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The Herring Gull Complex (Larus Argentatus - Fuscus - Cachinnans) As a Model Group for Recent Holarctic Vertebrate Radiations
The Herring Gull Complex (Larus argentatus - fuscus - cachinnans) as a Model Group for Recent Holarctic Vertebrate Radiations Dorit Liebers-Helbig, Viviane Sternkopf, Andreas J. Helbig{, and Peter de Knijff Abstract Under what circumstances speciation in sexually reproducing animals can occur without geographical disjunction is still controversial. According to the ring species model, a reproductive barrier may arise through “isolation-by-distance” when peripheral populations of a species meet after expanding around some uninhabitable barrier. The classical example for this kind of speciation is the herring gull (Larus argentatus) complex with a circumpolar distribution in the northern hemisphere. An analysis of mitochondrial DNA variation among 21 gull taxa indicated that members of this complex differentiated largely in allopatry following multiple vicariance and long-distance colonization events, not primarily through “isolation-by-distance”. In a recent approach, we applied nuclear intron sequences and AFLP markers to be compared with the mitochondrial phylogeography. These markers served to reconstruct the overall phylogeny of the genus Larus and to test for the apparent biphyletic origin of two species (argentatus, hyperboreus) as well as the unex- pected position of L. marinus within this complex. All three taxa are members of the herring gull radiation but experienced, to a different degree, extensive mitochon- drial introgression through hybridization. The discrepancies between the mitochon- drial gene tree and the taxon phylogeny based on nuclear markers are illustrated. 1 Introduction Ernst Mayr (1942), based on earlier ideas of Stegmann (1934) and Geyr (1938), proposed that reproductive isolation may evolve in a single species through D. Liebers-Helbig (*) and V. Sternkopf Deutsches Meeresmuseum, Katharinenberg 14-20, 18439 Stralsund, Germany e-mail: [email protected] P. -
The Primary Moult in Four Gull Species Near Amsterdam J. Walters
THE PRIMARY MOULT IN FOUR GULL SPECIES NEAR AMSTERDAM J. WALTERS Received IS April 1978 CONTENTS I. Introduction and study area ... 32 2. Methods and materials ..... 33 2.1. Moult observations on dead gulls 33 2.2. Collection of newly shed primaries 34 2.3. Identification of primaries collected 34 2.4. "Average" primary per date of collection 36 2.5. Average date of shedding per primary 36 2.6. Sampling ......... 37 3. Results ............ 37 3.1. Black-headed Gull, adult .. 37 3.2. Black-headed Gull, sub-adult 38 3.3. Common Gull, adult .. 39 3.4. Common Gull, sub-adult .. 40 3.5. Herring Gull, adult 40 3.6. Herring Gull, sub-adult ... 42 3.7. Great Black-backed Gull, adult 42 3.8. Primary moult and reproduction 42 4. Discussion 43 5. Acknowledgements 46 6. Summary .. 46 7. References . 46 8. Samenvatting ... 46 1. INTRODUCTION AND STUDY AREA This paper deals with the primary moult in Black-headed Gull Larus ridibundus, Common Gull Larus canus, Herring Gull Larus argentatus and Great Black-backed Gull Larus marinus in areas near Amsterdam. Though a few studies on the moult of gulls already exist for the NW-European area, e.g. Stresemann & Stresemann (1966), Stresemann (1971), Barth (1975), Harris (1971), Ingolfsson (1970) and Verbeek (1977), such a study has not been published for a Dutch area and not for four Larus species at the same time and in the same restricted region. Moreover, for the three first mentioned species new information could be obtained on the timing of moult in relation to the incubation period. -
Field Identification of West Palearctic Gulls P
