植物研究雑誌 J. J. Jpn. Bo t. 80: 80: 323-333 (2005)

Taxonomic Notes on Vascular in Japan and Its Its Adjacent Regions (1). -New Combinations and New Names of Japanese Plants-

K 吋i YONEKURA

Botanical Botanical Gardens ，Tohoku University ，12-2 ，Kawauchi ，Sendai 980 ・0862 JAPAN E-mail: E-mail: yonekura@ma 幻ai l.tぬ凶 a幻i泊nS.tωohoku.a 配吋Cι .j

(Received (Received on May 凶16，2005)

Eighty-one Eighty-one new combinations ， five new names and one new form for mostly

J apanese seed plants ， which are needed during prep 紅 ation of database of Japanese vascu- lar lar plants 紅 e proposed. Some taxonomi and nomenclatural comments are made on Geranium onoei (Geraniaceae) ， pubescens () ， Kalimeris yomena (Asteraceae) and Ixeridium dentatum (Asteraceae).

Key words: Database ，Japan ，new combinations ，new names ，vascular plants.

During prep 紅 ation of a database of (3) Picea alcocquiana (Veitch ex Lind l.)

J apanese and scientific n倒 nes of indigenous ， C 訂 ri とre f. chlorocarpa (Hayashi) Yone k.， naturalized naturalized and cultivated vascular plants in comb. nov. - P. bicolor (Maxim.) Mayr f. Japan (BG Plants- YLis t: Japanese chlorocarpa Hayashi in Bull. Gov t. For. name index; http:/ /b ean.bio.chiba-u.jp/ Exp. Sta t. (77): 25 ， photo 12 (1 955). bgplants/) bgplants/) numerous new names and new Nom. Jap.: Midori-iramomi. combinations 訂 e needed for the publication. Here 1 propose 81 new combinations ， five (4) Picea torano (Siebold ex K. Koch) new names and one new form. Koehne f. rubriflora (Y oshio Kobay.) Yone k.， comb. nov. - P. polita (Siebold & Gymnospermae Zucc.) Carri とre f. rubriflora Y oshio Kobay. Taxaceae in J. Jpn. Bo t. 40: 191 (1 965). (1) (1) Torreya nucifera Siebold & Zucc. f. Nom. Jap.: Benibana-harimomi. sphaerica (K.i mura) Yone k.， sta t. nov. - T. nucifera nucifera v訂 . sphaerica K.i mura in Oinuma in Angiospermae - Dicotyledoneae Jpn. Jpn. J. Bo t. 13: 519 ， in adno t.， f. 1，2 & 7 Crassulaceae (1 949). (5) Hylotelephium verticillatum (L.) H. Nom. Jap.: M 訂 umi-gaya. Ohba f. viviparum (N. Yonez.) Yone k.， comb. nov. - Sedum verticillatum L. f. Pinaceae Pinaceae viviparum N. Y onez. in J. Phytogeog r. (2) (2) Larix kaempferi Carri とre f. pendula Taxon. 34: 69 ， photos 1-2 (1 986). (Sugim.) (Sugim.) Yone k.， comb. nov. - L. leptolepis Nom. Jap.: Komochi-mitsuba-benkeiso. (Siebold (Siebold & Zucc.) Gordon f. pendula Sugim. in in J. Geobo t. (Kanazawa) 6: 27 (1 957). Rosaceae Nom. Jap.: Shidare-karamatsu. (6) Aria alnifolia (Siebold & Zucc.) Decne.

-323- 324 植物研究雑誌第80 巻第6号 平成17 年12 月 f. f. lobulata (Koidz.) Y onek. ，comb. nov. - Okuyama in J. Jpn. Bo t. 30: 44 (1955). Micromeles alnifolia (Siebold & Zucc.) P. sargentii f. nagaokae Tatew. ex Koji Koehne var. s. lobulat αKoidz.， in J. Coll. It o in J. Geobo t. (Kanazawa) 13: 2 (1964) ， Sci. Sci. Imp. Univ. Tokyo 34(2): 69 (1913) - syn. nov. Sorbus Sorbus albifolia (Siebold & Zucc.) K. Koch var. var. lobulata (Koidz.) Rehder in Sarg. ， P l. (1 2) Cerasus xtakinoensis (Kawas.)

Wilson. Wilson. 2: 275 (1 915). Yone k.， comb. nov. - Prunus t α， kinoensis Nom. Jap.: Fugire-azukinashi. Kawas. in J. Jpn. Bo t. 45: 137 ，figs. 1-2 (1 970). (7) (7) Aria japonica Decne. f. chlorocarpa Nom. Jap.: Takino-zakura. (Rehder) (Rehder) Yone k.， comb. nov. - Sorbus j， α- ponica ponica (Decne.) Hed l. var. chlorocarpa (13) Filipendula glaberrima N akai f. alba Rehder in S 紅 g. ， Pl. Wilson. 2: 276 (1915). (Nak 泊) Yonek. ，comb. nov. - F. multijuga Nom. Jap.: Ki mi-no-urajironoki. Maxim. var. alba Nakai in Reper t. Spec. f. f. denudata (Nakai) Yone k.， comb. Nov. Regni Veg. 13: 274 (1 914) - F. nov. nov. - Sorbus japonica var. denudata Nakai yezoensis H. Hara f. alba (Nakai) T. Shimizu in in Bull. Natn. Sci. Mus. Tokyo (33): 11 in J. Fac. Tex t. Sci. Tech. Shinshu Univ. no. (1 953) - S. japonica f. denudata (Nakai) 26 ， se r. A. Bio l. (1 0): 9 (1 961). Hiyama in J. Jpn. Bo t. 35: 309 (1 960). Nom. Jap.: Shirobana-ezo-no-shimo- Nom. Jap.: Komono-urajironok i. tsukeso.

