Novelties in Bunchosia (Malpighiaceae)

FREDERICK H. DOBSONIII P.O. Box 71, Huntington,Vermont 05462

ABSTRACT. Bunchosia quaesitor has inflorescencesterminating main axes, a characteristicunique to this endemic to southwesternHaiti. Bunchosia pauciflora fromnorthern Haiti was confused by Niedenzu with B. haitiensisUrban & Niedenzu, but B. haitiensisis two-carpellatewith a glabrous oyaryand connate styles,and B. pauciflorais three-carpellatewith a pilose ovary and distinctstyles. Bunchosia articulata, endemic to Cuba, is distinguished from its closest relative, B. swartziana Griseb.,by its petals with glandular limb margins,basally articulatedpeduncles, and elliptic leaf blades. Bunchosia macilenta, fromBahia, Brazil,can be divorcedfrom Bunchosia acuminata,another new species fromthe same region, by its smaller, more meagre inflorescenceswith fewer flowers and smaller leaves with 4-6 abaxial glands. Bunchosia systyla (Niedenzu) Dobson was originally described as a formof B. fluminensis(Griseb.) Griseb. but is given species status here.

NEW ANTILLEAN TAXA 2 mm long, 1.7 mm diam., the styles connate in the bud. Fruitsnot seen. Bunchosia quaesitor Dobson, sp. nov. (fig. 1).- This species is endemic to southwesternHai- TYPE: Haiti, Massif de la Hotte, Western ti. The specificepithet quaesitor, "the searcher," Group,Pestel on the road to Delcour, "hard is dedicated to ErikLeonard Ekman (1883-1931), limestone,ca. 400 m, rare shrub,puzzling, whose field work is responsible for much of flowers not well developed, yellowish. I what we know of the flora of Haiti. The type really ought to had stayedhe&re and looked is the only known collection. Unfortunately, for more but we wanted to reach Delcour the specimen had only one mature flowerand and it was late," 26 Aug 1927, E. L. Ekman that lacked petals and stamens; its style was H 8976 (holotype: S!). broken off,but the three-carpellateovary was Inter omnia species inflorescentiistermina- preserved (fig. 1). Niedenzu examined this libus axibus principalibusinsignis, inflorescen- specimen in 1929 and placed it in B. punicifolia tiis simpliciter racemosis paucifloris eglandu- (annotation label on specimen), another Hai- losis, ovario 3-carpellatoglabro. tian endemic. This species is easily distin- Shrubs. Branchletsgreen, with minute, scat- guished fromB. punicifolia,however, by inflo- tered, T-shaped trichomes 0.2-0.25 mm long. rescencesterminating main axes and by bearing Older branchessoon glabrous and whitishwith primarybracteoles, a glabrous ovary, broadly a corkysurface, the lenticels neitherprominent elliptic leaves with minute glandular apical si- nor conspicuous. Leaf blades elliptic, 2.5-5 cm nuses, and two conspicuous abaxial glands on long, 1.5-3 cm broad, the base obtuse, with a the lower third of the lamina (fig. 1). Inflores- minute shallow glandular sinus at the obtuse cences terminatingthe main axes is a charac- apex, the blades with 2 conspicuous circular teristicunique to this species. abaxial glands on the lower third,dark green, glabrous and with 3-5 pairs of evident lateral veins above, paler and glabrous beneath, the Bunchosia pauciflora Dobson, sp. nov. (fig. marginsbarely revolute.Petioles 2-3 mm long, 2).-TYPE: Haiti, Massif du Nord, Anse-a- glabrous or with scatteredtrichomes. Inflores- foleur, Morne Cheneau, steep mountain- cences simple-racemose,eglandular, with scat- side, ca. 600 m, "Same as m. Brigand plant. tered trichomes,terminal on the main axes, 2- Obs. All Bunchosia blacken the paper in 4-flowered,2 primarybracteoles at the base of 'which they lie," 25 Jun 1925, E. L. Ekman the inflorescenceaxis, the axes 9-11 mm long, H 4396 (holotype:S!; isotypes:IJ!, NY!, TEX!, the secondary bracteoles to 1 mm long, the pe- US!). duncles basally articulated,to 3.5 mm long, the Inflorescentiis axillaribus simpliciter race- tertiarybracteoles up to 1 mm long from the mosis paucifloriseglandulosis, habitu simili B. articulation.Sepals triangular,2.5-3 mm long, haitiensissed ab eo ovario 3-carpellato piloso ciliate, the glands 8, 1.5-2 mm long. Petals and stylisdistinctis facile distinguendo. stamens not seen. Ovary 3-carpellate,glabrous, Shrubs 1.5-2 m high. Branchletsappressed- 269 270 SYSTEMATIC BOTANY [Volume 8

