Invasive Exotic Shrub Honeysuckles Honeysuckles Shrub Exotic Invasive Year
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Lonicera Maackii Amur Honeysuckle
Lonicera maackii (F.J. Ruprecht) C.J. Maximowicz Amur Honeysuckle (Caprifolium maackii, Xylosteon maackii) • Lonicera maackii is also known as Bush Honeysuckle; Amur Honeysuckle, which is a native of temperate Asia, forms a large deciduous irregular twiggy mound 12 to 15 tall with a similar or greater spread; growth rates can be rapid and plants can crowd out native vegetation; this is facilitated by the ability of Amur Honeysuckle to tolerate heavy shade until released; the gray pubescent buds of L. maackii can be distinguished from those of L. tatarica which are flattened and glabrous; the bluish green to medium green leaves of L. maackii are longer, 2 to 3 long, have mostly broadly cuneate to occasionally rounded bases, and acute to long acuminate tips while the medium to dark green leaves of L. tatarica are on average shorter, 1½ to 2½ long, with rounded to slightly cordate bases, and acute to short acuminate tips; new twigs have a whitish pubescence and may be flushed purple-red, later maturing to a gray-brown; excavated brown pith is present between the solid nodes; fall colors vary from almost nonexistent to a poor yellow; the tan-brown to gray- brown bark of old trunks develops vertical thin exfoliations. • A profusion of ¾ to 1 long white flowers are borne in spring, fading to a creamy yellow as they mature; flowers are generally reminiscent of those of L. japonica, flowers are produced in spring, occurring during or shortly after the foliage emerges; the flowers are followed by shiny, bright red, ¼to ½ diameter berries, produced mostly in one to two pairs per node, which hold into late winter if not eaten by birds or other wildlife. -
Bumble Bee Clearwing Moths
Colorado Insects of Interest “Bumble Bee Clearwing” Moths Scientific Names: Hemaris thysbe (F.) (hummingbird clearwing), Hemaris diffinis (Boisduval) (snowberry clearwing), Hemaris thetis (Boisduval) (Rocky Mountain clearwing), Amphion floridensis (Nessus sphinx) Figure 1. Hemaris thysbe, the hummingbird clearwing. Photograph courtesy of David Order: Lepidoptera (Butterflies, Moths, and Cappaert. Skippers) Family: Sphingidae (Sphinx Moths, Hawk Moths, Hornworms) Identification and Descriptive Features: Adults of these insects are moderately large moths that have some superficial resemblance to bumble bees. They most often attract attention when they are seen hovering at flowers in late spring and early summer. It can be difficult to distinguish the three “bumble bee clearwing” moths that occur in Colorado, particularly when they are actively moving about plants. The three species are approximately the same size, with wingspans that range between 3.2 to 5.5cm. The hummingbird clearwing is the largest and distinguished by having yellow legs, an Figure 2. Amphion floridensis, the Nessus olive/olive yellow thorax and dark abdomen with sphinx. small patches. The edges of the wings have a thick bordering edge of reddish brown. The snowberry clearwing has black legs, a black band that runs through the eye and along the thorax, a golden/olive golden thorax and a brown or black abdomen with 1-2 yellow bands. The head and thorax of the Rocky Mountain clearwing is brownish olive or olive green and the abdomen black or olive green above, with yellow underside. Although the caterpillar stage of all the clearwing sphinx moths feed on foliage of various shrubs and trees, damage is minimal, none are considered pest species. -
Lonicera Spp
