4 Keiichi Kakui Bull. Kitakyushu Mus. Nat. Hist. Hum. Hist., Ser. A, 18: 5–42, March 31, 2020 5 dation in the water column. arboreal life in oribatid mites indicates the primacy of larvae and juveniles in marine soft-bottom communities. List of the Adoliadini (, ) ecology. Proceedings of the Royal Society B, 276: Marine Ecology Progress Series, 7: 179–184. of the Philippine Islands (Part 1) 3219–3227. Puitika, T., Kasahara, Y., Miyoshi, N., Sato, Y. and Shimano, ACKNOWLEDGMENTS Müller, C. H. G., Harzsch, S. and Perez, Y. 2018. Chaetognatha. S. 2007. A taxon-specific oligonucleotide primer set for In: Schmidt-Rhaesa, A. (ed.), Miscellaneous Invertebrates, PCR-based detection of soil ciliate. Microbes and Takashi Yokochi1 and Fidel Bendanillo2 I thank Taeko Kimura for providing the specimen used in De Gruyter, Berlin, pp. 163–282. Environments, 22: 78–81. this study; Captain Yoichi Maekawa and the crew of TR/V Nakayama, T., Watanabe, S., Mitsui, K., Uchida, H. and Inoue, Shaffer, P. L. 1979. The feeding biology of Podarke pugettensis Seisui-maru, and researchers aboard, for support during the I. 1996. The phylogenetic relationship between the (Polychaeta: Hesionidae). Biological Bulletin, 156: 1 Asian Research Society (AIRS). 1-10-26, Shonan, Owariasahi, Aichi, 488-0823, Japan. cruise in 2019; Matthew H. Dick for reviewing the manuscript Chlamydomonadales and Chlorococcales inferred from 343–355. E-mail: [email protected] 2 and editing the English; and Taichiro Goto and one anonymous 18SrDNA sequence data. Phycological Research, 44: Tomioka, S., Kondoh, T., Sato-Okoshi, W., Ito, K., Kakui, K. University of San Carlos Biological Museum. Learning Resource Center, reviewer for critical comments on the manuscript. This study 47–55. and Kajihara, H. 2016. Cosmopolitan or cryptic ? Josef Baumgartner Bldg. Talamban Campus, , Republic of . E-mail: [email protected] was supported in part by a KAKENHI grant (JP19K06800) Nomura, H., Aihara, K. and Ishimaru, T. 2007. Feeding of A case study of Capitella teleta (Annelida: Capitellidae). from the Japan Society for the Promotion of Science (JSPS). the chaetognath Sagitta crassa Tokioka in heavily Zoological Science, 33: 545–554. eutrophicated Tokyo Bay, Japan. Plankton & Benthos Tonnesson, K. and Tiselius, P. 2005. Diet of the chaetognaths (Received June 30, 2019; accepted September 6, 2019) Research, 2: 120–127. Sagitta setosa and S. elegans in relation to prey abundance REFERENCES Oliver, J. S., Oakden, J. M. and Slattery, P. N. 1982. and vertical distribution. Marine Ecology Progress Series, ABSTRACT − The type-specimens of one-hundred and twenty taxa of the tribe Adoliadini from Philippines Phoxocephalid amphipod crustaceans as predators on 289: 177–190. are examined and the classification of species-subspecies relations is revised. With information of the type Altschul, S. F., Gish, W., Miller, W., Myers, E. W. and locality and the current repository of the type or syntypes all of the taxonomic names are shown alphabetically in Lipman, D. J. 1990. Basic local alignment search tool. part 1. The annotated synonymic list is present in part 2. Journal of Molecular Biology, 215: 403–410. RESULTS AND DISCUSSION Błażewicz-Paszkowycz, M., Jenning, R. M., Jeskulke, K. and Brix, S. 2014. Discovery of swimming males of KEY WORDS: Lepidoptera, Rhopalocera, Nymphalidae, Adoliadini, Philippines, , Cynthia, Dophla, The tanaidacean in the chaetognath gut (Fig. 1) was very Paratanaoidea (Tanaidacea). Polish Polar Research, 35: , Lexias, , aconthea, adonia, albusequus, alpheda, amlana, anosia, antiquea, aquamarina, aruna, INTRODUCTION MATERIALS AND METHODS fragile, and most of its appendages were broken or lost during 415–453. balabacana, balabacensis, boholensis, bongaoensis, borromeoi, cabigoni, calliphorus, candida, canescens, dissection, but I observed that it has 1) a ring-shaped pereonite Feigenbaum, D. L. and Maris, R. C. 1984. Feeding in the cavarna, chloe, circe, cocytina, cocytus, cusama, dacasini, damalis, darani, dhayma, dinagatensis, dinorah, dirtea, Tanaidaceans are typically benthic, free-living crusta- The chaetognath was collected on 23 April 2019 during 1 (p1 in Fig. 1); 2) the maxillipedal endites partly fused; 3) a Chaetognatha. Oceanography and Marine Biology, An djata, dodong, dohertyi, dumaranensis, dunya, ellora, eson, eva, evelina, exul, felicitacionae, gahiti, galoa, ceans; most are small, up to a few millimeters long. They are the cruise 1903 by TR/V Seisui-maru, at Station 3B in Tanabe ventrodistal blade-like seta on a pereopod (not pereopod 1, but Annual Review, 22: 343–392. garuda, godartii, goertzi, hikarugenzi, hinunanganensis, hiwaga, howarthi, imperator, ingae, irauana, jama, highly abundant in shallow-water and deep-sea bottom habi- Bay (33°32.66'N 135°7.2'E to 33°33.67'N 135°6.07'E; 646–752 one of pereopods 2–6); 4) a chelipedal fixed finger with one Folmer, O., Black, M., Hoeh, W., Lutz, R. and Vrijenhoek, R. joloana, kayumanggia, kulaya, laetitiae, leopardina, leucotaenia, leytana, leytensis, liaoi, lubentina, ludonia, tats, and play an important role in ecosystems (Larsen et al., m depth), in one of two small plankton nets attached inside a ventral simple seta; and 5) a developing oostegite (part of the 1994. DNA primers for amplification of mitochondrial 2015). Species in various groups have been reported as larger beam trawl; the sample from Station 3B contained many brood pouch). Although identification to species was not cytochrome c oxidase subunit I from diverse metazoan lupina, lusiada, macer, mahadeva, malissia, marinduque, medaga, merta, mindanaensis, mindorana, mindorensis, their consumers, including crustaceans, polychaetes, sea anem- other chaetognaths and tanaidaceans. The specimen was possible, these features allowed identification of the female invertebrates. Molecular Marine Biology and Biotechnology, miscus, monara, monina, nadenya, negrosiana, nirodha, obatrata, ormocana, ornata, palawana, pallida, ones, wading birds, and fishes (Shaffer, 1979; Oliver et al., photographed and then fixed and preserved in 80% ethanol. individual as Pseudotanais sp., in the family Pseudotanaidae. I 3: 294–299. panayana, panopus, pardalis, pelea, phelada, philippensis, phlegethon, pinamalayana, piratica, placida, princesa, 1982; Larsen, 2005). The tanaidacean was extracted from the gut of the chaetognath morphologically identified the chaetognath specimen as Heard, R. W. and Anderson, G. 2009. Tanaidacea (Crustacea) proditrix, producta, ramada, rhamases, rodriguezi, romeo, rudraca, saidja, samarensis, sarmana, satrapes, Planktonic arrow worms (phylum Chaetognatha) are by using sharpened needles, mounted on a glass slide in belonging to the family Sagittidae, one of planktonic families of the Gulf of Mexico. In: Felder, D. L. and Camp, D. K. schoenigi, semperi, senilis, sibutana, sibuyana, smaragdifera, soloni, soregina, subpiratica, suluana, susoni, strictly carnivorous and occupy an important position in the glycerin, and observed with an Olympus BX51 microscope. (Müller et al., 2018); this was supported by the BLAST result (eds.), Gulf of Mexico Origin, Waters, and Biota, Volume tanagra, tawitawia, tethys, teuta, trapesa, treadawayi, tsukada, tyawena, uposatha, virginalis, visayana, vistrica, marine food web (Feigenbaum and Maris, 1984). As predators, The chaetognath and tanaidacean were subsequently deposited of its 18S sequence (1620 nt long; INSD accession number 1, Biodiversity, Texas A&M University Press, College volupia, waltraudae, yoshiroui, yui, holotype, paratype, syntype, lectotype, homonym, synonym, . they consume mainly copepods but are not copepod specialists, in the Kitakyushu Museum of Natural History & Human LC494513) which determined a sagittid sequence in INSD as Station, pp. 987–1000. and various other organisms have also been reported from their History (KMNH IvR 600,001 and 600,002, respectively). the most similar to the present sequence (the COI sequence Kehayias, G., Lykakis, J. and Fragopoulu, N. 1996. The diets gut contents, including chaetognaths, polychaete larvae, roti- The tanaidacean was in too poor condition to obtain tissue could not be amplified). of the chaetognaths Sagitta enflata, S. serratodentata fers, cladocerans, cirripedian nauplii, ostracods, amphipods, for DNA extraction. Total DNA was extracted from a piece of The chaetognath I observed was collected in one of two atlantica and S. bipunctata at different seasons in Eastern INDEX decapod larvae, euphausiids, appendicularians, echinoderm the chaetognath’s body muscle by using a NucleoSpin Tissue small plankton nets attached inside a larger beam trawl. Both Mediterranean coastal waters. ICES Journal of Marine larvae, ciliates, diatoms, dinoflagellates, infusorians, and fish XS kit (TaKaRa Bio, Japan). Attempts were made to amplify nets had picked up bottom sediments (Fig. 2). As female Science, 53: 837–846. (PART 1) larvae and postlarvae (e.g., Feigenbaum and Maris, 1984; part of the cytochrome c oxidase subunit I (COI) and 18S pseudotanaids are typically benthic tube-dwellers (Heard and Larsen, K. 2005. Deep-sea Tanaidacea (Peracarida) from the 1. INTRODUCTION Kehayias et al., 1996; Tonnesson and Tiselius, 2005; Nomura rRNA (18S) genes by PCR using the primers listed in Table 1. Anderson, 2009), whereas sagittid chaetognaths are planktonic, Gulf of Mexico. Brill, Leiden. 2. ABBREVIATIONS et al., 2007). PCR amplification conditions for COI with TaKaRa Ex Taq this case of chaetognath predation on a tanaidacean may have Larsen, K., Guţu, M. and Sieg, J. 2015. Order Tanaidacea Dana, 3. LIST OF DESCRIBED SPECIES AND SUBSPECIES NAME During a cruise of the TR/V Seisui-maru (Mie University, DNA polymerase (TaKaRa Bio) were 94°C for 1 min; 35 cycles resulted from cod-end feeding in the net. 1849. In: Vaupel Klein, J. C. von, Charmantier-Daures, M. Japan) in 2019, one chaetognath individual containing a of 98°C for 10 sec, 50°C for 30 sec, and 72°C for 50 sec; and 72°C Although chaetognaths are probably uncommon preda- and Schram, F. R. (eds.), The Crustacea. Revised and (PART 2) tanaidacean in its gut was collected. As tanaidaceans had not for 2 min. Conditions for 18S amplification with KOD FX Neo tors on tanaidaceans as they usually occupy different oceanic Updated, as well as Extended from the Traité de Zoologie 4. DESCRIPTIONS OF SPECIES AND SUBSPECIES been previously noted among chaetognath prey items, I report polymerase (Toyobo, Japan) were 94°C for 2 min; 45 cycles of were as described by Tomioka et al. (2016). The nucleotide of Japan (DDBJ). BLAST searches (Altschul et al., 1990) zones, species in several tanaidacean families produce 5, Brill, Leiden, pp. 249–329. 5. DISCUSSION this observation here. 98°C for 10 sec, 65°C for 30 sec, and 68°C for 1 min; and 68°C sequence obtained was deposited in the International Nucle- were used to determine the nucleotide sequences in the INSD non-benthic, swimming males (Błażewicz-Paszkowycz et al., Maraun, M., Erdmann, G., Schulz, G., Norton, R. A., Scheu, S. 6. ACKNOWLEDGMENTS for 2 min. Methods for sequencing and sequence assembly otide Sequence Database (INSD) through the DNA Data Bank most similar to the sequence obtained. 2014), and these males would be exposed to chaetognath pre- and Domes, K. 2009. Multiple convergent evolution of 7. REFERENCES 6 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 7

1. INTRODUCTION Limbach, about 250 km from . It is an undoubted fact amlana Jumalon, 1970 (Fig. 3) cabigoni Schröder, Treadaway, A. Mohagan & N. Mohagan, dhayma Fruhstorfer, 1913 (Fig. 21) exul Tsukada & Nishiyama, 1980 (Fig. 31) ingae (Euthalia) Schröder & Treadaway, 1990 (Fig. 42) leytensis Jumalon, 1975 (Fig. 51) TS (TP): ST 2♂1♀ (BMNH). ormocana Jumalon, 1970 (Fig. 69) 11.11.81 / 110 / Coll. C. Semper / Lectotype ♀ Euthalia TS (TP): LT ♂ (ZMHU). sibutana Schröder & Treadaway, 1990 (Fig. 95) Note: Attached label to the holotype specimen shows TS (TP): HT ♂ (SMFL), PT 10♂2♀. waltraudae Schröder & Treadaway, 1990 (Fig. 114) that Mr. Treadaway and Dr. H. Schröder of Senckenberg TL: Amlan, Negros Oriental. 2009 (Fig. 12) TL: Archipelago. TL: Bo. Melgar, Dinagat. TL: Peak, 200 m, Bongao, Tawitawi Group. TL: Lake Danao, Ormoc , Leyte. Note: The figured female (Fig. 61) is preserved in the TL: Lake Danao, Ormoc City, Leyte. phelada Semper [87]:1888 Designated by T. Yokochi, 1997”). Note: Description is based on males, and four males are TL: Cavan Cavan, 5 m, Sibutu, Tawitawi Group. “Balabac”, but the correct type locality is . TL: Indalawan, Balabac. The Philippine Islands are situated in the northern end of thoroughly deserve credit for working further on systematic TS (TP): HT ♂ (USC), PT10 ♂1♀ (USC, JF). TL: Balut Is, Sarangani Group. TS (TP): HT ♀ (BMNH). TS (TP): HT ♂ (ET), PT 3♂6♀. TS (TP): HT ♀ (SMFL), PT 10♂12♀. TS (TP): HT ♀ (USC). Rothschild collection (Ro 98/7), BMNH. It might be one of the TS (TP): HT ♂ (USC), PT 1♀. preserved in ZMHU. The lectotype designation was carried out TS (TP): HT ♂ (SMFL), PT 17♂15♀. vistrica Fruhstorfer, 1913 (Fig. 112) TS (TP): HT ♀ (SMFL), PT 2♂2♀. the Malay Archipelago and are composed of more than 7,000 classification in Philippines Rhopalocera. TS (TP): HT ♂ (SMFL), PT 2♂1♀. Note: Though the original description does not mention the types, although the attached label is only “”. No other philippensis Fruhstorfer, 1899 (Fig. 78) by Yokochi (1999). trapesa G. Semper, [1887] (Fig. 106) TL: locality unknown, presumably one of the Central islands, large and small, extending from north to south about Here we revised the group of the tribe Adoliadini antiquea Schröder & Treadaway, 1980 (Fig. 4) number of type series, one female would be the holotype. felicitacionae Jumalon, 1989 (Fig. 32) irauana Jumalon, 1975 (Fig. 43) liaoi Schröder & Treadaway, 1982 (Fig. 52) specimen is found that corresponded with the type series. ornata Schröder & Treadaway, 1979 (Fig. 70) TL: . sibuyana Schröder & Treadaway, 1982 (Fig. 96) TL: Camiguin de . Philippine islands [recte Dinagat]. yoshiroui K. Okano, 1992 (Fig. 115) 1,800 km and from east to west about 1,100 km. The distributed in the Philippines, based on the checklist by TL: Culasi, 500 m, Antique, Panay. calliphorus C. & R. Felder, 1861 (Fig. 13) TL: Kantakpal Creek, Awihao, Ormoc, Western Leyte. TL: Irauan [Irawan], Puerto Princesa, Palawan. TL: Mt. Madja-an [Mt. Madja-as], 500 m, Antique, Panay. TL: Romblon. TS (TP): LT ♂ (MNHN). rodriguezi Schröder & Treadaway, 1982 (Fig. 86) TL: Sibuyan, Romblon Group. TS (TP): LT ♂ (SMFL). TS (TP): ST ♂ (BMNH). TL: Babuyan. fauna is rich in diversity and the tribe Adoliadini, Nymphalidae, Treadaway (1995). One hundred-twenty taxa of type specimens TS (TP): HT ♂ (SMFL). TL: . dinagatensis Tsukada, 1991 (Fig. 22) TS (TP): HT ♂ (JF). TS (TP): HT ♀ (JF), PT 1♂1♀ (USC). TS (TP): HT ♂ (SMFL), PT 1♂2♀. mindorana (Euthalia) Fruhstorfer, 1899 (Fig. 62) TS (TP): HT ♂ (SMFL), PT 1♂2♀. Note: Description is based on one male from Basilan TL: 28 km W of Puerto Princesa, Palawan. TS (TP): HT ♂ (SMFL), PT 2♂5♀. Note: Description is based on seven materials from E. Note: Tsukada (1991) suspected this taxon is distributed TS (TP): HT ♂ (KMNH). Lepidoptera, the main group of this study, is no exception. The founded in the Philippines are examined and revised the TS (TP): ST ♂ (BMNH). TL: Dinagat. Note: The holotype is deposited in Jumalon family collec- TL: Mindoro. (MNHN) and one female from Luzon (ZMHU). As the TS (TP): HT ♂ (SMFL), PT 2♂2♀. Mindanao. The lectotype designation was carried out for the in Basilan, but it presumably comes from Dinagat. important works concerned with Philippines were, back in classification of species-subspecies relation. This paper is aquamarina Fruhstorfer, 1913 (Fig. 5) TS (TP): HT ♂ (ET), PT 2♂1♀. gahiti M. & T. Okano, 1990 (unavailable name) (Fig. 33) tion. ludonia Staudinger, 1889 (Fig. 53) TS (TP): ST (MNHN). palawana (Lexias) Moore, 1897 (Fig. 71) Luzon-group should be classified as nadenya. We select a smaragdifera Fruhstorfer, 1913 (Fig. 97) Camiguin de Mindanao specimen by Yokochi (1998). yui Yokochi, 1994 (Fig. 116) history, “Diagnosen einiger neuer Tagfalter von den Philippinen published in two parts. All the taxa described before are TL: Mindanao. candida Schröder, Treadaway & Nuyda, 1990 (Fig. 14) TL: Bilar, Bohol. TL: Palawan. TL: Palawan. male (Fig. 78) deposited in MNHN as the lectotype, labelled, romeo Schröder & Treadaway, 1987 (Fig. 87) TL: Locality unknown, presumably Mindoro [Mindoro]. volupia Tsukada & Nishiyama, 1981 (Fig. 113) TL: Dumaran. und Bemerkungen über die Verbreitung einiger derselben” (G. examined and figured as much as possible in part 1. In the TS (TP): LT ♂ (SMFL). TL: 30 m NN, Calamian Group. dinorah Fruhstorfer, 1899 (Fig. 23) TS (TP): HT ♀ (FASU). joloana Staudinger, 1889 (Fig. 44) TS (TP): LT ♂ (ZMHU). mindorana (Euthalia) Tsukada, 1991 (permanently invalid) TS (TP): HT ♂ (BMNH). “Type (red) / lubentina philippensis Fruhst. / Philippinen TL: Mt. Halcon, N. Mindoro. TS (TP): ST (MNHN). treadawayi Tsukada, 1991 (Fig. 107) TL: Camiguin. TS (TP): HT ♂ (TY), PT 11♂4♀. Semper, 1886–1892), and then “Die Gross-Schmetterlinge der second part, synonymic lists and some important specimens Note: Description is based on some specimens. The lecto- TS (TP): HT ♂ (SMFL), PT10 ♂. TL: Basilan. Note: As this taxon gahiti was described as a form of TL: Sulu. Note: Description is based on four males and six females, Note: The taxon mindorana described under the Bazilan II-III, 98 Doherty ex. coll. H. Fruhstorfer / TS (TP): HT ♀ (SMFL), PT 1♀. Note: Though the original description does not mention TL: Romblon, Sibuyanes. TS (TP): HT ♂ (ET), PT 2♂5♀. (to be continued) Erde, 9” (1908–1927) edited by A. Seitz. Recently, a Japanese will be shown, affixing several discussions. type designation was carried out by Yokochi (1998). TS (TP): ST 2♂2♀ (MNHN). Lexias panopus boholensis (infrasubspecific name), it should TS (TP): LT ♂ (ZMHU). and two pairs are preserved in ZMHU. The lectotype designa- Euthalia is a junior primary homonym of Euthalia lusiada palawana (Tanaecia) Staudinger, 1889 (Fig. 72) MUSEUM PARIS 1934 COLL H Fruhstorfer”. the number and sex, a syntype specimen in MNHN is a male. TS (TP): HT ♂ (SMFL). entomologist, E. Tsukada, had compiled a great series of cavarna Fruhstorfer, 1913 (Fig. 15) be treated as an unavailable name. Note: Description is based on two males and one female, tion was carried out by Yokochi (1999). mindorana Fruhstorfer, 1899 (permanently invalid name). TL: Palawan. rudraca Fruhstorfer, 1913 (Fig. 88) “ of the South East Asian Islands” (1980–1991). balabacana (Bassarona) Tsukada, 1991 (Fig. 6) TL: Balabas [Balabac]. dodong Schröder & Treadaway, 1978 (Fig. 24) and all of them are preserved in ZMHU. The lectotype desig- Now it is replaced by tsukada Koçak (1996). TS (TP): LT ♂ (ZMHU). phlegethon G. Semper, [1887] (Fig. 79) TL: Balabac. soloni M. & T. Okano, 1990 (Fig. 98) tsukada Koçak, 1996 (Fig. 80) The late Prof. Julian N. Jumalon (1909–2000) (Fig. 1, TL: Balabac. TS (TP): LT ♀ (ZMHU). TL: Mobo, 1000 ft, Masbate. galoa Fruhstorfer, 1913 (Fig. 34) nation was carried out by Yokochi (1999). lupina Druce, 1874 (Fig. 54) Note: Description is based on males and females, and six TL: Mindanao. TS (TP): LT ♂ (ZMHU). TL: Bilar, Bohol. TL: Mindoro. left), the Lepidoptera Research Associate and Lecturer of 2. ABBREVIATIONS TS (TP): HT ♂ (ET), PT 8♂6♀. Note: Description is based on a few females, and one spec- TS (TP): HT ♂ (SMFL), PT 3♂7♀. TL: Mindoro. TL: Philippine Islands [Sulu]. mindorensis Schröder & Treadaway, 1982 (Fig. 63) males and three females are preserved in ZMHU. The lecto- TS (TP): LT ♀ (SMFL). Note: Description is based on some specimens, and two TS (TP): HT ♂ (FASU), PT 2♂3♀. TS (TP): HT ♂ (ET), PT 10♂11♀. Department of Biology, University of San Carlos, published a imen is preserved in ZMHU. The lectotype designation was TS (TP): ST (BMNH). kayumanggia Treadaway & Nuyda, 1994 (Fig. 45) TS (TP): HT ♀ (BMNH). TL: Mindoro. type designation was carried out by Yokochi (1999). Note: Description is based on twenty-one males and seven males and two females are preserved in ZMHU. The lectotype Note: mindorana Tsukada, 1991 is a paper of “New Philippine butterflies” in 1975, and many new Public Institution (Museum, University). BMNH: The balabacana (Dophla) Tsukada, 1991 (secondary homonym) carried out by Yokochi (1999). dohertyi Butler, 1901 (Fig. 25) Note: Though the original description does not mention TL: Mt. Bayog, S. Palawan. TS (TP): HT ♂ (SMFL), PT 6♂7♀. females from Samar, Camiguin de Mindanao, N. Mindanao, E. designation was carried out by Yokochi (1999). soregina Fruhstorfer, 1913 (Fig. 99) permanently invalid name (nom. nov. pro mindorana Tsukada, taxonomic names concerned with various families were Natural History Museum, London, United Kingdom; FASU: (Fig. 7) TL: Sulu Archipelago. the number and sex, syntype specimen in BMNH is a male. TS (TP): HT ♂ (SMFL), PT 13♂4♀. lusiada C. & R. Felder, 1863 (Fig. 55) palawana (Euthalia) Staudinger, 1889 (Fig. 73) Mindanao, SE. Mindanao, and several materials are preserved TL: Sulu Archipelago. 