British Birds VOLUME 71 NUMBER 4 APRIL 1978 Field identification of west Palearctic gulls P. J. Grant The west Palearctic list includes 23 species of gulls: more than half the world total. Field guides—because of their concise format—provide inad equate coverage of identification and ageing, which has probably fostered the indifference felt by many bird watchers towards gulls. This five- part series aims to change that attitude nterest in identifying gulls is growing, as part of the recent general I improvement in identification standards, but doubtless also stimulated by the addition to the British and Irish list of no less than three Nearctic species in little over a decade (Laughing Gull Larus atricilla in 1966, Franklin's Gull L. pipixcan in 1970 and Ring-billed Gull L. delaivarensis in 1973). The realisation is slowly dawning that regular checking through flocks of gulls can be worthwhile. Just as important as identification is the ability to recognise the age of individual immatures. This is obviously necessary in studies of popula tion, distribution and migration, but is also a challenge in its own right to the serious bird-identifier. Indeed, identification and ageing go hand- in-hand, for it is only by practising his recognition skills on the common species—of all ages—that an observer will acquire the degree of familiarity necessary for the confident identification of the occasional rarity. The enormous debt owed to D.J. Dwight's The Gulls of the World (1925) is readily acknowledged. That work, however, has long been out of print and its format was designed for the museum and taxonomic worker; the present series of papers will provide a reference more suited to field observers. -
Have We All Missed the Point About Seagulls? Written by Joe Reynolds, Save Coastal Wildlife, Published: 20 February 2020
Have We All Missed the Point About Seagulls? Written by Joe Reynolds, Save Coastal Wildlife, Published: 20 February 2020 Along the picturesque Jersey Shore, a remarkable drama plays out almost every time someone visits a beach. No matter the season, from summer to spring, people will encounter gulls, erroneously known as seagulls. For me, I have a soft spot in my heart for these largely grey-and-white birds. They can be awe-inspiring sea creatures when soaring over the open ocean and dropping down out for the sapphire sky to catch a slimy fish or crusty crab. The sight of them brings to mind a sense of the long soft sand shorelines and sweeping winds and waves over a sun-and-shelled filled beach. Gulls are extraordinary birds. They are able to fly long distances and glide over the open ocean for hours in search of food. Gulls can fly as fast as 28 mph. They can even drink salty ocean water when thirsty. The birds have evolved to have a special pair of glands right above their eyes to flush the salt from their body through openings in their bill. This enables a gull to spend several days foraging for food atop salty ocean waters without needing to return to land just to get a drink of freshwater. James Gorman in 2019 wrote an article in The New York Times entitled: “In Defense of Sea Gulls: They’re Smart, and They Co-Parent, 50/50 All the Way.” He interviewed ornithologist Christopher Elphick from the University of Connecticut who also has a soft spot for gulls. -
Template for Submission of Scientific Information to Describe Areas Meeting Scientific Criteria for Ecologically Or Biologically Significant Marine Areas
Template for Submission of Scientific Information to Describe Areas Meeting Scientific Criteria for Ecologically or Biologically Significant Marine Areas Title/Name of the area: Ukrainian bays Abstract (in less than 150 words) Area located in the northern coast of the Black Sea, that encompasses 2 marine Important Bird and Biodiversity Areas mostly designated for the importance for several species of gulls, terns and seaducks. More than 1% of the global/biogeographic population of 10 seabird species occur in the area. The area encompasses also the non-breeding distribution of two Vulnerable seabirds – the Velvet Scoter Melanitta fusca and the Horned Grebe Podiceps auritus. Introduction Area located in the northern coast of the Black Sea (Figure 1), in the shallow waters (< 100 m deep) of the Ukranian bays of Yagorlyts'ka, Tendrivs'ka, Karkinits'ka and Dzharylgats'ka. The area includes 2 marine Important Bird and Biodiversity Areas (Figure 1), mostly designated for the importance for several species of gulls, terns and seaducks. More than 1% of the global/biogeographic population of 10 seabird species occur in the area (Greater Scaup Aythya marila, Great White Pelican Pelecanus onocrotalus, Great Cormorant Phalacrocorax carbo, Caspian Gull Larus cachinnans, Little Gull Hydrocoloeus minutus, Slender-billed Gull Larus genei, Mediterranean Gull Larus melanocephalus, Common Tern Sterna hirundo, Sandwich Tern Thalasseus sandvicensis and Caspian Tern Hydroprogne caspia: BirdLife International 2017a, 2017b). The area encompasses also the non- breeding distribution of two Vulnerable seabirds – the Velvet Scoter Melanitta fusca and the Horned Grebe Podiceps auritus (BirdLife International 2017c; Figure 2). Location The area is located in the North coast of Black Sea (ca. -