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I L FIG. 1. Bunchosiaquaesitor. a. Habit (scale 1 cm). b. Abaxial leaf surfaceshowing glands on the lower 1/3 of the lamina and the glandular sinus at the apex. c. Inflorescenceterminal on the main axis, note the primarybracteoles at the base of the inflorescenceaxis. d. Detail of indumenton inflorescenceaxis. Relative size of cut-outis indicated by two parallel lines on the whole structurein c. e. Sepal with glands (abaxial view). f. Transverse section near base of 3-carpellate ovary. Scales for b and c= 5 mm; e and f = 1 mm. Drawn fromthe holotype, EkmanH 8976. There has been some restorationof flowersto the naked axis on the specimen. 1983] DOBSON: BUNCHOSIA 271

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FIG. 2. Bunchosiapauciflora. a. Habit (scale= 1 cm). b. A portion of inflorescenceaxis showing two floweringpeduncles (in bud). c. Indument on the inflorescenceaxis. Relative size of cut-outc is indicated by two parallel lines on the whole structurein b. d. Sepal with glands (abaxial view). e. Petal with erose limb margin (drawn from bud). f. Pistil, note appressed-pilose ovary and connivent styles (drawn from bud). g. Fruitingpeduncle, note persistentstyle bases. Scales for b, e, and g = 5 mm; for d and f, 1 mm. a- f fromthe holotype; g fromEkman H 2865. 272 SYSTEMATIC BOTANY [Volume 8 pilose (T-shaped, rust-coloredtrichomes 0.28- distinguish B. paucifloraare its two- to four- 0.55 mm long). Older branches glabrous and flowered inflorescences,fruits with three dis- lenticellate. Leaf blades elliptic to obovate, 2- tinct persistentstyle bases, and northern dis- 8.5 cm long, 1-3.5 cm broad, the base acute to tribution. shortlyattenuate, apex acute, the blades with 2 minute abaxial glands near the margin on the Bunchosia articulataDobson, sp. nov. (fig.3).- lower third,green glabrous and with 3-4 pairs TYPE: Cuba, Havana, hill near Mariano, of evident lateral veins above, paler green, gla- about 8-10 ft high, 25 Jun 1908, Leon 527 brous but the midvein with scatteredtrichomes (holotype: NY!). beneath, the margins barely revolute. Petioles Inflorescentiis axillaribus simpliciter race- glabrescent,2-5 mm long, the stipules to 1 mm mosis, pedunculis florum5-9 mm longis basi long inconspicuous. Inflorescences axillary, articulatis,petalis tertiusvel quartus interiori- simple-racemose, leafless, eglandular, ap- bus glandulosis prope unguem et margine pe- pressed-pilose or with scatteredtrichomes, the tali intimibasaliter vel omnino glandulosis. primarybracteoles absent, axes 1.3-3.5 cm long, Shrubs or trees 2-4 m high. Branchletswith 2-4-flowered, the secondary bracteoles 1-1.5 scatteredT-shaped trichomes0.26-0.7 mm long mm long, the peduncles basally articulated2.5- to glabrescent. Older branches glabrous and 4.5 mm long, the tertiarybracteoles paired 1- lenticellate.Leaf blades elliptic to obovate, 3.5- 1.5 mm long from the articulation. Mature 12 cm long, 1.5-5 cm broad, the base acute to flowers not seen. Flower buds 3-4 mm diam. shortly attenuate, the apex obtuse to shortly Sepals triangular,3-3.8 mm long, ciliate, the acuminate,sometimes retuse-tipped, the blades glands 8, 2-2.5 mm long. Petals, all 5 with erose with 2(-4) abaxial glands abuttingthe midvein limb margins,3.5-4 mm long, the.claws 1.5-2 fromthe base up to the lower 1/3 of the lamina, mm long, the limbs 2-3 mm long, 2-3 mm diam. glabrous and with 3-5 pairs of evident lateral Stamens 2.5 mm long, connate for 1/3 their veins above, glabrescentbeneath, the margins length,anthers 1 mm long, the connective yel- barelyrevolute. Petioles glabrous,5-9 mm long, low. Pistil up to 3 mm long, the ovary 3-car- the stipules to 1 mm long, inconspicuous. In- pellate, appressed-pilose, 1-1.5 mm long, 1-1.5 florescencesaxillary, simple-racemose, leafless, mm diam., the stylesconnivent with persistent with scattered