Species: Lonicera spp. http://www.fs.fed.us/database/feis/plants/shrub/lonspp/all.html SPECIES: Lonicera spp. Choose from the following categories of information. Introductory Distribution and occurrence Botanical and ecological characteristics Fire ecology Fire effects Fire case studies Management considerations References INTRODUCTORY SPECIES: Lonicera spp. AUTHORSHIP AND CITATION FEIS ABBREVIATION SYNONYMS NRCS PLANT CODE COMMON NAMES TAXONOMY LIFE FORM FEDERAL LEGAL STATUS OTHER STATUS AUTHORSHIP AND CITATION: Munger, Gregory T. 2005. Lonicera spp. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/ [2007, September 24]. FEIS ABBREVIATIONS: LONSPP LONFRA LONMAA LONMOR LONTAT LONXYL LONBEL SYNONYMS: None NRCS PLANT CODES [172]: LOFR LOMA6 LOMO2 LOTA LOXY 1 of 67 9/24/2007 4:44 PM Species: Lonicera spp. http://www.fs.fed.us/database/feis/plants/shrub/lonspp/all.html LOBE COMMON NAMES: winter honeysuckle Amur honeysuckle Morrow's honeysuckle Tatarian honeysuckle European fly honeysuckle Bell's honeysuckle TAXONOMY: The currently accepted genus name for honeysuckle is Lonicera L. (Caprifoliaceae) [18,36,54,59,82,83,93,133,161,189,190,191,197]. This report summarizes information on 5 species and 1 hybrid of Lonicera: Lonicera fragrantissima Lindl. & Paxt. [36,82,83,133,191] winter honeysuckle Lonicera maackii Maxim. [18,27,36,54,59,82,83,131,137,186] Amur honeysuckle Lonicera morrowii A. Gray [18,39,54,60,83,161,186,189,190,197] Morrow's honeysuckle Lonicera tatarica L. [18,38,39,54,59,60,82,83,92,93,157,161,186,190,191] Tatarian honeysuckle Lonicera xylosteum L. -
Invasive Plants of the Southeast Flyer
13 15 5 1 19 10 6 18 8 7 T o p 2 0 I n v a s i v e S p e c i e s 1. Chinese Privet, Ligustrum sinense 2. Nepalese Browntop, Microstegium vimineum 3. Autumn Olive, Elaeagnus umbellata 4. Chinese Wisteria, Wisteria sinensis & Japanese Wisteria, W. floribunda 5. Mimosa, Albizia julibrissin 6. Japanese Honeysuckle, Lonicera japonica 7. Amur Honeysuckle, Lonicera maackii 8. Multiflora Rose, Rosa multiflora 9. Hydrilla, Hydrilla verticillata 10. Kudzu, Pueraria montana 11. Golden Bamboo, Phyllostachys aurea 12. Oriental Bittersweet, Celastrus orbiculatus 13. English Ivy, Hedera helix 14. Tree-of-Heaven, Ailanthus altissima 15. Chinese Tallow, Sapium sebiferum 16. Chinese Princess Tree, Paulownia tomentosa 17. Japanese Knotweed, Polygonum cuspidatum 18. Silvergrass, Miscanthus sinensis 19. Thorny Olive, Elaeagnus pungens 20. Nandina, Nandina domestica The State Botanical Garden of Georgia and The Georgia Plant Conservation A l l i a n c e d e f i n i t i o n s you can help n a t i ve Avoid disturbing natural areas, including clearing of native vegetation. A native species is one that occurs in a particular region, ecosystem or habitat Know your plants. Find out if plants you without direct or indirect human action. grow have invasive tendencies. Do not use invasive species in landscaping, n o n - n a t i ve restoration, or for erosion control; use (alien, exotic, foreign, introduced, plants known not to be invasive in your area. non-indigenous) A species that occurs artificially in locations Control invasive plants on your land by beyond its known historical removing or managing them to prevent natural range. -
ON the WOODY PLANTS of Llasca PARK a MAJOR REPORT
