Of New Species. by DE KIMMINS, Department Of
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602 Mr. D. E. Kimmins on the LX1I.-Notes on the Genera Megalomus Rambur and Nesobiella, gen. nov. (Neuroptera), with Descriptions of new Species. By D. E. KIMMINS,Department of Entomology, British Museum (Natural History ). THIS paper was originally intended to contain de- scriptions of new species only, but after a comparison of the male genitalia of the new species with that of existing European species had been made, it was decided to enlarge its scope, and to figure and redescribe the genitalia of all the old species of which material was available. This paper thus contains details of three new and of six old species and also of a new genus formed to receive a species which can no longer be retained within the genus Megalomus. Rambur, in 1842, described the genus Megalomus, and included in it four species, Hemerobius phalmnoides L., Megalomus tortricoides Ramb., M. pyraloides Ramb., and M. tineoides Ramb. The first had already had a genus (Drepanepteryx Leach) created for it, and from the remaining three Banks, in 1906, selected tortricoides as the type of the genus. He was in error, however, in stating that tortricoides Rambur was a synonym of hirtus L. Sinoe Rambur the number of described species has been increased to over thirty, giving a wide distribution, although I know of no records of the genus from Australia, New Zealand, or Africa (other than N. Africa). Until recently little use has been made in systematics of the male genitalia in the Hemerobiidze apart from the superior appendages and the ninth sternite. Tjeder, in his paper on the Boriomyia persica group, showed that valuable characters were to be found in the other genital structures, even where the superior appendages were closely related in form. For the purpose of studying the male genitalia the abdomen was removed and cleared in KOH and stained, when necessary, to render the delicate structures more visible. The drawings were made with camera lucida while the specimens were in fluid, so that different aspects of the same example could be studied. The preparations were afterwards mounted in Canada balsam. The drawings Genera Megalomus and Nesobiella. 603 of the various species are not all to the same scale, as variations in size made this somewhat impracticable. The $ genitalia in the genus Megalomus consists of the following parts :-A pair of " superior appendages " (" claspers," " cerci "), attached to the ninth tergite ; an arch-shaped chitinous framework, the tenth sternite, carrying a median process terminating in a pair of hooks, and on each side of the median process a slender spine ; a pair of elongate finger-like parameres, generally fused at their bases ; and a thin hypandrium, sagittate, in side view somewhat resembling the prow of a boat. I am much indebted to Mr. K. J. Morton for the loan of preparations and sketches which, in the early stages of this paper, enabled me to verify my deter- minations of our European species. Megalomus hirtus (L.). (Fig. 1.) Attached to the ninth tergite are a pair of " superior appendages " or " claspers." These are broad at their bases, subtriangular, their upper margins sinuate, and towards their apices bearing a few stout teeth. Ninth sternite short, outer margin rounded. The tenth sternite arch-shaped, made up of a pair of lateral upright plates, joined above by a narrow bridge. Seen from the side each lateral plate is rounded, its posterior margin strangly excised, the lower angle of the excision being expanded and compressed dorso-ventrally. From above the con- necting bridge shows a deep rectangular excision in the centre of the tailward margin. Attached to the upper portion of the arch is a median process which in-the genus Megalornus generally bears two hooks at its apex. It is a rectangular plate, suspended by its upper angles from the arch, the longitudinal median portion convex and its lower edge curved slightly upwards. From this margin there arise two hooks, fused at their bases and slightly divergent. In side view each hook is directed upward for about half its length, where it is bent outward and suddenly narrowed, terminating in a somewhat enlarged and obliquely truncated apex. At the base of the median process, on each side of it, there is a slender sword-like " lateral spine," gradually tapering and bent upwards in its outer half. Parameres situated within the arch, curved, finger-like, fused at their bases, where 604 Mr. D. E- Kimmins on the the margins are expanded upwards and outwards. On the lower surface, at the base, is a thin keel, and the apical portion of each paramere is membranous and covered with microscopic hairs. Hypandrium very lightly chitin- ized, sagittate, apex blunt, slightly excised ; a shallow keel ventrally along the median line. Fig. 1. A, genitalia from side (hypandrium omitted) ; B, tenth sternite from above and behind ; C, tenth sternite and parameres fmm behind ; D, parameres from beneath ; E, hypandriurn from above. Hegalomus tortricoides Rambur. (Fig. 2.) $.