trichomes, the primary brac- bases. Fruits,9-13 mm long, 8-11 mm diam., teoles absent, the axes 2.5-8 cm long, 4-10- with widely scattered,minute trichomes or gla- flowered,the secondary bracteoles 0.7-2.5 mm brous. Seeds 3, or 2 by abortion. long, the peduncles basally articulated5-9 mm Other specimensexamined. Haiti, Massifdu long, the tertiarybracteoles paired, 1-1.5 mm Nord, Bayeux, Morne Brigand, E. L. EkmanH long from the articulation,one or neither of 2865 (A, IJ,NY 2, TEX, US). the pair glandular. Mature flowers 13-22 mm Bunchosiapauciflora is presumablyendemic to diam. Sepals ovate, 2.5-4 mm long, ciliate, the northernHaiti. Niedenzu examined the holo- glands 8, 1.5-2 mm long. The 4 outermostpet- typein 1926 (annotationlabel on specimen) and als with erose limb margins,or the 3rd and 4th placed it in B. haitiensis,which is two-carpellate inner, glandular near the claw, 7-10 mm long, with a glabrous ovary and connate styles; the claws 2.5-4 mm long, the limbs 5-7 mm whereas B. pauciflorais three-carpellatewith an long, 5-8 mm diam., the innermost 7-10 mm ovary that is appressed-pilose and has conni- long, the claw 3-4 mm long, the limb basally vent styles.Niedenzu and Urban indicated two to completelyglandular, the limb 4-6 mm long, connate stylesin B. haitiensis(1921) in the orig- 4-6 mm diam. Stamens 3-4.5 mm long, connate inal description,but in 1928 Niedenzu emend- for 1/4to 1/2their length, the anthers 1.2-1.6 mm ed and described the ovary as three- long, the connectiveyellow. Pistil4-5 mm long, locular and glabrous and cited the following the ovary3-carpellate, glabrous, 1.5-2 mm long, specimens: E. L. EkmanH 488 (type of B. haitien- 1-2 mm diam., the styles connate with persis- sis); H 4396 (type of B. pauciflora);and H 2865 tent bases. Fruitsred 6-10 mm long, 6-10 mm (also B. pauciflora).This suggests that Niedenzu diam. Seeds 3, or 2 by abortion. either overlooked or chose to disregard carpel Other specimens examined. CUBA. Havan- number and indument of the ovary as impor- na: Bejucal (Cacahual), Alain 4014 (GH, IJ); tant specific characters.Other characters that Caimito, Webster3691 (GH); Cojimar, Ekman 1983] DOBSON: BUNCHOSIA 273

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FIG. 3. Bunchosiaarticulata. a. Habit (scale 1 cm). b. Abaxial leaf.surface.showing.glands oni the.lower 1/3 of lamina. c. Flowering peduncle. d. Indument on the inflorescenceaxis. Relative size of cut-outd is indicated by two parallel lines on the whole structurein c. e. Sepal with glands (abaxial view). f. Petals- limb margins of 3rd and 4th outermostbasally glandular (near the claw); the innermostwith completely glandular limb margin. g. Fruitingpeduncle. Scales for b, c, f, and g = 5 mm; for e, 1 mm. a-f fromthe holotype; g fromN. L. Britton410. 274 SYSTEMATIC BOTANY [Volume 8

13315 (A, S, US); Jamaica,Leon and Roig 11458 NEWBRAZILIAN TAXA (GH, NY, US); Lomas de Comoa, Ekman13447 (S); 14124 (BM, F, G, S); Santiago de las Vegas, Bunchosia macilenta Dobson, sp. nov.-TYPE: Leon 14902(GH, US); Sierra de Camoa, Leonand Brazil, Bahia, Blanchet 1512 (holotype: G!; Albear 7319 (NY); between Vedado and Mari- isotypes: F!, P!). ano, Ekman831 (NY, S). Mantazas: Mantazas, Inflorescentiis axillaribus simpliciter race- Brittonand Wilson410 (NY 2); Pinar del Rio, mosis 6-8 floribus, marginibus limborum pe- Sierra de Anate, Ekman13028 (G, S); Sierra de talorum omnino glandulosis, foliis ellipticis vel los Organos, Ekman 12946 (S). Santa Clara: obovatis 4-6 glandibus abaxialibus dimidio in- Cienfuegos, Combs362 (F, GH, MO, NY, US); feriore laminae, ovario 3-carpellato glabro, sty- Trinidad Hills, Buenos Aires, Glen Ames (Mt. lis distinctis. Harvard), Jack8079 (A, NY, US). No locality: Habit unknown. Branchlets appressed-pilose Firtuns.n. (NY); C. Wright2149 (G 2, GH, MO). (T-shaped trichomes 0.7-1.25 mm long). Older Bunchosiaarticulata is endemic to Cuba. It is branches glabrescent to scattered trichomes, characterizedby its three-carpellateovary, pet- lenticellate. Leaf blades elliptic to obovate, 7- als with