FAMILY Bythitidae Gill, 1863 - viviparous brotulas [=Brosmophycinae, Lucifugae, Pteridiidae, Protulina, Hephthocarinae] GENUS Acarobythites Machida, 2000 - viviparous brotulas Species Acarobythites larsonae Machida, 2000 - Larson's cusk GENUS Anacanthobythites Anderson, 2008 - viviparous brotulas Species Acanthobythites platycephalus Anderson, 2008 - Lucky Bay brotula Species Acanthobythites tasmaniensis Anderson, 2008 - Tasmanian brotula GENUS Aphyonus Gunther, 1878 - brotulas Species Aphyonus gelatinosus Gunther, 1878 - brotulas [=mollis] GENUS Barathronus Goode & Bean, 1886 - brotulas [=Alexeterion] Species Barathronus affinis Brauer, 1906 - Chagos brotula Species Barathronus bicolor Goode and Bean, 1886 - Guadeloupe Island brotula Species Barathronus bruuni Nielsen, 1969 - Bruun's brotula Species Barathronus diaphanus Brauer, 1906 - Valdivia brotula Species Barathronus linsi Nielsen et al., 2015 - Potiguar brotula Species Barathronus maculatus Shcherbachev, 1976 - Mozambique brotula Species Barathronus multidens Nielsen, 1984 - multidens brotula Species Barathronus pacificus Nielsen & Eagle, 1974 - Pacific brotula Species Barathronus parfaiti (Vaillant, 1888) - Parfait's brotula Species Barathronus unicolor Nielsen, 1984 - unicolor brotula GENUS Bellottia Giglioli, 1883 - viviparous brotulas [=Xenobythites] Species Bellottia apoda Giglioli, 1883 - apoda brotula Species Bellottia armiger (Smith & Radcliffe, in Radcliffe, 1913) - Macajalar Bay brotula Species Bellottia cryptica Nielson et al., 2009 - cryptic brotula Species Bellottia galatheae Nielsen & Moller, 2008 - Solomon brotula Species Bellottia robusta Nielsen et al., 2009 - robust brotula GENUS Bidenichthys Barnard, 1934 - viviparous brotulas [=Fiordichthys] Species Bidenichthys beeblebroxi Paulin, 1995 - Hinemoa brotula Species Bidenichthys capensis Barnard, 1934 - freetail brotula Species Bidenichthys consobrinus (Hutton, 1876) - grey brotula Species Bidenichthys paxtoni (Nielsen & Cohen, 1986) - Paxton's brotula Species Bidenichthys slartibartfasti (Paulin, 1995) - Fiordland brotula GENUS Brosmodorsalis Paulin & Roberts, 1989 - viviparous brotulas Species Brosmodorsalis persicinus Paulin & Roberts, 1989 - pink brotula GENUS Brosmophyciops Schultz, in Schultz et al., 1960 - viviparous brotulas Species Brosmophyciops pautzkei Schultz, in Schultz et al., 1960 - slimy cusk eel GENUS Brosmophycis Gill, 1861 - viviparous brotulas [=Halias] Species Brosmophycis marginata (Ayres, 1854) - red brotula GENUS Bythites Reinhardt, 1835 - viviparous brotulas Species Bythites fuscus Reinhardt, 1837 - Arctic brotula Species Bythites gerdae Nielsen & Cohen, 1973 - Gerda brotula Species Bythites islandicus Nielsen & Cohen, 1973 - Iceland brotula GENUS Calamopteryx Bohlke & Cohen, 1966 - viviparous brotulas Species Calamopteryx goslinei Bohlke & Cohen, 1966 - longarm brotula Species Calamopteryx jeb Cohen, 1973 - Jeb brotula Species Calamopteryx robinsorum Cohen, 1973 - teacher brotula GENUS Cataetyx Gunther, 1887 - viviparous brotulas [=Abythites, Oculospinus] Species Cataetyx alleni (Byrne, 1906) - Allen's brotula [=brevis] Species Cataetyx bruuni (Nielsen & Nybelin, 1963) - Brunn's brotula Species Cataetyx chthamalorhynchus Cohen, 1981 - chthamalorhynchus brotula Species Cataetyx hawaiiensis Gosline, 1954 - Gosline's Hawaiian brotula Species Cataetyx laticeps Koefoed, 1927 - Koefoed's brotula [=memorabilis] Species Cataetyx lepidogenys (Smith & Radcliffe, in Radcliffe, 1913) - Masbate brotula [=matsubarai] Species Cataetyx messieri (Gunther, 1878) - hairlip brotula Species Cataetyx nielseni Balushkin & Prokofiev, 2005 - Nielsen's catetyx Species Cataetyx niki Cohen, 1981 - brown