Germany) and Its Bearing on the Tectonostratigraphic History of the Saxothuringian Domain
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Available Generic Names for Trilobites
AVAILABLE GENERIC NAMES FOR TRILOBITES P.A. JELL AND J.M. ADRAIN Jell, P.A. & Adrain, J.M. 30 8 2002: Available generic names for trilobites. Memoirs of the Queensland Museum 48(2): 331-553. Brisbane. ISSN0079-8835. Aconsolidated list of available generic names introduced since the beginning of the binomial nomenclature system for trilobites is presented for the first time. Each entry is accompanied by the author and date of availability, by the name of the type species, by a lithostratigraphic or biostratigraphic and geographic reference for the type species, by a family assignment and by an age indication of the type species at the Period level (e.g. MCAM, LDEV). A second listing of these names is taxonomically arranged in families with the families listed alphabetically, higher level classification being outside the scope of this work. We also provide a list of names that have apparently been applied to trilobites but which remain nomina nuda within the ICZN definition. Peter A. Jell, Queensland Museum, PO Box 3300, South Brisbane, Queensland 4101, Australia; Jonathan M. Adrain, Department of Geoscience, 121 Trowbridge Hall, Univ- ersity of Iowa, Iowa City, Iowa 52242, USA; 1 August 2002. p Trilobites, generic names, checklist. Trilobite fossils attracted the attention of could find. This list was copied on an early spirit humans in different parts of the world from the stencil machine to some 20 or more trilobite very beginning, probably even prehistoric times. workers around the world, principally those who In the 1700s various European natural historians would author the 1959 Treatise edition. Weller began systematic study of living and fossil also drew on this compilation for his Presidential organisms including trilobites. -
Th TRILO the Back to the Past Museum Guide to TRILO BITES
With regard to human interest in fossils, trilobites may rank second only to dinosaurs. Having studied trilobites most of my life, the English version of The Back to the Past Museum Guide to TRILOBITES by Enrico Bonino and Carlo Kier is a pleasant treat. I am captivated by the abundant color images of more than 600 diverse species of trilobites, mostly from the authors’ own collections. Carlo Kier The Back to the Past Museum Guide to Specimens amply represent famous trilobite localities around the world and typify forms from most of the Enrico Bonino Enrico 250-million-year history of trilobites. Numerous specimens are masterpieces of modern professional preparation. Richard A. Robison Professor Emeritus University of Kansas TRILOBITES Enrico Bonino was born in the Province of Bergamo in 1966 and received his degree in Geology from the Depart- ment of Earth Sciences at the University of Genoa. He currently lives in Belgium where he works as a cartographer specialized in the use of satellite imaging and geographic information systems (GIS). His proficiency in the use of digital-image processing, a healthy dose of artistic talent, and a good knowledge of desktop publishing software have provided him with the skills he needed to create graphics, including dozens of posters and illustrations, for all of the displays at the Back to the Past Museum in Cancún. In addition to his passion for trilobites, Enrico is particularly inter- TRILOBITES ested in the life forms that developed during the Precambrian. Carlo Kier was born in Milan in 1961. He holds a degree in law and is currently the director of the Azul Hotel chain. -
An Inventory of Trilobites from National Park Service Areas
