715. Fenniae. VI. Dipera. Helsingin Hyönteisvaih- Lyneborg, L. 1965: A Revised List of Danish toyhdistys. p. 1–31. Bombyliidae, with a Subspecific Division of Frey, R. 1950: Dipterafaunan vid Tana älv i Uts- Villa circumdata Meig. — Entomologiske joki sommaren 1949. — Notulae Entomol. Meddelelser 34: 155–166. 30: 5–18. McAlpine, J. F. (ed.) 1989: Manual of Nearctic Haarto, A. 2000: Taxa new for Finland (Diptera). Diptera. Vol. 3. viii + pp. 1333–1581. Scenopinus vitripennis Meigen (Scenopini- Nartshuk, E. P. 1995: Taxonomic and faunistic data on the Rhagionidae (Diptera, Brachyce- dae). — Entomol. Fenn. 11: 1. ra) of the northern Palaearctic. — Acta Zool. Haarto, A. & Kerppola, S. 2004: Checklist of Fenn. 199: 17–24. Finnish Hoverflies (Diptera, Syrphidae). Rozkošný, R. 1973: The Stratiomyioidea (Dip- — Sahlbergia 9(2): 147–164. tera) of Fennoskandia and Denmark. Fauna © Sahlbergia Vol. 10: 10–27, 2005 19

Ent. Scand. 1. Scandinavian Science Press Soós, Á. & Papp, L. (eds.) 1984-1994: Catalogue ltd., Gadstrup, Denmark. of Palaearctic Diptera. 14 volumes. Akad. Ki- adó Budapest. Schlinger, E. I. 1960: A revision of the genus Og- Tuomikoski, R. 1958: Mitteilungen über die Em- codes Latreille with particular reference to pididen Finnlands. V. Die Gattung Iteaphila species of the western hemisphere. — Proc. Zett. s. str. — Ann. Ent. Fenn. 24(3): 125– U. S. National Mus. 111: 227–336. 133. Vuorimies, J. 1984: Kaksi maalle uutta Haemato- Sinclair, B. J. 1997: Review of the Nearctic spe- pota-lajia. — Notulae Entomol. 64: 203. cies of Wiedemannia Zetterstedt. — Studia Zaitzev, V. F. 1989: Family Bombyliidae. In Soós dipt. 4: 337–352. & Papp (q.v.), Vol. 6. p. 43-169.

APPENDIX A: Checklist of Finnish Diptera, Rhagio Fabricius, 1775 families Xylophagidae to Microphoridae. annulatus (De Geer, 1776) lineola Fabricius, 1794 Order DIPTERA maculatus (De Geer, 1776) Suborder BRACHYCERA notatus (Meigen, 1820) * Infraorder XYLOPHAGOMORPHA scolopaceus (Linnaeus, 1758) Superfamily XYLOPHAGOIDEA tringarius (Linnaeus, 1758) XYLOPHAGIDAE Spania Meigen, 1830 Xylophagus Meigen, 1803 nigra Meigen, 1830 ater Meigen, 1804 * Symphoromyia von Frauenfeld, 1867 = compeditus Wiedemann in Meigen, 1820 sg. Paraphoromyia Becker, 1921 cinctus (De Geer, 1776) crassicornis (Panzer, 1806) junki (Szilády, 1932) kowarzi (Pleske, 1925) Doubtful records in Rhagionidae: Chrysophilus = ater authors, misident. suomianus Szilády, 1934 *. matsumurae Miyatake, 1965 * = maculatus Matsumura, 1916, preocc. TABANIDAE subfamily Chrysopsinae Infraorder tribus Chrysopsini Superfamily TABANOIDEA Chrysops Meigen, 1803 caecutiens (Linnaeus, 1758) ATHERICIDAE divaricatus Loew, 1858 Atherix Meigen, 1803 nigripes Zetterstedt, 1838 ibis (Fabricius, 1798) = lapponicus Loew, 1858 relictus Meigen, 1820 RHAGIONIDAE = melanopleurus Wahlberg, 1848 Chrysopilus Macquart, 1826 rufipes Meigen, 1820 cristatus (Fabricius, 1775) sepulcralis (Fabricius, 1794) = auratus (Fabricius, 1805) = maurus Siebke, 1863 luteolus (Fallén, 1814) viduatus (Fabricius, 