.. \ PATTERNS AND INTEI~ SITY OF WHI'l'E•TAILED DEER BROWSING ON THE WOODY PLANTS OF llASCA PARK ~ -·• .-:~~ -"7,"l!'~C .-'";"'l.;'3. • ..., 1; ;-:_ . .. ..... .. ;_., ( •• ~ -· ., : A MAJOR REPORT ~ i ..,; ·:-.:.. _, j i... ...... .. .' J £70 q ... c a:s·:, >rse·- _....,. •e '~· SUBMITTED TO THE.. SCHOOL OF FORESTRY By DONALD N. ORKE •' IN: PARTIAL FULFILU1ENT OF THE REQUIREMENTS. FOR THE DEGREE OF MASTER OF SCIENCE OCTOBER, 1966 - .. ,,..\ ., ..- ./ ACKNOWLEDGEMENTS The author wishes to thank the School of Forestry . :-~~ ..·,. for providing the, financial assistance needed to conduct the study. Also, special thf.llks ·are due to Dr. Henry L. Hansen who provided help:f'ul advice in th• preparation of. this report • .. - ' TABLE OF CONTENTS INTRODUCTION ••••••••••••••••••••••••••••••••••••••••• 1 HISTORY OF THE ITASCA DEER HERD AND ITS EFFECTS ON THE VEGETATION ••••••••••••••••••••••••••• •••••••••• 2 LITERATURE REVIEW •••••••••••••••••••••••••••••••••••• 5 STUDY AREA ••••••••••••••••••••••••••••••••••••••••• • 11 METH<D.DS:i •••• . .. .. .. .. .. .. .. .. .. .•• 13 RESULTS BROWSE PREFERENCES ••••••••••••••••••••••••••• 17 BROWSING ON KONIFER SEEDLINGS •••••••••••••••• 40 DISCUSSION BROWSE PREFERENCES ••••••••••••••••••••••••••• 51 BROWSING ON CONIFER SEEDLINGS •• • • • • • • • • • • • • • .54 SUGGESTIONS FOR FUTURE STUDIES ••••••••••••••• 56 SUMMA.RY • •••••••••••••••••••••••••••••••••••••••••••• 5 7 LITERATURE CITED ••••••••••••••••••••••••••••••• a.• .. 59 APPENDIX •••••••••••••••••••••• e ••••••••••••••••••••• 61 ,-·. INTRODUCTION -
State of Delaware Invasive Plants Booklet
Planting for a livable Delaware Widespread and Invasive Growth Habit 1. Multiflora rose Rosa multiflora S 2. Oriental bittersweet Celastrus orbiculata V 3. Japanese stilt grass Microstegium vimineum H 4. Japanese knotweed Polygonum cuspidatum H 5. Russian olive Elaeagnus umbellata S 6. Norway maple Acer platanoides T 7. Common reed Phragmites australis H 8. Hydrilla Hydrilla verticillata A 9. Mile-a-minute Polygonum perfoliatum V 10. Clematis Clematis terniflora S 11. Privet Several species S 12. European sweetflag Acorus calamus H 13. Wineberry Rubus phoenicolasius S 14. Bamboo Several species H Restricted and Invasive 15. Japanese barberry Berberis thunbergii S 16. Periwinkle Vinca minor V 17. Garlic mustard Alliaria petiolata H 18. Winged euonymus Euonymus alata S 19. Porcelainberry Ampelopsis brevipedunculata V 20. Bradford pear Pyrus calleryana T 21. Marsh dewflower Murdannia keisak H 22. Lesser celandine Ranunculus ficaria H 23. Purple loosestrife Lythrum salicaria H 24. Reed canarygrass Phalaris arundinacea H 25. Honeysuckle Lonicera species S 26. Tree of heaven Alianthus altissima T 27. Spotted knapweed Centaruea biebersteinii H Restricted and Potentially-Invasive 28. Butterfly bush Buddleia davidii S Growth Habit: S=shrub, V=vine, H=herbaceous, T=tree, A=aquatic THE LIST • Plants on The List are non-native to Delaware, have the potential for widespread dispersal and establishment, can out-compete other species in the same area, and have the potential for rapid growth, high seed or propagule production, and establishment in natural areas. • Plants on Delaware’s Invasive Plant List were chosen by a committee of experts in environmental science and botany, as well as representatives of State agencies and the Nursery and Landscape Industry. -
Japanese Honeysuckle Wildland (Lonicera Japonica Thunb.) Gary N
Japanese Honeysuckle Wildland (Lonicera japonica Thunb.) Gary N. Ervin, Ph.D., Associate Professor, Mississippi State University John D. Madsen, Ph.D., Extension/Research Professor, Mississippi State University Ryan M. Wersal, Research Associate, Mississippi State University Fig. 1. Japanese honeysuckle vine climbing up a tree. Fig. 2. Flowers of Japanese honeysuckle. Introduction Problems Created Japanese honeysuckle was introduced from Japan in the early 1800s and now is one of the most commonly encoun- tered exotic weeds in the Mid-South. This species