Sullivan, R.M. and Lucas, S.G., eds., 2016, Fossil Record 5. New Mexico Museum of Natural History and Science Bulletin 74. 179 AN INVENTORY OF TRILOBITES FROM NATIONAL PARK SERVICE AREAS MEGAN R. NORR¹, VINCENT L. SANTUCCI1 and JUSTIN S. TWEET2 1National Park Service. 1201 Eye Street NW, Washington, D.C. 20005; -email: [email protected]; 2Tweet Paleo-Consulting. 9149 79th St. S. Cottage Grove. MN 55016; Abstract—Trilobites represent an extinct group of Paleozoic marine invertebrate fossils that have great scientific interest and public appeal. Trilobites exhibit wide taxonomic diversity and are contained within nine orders of the Class Trilobita. A wealth of scientific literature exists regarding trilobites, their morphology, biostratigraphy, indicators of paleoenvironments, behavior, and other research themes. An inventory of National Park Service areas reveals that fossilized remains of trilobites are documented from within at least 33 NPS units, including Death Valley National Park, Grand Canyon National Park, Yellowstone National Park, and Yukon-Charley Rivers National Preserve. More than 120 trilobite hototype specimens are known from National Park Service areas. INTRODUCTION Of the 262 National Park Service areas identified with paleontological resources, 33 of those units have documented trilobite fossils (Fig. 1). More than 120 holotype specimens of trilobites have been found within National Park Service (NPS) units. Once thriving during the Paleozoic Era (between ~520 and 250 million years ago) and becoming extinct at the end of the Permian Period, trilobites were prone to fossilization due to their hard exoskeletons and the sedimentary marine environments they inhabited. While parks such as Death Valley National Park and Yukon-Charley Rivers National Preserve have reported a great abundance of fossilized trilobites, many other national parks also contain a diverse trilobite fauna. -
Paradoxides Œlandicus Beds of Öland, with the Account of a Diamond
SVERIGES GEOLOGISKA UNDERSÖKNING SER. c. Avhandlingar och uppsatser. N:o 394· ÅRSBOK 30 (1936) N:o 1. PARADOXIDEs CELANDICUS BEDS OF ÖLAND WITH THE ACCOUNT OF A DIAMOND BORING THROUGH THE CAMBRIAN AT MOSSBERGA BY A. H. WESTERGÅRD Witlt Twelve Plates Pris 3:- kr. STOCKHOLM 1936 KUNGL. BOKTRYCKERIET. P. A. NORSTEDT & SÖNER 36!789 SVERIGES GEOLOGISKA UNDERS ÖKNING SER. c. Avhandlingar och uppsatser. N:o 394· ÅRSBOK 30 (1936) N:o r. PARADOXIDEs CELANDICUS BEDS OF ÖLAND WITH THE ACCOUNT OF A DIAMOND BORING THROUGH THE CAMBRIAN AT MOSSBERGA BY A. H. WESTERGÅRD With Twelve Plates STOCKI-IOLM 1936 KUNGL. BOKTRYCKERIET. P. A. NORSTEDT & SÖNER CONTENT S. Page I. A Diamond Boring through the Cambrian at Mossberga . 5 Quartzite . ... 5 Lower Cambrian Deposits and the Sub-Cambrian Land Surface 7 Paradoxides oolandicus Beds !3 Comparison of the Mossberga and Borgholm Profiles 16 II. Paradoxides oolandicus Beds of Öland . 17 Introduction and History . 17 Distribution, Thickness, and Stratigraphy 19 Acknowledgements 22 Fauna ..... 22 Brachiopoda . 22 M icromitra . 2 2 Lingulella 23 Acrothele . 24 Acrotreta .. 24 Orthoid brachiopod 25 Gastropoda 25 ffilandia .. 25 Hyolithidae 26 Trilobita . 27 Condylopyge 27 Peronopsis 28 Agnostus . 29 Calodiscus 30 Burlingia 32 Paradoxides 33 Ontogeny of Paradoxides 45 Ellipsocephalus 56 Bailiella . 58 Solenopleura . 59 Phyllocarida. Hymenocaris (?) 61 Geographical and Stratigraphical Distribution of the Fauna Elements 62 Bibliography . 64 Explanation of Plates . 67 I. A Diamond Boring through the Cambrian at Mossberga. As a part of the work carried out by the Electrical Prospecting Co. (A.-B. Elektrisk Malmletning) in order to search for gas in the Lower Cambrian of Öland, a deep boring was made in the autumn of 1933 into a little flat dome at Mossberga, about 12 km S of Borgholm. -
10. Brief Notes on the Eodiscids 11, Phylogeny of the Dawsonidea