1991, non Fruhstorfer,1899). New name tsukada was proposed described there. He played a leading part of the study of Exhibition Hall of Food, Agricultural and Environmental Note: See below balabacensis. chloe Schröder & Treadaway, 1990 (Fig. 16) TS (TP): HT ♂ (BMNH). TL: Luzon. miscus Fruhstorfer, 1913 (Fig. 64) TL: Palawan. in SMFL. The lectotype designation was carried out for the samarensis Tsukada, 1991 (Fig. 89) TS (TP): ST ♀ (BMNH). by Koçak (1996). Philippine butterflies and contributed greatly to entomology. Sciences, Faculty of Agriculture, Shinshu University, Nagano, TL: Jao Asan, 200 m, Sibuyan, Romblon Group. goertzi Jumalon, 1975 (Fig. 35) kulaya Treadaway & Nuyda, 1994 (Fig. 46) TS (TP): ST ♂ (SMFL). TL: Mindanao. TS (TP): LT ♂ (ZMHU). Mindanao specimen by Yokochi (1998). TL: San Juan, Samar. Note: According to the original description, the type series In 1995, the late Mr. Colin G. Treadaway (Fig. 1, right, Japan; KMNH: Kitakyushu Museum of Natural History & balabacensis nom. nov. (Fig. 7) TS (TP): HT ♂ (SMFL), PT 11♂9♀. dumaranensis Hanafusa, 1992 (Fig. 26) TL: Amlan, Negros Oriental. TL: Magellanes, Homonhon. Note: Though the original description does not mention TS (TP): ST (MNHN, BMNH). Note: Description is based on one male and six females, TS (TP): HT ♂ (ET), PT 3♂2♀. are exactly female, but the number is unknown. tyawena Fruhstorfer, 1913 (Fig. 108) died in 2019) published “Checklist of the butterflies of the Human History, Fukuoka, Japan; MNHN: Museum National TL: Balabac. TL: Dumaran. TS (TP): HT ♀ (ET). TS (TP): HT ♂ (SMFL), PT 14♂10♀. the number of types, a male syntype is preserved in SMFL. Note: According to the original description, types and one male and four females are preserved in ZMHU. The pinamalayana Yokochi, 1997 (Fig. 80) TL: unknown [Palawan]. Philippine Islands (Lepidoptera: Rhopalocera)”. Although, 25 d'Histoire Naturelle Entomologie, Paris, ; SMFL: TS (TP): HT ♂ (ET), PT 2♂2♀. circe Schröder & Treadaway, 1990 (Fig. 17) TS (TP): HT ♂ (HH). Note: Though the original description mentions the Note: The name kulaya is spelt “kúlaya” in the original (syntypes) are preserved in Fruhstorfer collection (now in lectotype designation was carried out by Yokochi (1999). Note: The new name pinamalayana was proposed by sarmana Fruhstorfer, 1913 (Fig. 90) subpiratica Schröder & Treadaway, 1987 (Fig. 100) TS (TP): LT ♀ (ZMHU). years have passed since his paper presented, several new Senckenberg Forschungsinstitut und Naturmuseum Frankfurt, Note: The taxon balabacana was described under the TL: Cavan Cavan, Sibutu, Tawitawi Group. holotype is preserved in USC, it is deposited in ET. description. macer Tsukada & Nishiyama, 1980 (Fig. 56) MNHN) and BMNH. Yokochi (1997), because mindorana Tsukada, 1991 was a TL: Basilan. TL: Nueva Ecija, Bongabon, Centr. Luzon. Note: Description is based on some specimens, and one findings are still remained unpublished — it can be followed Frankfurt-am-Main, ; USC: University of San Carlos, genus Dophla. As Yokochi (2010) assigned both Dophla and TS (TP): HT ♂ (SMFL), PT 9♂10♀. dunya Doubleday, [1848] (Fig. 27) TL: Bo. Melgar, Dinagat. pallida Schröder, Treadaway & Nuyda, 1990 (Fig. 74) permanently invalid name. However, pinamalayana is a junior TS (TP): ST (MNHN). TS (TP): HT ♂ (SMFL), PT1♀. male and one female are preserved in ZMHU. The lectotype by his own handwriting on the glass surface of his specimen Cebu, Philippines; ZMHU: Zoologisches Museum, Humboldt Bassarona to the genus Euthalia, balabacana (Dophla) is a TL: Assam, India. hikarugenzi Tsukada & Nishiyama, 1980 (Fig. 36) laetitiae Schröder & Treadaway, 2000 (Fig. 47) TS (TP): HT ♂ (ET), PT 3♂2♀. monara Fruhstorfer, 1913 (Fig. 65) TL: 30 m NN, Calamian Group. synonym of tsukada Koçak (1996). Note: Though the original description does not mention designation was carried out by Yokochi (1999). cases —. The first author visited Senckenberg Museum in Universität, Berlin, Germany. junior secondary homonym of Bassarona teuta balabacana cusama Fruhstorfer, 1913 (Fig. 18) TS (TP): ST ♂ (BMNH). TL: near Baguio, N. Luzon. TL: Cavan Cavan, 5 m, Sibutu, Tawitawi Group. TL: N. . TS (TP): HT ♂ (SMFL), PT 11♂16♀. the number and sex, a syntype specimen in MNHN is a male. suluana Fruhstorfer, 1902 (Fig. 101) Frankfurt to research on the Treadaway collection on March Private collections. ET: Etsuzo Tsukada, Yamanashi, Tsukada, 1991. Here a new name balabacensis is replaced for TL: Mindanao, Camiguin de Mindanao. TS (TP): HT ♂ (MKN), PT 1♀ (MKN). TS (TP): HT ♂ (SMFL), PT 4♂. malissia Fruhstorfer, 1913 (Fig. 57) TS (TP): ST (MNHN). piratica G. Semper, [1887] (Fig. 81) TL: Sulu Archipelago. uposatha Fruhstorfer, 1913 (Fig. 109) 2019, and just then was informed that he had died several Japan; HH: Hiroto Hanafusa, Tottori, Japan; JF: Jumalon Dophla evelina balabacana Tsukada, 1991. TS (TP): ST ♂ (MNHN), ♀ (SMFL). ellora Fruhstorfer, 1898 (Fig. 28) TL: Basilan. panayana Schröder & Treadaway, 1980 (Fig. 75) TL: Camiguin de Mindanao. satrapes C. & R. Felder, 1861 (Fig. 91) TS (TP): HT ♀ (BMNH). TL: Banggi (Banguey), E. . weeks ago at his home in England. In the middle 1990’s, every Family, Cebu, Philippines; MKN: Misao Kaneko, Tokyo, Euthalia (Dophla) evelina balabacensis nom. nov. Note: Figured specimen is a syntype female in SMFL. TL: Mindoro. hinunanganensis M. & T. Okano, 1986 (Fig. 37) leopardina Fruhstorfer, 1913 (Fig. 48) TS (TP): ST♂ (MNHN). nadenya Fruhstorfer, 1913 TL: Culasi, 800 ft, Antique, Panay. TS (TP): LT ♂ (SMFL). TL: Luzon. TS (TP): LT ♂ (ZMHU). time the first author visited to Senckenberg, he came bringing Japan; SK: Shun-ichi Kawamura, Tokyo, Japan; STA: Shinji pro Dophla evelina balabacana Tsukada, 1991, non TS (TP): ST ♂ (MNHN). TL: Hinunangan, Leyte. TL: Sulu Islands. Note: Though the original description does not mention TL: Luzon. TS (TP): HT ♂ (SMFL), PT 4♂4♀. Note: Description is based on six materials from Luzon, TS (TP): ST ♂ (BMNH). susoni Jumalon, 1975 (Fig. 102) Note: According to the original description, the type series his described type specimens, with his wife driving from Taga, Tokyo, Japan; TY: Takashi Yokochi, Aichi, Japan. Bassarona teuta balabacana Tsukada, 1991 dacasini Hanafusa, 1990 (Fig. 19) Note: Though the original description does not mention TS (TP): HT ♂ (FASU), PT 1♂. TS (TP): HT ♀ (BMNH). the number of types, a male syntype is preserved in MNHN. TS (TP): HT ♀ [untraced, not examined]. Mindoro, Camiguin de Mindanao, N. Mindanao. The lectotype Note: According to the original description, the type series TL: Camp 7, Minglanilla, Cebu. (male and female, but the number is unknown) are deposited in Specimens concerned. FW: Length of Forewing; HT: TL: Balabac. the number and sex, a syntype specimen in MNHN is a male. Note: Though the original description indicates that the panopus C. & R. Felder, 1861 (Fig. 76) designation was carried out for the Camiguin de Mindanao (number is unknown) are deposited in Semper and Felder TS (TP): HT ♂ (USC), PT 9♂9♀ (USC, JF, SMFL, TY). Staudinger collection (ZMHU). The lectotype designation was Holotype; LT: Lectotype; PLT: Paralectotype; PT: Paratype; boholensis (Lexias) M. & T. Okano, 1988 (Fig. 8) TS (TP): HT ♂ (HH), PT 1♀. hiwaga Nuyda & Kawamura, 1989 (Fig. 38) leucotaenia G. Semper, 1878 (Fig. 49) marinduque K. Okano, 1992 (Fig. 58) type female should be preserved in ZMHU, the specimen TL: Luzon. specimen by Schröder & Treadaway (1987). collections. carried out by Yokochi (1999). TL: Type Locality; TP: Type Preservation; TS: Type Series; TL: Bilar, Bohol. eson de Nicéville, 1894 (Fig. 29) TL: Camiguin, Babuyan. TL: Camotes. TL: Marinduque. corresponding with nadenya could not be found there. No TS (TP): ST ♂ (BMNH). tanagra Staudinger, 1889 (Fig. 103) ST: Syntype. TS (TP): HT ♂ (FASU), PT 6♂2♀. damalis Erichson, 1834 (Fig. 117) TL: Palawan. TS (TP): HT ♂ (SK), PT 7♂4♀ (SK, ET). TS (TP): LT ♂ (SMFL). TS (TP): HT ♀ (KMNH), PT 1♂ (KMNH). figure is presented in the original paper. Note: The original description does not mention the princesa Fruhstorfer, 1899 (Fig. 82) schoenigi Schröder & Treadaway, 1978 (Fig. 92) TL: Palawan. virginalis Fruhstorfer, 1913 TL: Luzon. TS (TP): LT ♂ (SMFL), PLT 1♀. Note: This taxon was described based on sixty-nine number of type series. TL: Palawan. TL: Amlan, 600 m, Negros. TS (TP): LT ♂ (ZMHU). TL: locality not given. boholensis (Euthalia) M. & T. Okano, 1990 (Fig. 9) TS (TP): ST ♂ [untraced, not examined]. Note: Description is based on a single pair, and all howarthi Jumalon, 1975 (Fig. 39) materials from Camotes, Bohol, Leyte, Panaon, E. Mindanao, medaga Fruhstorfer, 1913 (Fig. 59) negrosiana Schröder & Treadaway, 1987 (Fig. 66) TS (TP): HT ♀ (MNHN). TS (TP): HT ♀ (SMFL). Note: Description is based on seven males and eleven TS (TP): ST ♀ (BMNH) [not examined]. 3. LIST OF DESCRIBED SPECIES AND TL: Bilar, Bohol. Note: The type series should be preserved in ZMHU, but materials are preserved in SMFL. The lectotype designation TL: Amlan, Negros Oriental. and N. Mindanao. Yokochi (1998) mentioned the type series of TL: Camiguin de Mindanao. TL: Negros Or. [Oriental]. phelada G. Semper, [1887] (Fig. 77) females, and two males and three females are preserved in Note: This is a female form of panopus. The type locality SUBSPECIES NAME TS (TP): HT ♀ (FASU), PT 4♀. the specimens that corresponded with the types could not be was carried out by Yokochi (1998). TS (TP): HT ♂ (USC), PT 4♀ (USC, JF). leucotaenia was untraced so designated the lectotype for the TS (TP): ST ♂ (MNHN). TS (TP): HT ♂ (SMFL), PT 1♂1♀. TL: Luzon. proditrix Fruhstorfer, 1913 (Fig. 83) semperi Staudinger, 1889 (Fig. 93) ZMHU. The lectotype designation was carried out by Yokochi suspected is Mindanao, though the specific location is not found. The Fig. 117 is reproduced from the original description figure of the original description, but the type series specimen Note: Though the original description does not mention TS (TP): LT ♀ (SMFL). TL: Basilan. TL: Palawan. (1999). shown. The type series are deposited in the Rothschild collection The following list contains the taxonomic names bongaoensis Schröder & Treadaway, 1990 (Fig. 10) (pl. 50, fig. 4, ♂). eva C. & R. Felder, [1867] (Fig. 30) imperator Hewitson, 1863 (Fig. 40) was found after the third investigation in SMFL. We designate the number of types, a male syntype is preserved in MNHN. nirodha Fruhstorfer, 1913 (Fig. 67) Note: This taxon was described based on five materials TS (TP): ST ♂ (MNHN). TS (TP): LT ♂ (ZMHU). (BMNH, London), but we have not located them at present. concerned with Philippines. The names which are described TL: Bongao Peak, Bongao, Tawitawi Group. TL: Luzon. TL: Luzon. here again the lectotype for a male specimen shown in Fig. 49 TL: Basilan. from Luzon, Camiguin de Mindanao, and SE. Mindanao. Note: Description is based on males and females, and four tawitawia Treadaway & Nuyda, 1994 (Fig. 104) from the other areas are also included. TS (TP): HT ♀ (SMFL), PT 18♂20♀. darani Fruhstorfer, 1913 (Fig. 20) TS (TP): ST ♂ (BMNH). TS (TP): HT ♀ (BMNH). (labelled “Typus (red) / Camotes / Bohol / 113 / 211 / Coll. C. mindanaensis Schröder & Treadaway, 1978 (Fig. 60) TS (TP): ST ♂♀ (MNHN). Yokochi (1998) mentioned the type series of phelada was producta Butler, 1901 (Fig. 84) males and four females are preserved in ZMHU. The lectotype TL: Magsaggao, Tawitawi. visayana (Lexias) Schröder & Treadaway, 1987 (Fig. 110) TL: Sulu Archipelago. Note: According to the original description, eva is distrib- Semper”). TL: del Sur, Mindanao. untraced so designated the lectotype for the figure of the TL: Philippines [recte W. Malaysia]. designation was carried out by Yokochi (1999). TS (TP): HT ♂ (SMFL). TL: Catmon, S. Leyte. albusequus Nihira & Kawamura, 1986 (Fig. 2) borromeoi Schröder, 1977 (Fig. 11) TS (TP): HT ♀ (BMNH). uted in Assam, Luzon, and Celebes. The details of type series ingae (Lexias) Schröder & Treadaway, 1987 (Fig. 41) TS (TP): HT ♂ (SMFL). obatrata Yokochi, 1994 (Fig. 68) original description, but the target specimen was found after the TS (TP): ST ♀ (BMNH). TS (TP): HT ♂ (SMFL), PT 6♂4♀. TL: Lakit lakit, Sanga Sanga, Tawitawi. TL: Espana, 2000 ft, Sibuyan. Note: Though the original description does not mention are not mentioned. Staudinger (1886) is the first revisory of TL: Negros Or. [Oriental], 800 m. leytana Schröder & Treadaway, 1982 (Fig. 50) TL: Dumaran. third investigation in SMFL. We designate here again the senilis Hanafusa, 1992 (Fig. 94) tethys Tsukada, 1991 (Fig. 105) Fig. 1. Younger age of Julian N. Jumalon (left) and Colin G. TS (TP): HT ♀ (SK). TS (TP): HT ♂ (SMFL), PT 1♀. the number of type series, one female would be the holotype. which Luzon is recognized as a locality of eva. TS (TP): HT ♂ (SMFL), PT 5♂5♀. TL: St. Bernard, Calpio, 280 m, S. Leyte. mindorana (Tanaecia) Fruhstorfer, 1904 (Fig. 61) TS (TP): HT ♀ (HH), PT 1♂1♀. lectotype for a female specimen shown in Fig. 77 (labelled rhamases Staudinger, 1889 (Fig. 85) TL: Dumaran. TL: N. Palawan. visayana (Cynitia) Schröder & Treadaway, 1981 (Fig. 111) Treadaway (right). TS (TP): HT ♂ (SMFL), PT 18♂10♀. TL: Mindoro. “Typus (red) / Euth. phelada typ. ♀ Semper / Marivcles (?) TL: Palawan. TS (TP): HT ♂ (HH), PT 1♂. TS (TP): HT ♂ (ET), PT 10♂12♀. TL: St. Bernard, Catmon, S. Leyte. 6 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 7

1. INTRODUCTION Limbach, about 250 km from Frankfurt. It is an undoubted fact amlana Jumalon, 1970 (Fig. 3) cabigoni Schröder, Treadaway, A. Mohagan & N. Mohagan, dhayma Fruhstorfer, 1913 (Fig. 21) exul Tsukada & Nishiyama, 1980 (Fig. 31) ingae (Euthalia) Schröder & Treadaway, 1990 (Fig. 42) leytensis Jumalon, 1975 (Fig. 51) TS (TP): ST 2♂1♀ (BMNH). ormocana Jumalon, 1970 (Fig. 69) 11.11.81 / 110 / Coll. C. Semper / Lectotype ♀ Euthalia TS (TP): LT ♂ (ZMHU). sibutana Schröder & Treadaway, 1990 (Fig. 95) Note: Attached label to the holotype specimen shows TS (TP): HT ♂ (SMFL), PT 10♂2♀. waltraudae Schröder & Treadaway, 1990 (Fig. 114) that Mr. Treadaway and Dr. H. Schröder of Senckenberg TL: Amlan, Negros Oriental. 2009 (Fig. 12) TL: Sulu Archipelago. TL: Bo. Melgar, Dinagat. TL: Bongao Peak, 200 m, Bongao, Tawitawi Group. TL: Lake Danao, Ormoc City, Leyte. Note: The figured female (Fig. 61) is preserved in the TL: Lake Danao, Ormoc City, Leyte. phelada Semper [87]:1888 Designated by T. Yokochi, 1997”). Note: Description is based on males, and four males are TL: Cavan Cavan, 5 m, Sibutu, Tawitawi Group. “Balabac”, but the correct type locality is Palawan. TL: Indalawan, Balabac. The Philippine Islands are situated in the northern end of thoroughly deserve credit for working further on systematic TS (TP): HT ♂ (USC), PT10 ♂1♀ (USC, JF). TL: Balut Is, Sarangani Group. TS (TP): HT ♀ (BMNH). TS (TP): HT ♂ (ET), PT 3♂6♀. TS (TP): HT ♀ (SMFL), PT 10♂12♀. TS (TP): HT ♀ (USC). Rothschild collection (Ro 98/7), BMNH. It might be one of the TS (TP): HT ♂ (USC), PT 1♀. preserved in ZMHU. The lectotype designation was carried out TS (TP): HT ♂ (SMFL), PT 17♂15♀. vistrica Fruhstorfer, 1913 (Fig. 112) TS (TP): HT ♀ (SMFL), PT 2♂2♀. the Malay Archipelago and are composed of more than 7,000 classification in Philippines Rhopalocera. TS (TP): HT ♂ (SMFL), PT 2♂1♀. Note: Though the original description does not mention the types, although the attached label is only “Mindoro”. No other philippensis Fruhstorfer, 1899 (Fig. 78) by Yokochi (1999). trapesa G. Semper, [1887] (Fig. 106) TL: locality unknown, presumably one of the Central islands, large and small, extending from north to south about Here we revised the group of the tribe Adoliadini antiquea Schröder & Treadaway, 1980 (Fig. 4) number of type series, one female would be the holotype. felicitacionae Jumalon, 1989 (Fig. 32) irauana Jumalon, 1975 (Fig. 43) liaoi Schröder & Treadaway, 1982 (Fig. 52) specimen is found that corresponded with the type series. ornata Schröder & Treadaway, 1979 (Fig. 70) TL: Basilan. sibuyana Schröder & Treadaway, 1982 (Fig. 96) TL: Camiguin de Mindanao. Philippine islands [recte Dinagat]. yoshiroui K. Okano, 1992 (Fig. 115) 1,800 km and from east to west about 1,100 km. The insect distributed in the Philippines, based on the checklist by TL: Culasi, 500 m, Antique, Panay. calliphorus C. & R. Felder, 1861 (Fig. 13) TL: Kantakpal Creek, Awihao, Ormoc, Western Leyte. TL: Irauan [Irawan], Puerto Princesa, Palawan. TL: Mt. Madja-an [Mt. Madja-as], 500 m, Antique, Panay. TL: Romblon. TS (TP): LT ♂ (MNHN). rodriguezi Schröder & Treadaway, 1982 (Fig. 86) TL: Sibuyan, Romblon Group. TS (TP): LT ♂ (SMFL). TS (TP): ST ♂ (BMNH). TL: Babuyan. fauna is rich in diversity and the tribe Adoliadini, Nymphalidae, Treadaway (1995). One hundred-twenty taxa of type specimens TS (TP): HT ♂ (SMFL). TL: Luzon. dinagatensis Tsukada, 1991 (Fig. 22) TS (TP): HT ♂ (JF). TS (TP): HT ♀ (JF), PT 1♂1♀ (USC). TS (TP): HT ♂ (SMFL), PT 1♂2♀. mindorana (Euthalia) Fruhstorfer, 1899 (Fig. 62) TS (TP): HT ♂ (SMFL), PT 1♂2♀. Note: Description is based on one male from Basilan TL: 28 km W of Puerto Princesa, Palawan. TS (TP): HT ♂ (SMFL), PT 2♂5♀. Note: Description is based on seven materials from E. Note: Tsukada (1991) suspected this taxon is distributed TS (TP): HT ♂ (KMNH). Lepidoptera, the main group of this study, is no exception. The founded in the Philippines are examined and revised the TS (TP): ST ♂ (BMNH). TL: Dinagat. Note: The holotype is deposited in Jumalon family collec- TL: Mindoro. (MNHN) and one female from Luzon (ZMHU). As the TS (TP): HT ♂ (SMFL), PT 2♂2♀. Mindanao. The lectotype designation was carried out for the in Basilan, but it presumably comes from Dinagat. important works concerned with Philippines were, back in classification of species-subspecies relation. This paper is aquamarina Fruhstorfer, 1913 (Fig. 5) TS (TP): HT ♂ (ET), PT 2♂1♀. gahiti M. & T. Okano, 1990 (unavailable name) (Fig. 33) tion. ludonia Staudinger, 1889 (Fig. 53) TS (TP): ST (MNHN). palawana (Lexias) Moore, 1897 (Fig. 71) Luzon-group should be classified as nadenya. We select a smaragdifera Fruhstorfer, 1913 (Fig. 97) Camiguin de Mindanao specimen by Yokochi (1998). yui Yokochi, 1994 (Fig. 116) history, “Diagnosen einiger neuer Tagfalter von den Philippinen published in two parts. All the taxa described before are TL: Mindanao. candida Schröder, Treadaway & Nuyda, 1990 (Fig. 14) TL: Bilar, Bohol. TL: Palawan. TL: Palawan. male (Fig. 78) deposited in MNHN as the lectotype, labelled, romeo Schröder & Treadaway, 1987 (Fig. 87) TL: Locality unknown, presumably Mindoro [Mindoro]. volupia Tsukada & Nishiyama, 1981 (Fig. 113) TL: Dumaran. und Bemerkungen über die Verbreitung einiger derselben” (G. examined and figured as much as possible in part 1. In the TS (TP): LT ♂ (SMFL). TL: 30 m NN, Calamian Group. dinorah Fruhstorfer, 1899 (Fig. 23) TS (TP): HT ♀ (FASU). joloana Staudinger, 1889 (Fig. 44) TS (TP): LT ♂ (ZMHU). mindorana (Euthalia) Tsukada, 1991 (permanently invalid) TS (TP): HT ♂ (BMNH). “Type (red) / lubentina philippensis Fruhst. / Philippinen TL: Mt. Halcon, N. Mindoro. TS (TP): ST (MNHN). treadawayi Tsukada, 1991 (Fig. 107) TL: Camiguin. TS (TP): HT ♂ (TY), PT 11♂4♀. Semper, 1886–1892), and then “Die Gross-Schmetterlinge der second part, synonymic lists and some important specimens Note: Description is based on some specimens. The lecto- TS (TP): HT ♂ (SMFL), PT10 ♂. TL: Basilan. Note: As this taxon gahiti was described as a form of TL: Sulu. Note: Description is based on four males and six females, Note: The taxon mindorana described under the genus Bazilan II-III, 98 Doherty ex. coll. H. Fruhstorfer / TS (TP): HT ♀ (SMFL), PT 1♀. Note: Though the original description does not mention TL: Romblon, Sibuyanes. TS (TP): HT ♂ (ET), PT 2♂5♀. (to be continued) Erde, 9” (1908–1927) edited by A. Seitz. Recently, a Japanese will be shown, affixing several discussions. type designation was carried out by Yokochi (1998). TS (TP): ST 2♂2♀ (MNHN). Lexias panopus boholensis (infrasubspecific name), it should TS (TP): LT ♂ (ZMHU). and two pairs are preserved in ZMHU. The lectotype designa- Euthalia is a junior primary homonym of Euthalia lusiada palawana (Tanaecia) Staudinger, 1889 (Fig. 72) MUSEUM PARIS 1934 COLL H Fruhstorfer”. the number and sex, a syntype specimen in MNHN is a male. TS (TP): HT ♂ (SMFL). entomologist, E. Tsukada, had compiled a great series of cavarna Fruhstorfer, 1913 (Fig. 15) be treated as an unavailable name. Note: Description is based on two males and one female, tion was carried out by Yokochi (1999). mindorana Fruhstorfer, 1899 (permanently invalid name). TL: Palawan. rudraca Fruhstorfer, 1913 (Fig. 88) “Butterflies of the South East Asian Islands” (1980–1991). balabacana (Bassarona) Tsukada, 1991 (Fig. 6) TL: Balabas [Balabac]. dodong Schröder & Treadaway, 1978 (Fig. 24) and all of them are preserved in ZMHU. The lectotype desig- Now it is replaced by tsukada Koçak (1996). TS (TP): LT ♂ (ZMHU). phlegethon G. Semper, [1887] (Fig. 79) TL: Balabac. soloni M. & T. Okano, 1990 (Fig. 98) tsukada Koçak, 1996 (Fig. 80) The late Prof. Julian N. Jumalon (1909–2000) (Fig. 1, TL: Balabac. TS (TP): LT ♀ (ZMHU). TL: Mobo, 1000 ft, Masbate. galoa Fruhstorfer, 1913 (Fig. 34) nation was carried out by Yokochi (1999). lupina Druce, 1874 (Fig. 54) Note: Description is based on males and females, and six TL: Mindanao. TS (TP): LT ♂ (ZMHU). TL: Bilar, Bohol. TL: Mindoro. left), the Lepidoptera Research Associate and Lecturer of 2. ABBREVIATIONS TS (TP): HT ♂ (ET), PT 8♂6♀. Note: Description is based on a few females, and one spec- TS (TP): HT ♂ (SMFL), PT 3♂7♀. TL: Mindoro. TL: Philippine Islands [Sulu]. mindorensis Schröder & Treadaway, 1982 (Fig. 63) males and three females are preserved in ZMHU. The lecto- TS (TP): LT ♀ (SMFL). Note: Description is based on some specimens, and two TS (TP): HT ♂ (FASU), PT 2♂3♀. TS (TP): HT ♂ (ET), PT 10♂11♀. Department of Biology, University of San Carlos, published a imen is preserved in ZMHU. The lectotype designation was TS (TP): ST (BMNH). kayumanggia Treadaway & Nuyda, 1994 (Fig. 45) TS (TP): HT ♀ (BMNH). TL: Mindoro. type designation was carried out by Yokochi (1999). Note: Description is based on twenty-one males and seven males and two females are preserved in ZMHU. The lectotype Note: Euthalia lubentina mindorana Tsukada, 1991 is a paper of “New Philippine butterflies” in 1975, and many new Public Institution (Museum, University). BMNH: The balabacana (Dophla) Tsukada, 1991 (secondary homonym) carried out by Yokochi (1999). dohertyi Butler, 1901 (Fig. 25) Note: Though the original description does not mention TL: Mt. Bayog, S. Palawan. TS (TP): HT ♂ (SMFL), PT 6♂7♀. females from Samar, Camiguin de Mindanao, N. Mindanao, E. designation was carried out by Yokochi (1999). soregina Fruhstorfer, 1913 (Fig. 99) permanently invalid name (nom. nov. pro mindorana Tsukada, taxonomic names concerned with various families were Natural History Museum, London, United Kingdom; FASU: (Fig. 7) TL: Sulu Archipelago. the number and sex, syntype specimen in BMNH is a male. TS (TP): HT ♂ (SMFL), PT 13♂4♀. lusiada C. & R. Felder, 1863 (Fig. 55) palawana (Euthalia) Staudinger, 1889 (Fig. 73) Mindanao, SE. Mindanao, and several materials are preserved TL: Sulu Archipelago. 1991, non Fruhstorfer,1899). New name tsukada was proposed described there. He played a leading part of the study of Exhibition Hall of Food, Agricultural and Environmental Note: See below balabacensis. chloe Schröder & Treadaway, 1990 (Fig. 16) TS (TP): HT ♂ (BMNH). TL: Luzon. miscus Fruhstorfer, 1913 (Fig. 64) TL: Palawan. in SMFL. The lectotype designation was carried out for the samarensis Tsukada, 1991 (Fig. 89) TS (TP): ST ♀ (BMNH). by Koçak (1996). Philippine butterflies and contributed greatly to entomology. Sciences, Faculty of Agriculture, Shinshu University, Nagano, TL: Jao Asan, 200 m, Sibuyan, Romblon Group. goertzi Jumalon, 1975 (Fig. 35) kulaya Treadaway & Nuyda, 1994 (Fig. 46) TS (TP): ST ♂ (SMFL). TL: Mindanao. TS (TP): LT ♂ (ZMHU). Mindanao specimen by Yokochi (1998). TL: San Juan, Samar. Note: According to the original description, the type series In 1995, the late Mr. Colin G. Treadaway (Fig. 1, right, Japan; KMNH: Kitakyushu Museum of Natural History & balabacensis nom. nov. (Fig. 7) TS (TP): HT ♂ (SMFL), PT 11♂9♀. dumaranensis Hanafusa, 1992 (Fig. 26) TL: Amlan, Negros Oriental. TL: Magellanes, Homonhon. Note: Though the original description does not mention TS (TP): ST (MNHN, BMNH). Note: Description is based on one male and six females, TS (TP): HT ♂ (ET), PT 3♂2♀. are exactly female, but the number is unknown. tyawena Fruhstorfer, 1913 (Fig. 108) died in 2019) published “Checklist of the butterflies of the Human History, Fukuoka, Japan; MNHN: Museum National TL: Balabac. TL: Dumaran. TS (TP): HT ♀ (ET). TS (TP): HT ♂ (SMFL), PT 14♂10♀. the number of types, a male syntype is preserved in SMFL. Note: According to the original description, types and one male and four females are preserved in ZMHU. The pinamalayana Yokochi, 1997 (Fig. 80) TL: unknown [Palawan]. Philippine Islands (Lepidoptera: Rhopalocera)”. Although, 25 d'Histoire Naturelle Entomologie, Paris, France; SMFL: TS (TP): HT ♂ (ET), PT 2♂2♀. circe Schröder & Treadaway, 1990 (Fig. 17) TS (TP): HT ♂ (HH). Note: Though the original description mentions the Note: The name kulaya is spelt “kúlaya” in the original (syntypes) are preserved in Fruhstorfer collection (now in lectotype designation was carried out by Yokochi (1999). Note: The new name pinamalayana was proposed by sarmana Fruhstorfer, 1913 (Fig. 90) subpiratica Schröder & Treadaway, 1987 (Fig. 100) TS (TP): LT ♀ (ZMHU). years have passed since his paper presented, several new Senckenberg Forschungsinstitut und Naturmuseum Frankfurt, Note: The taxon balabacana was described under the TL: Cavan Cavan, Sibutu, Tawitawi Group. holotype is preserved in USC, it is deposited in ET. description. macer Tsukada & Nishiyama, 1980 (Fig. 56) MNHN) and BMNH. Yokochi (1997), because mindorana Tsukada, 1991 was a TL: Basilan. TL: Nueva Ecija, Bongabon, Centr. Luzon. Note: Description is based on some specimens, and one findings are still remained unpublished — it can be followed Frankfurt-am-Main, Germany; USC: University of San Carlos, genus Dophla. As Yokochi (2010) assigned both Dophla and TS (TP): HT ♂ (SMFL), PT 9♂10♀. dunya Doubleday, [1848] (Fig. 27) TL: Bo. Melgar, Dinagat. pallida Schröder, Treadaway & Nuyda, 1990 (Fig. 74) permanently invalid name. However, pinamalayana is a junior TS (TP): ST (MNHN). TS (TP): HT ♂ (SMFL), PT1♀. male and one female are preserved in ZMHU. The lectotype by his own handwriting on the glass surface of his specimen Cebu, Philippines; ZMHU: Zoologisches Museum, Humboldt Bassarona to the genus Euthalia, balabacana (Dophla) is a TL: Assam, India. hikarugenzi Tsukada & Nishiyama, 1980 (Fig. 36) laetitiae Schröder & Treadaway, 2000 (Fig. 47) TS (TP): HT ♂ (ET), PT 3♂2♀. monara Fruhstorfer, 1913 (Fig. 65) TL: 30 m NN, Calamian Group. synonym of tsukada Koçak (1996). Note: Though the original description does not mention designation was carried out by Yokochi (1999). cases —. The first author visited Senckenberg Museum in Universität, Berlin, Germany. junior secondary homonym of Bassarona teuta balabacana cusama Fruhstorfer, 1913 (Fig. 18) TS (TP): ST ♂ (BMNH). TL: near Baguio, N. Luzon. TL: Cavan Cavan, 5 m, Sibutu, Tawitawi Group. TL: N. Borneo. TS (TP): HT ♂ (SMFL), PT 11♂16♀. the number and sex, a syntype specimen in MNHN is a male. suluana Fruhstorfer, 1902 (Fig. 101) Frankfurt to research on the Treadaway collection on March Private collections. ET: Etsuzo Tsukada, Yamanashi, Tsukada, 1991. Here a new name balabacensis is replaced for TL: Mindanao, Camiguin de Mindanao. TS (TP): HT ♂ (MKN), PT 1♀ (MKN). TS (TP): HT ♂ (SMFL), PT 4♂. malissia Fruhstorfer, 1913 (Fig. 57) TS (TP): ST (MNHN). piratica G. Semper, [1887] (Fig. 81) TL: Sulu Archipelago. uposatha Fruhstorfer, 1913 (Fig. 109) 2019, and just then was informed that he had died several Japan; HH: Hiroto Hanafusa, Tottori, Japan; JF: Jumalon Dophla evelina balabacana Tsukada, 1991. TS (TP): ST ♂ (MNHN), ♀ (SMFL). ellora Fruhstorfer, 1898 (Fig. 28) TL: Basilan. panayana Schröder & Treadaway, 1980 (Fig. 75) TL: Camiguin de Mindanao. satrapes C. & R. Felder, 1861 (Fig. 91) TS (TP): HT ♀ (BMNH). TL: Banggi (Banguey), E. Malaysia. weeks ago at his home in England. In the middle 1990’s, every Family, Cebu, Philippines; MKN: Misao Kaneko, Tokyo, Euthalia (Dophla) evelina balabacensis nom. nov. Note: Figured specimen is a syntype female in SMFL. TL: Mindoro. hinunanganensis M. & T. Okano, 1986 (Fig. 37) leopardina Fruhstorfer, 1913 (Fig. 48) TS (TP): ST♂ (MNHN). nadenya Fruhstorfer, 1913 TL: Culasi, 800 ft, Antique, Panay. TS (TP): LT ♂ (SMFL). TL: Luzon. TS (TP): LT ♂ (ZMHU). time the first author visited to Senckenberg, he came bringing Japan; SK: Shun-ichi Kawamura, Tokyo, Japan; STA: Shinji pro Dophla evelina balabacana Tsukada, 1991, non TS (TP): ST ♂ (MNHN). TL: Hinunangan, Leyte. TL: Sulu Islands. Note: Though the original description does not mention TL: Luzon. TS (TP): HT ♂ (SMFL), PT 4♂4♀. Note: Description is based on six materials from Luzon, TS (TP): ST ♂ (BMNH). susoni Jumalon, 1975 (Fig. 102) Note: According to the original description, the type series his described type specimens, with his wife driving from Taga, Tokyo, Japan; TY: Takashi Yokochi, Aichi, Japan. Bassarona teuta balabacana Tsukada, 1991 dacasini Hanafusa, 1990 (Fig. 19) Note: Though the original description does not mention TS (TP): HT ♂ (FASU), PT 1♂. TS (TP): HT ♀ (BMNH). the number of types, a male syntype is preserved in MNHN. TS (TP): HT ♀ [untraced, not examined]. Mindoro, Camiguin de Mindanao, N. Mindanao. The lectotype Note: According to the original description, the type series TL: Camp 7, Minglanilla, Cebu. (male and female, but the number is unknown) are deposited in Specimens concerned. FW: Length of Forewing; HT: TL: Balabac. the number and sex, a syntype specimen in MNHN is a male. Note: Though the original description indicates that the panopus C. & R. Felder, 1861 (Fig. 76) designation was carried out for the Camiguin de Mindanao (number is unknown) are deposited in Semper and Felder TS (TP): HT ♂ (USC), PT 9♂9♀ (USC, JF, SMFL, TY). Staudinger collection (ZMHU). The lectotype designation was Holotype; LT: Lectotype; PLT: Paralectotype; PT: Paratype; boholensis (Lexias) M. & T. Okano, 1988 (Fig. 8) TS (TP): HT ♂ (HH), PT 1♀. hiwaga Nuyda & Kawamura, 1989 (Fig. 38) leucotaenia G. Semper, 1878 (Fig. 49) marinduque K. Okano, 1992 (Fig. 58) type female should be preserved in ZMHU, the specimen TL: Luzon. specimen by Schröder & Treadaway (1987). collections. carried out by Yokochi (1999). TL: Type Locality; TP: Type Preservation; TS: Type Series; TL: Bilar, Bohol. eson de Nicéville, 1894 (Fig. 29) TL: Camiguin, Babuyan. TL: Camotes. TL: Marinduque. corresponding with nadenya could not be found there. No TS (TP): ST ♂ (BMNH). tanagra Staudinger, 1889 (Fig. 103) ST: Syntype. TS (TP): HT ♂ (FASU), PT 6♂2♀. damalis Erichson, 1834 (Fig. 117) TL: Palawan. TS (TP): HT ♂ (SK), PT 7♂4♀ (SK, ET). TS (TP): LT ♂ (SMFL). TS (TP): HT ♀ (KMNH), PT 1♂ (KMNH). figure is presented in the original paper. Note: The original description does not mention the princesa Fruhstorfer, 1899 (Fig. 82) schoenigi Schröder & Treadaway, 1978 (Fig. 92) TL: Palawan. virginalis Fruhstorfer, 1913 TL: Luzon. TS (TP): LT ♂ (SMFL), PLT 1♀. Note: This taxon was described based on sixty-nine number of type series. TL: Palawan. TL: Amlan, 600 m, Negros. TS (TP): LT ♂ (ZMHU). TL: locality not given. boholensis (Euthalia) M. & T. Okano, 1990 (Fig. 9) TS (TP): ST ♂ [untraced, not examined]. Note: Description is based on a single pair, and all howarthi Jumalon, 1975 (Fig. 39) materials from Camotes, Bohol, Leyte, Panaon, E. Mindanao, medaga Fruhstorfer, 1913 (Fig. 59) negrosiana Schröder & Treadaway, 1987 (Fig. 66) TS (TP): HT ♀ (MNHN). TS (TP): HT ♀ (SMFL). Note: Description is based on seven males and eleven TS (TP): ST ♀ (BMNH) [not examined]. 3. LIST OF DESCRIBED SPECIES AND TL: Bilar, Bohol. Note: The type series should be preserved in ZMHU, but materials are preserved in SMFL. The lectotype designation TL: Amlan, Negros Oriental. and N. Mindanao. Yokochi (1998) mentioned the type series of TL: Camiguin de Mindanao. TL: Negros Or. [Oriental]. phelada G. Semper, [1887] (Fig. 77) females, and two males and three females are preserved in Note: This is a female form of panopus. The type locality SUBSPECIES NAME TS (TP): HT ♀ (FASU), PT 4♀. the specimens that corresponded with the types could not be was carried out by Yokochi (1998). TS (TP): HT ♂ (USC), PT 4♀ (USC, JF). leucotaenia was untraced so designated the lectotype for the TS (TP): ST ♂ (MNHN). TS (TP): HT ♂ (SMFL), PT 1♂1♀. TL: Luzon. proditrix Fruhstorfer, 1913 (Fig. 83) semperi Staudinger, 1889 (Fig. 93) ZMHU. The lectotype designation was carried out by Yokochi suspected is Mindanao, though the specific location is not found. The Fig. 117 is reproduced from the original description figure of the original description, but the type series specimen Note: Though the original description does not mention TS (TP): LT ♀ (SMFL). TL: Basilan. TL: Palawan. (1999). shown. The type series are deposited in the Rothschild collection The following list contains the taxonomic names bongaoensis Schröder & Treadaway, 1990 (Fig. 10) (pl. 50, fig. 4, ♂). eva C. & R. Felder, [1867] (Fig. 30) imperator Hewitson, 1863 (Fig. 40) was found after the third investigation in SMFL. We designate the number of types, a male syntype is preserved in MNHN. nirodha Fruhstorfer, 1913 (Fig. 67) Note: This taxon was described based on five materials TS (TP): ST ♂ (MNHN). TS (TP): LT ♂ (ZMHU). (BMNH, London), but we have not located them at present. concerned with Philippines. The names which are described TL: Bongao Peak, Bongao, Tawitawi Group. TL: Luzon. TL: Luzon. here again the lectotype for a male specimen shown in Fig. 49 TL: Basilan. from Luzon, Camiguin de Mindanao, and SE. Mindanao. Note: Description is based on males and females, and four tawitawia Treadaway & Nuyda, 1994 (Fig. 104) from the other areas are also included. TS (TP): HT ♀ (SMFL), PT 18♂20♀. darani Fruhstorfer, 1913 (Fig. 20) TS (TP): ST ♂ (BMNH). TS (TP): HT ♀ (BMNH). (labelled “Typus (red) / Camotes / Bohol / 113 / 211 / Coll. C. mindanaensis Schröder & Treadaway, 1978 (Fig. 60) TS (TP): ST ♂♀ (MNHN). Yokochi (1998) mentioned the type series of phelada was producta Butler, 1901 (Fig. 84) males and four females are preserved in ZMHU. The lectotype TL: Magsaggao, Tawitawi. visayana (Lexias) Schröder & Treadaway, 1987 (Fig. 110) TL: Sulu Archipelago. Note: According to the original description, eva is distrib- Semper”). TL: Surigao del Sur, Mindanao. untraced so designated the lectotype for the figure of the TL: Philippines [recte W. Malaysia]. designation was carried out by Yokochi (1999). TS (TP): HT ♂ (SMFL). TL: Catmon, S. Leyte. albusequus Nihira & Kawamura, 1986 (Fig. 2) borromeoi Schröder, 1977 (Fig. 11) TS (TP): HT ♀ (BMNH). uted in Assam, Luzon, and Celebes. The details of type series ingae (Lexias) Schröder & Treadaway, 1987 (Fig. 41) TS (TP): HT ♂ (SMFL). obatrata Yokochi, 1994 (Fig. 68) original description, but the target specimen was found after the TS (TP): ST ♀ (BMNH). TS (TP): HT ♂ (SMFL), PT 6♂4♀. TL: Lakit lakit, Sanga Sanga, Tawitawi. TL: Espana, 2000 ft, Sibuyan. Note: Though the original description does not mention are not mentioned. Staudinger (1886) is the first revisory of TL: Negros Or. [Oriental], 800 m. leytana Schröder & Treadaway, 1982 (Fig. 50) TL: Dumaran. third investigation in SMFL. We designate here again the senilis Hanafusa, 1992 (Fig. 94) tethys Tsukada, 1991 (Fig. 105) Fig. 1. Younger age of Julian N. Jumalon (left) and Colin G. TS (TP): HT ♀ (SK). TS (TP): HT ♂ (SMFL), PT 1♀. the number of type series, one female would be the holotype. which Luzon is recognized as a locality of eva. TS (TP): HT ♂ (SMFL), PT 5♂5♀. TL: St. Bernard, Calpio, 280 m, S. Leyte. mindorana (Tanaecia) Fruhstorfer, 1904 (Fig. 61) TS (TP): HT ♀ (HH), PT 1♂1♀. lectotype for a female specimen shown in Fig. 77 (labelled rhamases Staudinger, 1889 (Fig. 85) TL: Dumaran. TL: N. Palawan. visayana (Cynitia) Schröder & Treadaway, 1981 (Fig. 111) Treadaway (right). TS (TP): HT ♂ (SMFL), PT 18♂10♀. TL: Mindoro. “Typus (red) / Euth. phelada typ. ♀ Semper / Marivcles (?) TL: Palawan. TS (TP): HT ♂ (HH), PT 1♂. TS (TP): HT ♂ (ET), PT 10♂12♀. TL: St. Bernard, Catmon, S. Leyte. 8 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 9

1. INTRODUCTION Limbach, about 250 km from Frankfurt. It is an undoubted fact amlana Jumalon, 1970 (Fig. 3) cabigoni Schröder, Treadaway, A. Mohagan & N. Mohagan, dhayma Fruhstorfer, 1913 (Fig. 21) exul Tsukada & Nishiyama, 1980 (Fig. 31) ingae (Euthalia) Schröder & Treadaway, 1990 (Fig. 42) leytensis Jumalon, 1975 (Fig. 51) TS (TP): ST 2♂1♀ (BMNH). ormocana Jumalon, 1970 (Fig. 69) 11.11.81 / 110 / Coll. C. Semper / Lectotype ♀ Euthalia TS (TP): LT ♂ (ZMHU). sibutana Schröder & Treadaway, 1990 (Fig. 95) Note: Attached label to the holotype specimen shows TS (TP): HT ♂ (SMFL), PT 10♂2♀. waltraudae Schröder & Treadaway, 1990 (Fig. 114) that Mr. Treadaway and Dr. H. Schröder of Senckenberg TL: Amlan, Negros Oriental. 2009 (Fig. 12) TL: Sulu Archipelago. TL: Bo. Melgar, Dinagat. TL: Bongao Peak, 200 m, Bongao, Tawitawi Group. TL: Lake Danao, Ormoc City, Leyte. Note: The figured female (Fig. 61) is preserved in the TL: Lake Danao, Ormoc City, Leyte. phelada Semper [87]:1888 Designated by T. Yokochi, 1997”). Note: Description is based on males, and four males are TL: Cavan Cavan, 5 m, Sibutu, Tawitawi Group. “Balabac”, but the correct type locality is Palawan. TL: Indalawan, Balabac. The Philippine Islands are situated in the northern end of thoroughly deserve credit for working further on systematic TS (TP): HT ♂ (USC), PT10 ♂1♀ (USC, JF). TL: Balut Is, Sarangani Group. TS (TP): HT ♀ (BMNH). TS (TP): HT ♂ (ET), PT 3♂6♀. TS (TP): HT ♀ (SMFL), PT 10♂12♀. TS (TP): HT ♀ (USC). Rothschild collection (Ro 98/7), BMNH. It might be one of the TS (TP): HT ♂ (USC), PT 1♀. preserved in ZMHU. The lectotype designation was carried out TS (TP): HT ♂ (SMFL), PT 17♂15♀. vistrica Fruhstorfer, 1913 (Fig. 112) TS (TP): HT ♀ (SMFL), PT 2♂2♀. the Malay Archipelago and are composed of more than 7,000 classification in Philippines Rhopalocera. TS (TP): HT ♂ (SMFL), PT 2♂1♀. Note: Though the original description does not mention the types, although the attached label is only “Mindoro”. No other philippensis Fruhstorfer, 1899 (Fig. 78) by Yokochi (1999). trapesa G. Semper, [1887] (Fig. 106) TL: locality unknown, presumably one of the Central islands, large and small, extending from north to south about Here we revised the group of the tribe Adoliadini antiquea Schröder & Treadaway, 1980 (Fig. 4) number of type series, one female would be the holotype. felicitacionae Jumalon, 1989 (Fig. 32) irauana Jumalon, 1975 (Fig. 43) liaoi Schröder & Treadaway, 1982 (Fig. 52) specimen is found that corresponded with the type series. ornata Schröder & Treadaway, 1979 (Fig. 70) TL: Basilan. sibuyana Schröder & Treadaway, 1982 (Fig. 96) TL: Camiguin de Mindanao. Philippine islands [recte Dinagat]. yoshiroui K. Okano, 1992 (Fig. 115) 1,800 km and from east to west about 1,100 km. The insect distributed in the Philippines, based on the checklist by TL: Culasi, 500 m, Antique, Panay. calliphorus C. & R. Felder, 1861 (Fig. 13) TL: Kantakpal Creek, Awihao, Ormoc, Western Leyte. TL: Irauan [Irawan], Puerto Princesa, Palawan. TL: Mt. Madja-an [Mt. Madja-as], 500 m, Antique, Panay. TL: Romblon. TS (TP): LT ♂ (MNHN). rodriguezi Schröder & Treadaway, 1982 (Fig. 86) TL: Sibuyan, Romblon Group. TS (TP): LT ♂ (SMFL). TS (TP): ST ♂ (BMNH). TL: Babuyan. fauna is rich in diversity and the tribe Adoliadini, Nymphalidae, Treadaway (1995). One hundred-twenty taxa of type specimens TS (TP): HT ♂ (SMFL). TL: Luzon. dinagatensis Tsukada, 1991 (Fig. 22) TS (TP): HT ♂ (JF). TS (TP): HT ♀ (JF), PT 1♂1♀ (USC). TS (TP): HT ♂ (SMFL), PT 1♂2♀. mindorana (Euthalia) Fruhstorfer, 1899 (Fig. 62) TS (TP): HT ♂ (SMFL), PT 1♂2♀. Note: Description is based on one male from Basilan TL: 28 km W of Puerto Princesa, Palawan. TS (TP): HT ♂ (SMFL), PT 2♂5♀. Note: Description is based on seven materials from E. Note: Tsukada (1991) suspected this taxon is distributed TS (TP): HT ♂ (KMNH). Lepidoptera, the main group of this study, is no exception. The founded in the Philippines are examined and revised the TS (TP): ST ♂ (BMNH). TL: Dinagat. Note: The holotype is deposited in Jumalon family collec- TL: Mindoro. (MNHN) and one female from Luzon (ZMHU). As the TS (TP): HT ♂ (SMFL), PT 2♂2♀. Mindanao. The lectotype designation was carried out for the in Basilan, but it presumably comes from Dinagat. important works concerned with Philippines were, back in classification of species-subspecies relation. This paper is aquamarina Fruhstorfer, 1913 (Fig. 5) TS (TP): HT ♂ (ET), PT 2♂1♀. gahiti M. & T. Okano, 1990 (unavailable name) (Fig. 33) tion. ludonia Staudinger, 1889 (Fig. 53) TS (TP): ST (MNHN). palawana (Lexias) Moore, 1897 (Fig. 71) Luzon-group should be classified as nadenya. We select a smaragdifera Fruhstorfer, 1913 (Fig. 97) Camiguin de Mindanao specimen by Yokochi (1998). yui Yokochi, 1994 (Fig. 116) history, “Diagnosen einiger neuer Tagfalter von den Philippinen published in two parts. All the taxa described before are TL: Mindanao. candida Schröder, Treadaway & Nuyda, 1990 (Fig. 14) TL: Bilar, Bohol. TL: Palawan. TL: Palawan. male (Fig. 78) deposited in MNHN as the lectotype, labelled, romeo Schröder & Treadaway, 1987 (Fig. 87) TL: Locality unknown, presumably Mindoro [Mindoro]. volupia Tsukada & Nishiyama, 1981 (Fig. 113) TL: Dumaran. und Bemerkungen über die Verbreitung einiger derselben” (G. examined and figured as much as possible in part 1. In the TS (TP): LT ♂ (SMFL). TL: 30 m NN, Calamian Group. dinorah Fruhstorfer, 1899 (Fig. 23) TS (TP): HT ♀ (FASU). joloana Staudinger, 1889 (Fig. 44) TS (TP): LT ♂ (ZMHU). mindorana (Euthalia) Tsukada, 1991 (permanently invalid) TS (TP): HT ♂ (BMNH). “Type (red) / lubentina philippensis Fruhst. / Philippinen TL: Mt. Halcon, N. Mindoro. TS (TP): ST (MNHN). treadawayi Tsukada, 1991 (Fig. 107) TL: Camiguin. TS (TP): HT ♂ (TY), PT 11♂4♀. Semper, 1886–1892), and then “Die Gross-Schmetterlinge der second part, synonymic lists and some important specimens Note: Description is based on some specimens. The lecto- TS (TP): HT ♂ (SMFL), PT10 ♂. TL: Basilan. Note: As this taxon gahiti was described as a form of TL: Sulu. Note: Description is based on four males and six females, Note: The taxon mindorana described under the genus Bazilan II-III, 98 Doherty ex. coll. H. Fruhstorfer / TS (TP): HT ♀ (SMFL), PT 1♀. Note: Though the original description does not mention TL: Romblon, Sibuyanes. TS (TP): HT ♂ (ET), PT 2♂5♀. (to be continued) Erde, 9” (1908–1927) edited by A. Seitz. Recently, a Japanese will be shown, affixing several discussions. type designation was carried out by Yokochi (1998). TS (TP): ST 2♂2♀ (MNHN). Lexias panopus boholensis (infrasubspecific name), it should TS (TP): LT ♂ (ZMHU). and two pairs are preserved in ZMHU. The lectotype designa- Euthalia is a junior primary homonym of Euthalia lusiada palawana (Tanaecia) Staudinger, 1889 (Fig. 72) MUSEUM PARIS 1934 COLL H Fruhstorfer”. the number and sex, a syntype specimen in MNHN is a male. TS (TP): HT ♂ (SMFL). entomologist, E. Tsukada, had compiled a great series of cavarna Fruhstorfer, 1913 (Fig. 15) be treated as an unavailable name. Note: Description is based on two males and one female, tion was carried out by Yokochi (1999). mindorana Fruhstorfer, 1899 (permanently invalid name). TL: Palawan. rudraca Fruhstorfer, 1913 (Fig. 88) “Butterflies of the South East Asian Islands” (1980–1991). balabacana (Bassarona) Tsukada, 1991 (Fig. 6) TL: Balabas [Balabac]. dodong Schröder & Treadaway, 1978 (Fig. 24) and all of them are preserved in ZMHU. The lectotype desig- Now it is replaced by tsukada Koçak (1996). TS (TP): LT ♂ (ZMHU). phlegethon G. Semper, [1887] (Fig. 79) TL: Balabac. soloni M. & T. Okano, 1990 (Fig. 98) tsukada Koçak, 1996 (Fig. 80) The late Prof. Julian N. Jumalon (1909–2000) (Fig. 1, TL: Balabac. TS (TP): LT ♀ (ZMHU). TL: Mobo, 1000 ft, Masbate. galoa Fruhstorfer, 1913 (Fig. 34) nation was carried out by Yokochi (1999). lupina Druce, 1874 (Fig. 54) Note: Description is based on males and females, and six TL: Mindanao. TS (TP): LT ♂ (ZMHU). TL: Bilar, Bohol. TL: Mindoro. left), the Lepidoptera Research Associate and Lecturer of 2. ABBREVIATIONS TS (TP): HT ♂ (ET), PT 8♂6♀. Note: Description is based on a few females, and one spec- TS (TP): HT ♂ (SMFL), PT 3♂7♀. TL: Mindoro. TL: Philippine Islands [Sulu]. mindorensis Schröder & Treadaway, 1982 (Fig. 63) males and three females are preserved in ZMHU. The lecto- TS (TP): LT ♀ (SMFL). Note: Description is based on some specimens, and two TS (TP): HT ♂ (FASU), PT 2♂3♀. TS (TP): HT ♂ (ET), PT 10♂11♀. Department of Biology, University of San Carlos, published a imen is preserved in ZMHU. The lectotype designation was TS (TP): ST (BMNH). kayumanggia Treadaway & Nuyda, 1994 (Fig. 45) TS (TP): HT ♀ (BMNH). TL: Mindoro. type designation was carried out by Yokochi (1999). Note: Description is based on twenty-one males and seven males and two females are preserved in ZMHU. The lectotype Note: Euthalia lubentina mindorana Tsukada, 1991 is a paper of “New Philippine butterflies” in 1975, and many new Public Institution (Museum, University). BMNH: The balabacana (Dophla) Tsukada, 1991 (secondary homonym) carried out by Yokochi (1999). dohertyi Butler, 1901 (Fig. 25) Note: Though the original description does not mention TL: Mt. Bayog, S. Palawan. TS (TP): HT ♂ (SMFL), PT 6♂7♀. females from Samar, Camiguin de Mindanao, N. Mindanao, E. designation was carried out by Yokochi (1999). soregina Fruhstorfer, 1913 (Fig. 99) permanently invalid name (nom. nov. pro mindorana Tsukada, taxonomic names concerned with various families were Natural History Museum, London, United Kingdom; FASU: (Fig. 7) TL: Sulu Archipelago. the number and sex, syntype specimen in BMNH is a male. TS (TP): HT ♂ (SMFL), PT 13♂4♀. lusiada C. & R. Felder, 1863 (Fig. 55) palawana (Euthalia) Staudinger, 1889 (Fig. 73) Mindanao, SE. Mindanao, and several materials are preserved TL: Sulu Archipelago. 1991, non Fruhstorfer,1899). New name tsukada was proposed described there. He played a leading part of the study of Exhibition Hall of Food, Agricultural and Environmental Note: See below balabacensis. chloe Schröder & Treadaway, 1990 (Fig. 16) TS (TP): HT ♂ (BMNH). TL: Luzon. miscus Fruhstorfer, 1913 (Fig. 64) TL: Palawan. in SMFL. The lectotype designation was carried out for the samarensis Tsukada, 1991 (Fig. 89) TS (TP): ST ♀ (BMNH). by Koçak (1996). Philippine butterflies and contributed greatly to entomology. Sciences, Faculty of Agriculture, Shinshu University, Nagano, TL: Jao Asan, 200 m, Sibuyan, Romblon Group. goertzi Jumalon, 1975 (Fig. 35) kulaya Treadaway & Nuyda, 1994 (Fig. 46) TS (TP): ST ♂ (SMFL). TL: Mindanao. TS (TP): LT ♂ (ZMHU). Mindanao specimen by Yokochi (1998). TL: San Juan, Samar. Note: According to the original description, the type series In 1995, the late Mr. Colin G. Treadaway (Fig. 1, right, Japan; KMNH: Kitakyushu Museum of Natural History & balabacensis nom. nov. (Fig. 7) TS (TP): HT ♂ (SMFL), PT 11♂9♀. dumaranensis Hanafusa, 1992 (Fig. 26) TL: Amlan, Negros Oriental. TL: Magellanes, Homonhon. Note: Though the original description does not mention TS (TP): ST (MNHN, BMNH). Note: Description is based on one male and six females, TS (TP): HT ♂ (ET), PT 3♂2♀. are exactly female, but the number is unknown. tyawena Fruhstorfer, 1913 (Fig. 108) died in 2019) published “Checklist of the butterflies of the Human History, Fukuoka, Japan; MNHN: Museum National TL: Balabac. TL: Dumaran. TS (TP): HT ♀ (ET). TS (TP): HT ♂ (SMFL), PT 14♂10♀. the number of types, a male syntype is preserved in SMFL. Note: According to the original description, types and one male and four females are preserved in ZMHU. The pinamalayana Yokochi, 1997 (Fig. 80) TL: unknown [Palawan]. Philippine Islands (Lepidoptera: Rhopalocera)”. Although, 25 d'Histoire Naturelle Entomologie, Paris, France; SMFL: TS (TP): HT ♂ (ET), PT 2♂2♀. circe Schröder & Treadaway, 1990 (Fig. 17) TS (TP): HT ♂ (HH). Note: Though the original description mentions the Note: The name kulaya is spelt “kúlaya” in the original (syntypes) are preserved in Fruhstorfer collection (now in lectotype designation was carried out by Yokochi (1999). Note: The new name pinamalayana was proposed by sarmana Fruhstorfer, 1913 (Fig. 90) subpiratica Schröder & Treadaway, 1987 (Fig. 100) TS (TP): LT ♀ (ZMHU). years have passed since his paper presented, several new Senckenberg Forschungsinstitut und Naturmuseum Frankfurt, Note: The taxon balabacana was described under the TL: Cavan Cavan, Sibutu, Tawitawi Group. holotype is preserved in USC, it is deposited in ET. description. macer Tsukada & Nishiyama, 1980 (Fig. 56) MNHN) and BMNH. Yokochi (1997), because mindorana Tsukada, 1991 was a TL: Basilan. TL: Nueva Ecija, Bongabon, Centr. Luzon. Note: Description is based on some specimens, and one findings are still remained unpublished — it can be followed Frankfurt-am-Main, Germany; USC: University of San Carlos, genus Dophla. As Yokochi (2010) assigned both Dophla and TS (TP): HT ♂ (SMFL), PT 9♂10♀. dunya Doubleday, [1848] (Fig. 27) TL: Bo. Melgar, Dinagat. pallida Schröder, Treadaway & Nuyda, 1990 (Fig. 74) permanently invalid name. However, pinamalayana is a junior TS (TP): ST (MNHN). TS (TP): HT ♂ (SMFL), PT1♀. male and one female are preserved in ZMHU. The lectotype by his own handwriting on the glass surface of his specimen Cebu, Philippines; ZMHU: Zoologisches Museum, Humboldt Bassarona to the genus Euthalia, balabacana (Dophla) is a TL: Assam, India. hikarugenzi Tsukada & Nishiyama, 1980 (Fig. 36) laetitiae Schröder & Treadaway, 2000 (Fig. 47) TS (TP): HT ♂ (ET), PT 3♂2♀. monara Fruhstorfer, 1913 (Fig. 65) TL: 30 m NN, Calamian Group. synonym of tsukada Koçak (1996). Note: Though the original description does not mention designation was carried out by Yokochi (1999). cases —. The first author visited Senckenberg Museum in Universität, Berlin, Germany. junior secondary homonym of Bassarona teuta balabacana cusama Fruhstorfer, 1913 (Fig. 18) TS (TP): ST ♂ (BMNH). TL: near Baguio, N. Luzon. TL: Cavan Cavan, 5 m, Sibutu, Tawitawi Group. TL: N. Borneo. TS (TP): HT ♂ (SMFL), PT 11♂16♀. the number and sex, a syntype specimen in MNHN is a male. suluana Fruhstorfer, 1902 (Fig. 101) Frankfurt to research on the Treadaway collection on March Private collections. ET: Etsuzo Tsukada, Yamanashi, Tsukada, 1991. Here a new name balabacensis is replaced for TL: Mindanao, Camiguin de Mindanao. TS (TP): HT ♂ (MKN), PT 1♀ (MKN). TS (TP): HT ♂ (SMFL), PT 4♂. malissia Fruhstorfer, 1913 (Fig. 57) TS (TP): ST (MNHN). piratica G. Semper, [1887] (Fig. 81) TL: Sulu Archipelago. uposatha Fruhstorfer, 1913 (Fig. 109) 2019, and just then was informed that he had died several Japan; HH: Hiroto Hanafusa, Tottori, Japan; JF: Jumalon Dophla evelina balabacana Tsukada, 1991. TS (TP): ST ♂ (MNHN), ♀ (SMFL). ellora Fruhstorfer, 1898 (Fig. 28) TL: Basilan. panayana Schröder & Treadaway, 1980 (Fig. 75) TL: Camiguin de Mindanao. satrapes C. & R. Felder, 1861 (Fig. 91) TS (TP): HT ♀ (BMNH). TL: Banggi (Banguey), E. Malaysia. weeks ago at his home in England. In the middle 1990’s, every Family, Cebu, Philippines; MKN: Misao Kaneko, Tokyo, Euthalia (Dophla) evelina balabacensis nom. nov. Note: Figured specimen is a syntype female in SMFL. TL: Mindoro. hinunanganensis M. & T. Okano, 1986 (Fig. 37) leopardina Fruhstorfer, 1913 (Fig. 48) TS (TP): ST♂ (MNHN). nadenya Fruhstorfer, 1913 TL: Culasi, 800 ft, Antique, Panay. TS (TP): LT ♂ (SMFL). TL: Luzon. TS (TP): LT ♂ (ZMHU). time the first author visited to Senckenberg, he came bringing Japan; SK: Shun-ichi Kawamura, Tokyo, Japan; STA: Shinji pro Dophla evelina balabacana Tsukada, 1991, non TS (TP): ST ♂ (MNHN). TL: Hinunangan, Leyte. TL: Sulu Islands. Note: Though the original description does not mention TL: Luzon. TS (TP): HT ♂ (SMFL), PT 4♂4♀. Note: Description is based on six materials from Luzon, TS (TP): ST ♂ (BMNH). susoni Jumalon, 1975 (Fig. 102) Note: According to the original description, the type series his described type specimens, with his wife driving from Taga, Tokyo, Japan; TY: Takashi Yokochi, Aichi, Japan. Bassarona teuta balabacana Tsukada, 1991 dacasini Hanafusa, 1990 (Fig. 19) Note: Though the original description does not mention TS (TP): HT ♂ (FASU), PT 1♂. TS (TP): HT ♀ (BMNH). the number of types, a male syntype is preserved in MNHN. TS (TP): HT ♀ [untraced, not examined]. Mindoro, Camiguin de Mindanao, N. Mindanao. The lectotype Note: According to the original description, the type series TL: Camp 7, Minglanilla, Cebu. (male and female, but the number is unknown) are deposited in Specimens concerned. FW: Length of Forewing; HT: TL: Balabac. the number and sex, a syntype specimen in MNHN is a male. Note: Though the original description indicates that the panopus C. & R. Felder, 1861 (Fig. 76) designation was carried out for the Camiguin de Mindanao (number is unknown) are deposited in Semper and Felder TS (TP): HT ♂ (USC), PT 9♂9♀ (USC, JF, SMFL, TY). Staudinger collection (ZMHU). The lectotype designation was Holotype; LT: Lectotype; PLT: Paralectotype; PT: Paratype; boholensis (Lexias) M. & T. Okano, 1988 (Fig. 8) TS (TP): HT ♂ (HH), PT 1♀. hiwaga Nuyda & Kawamura, 1989 (Fig. 38) leucotaenia G. Semper, 1878 (Fig. 49) marinduque K. Okano, 1992 (Fig. 58) type female should be preserved in ZMHU, the specimen TL: Luzon. specimen by Schröder & Treadaway (1987). collections. carried out by Yokochi (1999). TL: Type Locality; TP: Type Preservation; TS: Type Series; TL: Bilar, Bohol. eson de Nicéville, 1894 (Fig. 29) TL: Camiguin, Babuyan. TL: Camotes. TL: Marinduque. corresponding with nadenya could not be found there. No TS (TP): ST ♂ (BMNH). tanagra Staudinger, 1889 (Fig. 103) ST: Syntype. TS (TP): HT ♂ (FASU), PT 6♂2♀. damalis Erichson, 1834 (Fig. 117) TL: Palawan. TS (TP): HT ♂ (SK), PT 7♂4♀ (SK, ET). TS (TP): LT ♂ (SMFL). TS (TP): HT ♀ (KMNH), PT 1♂ (KMNH). figure is presented in the original paper. Note: The original description does not mention the princesa Fruhstorfer, 1899 (Fig. 82) schoenigi Schröder & Treadaway, 1978 (Fig. 92) TL: Palawan. virginalis Fruhstorfer, 1913 TL: Luzon. TS (TP): LT ♂ (SMFL), PLT 1♀. Note: This taxon was described based on sixty-nine number of type series. TL: Palawan. TL: Amlan, 600 m, Negros. TS (TP): LT ♂ (ZMHU). TL: locality not given. boholensis (Euthalia) M. & T. Okano, 1990 (Fig. 9) TS (TP): ST ♂ [untraced, not examined]. Note: Description is based on a single pair, and all howarthi Jumalon, 1975 (Fig. 39) materials from Camotes, Bohol, Leyte, Panaon, E. Mindanao, medaga Fruhstorfer, 1913 (Fig. 59) negrosiana Schröder & Treadaway, 1987 (Fig. 66) TS (TP): HT ♀ (MNHN). TS (TP): HT ♀ (SMFL). Note: Description is based on seven males and eleven TS (TP): ST ♀ (BMNH) [not examined]. 