(8) (8) Cerasus chichibuensis (H. Kubota & (1 4) Rosa luciae Franch. & Rochebr. ex Moriya) Moriya) H. Ohba nothovar. aizuensis Crep. f. ohwii (Oishi) Yone k.， comb. nov. - (Kawas.) (Kawas.) Yone k.， comb. nov. - Prunus R. wichuraiana Crep. f. ohwii Oishi in J. chichibuensis chichibuensis H. Kubota & Moriya nothova r. Phytogeogr. Taxon. 28: 62 (1 980). aizuensis aizuensis Kawas. in J. Phytogeogr. Taxon. Nom. Jap.: Togenashi-teriha-no-ibara. 49: 49: 3 ，fig. 1 & 2 (2001). f. plena (Honda) Yonek. ，comb. nov. - R. Nom. Jap.: Aizu-shidare. wichuraiana Crep. var. plen αHonda in Bo t. Mag. Tokyo 50: 435 (1936). (9) (9) Cerasus tamaclivorum (T. Oohara ， Nom. Jap.: Yae-teriha-no-ibara. Seriz. Seriz. & Wakab.) Yone k.， comb. nov. -

Prunus tama ・clivorum T. Oohara ， Seriz. & (1 5) Rosa mu It iflora Thunb. v紅. mu It iflora Wakab. in J. Jpn. Bo t. 79: 343 ，figs. 1，3 f. rosipetala (Honda) Yonek. ，comb. nov. - (2004). (2004). Rosa poly α ntha Siebold & Zucc. f. Nom. Jap.: Hoshi-zakura. rosipetala Honda in Bo t. Mag. Tokyo 52: 518 (1 938). (10) (10) Cerasus moniwana (Kawas.) Yonek. ， Nom. Jap.: Usuaka-no-ibara. comb. nov. - Prunus moniwana Kawas. in J. Jpn. Jpn. Bo t. 38: 94 ，figs. 1-2 (1 963). (1 6) Rosa onoei Makino var. oligantha Nom. Jap.: Moniwa-zakura. (Franch. & Sav.) H. Ohba f. rosea (Hayashi) Yone k.， comb. nov. - R. luciae Franch. & (1 1) Cerasus sargentii (Rehder) H. Ohba Rochebr. ex Crep. f. rosea Hayashi in J. V ぽ. sargentii f. pendula (Honda) Yonek. ， Geobo t. (Kanazawa) 25(3): 55 (1 977). comb. nov. - Prunus sα rgentii Rehder var. Nom. Jap.: Akebono-o-fuji-ibara. pendula pendula Honda in Bo t. Mag. Tokyo 54: 2 (1940) (1940) - P. sargentii f. pendula (Honda) Fabaceae (Leguminosae) December 2005 Journal of Japanese Botany Vo l. 80 No. 6 325

(17) (17) Wisteria japonica Siebold & Zucc. f. The commonest form of Geranium onoei alborosea alborosea (Sakata) Yone k.， comb. nov. - in J apan has pale purple flowers and leaves Millettiajaponica Millettiajaponica (Siebold & Zucc.) A. Gray hairy throughou t. The name G. onoei has f. f. alborosea Sakata in J. Jpn. Bo t. 33: 30 been applied to a form growing in alpine re- (1 958). gions of Central Japan ， with deep purple Nom. Jap.: Ak ebono-natsu-fuji. flowers and usually less hairy leaves (Nakai 1912). 1912). Nakai (1935) has previously pointed Geraniaceae Geraniaceae out that the type specimen of G. onoei repre- (1 8) New combinations and a new form of sented the commonest fOfffi. 1 recognize the Geranium onoei alpine form as a new form exclusively based Geranium reinii and G. onoei both de- on the difference in flower colour because scribed scribed from Japan by Franchet and Savatier leaf hairiness is variable as pointed out by (1 878) have been treated as infraspecific taxa Hara (1 948). of of G. eriostemon Fisch. ex DC. (1 824) de- Geranium onoei Franch. & Sav. in Enum. scribed scribed from eastem Siberia. The latter name P1. Jap. 2: 303 (1878) ，‘ Onaei'. is ， however ， a later homonym of G. var.onoel eriostemum eriostemum Poir. (1 811) (Yeo 1992). Distr.: Japan (Hokkaido and Honshu). Accordingly ， either G. reinii or G. onoei f.onoe i. should should be used for the name of the species. G. reinii Franch. & Sav. ，Enum. P1. Jap. As Nakai (1 935) synonymized G. reinii with 2: 304 (1 878). G. G. onoei ， the latter has priority. Geranium Nom. Jap.: Gunnai-furo ， Ibuki-gunnai- japonicum Franch. & Sav. (1878) which was furo. misapplied misapplied to G. krameri Franch. & Sav. at f. alpinum Yonek. ，f. nov. past past is also identical with G. onoei (Hara A typo floris purpureo- violaceis differt. 1955) 1955) although G. onoei is not recorded in Type: JAPAN: Honshu ，Nagano Pref. the the type locality of G. japonicum (Hakone in (Shinano) ， M t. Yatsugatake (M. Okamoto s. Kanagawa Pref.) in cu 町 ent floras. n. (TNS exsicc. no. 1565) ，23 Ju 1. 1957 ，TUS Yeo (1 992) regarded continental 22794 -holotype; TNS -isotype). Geranium eriostemon (nom. illeg.) as dis- Misapplied names: tinct tinct from G. onoei and used the name G. G. eriostemon var. y. onoei seusu Nakai in platyanthum platyanthum Duthie (1 906) for the former. Bo t. Mag. Tokyo 26: 255 (1 912) ，non G. However ，a wide range of mo 中hological onoei Franch. & Sav. variation variation in N orth Korea (N akai 1912 ，Hara G. eriostemon var. reinii f. onoei sensu 1948) 1948) makes it difficult to distinguish G. Ohwi ， F 1. Japan: 702 (1953); Kitam. & pla 砂anthum from G. onoei at the specific Murata ，Co 1. 11 1. Herb. P1. Jap. 2: 89 (1 961); leve 1. Park and Ki m (1997) included G. Ak iyama in K. Iwats. & a1.， F1. Japan 2b: onoei onoei in G. eriostemon apparently unaware 288 ， in nota (2001). of of the illegitimacy of the latter. The quantity Nom. Jap.: Takane-gunnai-furo. of of materials seen by them seems insufficient f. albiflorum (N. Yonez.) Yonek. ，comb. for for the inclusion. 1 think both can be distin- nov. - G. eriostemon var. reinii (Franch. & guished guished from each other at varietal ran k. Sav.) Maxim. f. albiflorum N. Y onez. in J. Geranium eriostemon var. glabrescens Phytogeogr. Taxon. 29: 110 ， photo (1 98 1). Nakai ex H. Hara (1 948) ，which is here Nom. Jap.: Shirobana-gunnai-furo. transferred transferred to a variety of G. onoei ， seems to f. yezoense (H. Hara) Yone k.， comb. be be another distinct form restricted to the high nov. - G. eriostemon Fisch. ex DC. var. mountains mountains of North Korea. reinii (Franch. & Sav.) Maxim. f. yezoense 326 326 植物研究雑誌第80 巻第6号 平成17 年12 月

H. Hara in J. Jpn. Bo t. 22: 166 (1 948). Jpn. Bo t. 12: 876 (1936) ，non Nakai (1 919). Nom. Jap.: Ezo-gunnai-furo. Z. ailanthoides f. inermis (Hatus.) H. var. var. glabrescens (Nakai ex H. Hara) Ohba in K. Iwats. et al.， Fl. Japan 2c: 39 Yonek. ，comb. nov. - G. eriostemon Fisch. (1 999) ，non Z. ailanthoides v訂 inermis ex DC. var. glabrescens Nakai ex H. Hara in Rehder & E. H. Wilson (1919) ， nec T. B. J. J. Jpn. Bo t. 22: 171 (1 948). Lee (1 966).