1794) nubecula (Fallén, 1814) = pictus Meigen, 1820 Ptiolina Zetterstedt, 1842 = quadratus Meigen, 1820 ? = Omphalophora Becker, 1900 subfamily Tabaninae nigra Zetterstedt, 1842 tribus Haematopotini nigrina Wahlgren, 1854 Haematopota Meigen, 1803 nitida Wahlgren, 1854 crassicornis Wahlberg, 1848 obscura (Fallén, 1814) pluvialis (Linnaeus, 1758) oculata (Becker, 1900) * = italica Vuorimies, 1984, misident. * = lapponica (Frey, 1911) subcylindrica Pandellé, 1883 20 © Sahlbergia Vol. 10: 10–27, 2005

Heptatoma Meigen, 1803 maculicornis Zetterstedt, 1842 pellucens Pandellé, 1883 miki Brauer, 1880 tribus Tabanini sudeticus Zeller, 1842 Atylotus Osten Sacken, 1876 fulvus (Meigen, 1820) Superfamily STRATIOMYIOIDEA plebeius (Fallén, 1817) STRATIOMYIDAE rusticus (Linnaeus, 1767) subfamily Beridinae sublunaticornis (Zetterstedt, 1842) Beris Latreille, 1802 Hybomitra Enderlein, 1922 chalybata (Forster, 1771) arpadi (Szilády, 1923) = fuscipes Frey, 1911, misident. astuta (Osten Sacken, 1876) * clavipes (Linnaeus, 1767) = polaris (Frey, 1915) fuscipes Meigen, 1820 auripila (Meigen, 1820) = geniculata Frey, 1941, misident. = aterrima Frey, 1911, misident. morrisii Dale, 1841 bimaculata (Macquart, 1826) strobli Dušek & Rozkošný, 1968 = tropica authors, misident. = geniculata Frey, 1911, misident. = confinis (Zetterstedt, 1838) = fuscipes Frey, 1911, misident. borealis (Fabricius, 1781) subfamily Clitellariinae = lapponicus (Wahlberg, 1848) Nemotelus Geoffroy, 1762 ciureai (Séguy, 1937) sg. Camptopelta Williston, 1917 = solstitialis authors, misident. nigrinus Fallén, 1817 = schineri Lyneborg, 1959 sg. Nemotelus Geoffroy, 1762 distinguenda (Verrall, 1909) notatus Zetterstedt, 1842 kaurii Chvála & Lyneborg, 1970 uliginosus (Linnaeus, 1767) = borealis authors, misident. Meigen, 1803 lundbecki Lyneborg, 1959 centralis Loew, 1863 = tropica authors, misident. = centralis Frey, 1911, preocc. = fulvicornis authors, misident. = freyi Lindner, 1938 lurida (Fallén, 1817) dives Loew, 1845 montana (Meigen, 1820) trilineata (Linnaeus, 1767) = tropica authors, misident. subfamily Pachygasterinae muehlfeldi (Brauer, 1880) Berkshiria Johnson, 1914 = tropica authors, misident. hungarica (Kertész, 1921) = solstitialis authors, misident. ? = albistylum Johnson, 1914 nigricornis (Zetterstedt, 1842) = barovskii (Pleske, 1922) nitidifrons (Szilády, 1914) ssp. confiformis (Chvála & Neopachygaster Austen, 1901 Moucha, 1971) meromelas (Dufour, 1841) = confinis authors, misident. = meromelaena (Austen, 1901), emend. = conformis (Frey, 1916) = orbitalis (Wahlberg, 1854) sexfasciata (Hine, 1923) Zabrachia Coquillett, 1901 = borealis authors, misident. minutissima (Zetterstedt, 1838) = [borealis] ssp. anderi Kauri, 1951 subfamily Sarginae solstitialis (Meigen, 1820) Duncan, 1837 tarandina (Linnaeus, 1758) formosa (Scopoli, 1763) tropica (Linnaeus, 1758) Loew, 1855 = montana authors, misident. cyaneiventris (Zetterstedt, 1842) = fulvicornis authors, misident. flavicornis (Meigen, 1822) = tuxeni Lyneborg, 1959 polita (Linnaeus, 1758) Tabanus Linnaeus, 1758 Fabricius, 1798 autumnalis Linnaeus, 1761 cuprarius (Linnaeus, 1758) bovinus Linnaeus, 1758 flavipes Meigen, 1822 bromius Linnaeus, 1758 = nigripes Zetterstedt, 1842 cordiger Meigen, 1820 = splendens Hackman, 1972, misident. © Sahlbergia Vol. 10: 10–27, 2005 21