3. LIST OF DESCRIBED SPECIES AND TL: Bilar, Bohol. Note: The type series should be preserved in ZMHU, but materials are preserved in SMFL. The lectotype designation TL: Amlan, Negros Oriental. and N. Mindanao. Yokochi (1998) mentioned the type series of TL: Camiguin de Mindanao. TL: Negros Or. [Oriental]. phelada G. Semper, [1887] (Fig. 77) females, and two males and three females are preserved in Note: This is a female form of panopus. The type locality SUBSPECIES NAME TS (TP): HT ♀ (FASU), PT 4♀. the specimens that corresponded with the types could not be was carried out by Yokochi (1998). TS (TP): HT ♂ (USC), PT 4♀ (USC, JF). leucotaenia was untraced so designated the lectotype for the TS (TP): ST ♂ (MNHN). TS (TP): HT ♂ (SMFL), PT 1♂1♀. TL: Luzon. proditrix Fruhstorfer, 1913 (Fig. 83) semperi Staudinger, 1889 (Fig. 93) ZMHU. The lectotype designation was carried out by Yokochi suspected is Mindanao, though the specific location is not found. The Fig. 117 is reproduced from the original description figure of the original description, but the type series specimen Note: Though the original description does not mention TS (TP): LT ♀ (SMFL). TL: Basilan. TL: Palawan. (1999). shown. The type series are deposited in the Rothschild collection The following list contains the taxonomic names bongaoensis Schröder & Treadaway, 1990 (Fig. 10) (pl. 50, fig. 4, ♂). eva C. & R. Felder, [1867] (Fig. 30) imperator Hewitson, 1863 (Fig. 40) was found after the third investigation in SMFL. We designate the number of types, a male syntype is preserved in MNHN. nirodha Fruhstorfer, 1913 (Fig. 67) Note: This taxon was described based on five materials TS (TP): ST ♂ (MNHN). TS (TP): LT ♂ (ZMHU). (BMNH, London), but we have not located them at present. concerned with Philippines. The names which are described TL: Bongao Peak, Bongao, Tawitawi Group. TL: Luzon. TL: Luzon. here again the lectotype for a male specimen shown in Fig. 49 TL: Basilan. from Luzon, Camiguin de Mindanao, and SE. Mindanao. Note: Description is based on males and females, and four tawitawia Treadaway & Nuyda, 1994 (Fig. 104) from the other areas are also included. TS (TP): HT ♀ (SMFL), PT 18♂20♀. darani Fruhstorfer, 1913 (Fig. 20) TS (TP): ST ♂ (BMNH). TS (TP): HT ♀ (BMNH). (labelled “Typus (red) / Camotes / Bohol / 113 / 211 / Coll. C. mindanaensis Schröder & Treadaway, 1978 (Fig. 60) TS (TP): ST ♂♀ (MNHN). Yokochi (1998) mentioned the type series of phelada was producta Butler, 1901 (Fig. 84) males and four females are preserved in ZMHU. The lectotype TL: Magsaggao, Tawitawi. visayana (Lexias) Schröder & Treadaway, 1987 (Fig. 110) TL: Sulu Archipelago. Note: According to the original description, eva is distrib- Semper”). TL: Surigao del Sur, Mindanao. untraced so designated the lectotype for the figure of the TL: Philippines [recte W. Malaysia]. designation was carried out by Yokochi (1999). TS (TP): HT ♂ (SMFL). TL: Catmon, S. Leyte. albusequus Nihira & Kawamura, 1986 (Fig. 2) borromeoi Schröder, 1977 (Fig. 11) TS (TP): HT ♀ (BMNH). uted in Assam, Luzon, and Celebes. The details of type series ingae (Lexias) Schröder & Treadaway, 1987 (Fig. 41) TS (TP): HT ♂ (SMFL). obatrata Yokochi, 1994 (Fig. 68) original description, but the target specimen was found after the TS (TP): ST ♀ (BMNH). TS (TP): HT ♂ (SMFL), PT 6♂4♀. TL: Lakit lakit, Sanga Sanga, Tawitawi. TL: Espana, 2000 ft, Sibuyan. Note: Though the original description does not mention are not mentioned. Staudinger (1886) is the first revisory of TL: Negros Or. [Oriental], 800 m. leytana Schröder & Treadaway, 1982 (Fig. 50) TL: Dumaran. third investigation in SMFL. We designate here again the senilis Hanafusa, 1992 (Fig. 94) tethys Tsukada, 1991 (Fig. 105) TS (TP): HT ♀ (SK). TS (TP): HT ♂ (SMFL), PT 1♀. the number of type series, one female would be the holotype. which Luzon is recognized as a locality of eva. TS (TP): HT ♂ (SMFL), PT 5♂5♀. TL: St. Bernard, Calpio, 280 m, S. Leyte. mindorana (Tanaecia) Fruhstorfer, 1904 (Fig. 61) TS (TP): HT ♀ (HH), PT 1♂1♀. lectotype for a female specimen shown in Fig. 77 (labelled rhamases Staudinger, 1889 (Fig. 85) TL: Dumaran. TL: N. Palawan. visayana (Cynitia) Schröder & Treadaway, 1981 (Fig. 111) TS (TP): HT ♂ (SMFL), PT 18♂10♀. TL: Mindoro. “Typus (red) / Euth. phelada typ. ♀ Semper / Marivcles (?) TL: Palawan. TS (TP): HT ♂ (HH), PT 1♂. TS (TP): HT ♂ (ET), PT 10♂12♀. TL: St. Bernard, Catmon, S. Leyte. 8 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 9

1. INTRODUCTION Limbach, about 250 km from Frankfurt. It is an undoubted fact amlana Jumalon, 1970 (Fig. 3) cabigoni Schröder, Treadaway, A. Mohagan & N. Mohagan, dhayma Fruhstorfer, 1913 (Fig. 21) exul Tsukada & Nishiyama, 1980 (Fig. 31) ingae (Euthalia) Schröder & Treadaway, 1990 (Fig. 42) leytensis Jumalon, 1975 (Fig. 51) TS (TP): ST 2♂1♀ (BMNH). ormocana Jumalon, 1970 (Fig. 69) 11.11.81 / 110 / Coll. C. Semper / Lectotype ♀ Euthalia TS (TP): LT ♂ (ZMHU). sibutana Schröder & Treadaway, 1990 (Fig. 95) Note: Attached label to the holotype specimen shows TS (TP): HT ♂ (SMFL), PT 10♂2♀. waltraudae Schröder & Treadaway, 1990 (Fig. 114) that Mr. Treadaway and Dr. H. Schröder of Senckenberg TL: Amlan, Negros Oriental. 2009 (Fig. 12) TL: Sulu Archipelago. TL: Bo. Melgar, Dinagat. TL: Bongao Peak, 200 m, Bongao, Tawitawi Group. TL: Lake Danao, Ormoc City, Leyte. Note: The figured female (Fig. 61) is preserved in the TL: Lake Danao, Ormoc City, Leyte. phelada Semper [87]:1888 Designated by T. Yokochi, 1997”). Note: Description is based on males, and four males are TL: Cavan Cavan, 5 m, Sibutu, Tawitawi Group. “Balabac”, but the correct type locality is Palawan. TL: Indalawan, Balabac. The Philippine Islands are situated in the northern end of thoroughly deserve credit for working further on systematic TS (TP): HT ♂ (USC), PT10 ♂1♀ (USC, JF). TL: Balut Is, Sarangani Group. TS (TP): HT ♀ (BMNH). TS (TP): HT ♂ (ET), PT 3♂6♀. TS (TP): HT ♀ (SMFL), PT 10♂12♀. TS (TP): HT ♀ (USC). Rothschild collection (Ro 98/7), BMNH. It might be one of the TS (TP): HT ♂ (USC), PT 1♀. preserved in ZMHU. The lectotype designation was carried out TS (TP): HT ♂ (SMFL), PT 17♂15♀. vistrica Fruhstorfer, 1913 (Fig. 112) TS (TP): HT ♀ (SMFL), PT 2♂2♀. the Malay Archipelago and are composed of more than 7,000 classification in Philippines Rhopalocera. TS (TP): HT ♂ (SMFL), PT 2♂1♀. Note: Though the original description does not mention the types, although the attached label is only “Mindoro”. No other philippensis Fruhstorfer, 1899 (Fig. 78) by Yokochi (1999). trapesa G. Semper, [1887] (Fig. 106) TL: locality unknown, presumably one of the Central islands, large and small, extending from north to south about Here we revised the group of the tribe Adoliadini antiquea Schröder & Treadaway, 1980 (Fig. 4) number of type series, one female would be the holotype. felicitacionae Jumalon, 1989 (Fig. 32) irauana Jumalon, 1975 (Fig. 43) liaoi Schröder & Treadaway, 1982 (Fig. 52) specimen is found that corresponded with the type series. ornata Schröder & Treadaway, 1979 (Fig. 70) TL: Basilan. sibuyana Schröder & Treadaway, 1982 (Fig. 96) TL: Camiguin de Mindanao. Philippine islands [recte Dinagat]. yoshiroui K. Okano, 1992 (Fig. 115) 1,800 km and from east to west about 1,100 km. The insect distributed in the Philippines, based on the checklist by TL: Culasi, 500 m, Antique, Panay. calliphorus C. & R. Felder, 1861 (Fig. 13) TL: Kantakpal Creek, Awihao, Ormoc, Western Leyte. TL: Irauan [Irawan], Puerto Princesa, Palawan. TL: Mt. Madja-an [Mt. Madja-as], 500 m, Antique, Panay. TL: Romblon. TS (TP): LT ♂ (MNHN). rodriguezi Schröder & Treadaway, 1982 (Fig. 86) TL: Sibuyan, Romblon Group. TS (TP): LT ♂ (SMFL). TS (TP): ST ♂ (BMNH). TL: Babuyan. fauna is rich in diversity and the tribe Adoliadini, Nymphalidae, Treadaway (1995). One hundred-twenty taxa of type specimens TS (TP): HT ♂ (SMFL). TL: Luzon. dinagatensis Tsukada, 1991 (Fig. 22) TS (TP): HT ♂ (JF). TS (TP): HT ♀ (JF), PT 1♂1♀ (USC). TS (TP): HT ♂ (SMFL), PT 1♂2♀. mindorana (Euthalia) Fruhstorfer, 1899 (Fig. 62) TS (TP): HT ♂ (SMFL), PT 1♂2♀. Note: Description is based on one male from Basilan TL: 28 km W of Puerto Princesa, Palawan. TS (TP): HT ♂ (SMFL), PT 2♂5♀. Note: Description is based on seven materials from E. Note: Tsukada (1991) suspected this taxon is distributed TS (TP): HT ♂ (KMNH). Lepidoptera, the main group of this study, is no exception. The founded in the Philippines are examined and revised the TS (TP): ST ♂ (BMNH). TL: Dinagat. Note: The holotype is deposited in Jumalon family collec- TL: Mindoro. (MNHN) and one female from Luzon (ZMHU). As the TS (TP): HT ♂ (SMFL), PT 2♂2♀. Mindanao. The lectotype designation was carried out for the in Basilan, but it presumably comes from Dinagat. important works concerned with Philippines were, back in classification of species-subspecies relation. This paper is aquamarina Fruhstorfer, 1913 (Fig. 5) TS (TP): HT ♂ (ET), PT 2♂1♀. gahiti M. & T. Okano, 1990 (unavailable name) (Fig. 33) tion. ludonia Staudinger, 1889 (Fig. 53) TS (TP): ST (MNHN). palawana (Lexias) Moore, 1897 (Fig. 71) Luzon-group should be classified as nadenya. We select a smaragdifera Fruhstorfer, 1913 (Fig. 97) Camiguin de Mindanao specimen by Yokochi (1998). yui Yokochi, 1994 (Fig. 116) history, “Diagnosen einiger neuer Tagfalter von den Philippinen published in two parts. All the taxa described before are TL: Mindanao. candida Schröder, Treadaway & Nuyda, 1990 (Fig. 14) TL: Bilar, Bohol. TL: Palawan. TL: Palawan. male (Fig. 78) deposited in MNHN as the lectotype, labelled, romeo Schröder & Treadaway, 1987 (Fig. 87) TL: Locality unknown, presumably Mindoro [Mindoro]. volupia Tsukada & Nishiyama, 1981 (Fig. 113) TL: Dumaran. und Bemerkungen über die Verbreitung einiger derselben” (G. examined and figured as much as possible in part 1. In the TS (TP): LT ♂ (SMFL). TL: 30 m NN, Calamian Group. dinorah Fruhstorfer, 1899 (Fig. 23) TS (TP): HT ♀ (FASU). joloana Staudinger, 1889 (Fig. 44) TS (TP): LT ♂ (ZMHU). mindorana (Euthalia) Tsukada, 1991 (permanently invalid) TS (TP): HT ♂ (BMNH). “Type (red) / lubentina philippensis Fruhst. / Philippinen TL: Mt. Halcon, N. Mindoro. TS (TP): ST (MNHN). treadawayi Tsukada, 1991 (Fig. 107) TL: Camiguin. TS (TP): HT ♂ (TY), PT 11♂4♀. Semper, 1886–1892), and then “Die Gross-Schmetterlinge der second part, synonymic lists and some important specimens Note: Description is based on some specimens. The lecto- TS (TP): HT ♂ (SMFL), PT10 ♂. TL: Basilan. Note: As this taxon gahiti was described as a form of TL: Sulu. Note: Description is based on four males and six females, Note: The taxon mindorana described under the genus Bazilan II-III, 98 Doherty ex. coll. H. Fruhstorfer / TS (TP): HT ♀ (SMFL), PT 1♀. Note: Though the original description does not mention TL: Romblon, Sibuyanes. TS (TP): HT ♂ (ET), PT 2♂5♀. (to be continued) Erde, 9” (1908–1927) edited by A. Seitz. Recently, a Japanese will be shown, affixing several discussions. type designation was carried out by Yokochi (1998). TS (TP): ST 2♂2♀ (MNHN). Lexias panopus boholensis (infrasubspecific name), it should TS (TP): LT ♂ (ZMHU). and two pairs are preserved in ZMHU. The lectotype designa- Euthalia is a junior primary homonym of Euthalia lusiada palawana (Tanaecia) Staudinger, 1889 (Fig. 72) MUSEUM PARIS 1934 COLL H Fruhstorfer”. the number and sex, a syntype specimen in MNHN is a male. TS (TP): HT ♂ (SMFL). entomologist, E. Tsukada, had compiled a great series of cavarna Fruhstorfer, 1913 (Fig. 15) be treated as an unavailable name. Note: Description is based on two males and one female, tion was carried out by Yokochi (1999). mindorana Fruhstorfer, 1899 (permanently invalid name). TL: Palawan. rudraca Fruhstorfer, 1913 (Fig. 88) “Butterflies of the South East Asian Islands” (1980–1991). balabacana (Bassarona) Tsukada, 1991 (Fig. 6) TL: Balabas [Balabac]. dodong Schröder & Treadaway, 1978 (Fig. 24) and all of them are preserved in ZMHU. The lectotype desig- Now it is replaced by tsukada Koçak (1996). TS (TP): LT ♂ (ZMHU). phlegethon G. Semper, [1887] (Fig. 79) TL: Balabac. soloni M. & T. Okano, 1990 (Fig. 98) tsukada Koçak, 1996 (Fig. 80) The late Prof. Julian N. Jumalon (1909–2000) (Fig. 1, TL: Balabac. TS (TP): LT ♀ (ZMHU). TL: Mobo, 1000 ft, Masbate. galoa Fruhstorfer, 1913 (Fig. 34) nation was carried out by Yokochi (1999). lupina Druce, 1874 (Fig. 54) Note: Description is based on males and females, and six TL: Mindanao. TS (TP): LT ♂ (ZMHU). TL: Bilar, Bohol. TL: Mindoro. left), the Lepidoptera Research Associate and Lecturer of 2. ABBREVIATIONS TS (TP): HT ♂ (ET), PT 8♂6♀. Note: Description is based on a few females, and one spec- TS (TP): HT ♂ (SMFL), PT 3♂7♀. TL: Mindoro. TL: Philippine Islands [Sulu]. mindorensis Schröder & Treadaway, 1982 (Fig. 63) males and three females are preserved in ZMHU. The lecto- TS (TP): LT ♀ (SMFL). Note: Description is based on some specimens, and two TS (TP): HT ♂ (FASU), PT 2♂3♀. TS (TP): HT ♂ (ET), PT 10♂11♀. Department of Biology, University of San Carlos, published a imen is preserved in ZMHU. The lectotype designation was TS (TP): ST (BMNH). kayumanggia Treadaway & Nuyda, 1994 (Fig. 45) TS (TP): HT ♀ (BMNH). TL: Mindoro. type designation was carried out by Yokochi (1999). Note: Description is based on twenty-one males and seven males and two females are preserved in ZMHU. The lectotype Note: Euthalia lubentina mindorana Tsukada, 1991 is a paper of “New Philippine butterflies” in 1975, and many new Public Institution (Museum, University). BMNH: The balabacana (Dophla) Tsukada, 1991 (secondary homonym) carried out by Yokochi (1999). dohertyi Butler, 1901 (Fig. 25) Note: Though the original description does not mention TL: Mt. Bayog, S. Palawan. TS (TP): HT ♂ (SMFL), PT 6♂7♀. females from Samar, Camiguin de Mindanao, N. Mindanao, E. designation was carried out by Yokochi (1999). soregina Fruhstorfer, 1913 (Fig. 99) permanently invalid name (nom. nov. pro mindorana Tsukada, taxonomic names concerned with various families were Natural History Museum, London, United Kingdom; FASU: (Fig. 7) TL: Sulu Archipelago. the number and sex, syntype specimen in BMNH is a male. TS (TP): HT ♂ (SMFL), PT 13♂4♀. lusiada C. & R. Felder, 1863 (Fig. 55) palawana (Euthalia) Staudinger, 1889 (Fig. 73) Mindanao, SE. Mindanao, and several materials are preserved TL: Sulu Archipelago. 1991, non Fruhstorfer,1899). New name tsukada was proposed described there. He played a leading part of the study of Exhibition Hall of Food, Agricultural and Environmental Note: See below balabacensis. chloe Schröder & Treadaway, 1990 (Fig. 16) TS (TP): HT ♂ (BMNH). TL: Luzon. miscus Fruhstorfer, 1913 (Fig. 64) TL: Palawan. in SMFL. The lectotype designation was carried out for the samarensis Tsukada, 1991 (Fig. 89) TS (TP): ST ♀ (BMNH). by Koçak (1996). Philippine butterflies and contributed greatly to entomology. Sciences, Faculty of Agriculture, Shinshu University, Nagano, TL: Jao Asan, 200 m, Sibuyan, Romblon Group. goertzi Jumalon, 1975 (Fig. 35) kulaya Treadaway & Nuyda, 1994 (Fig. 46) TS (TP): ST ♂ (SMFL). TL: Mindanao. TS (TP): LT ♂ (ZMHU). Mindanao specimen by Yokochi (1998). TL: San Juan, Samar. Note: According to the original description, the type series In 1995, the late Mr. Colin G. Treadaway (Fig. 1, right, Japan; KMNH: Kitakyushu Museum of Natural History & balabacensis nom. nov. (Fig. 7) TS (TP): HT ♂ (SMFL), PT 11♂9♀. dumaranensis Hanafusa, 1992 (Fig. 26) TL: Amlan, Negros Oriental. TL: Magellanes, Homonhon. Note: Though the original description does not mention TS (TP): ST (MNHN, BMNH). Note: Description is based on one male and six females, TS (TP): HT ♂ (ET), PT 3♂2♀. are exactly female, but the number is unknown. tyawena Fruhstorfer, 1913 (Fig. 108) died in 2019) published “Checklist of the butterflies of the Human History, Fukuoka, Japan; MNHN: Museum National TL: Balabac. TL: Dumaran. TS (TP): HT ♀ (ET). TS (TP): HT ♂ (SMFL), PT 14♂10♀. the number of types, a male syntype is preserved in SMFL. Note: According to the original description, types and one male and four females are preserved in ZMHU. The pinamalayana Yokochi, 1997 (Fig. 80) TL: unknown [Palawan]. Philippine Islands (Lepidoptera: Rhopalocera)”. Although, 25 d'Histoire Naturelle Entomologie, Paris, France; SMFL: TS (TP): HT ♂ (ET), PT 2♂2♀. circe Schröder & Treadaway, 1990 (Fig. 17) TS (TP): HT ♂ (HH). Note: Though the original description mentions the Note: The name kulaya is spelt “kúlaya” in the original (syntypes) are preserved in Fruhstorfer collection (now in lectotype designation was carried out by Yokochi (1999). Note: The new name pinamalayana was proposed by sarmana Fruhstorfer, 1913 (Fig. 90) subpiratica Schröder & Treadaway, 1987 (Fig. 100) TS (TP): LT ♀ (ZMHU). years have passed since his paper presented, several new Senckenberg Forschungsinstitut und Naturmuseum Frankfurt, Note: The taxon balabacana was described under the TL: Cavan Cavan, Sibutu, Tawitawi Group. holotype is preserved in USC, it is deposited in ET. description. macer Tsukada & Nishiyama, 1980 (Fig. 56) MNHN) and BMNH. Yokochi (1997), because mindorana Tsukada, 1991 was a TL: Basilan. TL: Nueva Ecija, Bongabon, Centr. Luzon. Note: Description is based on some specimens, and one findings are still remained unpublished — it can be followed Frankfurt-am-Main, Germany; USC: University of San Carlos, genus Dophla. As Yokochi (2010) assigned both Dophla and TS (TP): HT ♂ (SMFL), PT 9♂10♀. dunya Doubleday, [1848] (Fig. 27) TL: Bo. Melgar, Dinagat. pallida Schröder, Treadaway & Nuyda, 1990 (Fig. 74) permanently invalid name. However, pinamalayana is a junior TS (TP): ST (MNHN). TS (TP): HT ♂ (SMFL), PT1♀. male and one female are preserved in ZMHU. The lectotype by his own handwriting on the glass surface of his specimen Cebu, Philippines; ZMHU: Zoologisches Museum, Humboldt Bassarona to the genus Euthalia, balabacana (Dophla) is a TL: Assam, India. hikarugenzi Tsukada & Nishiyama, 1980 (Fig. 36) laetitiae Schröder & Treadaway, 2000 (Fig. 47) TS (TP): HT ♂ (ET), PT 3♂2♀. monara Fruhstorfer, 1913 (Fig. 65) TL: 30 m NN, Calamian Group. synonym of tsukada Koçak (1996). Note: Though the original description does not mention designation was carried out by Yokochi (1999). cases —. The first author visited Senckenberg Museum in Universität, Berlin, Germany. junior secondary homonym of Bassarona teuta balabacana cusama Fruhstorfer, 1913 (Fig. 18) TS (TP): ST ♂ (BMNH). TL: near Baguio, N. Luzon. TL: Cavan Cavan, 5 m, Sibutu, Tawitawi Group. TL: N. Borneo. TS (TP): HT ♂ (SMFL), PT 11♂16♀. the number and sex, a syntype specimen in MNHN is a male. suluana Fruhstorfer, 1902 (Fig. 101) Frankfurt to research on the Treadaway collection on March Private collections. ET: Etsuzo Tsukada, Yamanashi, Tsukada, 1991. Here a new name balabacensis is replaced for TL: Mindanao, Camiguin de Mindanao. TS (TP): HT ♂ (MKN), PT 1♀ (MKN). TS (TP): HT ♂ (SMFL), PT 4♂. malissia Fruhstorfer, 1913 (Fig. 57) TS (TP): ST (MNHN). piratica G. Semper, [1887] (Fig. 81) TL: Sulu Archipelago. uposatha Fruhstorfer, 1913 (Fig. 109) 2019, and just then was informed that he had died several Japan; HH: Hiroto Hanafusa, Tottori, Japan; JF: Jumalon Dophla evelina balabacana Tsukada, 1991. TS (TP): ST ♂ (MNHN), ♀ (SMFL). ellora Fruhstorfer, 1898 (Fig. 28) TL: Basilan. panayana Schröder & Treadaway, 1980 (Fig. 75) TL: Camiguin de Mindanao. satrapes C. & R. Felder, 1861 (Fig. 91) TS (TP): HT ♀ (BMNH). TL: Banggi (Banguey), E. Malaysia. weeks ago at his home in England. In the middle 1990’s, every Family, Cebu, Philippines; MKN: Misao Kaneko, Tokyo, Euthalia (Dophla) evelina balabacensis nom. nov. Note: Figured specimen is a syntype female in SMFL. TL: Mindoro. hinunanganensis M. & T. Okano, 1986 (Fig. 37) leopardina Fruhstorfer, 1913 (Fig. 48) TS (TP): ST♂ (MNHN). nadenya Fruhstorfer, 1913 TL: Culasi, 800 ft, Antique, Panay. TS (TP): LT ♂ (SMFL). TL: Luzon. TS (TP): LT ♂ (ZMHU). time the first author visited to Senckenberg, he came bringing Japan; SK: Shun-ichi Kawamura, Tokyo, Japan; STA: Shinji pro Dophla evelina balabacana Tsukada, 1991, non TS (TP): ST ♂ (MNHN). TL: Hinunangan, Leyte. TL: Sulu Islands. Note: Though the original description does not mention TL: Luzon. TS (TP): HT ♂ (SMFL), PT 4♂4♀. Note: Description is based on six materials from Luzon, TS (TP): ST ♂ (BMNH). susoni Jumalon, 1975 (Fig. 102) Note: According to the original description, the type series his described type specimens, with his wife driving from Taga, Tokyo, Japan; TY: Takashi Yokochi, Aichi, Japan. Bassarona teuta balabacana Tsukada, 1991 dacasini Hanafusa, 1990 (Fig. 19) Note: Though the original description does not mention TS (TP): HT ♂ (FASU), PT 1♂. TS (TP): HT ♀ (BMNH). the number of types, a male syntype is preserved in MNHN. TS (TP): HT ♀ [untraced, not examined]. Mindoro, Camiguin de Mindanao, N. Mindanao. The lectotype Note: According to the original description, the type series TL: Camp 7, Minglanilla, Cebu. (male and female, but the number is unknown) are deposited in Specimens concerned. FW: Length of Forewing; HT: TL: Balabac. the number and sex, a syntype specimen in MNHN is a male. Note: Though the original description indicates that the panopus C. & R. Felder, 1861 (Fig. 76) designation was carried out for the Camiguin de Mindanao (number is unknown) are deposited in Semper and Felder TS (TP): HT ♂ (USC), PT 9♂9♀ (USC, JF, SMFL, TY). Staudinger collection (ZMHU). The lectotype designation was Holotype; LT: Lectotype; PLT: Paralectotype; PT: Paratype; boholensis (Lexias) M. & T. Okano, 1988 (Fig. 8) TS (TP): HT ♂ (HH), PT 1♀. hiwaga Nuyda & Kawamura, 1989 (Fig. 38) leucotaenia G. Semper, 1878 (Fig. 49) marinduque K. Okano, 1992 (Fig. 58) type female should be preserved in ZMHU, the specimen TL: Luzon. specimen by Schröder & Treadaway (1987). collections. carried out by Yokochi (1999). TL: Type Locality; TP: Type Preservation; TS: Type Series; TL: Bilar, Bohol. eson de Nicéville, 1894 (Fig. 29) TL: Camiguin, Babuyan. TL: Camotes. TL: Marinduque. corresponding with nadenya could not be found there. No TS (TP): ST ♂ (BMNH). tanagra Staudinger, 1889 (Fig. 103) ST: Syntype. TS (TP): HT ♂ (FASU), PT 6♂2♀. damalis Erichson, 1834 (Fig. 117) TL: Palawan. TS (TP): HT ♂ (SK), PT 7♂4♀ (SK, ET). TS (TP): LT ♂ (SMFL). TS (TP): HT ♀ (KMNH), PT 1♂ (KMNH). figure is presented in the original paper. Note: The original description does not mention the princesa Fruhstorfer, 1899 (Fig. 82) schoenigi Schröder & Treadaway, 1978 (Fig. 92) TL: Palawan. virginalis Fruhstorfer, 1913 TL: Luzon. TS (TP): LT ♂ (SMFL), PLT 1♀. Note: This taxon was described based on sixty-nine number of type series. TL: Palawan. TL: Amlan, 600 m, Negros. TS (TP): LT ♂ (ZMHU). TL: locality not given. boholensis (Euthalia) M. & T. Okano, 1990 (Fig. 9) TS (TP): ST ♂ [untraced, not examined]. Note: Description is based on a single pair, and all howarthi Jumalon, 1975 (Fig. 39) materials from Camotes, Bohol, Leyte, Panaon, E. Mindanao, medaga Fruhstorfer, 1913 (Fig. 59) negrosiana Schröder & Treadaway, 1987 (Fig. 66) TS (TP): HT ♀ (MNHN). TS (TP): HT ♀ (SMFL). Note: Description is based on seven males and eleven TS (TP): ST ♀ (BMNH) [not examined]. 