Distr.: Distr.: N. Korea. Nom. Jap.: Togenashi 闇 karasu-zansho.

Polygalaceae Polygalaceae Violaceae (19) (19) Polygala reinii Franch. & Sav. f. (22) Viola inconspicua Blume subsp. stenophylla stenophylla Yone k.， nom. nov. nagasakiensis (W. Becker) J. C. Wang & T. Based on: P. reinii v訂 . angustifolia T. It o， C. Huang f. albescens (Taken.) Yonek. ， Encyc l. Jap. 2: 612 (1 909). comb. nov. - V. minor (Makino) Makino var. Synonyms: albescens Taken. in Bio l. Mag. Fukuoka 2: P. P. reinii f. angustifolia (T. It o) K. Iwats. 195 (1 938) - V. minor f. albescens (Taken.) & H. Ohba in K. Iwats. & al.， Fl. Japan 2c: F. Mae k. in Hara ，Enum. Sperm. Jap. 3: 209 54 ， in adno t. (1 999) ，nom. illeg. (1 954). P. P. reinii v訂 . angustifolia Makino in J. Nom. Jap.: Usuiro-hime-sumire. Jpn. Jpn. Bo t. 3: 18 (1 926) ，non T. It o (1909). f. pilosa (F. Mae k.) Yone k.， comb. P. P. reinii f. angustifoli α(Makino) Ohwi in nov. - V. minor (Makino) Makino f. pilosa Bull. Bull. Natn. Sci. Mus. Tokyo (33): 78 (1 953) ， F. Mae k. in H. Hara ，Enum. Sperm. Jap. 3: nom. illeg. 209 (1 954). Nom. Jap.: Nagaba-kakinohagusa. Nom. Jap.: Ke-hime-sumire.

Rutaceae Rutaceae Onagraceae (20) (20) Melicope grisea (Planch.) T. Hartley (23) Chamerion angustifolium (L.) Holub var. var. crassifolia (Nak 泊) Yone k.， comb. f. pleniflorum (Nakai) Yone k.， comb. nov. nov. Boninia glabra Planch. var. nov. - Epilobium angustifolium L. v紅. crassifolia crassifolia Nakai in Bo t. Mag. Tokyo 36: pleniflorum Nakai in Bo t. Mag. Tokyo 22: 124 (1 922) - B. grisea Planch. var. 75 (1 908). crassifolia crassifolia (Nakai) T. Yamaz. ex H. Ohba in Nom. Jap.: Yaezaki-yanagiran. K. K. Iwats. & a l.， Fl. Japan 2c: 34 (1 999). Nom. Jap.: Atsuba-shirotetsu. (24) Epilobium amurense Hausskn. subsp. cephalostigma (Hausskn.) C. J. Chen ，Hoch (21) (21) Zanthoxylum ailanthoides Siebold & & Raven f. leucanthum (Honda) Y one k.， Zucc. Zucc. v訂. ailanthoides f. espinosum comb. nov. - E. ceph α lostigma Hausskn. f. Yonek. ，nom. nov. leucanthum Honda in Bo t. Mag. Tokyo 54: 2 Based on.: Fagara ailanthoides (Siebold (1 940). & Zucc.) Eng l. v紅 inermis Nakai ， Rep. Nom. Jap.: Shirobana-iwa-akabana. Veg. Ooryongto: 21 (nom. nud.) & 68 ， in clavi clavi (1 919) ，cum diagn. Jap. Araliaceae Synonyms: (25) Oplopanax japonicus (Nakai) Nakai Z. Z. ailanthoides var. inermis (Nakai) T. B. va r. brevilobus (H. Hara) Y one k.， comb. Lee ，Woody Pl. Ko r.: 297 (1966) ，non nov. - o. horridus (Sm.) Miq. v紅. Rehder & E. H. Wilson (1 919). brevilobus H. Hara in J. Jpn. Bo t. 30: 70 F. F. ailanthoides v紅 . inermis Hatus. in 1. (1 955). December 2005 Journal of Japanese Botany Vo l. 80 No. 6 327

