The Type Specimens of Anthomyiidae, Muscidae and Sarcophagidae (Diptera: Calyptratae) Housed at the Instituto – Fundación Miguel Lillo, Tucumán, Argentina
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154Acta zoológicaL. lilloanaD. Patitucci 55 (2): et 154–170,al.: The type 2011 specimens of Anthomyiidae, Muscidae and Sarcophagidae154 The type specimens of Anthomyiidae, Muscidae and Sarcophagidae (Diptera: Calyptratae) housed at the Instituto – Fundación Miguel Lillo, Tucumán, Argentina Patitucci, Luciano Damián; Mulieri, Pablo Ricardo; Mariluis, Juan Carlos Consejo Nacional de Investigaciones Científicas y Técnicas, Buenos Aires, Argentina. ANLIS Dr. Carlos G. Malbrán, Departamento Vectores – CeNDIE, Av. Vélez Sarsfield 563, (1281) Buenos Aires, Argentina. E-mail: [email protected]; [email protected]; [email protected] R e s u m e n — Se estudiaron los ejemplares tipo de Anthomyiidae, Muscidae y Sarcopha- gidae depositados en la Colección Instituto - Fundación Miguel Lillo (IFML). Un total de 13 especímenes tipo de Anthomyiidae (cuatro especies nominales), 271 especímenes tipo de Muscidae (36 especies nominales) y seis especímenes tipos de Sarcophagidae (cuatro espe- cies nominales) fueron examinados. Se provee información para cada taxón. Palabras clave: Anthomyiidae, Muscidae, Sarcophagidae, especímenes tipo, Argentina. A b s t r a c t — The type specimens of Anthomyiidae, Muscidae and Sarcophagidae deposited in the Instituto-Fundación Miguel Lillo (IFML) are revised. A total of 13 type specimens of Anthomyiidae (four nominal species), 271 type specimens of Muscidae (36 nominal species), and six type specimens of Sarcophagidae (four nominal species) were examined. Information for each taxa is provided. Keywords: Anthomyiidae, Muscidae, Sarcophagidae, type specimens, Argentina. INTRODUCTION specimens of Calyptratae deposited in argen- tinean institutions, a list of Anthomyiidae, Calyptratae have long been recognized as Muscidae and Sarcophagidae, kept in Insti- a major lineage of higher Diptera, and con- tuto - Fundación Miguel Lillo, are presented. tain some of the more diverse and successful The insect collection housed in the “Insti- families including Anthomyiidae, Calli- tuto - Fundación Miguel Lillo” (IFML) (Tu- phoridae, Fanniidae, Muscidae, Sarcoph- cumán, Argentina) is one of the most impor- agidae and Tachinidae, among others tant collections of Argentina and South (Yeates et al., 2007). Several type speci- America. Hayward & Golbach (1963) pub- mens of Calyptratae species are deposited in lished a list of the “primary types” of Diptera different institutions in Argentina. The infor- deposited in IFML, with about 1060 type mation on type specimens is useful not only specimens. The entomological collection of in taxonomic research, but also for phyloge- the IFML houses 13 type specimens belonging netic, ecological and conservation studies. to four nominal species of Anthomyiidae, Moreover, the 4th edition of the International 271 type specimens belonging to 36 nominal Code of Zoological Nomenclature recom- species of Muscidae, and six type specimens mends the publication of type material belonging to four nominal species of Sar- housed in different institutions and museums cophagidae. (Article 72 F 4, ICZN, 1999). Recently, a cat- The types of Anthomyiidae and Muscidae alogue of the type specimens of Sarcoph- belong to species described by Snyder agidae housed in the Museo Argentino de (1957). The author deposited holotypes and Ciencias Naturales “Bernardino Rivadavia” some paratypes in the IFML, whereas the re- was presented (Mulieri et al., 2010). Here, maining paratypes were deposited at the as part of a continuous work on listing type American Museum of Natural History (AMNH), New York, USA. Most of these Recibido: 15/04/11 – Aceptado: 29/08/11 specimens were collected at some locations Acta zoológica lilloana 55 (2): 154–170, 2011 155 near the city of Tucumán by Dr. Martin L. Labels: “R. A. Tucuman / Queb. La Toma Aczel and Dr. Rodolfo Golbach. / 21-XII-950 / coll: R. Golbach” handwritten The type specimens of Sarcophagidae are on white paper, black frame; “Holotype Ÿ / species described by Lopes (1969). Weyrauch Emmesomyia / argentina / Snyder” hand- was a German malacologist who worked for written on red paper; “Colección / Inst. – 30 years at the Universidad Nacional Mayor Fund. M. Lillo / (4000) – S. M. Tucuman / de San Marcos, Perú. Later he moved to Ar- Tucuman – Argentina” printed on white pa- gentina (1962-1970), and worked in the per. malacological collection from the IFML. PARATYPE female [TDIP004]. Weyrauch had collected several specimens of Labels: “R. A. Tucuman / Queb. La Toma Sarcophagidae while living in Perú, material / 21-XII-950 / coll: R. Golbach” handwritten which he then sent to Dr. Hugo Souza Lopes on white paper, black frame; “Allotype ÿ / from Brazil. Emmesomyia / argentina / Snyder” hand- written on red paper. MATERIALS & METHODS Notes: The holotype is damaged. Missing parts: head, mid and hind right legs. The The species are listed within their corre- paratype is in good condition. sponding genus, by family, in alphabetical order, following their original combination. Emmesomyia spadibasis Snyder, For each specimen, the following informa- 1957: 442. tion was provided: species name, type cate- gories, sex and number of specimens are in- HOLOTYPE male [TDIP005]. dicated followed by collection data (country, Argentina: Tucumán, Quebrada La Toma. province or state, locality and collector). The R. Golbach leg. code number of the collection of IFML (e.g. Labels: “R. A. Tucuman / Queb. La Toma “TDIP003”), is given in brackets. Labels with / 21-XII-950 / coll: R. Golbach” handwritten the code number (T = type, DI = diptera, P on white paper, black frame; “Holotype Ÿ / = project) are present in the holotypes and Emmesomyia / spadibasis / Snyder” hand- some paratypes, the rest of the paratypes written on red paper; “Colección / Inst. – have no such label. Fund. M. Lillo / (4000) – S.M. Tucuman / Labels are cited verbatim, lines separated Tucuman – Argentina” printed on white pa- by a slash, different labels by semicolon and per; “FML” printed on white paper. comments are given in brackets. The infor- PARATYPE female [TDIP006]. mation on the specimens has been taken Labels: “R. A. Tucuman / Queb. La Toma from labels and compared with the original / 21-XII-950 / coll: R. Golbach” handwritten descriptions. All the specimens are pinned. on white paper, black frame; “Allotype ÿ / Notes are included to describe the conditions Emmesomyia / spadibasis / Snyder” hand- of the specimens, the valid names of each written on red paper; “Colección / Inst. – taxon, and bibliography referred to synony- Fund. M. Lillo / (4000) – S.M. Tucuman / my or new combination. Tucuman – Argentina” printed on white pa- per; “FML” printed on white paper. RESULTS PARATYPE male [TDIP007]. Argentina: Tucumán, Quebrada La Toma. ANTHOMYIIDAE R. Golbach leg. Emmesomyia argentina Snyder, Labels: “R. A. Tucuman / Queb. La Toma 1957: 443. / 21-XII-950 / coll: R. Golbach” handwritten on white paper, black frame; “Emmesomyia HOLOTYPE male [TDIP003]. / spadibasis / Snyder [handwritten] / Argentina: Tucumán, Quebrada La Toma. Paratype [printed]” on white paper; “Parati- R. Golbach leg. po” printed on white paper, black frame; “An- 156 L. D. Patitucci et al.: The type specimens of Anthomyiidae, Muscidae and Sarcophagidae thomyiidae / Emmesomyia / spadibasis” Argentina: Tucumán, Burruyacú, Villa printed on white paper, black frame. Padre Monti. R. Golbach leg. PARATYPES 1 male, 2 females [TDIP008 - Labels: “V. Padre Monti (R.A. / Tucuman TDIP010]. – Burruyacu) / 17.I – 7.II.1948 / coll. R. Labels: “R. A. Tucuman / Queb. La Toma Golbach” printed on white paper; “Allotype / / 21-XII-950 / coll: R. Golbach” handwritten Pegomya / trifumosa / Snyder” handwritten on white paper, black frame; “Emmesomyia on red paper; “Muscidae” printed on white / spadibasis / Snyder [handwritten] / paper. Paratype [printed]” on white paper; “Parati- Notes: Head missing in holotype. Snyder po” printed on white paper, black frame. (1957) described the holotype as teneral. PARATYPES 2 males. The paratype is in good condition. Labels: “R. A. Tucuman / Queb. La Toma / 21-XII-950 / coll: R. Golbach” handwritten MUSCIDAE on white paper, black frame; “Emmesomyia Angolia albibarba Snyder, 1957: 446. / spadibasis / Snyder [handwritten] / Paratype [printed]” on white paper; “Parati- HOLOTYPE male [TDIP134]. po” printed on white paper, black frame; Argentina: Tucumán, Burruyacú, Villa “Colección / Inst. – Fund. M. Lillo / (4000) Padre Monti. R. Golbach leg. – S.M. Tucuman / Tucuman – Argentina” Labels: “V. Padre Monti (R.A. / Tucuman printed on white paper. – Burruyacu) / 17.I – 7.II.1948 / coll. R. Notes: The holotype and paratypes are in Golbach” printed on white paper; “Holotype good condition. Ÿ / Angolia / albibarbra [sic] / Snyder” handwritten on red paper; “Colección / Inst. Pegomya neopalposa Snyder, – Fund. M. Lillo / (4000) – S.M. Tucuman / 1957: 444. Tucuman – Argentina” printed on white pa- per. HOLOTYPE male [TDIP011]. Notes: Head missing in holotype. The ter- Argentina: Tucumán, Quebrada La Toma. minalia is in a plastic vial pinned with holo- R. Golbach leg. type. Synonymized with Stomopogon albise- Labels: “R. A. Tucuman / Queb. La Toma ta (Stein) by Couri (2002). / 21-XII-950 / coll: R. Golbach” handwritten on white paper, black frame; “Holotype Ÿ / Angolia argentina Snyder, Pegomya / neopalposa / Snyder” handwrit- 1957: 447. ten on red paper; “Mucidae[sic!]” printed on white paper. HOLOTYPE male [TDIP135]. Notes: The holotype is in good condition. Argentina: Neuquén, San Martín de los Andes. Pegomya trifumosa Snyder, Labels: “San Martín