iridatus (Scopoli, 1763) Philonicus Loew, 1849 rufipes Wahlberg, 1854 albiceps (Meigen, 1820) subfamily Rhadiurgus Loew, 1849 Odontomyia Meigen, 1803 variabilis (Zetterstedt, 1838) angulata (Panzer, 1798) Tolmerus Loew, 1849 argentata (Fabricius, 1794) atricapillus (Fallén, 1814) microleon (Linnaeus, 1758) pyragra (Zeller, 1840) * Rondani, 1863 subfamily Dioctriinae viridula (Fabricius, 1775) tribus Dioctriini Geoffroy, 1762 Dioctria Meigen, 1803 singularior (Harris, 1778) atricapilla Meigen, 1804 = furcata Fabricius, 1794 cothurnata Meigen, 1820 = reinhardi Meigen, 1820 XYLOMYIDAE hyalipennis (Fabricius, 1794) Xylomya Rondani, 1861 oelandica (Linnaeus, 1758) czekanovskii (Pleske, 1925) rufipes (De Geer, 1776) = interrupta (Pleske, 1925) subfamily Laphriinae = maculata Sahlberg, 1839, misident. tribus Andrenosomatini = maculata var. sahlbergi Frey, 1960 Andrenosoma Rondani, 1856 albibarbe (Meigen, 1820) Infraorder tribus Laphriini Superfamily NEMESTRINOIDEA Choerades Walker, 1851 ACROCERIDAE fuliginosa (Panzer, 1798) subfamily Acrocerinae gilva (Linnaeus, 1758) Meigen, 1803 ignea (Meigen, 1820) orbiculus (Fabricius, 1787) lapponica (Zetterstedt, 1842) = globulus (Panzer, 1804) marginata (Linnaeus, 1758) = borealis Zetterstedt, 1838 ursula (Loew, 1851) * Ogcodes Latreille, 1796 Laphria Meigen, 1803 borealis Cole, 1919 * flava (Linnaeus, 1761) gibbosus (Linnaeus, 1758) gibbosa (Linnaeus, 1758) nigripes (Zetterstedt, 1838) subfamily Leptogastrinae pallipes Latreille, 1812 Leptogaster Meigen, 1803 cylindrica (De Geer, 1776) Superfamily ASILOIDEA guttiventris Zetterstedt, 1842 ASILIDAE subfamily Stenopogoninae subfamily Asilinae tribus Stegopogonini Asilus Linnaeus, 1758 Cyrtopogon Loew, 1847 crabroniformis Linnaeus, 1758 flavimanus (Meigen, 1820) Dysmachus Loew, 1860 lapponicus (Zetterstedt, 1838) picipes (Meigen, 1820) lateralis (Fallén, 1814) trigonus (Meigen, 1804) luteicornis (Zetterstedt, 1838) Machimus Loew, 1849 pulchripes Loew, 1871 setibarbis Loew, 1849 subfamily Stichopogoninae = intermedius (Holmgren, 1852) tribus Stichopogonini Neoitamus Osten Sacken, 1878 Lasiopogon Loew, 1847 cothurnatus (Meigen, 1820) cinctus (Fabricius, 1781) cyanurus (Loew, 1849) socius (Loew, 1871) BOMBYLIIDAE Neomochtherus Osten Sacken, 1878 subfamily pallipes (Meigen, 1820) tribus Anthracini Pamponerus Loew, 1849 Anthrax Scopoli, 1763 germanicus (Linnaeus, 1758) anthrax (Schrank, 1781) 22 © Sahlbergia Vol. 10: 10–27, 2005