Nom. Jap.: Hiroha-haribuki. nov. - R. sinense (Lodd.) Sweet f. flavum Miyoshi ， Saishin Shokubutsugaku Kogi 3: t. (26) (26) Eleutherococcus spinosus (L. f.) S.Y. 32 (1 921). Hu f. espinosus (H. Hara) Y onek. ，comb. Nom. Jap.: Ki -renge-tsutsuji. nov. nov. - Acanthopanax spinosus (L. f.) Miq. f. f. glaucophyllum (Nakai) Yonek. ，comb. espinosus espinosus H. H ぽ a，Enum. Sperm. Jap. 3: 280 nov. - R. jα rponicum var. glaucophyllum (1 954). Nakai ， Trees & Shrubs Jap. rev. ed. 1: 82 A. A. spinosus var. inermis Makino ex F. (1 927) - R. japonicum f. glaucophyllum Mae k. in J. Jpn. Bo t. 14: 72 (1938) ， (Nakai) H. Hara ，Enum. Sperm. Jap. 1: 36 'inerme' ，non f. inerme Matsuda (1 912) - E. (1 949). spinosus spinosus f. inermis (Makino ex F. Maek.) H. Nom. Jap.: Urajiro-renge-tsutsuji. Ohashi in J. Jpn. Bo t. 62: 357 (1987). f. multifidum (Nakai) Yonek. ，comb. nov. Nom. Jap.: Togenashi-ukogi. - R. japonicum f. multifidum N akai ， Trees & Shrubs Shrubs Jap. rev. ed. 1: 82 (1 927). Apiaceae (Umbelliferae) Nom. Jap.: Renge-botan. (27) (27) Cryptotaenia canadensis DC. subsp. japonica (Hass k.) Hand.-Mazz. f. (30) Vaccinium microcarpum Turcz. ex atropurpurea (Makino) Yone k.， comb. nov. Rupr. f. kirigaminense (Honda & Tobita) - C. japonica va r. α tropurpurea Makino in J. Yone k.， comb. nov. - 0.刀 coccus micro- Jpn. Jpn. Bo t. 3(2): 8 (1926) 一仁 japonica f. cα rpus Turcz. ex Rupr. var. kirigaminensis atropurpure α(Makino) Ohwi [F 1. J apan: 845 Honda & Tobita in Honda in Bo t. Mag. (1 953) ，comb. nud.] ex H. Hara ，Enum. Tokyo 54: 223 (1940). Sperm. Jap. 3: 309 (1954). Nom. Jap.: Shirobana-hime-tsurukoke- Nom. Jap.: Murasaki-mitsuba. momo. f. f. warabiana (Makino) Y one k.， comb. nov. nov. - C. japonica v紅 . warabiana Makino in (31) Vaccinium oxycoccos L. f. J. J. Jpn. Bo t. 8: 46 (1933) - C. japonica f. leucanthum (T. Shimizu) Yone k.， comb. warabiana (Makino) H. Hara ，Enum. Sperm. nov. - 0.刀 coccus quadripet α， lus Gilib. f. Jap. Jap. 3: 309 (1 954). leucanthus T. Shimizu ，New Alpine F1. Jap. Nom. Jap.: Haniya-mitsuba. Co 1. 1: 312 (1 982). Nom. Jap.: Shirobana-tsurukokemomo. Ericaceae Ericaceae (28) (28) Elliottia paniculata (Siebold & Zucc.) Symplocaceae Hook. f. albiflora (Y. Ki mura) Yonek. ， (3 2) Symplocos sawafutagi Nagam. f. comb. nov. - Tripetaleia paniculata Siebold leucocarpa (Naka i) Yonek. ，comb. nov. - S. & Zucc. f. alb{ 月ora Y. Ki mura in 1. Jpn. Bo t. paniculata (Thunb.) D. Don v紅. s. 16: 16: 59 (1940). leucocarpa N akai in Bo t.民 1ag. Tokyo 32: Nom. Jap.: Shirobana-hotsutsuji. 227 (1 918). Nom. Jap.: Shiromi-no-sawafutagi. (29) (29) Rhododendron molle G. Don subsp. japonicum (A. Gray) K. Kr on f. canescens Oleaceae (Sugim.) (Sugim.) Yone k.， comb. nov. - R. japonicum (33) Fraxinus platypoda Oliv. f. nipponica (A. (A. Gray) Suringer f. canescens Sugim. in J. (Koidz.) Yone k.， comb. nov. - F. nipponica Geobo t. (Kanazawa) 22(4): 52 (1 975). Koidz. in Bo t. Mag. Tokyo 28: 286 Nom. Jap.: Bi 廿rδ do- イrenge- 帽却 (1914) - F. sphaethian αLingelsh. f. f. f. flavum (Miyoshi) Yonek. ， comb. nipponica (Koidz ，) Sa. Kurata ， 11 1. Impor t. 328 植物研究雑誌第80 巻第6号 平成17 年12 月

Forest Forest Trees Jap. 1: 196 (1 964). ca11ed plants C. japonica Choisy (1854) in Nom. Jap.: Yama-shioji. J apan. Hara (1 957) considered C. pubescens as as specifica11y different from C. japonica Gentianaceae Gentianaceae and reduced the former to a form of C. (34) (34) Gentiana yakumontana Masam. f. dahurica (Herb.) Choisy (1 845). However viridis viridis (Sugim.) Yone k.， comb. nov. - G. the application of the name C. dahurica is zollingeri zollingeri Fawc. var. yakumontana (Masam.) uncertain (Fang and Brummit 1995). Sugim. Sugim. f. viridis Sugim. ，Keys Herb. Pl. Jap. Although typical C. pubescens Lind l. (a 1: 1: 427 & 737 (1 965). cultivar ‘Flore Pleno') which came from Nom. Jap.: Midori-yakushima-koke-rindo. China differs from J apanese C. japonica in having having hairier stems and truncate leaf-bases (35) (35) Tripterospermum trinervium as we11 as the double-form flowers (Hara (Thunb.) (Thunb.) H. Ohashi & H. Nakai va r. 1957) ， these seem conspecific if the entire trinervium trinervium f. albiflorum (Honda) Yone k.， range of variation in China is considered. comb. nov. - T. japonicum (Siebold & Accordingly ，1 仕eat wild C. japonica with Zucc.) Zucc.) Maxim. f. αlbiflorum Honda in J. Jpn. normal pink flowers and C. japonica with Bo t. 30: 170 (1 955). normal white flowers as forms of C. Nom. Jap.: Shiroban 仕 tsuru-rindo. pubescens. V 紅. trinervium f. leucocarpum (Honda) Calystegia sepium va r. japonica f. major Yone k.， comb. nov. - T. japonicum f. Makino (1 908) is the earliest name published leucocarpum leucocarpum Honda in J. Jpn. Bo t. 30: 170 at forma rank applicable to typical C. japon- (1955). (1955). ica. Although Hara (1949) recognized it as a Nom. Jap.: Shiromi-no-tsuru-rindo. distinct form of C. japonica ，it seems indis- V 征. involubile (N. Yonez.) H. Ohashi & tinguishable from the latter as leaf shape and H. H. Nakai f. album (T. Shimizu) Yonek. ， size of this species vary continuously. The comb. nov. - T. japonicum var. involubile co 町 ect name of C. japonica becomes as fol- (N. (N. Yonez.) J. Murata f. album T. Shimizu in lows: 1. 1. Phytogeogr. Taxon. 46: 115 (1 998). Calystegia pubescens Lind l. f. major T. T. japonicum var. involubile f. albiflorum (Makino) Yone k.， comb. nov. - C. sepium T. T. Shimizu ， F l. Nagano Pre f.: 1509 (1 997) ， (L.) R. B r. var. japonica (Choisy) Makino f. non Honda (1 955). major Makino in Bo t. Mag. Tokyo 21: 162 Nom. Jap.: Shirobana-tengu-no-kozuchi. (1 907). Other Other synonyms: Asclepiadaceae Asclepiadaceae C. japonica Choisy in Zo11. ，Sys t. Verz. (36) (36) Vincetoxicum sublanceolatum (Miq.) Ind. Ar ch. 2: 130 & 132 (1 854). Maxim. var. kinokuniense (T. Yamaz.) 仁 sepium v紅 . japonica var.α ngustifolia Y onek. ， comb. nov. - Cynanchum sub- Nakai in Bo t. Mag. Tokyo 23: 107 (1 909). lanceolatum lanceolatum (Miq.) Matsum. v紅. C. japonica f. vulgaris H. Hara ，Enum. kinokuniense kinokuniense T. Yamaz. in K. Iwats. & al.， Sperm. Jap. 1: 160 (1949). l. Fl. Japan 3a: 174 (1993). Nom. Jap.: Hirugao. Nom. Jap.: Kinokuni-kamomezuru. f. albiflora (Makino) Yone k.， comb. nov. - C. japonica var.α lb 抑ora Makino in J. Jpn. Convolvulaceae Convolvulaceae Bo t. 3: 1 (1923) - C. japonica f. αlbiflora (37) (37) New combinations in Calystegia (Makino) H. Hara ，Enum. Sperm. Jap. 1: 160 Fang and Brummit (1 995) applied (1949) - C. sepium v紅 . japonica f. albiflora Calystegia Calystegia pubescens Lind l. (1846) to the (Makino) T. Yamaz. in K. Iwats. & al.， Fl. December 2005 Joumal of Japanese Botany Vo l. 80 No. 6 329