Laridaerefspart1 V1.2.Pdf
Introduction This is the first of two Gull Reference lists. It includes all those species of Gull that are not included in the genus Larus. I have endeavoured to keep typos, errors, omissions etc in this list to a minimum, however when you find more I would be grateful if you could mail the details during 2014 & 2015 to: [email protected]. Grateful thanks to Wietze Janse (http://picasaweb.google.nl/wietze.janse) and Dick Coombes for the cover images. All images © the photographers. Joe Hobbs Index The general order of species follows the International Ornithologists' Union World Bird List (Gill, F. & Donsker, D. (eds.) 2014. IOC World Bird List. Available from: http://www.worldbirdnames.org/ [version 4.2 accessed April 2014]). Cover Main image: Mediterranean Gull. Hellegatsplaten, South Holland, Netherlands. 30th April 2010. Picture by Wietze Janse. Vignette: Ivory Gull. Baltimore Harbour, Co. Cork, Ireland. 4th March 2009. Picture by Richard H. Coombes. Version Version 1.2 (August 2014). Species Page No. Andean Gull [Chroicocephalus serranus] 19 Audouin's Gull [Ichthyaetus audouinii] 37 Black-billed Gull [Chroicocephalus bulleri] 19 Black-headed Gull [Chroicocephalus ridibundus] 21 Black-legged Kittiwake [Rissa tridactyla] 6 Bonaparte's Gull [Chroicocephalus philadelphia] 16 Brown-headed Gull [Chroicocephalus brunnicephalus] 20 Brown-hooded Gull [Chroicocephalus maculipennis] 20 Dolphin Gull [Leucophaeus scoresbii] 31 Franklin's Gull [Leucophaeus pipixcan] 34 Great Black-headed Gull [Ichthyaetus ichthyaetus] 41 Grey Gull [Leucophaeus -
Identification of the Larus Canus Complex
Identification of the Larus canus complex Peter Adriaens & Chris Gibbins ecent decades have seen a wealth of new in- brachyrhynchus) in western North America (for- Rformation published on the identification and merly, the name Mew Gull was often reserved for taxonomy of the so-called ‘large white-headed the Nearctic taxon only). Treatment of these four gulls’. In contrast, the smaller taxa have been rath- taxa in recent literature is often minimal, and er neglected. This is certainly the case with Mew some information is even misleading; for instance, Gull Larus canus, a familiar species that occurs in the two most recent gull monographs heinei is throughout the Northern Hemisphere but whose pictured as a bird that is more or less inseparable geographical variation and field identification re- from canus (Olsen & Larsson 2003, Howell & main poorly understood. Despite its wide distri- Dunn 2007). This hinders identification in various bution, Mew Gull has been treated as a single places around the world; eg, how should we ap- species, consisting of four subspecies: nominate proach the identification of a vagrant kamtschat- L c canus (Common Gull; hereafter canus) in schensis when we do not know what heinei looks Europe (including parts of European Russia), L c like? In some ways, heinei is a key taxon but one heinei (Russian Common Gull; hereafter heinei) that has remained something of a mystery. Another throughout Russia, including large parts of Siberia, problem is that variation in nominate canus is still L c kamtschatschensis (Kamchatka Gull; hereafter not widely understood and is sometimes under- kamtschatschensis) in eastern Siberia, and L c estimated. -
Mytern Booklet