-Superior appendages from the side roughly tri- angular, upper margin sinuous, with a rounded projection towards the apex, covered with setiferous tubercles. Apex of the appendage obliquely truncate, armed with Genera Megalomus and Nesiobella. 605 two or three stout teeth. Ninth sternite from the side only slightly produced, from beneath transverse. Tenth sternite with the sides broad, rounded basally, lower Fig. 2. Megalomus tortricoides Ramb., 2. A, apex of abdomen, lateral ; B, tenth st,ernite, lateral ; C, parameres, lateral ; D, tent,h sternite from behind ; E:, hypandrium, dorsal ; F, hypandrium, lateral. apical margin triangularly produced. Median process short, rectangular, attached to the arch by a pair of divergent arms. Apical hooks long, fused for their Ann. & Mag. N. Hist. Ser. 10. Vo1. xvi. 41 606 Mr. D. E. Kimmins on, the basal two-thirds, a deep median channel between them in the fused portion. Apices narrow, abruptly angled basalwards. Parameres fused in the basal half, narrow, curved upwards, apices finely serrate beneath, a semi- circular expansion on each side at the base, and a small Fig. 3. Megalomus tincoidcs Ramb., 3. A, apex of abdomen, lateral ; B, tenth sternite and parameres, latoral ; C, tenth stornite, dorsal ; D, hypanclrium, dorsal. keel ventrally. Hypandrium sagittate, concave, lateral margins thickened, reflexed, and projecting beyond the truncate apex to enclose a rectangular excision. Genera Megalomus and Nesobiella. 607 Megalomus tineoides Rambur. (Fig. 3.) Superior appendages triangular, apices produced and rounded, furnished with a number of stout teeth. Ninth sternite short, apical margin somewhat produced. Tenth sternite with broad rounded side-lobes, the connecting bridge with its apical margin produced and cleft centrally so as to form two rounded lobes. Median process in lateral view with convex margins, the apical hooks long, broad and fused at their bases, apices turned downward and outward. Lateral spines slender, bent upward, and in the only example before me crossed. Parameres long, upcurved, their bases fused and dilated outward and upward, apices slightly dilated, and bearing a narrow patch of short hairs externally. Hypandrium sagittate, margins reflexed. Megalomus setulosus Walker. (Fig. 4.) 8.-Superior appendages from the side pear-shaped, broadest below. Prom the lower margin there arises a short, stout, somewhat inwardly-directed, finger-like process, its apex bearing a slender bristle. Lower apical angle on its inner side with two or three stout black spines. Ninth sternite from the side triangular, from below transverse, margin scarcely produced. Tenth sternite produced above at its centre in a strong bifurcate process. Lateral plates with their portions dilated tailward and twisted somewhat outward. Median Aprocess long and narrow, with a thin keel along its lower surface, angled near its apex, which bears a pair of stout hooks, fused at their bases. From the side these hooks are straight on their inner and convex on their outer sides. Lateral spines slender, bent upwards at about a right- angle midway, their apices obliquely truncate and divergent. Parameres broad and fused at their bases, in side view tapering for about a quarter of their length, then parallel and directed upward, apices bearing a number of microsco~ichairs. A rounded keel at the base ventrally. Hypandrium sagittate, concave, apex excised, outer margins thickened. Kashmir, near Srinagar, 5200 ft., Apr.-June, 1923 (F.J. Mitchell). Walker's type appears to be a female. 41* Mr. D. E. Kimmins on the Fig. 4. Megalomus setulosus Walk., 3. A, apex of abdomen, lateral ; B, tenth sternite and parameres, lateral ; C, tenth sternite from behind ; D, hypandrium, dorsal ; E, hypan- drium, lateral. Genera Megalornus and Nesobiella. 609 Megalomus mastus Banks. (Fig. 5.) Superior appendages triangular, directed downward, apices acute, the upper sides somewhat sinuous. Ninth sternite produced in the centre. Tenth sternite with broad oval side-lobes and a narrow connecting bridge. Median process broad, margins raised, its apex bearing the usual pair of fused hooks. These are slender, tapering and curving downward, their apices curled under and Fig. 5. A, apex of abtlomcn, lateral ; U, tcnth sternite, lateral ; C, parameres, lateral ; D, tenth sternitc, dorsal ; E, hyparitlrium, dorsal. outward. Lateral spines slender, acute, angled upward at about mid-length. Pararrieres slender, curved upward and lightly fused towards their bases. The actual bases are free, divergent, strongly upcurved, their lower margins with a clavate dilatation. Apices rounded in lateral, acute in dorsal view, their upper and outer margins finely serrate. Hypandrium sagittate, lateral margins 610 Mr. D. E. Kimmins on the reflexed and produced apically to form rounded lobes enclosing a narrow excision. Drawn from an example in the British Museum which was collected at Santa FB (the type-locality) by Prof. Cockerell. The wings agree very well with those of a female determined by Dr.