Japan Japan 3a: 199 (1 993). Sav.) Honda f. kimurae (Makino) Yone k.， Nom. Jap.: Shirobana-hirugao. comb. nov. - P. kimurae Makino in J. Jpn. Bo t. 3: 37 (1 926) ，‘ Kimurai' - P. sα vatieri Boraginaceae Boraginaceae Makino f. kimurae (Makino) Ohwi ， F 1. Jap.: (38) (38) Cynoglossum furcatum Wall. va r. 1026 (1 953) ，comb. nud. ， ed. Eng 1.: 789 villosulum villosulum (Nakai) Riedl f. albi f1 0rum (H. (1 965) ，comb. nud.; Kitam. & Murata ，Co 1. Hara) Hara) Yone k.， comb. nov. - C. zeylanicum 11 1. Herb. P1. Jap. 1: 156 (1 957) ，comb. nud. (Vahl (Vahl ex Homem.) Thunb. ex Lehm. f. Nom. Jap.: Takao-hozuki. albiflorum albiflorum H. H 訂 a，Enum. Sperm. Jap. 1: 173 173 (1 949). Scrophul 紅 iaceae Nom. Jap.: Shirobana-o-ruriso. (43) Lindernia micrantha D. Don f. leucantha leucantha (Hiyama) Yone k.， comb. nov. - (39) (39) Cynoglossum lanceolatum Forss k. va r. Vandellia angust 約li αBenth. f. leucantha formosanum (Nakai) H. Hara f. albi f1 0rum Hiyama in J. Jpn. Bo t. 32: 57 (1 957). (Masam.) (Masam.) Yonek. ，comb. nov. - C. formosa- Nom. Jap.: Shirobana-aze-togarash i. num Nakai f. albiflorum Masam. in J. Soc. Trop. Trop. Agric. Taiwan 2: 152 (1 930). (44) Mazus pumilus (Burm. f.) Steenis f. Nom. Jap.: Shirobana-shima-ruriso. senanensis (Asai) Yone k.， comb. nov. - Mazus japonicus (Thunb.) Kuntze f. (40) (40) Omphalodes krameri Franch. & Sav. senanensis Asai in J. Jpn. Bo t. 38: 168 var. var. laevisperma (Nakai) Ohwi f. leucantha (1 963). Yonek. ，nom. nov. Nom. Jap.: Shirobana-tokiwahaze. Based on: O. krameri var. laevisperma f. albiflora albiflora Satomi in J. Phytogeog r. Taxon. (45) Veronicastrum sibiricum (L.) Pennell

31: 31: 30 (1983) ，non O. krameri v紅 .α lbiflora subsp. yezoense (H. H 紅 a) T. Yamaz. f. H. Hara (1934). candidum Yone k.， nom. nov. Nom. Jap.: Shirobana-echigo-ruriso. V. sibiricum var. yezoense H. Hara f. albiflorum albiflorum Hideki Takah. in J. Phytogeogr. Lamiaceae (Labiatae) Taxon.52: 191 ，f. l (‘ 2004' pub 1. 2005) ，non (41) (41) Prunella vulgaris L. subsp. asiatica V. sibiricum var. japonicum f. alb{ 月orum (Nakai) (Nakai) H. Hara f. candida Yonek. ，nom. Akasawa (1 955) nec Hayashi (1 960) nec N. nov. nov. Y onez. (1 984). Based on: Brunella vulgaris L. var. Nom. Jap.: Shirobana-ezo-kugaiso. albiflora albiflora Koidz. in Bo t. Mag. Tokyo 29: 310 (1915). (1915). Acanthaceae Other Other synonyms: (46) Peristrophe japonica (Thunb.) P. P. vulgaris f.α lbiflora (Koidz.) Kudo in Breme k. f. leucantha (Hiyama) Y one k.， Mem. Fac. Sci. Taihoku Imp. Univ. 2: 308 comb. nov. - Dicliptera japonica (Thunb.) (1 929) ，non Nakai (1 921). Makino f. albiflora Hiyama in 1. Jpn. Bo t. P. P. vulgaris var. elongata Benth. f. 28: 94 (1 953). albiflora albiflora Nakai in Bo t. Mag. Tokyo 35: 192 Nom. Jap.: Shirobana-haguroso. (1 921) ，non var. albiflora Koidz. (1915). Nom. Jap.: Shirobana-utsubogusa. Campanulaceae (47) (47) Adenophora nikoensis Franch. & Sav. Solanaceae Solanaceae va r. teramotoi (Hurus. & T. Yamaz.) J. (42) (42) Physaliastrum japonicum (Franch. & Okazaki & T. Shimizu f. hispidula (T. 330 植物研究雑誌第80 巻第6号 平成17 年 12 月