No. 1.] 43 10. Brief Notes on the Eodiscids 11, Phylogeny of the Dawsonidea. By Teiichi KoBAYASHI. (Comm. by H. YABE,M.I.A., Jan. 12, 1943.) Microdiscus has long been in common use for eodiscids, but as pointed out by Barrande in 1881, Microdiscus quadricostatus Emmons which is the type species of the genus, was a larval form of Trinucleus which was 'found in association with Ordovician graptolites and pele- cypods. Because Microdiscus Emmons, 1885, and the Microdiscidae Koken or Coquin, 196, lost their standing, Raymond adopted Eodiscus Hartt in place of Microdiscus and established Eodiscidae to include, beside the genus, two new ones, Goniodiscus and Weymouthia. While the three genera were all blind, Delgado described eodiscids with eyes in 1904 and Vogdes founded Delgadoia on Microdiscus caudatus Delgado in 1917. One year prior to this, Walcott had described proparian Pagetia as a member of the Eodiscidae, which became a subject of debate among trilobite students. Poulsen mentioned in 1927 that the sole reason for the resemblance of Pagetia to hypoparian eodiscids is adaptative convergence. This opinion is intenable because it is now known in Pagetia, Delgadoia or Al emtejoia that the two forms of each genus which are almost identical except for the presence or the absence of eyes and facial sutures, are found at the same locality. Supporting Beecher's view on the migration of the facial suture, Raymond emphasized in 1917, that Pagetia shows the stage in which the suture just came up to the dorsal side from the ventral in the agnostidian stage, notwithstanding the fact that Dindstrom and others have proven that the agnostid has no suture on the ventral side of its cephalon. -
Paradoxides CELANDICUS BEDS of ÖLAND
SVERIGES GEOLOGISKA UNDERSÖKNING SER. c. Avhandlingar och uppsatser. N:o 394· ÅRSBOK 30 (1936) N:o r. PARADOXIDEs CELANDICUS BEDS OF ÖLAND WITH THE ACCOUNT OF A DIAMOND BORING THROUGH THE CAMBRIAN AT MOSSBERGA BY A. H. WESTERGÅRD With T w elve Flates S TO C KH O LM 19 36 KUNGL . BOKTRYCKERIET . P. A. NORSTEDT & SÖNER 36!789 CON TENTS. Page I. A Diamond Boring through the Cambrian at Mossberga . 5 Quartzite . 5 Lower Cambrian Deposits and the Sub-Cambrian Land Surfa ~e 7 Paradoxides relandicus Beds . 13 Comparison of t he Mossberga a nd Borgholm Profiles r 6 II. Paradoxides relandicus Beds of Öland . I7 Introduction and History . I7 Distribution, Thickness, and Stratigraphy I g Acknow ledgements 22 Fauna . ... 22 Brachiopoda . 22 M icromitra. 22 Lingulella 23 Acrothele . 24 Aerotre/a . 2 4 Orthoid brachiopod 25 Gastropoda 25 CElandia .. 25 H yolithidae 26 Trilobita . 27 Condylopyge 27 Peronopsis 28 Agnostus . 29 Calodiscus 30 Burlingia 32 Paradoxides 33 Ontogeny of Paradoxides 45 Ellipsocephalus 56 Bailiella . ss Solenopleura . 59 Phyllocarida. Hymenocaris (?) 6r Geographical and Stratigraphical Distribution of the F auna E lements 62 Bibliography . 64 Explanation of Flates . .•. 67 I. A Diamond Boring through the Cambrian at Mossberga. As a part of the work carried out by the Electrical Prospecting Co. (A.-B. Elektrisk Malmletning) in order to search for gas in the Lower Cambrian of Öland, a deep boring was made in the autumn of 1933 into a little flat dome at Mossberga, about 12 km S of Borgholm. The Company had the courtesy to present the core to the Geological Survey, and thus the present writer had the opportunity of subjecting it to a doser investigation, the results of which are given below. -
Early Cambrian Trilobite Faunas and Correlations, Labrador Group, Southern Labrador—Western Newfoundland