basally to completely glandular limb 13 cm long, 3-5 cm broad, the base acute to margins, basally articulated peduncles, few- shortly attenuate, the apex briefly acuminate, floweredinflorescences, and usually ellipticleaf the blades with (2-)4-6 abaxial glands on the blades with two abaxial glands abutting the lower half, with widely scattered trichomes and midvein fromthe base to the lower thirdof the 4-6 pairs of evident lateral veins above, and lamina. This combination of characterseasily scattered trichomes beneath. Petioles ap- divorces it fromB. swartzianaGriseb. to which pressed-pilose, 3-7 mm long, the stipules to 1 it is morphologicallysimilar. mm long, inconspicuous. Inflorescences, axil- Niedenzu confused this species with B. emar- lary, simple-racemose, leafless, appressed-pi- ginataRegel and B. media(Dryander in Aiton) lose, the primary bracteoles absent, the axes 2- DC. In 1898 and later (1914, 1928), he described 3 cm long, 6-8-flowered, the secondary brac- B. emarginataand B. media as three-carpellate teoles 1-2 mm long, the tertiary bracteoles and cited Wright2149 under both species. He paired, 0.7-1.5 mm long, one or neither of the also published (1898) B. emarginatavar. martin- pair glandular. Mature flowers not seen. Sepals icensisUrban & Niedenzu and described it as ovate, 3-3.5 mm long, with scattered tri- two-carpellate.From this treatmentit is diffi- chomes, ciliate, the glands 8, 1.5-2 mm long. cult to assess Niedenzu's concept of B. emargin- Petals, all 5 with completely glandular limb ata Regel. It is doubtfulthat Niedenzu saw ma- margins, the outermost 4 petals 4-6 mm long, terial of this species described by Regel from the claws 1 mm long, the limbs 3-5 mm long, materialcultivated at St. Petersburg.I have seen 3-5 mm diam., the innermost 5 mm long, the Leningrad materialthat fits Regel's original de- claw 2 mm long, the limb 4 mm long, 4 mm scription(1856) as two-carpellate,"stigmate vix diam. Stamens 3-3.5 mm long, connate for 1/3 bilobo." This is the only specimen known to to 1/2their length, the anthers 1 mm long, the me, and I am designating it here as the lecto- connective yellow. Pistil 3.5-4 mm long, the type of B. emarginataRegel. The specimen bears ovary 3-carpellate, glabrous, 1.5 mm long, 1.2 no locality data and has only a single flower, mm diam., the styles distinct. Fruits not seen. which lacks petals and stamens. Other specimen examined. Brazil, Bahia, Thus I have characterized B. emarginataas Blanchet1542 (G). two-carpellate,endemic to Cuba and known Bunchosiamacilenta is known only from Ba- only from the lectotype (LE). Wright2149 is hia, Brazil. The exact locality is not known be- three-carpellate and belongs to B. articulata. cause Blanchet's labels do not provide this in- Bunchosiamedia (Dryander in Aiton) DC. is two- formation and his itinerary is not known. carpellate and is probably a synonymof B. ni- Niedenzu (1928) was not sure of the affinities tida (Jacq.) DC. of this species; he placed the holotype (Blanchet Small (1910) also treated B. emarginataas 1512) under both B. nitida and B. tuberculata. three-carpellate,presumably following Nie- However, B. nitida is two-carpellate and occurs denzu, but he cited no specimen(s) and his in northern South America and the Lesser An- treatmentis doubtful. tilles while B. tuberculatais three-carpellate from 1983] DOBSON: BUNCHOSIA 275 the region of Caracas. This was discussed more fully in my dissertationunder speciesincertae A B. acuminota _, sedis(Dobson 1976). *B. moritimao16 Bunchosiamacilenta can be distinguishedfrom B. maritimaby its smaller, thinner or more 180 meagre inflorescences(hence its epithet: macilenta) with fewer flowers and smaller leaves with shorterpetioles. These same charactersdi- 200 vorce it fromB. acuminata,which, in addition, has glabrescentleaves with two abaxial glands while those of B. macilentausually have fourto 0 ~~~~~~~~~220 six abaxial glands and scatteredtrichomes. The 0 ~~~400 measurementsand dimensions of this species are frombuds; no mature flowerswere avail- 2 420 able.