Shimizu} Shimizu} Yone k.， comb. et sta t. nov. - A. (5 1) A new status for Kalimeris yomena teramotoi teramotoi Hurus. & T. Yamaz. v紅 . hispidula subsp.α ngustifolia T. T. Shimizu in 1. Fac. Tex t. Sci. Tech. J apanese plants hitherto referred to Shinshu Shinshu Univ. no. 36 ，ser. A (Bio l.) (1 2): 66 Kalimeris incisa DC. (Aster incisus Fisch.) (1 963) et in Acta Phytotax. Geobo t. 21: 28 in continental Asia are different from the (1 964). mainland plants in having elongate stoloni- Nom. Jap.: Ke-shiraiwa-shajin. ferous rhizomes (Gu and Hoch 1997). Gu and and Hoch (1 997) regarded them as a distinct Asteraceae Asteraceae (Compositae) subspecies of K. yomena endemic to J apan (48) (48) Ainsliaea uniflora Sch. Bip. f. nivea (subsp. angustifolia (Nakai) H. Y. Gu) (M. (M. Mizush.) Yone k.， comb. nov. - because of the exsistence of intermediate Diaspananthus Diaspananthus uniflorus (Sch. Bip.) Kitam. plants. On the other hand ， K. yomena is be- f. f. niveus M. Mizush. in J. Jpn. Bo t. 30: 95 lieved to have originated as a hybrid between (1 955). K. indic α (Aster indicus) and subsp. Nom. Jap.: Yuki-yatsude. angustifolia (Shimotomai and Inoue 1951). Difficulty Difficulty for distinguishing the subsp. (49) (49) Aster hispidus Thunb. v紅. hispidus f. angustifolia from K. yomena has been ex- albidiflorus albidiflorus (Akasawa) Yone k.， comb. nov. plained as a result of back-crossing with - Heteropappus hispidus (Thunb.) Less. f. parental species (Shindo 1966a ， 1966b). albidiflorus albidiflorus Ak asawa in Bull. Kochi When the origin of K. yomena is considered ， Women's Univ. ，ser. Na t. Sci. 27: 5 (1 979) ， it is natural to regard Gu and Hoch (1997)' s as as "albidoflorus". subsp.α ngustifolia as a distinct species. Nom. Jap.: Shirobana-areno-nogiku. As Kalimeris seems insufficiently distinct from from Aster (s. s甘.) at generic rank (lto et al. (50) (50) Aster microcephalus (Miq.) Franch. & 1995) ，1 make the following combination Sav.v 紅. ovatus (Franch. & Sav.) So 吋ima& under the genus Aster: Mo t. It o f. ovatus (Franch. & Sav.) Y one k.， Aster robustus (Makino) Y one k.， stat. comb. comb. nov. - A. trinervius Roxb. ex D. Don nov. - Aster pinnatifidus (Maxim. ex V 訂. s. ovatus Franch. & Sav. ，Enum. Pl. Jap. Makino) Makino f. robustus Makino in Bo t. 1: 1: 222 (1 875) ，‘ ovata'. Mag. Tokyo 27: 115 (1 913). Nom. Jap.: Nokongiku. Other synonyms: f. f. albidus (Kitam.) Yone k.， comb. nov. - Kalimeris incisa (Fisch.) DC. var. robusta A. αgeratoides subsp. ov α tus (Franch. & (M 紘 ino) Ki tag. ， Neo-Lineam. Fl. Mansh.: Sav.) Sav.) Ki tam. f.α lbidus Kitam. in Mem. Coll. 653 (1 979) et in Ohwi & Ki tag. ，New Fl. Sci. Sci. Kyoto Imp. Univ. ser. B ， 13: 352 (1 937). Japan: 1460 (1 983). Nom. Jap.: Shirobana-nokongiku. K. pinnatifida (Maxim. ex Makino) Kitam. f. f. humilis (Nakai) Yonek. ， comb. nov. - var. angustifolia Nakai in Bull. Natn. Sci. A. A. ageratoides Turcz. var. humilis Nakai ， Mus. Tokyo (33): 26 (1953). Ca t. Sem. Spor. Hort. Bo t. Univ. Tokyo K. yomena (Ki tam.) Kitam. subsp. [1916: [1916: 31 (1 916) ， nom. nud.] 1920: 39 angustifolia (Nakai) H. Y. Gu in H. Y. Gu & (1 920). Hoch in Ann. Missouri Bo t. Gard. 84: 799 ，f. Nom. Jap.: Komachi-giku. 17D (1 997). f. f. vernalis (Honda) Yone k.， comb. nov. Misapplied names: - Aster ageratoides Turcz. var. vernalis Aster incisus auc t. non Fisch.: Nakai in Honda in Bo t. Mag. Tokyo 45: 45 (1 931). Bo t. Mag. Tokyo 33: 214 (1919) ， p. p.; H. Nom. Jap.: Haru-nokongiku. Hara ，Enum. Sperm. Jap. 2: 131 (1 952); Mo t. December 2005 Journal of Japanese Botany Vo l. 80 No. 6 331