frequently overtops and displaces native plants and forestry species in any habitat, but particularly where natural or human activities creates edges. Japanese honeysuckle also is somewhat shade tolerant and can be found in relatively densely canopied forest. This species perenniates with the aid of well de- veloped root and rhizome systems, by which it also is capable of spreading vegetatively, in addition to rooting at nodes along aboveground stems. Both features contribute substantially to its rapid dominance over native vegetation. Regulations Japanese honeysuckle is listed as a noxious weed in CT, MA, NH, and VT. In the southeast, Japanese honeysuckle is considered a severe invasion threat in KY, SC, and TN. It is considered one of the top ten invasive plants in GA, and is listed as a category one invasive in Florida. It is not presently listed in any noxious weed legislation in southern states. Description Vegetative Growth Japanese honeysuckle exhibits a semi-evergreen to evergreen life cycle and is readily identified during winter by its per- sistent green foliage. Its vines may climb and/or spread along the ground to lengths of 80’. -
Phylogeny and Phylogenetic Taxonomy of Dipsacales, with Special Reference to Sinadoxa and Tetradoxa (Adoxaceae)
PHYLOGENY AND PHYLOGENETIC TAXONOMY OF DIPSACALES, WITH SPECIAL REFERENCE TO SINADOXA AND TETRADOXA (ADOXACEAE) MICHAEL J. DONOGHUE,1 TORSTEN ERIKSSON,2 PATRICK A. REEVES,3 AND RICHARD G. OLMSTEAD 3 Abstract. To further clarify phylogenetic relationships within Dipsacales,we analyzed new and previously pub- lished rbcL sequences, alone and in combination with morphological data. We also examined relationships within Adoxaceae using rbcL and nuclear ribosomal internal transcribed spacer (ITS) sequences. We conclude from these analyses that Dipsacales comprise two major lineages:Adoxaceae and Caprifoliaceae (sensu Judd et al.,1994), which both contain elements of traditional Caprifoliaceae.Within Adoxaceae, the following relation- ships are strongly supported: (Viburnum (Sambucus (Sinadoxa (Tetradoxa, Adoxa)))). Combined analyses of C ap ri foliaceae yield the fo l l ow i n g : ( C ap ri folieae (Diervilleae (Linnaeeae (Morinaceae (Dipsacaceae (Triplostegia,Valerianaceae)))))). On the basis of these results we provide phylogenetic definitions for the names of several major clades. Within Adoxaceae, Adoxina refers to the clade including Sinadoxa, Tetradoxa, and Adoxa.This lineage is marked by herbaceous habit, reduction in the number of perianth parts,nectaries of mul- ticellular hairs on the perianth,and bifid stamens. The clade including Morinaceae,Valerianaceae, Triplostegia, and Dipsacaceae is here named Valerina. Probable synapomorphies include herbaceousness,presence of an epi- calyx (lost or modified in Valerianaceae), reduced endosperm,and distinctive chemistry, including production of monoterpenoids. The clade containing Valerina plus Linnaeeae we name Linnina. This lineage is distinguished by reduction to four (or fewer) stamens, by abortion of two of the three carpels,and possibly by supernumerary inflorescences bracts. Keywords: Adoxaceae, Caprifoliaceae, Dipsacales, ITS, morphological characters, phylogeny, phylogenetic taxonomy, phylogenetic nomenclature, rbcL, Sinadoxa, Tetradoxa. -
Japanese Honeysuckle)