brotula Species Cataetyx platyrhynchus Machida, in Okamura & Kitajima, 1984 - Okinawa cataetyx Species Cataetyx rubrirostris Gilbert, 1890 - rubynose brotula Species Cataetyx simus Garman, 1899 - Panama Bay brotula GENUS Diplacanthopoma Gunther, 1887 - viviparous brotulas [=Myxocephalus, Sarcocara] Species Diplacanthopoma brachysoma Gunther, 1887 - Pernambuco brotula Species Diplacanthopoma brunnea Smith & Radcliffe, in Radcliffe, 1913 - Palawan brotula Species Diplacanthopoma japonicus (Steindachner & Doderlein, 1887) - Japanese diplacanthopoma Species Diplacanthopoma jordani Garman, 1899 - Jordan's Eastern Pacific brotula Species Diplacanthopoma kreffti Cohen & Nielsen, 2002 - deepbody cusk Species Diplacanthopoma nigripinnis Gilchrist & von Bonde, 1924 - blackfin brotula Species Diplacanthopoma raniceps Alcock, 1898 - Andaman diplacanthopoma Species Diplacanthopoma riversandersoni Alcock, 1895 - Arabian diplacanthopoma GENUS Ematops Schwarzhans & Nielsen, 2011 - viviparous brotulas Species Ematops randalli (Cohen & Wourms, 1976) - Randall's ematops GENUS Grammonus Gill, in Goode & Bean, 1896 - viviparous brotulas [=Bathystorreus, Eutyx, Gadopsis, Grammonoides, Leptopodus, Oligopodus, Oligopus, Verater J 343, Verater J 265] Species Grammonus ater (Risso, 1810) - Mediterranean grammonus [=niger, nigrita] Species Grammonus claudei (Torre y Huerta, 1930) - reef-cave brotula [=mowbrayi] Species Grammonus diagrammus (Heller & Snodgrass, 1903) - purple brotula Species Grammonus longhursti (Cohen, 1964) - Longhurst's brotula Species Grammonus minutus Nielsen & Prokofiev, 2010 - Vietnamese grammonus Species Grammonus nagaredai Randall & Hughes, 2008 - Nagareda's brotula Species Grammonus opisthodon Smith, 1934 - bighead brotula Species Grammonus robustus Smith & Radcliffe, in Radcliffe, 1913 - robust grammonus Species Grammonus thielei Nielsen & Cohen, 2004 - Tomia brotula Species Grammonus waikiki (Cohen, 1964) - Waikiki brotula [=nigra] Species Grammonus yunokawai Nielsen, 2007 - Yunakawa's brotula GENUS Hastatobythites Machida, 1997 - viviparous brotulas Species Hastatobythites arafurensis Machida, 1997 - spinyhead cusk GENUS Hephthocara Alcock, 1892 - viviparous brotulas Species Hephthocara crassiceps Smith & Radcliffe, in Radcliffe, 1913 - Buton brotula Species Hephthocara simum Alcock, 1892 - Manaar brotula GENUS Lucifuga Poey, 1858 - viviparous brotulas [=Stygicola] Species Lucifuga dentata Poey, 1858 - toothed Cuban cusk eel Species Lucifuga inopinata Cohen & McCosker, 1998 - Galapagos inopinata brotula Species Lucifuga lucayana Moller et al., 2006 - Lucaya cavebrotula Species Lucifuga simile Nalbant, 1981 - Nalbant's lucifuga Species Lucifuga spelaeotes Cohen & Robins, 1970 - New Providence cusk eel Species Lucifuga subterranea Poey, 1858 - Cuban cusk eel Species Lucifuga teresinarum Diaz Perez, 1988 - Artemisa brotula GENUS Megacataetyx Prokofiev, 2005 - brotulas Species Megacataetyx niki (Cohen, 1981) - niki brotula GENUS Melodichthys Nielsen & Cohen, 1986 - viviparous brotulas Species Melodichthys hadrocephalus Nielsen & Cohen, 1986 - Celtic Sea brotula GENUS Meteoria Nielsen, 1969 - brotulas Species Meteoria erythrops Nielsen, 1969 - Azores brotula Species Meteoria longidorsalis Nielsen, 2016 - Hatteras brotula Species Meteoria pauciradiatus (Nielsen, 1997) - Loyalty brotula GENUS Microbrotula Gosline, 1953 - viviparous brotulas Species Microbrotula andersoni Schwarzhans & Nielsen, 2011 - Christmas Island brotula Species Microbrotula bentleyi Anderson, 2005 - manyray cusk [=polyactis] Species Microbrotula geraldalleni Schwarzhans & Nielsen, 2012 - Cenderawasih brotula Species Microbrotula greenfieldi Anderson, 2007 - Greenfield's