It o & Soejima in K. Iwats. & a1.， F1. Japan thunbergii (A. Gray) Maxim. var. alb 伊ora 3b: 3b: 66 (1 995). Makino in Bo t. Mag. Tokyo 12: 48 (1 898). Kalimeris Kalimeris incisa auc t. non (Fisch.) DC.: f. amplifolium (Kitam.) H. N akai & H. Ki tam. in Mem. Col 1. Sci. Kyoto Imp. Univ. Ohashi ，comb. nov. - /xeris dentata var. ser. ser. B ， 13: 309 (1937) ， p. p. ，quoad specim. amplifolia Kitam. in Bo t. Mag. Tokyo 49: ex Jap. ， in Ohwi ， F 1. Jap. ed. Eng 1.: 869 285 (1 935). (1 965) et in Satake & a1.， Wild Flow. Jap. Nom. Jap.: Hana-nigana ， Obana-nigana. Herb. Herb. P1. 3: 191 (1 981); Ohwi ， F 1. Jap.: 1151 f. atropurpureum (Nakai) H. Nakai & H. (1 953); Kitam. & Murata ，Co 1. 11 1. Herb. P1. Ohashi ，comb. nov. - /xeris dentata var. Jap. Jap. 1: 85 (1 957). atropurpure αNakai in Bo t. Mag. Tokyo 42: Nom. Jap.: O-yuga-giku. 16 (1 928). Nom. Jap.: Kuro-nigana. (52) (52) Crepidiastrum denticulatum (Hout t.) f. leucanthum (H. Hara) H. Nakai & H. J. J. H. Pak & Kawano f. pallescens (Momiy. Ohashi ，comb. nov. - /xeris dentata var. & Tuyama) Yone k.， comb. nov. - Paraixeris octoradiata Nakai f. leucantha H. Hara in J. denticulata denticulata (Hout t.) N akai f. pallescens Jpn. Bo t. 10: 435 (1 934). Momiy. & Tuyama in Tuyama in J. Jpn. Bo t. Nom. Jap.: Shirobana-nigana. 21: 21: 112 (1 947). var. nipponicum (Nakai) H. Nakai & H. Nom. Jap.: Usuiro-yakushiso. Ohashi ，sta t. nov. - /xeris n伊'P onica N akai in Bo t. Mag. Tokyo 34: 154 (1 920). (53) (53) Crepidiastrum xsurugense (Hisauti) Nom. Jap.: Iso-nigana. Yone k.， comb. nov. - Paraixeris surugensis subsp. ozense (Sugim.) Yonek. ，comb. et Hisauti Hisauti in J. Jpn. Bo t. 10: 698 ， photo (1 934). sta t. nov. - 1 χeris dentata (Thunb.) Nakai Nom. Jap.: Yakushi-azetona. subsp. stolonifera Ki tam. v紅 . ozensis Sugim. in in J. Jpn. Bo t. 32: 64 (1957). (54) (54) New combinations in /xeridium Nom. Jap.: Oze-nigana. 1 agree with Pak and Kawano (1 992) on generic generic circumscriptions of /xeris and related (55) Nemosenecio nikoensis (Miq.) B. genera. genera. /xeridium dentatum is polymorphic Nord. f. albiflorus (Sugim.) Yonek. ，comb. in in Japan and many infraspecific taxa have nov. Senecio nikoensis Miq. f. albiflorus been recognized. Although Pak and Kawano Sugim. in J. Jpn. Bo t. 32: 64 (1 957). (1992) (1992) in cI uded almost all taxa growing in Nom. Jap.: Shirobana-sawagiku. lowlands lowlands in subsp. dentatum ， 1 follow Yahara (1 995)' s opinion who recognized (56) Parasenecio delphiniifolius (Siebold & two subspecies and several varieties in Zucc.) H. Koyama v紅. brevilobus (Sugim. subsp. subsp. dentatum sensu Pak and Kawano & Sugino) Yone k.， comb. nov. - Cacalia

(1992). (1992). delphiniifollia Siebold & Zucc. v紅. Ixeridium dentatum (Thunb.) Tzvelev breviloba Sugim. & Sugino in J. Geobo t. subsp. subsp. dentatum f. stoloniferum (Ki tam.) (Kanazawa) 26: 81 ， photo (1978). Yone k.， comb. et sta t. nov. - La ctuca Nom. Jap.: Ogasa-momijigasa. dentata dentata (Thunb.) C. B. Rob. v紅 . stolonifera Ki tam. ，Compos. Nov. Jap.: 26 (1 931). (57) Saussurea kudoana Tatew. & Kitam. Nom. Jap.: Hai-nigana. var. yuparensis (Ki tam.) Yone k.， comb. subsp. subsp. nipponicum (Nakai) 1. H. Pak & nov. - Saussurea riederi Herder subsp. Kawano var. albiflorum (M 紘 ino) H. N akai kudoan α(Tatew. & Ki tam.) Ki tam. v紅. & H. Ohashi ， comb. nov. - Lactuca yuparensis Kitam. in Mem. Coll. Sci. K yoto 332 植物研究雑誌第80 巻第6号 平成17 年12 月

Imp. Imp. Univ. ser. B ， 13: 173 (1 937). sphondylodes Trin. va r. subtrivi αlis Ohwi in Nom. Jap.: Yubari-kita-azami. Acta Phytotax. Geobo t. 10: 126 (1941). Nom. Jap.: Hiroha-ichigo-tsunag i. 島lonocotyledoneae Liliaceae Liliaceae (64) Urochloa villosa (Lam.) T.Q. Nguyen f. (58) (58) Hosta tsushimensis N. F吋ita f. glabriglumis (Ohwi) Yone k.， comb. nov. - albiflora albiflora (S. Toyama) Yone k.， comb. nov. Brachiaria villosa (Lam.) A. Camus v紅. - H. minor Nakai f. albiflora S. Toyama in glabriglumis Ohwi in Acta Phytotax. J. J. Jpn. Bo t. 36: 141 (1 96 1). Geobo t. 5: 51 (1 936) - Brachiaria villosa f. Nom. Jap.: Shirobana-tsushima-giboshi glabriglumis (Ohwi) Ohwi in Acta Phytotax. (nov よ Shirobana-keirin-giboshi (Toyama Geobo t. 11: 43 (1 942). 1961). 1961). Nom. Jap.: Rasha-kibi.

Poaceae (Gramineae) Cyperaceae (59) (59) Calamagrostis brachytricha Steud. (65) Scleria rugosa R. Br. var. onoei

V紅. inaequata (Hac k. ex Honda) Yone k.， (Franch. & Sav.) Yone k.， comb. nov. - S. comb. nov. - Calamagrostis arundinacea onoei Franch. & Sav. ，Enum. Pl. Jap. 2: [1 22 (L.) (L.) Roth var. inaequa ωHac k. ex Honda in (1 877) ，nom. nud.] 549 (1 878). Bo t. Mag. Tokyo 40: 441 (1 926). Nom. Jap.: Maneki-shinju-gaya. Nom. Jap.: Saishu-nogariyasu. 1 wish to express my sincere thanks to Dr. (60) (60) Elymus xnakashimae (Ohwi) Yone k.， Hideki N akai ， for making his large amount comb. nov. - Agropyron nak α shimae Ohwi of literature available ， for critical comments in in Acta Phytotax. Geobo t. 11: 257 (1 942). on manuscript and for his kind permission to publish publish new combinations of /xeridium pre- (6 1) Ischaemum aristatum L. var. pared by him. 1 am also indebted to Prof. crassipes crassipes (Steud.) Yonek. ，comb. nov. - l. Emeritus Hiroyoshi Ohashi ， Sendai and Dr. crassipes crassipes (Steud.) Thell. in Reper t. Spec. Tadashi K 司jita ，Chiba University for reading Nov. Regni Veg. 10: 289 (1 912); Nakai in the manuscrip t. My thanks 紅 e also due to Bo t. Mag. Tokyo 33: 1 (1 919) ，cum Var' Prof. Jin Murata ， University of Tokyo ， for typicum typicum Nakai. his encouragemen t. This study is supported