No. 09 March 2010 Lonicera japonica (Japanese Honeysuckle) Initial Introduction and Expansion in Range Introduced to the United States in the early to mid-1800s as an ornamental plant, Lonicera japonica is native to East Asia, including Japan and Korea. It is still promoted by some landscapes architects for its rapid growth and fragrant flowers that linger on the vine throughout most of the summer. Wildlife managers have promoted the plant as winter forage, particularly for deer. Still others are nostalgic about this plant, remembering the sweet nectar they enjoyed as children. Lonicera japonica is now found across the southern United States from California to New England and the Great Lakes Region. Lonicera japonica spreads locally by long aboveground runners and underground rhizomes. The runners develop roots at the nodes (stem and leaf junctions) so this plant often forms dense mats. Under high light conditions, the plants are able to flower and produce fruits that can be dispersed long distances primarily by birds. Description and Biology • Perennial trailing or twining vine. In North Carolina, it is considered semi-evergreen to evergreen. • Young stems are slender, while older stems are hollow and up to 2 inches in diameter with brownish bark that peels in long strips. • Oblong to oval shaped leaves are 1 to 2 and a half inches long arranged in opposite pairs along the stem. Mature leaves have smooth edges and young leaves are often lobed. • White and pale yellow, trumpet-shaped flowers occur in pairs from between the leaves and bloom from late April to August. • Small, nearly spherical, black fruits mature in autumn. -
(Betula Papyrifera) / Diervilla Lonicera Trembling Aspen (Paper Birch) / Northern Bush-Honeysuckle Peuplier Faux-Tremble (Bouleau À Papier) / Dièreville Chèvrefeuille
Forest / Forêt Association CNVC 00238 Populus tremuloides (Betula papyrifera) / Diervilla lonicera Trembling Aspen (Paper Birch) / Northern Bush-honeysuckle Peuplier faux-tremble (Bouleau à papier) / Dièreville chèvrefeuille Subassociations: 238a typic, 238b Alnus viridis, 238c Kalmia angustifolia CNVC Alliance: CA00014 Betula papyrifera – Populus tremuloides – Abies balsamea / Clintonia borealis CNVC Group: CG0007 Ontario-Quebec Boreal Mesic Paper Birch – Balsam Fir – Trembling Aspen Forest Type Description Concept: CNVC00238 is a boreal hardwood forest Association that ranges from Manitoba to Quebec. It has a closed canopy dominated by trembling aspen ( Populus tremuloides ), usually with paper birch ( Betula papyrifera ), overtopping a well-developed to dense shrub layer. The shrub layer includes a mix of regenerating tree species, primarily trembling aspen, balsam fir ( Abies balsamea ), paper birch and black spruce ( Picea mariana ), as well as low shrub species, such as velvet-leaved blueberry ( Vaccinium myrtilloides ), northern bush-honeysuckle ( Diervilla lonicera ) and early lowbush blueberry ( V. angustifolium ). The herb layer is well developed and typically includes bunchberry (Cornus canadensis ), wild lily-of-the-valley ( Maianthemum canadense ), wild sarsaparilla ( Aralia nudicaulis ), yellow clintonia ( Clintonia borealis ), twinflower ( Linnaea borealis ) and northern starflower Source: Natural Resources Canada - (Lysimachia borealis ). The forest floor cover is mainly broad-leaf litter so the moss layer is Canadian Forest Service sparse, with only minor cover of red-stemmed feathermoss ( Pleurozium schreberi) . CNVC00238 is an early seral condition that typically establishes after fire or harvesting. It occurs in a region with a continental boreal climate that grades from subhumid in the west to humid in the east and is usually found on mesic, nutrient-medium sites. -
Viburnum Opulus Var. Americanum