trifasciatus Meigen, 1804 THEREVIDAE = leucogaster Wiedemann in Meigen, 1820 subfamily Therevinae varius Fabricius, 1794 Acrosathe Irwin & Lyneborg, 1981 tribus annulata (Fabricius, 1805) Macquart, 1840 Dialineura Rondani, 1856 capucina (Fabricius, 1781) anilis (Linnaeus, 1761) Micomitra Bowden, 1964 Dichoglena Irwin & Lyneborg, 1981 stupida (Rossi, 1790) nigripennis (Ruthe, 1831) tribus Pandivirilia Irwin & Lyneborg, 1981 Loew, 1869 eximia (Meigen, 1820) maura (Linnaeus, 1758) Psilocephala Zetterstedt, 1838 morio (Linnaeus, 1758) imberbis (Fallén, 1814) Osten Sacken, 1886 Spiriverpa Irwin & Lyneborg, 1981 fenestratus (Fallén, 1814) lunulata (Zetterstedt, 1838) Villa Lioy, 1864 = clausa (Frey, 1911) cingulata (Meigen, 1804) Thereva Latreille, 1796 = circumdatus Pipping, 1858, misident. * cinifera (Meigen, 1830) hottentotta (Linnaeus, 1758) = subfasciata Schummel, 1839 = cingulatus Frey, 1911, misident. fuscinervis Zetterstedt, 1838 modesta (Meigen, 1820) * handlirschi Kröber, 1912 = paniscus Frey, 1911, misident. * = praestans Collin, 1948 occulta (Wiedemann in Meigen, 1820) inornata Verrall, 1909 subfamily Bombyliinae lanata Zetterstedt, 1838 tribus Bombyliini microcephala Loew, 1847 Bombylius Linnaeus, 1758 nobilitata (Fabricius, 1775) discolor Mikan, 1796 plebeja (Linnaeus, 1758) major Linnaeus, 1758 strigata (Fabricius, 1794) minor Linnaeus, 1758 unica (Harris, 1780) = allibarbis Zetterstedt, 1842 = bipunctata Meigen, 1820 = albibarbis, emend. valida Loew, 1847 Systoechus Loew, 1855 = circumscripta Bonsdorff, 1861, misident. ctenopterus (Mikan, 1796) = sulphureus (Mikan, 1796) Superfamily EMPIDOIDEA gradatus (Wiedemann in Meigen, 1820) ATELESTIDAE = leucophaeus Wiedemann in Meigen, 1820 Atelestus Walker, 1837 subfamily Phthiriinae pulicarius (Fallén, 1816) tribus Phthiriini Meghyperus Loew, 1850 Phthiria Meigen, 1803 sudeticus Loew, 1850 * pulicaria (Mikan, 1796) EMPIDIDAE Doubtful records in Bombyliidae: Villa halteralis subfamily Clinocerinae (Kowarz, 1883) *. Clinocera Meigen, 1803 sg. Clinocera Meigen, 1803 MYTHICOMYIIDAE appendiculata (Zetterstedt, 1838) Bezzi, 1902 = var. simplicinervis Frey, 1913 arctica (Zetterstedt, 1838) aucta (Zetterstedt, 1849) SCENOPINIDAE sg. Hydrodromia Macquart, 1835 Scenopinus Latreille, 1802 nivalis (Zetterstedt, 1838) fenestralis (Linnaeus, 1758) stagnalis (Haliday, 1833) = glabrifrons Frey, 1911, misident. wesmaeli (Macquart, 1835) niger (De Geer, 1776) = fontinalis Frey, 1916, misident. sp. * Dolichocephala Macquart, 1823 = vitripennis Haarto, 2000, misident. irrorata (Fallén, 1815) © Sahlbergia Vol. 10: 10–27, 2005 23

ocellata (Costa, 1854) biseta Collin, 1927 * = novemguttata (Strobl, 1893) bistriata Zetterstedt, 1842 Wiedemannia Zetterstedt, 1838 brevistyla Collin, 1927 sg. Eucelidia Mik, 1881 campinosensis Niesiolowski, 1986 * zetterstedti (Fallén, 1826) canescens Zetterstedt, 1849 sg. Philolutra Mik, 1881 chorica (Fallén, 1816) bohemani (Zetterstedt, 1838) clavipes (Harris, 1776) simplex (Loew, 1862) = spinimana Zetterstedt, 1838 = impudica Mik, 1880 clypeata Meigen, 1822 = appendiculata Frey, 1913, misident. coracina Oldenberg, 1916 * = fallaciosa Frey, 1924, misident. = quadrifaria Frey, 1913, misident. sg. Wiedemannia Zetterstedt, 1838 cornicula Loew, 1873 bistigma (Curtis, 1834) discoidalis Lundbeck, 1910 subfamily Empidinae