MyTern AUSTRALIA A po ck et g ui de t o th e te rn s of A us tr al ia Caspian Terns, Glenn Ehmke MyTern.indd 1 22/5/19 10:44 am Co nt en ts Introduction ......................................................... 3 General habitat .....................................................4 Nesting habitat ..................................................... 5 Chicks ................................................................. 6 Feeding strategies ................................................. 7 Threats ............................................................... 8 Tips for identifying similar-looking terns.................... 9 !"#$%#&R(")*[,#&(--------------------------------------------------(./ 0#)1&(%1(\%345(-----------------------------------------------------(67 8*1%5*)%13(5#)1&---------------------------------------------------9/ Counting techniques ............................................ 32 BirdLife Australia projects .....................................38 Ke y fo r sp ec ie s’ p ro fil es bp = breeding plumage nbp = non-breeding plumage juv = juvenile plumage Note: the size (cm) of each species refers to body length, the images are not drawn to scale, and the distribution maps are based on all species’ records in Australia (from BirdLife Australia Atlas data). 2 MyTern.indd 2 22/5/19 10:44 am In tr od uc ti on 04%&(:**;,#5(%&(<([#,=($*>"<1%*1(?*)(>*1%5*)%13(@A&5)<,%<1( terns. MyTern contains key information about the habitat, =%&5)%:A5%*1B(1#&5%13()#CA%)#>#15&(<1=(%=#15%[$<5%*1(*?(5#)1&- Terns are in the subfamily Sterninae and are closely related to gulls, but are smaller, slimmer and longer-tailed. Noddies are very closely related and appear similar to terns (but have a wedge-shaped tail and inverse colouration: light cap and dark body). Twenty species of terns and three species of noddies have been recorded in Australia, including migratory &"#$%#&( <1=( D<3)<15&-( 8*&5( 5#)1&( 4A15( [&4( *)( <)54)*"*=&( (insects and crustaceans) by diving, skimming the surface of the water, or on the wing. -
An Atlas of Movements of Southwest Siberian Waterbirds
An atlas of movements of Southwest Siberian waterbirds J. Veen1, A.K. Yurlov2, S.N. Delany3, A.I. Mihantiev2, M.A. Selivanova2 & G.C. Boere4 1 VEDA consultancy, Wieselseweg 110, 7345 CC Wenum Wiesel, The Netherlands 2 Institute for Systematics and Animal Ecology, Siberian Branch of the Russian Academy of Sciences, 11 Frunze Street, Novosibirsk 91, Siberia, Russian Federation 3 Wetlands International, Droevendaalsesteeg 3A, PO Box 471, 6700 AL Wageningen, The Netherlands 4 Dorrewold 22, 7213 TG Gorssel, The Netherlands Wetlands International – 2005 Atlas of movements of Southwest Siberian waterbirds Garganey – Chirok-treskunok Anas querquedula sites (circles), in India and Africa, (2) moulting sites (tri- angles) in Kazakhstan and (3) sites where the species was ringed during summer or migration (squares) in western Europe and Iran. Birds ringed in Europe and Iran (mainly ringed in March and August) may refer to passage migrants moving between Southwest Siberian breeding areas and African winter quarters, as sug- Ringing data gested by Cramp et al. (1977). Figure 24 shows recoveries of Garganeys ringed within Figure 26 shows the distribution of recoveries within the the study area as breeding birds (adults and their young) study area of birds ringed in western Europe, Africa- or during wing moult. There are 22 recoveries (9 breed- Caspian Sea, and India. Recoveries of European ringed ing birds and 13 moulting birds) within the study area, birds have an unexpected mean longitudinal position which have not been depicted. They refer to individuals between African and Indian ringed birds (see mean lati- recovered in the post-breeding, post-fledging or post tudinal positions indicated on map). -
Franklin's Gull in Hampshire: a Species New to Britain and Ireland D
Franklin's Gull in Hampshire: a species new to Britain and Ireland D. F. Billett and P.J. Grant Plates 50-51 At 16.40 on 21st February 1970 J. T. Smith and D.F.B. were at the southern tip of Farlington Marshes, Hampshire, when a small, dark- winged gull approached low along the shore and settled immediately in front of them about 100 yards away. They were able to watch it for about ten minutes before it circled and flew off. During this time it made several short flights of a yard or two when harassed by Black-headed Gulls Larus ridibundus near-by. Several features suggested that it was something unusual. It was slightly smaller than the Black- headed Gulls and its mantle was rather dark grey. Its bill was short, stout and blackish; its dark red legs were also short and this accen