3. LIST OF DESCRIBED SPECIES AND TL: Bilar, Bohol. Note: The type series should be preserved in ZMHU, but materials are preserved in SMFL. The lectotype designation TL: Amlan, Negros Oriental. and N. Mindanao. Yokochi (1998) mentioned the type series of TL: Camiguin de Mindanao. TL: Negros Or. [Oriental]. phelada G. Semper, [1887] (Fig. 77) females, and two males and three females are preserved in Note: This is a female form of panopus. The type locality SUBSPECIES NAME TS (TP): HT ♀ (FASU), PT 4♀. the specimens that corresponded with the types could not be was carried out by Yokochi (1998). TS (TP): HT ♂ (USC), PT 4♀ (USC, JF). leucotaenia was untraced so designated the lectotype for the TS (TP): ST ♂ (MNHN). TS (TP): HT ♂ (SMFL), PT 1♂1♀. TL: Luzon. proditrix Fruhstorfer, 1913 (Fig. 83) semperi Staudinger, 1889 (Fig. 93) ZMHU. The lectotype designation was carried out by Yokochi suspected is Mindanao, though the specific location is not found. The Fig. 117 is reproduced from the original description figure of the original description, but the type series specimen Note: Though the original description does not mention TS (TP): LT ♀ (SMFL). TL: Basilan. TL: Palawan. (1999). shown. The type series are deposited in the Rothschild collection The following list contains the taxonomic names bongaoensis Schröder & Treadaway, 1990 (Fig. 10) (pl. 50, fig. 4, ♂). eva C. & R. Felder, [1867] (Fig. 30) imperator Hewitson, 1863 (Fig. 40) was found after the third investigation in SMFL. We designate the number of types, a male syntype is preserved in MNHN. nirodha Fruhstorfer, 1913 (Fig. 67) Note: This taxon was described based on five materials TS (TP): ST ♂ (MNHN). TS (TP): LT ♂ (ZMHU). (BMNH, London), but we have not located them at present. concerned with Philippines. The names which are described TL: Bongao Peak, Bongao, Tawitawi Group. TL: Luzon. TL: Luzon. here again the lectotype for a male specimen shown in Fig. 49 TL: Basilan. from Luzon, Camiguin de Mindanao, and SE. Mindanao. Note: Description is based on males and females, and four tawitawia Treadaway & Nuyda, 1994 (Fig. 104) from the other areas are also included. TS (TP): HT ♀ (SMFL), PT 18♂20♀. darani Fruhstorfer, 1913 (Fig. 20) TS (TP): ST ♂ (BMNH). TS (TP): HT ♀ (BMNH). (labelled “Typus (red) / Camotes / Bohol / 113 / 211 / Coll. C. mindanaensis Schröder & Treadaway, 1978 (Fig. 60) TS (TP): ST ♂♀ (MNHN). Yokochi (1998) mentioned the type series of phelada was producta Butler, 1901 (Fig. 84) males and four females are preserved in ZMHU. The lectotype TL: Magsaggao, Tawitawi. visayana (Lexias) Schröder & Treadaway, 1987 (Fig. 110) TL: Sulu Archipelago. Note: According to the original description, eva is distrib- Semper”). TL: Surigao del Sur, Mindanao. untraced so designated the lectotype for the figure of the TL: Philippines [recte W. Malaysia]. designation was carried out by Yokochi (1999). TS (TP): HT ♂ (SMFL). TL: Catmon, S. Leyte. albusequus Nihira & Kawamura, 1986 (Fig. 2) borromeoi Schröder, 1977 (Fig. 11) TS (TP): HT ♀ (BMNH). uted in Assam, Luzon, and Celebes. The details of type series ingae (Lexias) Schröder & Treadaway, 1987 (Fig. 41) TS (TP): HT ♂ (SMFL). obatrata Yokochi, 1994 (Fig. 68) original description, but the target specimen was found after the TS (TP): ST ♀ (BMNH). TS (TP): HT ♂ (SMFL), PT 6♂4♀. TL: Lakit lakit, Sanga Sanga, Tawitawi. TL: Espana, 2000 ft, Sibuyan. Note: Though the original description does not mention are not mentioned. Staudinger (1886) is the first revisory of TL: Negros Or. [Oriental], 800 m. leytana Schröder & Treadaway, 1982 (Fig. 50) TL: Dumaran. third investigation in SMFL. We designate here again the senilis Hanafusa, 1992 (Fig. 94) tethys Tsukada, 1991 (Fig. 105) TS (TP): HT ♀ (SK). TS (TP): HT ♂ (SMFL), PT 1♀. the number of type series, one female would be the holotype. which Luzon is recognized as a locality of eva. TS (TP): HT ♂ (SMFL), PT 5♂5♀. TL: St. Bernard, Calpio, 280 m, S. Leyte. mindorana (Tanaecia) Fruhstorfer, 1904 (Fig. 61) TS (TP): HT ♀ (HH), PT 1♂1♀. lectotype for a female specimen shown in Fig. 77 (labelled rhamases Staudinger, 1889 (Fig. 85) TL: Dumaran. TL: N. Palawan. visayana (Cynitia) Schröder & Treadaway, 1981 (Fig. 111) TS (TP): HT ♂ (SMFL), PT 18♂10♀. TL: Mindoro. “Typus (red) / Euth. phelada typ. ♀ Semper / Marivcles (?) TL: Palawan. TS (TP): HT ♂ (HH), PT 1♂. TS (TP): HT ♂ (ET), PT 10♂12♀. TL: St. Bernard, Catmon, S. Leyte. 10 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 11

1. INTRODUCTION Limbach, about 250 km from Frankfurt. It is an undoubted fact amlana Jumalon, 1970 (Fig. 3) cabigoni Schröder, Treadaway, A. Mohagan & N. Mohagan, dhayma Fruhstorfer, 1913 (Fig. 21) exul Tsukada & Nishiyama, 1980 (Fig. 31) ingae (Euthalia) Schröder & Treadaway, 1990 (Fig. 42) leytensis Jumalon, 1975 (Fig. 51) TS (TP): ST 2♂1♀ (BMNH). ormocana Jumalon, 1970 (Fig. 69) 11.11.81 / 110 / Coll. C. Semper / Lectotype ♀ Euthalia TS (TP): LT ♂ (ZMHU). sibutana Schröder & Treadaway, 1990 (Fig. 95) Note: Attached label to the holotype specimen shows TS (TP): HT ♂ (SMFL), PT 10♂2♀. waltraudae Schröder & Treadaway, 1990 (Fig. 114) that Mr. Treadaway and Dr. H. Schröder of Senckenberg TL: Amlan, Negros Oriental. 2009 (Fig. 12) TL: Sulu Archipelago. TL: Bo. Melgar, Dinagat. TL: Bongao Peak, 200 m, Bongao, Tawitawi Group. TL: Lake Danao, Ormoc City, Leyte. Note: The figured female (Fig. 61) is preserved in the TL: Lake Danao, Ormoc City, Leyte. phelada Semper [87]:1888 Designated by T. Yokochi, 1997”). Note: Description is based on males, and four males are TL: Cavan Cavan, 5 m, Sibutu, Tawitawi Group. “Balabac”, but the correct type locality is Palawan. TL: Indalawan, Balabac. The Philippine Islands are situated in the northern end of thoroughly deserve credit for working further on systematic TS (TP): HT ♂ (USC), PT10 ♂1♀ (USC, JF). TL: Balut Is, Sarangani Group. TS (TP): HT ♀ (BMNH). TS (TP): HT ♂ (ET), PT 3♂6♀. TS (TP): HT ♀ (SMFL), PT 10♂12♀. TS (TP): HT ♀ (USC). Rothschild collection (Ro 98/7), BMNH. It might be one of the TS (TP): HT ♂ (USC), PT 1♀. preserved in ZMHU. The lectotype designation was carried out TS (TP): HT ♂ (SMFL), PT 17♂15♀. vistrica Fruhstorfer, 1913 (Fig. 112) TS (TP): HT ♀ (SMFL), PT 2♂2♀. the Malay Archipelago and are composed of more than 7,000 classification in Philippines Rhopalocera. TS (TP): HT ♂ (SMFL), PT 2♂1♀. Note: Though the original description does not mention the types, although the attached label is only “Mindoro”. No other philippensis Fruhstorfer, 1899 (Fig. 78) by Yokochi (1999). trapesa G. Semper, [1887] (Fig. 106) TL: locality unknown, presumably one of the Central islands, large and small, extending from north to south about Here we revised the group of the tribe Adoliadini antiquea Schröder & Treadaway, 1980 (Fig. 4) number of type series, one female would be the holotype. felicitacionae Jumalon, 1989 (Fig. 32) irauana Jumalon, 1975 (Fig. 43) liaoi Schröder & Treadaway, 1982 (Fig. 52) specimen is found that corresponded with the type series. ornata Schröder & Treadaway, 1979 (Fig. 70) TL: Basilan. sibuyana Schröder & Treadaway, 1982 (Fig. 96) TL: Camiguin de Mindanao. Philippine islands [recte Dinagat]. yoshiroui K. Okano, 1992 (Fig. 115) 1,800 km and from east to west about 1,100 km. The insect distributed in the Philippines, based on the checklist by TL: Culasi, 500 m, Antique, Panay. calliphorus C. & R. Felder, 1861 (Fig. 13) TL: Kantakpal Creek, Awihao, Ormoc, Western Leyte. TL: Irauan [Irawan], Puerto Princesa, Palawan. TL: Mt. Madja-an [Mt. Madja-as], 500 m, Antique, Panay. TL: Romblon. TS (TP): LT ♂ (MNHN). rodriguezi Schröder & Treadaway, 1982 (Fig. 86) TL: Sibuyan, Romblon Group. TS (TP): LT ♂ (SMFL). TS (TP): ST ♂ (BMNH). TL: Babuyan. fauna is rich in diversity and the tribe Adoliadini, Nymphalidae, Treadaway (1995). One hundred-twenty taxa of type specimens TS (TP): HT ♂ (SMFL). TL: Luzon. dinagatensis Tsukada, 1991 (Fig. 22) TS (TP): HT ♂ (JF). TS (TP): HT ♀ (JF), PT 1♂1♀ (USC). TS (TP): HT ♂ (SMFL), PT 1♂2♀. mindorana (Euthalia) Fruhstorfer, 1899 (Fig. 62) TS (TP): HT ♂ (SMFL), PT 1♂2♀. Note: Description is based on one male from Basilan TL: 28 km W of Puerto Princesa, Palawan. TS (TP): HT ♂ (SMFL), PT 2♂5♀. Note: Description is based on seven materials from E. Note: Tsukada (1991) suspected this taxon is distributed TS (TP): HT ♂ (KMNH). Lepidoptera, the main group of this study, is no exception. The founded in the Philippines are examined and revised the TS (TP): ST ♂ (BMNH). TL: Dinagat. Note: The holotype is deposited in Jumalon family collec- TL: Mindoro. (MNHN) and one female from Luzon (ZMHU). As the TS (TP): HT ♂ (SMFL), PT 2♂2♀. Mindanao. The lectotype designation was carried out for the in Basilan, but it presumably comes from Dinagat. important works concerned with Philippines were, back in classification of species-subspecies relation. This paper is aquamarina Fruhstorfer, 1913 (Fig. 5) TS (TP): HT ♂ (ET), PT 2♂1♀. gahiti M. & T. Okano, 1990 (unavailable name) (Fig. 33) tion. ludonia Staudinger, 1889 (Fig. 53) TS (TP): ST (MNHN). palawana (Lexias) Moore, 1897 (Fig. 71) Luzon-group should be classified as nadenya. We select a smaragdifera Fruhstorfer, 1913 (Fig. 97) Camiguin de Mindanao specimen by Yokochi (1998). yui Yokochi, 1994 (Fig. 116) history, “Diagnosen einiger neuer Tagfalter von den Philippinen published in two parts. All the taxa described before are TL: Mindanao. candida Schröder, Treadaway & Nuyda, 1990 (Fig. 14) TL: Bilar, Bohol. TL: Palawan. TL: Palawan. male (Fig. 78) deposited in MNHN as the lectotype, labelled, romeo Schröder & Treadaway, 1987 (Fig. 87) TL: Locality unknown, presumably Mindoro [Mindoro]. volupia Tsukada & Nishiyama, 1981 (Fig. 113) TL: Dumaran. und Bemerkungen über die Verbreitung einiger derselben” (G. examined and figured as much as possible in part 1. In the TS (TP): LT ♂ (SMFL). TL: 30 m NN, Calamian Group. dinorah Fruhstorfer, 1899 (Fig. 23) TS (TP): HT ♀ (FASU). joloana Staudinger, 1889 (Fig. 44) TS (TP): LT ♂ (ZMHU). mindorana (Euthalia) Tsukada, 1991 (permanently invalid) TS (TP): HT ♂ (BMNH). “Type (red) / lubentina philippensis Fruhst. / Philippinen TL: Mt. Halcon, N. Mindoro. TS (TP): ST (MNHN). treadawayi Tsukada, 1991 (Fig. 107) TL: Camiguin. TS (TP): HT ♂ (TY), PT 11♂4♀. Semper, 1886–1892), and then “Die Gross-Schmetterlinge der second part, synonymic lists and some important specimens Note: Description is based on some specimens. The lecto- TS (TP): HT ♂ (SMFL), PT10 ♂. TL: Basilan. Note: As this taxon gahiti was described as a form of TL: Sulu. Note: Description is based on four males and six females, Note: The taxon mindorana described under the genus Bazilan II-III, 98 Doherty ex. coll. H. Fruhstorfer / TS (TP): HT ♀ (SMFL), PT 1♀. Note: Though the original description does not mention TL: Romblon, Sibuyanes. TS (TP): HT ♂ (ET), PT 2♂5♀. (to be continued) Erde, 9” (1908–1927) edited by A. Seitz. Recently, a Japanese will be shown, affixing several discussions. type designation was carried out by Yokochi (1998). TS (TP): ST 2♂2♀ (MNHN). Lexias panopus boholensis (infrasubspecific name), it should TS (TP): LT ♂ (ZMHU). and two pairs are preserved in ZMHU. The lectotype designa- Euthalia is a junior primary homonym of Euthalia lusiada palawana (Tanaecia) Staudinger, 1889 (Fig. 72) MUSEUM PARIS 1934 COLL H Fruhstorfer”. the number and sex, a syntype specimen in MNHN is a male. TS (TP): HT ♂ (SMFL). entomologist, E. Tsukada, had compiled a great series of cavarna Fruhstorfer, 1913 (Fig. 15) be treated as an unavailable name. Note: Description is based on two males and one female, tion was carried out by Yokochi (1999). mindorana Fruhstorfer, 1899 (permanently invalid name). TL: Palawan. rudraca Fruhstorfer, 1913 (Fig. 88) “Butterflies of the South East Asian Islands” (1980–1991). balabacana (Bassarona) Tsukada, 1991 (Fig. 6) TL: Balabas [Balabac]. dodong Schröder & Treadaway, 1978 (Fig. 24) and all of them are preserved in ZMHU. The lectotype desig- Now it is replaced by tsukada Koçak (1996). TS (TP): LT ♂ (ZMHU). phlegethon G. Semper, [1887] (Fig. 79) TL: Balabac. soloni M. & T. Okano, 1990 (Fig. 98) tsukada Koçak, 1996 (Fig. 80) The late Prof. Julian N. Jumalon (1909–2000) (Fig. 1, TL: Balabac. TS (TP): LT ♀ (ZMHU). TL: Mobo, 1000 ft, Masbate. galoa Fruhstorfer, 1913 (Fig. 34) nation was carried out by Yokochi (1999). lupina Druce, 1874 (Fig. 54) Note: Description is based on males and females, and six TL: Mindanao. TS (TP): LT ♂ (ZMHU). TL: Bilar, Bohol. TL: Mindoro. left), the Lepidoptera Research Associate and Lecturer of 2. ABBREVIATIONS TS (TP): HT ♂ (ET), PT 8♂6♀. Note: Description is based on a few females, and one spec- TS (TP): HT ♂ (SMFL), PT 3♂7♀. TL: Mindoro. TL: Philippine Islands [Sulu]. mindorensis Schröder & Treadaway, 1982 (Fig. 63) males and three females are preserved in ZMHU. The lecto- TS (TP): LT ♀ (SMFL). Note: Description is based on some specimens, and two TS (TP): HT ♂ (FASU), PT 2♂3♀. TS (TP): HT ♂ (ET), PT 10♂11♀. Department of Biology, University of San Carlos, published a imen is preserved in ZMHU. The lectotype designation was TS (TP): ST (BMNH). kayumanggia Treadaway & Nuyda, 1994 (Fig. 45) TS (TP): HT ♀ (BMNH). TL: Mindoro. type designation was carried out by Yokochi (1999). Note: Description is based on twenty-one males and seven males and two females are preserved in ZMHU. The lectotype Note: Euthalia lubentina mindorana Tsukada, 1991 is a paper of “New Philippine butterflies” in 1975, and many new Public Institution (Museum, University). BMNH: The balabacana (Dophla) Tsukada, 1991 (secondary homonym) carried out by Yokochi (1999). dohertyi Butler, 1901 (Fig. 25) Note: Though the original description does not mention TL: Mt. Bayog, S. Palawan. TS (TP): HT ♂ (SMFL), PT 6♂7♀. females from Samar, Camiguin de Mindanao, N. Mindanao, E. designation was carried out by Yokochi (1999). soregina Fruhstorfer, 1913 (Fig. 99) permanently invalid name (nom. nov. pro mindorana Tsukada, taxonomic names concerned with various families were Natural History Museum, London, United Kingdom; FASU: (Fig. 7) TL: Sulu Archipelago. the number and sex, syntype specimen in BMNH is a male. TS (TP): HT ♂ (SMFL), PT 13♂4♀. lusiada C. & R. Felder, 1863 (Fig. 55) palawana (Euthalia) Staudinger, 1889 (Fig. 73) Mindanao, SE. Mindanao, and several materials are preserved TL: Sulu Archipelago. 1991, non Fruhstorfer,1899). New name tsukada was proposed described there. He played a leading part of the study of Exhibition Hall of Food, Agricultural and Environmental Note: See below balabacensis. chloe Schröder & Treadaway, 1990 (Fig. 16) TS (TP): HT ♂ (BMNH). TL: Luzon. miscus Fruhstorfer, 1913 (Fig. 64) TL: Palawan. in SMFL. The lectotype designation was carried out for the samarensis Tsukada, 1991 (Fig. 89) TS (TP): ST ♀ (BMNH). by Koçak (1996). Philippine butterflies and contributed greatly to entomology. Sciences, Faculty of Agriculture, Shinshu University, Nagano, TL: Jao Asan, 200 m, Sibuyan, Romblon Group. goertzi Jumalon, 1975 (Fig. 35) kulaya Treadaway & Nuyda, 1994 (Fig. 46) TS (TP): ST ♂ (SMFL). TL: Mindanao. TS (TP): LT ♂ (ZMHU). Mindanao specimen by Yokochi (1998). TL: San Juan, Samar. Note: According to the original description, the type series In 1995, the late Mr. Colin G. Treadaway (Fig. 1, right, Japan; KMNH: Kitakyushu Museum of Natural History & balabacensis nom. nov. (Fig. 7) TS (TP): HT ♂ (SMFL), PT 11♂9♀. dumaranensis Hanafusa, 1992 (Fig. 26) TL: Amlan, Negros Oriental. TL: Magellanes, Homonhon. Note: Though the original description does not mention TS (TP): ST (MNHN, BMNH). Note: Description is based on one male and six females, TS (TP): HT ♂ (ET), PT 3♂2♀. are exactly female, but the number is unknown. tyawena Fruhstorfer, 1913 (Fig. 108) died in 2019) published “Checklist of the butterflies of the Human History, Fukuoka, Japan; MNHN: Museum National TL: Balabac. TL: Dumaran. TS (TP): HT ♀ (ET). TS (TP): HT ♂ (SMFL), PT 14♂10♀. the number of types, a male syntype is preserved in SMFL. Note: According to the original description, types and one male and four females are preserved in ZMHU. The pinamalayana Yokochi, 1997 (Fig. 80) TL: unknown [Palawan]. Philippine Islands (Lepidoptera: Rhopalocera)”. Although, 25 d'Histoire Naturelle Entomologie, Paris, France; SMFL: TS (TP): HT ♂ (ET), PT 2♂2♀. circe Schröder & Treadaway, 1990 (Fig. 17) TS (TP): HT ♂ (HH). Note: Though the original description mentions the Note: The name kulaya is spelt “kúlaya” in the original (syntypes) are preserved in Fruhstorfer collection (now in lectotype designation was carried out by Yokochi (1999). Note: The new name pinamalayana was proposed by sarmana Fruhstorfer, 1913 (Fig. 90) subpiratica Schröder & Treadaway, 1987 (Fig. 100) TS (TP): LT ♀ (ZMHU). years have passed since his paper presented, several new Senckenberg Forschungsinstitut und Naturmuseum Frankfurt, Note: The taxon balabacana was described under the TL: Cavan Cavan, Sibutu, Tawitawi Group. holotype is preserved in USC, it is deposited in ET. description. macer Tsukada & Nishiyama, 1980 (Fig. 56) MNHN) and BMNH. Yokochi (1997), because mindorana Tsukada, 1991 was a TL: Basilan. TL: Nueva Ecija, Bongabon, Centr. Luzon. Note: Description is based on some specimens, and one findings are still remained unpublished — it can be followed Frankfurt-am-Main, Germany; USC: University of San Carlos, genus Dophla. As Yokochi (2010) assigned both Dophla and TS (TP): HT ♂ (SMFL), PT 9♂10♀. dunya Doubleday, [1848] (Fig. 27) TL: Bo. Melgar, Dinagat. pallida Schröder, Treadaway & Nuyda, 1990 (Fig. 74) permanently invalid name. However, pinamalayana is a junior TS (TP): ST (MNHN). TS (TP): HT ♂ (SMFL), PT1♀. male and one female are preserved in ZMHU. The lectotype by his own handwriting on the glass surface of his specimen Cebu, Philippines; ZMHU: Zoologisches Museum, Humboldt Bassarona to the genus Euthalia, balabacana (Dophla) is a TL: Assam, India. hikarugenzi Tsukada & Nishiyama, 1980 (Fig. 36) laetitiae Schröder & Treadaway, 2000 (Fig. 47) TS (TP): HT ♂ (ET), PT 3♂2♀. monara Fruhstorfer, 1913 (Fig. 65) TL: 30 m NN, Calamian Group. synonym of tsukada Koçak (1996). Note: Though the original description does not mention designation was carried out by Yokochi (1999). cases —. The first author visited Senckenberg Museum in Universität, Berlin, Germany. junior secondary homonym of Bassarona teuta balabacana cusama Fruhstorfer, 1913 (Fig. 18) TS (TP): ST ♂ (BMNH). TL: near Baguio, N. Luzon. TL: Cavan Cavan, 5 m, Sibutu, Tawitawi Group. TL: N. Borneo. TS (TP): HT ♂ (SMFL), PT 11♂16♀. the number and sex, a syntype specimen in MNHN is a male. suluana Fruhstorfer, 1902 (Fig. 101) Frankfurt to research on the Treadaway collection on March Private collections. ET: Etsuzo Tsukada, Yamanashi, Tsukada, 1991. Here a new name balabacensis is replaced for TL: Mindanao, Camiguin de Mindanao. TS (TP): HT ♂ (MKN), PT 1♀ (MKN). TS (TP): HT ♂ (SMFL), PT 4♂. malissia Fruhstorfer, 1913 (Fig. 57) TS (TP): ST (MNHN). piratica G. Semper, [1887] (Fig. 81) TL: Sulu Archipelago. uposatha Fruhstorfer, 1913 (Fig. 109) 2019, and just then was informed that he had died several Japan; HH: Hiroto Hanafusa, Tottori, Japan; JF: Jumalon Dophla evelina balabacana Tsukada, 1991. TS (TP): ST ♂ (MNHN), ♀ (SMFL). ellora Fruhstorfer, 1898 (Fig. 28) TL: Basilan. panayana Schröder & Treadaway, 1980 (Fig. 75) TL: Camiguin de Mindanao. satrapes C. & R. Felder, 1861 (Fig. 91) TS (TP): HT ♀ (BMNH). TL: Banggi (Banguey), E. Malaysia. weeks ago at his home in England. In the middle 1990’s, every Family, Cebu, Philippines; MKN: Misao Kaneko, Tokyo, Euthalia (Dophla) evelina balabacensis nom. nov. Note: Figured specimen is a syntype female in SMFL. TL: Mindoro. hinunanganensis M. & T. Okano, 1986 (Fig. 37) leopardina Fruhstorfer, 1913 (Fig. 48) TS (TP): ST♂ (MNHN). nadenya Fruhstorfer, 1913 TL: Culasi, 800 ft, Antique, Panay. TS (TP): LT ♂ (SMFL). TL: Luzon. TS (TP): LT ♂ (ZMHU). time the first author visited to Senckenberg, he came bringing Japan; SK: Shun-ichi Kawamura, Tokyo, Japan; STA: Shinji pro Dophla evelina balabacana Tsukada, 1991, non TS (TP): ST ♂ (MNHN). TL: Hinunangan, Leyte. TL: Sulu Islands. Note: Though the original description does not mention TL: Luzon. TS (TP): HT ♂ (SMFL), PT 4♂4♀. Note: Description is based on six materials from Luzon, TS (TP): ST ♂ (BMNH). susoni Jumalon, 1975 (Fig. 102) Note: According to the original description, the type series his described type specimens, with his wife driving from Taga, Tokyo, Japan; TY: Takashi Yokochi, Aichi, Japan. Bassarona teuta balabacana Tsukada, 1991 dacasini Hanafusa, 1990 (Fig. 19) Note: Though the original description does not mention TS (TP): HT ♂ (FASU), PT 1♂. TS (TP): HT ♀ (BMNH). the number of types, a male syntype is preserved in MNHN. TS (TP): HT ♀ [untraced, not examined]. Mindoro, Camiguin de Mindanao, N. Mindanao. The lectotype Note: According to the original description, the type series TL: Camp 7, Minglanilla, Cebu. (male and female, but the number is unknown) are deposited in Specimens concerned. FW: Length of Forewing; HT: TL: Balabac. the number and sex, a syntype specimen in MNHN is a male. Note: Though the original description indicates that the panopus C. & R. Felder, 1861 (Fig. 76) designation was carried out for the Camiguin de Mindanao (number is unknown) are deposited in Semper and Felder TS (TP): HT ♂ (USC), PT 9♂9♀ (USC, JF, SMFL, TY). Staudinger collection (ZMHU). The lectotype designation was Holotype; LT: Lectotype; PLT: Paralectotype; PT: Paratype; boholensis (Lexias) M. & T. Okano, 1988 (Fig. 8) TS (TP): HT ♂ (HH), PT 1♀. hiwaga Nuyda & Kawamura, 1989 (Fig. 38) leucotaenia G. Semper, 1878 (Fig. 49) marinduque K. Okano, 1992 (Fig. 58) type female should be preserved in ZMHU, the specimen TL: Luzon. specimen by Schröder & Treadaway (1987). collections. carried out by Yokochi (1999). TL: Type Locality; TP: Type Preservation; TS: Type Series; TL: Bilar, Bohol. eson de Nicéville, 1894 (Fig. 29) TL: Camiguin, Babuyan. TL: Camotes. TL: Marinduque. corresponding with nadenya could not be found there. No TS (TP): ST ♂ (BMNH). tanagra Staudinger, 1889 (Fig. 103) ST: Syntype. TS (TP): HT ♂ (FASU), PT 6♂2♀. damalis Erichson, 1834 (Fig. 117) TL: Palawan. TS (TP): HT ♂ (SK), PT 7♂4♀ (SK, ET). TS (TP): LT ♂ (SMFL). TS (TP): HT ♀ (KMNH), PT 1♂ (KMNH). figure is presented in the original paper. Note: The original description does not mention the princesa Fruhstorfer, 1899 (Fig. 82) schoenigi Schröder & Treadaway, 1978 (Fig. 92) TL: Palawan. virginalis Fruhstorfer, 1913 TL: Luzon. TS (TP): LT ♂ (SMFL), PLT 1♀. Note: This taxon was described based on sixty-nine number of type series. TL: Palawan. TL: Amlan, 600 m, Negros. TS (TP): LT ♂ (ZMHU). TL: locality not given. boholensis (Euthalia) M. & T. Okano, 1990 (Fig. 9) TS (TP): ST ♂ [untraced, not examined]. Note: Description is based on a single pair, and all howarthi Jumalon, 1975 (Fig. 39) materials from Camotes, Bohol, Leyte, Panaon, E. Mindanao, medaga Fruhstorfer, 1913 (Fig. 59) negrosiana Schröder & Treadaway, 1987 (Fig. 66) TS (TP): HT ♀ (MNHN). TS (TP): HT ♀ (SMFL). Note: Description is based on seven males and eleven TS (TP): ST ♀ (BMNH) [not examined]. 