diversipes Strobl, 1892 Empis Linnaeus, 1758 = germanica Engel, 1941 sg. Anacrositchus Bezzi, 1909 eviana Straka, 1976 * lucida Zetterstedt, 1838 femorella Zetterstedt, 1842 sg. Coptophlebia Bezzi, 1909 gallica (Meigen, 1804) hyalipennis Fallén, 1816 griseola Zetterstedt, 1838 sg. Empis Linnaeus, 1758 hirta Strobl, 1892 * acinerea Chvála, 1985 = hirtella Thuneberg, 1967, misident. = cinerea Zetterstedt, 1855, preocc. hybrida Collin, 1961 * bicuspidata Collin, 1927 = nitidula Pipping, 1858, misident. chioptera Meigen, 1804 hyposeta Straka, 1976 * laminata Collin, 1927 implicata Collin, 1927 * nigripes Fabricius, 1794 intermedia (Fallén, 1816) = pennaria Fallén, 1816 = pubipes Loew, 1873 = vernalis Meigen, 1822 interstincta (Fallén, 1816) pennipes Linnaeus, 1758 lapponica Chvàla, 2002 * prodromus Loew, 1867 litorea (Fallén, 1816) staegeri Collin, 1963 longivittata Zetterstedt, 1842 = planetica Hackman, 1980, misident. = longevittata Frey, 1913, emend. syrovatkai Chvàla, 1985 = bivittata Strobl, 1892 = plumipes Zetterstedt, 1842, preocc. lurida (Fallén, 1816) sg. Euempis Frey, 1953 maura (Fabricius, 1776) picipes Meigen, 1804 longifurca Strobl, 1892 * = maculipes Zetterstedt, 1842 = monedula Collin, 1927 tessellata Fabricius, 1794 nigritarsis Zetterstedt, 1838 sg. Kritempis Collin, 1926 = infans Zetterstedt, 1842 livida Linnaeus, 1758 nitidula Zetterstedt, 1838 sg. Platyptera Meigen, 1803 pilipes Zetterstedt, 1838 borealis Linnaeus, 1758 pulchripes Frey, 1913 sg. Xanthempis Bezzi, 1909 quadrifasciata Chvála, 2002 diagramma Meigen, 1835 = quadrivittata Frey, 1913, misident. laetabilis Collin, 1926 sturmii Wiedemann in Meigen, 1822 * stercorea Linnaeus, 1761 = cingulata Frey, 1913, misident. univittata Loew, 1867 submaura Collin, 1927 Hilara Meigen, 1822 tanythrix Frey, 1913 abdominalis Zetterstedt, 1838 tenuinervis Zetterstedt, 1838 = obscuritarsis Zetterstedt, 1859 woodiella Chvála, 1999 * albitarsis von Roser, 1840 * Rhamphomyia Meigen, 1822 barbipes Frey, 1908 sg. Aclonempis Collin, 1926 beckeri Strobl, 1892 galactoptera Strobl, 1893 * 24 © Sahlbergia Vol. 10: 10–27, 2005

sg. Amydroneura Collin, 1926 geniculata Meigen, 1830 gibba (Fallén, 1816) = plumipes Pipping, 1858, misident. sg. Holoclera Schiner, 1860 griseola (Zetterstedt, 1838) caliginosa Collin, 1926 = dispar Zetterstedt, 1838 culicina (Fallén, 1816) ? = aperta Zetterstedt, 1859 nigripennis (Fabricius, 1794) helleni Frey, 1922 * sciarina (Fallén, 1816) lapponica Frey, 1955 trigemina Oldenberg, 1927 * = rufipes Zetterstedt, 1838, preocc. umbripennis Meigen, 1822 lividiventris Zetterstedt, 1838 variabilis (Fallén, 1816) lucidula Zetterstedt, 1842 = tenuirostris (Fallén, 1816) marginata (Fabricius, 1787) = pallidiventris (Fallén, 1816) modesta Wahlberg, 1844 sg. Lundstroemiella Frey, 1922 niveipennis Zetterstedt, 1838 dudai Oldenberg, 1927 obscura Zetterstedt, 1838 hybotina Zetterstedt, 1838 obscuripennis Meigen, 1830 sg. Megacyttarus Bigot, 1880 = nitidicollis Frey, 1913 anomalina Zetterstedt, 1838 physoprocta Frey, 1913 = metatarsata