EARLY CAMBRIAN TRILOBITE FAUNAS AND CORRELATIONS, LABRADOR GROUP, SOUTHERN LABRADOR—WESTERN NEWFOUNDLAND Douglas Boyce1, Ian Knight1, Christian Skovsted2 and Uwe Balthasar3 1 - GSNL; 2 - Swedish Museum of Natural History; 3 – University of Plymouth, England. Paleontological, lithological and mapping studies of mixed siliciclastic‒carbonate rocks of the Labrador Group have been ongoing, since 1976, by the GSNL. Recent systematic litho- and bio-stratigraphic studies with Drs. Skovsted and Balthasar focus on trilobite and small shelly fossils (SSF), mostly in the Forteau Formation. At least 34 co-eval sections have been measured, and 434 fossiliferous samples have been collected; publication of trilobite and SSF systematics is in preparation. Dyeran to Topazan biostratigraphy, southern Labrador and western Newfoundland. Early Cambrian and early Middle Cambrian global and Laurentian series and stages (Hollingsworth, 2011). Distribution of the Trilobite Faunas Early Cambrian trilobite faunas occur throughout the Forteau Formation and in the lowest strata of the overlying Hawke Bay Formation. They divide into two broad faunas – Olenelloidea, mostly occur in deep-water shale and mudrocks, and Corynexochida, are hosted in shallow-water limestone, including archeocyathid reefs. The Devils Cove member, basal Forteau Formation - a regionally widespread pink Bonnia sp. nov. Boyce: latex replica of limestone - hosts Calodiscus lobatus (Hall, 1847), Elliptocephala logani (Walcott, partial, highly ornamented, spinose 1910), and Labradoria misera (Billings, 1861a). Calodiscus lobatus and E. logani range cranidium, Route 432, Great Northern Zacanthopsis sp. A Boyce: latex replica of Peninsula (GNP); it also occurs in the Deer high in the formation regionally, but L. misera is restricted to the lower 20 m of the partial cranidium, Mackenzie Mill member, Arm limestone, Mackenzie Mill member, formation in Labrador (includes archeocyathid reefs). -
Upper Lower Cambrian (Provisional Cambrian Series 2) Trilobites from Northwestern Gansu Province, China
Estonian Journal of Earth Sciences, 2014, 63, 3, 123–143 doi: 10.3176/earth.2014.12 Upper lower Cambrian (provisional Cambrian Series 2) trilobites from northwestern Gansu Province, China a b c c Jan Bergström , Zhou Zhiqiang , Per Ahlberg and Niklas Axheimer a Department of Palaeozoology, Swedish Museum of Natural History, P.O. Box 5007, SE-104 05 Stockholm, Sweden b Xi’an Institute of Geology and Mineral Resources, 438 East You Yi Road, Xi’an 710054, Peoples Republic of China; [email protected] c Department of Geology, Lund University, Sölvegatan 12, SE-223 62 Lund, Sweden; [email protected], [email protected] Received 7 March 2014, accepted 24 June 2014 Abstract. Upper lower Cambrian (provisional Cambrian Series 2) trilobites are described from three sections through the Shuangyingshan Formation in the Beishan area, northwestern Gansu Province, China. The trilobite fauna is dominated by eodiscoid and ‘corynexochid’ trilobites, together representing at least ten genera: Serrodiscus, Tannudiscus, Calodiscus, Pagetides, Kootenia, Edelsteinaspis, Ptarmiganoides?, Politinella, Dinesus and Subeia. Eleven species are described, of which seven are identified with previously described taxa and four described under open nomenclature. The composition of the fauna suggests biogeographic affinity with Siberian rather than Gondwanan trilobite faunas, and the Cambrian Series 2 faunas described herein and from elsewhere in northwestern China seem to be indicative of the marginal areas of the Siberian palaeocontinent. This suggests that the Middle Tianshan–Beishan Terrane may have been located fairly close to Siberia during middle–late Cambrian Epoch 2. Key words: Trilobita, taxonomy, palaeobiogeography, lower Cambrian, Cambrian Series 2, Beishan, Gansu Province, China. -
Small Shelly Fossils from the Argillaceous Facies of the Lower Cambrian Forteau Formation of Western Newfoundland