Viburnum opulus L. var. americanum (Mill.) Ait. (American cranberrybush): A Technical Conservation Assessment Prepared for the USDA Forest Service, Rocky Mountain Region, Species Conservation Project May 8, 2006 James E. Nellessen Taschek Environmental Consulting 8901 Adams St. NE Ste D Albuquerque, NM 87113-2701 Peer Review Administered by Society for Conservation Biology Nellessen, J.E. (2006, May 8). Viburnum opulus L. var. americanum (Mill.) Ait. (American cranberrybush): a technical conservation assessment. [Online]. USDA Forest Service, Rocky Mountain Region. Available: http://www.fs.fed.us/r2/projects/scp/assessments/viburnumopulusvaramericanum.pdf [date of access]. ACKNOWLEDGMENTS Production of this assessment would not have been possible without the help of others. I wish to thank David Wunker for his help conducting Internet searches for information on Viburnum opulus var. americanum. I wish to thank Dr. Ron Hartman for supplying photocopies of herbarium specimen labels from the University of Wyoming Rocky Mountain Herbarium. Numerous other specimen labels were obtained through searches of on-line databases, so thanks go to those universities, botanic gardens, and agencies (cited in this document) for having such convenient systems established. I would like to thank local Region 2 botanists Bonnie Heidel of the Wyoming Natural Heritage Program, and Katherine Zacharkevics and Beth Burkhart of the Black Hills National Forest for supplying information. Thanks go to Paula Nellessen for proofing the draft of this document. Thanks go to Teresa Hurt and John Taschek of Taschek Environmental Consulting for supplying tips on style and presentation for this document. Thanks are extended to employees of the USDA Forest Service Region 2, Kathy Roche and Richard Vacirca, for reviewing, supplying guidance, and making suggestions for assembling this assessment. -
Maryum Bhatti1, Aaron Lee1, Hiba Rahman-Vyas1, Dianella G
NAC SUBFAMILY 1A GENE TREE Combining benchwork and bioinformatics to reconstruct the evolutionary history of CUP- SHAPED COTYLEDON in honeysuckles and relatives Maryum Bhatti1, Aaron Lee1, Hiba Rahman-Vyas1, Dianella G. Howarth2, Michael J. Donoghue3, Wendy L. Clement1 1Department of Biology, The College of New Jersey, Ewing, NJ; 2Department of Biological Sciences, St. John’s University, Jamaica, NY; 3Department of Ecology and Evolution, Yale University, New Haven, CT RECOVERING CUC FROM INTRODUCTION DIPSACALES Context Fusion among adjacent parts is a phenomena that occurs Sampling and Isolating CUC throughout flowering plants. CUP-SHAPED COTYLEDON, CUC, • 38 species were selected across the Dipsacales for a member of the NAC Subfamily 1a transcription factors, and has been shown to affect organ boundary formation.1,2 Honeysuckles, species that had no available genomic resources or Lonicera (Caprifoliaceae, Dipsacales), are known for fusing • Degenerate primers were designed to isolate exons petals into long tubes and also exhibit fusion among ovaries, bracts 1 and 2 of CUC1/2 and CUC3 based on reference and leaves. Variation in fusion across 160 species of Lonicera genomes (e.g., Arabidopsis, Petunia, Snapdragon) make them an excellent system to investigate the evolution of • CUC specific primers were also created including fusion. all introns and exons of CUC Goals • Successful reactions near 500 bp were isolated and • Recover CUC from the phylogenetic diversity of Dipsacales, cloned using PCR and cloning; sample target species in which no • Multiple clones per species were sequenced at the genomic data is available. Yale Sequencing on the Hill Facility • Reconstruct a gene tree for NAC Subfamily 1a and CUC using • Sequences were assembled and manually edited both direct sequences gained in this study and data extracted 3 from available genomic resources.