Zetterstedt, 1838 pilifer Meigen, 1838 anomalipennis Meigen, 1822 = dentipes Zetterstedt, 1842 = anomala Frey, 1939, misident. * = glaucella Zetterstedt, 1842 crassirostris (Fallén, 1816) = intermedia Frey, 1922 = nigripes Bonsdorff, 1861, misident. plumifera Zetterstedt, 1838 gufitar Frey, 1922 poplitea Wahlberg, 1844 maculipennis Zetterstedt, 1842 praestans Frey, 1913 = tephraea Frey, 1953, misident. pusilla Zetterstedt, 1838 = poissoni, misident. * simplex Zetterstedt, 1849 nodipes (Fallén, 1816) subglaucella Frey, 1922 = spissirostris (Fallén, 1816) tibiella Zetterstedt, 1842 paradoxa Wahlberg, 1844 tipularia (Fallén, 1816) sg. Pararhamphomyia Frey, 1922 unguiculata Frey, 1913 aethiops Zetterstedt, 1838 sg. Rhamphomyia Meigen, 1822 = caudata Frey, 1913, misident. * albosegmentata Zetterstedt, 1838 albidiventris Strobl, 1898 cinerascens (Meigen, 1804) * = woldstedti Frey, 1913 coracina Zetterstedt, 1849 albipennis (Fallén, 1816) dorsata Becker, 1915 albissima Frey, 1913 = phanerostigma Frey, 1918 alpina Zetterstedt, 1838 ignobilis Zetterstedt, 1859 amoena Loew, 1840 = attenuata Frey, 1913 angulifera Frey, 1913 laevipes (Fallén, 1816) * breviventris Frey, 1913 = tephraea Meigen, 1822 caesia Meigen, 1822 latifrons Frey, 1913 = filata Zetterstedt, 1842 morio Zetterstedt, 1838 caudata Zetterstedt, 1838 * nitidula Zetterstedt, 1842 = longestylata Frey, 1916 palmeni Frey, 1913 chibinensis Frey, 1922 plumipes (Meigen, 1804) cribrata Oldenberg, 1927 = vespertilio Zetterstedt, 1842 curvula Frey, 1913 reflexa Zetterstedt, 1838 dentata Oldenberg, 1910 spinipes (Fallén, 1816) fascipennis Zetterstedt, 1838 stigmosa Macquart, 1827 filicaudula Frey, 1949 = conformis Frey, 1913, misident. fuscipennis Zetterstedt, 1838 sulcata (Meigen, 1804) fuscula Zetterstedt, 1838 = tibialis authors, misident. © Sahlbergia Vol. 10: 10–27, 2005 25

sulcatina Collin, 1926 HYBOTIDAE = tibialis Frey, 1913, misident. subfamily Hybotinae = propinqua authors, misident. Hybos Meigen, 1803 vesiculosa (Fallén, 1816) culiciformis (Fabricius, 1775) subfamily Hemerodromiinae femoratus (Müller, 1776) Chelifera Macquart, 1823 = femoralis, emend. concinnicauda Collin, 1927 grossipes (Linnaeus, 1767) = lapponica Frey, 1950 Syndyas Loew, 1857 flavella (Zetterstedt, 1838) nigripes (Zetterstedt, 1842) = melanocephala Bonsdorff, 1861, misident. subfamily frigelii (Zetterstedt, 1838) tribus Ocydromiini precabunda Collin, 1961 Leptodromiella Tuomikoski, 1936 precatoria (Fallén, 1816) crassiseta (Tuomikoski, 1932) stigmatica (Schiner, 1862) Leptopeza Macquart, 1834 trapezina (Zetterstedt, 1838) borealis Zetterstedt, 1842 = melanocephala Bonsdorff, 1861, misident. flavipes (Meigen, 1820) Chelipoda Macquart, 1823 = ruficollis (Meigen, 1820) albiseta (Zetterstedt, 1838) Ocydromia Meigen, 1820 inexpectata Tuomikoski, 1966 glabricula (Fallén, 1816) vocatoria (Fallén, 1816) melanopleura Loew, 1840 Hemerodromia Meigen, 1822 tribus Oedaleini adulatoria Collin, 1927 * Allanthalia Melander, 1927 oratoria (Fallén, 1816) pallida (Zetterstedt, 