Small shelly fossils from the argillaceous facies of the Lower Cambrian Forteau Formation of western Newfoundland CHRISTIAN B. SKOVSTED and JOHN S. PEEL Skovsted, C.B. and Peel, J.S. 2007. Small shelly fossils from the argillaceous facies of the Lower Cambrian Forteau For− mation of western Newfoundland. Acta Palaeontologica Polonica 52 (4): 729–748. A diverse fauna of helcionelloid molluscs, hyoliths, and other small shelly fossils is described from limestone layers within the Forteau Formation of the Bonne Bay region in western Newfoundland. The fauna is dominated by internal moulds of various molluscs and tubular problematica, but also includes hyolith opercula, echinoderm ossicles, and other calcareous small shelly fossils preserved by phosphatisation. Originally organophosphatic shells are comparatively rare, but are represented by brachiopods, hyolithelminths, and tommotiids. The fauna is similar to other late Early Cambrian faunas from slope and outer shelf settings along the eastern margin of Laurentia and may be of middle Dyeran age. The similarity of these faunas indicates that at least by the late Early Cambrian, a distinctive and laterally continuous outer shelf fauna had evolved. The Forteau Formation also shares elements with faunas from other Early Cambrian provinces, strengthening ties between Laurentia and Australia, China, and Europe during the late Early Cambrian. Two new taxa of problematic fossil organisms are described, the conical Clavitella curvata gen. et sp. nov. and the wedge−shaped Sphenopteron boomerang gen. et sp. nov. Key words: Helcionellidae, Hyolitha, Brachiopoda, small shelly fossils, Cambrian, Laurentia, Newfoundland. Christian B. Skovsted [[email protected]], Centre for Ecostratigraphy and Palaeobiology, Macquarie University, NSW 2109, Marsfield, Sydney, Australia. -
The Early Cambrian Fauna of North-East Greenland
Christian Skovsted The Early Cambrian fauna of North-East Greenland Dissertation presented at Uppsala University to be publicly examined in Lecture Theatre, Paleontology building, Uppsala, Friday, January 16, 2004 at 13.00 for the degree of Doctor of Philosphy. The examination will be conducted in English. Abstract Skovsted, C.B. 2003. The Early Cambrian fauna of North-East Greenland. 22 pp. Uppsala. ISBN 91-506-1731-1 Small shelly fossils are common in sediments of Early Cambrian age, and include the earliest common representatives of metazoan animals with mineralized hard parts. The group include fossils of very different morphology, composition and ultrastructure, presumably representing skeletal remains of numerous animal groups, the biological affinity of which is sometimes unresolved. Although the nature of many small shelly fossils is obscure, the wide geographical range of many forms, yields a potential for enhancing biostratigraphic and palaeogeographic resolution in the Early Cambrian. Small shelly fossils have been studied extensively in many parts of the world, but our knowledge of their occurrence in Laurentia is still limited. The late Early Cambrian sequence of North-East Greenland has yielded a well preserved small shelly fossil assemblage of more than 88 species, representing a diversity which is unparalleled in Laurentian strata. The composition of the fauna, which also includes brachiopods and trilobites, is indicative of a middle Dyeran (Botoman equivalent) age. The recovered fossils include a number of species that are known previously from other Early Cambrian palaeocontinents, and particularly strong ties to late Early Cambrian faunas of Australia are documented. Among the widespread taxa are species belonging to very different animal groups such as brachiopods, molluscs, hyoliths, sponges, coeloscleritophorans, eodiscid trilobites, bivalved arthropods and problematic fossils. -
Cambrian Trilobites of East-Central Alaska