brotula Species Microbrotula hamata Schwarzhans & Nielsen, 2011 - New Caledonia microbrotula Species Microbrotula punicea Anderson, 2007 - Kermadec microbrotula Species Microbrotula queenslandica Anderson, 2005 - Queensland cusk Species Microbrotula rubra Gosline, 1953 - Kaneohe brotula GENUS Nybelinella Nielsen, 1972 - brotulas [=Nybelinia] Species Nybelinella brevinalis Nielsen, 2017 - Madeira brotula Species Nybelinella brevidorsalis Shcherbachev, 1976 - short-tailed brotula Species Nybelinella erikssoni (Nybelin, 1957) - Eriksson's brotula GENUS Parabrotula Zugmayer, 1911 - brotulas Species Parabrotula plagiophthalma Zugmayer, 1911 - Azores parabrotula [=dentiens] Species Parabrotula tanseimaru Miya & Nielsen, 1991 - Sagami brotula GENUS Paraphyonus Nielsen, 2015 - brotulas Species Paraphyonus bolini (Nielsen, 1974) - Bolin's brotula Species Paraphyonus brevidorsalis (Nielsen, 1969) - Durban brotula Species Paraphyonus iselini Nielsen, 2015 - Sargasso brotula Species Paraphyonus merretti Nielsen, 2015 - Moroccan brotula Species Paraphyonus rassi (Nielsen, 1975) - Rass' brotula Species Paraphyonus solomonensis (Nielsen & Moller, 2008) - Solomon Sea brotula GENUS Parasaccogaster Nielsen et al., 2004 - viviparous brotulas Species Parasaccogaster melanomycter (Cohen, 1981) - Gairaca Bay brotula Species Parasaccogaster normae (Cohen & Nielsen, 1972) - Norma's brotula Species Parasaccogaster rhamphidognatha (Cohen, 1987) - delta brotula GENUS Parasciadonus Nielsen, 1984 - brotulas Species Parasciadonus brevibrachium Nielsen, 1984 - brevibrachium brotula GENUS Petrotyx Heller & Snodgrass, 1903 - brotulas Species Petrotyx hopkinsi Heller & Snodgrass, 1903 - Barrington brotula Species Petrotyx sanguineus (Meek & Hildebrand, 1928) - Toro brotula GENUS Pseudonus Garman, 1899 - viviparous brotulas [=Leucochlamys] Species Pseudonus acutus Garman, 1899 - Garman's pseudonus Species Pseudonus squamiceps (Lloyd, 1907) - squamiceps brotula [=platycephalus] GENUS Saccogaster Alcock, 1889 - viviparous brotulas [=Barbuliceps] Species Saccogaster brayae Nielsen et al., 2012 - Bray brotula Species Saccogaster hawaii Cohen & Nielsen, 1972 - Maui brotula Species Saccogaster horrida Nielsen et al., 2012 - horrida brotula Species Saccogaster maculata Alcock, 1889 - Alcock's saccogaster Species Saccogaster nikoliviae Nielsen et al., 2012 - Nikolivia's brotula Species Saccogaster parva Cohen & Nielsen, 1972 - parva brotula Species Saccogaster staigeri Cohen & Nielsen, 1972 - Staiger's brotula Species Saccogaster tuberculata (Chan, 1966) - bagbelly cusk GENUS Sciadonus Garman, 1899 - brotulas Species Sciadonus cryptophthalmus (Zugmayer, 1911) - Spanish brotula Species Sciadonus galatheae (Nielsen, 1969) - Kermadec brotula Species Sciadonus jonassoni (Nybelin, 1957) - Jonasson's brotula Species Sciadonus pedicellaris Garman, 1899 - Cocos brotula [=kullenbergi] GENUS Stygnobrotula Bohlke, 1957 - viviparous brotulas Species Stygnobrotula latebricola Bohlke, 1957 - black widow [=tumidirostris] GENUS Thermichthys Nielsen & Cohen, 2005 - viviparous brotulas [=Gerhardia] Species Thermichthys hollisi (Cohen et al., 1990) - Hollis' brotulas GENUS Timorichthys Nielsen & Schwarzhans, 2011 - viviparous brotulas Species Timorichthys angustus Neilsen et al., 2013 - Japanese timorichthys Species Timorichthys disjunctus Nielsen & Schwarzhans, 2011 - Timor brotula GENUS Tuamotuichthys Moller et al., 2004 - viviparous brotulas Species Tuamotuichthys bispinosus Moller et al., 2004 - twospined slopebrotula Species Tuamotuichthys marshallensis Nielsen et al., 2006 - Marshall slopebrotula Species Tuamotuichthys schwarzhansi Nielsen & Moller, 2008 - Schwarzhan's brotula