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Salari 381..408 ivist stlin di leontologi e trtigrfi volume IPH noF Q I plF ppF QVIERHV heemer PHIR ri vei visygixi eixxsxi grewys pyw qye wye geyy @swf sxs wy xesxD gixev sevA IB P I viyxehy eves D wesy pF yvpy 8 gewivy iyxsy st eeivedX eptemer I D PHIRY eptedX xovemer PVD PHIR uey wordsX vte leistoeneD upiprD qrott wor gvorsoD of outhern iurope nd xer ist @from QSë to SHë xA stlyF @eulgnier et lF PHHVY rerrero et lF PHHVAD ut in the vte leistoene its spred ws muh widerD inluding estrtF his pper desries suEentireD prtilly rtiuE lowEltitude regions nd lso ostl res @wsini IWVSY lted skeleton of hmois found together with other verterte reE mins in silty lens with limestone lsts etween the IHR nd IHS wsini 8 vovri IWVVY wsseti 8 lri PHIPAF of qrott wor gvorso @vtiumD stlyAF his silty lens is he genus is urrently divided into two speies hronologilly orreltle to temperte osilltion of ws QF @wsini 8 vovri IWVVY qru PHHSAX the elpine hE he skull nd postErnil remins from qrott wor gvorso were moisD upipr rupipr @vinneusD IUSVAD divided ompred with the fossil remins of hmois from numerous vte into seven regionl suspeiesD spred from entoli leistoene nd older roloene sites of stly nd of yrenees nd serin eninsulF edditionl omprisons were mde with lrge smE nd the gusus to the elpsY tht of the yreneesD ples of extnt hmoisD upipr pyreni orntD F pyreni pyrE upipr pyreni fonprteD IVRSD widespred in eni nd F rupipr rupiprF worphologil nd morphometri the serin eninsulD with its suspeies from the yrE fetures of the skull nd the hornEoreD s well s other hrters of enees nd the gntrin wountinsD whih hs surE the metrpus suggest to refer the fossil remins from qrott wor gvorso to epennine hmoisD F pyreni orntF his is the erE vived up to present in stly only within the eruzzi liest sertined reord in gentrlEouthern stly of the orntElike xtionl rk in the form of the suspeies F pyreE hmoisD suspeies so fr known with ertinty only in the roloE ni ornt xeumnnD IVWW @see eulgnier et lF PHHVY eneD or possily in the vteglilF rerrero et lF PHHVAF eentlyD the epennine hmoisD F pyreni orntD hs een suessfully reintrodued in other xtionl rks in gentrl stly @wri 8 vovri sntrodution PHHWAF he erliest fossil reords of hmois in estern he vte leistoene one remins of hmois iuropeD referred to upipr fF pyreniD ome from from qrott wor gvorso @tenneD vtiumD stlyAD more lyer «p» of the gune de l9ergo @utvelD yreÂneÂesE speifilly suEentireD prtilly rtiulted skeletonD yrientlesD prneAD dted out RRHDHHH yers f nd re desried in this pperF orrelted with the wrine ssotope tge IP @ws IPA emins of upipr de flinvilleD IVIT our in @greÂgut IWUWY woigne et lF PHHTY greÂgutEfonnoure estern iurope sine the widdle leistoene nd re PHHUAF he sme txon ws lso reorded from the site reltively ommon in the vte leistoene of stly @wE of yrgn Q @erdeÁheD prneA in lyers dted etween sini 8 vovri IWVVY greÂgutEfonnoure PHHUY wsseti 8 QQWDHHH nd PWVDHHH yers f @ws WA @ivls PHHRAF lri PHIPD PHIPAF he finds then eome slightly more numerous in the gurrent distriution of this genus inludes severl portions of the middle to highEltitude mountin rnges finl stges of widdle leistoene @ws TA nd mong I B gorresponding uthorF hiprtimento di ienze dell errD pienz99 niversitÁ di omD izzle eldo woro SD sEHHIVS omD stlyF iEmilX leonrdoslridvirgilioFit P hiprtimento di ienze toriheD filosofioEsoiliD dei eni ulturli e del territorioD niversitÁ or ergt99D i golumi ID sEHHITQ omD stlyF QVP lri vFD olfo wF pF 8 etronio gF pigF I E qrott wor gvorsoX eA loE tion of the siteY fA prtil longitudinl setion @upA nd omplete pln @downA of the veF these we n mention other prenh fossil reminsX from nd the writime elpsD s well s in the xorthern the qrotte du gp de l fielle @xestierD rutesEyreÂE epennines @wsini IWVSY wsini 8 vovri IWVVAD or neÂesAD referred to upipr fF pyreniY from the possily slightly frther southD t lest up to isno qrotte des geÁdres @ve ln d9eupsD rA nd the qrotte wountD usnyD yrrhenin side @wsseti 8 lri du vzret @xieD elpesEwritimesAD ttriuted to F PHIPAF he epennine hmois ws onfined to porE rupiprY from the qrotte de l xihe @wontmurinD tion of peninsul etween northern glri nd the ruteEqronnAD ttriuted to F pyreni nd from iillini wountins @southern mriEwrhe epenE ominElEohe @ominD housAD referred to upiE ninesA @wsini IWVSY wsini 8 vovri IWVVAD or proE pr fF rupipr @see glot 8 wrsn IWVTY wsini 8 ly slightly frther northD up to the entrl mriE vovri IWVVY hefleur et lF IWVWY voso et lF IWWHY wrhe epenninesD edriti side @wsseti 8 lri greÂgutEfonnoure IWWPD PHHUY lensi PHHWY erouteÁre PHIPAF 8 queÂrin PHIHAF wsini @IWVSD with referenesA quoted QP stlin possil remins ttriuted to upipr in stly re sites with fossil nd suEfossil remins of hmois nd reltively numerous during the vte leistoeneD espeE wsseti 8 