3. LIST OF DESCRIBED SPECIES AND TL: Bilar, Bohol. Note: The type series should be preserved in ZMHU, but materials are preserved in SMFL. The lectotype designation TL: Amlan, Negros Oriental. and N. Mindanao. Yokochi (1998) mentioned the type series of TL: Camiguin de Mindanao. TL: Negros Or. [Oriental]. phelada G. Semper, [1887] (Fig. 77) females, and two males and three females are preserved in Note: This is a female form of panopus. The type locality SUBSPECIES NAME TS (TP): HT ♀ (FASU), PT 4♀. the specimens that corresponded with the types could not be was carried out by Yokochi (1998). TS (TP): HT ♂ (USC), PT 4♀ (USC, JF). leucotaenia was untraced so designated the lectotype for the TS (TP): ST ♂ (MNHN). TS (TP): HT ♂ (SMFL), PT 1♂1♀. TL: Luzon. proditrix Fruhstorfer, 1913 (Fig. 83) semperi Staudinger, 1889 (Fig. 93) ZMHU. The lectotype designation was carried out by Yokochi suspected is Mindanao, though the specific location is not found. The Fig. 117 is reproduced from the original description figure of the original description, but the type series specimen Note: Though the original description does not mention TS (TP): LT ♀ (SMFL). TL: Basilan. TL: Palawan. (1999). shown. The type series are deposited in the Rothschild collection The following list contains the taxonomic names bongaoensis Schröder & Treadaway, 1990 (Fig. 10) (pl. 50, fig. 4, ♂). eva C. & R. Felder, [1867] (Fig. 30) imperator Hewitson, 1863 (Fig. 40) was found after the third investigation in SMFL. We designate the number of types, a male syntype is preserved in MNHN. nirodha Fruhstorfer, 1913 (Fig. 67) Note: This taxon was described based on five materials TS (TP): ST ♂ (MNHN). TS (TP): LT ♂ (ZMHU). (BMNH, London), but we have not located them at present. concerned with Philippines. The names which are described TL: Bongao Peak, Bongao, Tawitawi Group. TL: Luzon. TL: Luzon. here again the lectotype for a male specimen shown in Fig. 49 TL: Basilan. from Luzon, Camiguin de Mindanao, and SE. Mindanao. Note: Description is based on males and females, and four tawitawia Treadaway & Nuyda, 1994 (Fig. 104) from the other areas are also included. TS (TP): HT ♀ (SMFL), PT 18♂20♀. darani Fruhstorfer, 1913 (Fig. 20) TS (TP): ST ♂ (BMNH). TS (TP): HT ♀ (BMNH). (labelled “Typus (red) / Camotes / Bohol / 113 / 211 / Coll. C. mindanaensis Schröder & Treadaway, 1978 (Fig. 60) TS (TP): ST ♂♀ (MNHN). Yokochi (1998) mentioned the type series of phelada was producta Butler, 1901 (Fig. 84) males and four females are preserved in ZMHU. The lectotype TL: Magsaggao, Tawitawi. visayana (Lexias) Schröder & Treadaway, 1987 (Fig. 110) TL: Sulu Archipelago. Note: According to the original description, eva is distrib- Semper”). TL: Surigao del Sur, Mindanao. untraced so designated the lectotype for the figure of the TL: Philippines [recte W. Malaysia]. designation was carried out by Yokochi (1999). TS (TP): HT ♂ (SMFL). TL: Catmon, S. Leyte. albusequus Nihira & Kawamura, 1986 (Fig. 2) borromeoi Schröder, 1977 (Fig. 11) TS (TP): HT ♀ (BMNH). uted in Assam, Luzon, and Celebes. The details of type series ingae (Lexias) Schröder & Treadaway, 1987 (Fig. 41) TS (TP): HT ♂ (SMFL). obatrata Yokochi, 1994 (Fig. 68) original description, but the target specimen was found after the TS (TP): ST ♀ (BMNH). TS (TP): HT ♂ (SMFL), PT 6♂4♀. TL: Lakit lakit, Sanga Sanga, Tawitawi. TL: Espana, 2000 ft, Sibuyan. Note: Though the original description does not mention are not mentioned. Staudinger (1886) is the first revisory of TL: Negros Or. [Oriental], 800 m. leytana Schröder & Treadaway, 1982 (Fig. 50) TL: Dumaran. third investigation in SMFL. We designate here again the senilis Hanafusa, 1992 (Fig. 94) tethys Tsukada, 1991 (Fig. 105) TS (TP): HT ♀ (SK). TS (TP): HT ♂ (SMFL), PT 1♀. the number of type series, one female would be the holotype. which Luzon is recognized as a locality of eva. TS (TP): HT ♂ (SMFL), PT 5♂5♀. TL: St. Bernard, Calpio, 280 m, S. Leyte. mindorana (Tanaecia) Fruhstorfer, 1904 (Fig. 61) TS (TP): HT ♀ (HH), PT 1♂1♀. lectotype for a female specimen shown in Fig. 77 (labelled rhamases Staudinger, 1889 (Fig. 85) TL: Dumaran. TL: N. Palawan. visayana (Cynitia) Schröder & Treadaway, 1981 (Fig. 111) TS (TP): HT ♂ (SMFL), PT 18♂10♀. TL: Mindoro. “Typus (red) / Euth. phelada typ. ♀ Semper / Marivcles (?) TL: Palawan. TS (TP): HT ♂ (HH), PT 1♂. TS (TP): HT ♂ (ET), PT 10♂12♀. TL: St. Bernard, Catmon, S. Leyte. 10 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 11

1. INTRODUCTION Limbach, about 250 km from Frankfurt. It is an undoubted fact amlana Jumalon, 1970 (Fig. 3) cabigoni Schröder, Treadaway, A. Mohagan & N. Mohagan, dhayma Fruhstorfer, 1913 (Fig. 21) exul Tsukada & Nishiyama, 1980 (Fig. 31) ingae (Euthalia) Schröder & Treadaway, 1990 (Fig. 42) leytensis Jumalon, 1975 (Fig. 51) TS (TP): ST 2♂1♀ (BMNH). ormocana Jumalon, 1970 (Fig. 69) 11.11.81 / 110 / Coll. C. Semper / Lectotype ♀ Euthalia TS (TP): LT ♂ (ZMHU). sibutana Schröder & Treadaway, 1990 (Fig. 95) Note: Attached label to the holotype specimen shows TS (TP): HT ♂ (SMFL), PT 10♂2♀. waltraudae Schröder & Treadaway, 1990 (Fig. 114) that Mr. Treadaway and Dr. H. Schröder of Senckenberg TL: Amlan, Negros Oriental. 2009 (Fig. 12) TL: Sulu Archipelago. TL: Bo. Melgar, Dinagat. TL: Bongao Peak, 200 m, Bongao, Tawitawi Group. TL: Lake Danao, Ormoc City, Leyte. Note: The figured female (Fig. 61) is preserved in the TL: Lake Danao, Ormoc City, Leyte. phelada Semper [87]:1888 Designated by T. Yokochi, 1997”). Note: Description is based on males, and four males are TL: Cavan Cavan, 5 m, Sibutu, Tawitawi Group. “Balabac”, but the correct type locality is Palawan. TL: Indalawan, Balabac. The Philippine Islands are situated in the northern end of thoroughly deserve credit for working further on systematic TS (TP): HT ♂ (USC), PT10 ♂1♀ (USC, JF). TL: Balut Is, Sarangani Group. TS (TP): HT ♀ (BMNH). TS (TP): HT ♂ (ET), PT 3♂6♀. TS (TP): HT ♀ (SMFL), PT 10♂12♀. TS (TP): HT ♀ (USC). Rothschild collection (Ro 98/7), BMNH. It might be one of the TS (TP): HT ♂ (USC), PT 1♀. preserved in ZMHU. The lectotype designation was carried out TS (TP): HT ♂ (SMFL), PT 17♂15♀. vistrica Fruhstorfer, 1913 (Fig. 112) TS (TP): HT ♀ (SMFL), PT 2♂2♀. the Malay Archipelago and are composed of more than 7,000 classification in Philippines Rhopalocera. TS (TP): HT ♂ (SMFL), PT 2♂1♀. Note: Though the original description does not mention the types, although the attached label is only “Mindoro”. No other philippensis Fruhstorfer, 1899 (Fig. 78) by Yokochi (1999). trapesa G. Semper, [1887] (Fig. 106) TL: locality unknown, presumably one of the Central islands, large and small, extending from north to south about Here we revised the group of the tribe Adoliadini antiquea Schröder & Treadaway, 1980 (Fig. 4) number of type series, one female would be the holotype. felicitacionae Jumalon, 1989 (Fig. 32) irauana Jumalon, 1975 (Fig. 43) liaoi Schröder & Treadaway, 1982 (Fig. 52) specimen is found that corresponded with the type series. ornata Schröder & Treadaway, 1979 (Fig. 70) TL: Basilan. sibuyana Schröder & Treadaway, 1982 (Fig. 96) TL: Camiguin de Mindanao. Philippine islands [recte Dinagat]. yoshiroui K. Okano, 1992 (Fig. 115) 1,800 km and from east to west about 1,100 km. The insect distributed in the Philippines, based on the checklist by TL: Culasi, 500 m, Antique, Panay. calliphorus C. & R. Felder, 1861 (Fig. 13) TL: Kantakpal Creek, Awihao, Ormoc, Western Leyte. TL: Irauan [Irawan], Puerto Princesa, Palawan. TL: Mt. Madja-an [Mt. Madja-as], 500 m, Antique, Panay. TL: Romblon. TS (TP): LT ♂ (MNHN). rodriguezi Schröder & Treadaway, 1982 (Fig. 86) TL: Sibuyan, Romblon Group. TS (TP): LT ♂ (SMFL). TS (TP): ST ♂ (BMNH). TL: Babuyan. fauna is rich in diversity and the tribe Adoliadini, Nymphalidae, Treadaway (1995). One hundred-twenty taxa of type specimens TS (TP): HT ♂ (SMFL). TL: Luzon. dinagatensis Tsukada, 1991 (Fig. 22) TS (TP): HT ♂ (JF). TS (TP): HT ♀ (JF), PT 1♂1♀ (USC). TS (TP): HT ♂ (SMFL), PT 1♂2♀. mindorana (Euthalia) Fruhstorfer, 1899 (Fig. 62) TS (TP): HT ♂ (SMFL), PT 1♂2♀. Note: Description is based on one male from Basilan TL: 28 km W of Puerto Princesa, Palawan. TS (TP): HT ♂ (SMFL), PT 2♂5♀. Note: Description is based on seven materials from E. Note: Tsukada (1991) suspected this taxon is distributed TS (TP): HT ♂ (KMNH). Lepidoptera, the main group of this study, is no exception. The founded in the Philippines are examined and revised the TS (TP): ST ♂ (BMNH). TL: Dinagat. Note: The holotype is deposited in Jumalon family collec- TL: Mindoro. (MNHN) and one female from Luzon (ZMHU). As the TS (TP): HT ♂ (SMFL), PT 2♂2♀. Mindanao. The lectotype designation was carried out for the in Basilan, but it presumably comes from Dinagat. important works concerned with Philippines were, back in classification of species-subspecies relation. This paper is aquamarina Fruhstorfer, 1913 (Fig. 5) TS (TP): HT ♂ (ET), PT 2♂1♀. gahiti M. & T. Okano, 1990 (unavailable name) (Fig. 33) tion. ludonia Staudinger, 1889 (Fig. 53) TS (TP): ST (MNHN). palawana (Lexias) Moore, 1897 (Fig. 71) Luzon-group should be classified as nadenya. We select a smaragdifera Fruhstorfer, 1913 (Fig. 97) Camiguin de Mindanao specimen by Yokochi (1998). yui Yokochi, 1994 (Fig. 116) history, “Diagnosen einiger neuer Tagfalter von den Philippinen published in two parts. All the taxa described before are TL: Mindanao. candida Schröder, Treadaway & Nuyda, 1990 (Fig. 14) TL: Bilar, Bohol. TL: Palawan. TL: Palawan. male (Fig. 78) deposited in MNHN as the lectotype, labelled, romeo Schröder & Treadaway, 1987 (Fig. 87) TL: Locality unknown, presumably Mindoro [Mindoro]. volupia Tsukada & Nishiyama, 1981 (Fig. 113) TL: Dumaran. und Bemerkungen über die Verbreitung einiger derselben” (G. examined and figured as much as possible in part 1. In the TS (TP): LT ♂ (SMFL). TL: 30 m NN, Calamian Group. dinorah Fruhstorfer, 1899 (Fig. 23) TS (TP): HT ♀ (FASU). joloana Staudinger, 1889 (Fig. 44) TS (TP): LT ♂ (ZMHU). mindorana (Euthalia) Tsukada, 1991 (permanently invalid) TS (TP): HT ♂ (BMNH). “Type (red) / lubentina philippensis Fruhst. / Philippinen TL: Mt. Halcon, N. Mindoro. TS (TP): ST (MNHN). treadawayi Tsukada, 1991 (Fig. 107) TL: Camiguin. TS (TP): HT ♂ (TY), PT 11♂4♀. Semper, 1886–1892), and then “Die Gross-Schmetterlinge der second part, synonymic lists and some important specimens Note: Description is based on some specimens. The lecto- TS (TP): HT ♂ (SMFL), PT10 ♂. TL: Basilan. Note: As this taxon gahiti was described as a form of TL: Sulu. Note: Description is based on four males and six females, Note: The taxon mindorana described under the genus Bazilan II-III, 98 Doherty ex. coll. H. Fruhstorfer / TS (TP): HT ♀ (SMFL), PT 1♀. Note: Though the original description does not mention TL: Romblon, Sibuyanes. TS (TP): HT ♂ (ET), PT 2♂5♀. (to be continued) Erde, 9” (1908–1927) edited by A. Seitz. Recently, a Japanese will be shown, affixing several discussions. type designation was carried out by Yokochi (1998). TS (TP): ST 2♂2♀ (MNHN). Lexias panopus boholensis (infrasubspecific name), it should TS (TP): LT ♂ (ZMHU). and two pairs are preserved in ZMHU. The lectotype designa- Euthalia is a junior primary homonym of Euthalia lusiada palawana (Tanaecia) Staudinger, 1889 (Fig. 72) MUSEUM PARIS 1934 COLL H Fruhstorfer”. the number and sex, a syntype specimen in MNHN is a male. TS (TP): HT ♂ (SMFL). entomologist, E. Tsukada, had compiled a great series of cavarna Fruhstorfer, 1913 (Fig. 15) be treated as an unavailable name. Note: Description is based on two males and one female, tion was carried out by Yokochi (1999). mindorana Fruhstorfer, 1899 (permanently invalid name). TL: Palawan. rudraca Fruhstorfer, 1913 (Fig. 88) “Butterflies of the South East Asian Islands” (1980–1991). balabacana (Bassarona) Tsukada, 1991 (Fig. 6) TL: Balabas [Balabac]. dodong Schröder & Treadaway, 1978 (Fig. 24) and all of them are preserved in ZMHU. The lectotype desig- Now it is replaced by tsukada Koçak (1996). TS (TP): LT ♂ (ZMHU). phlegethon G. Semper, [1887] (Fig. 79) TL: Balabac. soloni M. & T. Okano, 1990 (Fig. 98) tsukada Koçak, 1996 (Fig. 80) The late Prof. Julian N. Jumalon (1909–2000) (Fig. 1, TL: Balabac. TS (TP): LT ♀ (ZMHU). TL: Mobo, 1000 ft, Masbate. galoa Fruhstorfer, 1913 (Fig. 34) nation was carried out by Yokochi (1999). lupina Druce, 1874 (Fig. 54) Note: Description is based on males and females, and six TL: Mindanao. TS (TP): LT ♂ (ZMHU). TL: Bilar, Bohol. TL: Mindoro. left), the Lepidoptera Research Associate and Lecturer of 2. ABBREVIATIONS TS (TP): HT ♂ (ET), PT 8♂6♀. Note: Description is based on a few females, and one spec- TS (TP): HT ♂ (SMFL), PT 3♂7♀. TL: Mindoro. TL: Philippine Islands [Sulu]. mindorensis Schröder & Treadaway, 1982 (Fig. 63) males and three females are preserved in ZMHU. The lecto- TS (TP): LT ♀ (SMFL). Note: Description is based on some specimens, and two TS (TP): HT ♂ (FASU), PT 2♂3♀. TS (TP): HT ♂ (ET), PT 10♂11♀. Department of Biology, University of San Carlos, published a imen is preserved in ZMHU. The lectotype designation was TS (TP): ST (BMNH). kayumanggia Treadaway & Nuyda, 1994 (Fig. 45) TS (TP): HT ♀ (BMNH). TL: Mindoro. type designation was carried out by Yokochi (1999). Note: Description is based on twenty-one males and seven males and two females are preserved in ZMHU. The lectotype Note: Euthalia lubentina mindorana Tsukada, 1991 is a paper of “New Philippine butterflies” in 1975, and many new Public Institution (Museum, University). BMNH: The balabacana (Dophla) Tsukada, 1991 (secondary homonym) carried out by Yokochi (1999). dohertyi Butler, 1901 (Fig. 25) Note: Though the original description does not mention TL: Mt. Bayog, S. Palawan. TS (TP): HT ♂ (SMFL), PT 6♂7♀. females from Samar, Camiguin de Mindanao, N. Mindanao, E. designation was carried out by Yokochi (1999). soregina Fruhstorfer, 1913 (Fig. 99) permanently invalid name (nom. nov. pro mindorana Tsukada, taxonomic names concerned with various families were Natural History Museum, London, United Kingdom; FASU: (Fig. 7) TL: Sulu Archipelago. the number and sex, syntype specimen in BMNH is a male. TS (TP): HT ♂ (SMFL), PT 13♂4♀. lusiada C. & R. Felder, 1863 (Fig. 55) palawana (Euthalia) Staudinger, 1889 (Fig. 73) Mindanao, SE. Mindanao, and several materials are preserved TL: Sulu Archipelago. 1991, non Fruhstorfer,1899). New name tsukada was proposed described there. He played a leading part of the study of Exhibition Hall of Food, Agricultural and Environmental Note: See below balabacensis. chloe Schröder & Treadaway, 1990 (Fig. 16) TS (TP): HT ♂ (BMNH). TL: Luzon. miscus Fruhstorfer, 1913 (Fig. 64) TL: Palawan. in SMFL. The lectotype designation was carried out for the samarensis Tsukada, 1991 (Fig. 89) TS (TP): ST ♀ (BMNH). by Koçak (1996). Philippine butterflies and contributed greatly to entomology. Sciences, Faculty of Agriculture, Shinshu University, Nagano, TL: Jao Asan, 200 m, Sibuyan, Romblon Group. goertzi Jumalon, 1975 (Fig. 35) kulaya Treadaway & Nuyda, 1994 (Fig. 46) TS (TP): ST ♂ (SMFL). TL: Mindanao. TS (TP): LT ♂ (ZMHU). Mindanao specimen by Yokochi (1998). TL: San Juan, Samar. Note: According to the original description, the type series In 1995, the late Mr. Colin G. Treadaway (Fig. 1, right, Japan; KMNH: Kitakyushu Museum of Natural History & balabacensis nom. nov. (Fig. 7) TS (TP): HT ♂ (SMFL), PT 11♂9♀. dumaranensis Hanafusa, 1992 (Fig. 26) TL: Amlan, Negros Oriental. TL: Magellanes, Homonhon. Note: Though the original description does not mention TS (TP): ST (MNHN, BMNH). Note: Description is based on one male and six females, TS (TP): HT ♂ (ET), PT 3♂2♀. are exactly female, but the number is unknown. tyawena Fruhstorfer, 1913 (Fig. 108) died in 2019) published “Checklist of the butterflies of the Human History, Fukuoka, Japan; MNHN: Museum National TL: Balabac. TL: Dumaran. TS (TP): HT ♀ (ET). TS (TP): HT ♂ (SMFL), PT 14♂10♀. the number of types, a male syntype is preserved in SMFL. Note: According to the original description, types and one male and four females are preserved in ZMHU. The pinamalayana Yokochi, 1997 (Fig. 80) TL: unknown [Palawan]. Philippine Islands (Lepidoptera: Rhopalocera)”. Although, 25 d'Histoire Naturelle Entomologie, Paris, France; SMFL: TS (TP): HT ♂ (ET), PT 2♂2♀. circe Schröder & Treadaway, 1990 (Fig. 17) TS (TP): HT ♂ (HH). Note: Though the original description mentions the Note: The name kulaya is spelt “kúlaya” in the original (syntypes) are preserved in Fruhstorfer collection (now in lectotype designation was carried out by Yokochi (1999). Note: The new name pinamalayana was proposed by sarmana Fruhstorfer, 1913 (Fig. 90) subpiratica Schröder & Treadaway, 1987 (Fig. 100) TS (TP): LT ♀ (ZMHU). years have passed since his paper presented, several new Senckenberg Forschungsinstitut und Naturmuseum Frankfurt, Note: The taxon balabacana was described under the TL: Cavan Cavan, Sibutu, Tawitawi Group. holotype is preserved in USC, it is deposited in ET. description. macer Tsukada & Nishiyama, 1980 (Fig. 56) MNHN) and BMNH. Yokochi (1997), because mindorana Tsukada, 1991 was a TL: Basilan. TL: Nueva Ecija, Bongabon, Centr. Luzon. Note: Description is based on some specimens, and one findings are still remained unpublished — it can be followed Frankfurt-am-Main, Germany; USC: University of San Carlos, genus Dophla. As Yokochi (2010) assigned both Dophla and TS (TP): HT ♂ (SMFL), PT 9♂10♀. dunya Doubleday, [1848] (Fig. 27) TL: Bo. Melgar, Dinagat. pallida Schröder, Treadaway & Nuyda, 1990 (Fig. 74) permanently invalid name. However, pinamalayana is a junior TS (TP): ST (MNHN). TS (TP): HT ♂ (SMFL), PT1♀. male and one female are preserved in ZMHU. The lectotype by his own handwriting on the glass surface of his specimen Cebu, Philippines; ZMHU: Zoologisches Museum, Humboldt Bassarona to the genus Euthalia, balabacana (Dophla) is a TL: Assam, India. hikarugenzi Tsukada & Nishiyama, 1980 (Fig. 36) laetitiae Schröder & Treadaway, 2000 (Fig. 47) TS (TP): HT ♂ (ET), PT 3♂2♀. monara Fruhstorfer, 1913 (Fig. 65) TL: 30 m NN, Calamian Group. synonym of tsukada Koçak (1996). Note: Though the original description does not mention designation was carried out by Yokochi (1999). cases —. The first author visited Senckenberg Museum in Universität, Berlin, Germany. junior secondary homonym of Bassarona teuta balabacana cusama Fruhstorfer, 1913 (Fig. 18) TS (TP): ST ♂ (BMNH). TL: near Baguio, N. Luzon. TL: Cavan Cavan, 5 m, Sibutu, Tawitawi Group. TL: N. Borneo. TS (TP): HT ♂ (SMFL), PT 11♂16♀. the number and sex, a syntype specimen in MNHN is a male. suluana Fruhstorfer, 1902 (Fig. 101) Frankfurt to research on the Treadaway collection on March Private collections. ET: Etsuzo Tsukada, Yamanashi, Tsukada, 1991. Here a new name balabacensis is replaced for TL: Mindanao, Camiguin de Mindanao. TS (TP): HT ♂ (MKN), PT 1♀ (MKN). TS (TP): HT ♂ (SMFL), PT 4♂. malissia Fruhstorfer, 1913 (Fig. 57) TS (TP): ST (MNHN). piratica G. Semper, [1887] (Fig. 81) TL: Sulu Archipelago. uposatha Fruhstorfer, 1913 (Fig. 109) 2019, and just then was informed that he had died several Japan; HH: Hiroto Hanafusa, Tottori, Japan; JF: Jumalon Dophla evelina balabacana Tsukada, 1991. TS (TP): ST ♂ (MNHN), ♀ (SMFL). ellora Fruhstorfer, 1898 (Fig. 28) TL: Basilan. panayana Schröder & Treadaway, 1980 (Fig. 75) TL: Camiguin de Mindanao. satrapes C. & R. Felder, 1861 (Fig. 91) TS (TP): HT ♀ (BMNH). TL: Banggi (Banguey), E. Malaysia. weeks ago at his home in England. In the middle 1990’s, every Family, Cebu, Philippines; MKN: Misao Kaneko, Tokyo, Euthalia (Dophla) evelina balabacensis nom. nov. Note: Figured specimen is a syntype female in SMFL. TL: Mindoro. hinunanganensis M. & T. Okano, 1986 (Fig. 37) leopardina Fruhstorfer, 1913 (Fig. 48) TS (TP): ST♂ (MNHN). nadenya Fruhstorfer, 1913 TL: Culasi, 800 ft, Antique, Panay. TS (TP): LT ♂ (SMFL). TL: Luzon. TS (TP): LT ♂ (ZMHU). time the first author visited to Senckenberg, he came bringing Japan; SK: Shun-ichi Kawamura, Tokyo, Japan; STA: Shinji pro Dophla evelina balabacana Tsukada, 1991, non TS (TP): ST ♂ (MNHN). TL: Hinunangan, Leyte. TL: Sulu Islands. Note: Though the original description does not mention TL: Luzon. TS (TP): HT ♂ (SMFL), PT 4♂4♀. Note: Description is based on six materials from Luzon, TS (TP): ST ♂ (BMNH). susoni Jumalon, 1975 (Fig. 102) Note: According to the original description, the type series his described type specimens, with his wife driving from Taga, Tokyo, Japan; TY: Takashi Yokochi, Aichi, Japan. Bassarona teuta balabacana Tsukada, 1991 dacasini Hanafusa, 1990 (Fig. 19) Note: Though the original description does not mention TS (TP): HT ♂ (FASU), PT 1♂. TS (TP): HT ♀ (BMNH). the number of types, a male syntype is preserved in MNHN. TS (TP): HT ♀ [untraced, not examined]. Mindoro, Camiguin de Mindanao, N. Mindanao. The lectotype Note: According to the original description, the type series TL: Camp 7, Minglanilla, Cebu. (male and female, but the number is unknown) are deposited in Specimens concerned. FW: Length of Forewing; HT: TL: Balabac. the number and sex, a syntype specimen in MNHN is a male. Note: Though the original description indicates that the panopus C. & R. Felder, 1861 (Fig. 76) designation was carried out for the Camiguin de Mindanao (number is unknown) are deposited in Semper and Felder TS (TP): HT ♂ (USC), PT 9♂9♀ (USC, JF, SMFL, TY). Staudinger collection (ZMHU). The lectotype designation was Holotype; LT: Lectotype; PLT: Paralectotype; PT: Paratype; boholensis (Lexias) M. & T. Okano, 1988 (Fig. 8) TS (TP): HT ♂ (HH), PT 1♀. hiwaga Nuyda & Kawamura, 1989 (Fig. 38) leucotaenia G. Semper, 1878 (Fig. 49) marinduque K. Okano, 1992 (Fig. 58) type female should be preserved in ZMHU, the specimen TL: Luzon. specimen by Schröder & Treadaway (1987). collections. carried out by Yokochi (1999). TL: Type Locality; TP: Type Preservation; TS: Type Series; TL: Bilar, Bohol. eson de Nicéville, 1894 (Fig. 29) TL: Camiguin, Babuyan. TL: Camotes. TL: Marinduque. corresponding with nadenya could not be found there. No TS (TP): ST ♂ (BMNH). tanagra Staudinger, 1889 (Fig. 103) ST: Syntype. TS (TP): HT ♂ (FASU), PT 6♂2♀. damalis Erichson, 1834 (Fig. 117) TL: Palawan. TS (TP): HT ♂ (SK), PT 7♂4♀ (SK, ET). TS (TP): LT ♂ (SMFL). TS (TP): HT ♀ (KMNH), PT 1♂ (KMNH). figure is presented in the original paper. Note: The original description does not mention the princesa Fruhstorfer, 1899 (Fig. 82) schoenigi Schröder & Treadaway, 1978 (Fig. 92) TL: Palawan. virginalis Fruhstorfer, 1913 TL: Luzon. TS (TP): LT ♂ (SMFL), PLT 1♀. Note: This taxon was described based on sixty-nine number of type series. TL: Palawan. TL: Amlan, 600 m, Negros. TS (TP): LT ♂ (ZMHU). TL: locality not given. boholensis (Euthalia) M. & T. Okano, 1990 (Fig. 9) TS (TP): ST ♂ [untraced, not examined]. Note: Description is based on a single pair, and all howarthi Jumalon, 1975 (Fig. 39) materials from Camotes, Bohol, Leyte, Panaon, E. Mindanao, medaga Fruhstorfer, 1913 (Fig. 59) negrosiana Schröder & Treadaway, 1987 (Fig. 66) TS (TP): HT ♀ (MNHN). TS (TP): HT ♀ (SMFL). Note: Description is based on seven males and eleven TS (TP): ST ♀ (BMNH) [not examined]. 3. LIST OF DESCRIBED SPECIES AND TL: Bilar, Bohol. Note: The type series should be preserved in ZMHU, but materials are preserved in SMFL. The lectotype designation TL: Amlan, Negros Oriental. and N. Mindanao. Yokochi (1998) mentioned the type series of TL: Camiguin de Mindanao. TL: Negros Or. [Oriental]. phelada G. Semper, [1887] (Fig. 77) females, and two males and three females are preserved in Note: This is a female form of panopus. The type locality SUBSPECIES NAME TS (TP): HT ♀ (FASU), PT 4♀. the specimens that corresponded with the types could not be was carried out by Yokochi (1998). TS (TP): HT ♂ (USC), PT 4♀ (USC, JF). leucotaenia was untraced so designated the lectotype for the TS (TP): ST ♂ (MNHN). TS (TP): HT ♂ (SMFL), PT 1♂1♀. TL: Luzon. proditrix Fruhstorfer, 1913 (Fig. 83) semperi Staudinger, 1889 (Fig. 93) ZMHU. The lectotype designation was carried out by Yokochi suspected is Mindanao, though the specific location is not found. The Fig. 117 is reproduced from the original description figure of the original description, but the type series specimen Note: Though the original description does not mention TS (TP): LT ♀ (SMFL). TL: Basilan. TL: Palawan. (1999). shown. The type series are deposited in the Rothschild collection The following list contains the taxonomic names bongaoensis Schröder & Treadaway, 1990 (Fig. 10) (pl. 50, fig. 4, ♂). eva C. & R. Felder, [1867] (Fig. 30) imperator Hewitson, 1863 (Fig. 40) was found after the third investigation in SMFL. We designate the number of types, a male syntype is preserved in MNHN. nirodha Fruhstorfer, 1913 (Fig. 67) Note: This taxon was described based on five materials TS (TP): ST ♂ (MNHN). TS (TP): LT ♂ (ZMHU). (BMNH, London), but we have not located them at present. concerned with Philippines. The names which are described TL: Bongao Peak, Bongao, Tawitawi Group. TL: Luzon. TL: Luzon. here again the lectotype for a male specimen shown in Fig. 49 TL: Basilan. from Luzon, Camiguin de Mindanao, and SE. Mindanao. Note: Description is based on males and females, and four tawitawia Treadaway & Nuyda, 1994 (Fig. 104) from the other areas are also included. TS (TP): HT ♀ (SMFL), PT 18♂20♀. darani Fruhstorfer, 1913 (Fig. 20) TS (TP): ST ♂ (BMNH). TS (TP): HT ♀ (BMNH). (labelled “Typus (red) / Camotes / Bohol / 113 / 211 / Coll. C. mindanaensis Schröder & Treadaway, 1978 (Fig. 60) TS (TP): ST ♂♀ (MNHN). Yokochi (1998) mentioned the type series of phelada was producta Butler, 1901 (Fig. 84) males and four females are preserved in ZMHU. The lectotype TL: Magsaggao, Tawitawi. visayana (Lexias) Schröder & Treadaway, 1987 (Fig. 110) TL: Sulu Archipelago. Note: According to the original description, eva is distrib- Semper”). TL: Surigao del Sur, Mindanao. untraced so designated the lectotype for the figure of the TL: Philippines [recte W. Malaysia]. designation was carried out by Yokochi (1999). TS (TP): HT ♂ (SMFL). TL: Catmon, S. Leyte. albusequus Nihira & Kawamura, 1986 (Fig. 2) borromeoi Schröder, 1977 (Fig. 11) TS (TP): HT ♀ (BMNH). uted in Assam, Luzon, and Celebes. The details of type series ingae (Lexias) Schröder & Treadaway, 1987 (Fig. 41) TS (TP): HT ♂ (SMFL). obatrata Yokochi, 1994 (Fig. 68) original description, but the target specimen was found after the TS (TP): ST ♀ (BMNH). TS (TP): HT ♂ (SMFL), PT 6♂4♀. TL: Lakit lakit, Sanga Sanga, Tawitawi. TL: Espana, 2000 ft, Sibuyan. Note: Though the original description does not mention are not mentioned. Staudinger (1886) is the first revisory of TL: Negros Or. [Oriental], 800 m. leytana Schröder & Treadaway, 1982 (Fig. 50) TL: Dumaran. third investigation in SMFL. We designate here again the senilis Hanafusa, 1992 (Fig. 94) tethys Tsukada, 1991 (Fig. 105) TS (TP): HT ♀ (SK). TS (TP): HT ♂ (SMFL), PT 1♀. the number of type series, one female would be the holotype. which Luzon is recognized as a locality of eva. TS (TP): HT ♂ (SMFL), PT 5♂5♀. TL: St. Bernard, Calpio, 280 m, S. Leyte. mindorana (Tanaecia) Fruhstorfer, 1904 (Fig. 61) TS (TP): HT ♀ (HH), PT 1♂1♀. lectotype for a female specimen shown in Fig. 77 (labelled rhamases Staudinger, 1889 (Fig. 85) TL: Dumaran. TL: N. Palawan. visayana (Cynitia) Schröder & Treadaway, 1981 (Fig. 111) TS (TP): HT ♂ (SMFL), PT 18♂10♀. TL: Mindoro. “Typus (red) / Euth. phelada typ. ♀ Semper / Marivcles (?) TL: Palawan. TS (TP): HT ♂ (HH), PT 1♂. TS (TP): HT ♂ (ET), PT 10♂12♀. TL: St. Bernard, Catmon, S. Leyte. 12 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 13