1838) raptoria Meigen, 1830 Anthalia Zetterstedt, 1838 Phyllodromia Zetterstedt, 1837 schoenherri Zetterstedt, 1838 melanocephala (Fabricius, 1794) Euthyneura Macquart, 1836 subfamily Oreogetoninae albipennis (Zetterstedt, 1842) Anthepiscopus Becker, 1891 gyllenhali (Zetterstedt, 1838) oedalinus (Zetterstedt, 1838) myrtilli Macquart, 1836 Hormopeza Zetterstedt, 1838 = myricae Frey, 1913, misident. copulifera Melander, 1928 Oedalea Meigen, 1820 obliterata Zetterstedt, 1838 freyi Chvàla, 1983 Iteaphila Zetterstedt, 1838 holmgreni Zetterstedt, 1852 macquarti Zetterstedt, 1838 hybotina (Fallén, 1816) nitidula Zetterstedt, 1838 stigmatella Zetterstedt, 1842 = obscura Zetterstedt, 1849 zetterstedti Collin, 1926 = var. fuscipennis Frey, 1913 tribus Trichinini Ragas Walker, 1837 Bicellaria Macquart, 1823 unica Walker, 1837 austriaca Tuomikoski, 1955 subfamily bisetosa Tuomikoski, 1936 Gloma Meigen, 1822 intermedia Lundbeck, 1910 fuscipennis Meigen, 1822 nigra (Meigen, 1824) Heleodromia Haliday, 1833 pilosa Lundbeck, 1910 simplicipes (Zetterstedt, 1842) immaculata Haliday, 1833 spuria (Fallén, 1816) Trichopeza Rondani, 1856 subpilosa Collin, 1926 albocincta (Boheman, 1864) sulcata (Zetterstedt, 1942) = albicincta Frey, 1913 = ssp. vanella Tuomikoski, 1936 longicornis (Meigen, 1822) Trichina Meigen, 1830 bilobata Collin, 1926 Doubtful records in Empididae: Clinocera nigra clavipes Meigen, 1830 Meigen, 1804 *, Chelifera astigma Collin, 1927 *, elongata Haliday, 1833 Iteaphila furcata (Zetterstedt, 1842) *. = picipes Frey, 1956, misident. 26 © Sahlbergia Vol. 10: 10–27, 2005

opaca Loew, 1864 ater (Wahlberg, 1844) = picipes Tuomikoski, 1935 = atra ememd. pallipes (Zetterstedt, 1838) boreoalpinus Frey, 1943 = sexsetosa Frey, 1913 = var. nigricollis Frey, 1913 (pro parte) Trichinomyia Tuomikoski, 1959 brachystylus (Bezzi, 1892) flavipes (Meigen, 1830) = brunneitibia (Strobl, 1899) fuscipes (Zetterstedt, 1838) = autumnalis Frey, nomen nudum = elongata Frey, 1913, misident. brevicornis (Zetterstedt, 1842) subfamily Tachydromiinae calceatus (Meigen, 1822) tribus Drapetini candicans (Fallén, 1815) Chersodromia Haliday, 1851 ciliaris (Fallén, 1816) arenaria (Haliday, 1833) confiformis Chvàla, 1971 cursitans (Zetterstedt, 1819) confinis (Zetterstedt, 1942) = difficilis Frey, 1913, misident. cothurnatus Macquart, 1827 Crossopalpus Bigot, 1857 cryptospina (Frey, 1909) curvinervis (Zetterstedt, 1842) = tantulus (Collin, 1961) curvipes (Meigen, 1822) cursitans (Fabricius, 1775) = aterrima (Haliday in Curtis, 1834) = bicolor (Meigen, 1804) humilis (Frey, 1913) = denominatus (Frey, 1907) nigritellus (Zetterstedt, 1842) ecalceatus (Zetterstedt, 1838) setiger (Loew, 1859) exilis (Meigen, 1822) = setigera, emend. fenestrella Kovalev, 1971 Drapetis Meigen, 1822 flavicornis (Meigen, 1822) sg. Drapetis Meigen, 1822 fuscicornis (Zetterstedt, 1842) * arcuata Loew, 1859 hackmani Chvàla, 1972 assimilis (Fallén, 1815) infectus (Collin, 1926) exilis Meigen, 1822 interstinctus (Collin, 1926) infitialis Collin, 1961 = flavipes