Bunchosia acuminata Dobson, sp. nov.-TYPE: FIG. 4. Distributionof Bunchosiaacuminata and B. Brazil, Bahia, Munic. Itambe, Sapucaia, ba- maritima. sin of the Rio Pardo, shrub 3 m high on high land in hilly country,8 Nov 1942, Froes 12660/26(holotype: NY!). long, the claw 3-3.5 mm long, the limb 4-4.5 Inflorescentiis axillaribus simpliciter race- mm long, 3-4 mm diam. Stamens 3-3.5 nim mosis 12-16 floribus, marginibus petalorum long, connate for 1/3 to 1/2their length, the an- limborum omnino glandulosis, foliis ellipticis thers 1 mm long, the connective yellow. Pistil 2 glandibus abaxialibus prope basim tertioin- 3.5-3.7 mm long, the ovary 3-carpellate with feriorelaminae, ovario 3-carpellato glabro sty- scattered trichomes, 1.5 mm long, 1-1.3 mm lis distinctis. diam., the styles distinct.Fruits not seen. Shrubs 3 m high. Branchlets appressed-pi- Bunchosiaacuminata is a distinctive species lose (T-shaped trichomes 0.5-0.7 mm long). fromthe basin of the Rio Pardo in eastern Bra- Older branches glabrous, lenticels inconspic- zil (fig. 4). It is separated fromB. maritima(fig. uous. Leaf blades elliptic, 10.5-17.5 cm long, 3- 4) by generally fewer-floweredinflorescences, 4.5 cm broad, the base acute to narrowlyatten- narrowlytriangular to acuminate tertiarybrac- uate, the apex acuminate, the blades with 2 teoles, ovary with scattered trichomes and abaxial glands near the base, glabrous with 6- leaves with two abaxial glands on the lower 10 pairs of evident lateral veins above, glabres- third. It is also readily distinguished from B. cent beneath, except for scattered trichomes macilentaby longer, robust inflorescenceswith along the midvein. Petioles with scatteredtri- more flowers, longer petioles, larger leaves, chomes, 10-12 mm long, the stipules 2-2.5 mm glabrescent, with two abaxial glands on the long. Inflorescences,axillary, simple-racemose, lower third of the lamina and ovary with scat- leafless, appressed-pilose, the primary brac- tered trichomes. teoles absent, the axes 7-9.5 cm long, 12-16- flowered, the secondary bracteoles 2-3 mm Bunchosia systyla (Niedenzu) Dobson, stat. long, the peduncles basally articulated to 1.5 nov.-Bunchosia fluminensisf. 2 systylaNie- mm above the base, 5-7.5 mm long, the tertiary denzu, Arbeiten Bot. Inst. Kbnigl. Ly- bracteoles paired, narrowly triangular to -acu- ceums Hosianum Braunsberg5:41. 1914.- minate, 1.5-2 mm long fromthe articulation,1 TYPE: Brazil,Amazonas, Rio Jurua,Ule 5042 or none of the pair glandular. Mature flowers (G). 14-16 mm diam. Sepals triangular,3.5-4.5 mm Shrubs 3-7 m high. Branchletswith scattered long, ciliate, the glands 8, 2.5-3.5 mm long. T-shaped trichomes 0.35-0.6 mm long. Older Petals, all 5 with completely glandular limb branches glabrescent and lenticellate. Leaf margins,the outermost4 petals 6.5-8 mm long, blades ovate, 10.5-17 cm long, 3.5-7.5 cm broad, the claws 2-2.5 mm long, the limbs 4-5.5 mm the base broadly cuneate to obtuse, the apex long, 4-5.5 mm diam., the innermost7-7.5 mm obtuse to acuminate,the blades (2-)4-6 abaxial 276 