よcrassipes v紅 . glaucum Honda in J. Fac. by Grants-in Aid nos. 137002 & 147003 Sci. Sci. Imp. Univ. Tokyo sec t. 3，Bo t. 3: 354 from Japan Society for the Promotion of (1930) (1930) - l. aristatum var. glaucum (Honda) Science to Prof. Jin Murata ， University of T. T. Koyama in 1. Jpn. Bo t. 37: 239 (1 962). Tokyo. Nom. Jap.: Kamonohashi. References References (62) (62) Neomolinia japonica (Franch. & Sav.) Fang R.-c. and Brummitt R. K. 1995. Calystegia R. Proba t. f. musashiensis (Hiyama) Y one k.， Brown. In: Wu Z.-Y. and Raven P. (eds.) ，Fl ora of comb. nov. - Diarrhena japonica Franch. & China 16: 28 6- 289. Science Press ，Beijing and Missouri Missouri Botanical Garden Press ， S t. Louis. Sav. Sav. f. musashiensis Hiyama in 1. Jpn. Bo t. Gu H. ーY. and Hoch P. C. 1997. Systematics of 28: 28: 218 (1953). Kalimeris (Asteraceae: Astereae). Ann. Missouri Nom. Jap.: Yokoyama-tatsunohige. Bo t. Gard. 84(4): 762-814. Hara Hara H. 1948. Notes on the Japanese species of (63) (63) Poa ochotensis Trin. var. subtrivialis Geranium. J. Jpn. Bo t. 22 (1 0-12): 165-172 (in Japanese). Japanese). (Ohwi) Yonek. ， comb. nov. - P. December December 2005 Journal of Japanese Botany Vo l. 80 No. 6 333

一一一 1949. Enumeratio Spermatophytarum Japonica- Geranium ， subgenus Geranium of Asia rum 1: 160. Iwanami Shoten ， Tokyo. (Geraniaceae). Korean J. Pl. Taxon. 27(2): 195- 一一一 1955. Critical notes on some type specimens of 231 (in Korean). East-Asiatic East-Asiatic plants in foreign herbaria (1). J. Jpn. Shimotomai N. and Inoue S. 195 1. Morphological and Bo t. 30 (1): 19-26. cytological studies on Kalimeris yomena Ki tamura ， 一一一 1957. On some double-flowered cultivated plants a natural hybrid. J. Sci. Hiroshima Univ. ser. B ， of of old Chinese origin. J. Jpn. Bo t. 32 (1 0): 313-315 Div. 2， 6: 1-9 ，f. 1-3. (in (in Japanese). Shindo K. 1966a. Cytological ，mo 中hological and geo- It o M. ， Soejima A.， Hasebe M. and Watanabe K. 1995. graphical studies on the differentiation of species A chloroplast rDNA phylogeny of Kalimeris and of Kalimeris in Japan IV. Geographical distribution Aster ， with reference to the generic circumscrip- and chromosome number of Kalimeris in cI sa. Bo t. tion. tion. J. Pl. Res. 108: 93-96. Mag. Tokyo 79: 12 4- 130 (in Japanese). Nakai Nakai T. 1912. Notulae ad plantas Japoniae et Coreae 一一一 1966b. Cytological ， morphological and geo- VI I. Bo t. Mag. Tokyo 26(308): 251-266. graphical studies on the differentiation of species 一一一 1935. Notulae ad plantas Japoniae et Koreae of Kalimeris in Japan V. Kalimeris yomena. Bo t. XLV. Bo t. Mag. Tokyo 49(582): 345-35 1. Mag. Tokyo 79: 355-365 (in Japanese). Pak J.-H. and Kawano S. 1992. Biosystematic studies Yahara T. 1995. 9. /xeris Cass. /n: Iwatsuki K.， on the genus /xeris and its allied genera (Composi- Yamazaki T. ， Boufford D. E. and Ohba H. (eds よ tae-Lactuceae) tae-Lactuceae) IV. Taxonomic treatments and no- Fl ora of Japan 3b: 15-19. Kodansha Ltd. ， Tokyo. menclature. menclature. Mem. Fac. Sci. Kyoto Univ. (ser. Yeo P. F. 1992. A revision of Geranium L. in South- Bio l.) 15: 29-6 1. west China. Edinburgh J. Bo t. 49(2): 123-21 1. Park Park S. -J. and Ki m Y. -S. 1997. of

米倉浩司:日本とその周辺の植物に関する分類学 的研究 (1) 一日本産植物の新組合せおよび新名一 筆者は梶田 忠助教授(千葉大学)と共同で， of Japan IIa で、取り扱われる双子葉植物離弁花類の 日本に自生，帰化している全ての維管束植物と和 一部および札幌の中井秀樹博士が作成中の北海道 名のついている主要な栽培植物，有用植物につい 産維管束植物目録で取り扱われる予定の新組合せ て，和名と学名(戦後の文献に現れた異名も含む) を除いた 81 新組合せと 5 新名について発表する. およびその出典と，日本の主要植物誌(北村他編 また， (18) グンナイフウロ， (37) ヒルガオと 『原色日本植物図鑑JI (1957-1979) ，佐竹他編『日 (51) オオユウガギクについては本文にやや詳し 本の野生植物JI (1982-2003) ，大井・北JII r新日 い解説を付し，タカネグンナイフウロを新品種と 本植物誌JI (1983) および、 Iwatsuki & al. (eds.) して記載した.また， (54) ニガナについては， rFl ora of Japan Jl (1993-) にそれらの学名が現れ 中井秀樹博士の好意により 博士が上記目録のた るページ，および所属する科名を対応させたデー めに準備していた 5 個の新組合せ (H. Nakai & H. タベースを構築した (URL は本文を参照されたい) . Ohashi に帰属させられているもの)をここに発表 この構築の過程で，多数の新組合せや新名が必要 することができた.これらの組合せの著者名は， となり，またいくつかの植物群では学名の選択に 国際植物命名規約第46.2 条の規定により H. 関して筆者の基準を示す必要に迫られた.今回は， Nakai & H. Ohashi となり H. Nakai & H. Ohashi ex 主に命名規約上の理由により必要となった種子植 Yonek. で、はないので 引用の際は注意されたい. 物の新組合せと新名のうち 近く出版されるFl ora (東北大学植物園)