Cambrian Trilobites of East-Central Alaska GEOLOGICAL SURVEY PROFESSIONAL PAPER 559-B Cambrian Trilobites of East-Central Alaska By ALLISON R. PALMER LOWER PALEOZOIC PALEONTOLOGY AND STRATIGRAPHY OF EAST-CENTRAL ALASKA GEOLOGICAL SURVEY PROFESSIONAL PAPER 559-B Biostratigraphy and geological history of Cambrian deposits of east-central Alaska; 12 J species representing <?/ genera of trilobites of Cambrian age are described UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1968 UNITED STATES DEPARTMENT OF THE INTERIOR STEWART L. UDALL, Secretary GEOLOGICAL SURVEY William T. Pecora, Director Library of Congress catalog^card No. G<S 67-27i5 For sale by the Superintendent of Documents, U.S. Government Printing Office Washington, D.C. 20402 - Price $2 (paper cover) CONTENTS Page Page Abstract ___________________________________________ Bl Geological history of the Cambrian of eastern Alaska.___ B17 Introduction_______________________________________ 2 Problems for future investigation ______ 20 Acknowledgments.- _ ________________________________ 4 Systematic paleontology______________ 20 23 General stratigraphy and principal collecting areas______ 4 Agnostida______-___ _____________ Redlichiida _____________________ 38 Composition and correlation of the faunas _____________ 5 Corynexochida __________________ 41 Early Cambrian________________________________ 5 Pty chopariida __________________ 51 Middle Cambrian_______________________________ 8 Unassigned trilobites_____________ 82 Late Cambrian_--_-___________________-_--__.__ 10 Indeterminate or undescribed taxa. 104 Biogeographic significance of the Alaskan Cambrian References cited.____________________ 105 trilobites_ _ __-_----__-___-____-___-__________.__ 16 Index ___--_-_-____________-___.-__. 111 ILLUSTRATIONS fPlates 1-15 follow index; plate 16 is in pocket] PLATE 1, 2. Early Cambrian-1 fauna. 3. Early Cambrian-2 fauna. 4. Early Cambrian-3 fauna; Middle Cambrian-1 fauna. 5. Middle Cambian-1 fauna. -
Trilobite Evolutionary Rates Constrain the Duration of the Cambrian Explosion
Trilobite evolutionary rates constrain the duration of the Cambrian explosion John R. Patersona,1, Gregory D. Edgecombeb, and Michael S. Y. Leec,d aPalaeoscience Research Centre, School of Environmental & Rural Science, University of New England, Armidale, NSW 2351, Australia; bDepartment of Earth Sciences, The Natural History Museum, London SW7 5BD, United Kingdom; cCollege of Science and Engineering, Flinders University, SA 5001, Australia; and dEarth Sciences Section, South Australian Museum, Adelaide, SA 5000, Australia Edited by Andrew H. Knoll, Harvard University, Cambridge, MA, and approved January 9, 2019 (received for review November 12, 2018) Trilobites are often considered exemplary for understanding the phenotypic and genomic evolution (7, 8). Rapid morphological Cambrian explosion of animal life, due to their unsurpassed di- and molecular evolution during the earliest Cambrian almost versity and abundance. These biomineralized arthropods appear certainly underpinned the pronounced pulses of origination and abruptly in the fossil record with an established diversity, phyloge- diversification throughout the Terreneuvian (3, 9, 10). However, netic disparity, and provincialism at the beginning of Cambrian the question remains as to when evolutionary rates slowed to Series 2 (∼521 Ma), suggesting a protracted but cryptic earlier his- Phanerozoic norms, thus marking the end of the Cambrian ex- tory that possibly extends into the Precambrian. However, recent plosion. For instance, the calibrations used in ref. 7 were mostly analyses indicate elevated rates of phenotypic and genomic evolu- 488 Ma or younger; that analysis therefore only had weak power tion for arthropods during the early Cambrian, thereby shortening to constrain fast early rates further back than that time point.