1. INTRODUCTION Limbach, about 250 km from Frankfurt. It is an undoubted fact amlana Jumalon, 1970 (Fig. 3) cabigoni Schröder, Treadaway, A. Mohagan & N. Mohagan, dhayma Fruhstorfer, 1913 (Fig. 21) exul Tsukada & Nishiyama, 1980 (Fig. 31) ingae (Euthalia) Schröder & Treadaway, 1990 (Fig. 42) leytensis Jumalon, 1975 (Fig. 51) TS (TP): ST 2♂1♀ (BMNH). ormocana Jumalon, 1970 (Fig. 69) 11.11.81 / 110 / Coll. C. Semper / Lectotype ♀ Euthalia TS (TP): LT ♂ (ZMHU). sibutana Schröder & Treadaway, 1990 (Fig. 95) Note: Attached label to the holotype specimen shows TS (TP): HT ♂ (SMFL), PT 10♂2♀. waltraudae Schröder & Treadaway, 1990 (Fig. 114) that Mr. Treadaway and Dr. H. Schröder of Senckenberg TL: Amlan, Negros Oriental. 2009 (Fig. 12) TL: Sulu Archipelago. TL: Bo. Melgar, Dinagat. TL: Bongao Peak, 200 m, Bongao, Tawitawi Group. TL: Lake Danao, Ormoc City, Leyte. Note: The figured female (Fig. 61) is preserved in the TL: Lake Danao, Ormoc City, Leyte. phelada Semper [87]:1888 Designated by T. Yokochi, 1997”). Note: Description is based on males, and four males are TL: Cavan Cavan, 5 m, Sibutu, Tawitawi Group. “Balabac”, but the correct type locality is Palawan. TL: Indalawan, Balabac. The Philippine Islands are situated in the northern end of thoroughly deserve credit for working further on systematic TS (TP): HT ♂ (USC), PT10 ♂1♀ (USC, JF). TL: Balut Is, Sarangani Group. TS (TP): HT ♀ (BMNH). TS (TP): HT ♂ (ET), PT 3♂6♀. TS (TP): HT ♀ (SMFL), PT 10♂12♀. TS (TP): HT ♀ (USC). Rothschild collection (Ro 98/7), BMNH. It might be one of the TS (TP): HT ♂ (USC), PT 1♀. preserved in ZMHU. The lectotype designation was carried out TS (TP): HT ♂ (SMFL), PT 17♂15♀. vistrica Fruhstorfer, 1913 (Fig. 112) TS (TP): HT ♀ (SMFL), PT 2♂2♀. the Malay Archipelago and are composed of more than 7,000 classification in Philippines Rhopalocera. TS (TP): HT ♂ (SMFL), PT 2♂1♀. Note: Though the original description does not mention the types, although the attached label is only “Mindoro”. No other philippensis Fruhstorfer, 1899 (Fig. 78) by Yokochi (1999). trapesa G. Semper, [1887] (Fig. 106) TL: locality unknown, presumably one of the Central islands, large and small, extending from north to south about Here we revised the group of the tribe Adoliadini antiquea Schröder & Treadaway, 1980 (Fig. 4) number of type series, one female would be the holotype. felicitacionae Jumalon, 1989 (Fig. 32) irauana Jumalon, 1975 (Fig. 43) liaoi Schröder & Treadaway, 1982 (Fig. 52) specimen is found that corresponded with the type series. ornata Schröder & Treadaway, 1979 (Fig. 70) TL: Basilan. sibuyana Schröder & Treadaway, 1982 (Fig. 96) TL: Camiguin de Mindanao. Philippine islands [recte Dinagat]. yoshiroui K. Okano, 1992 (Fig. 115) 1,800 km and from east to west about 1,100 km. The insect distributed in the Philippines, based on the checklist by TL: Culasi, 500 m, Antique, Panay. calliphorus C. & R. Felder, 1861 (Fig. 13) TL: Kantakpal Creek, Awihao, Ormoc, Western Leyte. TL: Irauan [Irawan], Puerto Princesa, Palawan. TL: Mt. Madja-an [Mt. Madja-as], 500 m, Antique, Panay. TL: Romblon. TS (TP): LT ♂ (MNHN). rodriguezi Schröder & Treadaway, 1982 (Fig. 86) TL: Sibuyan, Romblon Group. TS (TP): LT ♂ (SMFL). TS (TP): ST ♂ (BMNH). TL: Babuyan. fauna is rich in diversity and the tribe Adoliadini, Nymphalidae, Treadaway (1995). One hundred-twenty taxa of type specimens TS (TP): HT ♂ (SMFL). TL: Luzon. dinagatensis Tsukada, 1991 (Fig. 22) TS (TP): HT ♂ (JF). TS (TP): HT ♀ (JF), PT 1♂1♀ (USC). TS (TP): HT ♂ (SMFL), PT 1♂2♀. mindorana (Euthalia) Fruhstorfer, 1899 (Fig. 62) TS (TP): HT ♂ (SMFL), PT 1♂2♀. Note: Description is based on one male from Basilan TL: 28 km W of Puerto Princesa, Palawan. TS (TP): HT ♂ (SMFL), PT 2♂5♀. Note: Description is based on seven materials from E. Note: Tsukada (1991) suspected this taxon is distributed TS (TP): HT ♂ (KMNH). Lepidoptera, the main group of this study, is no exception. The founded in the Philippines are examined and revised the TS (TP): ST ♂ (BMNH). TL: Dinagat. Note: The holotype is deposited in Jumalon family collec- TL: Mindoro. (MNHN) and one female from Luzon (ZMHU). As the TS (TP): HT ♂ (SMFL), PT 2♂2♀. Mindanao. The lectotype designation was carried out for the in Basilan, but it presumably comes from Dinagat. important works concerned with Philippines were, back in classification of species-subspecies relation. This paper is aquamarina Fruhstorfer, 1913 (Fig. 5) TS (TP): HT ♂ (ET), PT 2♂1♀. gahiti M. & T. Okano, 1990 (unavailable name) (Fig. 33) tion. ludonia Staudinger, 1889 (Fig. 53) TS (TP): ST (MNHN). palawana (Lexias) Moore, 1897 (Fig. 71) Luzon-group should be classified as nadenya. We select a smaragdifera Fruhstorfer, 1913 (Fig. 97) Camiguin de Mindanao specimen by Yokochi (1998). yui Yokochi, 1994 (Fig. 116) history, “Diagnosen einiger neuer Tagfalter von den Philippinen published in two parts. All the taxa described before are TL: Mindanao. candida Schröder, Treadaway & Nuyda, 1990 (Fig. 14) TL: Bilar, Bohol. TL: Palawan. TL: Palawan. male (Fig. 78) deposited in MNHN as the lectotype, labelled, romeo Schröder & Treadaway, 1987 (Fig. 87) TL: Locality unknown, presumably Mindoro [Mindoro]. volupia Tsukada & Nishiyama, 1981 (Fig. 113) TL: Dumaran. und Bemerkungen über die Verbreitung einiger derselben” (G. examined and figured as much as possible in part 1. In the TS (TP): LT ♂ (SMFL). TL: 30 m NN, Calamian Group. dinorah Fruhstorfer, 1899 (Fig. 23) TS (TP): HT ♀ (FASU). joloana Staudinger, 1889 (Fig. 44) TS (TP): LT ♂ (ZMHU). mindorana (Euthalia) Tsukada, 1991 (permanently invalid) TS (TP): HT ♂ (BMNH). “Type (red) / lubentina philippensis Fruhst. / Philippinen TL: Mt. Halcon, N. Mindoro. TS (TP): ST (MNHN). treadawayi Tsukada, 1991 (Fig. 107) TL: Camiguin. TS (TP): HT ♂ (TY), PT 11♂4♀. Semper, 1886–1892), and then “Die Gross-Schmetterlinge der second part, synonymic lists and some important specimens Note: Description is based on some specimens. The lecto- TS (TP): HT ♂ (SMFL), PT10 ♂. TL: Basilan. Note: As this taxon gahiti was described as a form of TL: Sulu. Note: Description is based on four males and six females, Note: The taxon mindorana described under the genus Bazilan II-III, 98 Doherty ex. coll. H. Fruhstorfer / TS (TP): HT ♀ (SMFL), PT 1♀. Note: Though the original description does not mention TL: Romblon, Sibuyanes. TS (TP): HT ♂ (ET), PT 2♂5♀. (to be continued) Erde, 9” (1908–1927) edited by A. Seitz. Recently, a Japanese will be shown, affixing several discussions. type designation was carried out by Yokochi (1998). TS (TP): ST 2♂2♀ (MNHN). Lexias panopus boholensis (infrasubspecific name), it should TS (TP): LT ♂ (ZMHU). and two pairs are preserved in ZMHU. The lectotype designa- Euthalia is a junior primary homonym of Euthalia lusiada palawana (Tanaecia) Staudinger, 1889 (Fig. 72) MUSEUM PARIS 1934 COLL H Fruhstorfer”. the number and sex, a syntype specimen in MNHN is a male. TS (TP): HT ♂ (SMFL). entomologist, E. Tsukada, had compiled a great series of cavarna Fruhstorfer, 1913 (Fig. 15) be treated as an unavailable name. Note: Description is based on two males and one female, tion was carried out by Yokochi (1999). mindorana Fruhstorfer, 1899 (permanently invalid name). TL: Palawan. rudraca Fruhstorfer, 1913 (Fig. 88) “Butterflies of the South East Asian Islands” (1980–1991). balabacana (Bassarona) Tsukada, 1991 (Fig. 6) TL: Balabas [Balabac]. dodong Schröder & Treadaway, 1978 (Fig. 24) and all of them are preserved in ZMHU. The lectotype desig- Now it is replaced by tsukada Koçak (1996). TS (TP): LT ♂ (ZMHU). phlegethon G. Semper, [1887] (Fig. 79) TL: Balabac. soloni M. & T. Okano, 1990 (Fig. 98) tsukada Koçak, 1996 (Fig. 80) The late Prof. Julian N. Jumalon (1909–2000) (Fig. 1, TL: Balabac. TS (TP): LT ♀ (ZMHU). TL: Mobo, 1000 ft, Masbate. galoa Fruhstorfer, 1913 (Fig. 34) nation was carried out by Yokochi (1999). lupina Druce, 1874 (Fig. 54) Note: Description is based on males and females, and six TL: Mindanao. TS (TP): LT ♂ (ZMHU). TL: Bilar, Bohol. TL: Mindoro. left), the Lepidoptera Research Associate and Lecturer of 2. ABBREVIATIONS TS (TP): HT ♂ (ET), PT 8♂6♀. Note: Description is based on a few females, and one spec- TS (TP): HT ♂ (SMFL), PT 3♂7♀. TL: Mindoro. TL: Philippine Islands [Sulu]. mindorensis Schröder & Treadaway, 1982 (Fig. 63) males and three females are preserved in ZMHU. The lecto- TS (TP): LT ♀ (SMFL). Note: Description is based on some specimens, and two TS (TP): HT ♂ (FASU), PT 2♂3♀. TS (TP): HT ♂ (ET), PT 10♂11♀. Department of Biology, University of San Carlos, published a imen is preserved in ZMHU. The lectotype designation was TS (TP): ST (BMNH). kayumanggia Treadaway & Nuyda, 1994 (Fig. 45) TS (TP): HT ♀ (BMNH). TL: Mindoro. type designation was carried out by Yokochi (1999). Note: Description is based on twenty-one males and seven males and two females are preserved in ZMHU. The lectotype Note: Euthalia lubentina mindorana Tsukada, 1991 is a paper of “New Philippine butterflies” in 1975, and many new Public Institution (Museum, University). BMNH: The balabacana (Dophla) Tsukada, 1991 (secondary homonym) carried out by Yokochi (1999). dohertyi Butler, 1901 (Fig. 25) Note: Though the original description does not mention TL: Mt. Bayog, S. Palawan. TS (TP): HT ♂ (SMFL), PT 6♂7♀. females from Samar, Camiguin de Mindanao, N. Mindanao, E. designation was carried out by Yokochi (1999). soregina Fruhstorfer, 1913 (Fig. 99) permanently invalid name (nom. nov. pro mindorana Tsukada, taxonomic names concerned with various families were Natural History Museum, London, United Kingdom; FASU: (Fig. 7) TL: Sulu Archipelago. the number and sex, syntype specimen in BMNH is a male. TS (TP): HT ♂ (SMFL), PT 13♂4♀. lusiada C. & R. Felder, 1863 (Fig. 55) palawana (Euthalia) Staudinger, 1889 (Fig. 73) Mindanao, SE. Mindanao, and several materials are preserved TL: Sulu Archipelago. 1991, non Fruhstorfer,1899). New name tsukada was proposed described there. He played a leading part of the study of Exhibition Hall of Food, Agricultural and Environmental Note: See below balabacensis. chloe Schröder & Treadaway, 1990 (Fig. 16) TS (TP): HT ♂ (BMNH). TL: Luzon. miscus Fruhstorfer, 1913 (Fig. 64) TL: Palawan. in SMFL. The lectotype designation was carried out for the samarensis Tsukada, 1991 (Fig. 89) TS (TP): ST ♀ (BMNH). by Koçak (1996). Philippine butterflies and contributed greatly to entomology. Sciences, Faculty of Agriculture, Shinshu University, Nagano, TL: Jao Asan, 200 m, Sibuyan, Romblon Group. goertzi Jumalon, 1975 (Fig. 35) kulaya Treadaway & Nuyda, 1994 (Fig. 46) TS (TP): ST ♂ (SMFL). TL: Mindanao. TS (TP): LT ♂ (ZMHU). Mindanao specimen by Yokochi (1998). TL: San Juan, Samar. Note: According to the original description, the type series In 1995, the late Mr. Colin G. Treadaway (Fig. 1, right, Japan; KMNH: Kitakyushu Museum of Natural History & balabacensis nom. nov. (Fig. 7) TS (TP): HT ♂ (SMFL), PT 11♂9♀. dumaranensis Hanafusa, 1992 (Fig. 26) TL: Amlan, Negros Oriental. TL: Magellanes, Homonhon. Note: Though the original description does not mention TS (TP): ST (MNHN, BMNH). Note: Description is based on one male and six females, TS (TP): HT ♂ (ET), PT 3♂2♀. are exactly female, but the number is unknown. tyawena Fruhstorfer, 1913 (Fig. 108) died in 2019) published “Checklist of the butterflies of the Human History, Fukuoka, Japan; MNHN: Museum National TL: Balabac. TL: Dumaran. TS (TP): HT ♀ (ET). TS (TP): HT ♂ (SMFL), PT 14♂10♀. the number of types, a male syntype is preserved in SMFL. Note: According to the original description, types and one male and four females are preserved in ZMHU. The pinamalayana Yokochi, 1997 (Fig. 80) TL: unknown [Palawan]. Philippine Islands (Lepidoptera: Rhopalocera)”. Although, 25 d'Histoire Naturelle Entomologie, Paris, France; SMFL: TS (TP): HT ♂ (ET), PT 2♂2♀. circe Schröder & Treadaway, 1990 (Fig. 17) TS (TP): HT ♂ (HH). Note: Though the original description mentions the Note: The name kulaya is spelt “kúlaya” in the original (syntypes) are preserved in Fruhstorfer collection (now in lectotype designation was carried out by Yokochi (1999). Note: The new name pinamalayana was proposed by sarmana Fruhstorfer, 1913 (Fig. 90) subpiratica Schröder & Treadaway, 1987 (Fig. 100) TS (TP): LT ♀ (ZMHU). years have passed since his paper presented, several new Senckenberg Forschungsinstitut und Naturmuseum Frankfurt, Note: The taxon balabacana was described under the TL: Cavan Cavan, Sibutu, Tawitawi Group. holotype is preserved in USC, it is deposited in ET. description. macer Tsukada & Nishiyama, 1980 (Fig. 56) MNHN) and BMNH. Yokochi (1997), because mindorana Tsukada, 1991 was a TL: Basilan. TL: Nueva Ecija, Bongabon, Centr. Luzon. Note: Description is based on some specimens, and one findings are still remained unpublished — it can be followed Frankfurt-am-Main, Germany; USC: University of San Carlos, genus Dophla. As Yokochi (2010) assigned both Dophla and TS (TP): HT ♂ (SMFL), PT 9♂10♀. dunya Doubleday, [1848] (Fig. 27) TL: Bo. Melgar, Dinagat. pallida Schröder, Treadaway & Nuyda, 1990 (Fig. 74) permanently invalid name. However, pinamalayana is a junior TS (TP): ST (MNHN). TS (TP): HT ♂ (SMFL), PT1♀. male and one female are preserved in ZMHU. The lectotype by his own handwriting on the glass surface of his specimen Cebu, Philippines; ZMHU: Zoologisches Museum, Humboldt Bassarona to the genus Euthalia, balabacana (Dophla) is a TL: Assam, India. hikarugenzi Tsukada & Nishiyama, 1980 (Fig. 36) laetitiae Schröder & Treadaway, 2000 (Fig. 47) TS (TP): HT ♂ (ET), PT 3♂2♀. monara Fruhstorfer, 1913 (Fig. 65) TL: 30 m NN, Calamian Group. synonym of tsukada Koçak (1996). Note: Though the original description does not mention designation was carried out by Yokochi (1999). cases —. The first author visited Senckenberg Museum in Universität, Berlin, Germany. junior secondary homonym of Bassarona teuta balabacana cusama Fruhstorfer, 1913 (Fig. 18) TS (TP): ST ♂ (BMNH). TL: near Baguio, N. Luzon. TL: Cavan Cavan, 5 m, Sibutu, Tawitawi Group. TL: N. Borneo. TS (TP): HT ♂ (SMFL), PT 11♂16♀. the number and sex, a syntype specimen in MNHN is a male. suluana Fruhstorfer, 1902 (Fig. 101) Frankfurt to research on the Treadaway collection on March Private collections. ET: Etsuzo Tsukada, Yamanashi, Tsukada, 1991. Here a new name balabacensis is replaced for TL: Mindanao, Camiguin de Mindanao. TS (TP): HT ♂ (MKN), PT 1♀ (MKN). TS (TP): HT ♂ (SMFL), PT 4♂. malissia Fruhstorfer, 1913 (Fig. 57) TS (TP): ST (MNHN). piratica G. Semper, [1887] (Fig. 81) TL: Sulu Archipelago. uposatha Fruhstorfer, 1913 (Fig. 109) 2019, and just then was informed that he had died several Japan; HH: Hiroto Hanafusa, Tottori, Japan; JF: Jumalon Dophla evelina balabacana Tsukada, 1991. TS (TP): ST ♂ (MNHN), ♀ (SMFL). ellora Fruhstorfer, 1898 (Fig. 28) TL: Basilan. panayana Schröder & Treadaway, 1980 (Fig. 75) TL: Camiguin de Mindanao. satrapes C. & R. Felder, 1861 (Fig. 91) TS (TP): HT ♀ (BMNH). TL: Banggi (Banguey), E. Malaysia. weeks ago at his home in England. In the middle 1990’s, every Family, Cebu, Philippines; MKN: Misao Kaneko, Tokyo, Euthalia (Dophla) evelina balabacensis nom. nov. Note: Figured specimen is a syntype female in SMFL. TL: Mindoro. hinunanganensis M. & T. Okano, 1986 (Fig. 37) leopardina Fruhstorfer, 1913 (Fig. 48) TS (TP): ST♂ (MNHN). nadenya Fruhstorfer, 1913 TL: Culasi, 800 ft, Antique, Panay. TS (TP): LT ♂ (SMFL). TL: Luzon. TS (TP): LT ♂ (ZMHU). time the first author visited to Senckenberg, he came bringing Japan; SK: Shun-ichi Kawamura, Tokyo, Japan; STA: Shinji pro Dophla evelina balabacana Tsukada, 1991, non TS (TP): ST ♂ (MNHN). TL: Hinunangan, Leyte. TL: Sulu Islands. Note: Though the original description does not mention TL: Luzon. TS (TP): HT ♂ (SMFL), PT 4♂4♀. Note: Description is based on six materials from Luzon, TS (TP): ST ♂ (BMNH). susoni Jumalon, 1975 (Fig. 102) Note: According to the original description, the type series his described type specimens, with his wife driving from Taga, Tokyo, Japan; TY: Takashi Yokochi, Aichi, Japan. Bassarona teuta balabacana Tsukada, 1991 dacasini Hanafusa, 1990 (Fig. 19) Note: Though the original description does not mention TS (TP): HT ♂ (FASU), PT 1♂. TS (TP): HT ♀ (BMNH). the number of types, a male syntype is preserved in MNHN. TS (TP): HT ♀ [untraced, not examined]. Mindoro, Camiguin de Mindanao, N. Mindanao. The lectotype Note: According to the original description, the type series TL: Camp 7, Minglanilla, Cebu. (male and female, but the number is unknown) are deposited in Specimens concerned. FW: Length of Forewing; HT: TL: Balabac. the number and sex, a syntype specimen in MNHN is a male. Note: Though the original description indicates that the panopus C. & R. Felder, 1861 (Fig. 76) designation was carried out for the Camiguin de Mindanao (number is unknown) are deposited in Semper and Felder TS (TP): HT ♂ (USC), PT 9♂9♀ (USC, JF, SMFL, TY). Staudinger collection (ZMHU). The lectotype designation was Holotype; LT: Lectotype; PLT: Paralectotype; PT: Paratype; boholensis (Lexias) M. & T. Okano, 1988 (Fig. 8) TS (TP): HT ♂ (HH), PT 1♀. hiwaga Nuyda & Kawamura, 1989 (Fig. 38) leucotaenia G. Semper, 1878 (Fig. 49) marinduque K. Okano, 1992 (Fig. 58) type female should be preserved in ZMHU, the specimen TL: Luzon. specimen by Schröder & Treadaway (1987). collections. carried out by Yokochi (1999). TL: Type Locality; TP: Type Preservation; TS: Type Series; TL: Bilar, Bohol. eson de Nicéville, 1894 (Fig. 29) TL: Camiguin, Babuyan. TL: Camotes. TL: Marinduque. corresponding with nadenya could not be found there. No TS (TP): ST ♂ (BMNH). tanagra Staudinger, 1889 (Fig. 103) ST: Syntype. TS (TP): HT ♂ (FASU), PT 6♂2♀. damalis Erichson, 1834 (Fig. 117) TL: Palawan. TS (TP): HT ♂ (SK), PT 7♂4♀ (SK, ET). TS (TP): LT ♂ (SMFL). TS (TP): HT ♀ (KMNH), PT 1♂ (KMNH). figure is presented in the original paper. Note: The original description does not mention the princesa Fruhstorfer, 1899 (Fig. 82) schoenigi Schröder & Treadaway, 1978 (Fig. 92) TL: Palawan. virginalis Fruhstorfer, 1913 TL: Luzon. TS (TP): LT ♂ (SMFL), PLT 1♀. Note: This taxon was described based on sixty-nine number of type series. TL: Palawan. TL: Amlan, 600 m, Negros. TS (TP): LT ♂ (ZMHU). TL: locality not given. boholensis (Euthalia) M. & T. Okano, 1990 (Fig. 9) TS (TP): ST ♂ [untraced, not examined]. Note: Description is based on a single pair, and all howarthi Jumalon, 1975 (Fig. 39) materials from Camotes, Bohol, Leyte, Panaon, E. Mindanao, medaga Fruhstorfer, 1913 (Fig. 59) negrosiana Schröder & Treadaway, 1987 (Fig. 66) TS (TP): HT ♀ (MNHN). TS (TP): HT ♀ (SMFL). Note: Description is based on seven males and eleven TS (TP): ST ♀ (BMNH) [not examined]. 3. LIST OF DESCRIBED SPECIES AND TL: Bilar, Bohol. Note: The type series should be preserved in ZMHU, but materials are preserved in SMFL. The lectotype designation TL: Amlan, Negros Oriental. and N. Mindanao. Yokochi (1998) mentioned the type series of TL: Camiguin de Mindanao. TL: Negros Or. [Oriental]. phelada G. Semper, [1887] (Fig. 77) females, and two males and three females are preserved in Note: This is a female form of panopus. The type locality SUBSPECIES NAME TS (TP): HT ♀ (FASU), PT 4♀. the specimens that corresponded with the types could not be was carried out by Yokochi (1998). TS (TP): HT ♂ (USC), PT 4♀ (USC, JF). leucotaenia was untraced so designated the lectotype for the TS (TP): ST ♂ (MNHN). TS (TP): HT ♂ (SMFL), PT 1♂1♀. TL: Luzon. proditrix Fruhstorfer, 1913 (Fig. 83) semperi Staudinger, 1889 (Fig. 93) ZMHU. The lectotype designation was carried out by Yokochi suspected is Mindanao, though the specific location is not found. The Fig. 117 is reproduced from the original description figure of the original description, but the type series specimen Note: Though the original description does not mention TS (TP): LT ♀ (SMFL). TL: Basilan. TL: Palawan. (1999). shown. The type series are deposited in the Rothschild collection The following list contains the taxonomic names bongaoensis Schröder & Treadaway, 1990 (Fig. 10) (pl. 50, fig. 4, ♂). eva C. & R. Felder, [1867] (Fig. 30) imperator Hewitson, 1863 (Fig. 40) was found after the third investigation in SMFL. We designate the number of types, a male syntype is preserved in MNHN. nirodha Fruhstorfer, 1913 (Fig. 67) Note: This taxon was described based on five materials TS (TP): ST ♂ (MNHN). TS (TP): LT ♂ (ZMHU). (BMNH, London), but we have not located them at present. concerned with Philippines. The names which are described TL: Bongao Peak, Bongao, Tawitawi Group. TL: Luzon. TL: Luzon. here again the lectotype for a male specimen shown in Fig. 49 TL: Basilan. from Luzon, Camiguin de Mindanao, and SE. Mindanao. Note: Description is based on males and females, and four tawitawia Treadaway & Nuyda, 1994 (Fig. 104) from the other areas are also included. TS (TP): HT ♀ (SMFL), PT 18♂20♀. darani Fruhstorfer, 1913 (Fig. 20) TS (TP): ST ♂ (BMNH). TS (TP): HT ♀ (BMNH). (labelled “Typus (red) / Camotes / Bohol / 113 / 211 / Coll. C. mindanaensis Schröder & Treadaway, 1978 (Fig. 60) TS (TP): ST ♂♀ (MNHN). Yokochi (1998) mentioned the type series of phelada was producta Butler, 1901 (Fig. 84) males and four females are preserved in ZMHU. The lectotype TL: Magsaggao, Tawitawi. visayana (Lexias) Schröder & Treadaway, 1987 (Fig. 110) TL: Sulu Archipelago. Note: According to the original description, eva is distrib- Semper”). TL: Surigao del Sur, Mindanao. untraced so designated the lectotype for the figure of the TL: Philippines [recte W. Malaysia]. designation was carried out by Yokochi (1999). TS (TP): HT ♂ (SMFL). TL: Catmon, S. Leyte. albusequus Nihira & Kawamura, 1986 (Fig. 2) borromeoi Schröder, 1977 (Fig. 11) TS (TP): HT ♀ (BMNH). uted in Assam, Luzon, and Celebes. The details of type series ingae (Lexias) Schröder & Treadaway, 1987 (Fig. 41) TS (TP): HT ♂ (SMFL). obatrata Yokochi, 1994 (Fig. 68) original description, but the target specimen was found after the TS (TP): ST ♀ (BMNH). TS (TP): HT ♂ (SMFL), PT 6♂4♀. TL: Lakit lakit, Sanga Sanga, Tawitawi. TL: Espana, 2000 ft, Sibuyan. Note: Though the original description does not mention are not mentioned. Staudinger (1886) is the first revisory of TL: Negros Or. [Oriental], 800 m. leytana Schröder & Treadaway, 1982 (Fig. 50) TL: Dumaran. third investigation in SMFL. We designate here again the senilis Hanafusa, 1992 (Fig. 94) tethys Tsukada, 1991 (Fig. 105) TS (TP): HT ♀ (SK). TS (TP): HT ♂ (SMFL), PT 1♀. the number of type series, one female would be the holotype. which Luzon is recognized as a locality of eva. TS (TP): HT ♂ (SMFL), PT 5♂5♀. TL: St. Bernard, Calpio, 280 m, S. Leyte. mindorana (Tanaecia) Fruhstorfer, 1904 (Fig. 61) TS (TP): HT ♀ (HH), PT 1♂1♀. lectotype for a female specimen shown in Fig. 77 (labelled rhamases Staudinger, 1889 (Fig. 85) TL: Dumaran. TL: N. Palawan. visayana (Cynitia) Schröder & Treadaway, 1981 (Fig. 111) TS (TP): HT ♂ (SMFL), PT 18♂10♀. TL: Mindoro. “Typus (red) / Euth. phelada typ. ♀ Semper / Marivcles (?) TL: Palawan. TS (TP): HT ♂ (HH), PT 1♂. TS (TP): HT ♂ (ET), PT 10♂12♀. TL: St. Bernard, Catmon, S. Leyte. 12 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 13