SYSTEMATIC BOTANY [Volume 8 glands (barely visible to the naked eye) abut- peduncles articulated further from the base, ting the midvein or removed toward the mar- smaller tertiary bracteoles, leaf blades glabres- gin on the lower 1/3 of the lamina, glabrouswith cent beneath, and an ovary with connate styles, 4-6 pairs of evident lateral veins above, gla- which are found in B. systyla. brescentbeneath, the marginsbarely revolute. ACKNOWLEDGMENTS. This study representsa por- Petioles glabrous, 5-10 mm long, the stipules tion of a doctoral dissertationsubmitted to the De- 1.5-2 mm long. Inflorescences, axillary,simple- partment of Botany, University of Connecticut, in racemose, leafless, appressed-pilose, the pri- 1976. I thank Drs. H. W. Pfeifer,H. N. Andrews, and mary bracteoles absent, axes 3-4 cm long, 14- F. R. Trainor for their help; Mary Jane Gallo for the 24-flowered,the secondarybracteoles 1-2.5 mm excellent drawings; and Molly Hubbard and stafffor long, the peduncles articulated 1.5-3.5 mm maps and other illustrations. above the base, 6-11 mm long, the tertiary bracteoles paired, one of the pair glandular, to LITERATURECITED 1 mm long fromthe articulation.Sepals ovate, DOBSON,III, F. H. 1976. A taxonomicrevision in the 3-4.5 mm long, appressed-pilose,the glands 9, genus BunchosiaRich. ex H. B. K. (Malpighi- 3-4 mm long. Petals, all 5 with completely aceae)-the three-carpellatespecies. Ph.D. dis- glandular limb margins,the outermost4 petals sertation,Univ. Connecticut,Storrs. 5-7.5 mm long, the claws 2-3 mm long, the MACBRIDE,J. F. 1950. Bunchosia.(In Flora of Peru.) limbs 4-5.5 mm long, 3.5-6 mm diam., the in- Field Mus. Nat. Hist., Bot. Ser. 13 (part 3, no. 3): nermost6-6.5 mm long, the claw 3 mm long, 855-861. the limb 3-4 mm long, 3-4 mm diam. Stamens NIEDENZU,F. 1898. [De genere Bunchosia.] Index Lectionum 3-4 mm long, connate for 1/3 to 1/2their length, Lyceo Regio Hosiano Brunsbergensi.... anthers 1 mm long, the connective yellow. Pis- [Braunsberg:Typis Heyneanis.] 1914. Malpighiaceae til 3.5-4.5 mm long, the ovary 3-carpellate,gla- Americane III. Arbei- ten Bot. Inst. Kbnigl. Lyceums Hosianum brous, 1.5-2 mm 1-1.5 mm long, diam., the Braunsberg5:37-45. styles connate. Fruitsnot seen. . 1928. Bunchosia.Pp. 642-674 in Das Pflan- Niedenzu treatedthis species as a formof B. zenreich,IV.141, Heft 94, ed. A. Engler. Leipzig: fluminensis,which, according to Macbride (1950), Wilhelm Engelmann. is now more properly called B. maritima.Bun- and I. URBAN. 1921. Malpighiaceae (in Plan- chosiasystyla occurs along the drainage of the tae Haitienses ..., by I. Urban). Ark. Bot. 17(7): Rio Juruain the state of Amazonas in west cen- 35. tralBrazil, which is quite distantfrom the range REGEL,E. 1856. Bunchosiaemarginata. Index Sem. Hort. of B. maritimain eastern Brazil (fig. 4). Other Bot. Petrop. p. 35. SMALL,J. K. 1910. featuresthat separate B. systylafrom B. maritima Malpighiaceae. Pp. 160-166 in NorthAmerican Flora, vol. 25, part 2. New York: are the shortermore congested inflorescences, New York Bot. Gard.