lri @PHIPD with referenesA RQ lolities of illy from sites referred to the vst qlil @urm gentrlEouthern stly with the hmois finds referred È eutorumAD ut the txon proly rehed the peninE to the vte leistoene nd the older roloene ordE sul during the finl stges of the widdle leistoene ing to the literture fter IWVVF eprt from these oveD @ws TA @wsini IWVSY etronio et lF PHHUD PHIIY E in stly we n dd some preElpine northEestern nd lomo et lF PHIIAF he ssumed distriution of upiE vigurin sites @eFgFD gliozzo 8 gssoli IWWHD IWWPY pr in stly in the vte leistoene nd the older gssoli 8 gliozzo IWWID IWWRD IWWRY piore 8 E roloeneD sed on the urrent distriution of hmois gliozzo PHHSD with referenesY ierer 8 fosto PHHTY popultions nd the wellEpreserved remins of only SEU lensi PHHWD with referenesY qurioli et lF PHIHD sitesD seems to e s followsF elpine hmois ws wideE PHIHY fertolini et lF PHIPA nd the insulr finds of il spred in the elpine rhD etween the istern elps islnd @wsseti in litterisAF sndeedD exept for few sesD vte leistoene epennine hmois QVQ trtigrphi notes of qrott wor gvorso suh s qrott di iquiD wonte umr nd ip qrnde @wsini IWVSA nd ipro di pumne @gssoli 8 gliE qrott wor gvorso opens to UIS m FsFlF ner ozzo IWWIY omndini et lF PHIRAD the remins of hE tenne @vtiumD gentrl stlyA nd it lens out on the mois re reltively poor ndGor highly frgmentedF preE right slope of the upper eniene iver vlley inside the quently they re in suh stte of preservtion tht the imruini wountins egionl xturl rk @pigF IeAF lk of dignosti osteologil mterils does not llow his krst vity hold suession of rooms nd duts definite ttriutions to one or the other speiesF ghE @pigF IfA nd is rved in the glilutiti e glreniti mois is lso well represented from qrott delle flze del goninoEgmpnino99 formtion @gWEIIGQVD with one remins of lmost four young nd two dult pper greteousA @hmini et lF IWWVY olfo et lF individuls utD unfortuntelyD the hornEores with PHHWAF pollowing the fortuitous disovery of humn frontl ones mentioned in wsini 8 vovri @IWVVA ones inside the veD rheologil investigtions nd pulished y voreti 8 lerno @IWVWA were not strted in PHHT y or ergt99 niversity of omeD newfoundF hereforeD the suEentire skeleton from in greement with the oprintendenz per i feni erE qrott wor gvorso is relly n exeptionl nd interE heologii99 of vtium nd with the ollortion of esting se studyF pigF P E qrott wor gvorsoX strtiE grphy of the room IF BpE proximte position of the silty lens with the prtilly rtiulted skeleton of hE moisF QVR lri vFD olfo wF pF 8 etronio gF hk ulu99 speleologil groupF he inner rooms held tiulrly wirotus @wirotusA rvlis nd erviol mE the remins of t lest PI individulsD the lrgest xeoE phiius with rre lp eD F romnD hinolophus lithi humn ommunity of vtium nd one of the rihE ferrumequinumD wyotis ehsteiniiD wF emrgintusD est of iurope @olfo et lF PHHWD PHIHD PHIPD PHIPY winiopterus shreiersiiD wirotus @erriolA svii nd lri et lF PHIPAF sn the room ID just elow the disE epodemus sylvtiusGflviollis @exvtion PHHVY see tured surfe nd some roloene lyers @xeolithiD lri PHIRAF fronze ege nd historil geD see olfo et lF PHIHD felow the forementioned ontexts is the IHR D PHIPD PHIQY ilvestri et lF in pressAD the vte leistoene investigted etween PHHV nd PHIPF his lyer onsists sequene ws found @pigF PAF of inoherent soilD minly hrterized y entimetri he trtigrphi nits @ A IHWD IIHD IPHD limestone lstsF sn this D out TH m thikD no huE IQSD PSR nd PSS re hrterized y ompt redE mn rteftsD ut very undnt miroverterte reE dishErown silty ground with limestone lsts with mins together with severl one frgments of mroE shrp edges tht vry in dimeter from few millimeters mmml hve een foundF reliminry nlysis shows to one entimeterD where mny fossil remins of verteE tht the dominnt txon is wF @wFA rvlis @lmost RVH rte were found together with moderte numer of mndilesD out RH7 of determined remins nd more lithi rtefts referle to lte pper leolithi @finl thn UH7 of individuls mong the mmmlsA followed ipigrvettinAF prontl srper on retouhed lde of y severl remins of eF mphiius nd irdsD vepus spF jspered flintD frgments of ked lde letsD frgment nd wrtes mrtesD sre wF mrmotD eF sylvtiusG of ked point nd multiple urin re the findingsF sn flviollis nd gF iexD nd rre eptiliD F eD F this ipigrvettin lyerD out QH m thikD the most romnD orex smnitiusD wyodes glreolusD wustel undnt txon is wrmot mrmot @WR speimensD putoriusD upipr spF nd fos primigeniusF por strtiE out hlf of the determined remins nd one third of grphi position nd for the omposition of mmml A rvlis ompnied the individulsA followed y gervus elphusD gpr ssemlgeD dominted y wF @wF y old99 mmmlsD suh s mrmot nd iexD this lyer iexD upipr spF nd vepus spFD present with sigE ws orrelted with the vst qlil wximum @vqwA nifint
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