Bonsdorff, 1861, misident. ingrica Kovalev, 1972 laestadianorum (Frey, 1913) parilis Collin, 1926 = fuscicornis Frey, 1907, misident. pusilla Loew, 1859 lapponicus Frey, 1943 simulans Collin, 1961 longicornis (Meigen, 1822) sg. Elaphropeza Macquart, 1827 longiseta (Zetterstedt, 1842) ephippiata (Fallén, 1815) = extricatus (Collin, 1926) Stilpon Loew, 1859 luteicornis (Meigen, 1838) sg. Stilpon Loew, 1859 = difficilis (Frey, 1907) graminum (Fallén, 1815) = interjectus (Lundbeck, 1910) tribus Tachydromiini luteus (Meigen, 1804) Platypalpus Macquart, 1927 macula (Zetterstedt, 1842) agilis (Meigen, 1822) maculipes (Meigen, 1822) = hybridus (Frey, 1907) major (Zetterstedt, 1842) albicornis (Zetterstedt, 1842) melancholicus (Collin, 1961) albiseta (Panzer, 1806) minutus (Meigen, 1804) albocapillatus (Fallén, 1815) nigricoxa (Mik, 1884) alpinus Chvàla, 1971 * = poppiusi (Frey, 1913) nigritarsis (Fallén, 1816) alter (Collin, 1961) nigrosetosus (Strobl, 1893) annulatus (Fallén, 1815) nonstriatus (Strobl, 1901) = fascipes (Meigen, 1822) notatus (Meigen, 1822) = fulvipes (Meigen, 1822) pallidicornis (Collin, 1926) annulipes (Meigen, 1822) * pallidiventris (Meigen, 1822) articulatoides (Frey, 1918) = flavipes (Fabricius, 1794), preocc. articulatus Macquart, 1827 pallipes (Fallén, 1815) © Sahlbergia Vol. 10: 10–27, 2005 27

pectoralis (Fallén, 1815) arrogans (Linnaeus, 1761) pseudofulvipes (Frey, 1909) incompleta (Becker, 1900) = coarctatus (Collin, 1926) lundstroemi (Frey, 1913) pseudorapidus Kovalev, 1971 morio (Zetterstedt, 1838) pulicarius (Meigen, 1830) = connexa Bonsdorff, 1861, misident. pygmaeus (Meigen, 1838) punctifera (Becker, 1900) = pallidicoxa (Frey, 1913) = bimaculata Frey, 1941, misident. rapidus (Meigen, 1822) sabulosa Meigen, 1830 = fascipes Bonsdorff, 1861, misident. terricola Zetterstedt, 1819 sahlbergi (Frey, 1909) umbranum Haliday, 1833 = var. stramineipes Frey, 1907 = albitarsis Zetterstedt, 1838 = var. nigricollis Frey, 1913 (pro parte) = annulimana Pipping, 1859, misident. scandinavicus Chvàla, 1972 * Tachypeza Meigen, 1830 sordidus (Zetterstedt, 1838) fennica Tuomikoski, 1932 stabilis (Collin, 1861) fuscipennis (Fallén, 1815) stackelbergi Kovalenko, 1971 heeri (Zetterstedt, 1838) stigmatellus (Zetterstedt, 1842) nubila (Meigen, 1804) strigifrons (Zetterstedt, 1849) truncorum (Fallén, 1815) subbrevis (Frey, 1913) winthemi (Zetterstedt, 1838) sylvicola (Collin, 1826) tuomikoskii Chvàla, 1972 MICROPHORIDAE unguiculatus (Zetterstedt, 1838) subfamily Microphorinae verralli (Collin, 1826) tribus Microphorini vividus (Meigen, 1838) Microphor Macquart, 1827 = albisetoides Chvàla, 1973 anomalus (Meigen, 1824) * zetterstedti Chvàla, 1971 crassipes Macquart, 1827 Symballophthalmus Becker, 1889 = anomalus Frey, 1913, misident. dissimilis (Fallén, 1815) holosericeus (Meigen, 1804) fuscitarsis (Zetterstedt, 1859) = velutinus Macquart, 1827 = scapularis Collin, 1961 Tachydromia Meigen, 1803 Doubtful records in Microphoridae: Microphorella aemula (Loew, 1864) praecox (Loew, 1864) *.