tuated a hunched appearance. The bird had an incomplete black hood and was presumably acquiring summer plumage. In flight its tail appeared pale grey down the centre, contrasting with a pure white rump. Most startling of all, however, was its wing pattern in flight: a rather small black tip, with a white terminal spot or spots, separ ated from the dark grey of the remainder by a broad white area across the primaries which joined a continuous white border along the trailing edge. Both observers had previously seen the first-winter Laughing Gull L. atricilla which stayed in the Weymouth area of Dorset from 17th February to 6th October 1969 (Brit. Birds, 63: 279, plate 50) and at first they thought that this might be an adult or near-adult of that species. -
IDENTIFICATION of VULNERABLE SPECIES INCIDENTALLY CAUGHT in MEDITERRANEAN FISHERIES Seabirds
IDENTIFICATION OF VULNERABLE SPECIES INCIDENTALLY CAUGHT IN MEDITERRANEAN FISHERIES Seabirds Twenty-seven species of seven families are listed here as seabird taxa typical of the Mediterranean sea, which may either be encountered as bycatch in Mediterranean fisheries or could follow fishing vessels (at different ranges). SEABIRDS Source materials and references Text and artwork modified from: Lars Svensson (Author), Killian Mullarney (Illustrator), Dan Zetterström (Illustrator), Peter J.Grant (Contributor), David A Christie. 2011. Field Guide to Birds of Britain and Europe. PublisherHarperCollins, 448 pages. Bergier, P. ; Thévenot, M. ; Rihane, A. ; El Agbani, M.A. & Qninba, A. 2017. Liste des oiseaux du Maroc. Mise à jour mai 2017 (rév. 4.0). Go-South Bulletin 14 : 43-68. del Hoyo, J.and Collar, N. J. (2014). HBW and BirdLife International Illustrated Checklist of the Birds ofthe World. Lynx Edicions and BirdLife International, Barcelona, Spain and Cambridge, UK. Topography pictures by José Manuel Arcos. SEABIRD CHECKLIST (By Family) PROCELLARIIDAE ..................................................................................... 27 Calonectris diomedea, Scopoli’s shearwater ................................................ 27 Puffinus yelkouan, Yelkouan shearwater ................................................... 28 Puffinus mauretanicus, Balearic shearwater ................................................ 29 HYDROBATIDAE ...................................................................................... 30 Hydrobates pelagicus, -
The Lesser Black-Backed Gull Larus Fuscus in England: How to Resolve a Conservation Conundrum
The Lesser Black-backed Gull in England: how to resolve a conservation conundrum The Lesser Black-backed Gull Larus fuscus in England: how to resolve a conservation conundrum Viola H. Ross-Smith 1*, Robert A. Robinson 1, Alex N. Banks 2, Tim D. Frayling 2, Chris C. Gibson 3 and Jacquie A. Clark 1 *Correspondence author. Email: [email protected] 1 British Trust for Ornithology, The Nunnery, Thetford, Norfolk IP24 2PU, UK; 2 Natural England, Renslade House, Bonhay Road, Exeter EX4 3AW, UK; 3 Natural England, Mail Hub Block B, Whittington Road, Worcester WR5 2LQ, UK. Abstract Lesser Black-backed Gull Larus fuscus numbers in England have fluctuated in recent decades. Both breeding and wintering populations rose sharply in the latter half of the twentieth century, mostly due to increases at a small number of colonies and changes in migratory behaviour. However, there was a decline in breeding birds between 2000 and 2013 (largely because of losses at the same key colonies) and this species is on the Birds of Conservation Concern Amber List. Although protected at various sites in the breeding season, the Lesser Black-backed Gull can be taken under three General Licences issued under the Wildlife and Countryside Act 1981, allowing population control in certain circumstances without specific permission or reporting. There are no sites where the Lesser Black-backed Gull is a protected feature outside the breeding season, although numbers surpass the relevant thresholds at certain roosts. This review paper synthesises available information on the Lesser Black- backed Gull in England to help policy makers resolve this apparent legislative contra - diction and formulate a clearer conservation policy to guide future practice.