Received: 11 October 2019 | Revised: 10 February 2020 | Accepted: 18 March 2020 DOI: 10.1111/zsc.12422 ORIGINAL ARTICLE Densely sampled phylogenetic analyses of the Lesser Short- toed Lark (Alaudala rufescens) — Sand Lark (A. raytal) species complex (Aves, Passeriformes) reveal cryptic diversity Fatemeh Ghorbani1 | Mansour Aliabadian1,2 | Ruiying Zhang3 | Martin Irestedt4 | Yan Hao3 | Gombobaatar Sundev5 | Fumin Lei3 | Ming Ma6 | Urban Olsson7,8 | Per Alström3,9 1Department of Biology, Faculty of Science, Ferdowsi University of Mashhad, Mashhad, Iran 2Zoological Innovations Research Department, Institute of Applied Zoology, Faculty of Science, Ferdowsi University of Mashhad, Mashhad, Iran 3Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing, China 4Department of Bioinformatics and Genetics, Swedish Museum of Natural History, Stockholm, Sweden 5National University of Mongolia and Mongolian Ornithological Society, Ulaanbaatar, Mongolia 6Xinjiang Institute of Ecology and Geography, Chinese Academy of Sciences, Xinjiang, China 7Systematics and Biodiversity, Department of Biology and Environmental Sciences, University of Gothenburg, Göteborg, Sweden 8Gothenburg Global Biodiversity Centre, Gothenburg, Sweden 9Animal Ecology, Department of Ecology and Genetics, Evolutionary Biology Centre, Uppsala University, Uppsala, Sweden Correspondence Mansour Aliabadian, Department of Abstract Biology, Faculty of Science, Ferdowsi The taxonomy of the Lesser/Asian Short-toed Lark Alaudala rufescens–cheleensis University of Mashhad, Mashhad, Iran. complex has been debated for decades, mainly because of minor morphological dif- Email:
[email protected] ferentiation among the taxa within the complex, and different interpretations of the Per Alström, Animal Ecology, Department geographical pattern of morphological characters among different authors. In addi- of Ecology and Genetics, Evolutionary Biology Centre, Uppsala University, tion, there have been few studies based on non-morphological traits.