1. INTRODUCTION Limbach, about 250 km from Frankfurt. It is an undoubted fact amlana Jumalon, 1970 (Fig. 3) cabigoni Schröder, Treadaway, A. Mohagan & N. Mohagan, dhayma Fruhstorfer, 1913 (Fig. 21) exul Tsukada & Nishiyama, 1980 (Fig. 31) ingae (Euthalia) Schröder & Treadaway, 1990 (Fig. 42) leytensis Jumalon, 1975 (Fig. 51) TS (TP): ST 2♂1♀ (BMNH). ormocana Jumalon, 1970 (Fig. 69) 11.11.81 / 110 / Coll. C. Semper / Lectotype ♀ Euthalia TS (TP): LT ♂ (ZMHU). sibutana Schröder & Treadaway, 1990 (Fig. 95) Note: Attached label to the holotype specimen shows TS (TP): HT ♂ (SMFL), PT 10♂2♀. waltraudae Schröder & Treadaway, 1990 (Fig. 114) that Mr. Treadaway and Dr. H. Schröder of Senckenberg TL: Amlan, Negros Oriental. 2009 (Fig. 12) TL: Sulu Archipelago. TL: Bo. Melgar, Dinagat. TL: Bongao Peak, 200 m, Bongao, Tawitawi Group. TL: Lake Danao, Ormoc City, Leyte. Note: The figured female (Fig. 61) is preserved in the TL: Lake Danao, Ormoc City, Leyte. phelada Semper [87]:1888 Designated by T. Yokochi, 1997”). Note: Description is based on males, and four males are TL: Cavan Cavan, 5 m, Sibutu, Tawitawi Group. “Balabac”, but the correct type locality is Palawan. TL: Indalawan, Balabac. The Philippine Islands are situated in the northern end of thoroughly deserve credit for working further on systematic TS (TP): HT ♂ (USC), PT10 ♂1♀ (USC, JF). TL: Balut Is, Sarangani Group. TS (TP): HT ♀ (BMNH). TS (TP): HT ♂ (ET), PT 3♂6♀. TS (TP): HT ♀ (SMFL), PT 10♂12♀. TS (TP): HT ♀ (USC). Rothschild collection (Ro 98/7), BMNH. It might be one of the TS (TP): HT ♂ (USC), PT 1♀. preserved in ZMHU. The lectotype designation was carried out TS (TP): HT ♂ (SMFL), PT 17♂15♀. vistrica Fruhstorfer, 1913 (Fig. 112) TS (TP): HT ♀ (SMFL), PT 2♂2♀. the Malay Archipelago and are composed of more than 7,000 classification in Philippines Rhopalocera. TS (TP): HT ♂ (SMFL), PT 2♂1♀. Note: Though the original description does not mention the types, although the attached label is only “Mindoro”. No other philippensis Fruhstorfer, 1899 (Fig. 78) by Yokochi (1999). trapesa G. Semper, [1887] (Fig. 106) TL: locality unknown, presumably one of the Central islands, large and small, extending from north to south about Here we revised the group of the tribe Adoliadini antiquea Schröder & Treadaway, 1980 (Fig. 4) number of type series, one female would be the holotype. felicitacionae Jumalon, 1989 (Fig. 32) irauana Jumalon, 1975 (Fig. 43) liaoi Schröder & Treadaway, 1982 (Fig. 52) specimen is found that corresponded with the type series. ornata Schröder & Treadaway, 1979 (Fig. 70) TL: Basilan. sibuyana Schröder & Treadaway, 1982 (Fig. 96) TL: Camiguin de Mindanao. Philippine islands [recte Dinagat]. yoshiroui K. Okano, 1992 (Fig. 115) 1,800 km and from east to west about 1,100 km. The insect distributed in the Philippines, based on the checklist by TL: Culasi, 500 m, Antique, Panay. calliphorus C. & R. Felder, 1861 (Fig. 13) TL: Kantakpal Creek, Awihao, Ormoc, Western Leyte. TL: Irauan [Irawan], Puerto Princesa, Palawan. TL: Mt. Madja-an [Mt. Madja-as], 500 m, Antique, Panay. TL: Romblon. TS (TP): LT ♂ (MNHN). rodriguezi Schröder & Treadaway, 1982 (Fig. 86) TL: Sibuyan, Romblon Group. TS (TP): LT ♂ (SMFL). TS (TP): ST ♂ (BMNH). TL: Babuyan. fauna is rich in diversity and the tribe Adoliadini, Nymphalidae, Treadaway (1995). One hundred-twenty taxa of type specimens TS (TP): HT ♂ (SMFL). TL: Luzon. dinagatensis Tsukada, 1991 (Fig. 22) TS (TP): HT ♂ (JF). TS (TP): HT ♀ (JF), PT 1♂1♀ (USC). TS (TP): HT ♂ (SMFL), PT 1♂2♀. mindorana (Euthalia) Fruhstorfer, 1899 (Fig. 62) TS (TP): HT ♂ (SMFL), PT 1♂2♀. Note: Description is based on one male from Basilan TL: 28 km W of Puerto Princesa, Palawan. TS (TP): HT ♂ (SMFL), PT 2♂5♀. Note: Description is based on seven materials from E. Note: Tsukada (1991) suspected this taxon is distributed TS (TP): HT ♂ (KMNH). Lepidoptera, the main group of this study, is no exception. The founded in the Philippines are examined and revised the TS (TP): ST ♂ (BMNH). TL: Dinagat. Note: The holotype is deposited in Jumalon family collec- TL: Mindoro. (MNHN) and one female from Luzon (ZMHU). As the TS (TP): HT ♂ (SMFL), PT 2♂2♀. Mindanao. The lectotype designation was carried out for the in Basilan, but it presumably comes from Dinagat. important works concerned with Philippines were, back in classification of species-subspecies relation. This paper is aquamarina Fruhstorfer, 1913 (Fig. 5) TS (TP): HT ♂ (ET), PT 2♂1♀. gahiti M. & T. Okano, 1990 (unavailable name) (Fig. 33) tion. ludonia Staudinger, 1889 (Fig. 53) TS (TP): ST (MNHN). palawana (Lexias) Moore, 1897 (Fig. 71) Luzon-group should be classified as nadenya. We select a smaragdifera Fruhstorfer, 1913 (Fig. 97) Camiguin de Mindanao specimen by Yokochi (1998). yui Yokochi, 1994 (Fig. 116) history, “Diagnosen einiger neuer Tagfalter von den Philippinen published in two parts. All the taxa described before are TL: Mindanao. candida Schröder, Treadaway & Nuyda, 1990 (Fig. 14) TL: Bilar, Bohol. TL: Palawan. TL: Palawan. male (Fig. 78) deposited in MNHN as the lectotype, labelled, romeo Schröder & Treadaway, 1987 (Fig. 87) TL: Locality unknown, presumably Mindoro [Mindoro]. volupia Tsukada & Nishiyama, 1981 (Fig. 113) TL: Dumaran. und Bemerkungen über die Verbreitung einiger derselben” (G. examined and figured as much as possible in part 1. In the TS (TP): LT ♂ (SMFL). TL: 30 m NN, Calamian Group. dinorah Fruhstorfer, 1899 (Fig. 23) TS (TP): HT ♀ (FASU). joloana Staudinger, 1889 (Fig. 44) TS (TP): LT ♂ (ZMHU). mindorana (Euthalia) Tsukada, 1991 (permanently invalid) TS (TP): HT ♂ (BMNH). “Type (red) / lubentina philippensis Fruhst. / Philippinen TL: Mt. Halcon, N. Mindoro. TS (TP): ST (MNHN). treadawayi Tsukada, 1991 (Fig. 107) TL: Camiguin. TS (TP): HT ♂ (TY), PT 11♂4♀. Semper, 1886–1892), and then “Die Gross-Schmetterlinge der second part, synonymic lists and some important specimens Note: Description is based on some specimens. The lecto- TS (TP): HT ♂ (SMFL), PT10 ♂. TL: Basilan. Note: As this taxon gahiti was described as a form of TL: Sulu. Note: Description is based on four males and six females, Note: The taxon mindorana described under the genus Bazilan II-III, 98 Doherty ex. coll. H. Fruhstorfer / TS (TP): HT ♀ (SMFL), PT 1♀. Note: Though the original description does not mention TL: Romblon, Sibuyanes. TS (TP): HT ♂ (ET), PT 2♂5♀. (to be continued) Erde, 9” (1908–1927) edited by A. Seitz. Recently, a Japanese will be shown, affixing several discussions. type designation was carried out by Yokochi (1998). TS (TP): ST 2♂2♀ (MNHN). Lexias panopus boholensis (infrasubspecific name), it should TS (TP): LT ♂ (ZMHU). and two pairs are preserved in ZMHU. The lectotype designa- Euthalia is a junior primary homonym of Euthalia lusiada palawana (Tanaecia) Staudinger, 1889 (Fig. 72) MUSEUM PARIS 1934 COLL H Fruhstorfer”. the number and sex, a syntype specimen in MNHN is a male. TS (TP): HT ♂ (SMFL). entomologist, E. Tsukada, had compiled a great series of cavarna Fruhstorfer, 1913 (Fig. 15) be treated as an unavailable name. Note: Description is based on two males and one female, tion was carried out by Yokochi (1999). mindorana Fruhstorfer, 1899 (permanently invalid name). TL: Palawan. rudraca Fruhstorfer, 1913 (Fig. 88) “Butterflies of the South East Asian Islands” (1980–1991). balabacana (Bassarona) Tsukada, 1991 (Fig. 6) TL: Balabas [Balabac]. dodong Schröder & Treadaway, 1978 (Fig. 24) and all of them are preserved in ZMHU. The lectotype desig- Now it is replaced by tsukada Koçak (1996). TS (TP): LT ♂ (ZMHU). phlegethon G. Semper, [1887] (Fig. 79) TL: Balabac. soloni M. & T. Okano, 1990 (Fig. 98) tsukada Koçak, 1996 (Fig. 80) The late Prof. Julian N. Jumalon (1909–2000) (Fig. 1, TL: Balabac. TS (TP): LT ♀ (ZMHU). TL: Mobo, 1000 ft, Masbate. galoa Fruhstorfer, 1913 (Fig. 34) nation was carried out by Yokochi (1999). lupina Druce, 1874 (Fig. 54) Note: Description is based on males and females, and six TL: Mindanao. TS (TP): LT ♂ (ZMHU). TL: Bilar, Bohol. TL: Mindoro. left), the Lepidoptera Research Associate and Lecturer of 2. ABBREVIATIONS TS (TP): HT ♂ (ET), PT 8♂6♀. Note: Description is based on a few females, and one spec- TS (TP): HT ♂ (SMFL), PT 3♂7♀. TL: Mindoro. TL: Philippine Islands [Sulu]. mindorensis Schröder & Treadaway, 1982 (Fig. 63) males and three females are preserved in ZMHU. The lecto- TS (TP): LT ♀ (SMFL). Note: Description is based on some specimens, and two TS (TP): HT ♂ (FASU), PT 2♂3♀. TS (TP): HT ♂ (ET), PT 10♂11♀. Department of Biology, University of San Carlos, published a imen is preserved in ZMHU. The lectotype designation was TS (TP): ST (BMNH). kayumanggia Treadaway & Nuyda, 1994 (Fig. 45) TS (TP): HT ♀ (BMNH). TL: Mindoro. type designation was carried out by Yokochi (1999). Note: Description is based on twenty-one males and seven males and two females are preserved in ZMHU. The lectotype Note: Euthalia lubentina mindorana Tsukada, 1991 is a paper of “New Philippine butterflies” in 1975, and many new Public Institution (Museum, University). BMNH: The balabacana (Dophla) Tsukada, 1991 (secondary homonym) carried out by Yokochi (1999). dohertyi Butler, 1901 (Fig. 25) Note: Though the original description does not mention TL: Mt. Bayog, S. Palawan. TS (TP): HT ♂ (SMFL), PT 6♂7♀. females from Samar, Camiguin de Mindanao, N. Mindanao, E. designation was carried out by Yokochi (1999). soregina Fruhstorfer, 1913 (Fig. 99) permanently invalid name (nom. nov. pro mindorana Tsukada, taxonomic names concerned with various families were Natural History Museum, London, United Kingdom; FASU: (Fig. 7) TL: Sulu Archipelago. the number and sex, syntype specimen in BMNH is a male. TS (TP): HT ♂ (SMFL), PT 13♂4♀. lusiada C. & R. Felder, 1863 (Fig. 55) palawana (Euthalia) Staudinger, 1889 (Fig. 73) Mindanao, SE. Mindanao, and several materials are preserved TL: Sulu Archipelago. 1991, non Fruhstorfer,1899). New name tsukada was proposed described there. He played a leading part of the study of Exhibition Hall of Food, Agricultural and Environmental Note: See below balabacensis. chloe Schröder & Treadaway, 1990 (Fig. 16) TS (TP): HT ♂ (BMNH). TL: Luzon. miscus Fruhstorfer, 1913 (Fig. 64) TL: Palawan. in SMFL. The lectotype designation was carried out for the samarensis Tsukada, 1991 (Fig. 89) TS (TP): ST ♀ (BMNH). by Koçak (1996). Philippine butterflies and contributed greatly to entomology. Sciences, Faculty of Agriculture, Shinshu University, Nagano, TL: Jao Asan, 200 m, Sibuyan, Romblon Group. goertzi Jumalon, 1975 (Fig. 35) kulaya Treadaway & Nuyda, 1994 (Fig. 46) TS (TP): ST ♂ (SMFL). TL: Mindanao. TS (TP): LT ♂ (ZMHU). Mindanao specimen by Yokochi (1998). TL: San Juan, Samar. Note: According to the original description, the type series In 1995, the late Mr. Colin G. Treadaway (Fig. 1, right, Japan; KMNH: Kitakyushu Museum of Natural History & balabacensis nom. nov. (Fig. 7) TS (TP): HT ♂ (SMFL), PT 11♂9♀. dumaranensis Hanafusa, 1992 (Fig. 26) TL: Amlan, Negros Oriental. TL: Magellanes, Homonhon. Note: Though the original description does not mention TS (TP): ST (MNHN, BMNH). Note: Description is based on one male and six females, TS (TP): HT ♂ (ET), PT 3♂2♀. are exactly female, but the number is unknown. tyawena Fruhstorfer, 1913 (Fig. 108) died in 2019) published “Checklist of the butterflies of the Human History, Fukuoka, Japan; MNHN: Museum National TL: Balabac. TL: Dumaran. TS (TP): HT ♀ (ET). TS (TP): HT ♂ (SMFL), PT 14♂10♀. the number of types, a male syntype is preserved in SMFL. Note: According to the original description, types and one male and four females are preserved in ZMHU. The pinamalayana Yokochi, 1997 (Fig. 80) TL: unknown [Palawan]. Philippine Islands (Lepidoptera: Rhopalocera)”. Although, 25 d'Histoire Naturelle Entomologie, Paris, France; SMFL: TS (TP): HT ♂ (ET), PT 2♂2♀. circe Schröder & Treadaway, 1990 (Fig. 17) TS (TP): HT ♂ (HH). Note: Though the original description mentions the Note: The name kulaya is spelt “kúlaya” in the original (syntypes) are preserved in Fruhstorfer collection (now in lectotype designation was carried out by Yokochi (1999). Note: The new name pinamalayana was proposed by sarmana Fruhstorfer, 1913 (Fig. 90) subpiratica Schröder & Treadaway, 1987 (Fig. 100) TS (TP): LT ♀ (ZMHU). years have passed since his paper presented, several new Senckenberg Forschungsinstitut und Naturmuseum Frankfurt, Note: The taxon balabacana was described under the TL: Cavan Cavan, Sibutu, Tawitawi Group. holotype is preserved in USC, it is deposited in ET. description. macer Tsukada & Nishiyama, 1980 (Fig. 56) MNHN) and BMNH. Yokochi (1997), because mindorana Tsukada, 1991 was a TL: Basilan. TL: Nueva Ecija, Bongabon, Centr. Luzon. Note: Description is based on some specimens, and one findings are still remained unpublished — it can be followed Frankfurt-am-Main, Germany; USC: University of San Carlos, genus Dophla. As Yokochi (2010) assigned both Dophla and TS (TP): HT ♂ (SMFL), PT 9♂10♀. dunya Doubleday, [1848] (Fig. 27) TL: Bo. Melgar, Dinagat. pallida Schröder, Treadaway & Nuyda, 1990 (Fig. 74) permanently invalid name. However, pinamalayana is a junior TS (TP): ST (MNHN). TS (TP): HT ♂ (SMFL), PT1♀. male and one female are preserved in ZMHU. The lectotype by his own handwriting on the glass surface of his specimen Cebu, Philippines; ZMHU: Zoologisches Museum, Humboldt Bassarona to the genus Euthalia, balabacana (Dophla) is a TL: Assam, India. hikarugenzi Tsukada & Nishiyama, 1980 (Fig. 36) laetitiae Schröder & Treadaway, 2000 (Fig. 47) TS (TP): HT ♂ (ET), PT 3♂2♀. monara Fruhstorfer, 1913 (Fig. 65) TL: 30 m NN, Calamian Group. synonym of tsukada Koçak (1996). Note: Though the original description does not mention designation was carried out by Yokochi (1999). cases —. The first author visited Senckenberg Museum in Universität, Berlin, Germany. junior secondary homonym of Bassarona teuta balabacana cusama Fruhstorfer, 1913 (Fig. 18) TS (TP): ST ♂ (BMNH). TL: near Baguio, N. Luzon. TL: Cavan Cavan, 5 m, Sibutu, Tawitawi Group. TL: N. Borneo. TS (TP): HT ♂ (SMFL), PT 11♂16♀. the number and sex, a syntype specimen in MNHN is a male. suluana Fruhstorfer, 1902 (Fig. 101) Frankfurt to research on the Treadaway collection on March Private collections. ET: Etsuzo Tsukada, Yamanashi, Tsukada, 1991. Here a new name balabacensis is replaced for TL: Mindanao, Camiguin de Mindanao. TS (TP): HT ♂ (MKN), PT 1♀ (MKN). TS (TP): HT ♂ (SMFL), PT 4♂. malissia Fruhstorfer, 1913 (Fig. 57) TS (TP): ST (MNHN). piratica G. Semper, [1887] (Fig. 81) TL: Sulu Archipelago. uposatha Fruhstorfer, 1913 (Fig. 109) 2019, and just then was informed that he had died several Japan; HH: Hiroto Hanafusa, Tottori, Japan; JF: Jumalon Dophla evelina balabacana Tsukada, 1991. TS (TP): ST ♂ (MNHN), ♀ (SMFL). ellora Fruhstorfer, 1898 (Fig. 28) TL: Basilan. panayana Schröder & Treadaway, 1980 (Fig. 75) TL: Camiguin de Mindanao. satrapes C. & R. Felder, 1861 (Fig. 91) TS (TP): HT ♀ (BMNH). TL: Banggi (Banguey), E. Malaysia. weeks ago at his home in England. In the middle 1990’s, every Family, Cebu, Philippines; MKN: Misao Kaneko, Tokyo, Euthalia (Dophla) evelina balabacensis nom. nov. Note: Figured specimen is a syntype female in SMFL. TL: Mindoro. hinunanganensis M. & T. Okano, 1986 (Fig. 37) leopardina Fruhstorfer, 1913 (Fig. 48) TS (TP): ST♂ (MNHN). nadenya Fruhstorfer, 1913 TL: Culasi, 800 ft, Antique, Panay. TS (TP): LT ♂ (SMFL). TL: Luzon. TS (TP): LT ♂ (ZMHU). time the first author visited to Senckenberg, he came bringing Japan; SK: Shun-ichi Kawamura, Tokyo, Japan; STA: Shinji pro Dophla evelina balabacana Tsukada, 1991, non TS (TP): ST ♂ (MNHN). TL: Hinunangan, Leyte. TL: Sulu Islands. Note: Though the original description does not mention TL: Luzon. TS (TP): HT ♂ (SMFL), PT 4♂4♀. Note: Description is based on six materials from Luzon, TS (TP): ST ♂ (BMNH). susoni Jumalon, 1975 (Fig. 102) Note: According to the original description, the type series his described type specimens, with his wife driving from Taga, Tokyo, Japan; TY: Takashi Yokochi, Aichi, Japan. Bassarona teuta balabacana Tsukada, 1991 dacasini Hanafusa, 1990 (Fig. 19) Note: Though the original description does not mention TS (TP): HT ♂ (FASU), PT 1♂. TS (TP): HT ♀ (BMNH). the number of types, a male syntype is preserved in MNHN. TS (TP): HT ♀ [untraced, not examined]. Mindoro, Camiguin de Mindanao, N. Mindanao. The lectotype Note: According to the original description, the type series TL: Camp 7, Minglanilla, Cebu. (male and female, but the number is unknown) are deposited in Specimens concerned. FW: Length of Forewing; HT: TL: Balabac. the number and sex, a syntype specimen in MNHN is a male. Note: Though the original description indicates that the panopus C. & R. Felder, 1861 (Fig. 76) designation was carried out for the Camiguin de Mindanao (number is unknown) are deposited in Semper and Felder TS (TP): HT ♂ (USC), PT 9♂9♀ (USC, JF, SMFL, TY). Staudinger collection (ZMHU). The lectotype designation was Holotype; LT: Lectotype; PLT: Paralectotype; PT: Paratype; boholensis (Lexias) M. & T. Okano, 1988 (Fig. 8) TS (TP): HT ♂ (HH), PT 1♀. hiwaga Nuyda & Kawamura, 1989 (Fig. 38) leucotaenia G. Semper, 1878 (Fig. 49) marinduque K. Okano, 1992 (Fig. 58) type female should be preserved in ZMHU, the specimen TL: Luzon. specimen by Schröder & Treadaway (1987). collections. carried out by Yokochi (1999). TL: Type Locality; TP: Type Preservation; TS: Type Series; TL: Bilar, Bohol. eson de Nicéville, 1894 (Fig. 29) TL: Camiguin, Babuyan. TL: Camotes. TL: Marinduque. corresponding with nadenya could not be found there. No TS (TP): ST ♂ (BMNH). tanagra Staudinger, 1889 (Fig. 103) ST: Syntype. TS (TP): HT ♂ (FASU), PT 6♂2♀. damalis Erichson, 1834 (Fig. 117) TL: Palawan. TS (TP): HT ♂ (SK), PT 7♂4♀ (SK, ET). TS (TP): LT ♂ (SMFL). TS (TP): HT ♀ (KMNH), PT 1♂ (KMNH). figure is presented in the original paper. Note: The original description does not mention the princesa Fruhstorfer, 1899 (Fig. 82) schoenigi Schröder & Treadaway, 1978 (Fig. 92) TL: Palawan. virginalis Fruhstorfer, 1913 TL: Luzon. TS (TP): LT ♂ (SMFL), PLT 1♀. Note: This taxon was described based on sixty-nine number of type series. TL: Palawan. TL: Amlan, 600 m, Negros. TS (TP): LT ♂ (ZMHU). TL: locality not given. boholensis (Euthalia) M. & T. Okano, 1990 (Fig. 9) TS (TP): ST ♂ [untraced, not examined]. Note: Description is based on a single pair, and all howarthi Jumalon, 1975 (Fig. 39) materials from Camotes, Bohol, Leyte, Panaon, E. Mindanao, medaga Fruhstorfer, 1913 (Fig. 59) negrosiana Schröder & Treadaway, 1987 (Fig. 66) TS (TP): HT ♀ (MNHN). TS (TP): HT ♀ (SMFL). Note: Description is based on seven males and eleven TS (TP): ST ♀ (BMNH) [not examined]. 3. LIST OF DESCRIBED SPECIES AND TL: Bilar, Bohol. Note: The type series should be preserved in ZMHU, but materials are preserved in SMFL. The lectotype designation TL: Amlan, Negros Oriental. and N. Mindanao. Yokochi (1998) mentioned the type series of TL: Camiguin de Mindanao. TL: Negros Or. [Oriental]. phelada G. Semper, [1887] (Fig. 77) females, and two males and three females are preserved in Note: This is a female form of panopus. The type locality SUBSPECIES NAME TS (TP): HT ♀ (FASU), PT 4♀. the specimens that corresponded with the types could not be was carried out by Yokochi (1998). TS (TP): HT ♂ (USC), PT 4♀ (USC, JF). leucotaenia was untraced so designated the lectotype for the TS (TP): ST ♂ (MNHN). TS (TP): HT ♂ (SMFL), PT 1♂1♀. TL: Luzon. proditrix Fruhstorfer, 1913 (Fig. 83) semperi Staudinger, 1889 (Fig. 93) ZMHU. The lectotype designation was carried out by Yokochi suspected is Mindanao, though the specific location is not found. The Fig. 117 is reproduced from the original description figure of the original description, but the type series specimen Note: Though the original description does not mention TS (TP): LT ♀ (SMFL). TL: Basilan. TL: Palawan. (1999). shown. The type series are deposited in the Rothschild collection The following list contains the taxonomic names bongaoensis Schröder & Treadaway, 1990 (Fig. 10) (pl. 50, fig. 4, ♂). eva C. & R. Felder, [1867] (Fig. 30) imperator Hewitson, 1863 (Fig. 40) was found after the third investigation in SMFL. We designate the number of types, a male syntype is preserved in MNHN. nirodha Fruhstorfer, 1913 (Fig. 67) Note: This taxon was described based on five materials TS (TP): ST ♂ (MNHN). TS (TP): LT ♂ (ZMHU). (BMNH, London), but we have not located them at present. concerned with Philippines. The names which are described TL: Bongao Peak, Bongao, Tawitawi Group. TL: Luzon. TL: Luzon. here again the lectotype for a male specimen shown in Fig. 49 TL: Basilan. from Luzon, Camiguin de Mindanao, and SE. Mindanao. Note: Description is based on males and females, and four tawitawia Treadaway & Nuyda, 1994 (Fig. 104) from the other areas are also included. TS (TP): HT ♀ (SMFL), PT 18♂20♀. darani Fruhstorfer, 1913 (Fig. 20) TS (TP): ST ♂ (BMNH). TS (TP): HT ♀ (BMNH). (labelled “Typus (red) / Camotes / Bohol / 113 / 211 / Coll. C. mindanaensis Schröder & Treadaway, 1978 (Fig. 60) TS (TP): ST ♂♀ (MNHN). Yokochi (1998) mentioned the type series of phelada was producta Butler, 1901 (Fig. 84) males and four females are preserved in ZMHU. The lectotype TL: Magsaggao, Tawitawi. visayana (Lexias) Schröder & Treadaway, 1987 (Fig. 110) TL: Sulu Archipelago. Note: According to the original description, eva is distrib- Semper”). TL: Surigao del Sur, Mindanao. untraced so designated the lectotype for the figure of the TL: Philippines [recte W. Malaysia]. designation was carried out by Yokochi (1999). TS (TP): HT ♂ (SMFL). TL: Catmon, S. Leyte. albusequus Nihira & Kawamura, 1986 (Fig. 2) borromeoi Schröder, 1977 (Fig. 11) TS (TP): HT ♀ (BMNH). uted in Assam, Luzon, and Celebes. The details of type series ingae (Lexias) Schröder & Treadaway, 1987 (Fig. 41) TS (TP): HT ♂ (SMFL). obatrata Yokochi, 1994 (Fig. 68) original description, but the target specimen was found after the TS (TP): ST ♀ (BMNH). TS (TP): HT ♂ (SMFL), PT 6♂4♀. TL: Lakit lakit, Sanga Sanga, Tawitawi. TL: Espana, 2000 ft, Sibuyan. Note: Though the original description does not mention are not mentioned. Staudinger (1886) is the first revisory of TL: Negros Or. [Oriental], 800 m. leytana Schröder & Treadaway, 1982 (Fig. 50) TL: Dumaran. third investigation in SMFL. We designate here again the senilis Hanafusa, 1992 (Fig. 94) tethys Tsukada, 1991 (Fig. 105) TS (TP): HT ♀ (SK). TS (TP): HT ♂ (SMFL), PT 1♀. the number of type series, one female would be the holotype. which Luzon is recognized as a locality of eva. TS (TP): HT ♂ (SMFL), PT 5♂5♀. TL: St. Bernard, Calpio, 280 m, S. Leyte. mindorana (Tanaecia) Fruhstorfer, 1904 (Fig. 61) TS (TP): HT ♀ (HH), PT 1♂1♀. lectotype for a female specimen shown in Fig. 77 (labelled rhamases Staudinger, 1889 (Fig. 85) TL: Dumaran. TL: N. Palawan. visayana (Cynitia) Schröder & Treadaway, 1981 (Fig. 111) TS (TP): HT ♂ (SMFL), PT 18♂10♀. TL: Mindoro. “Typus (red) / Euth. phelada typ. ♀ Semper / Marivcles (?) TL: Palawan. TS (TP): HT ♂ (HH), PT 1♂. TS (TP): HT ♂ (ET), PT 10♂12♀. TL: St. Bernard, Catmon, S. Leyte. 14 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 15

Fig. 2. albusequus. Holotype, ♀, FW 54 mm, SK. 3. amlana. Holotype, ♂, FW 57 mm, USC. 4. antiquea. Holotype, ♂, FW 40.5 Fig. 6. balabacana (Bassarona). Holotype, ♂, FW 34 mm, ET. 7. balabacana (Dophla), balabacensis. Holotype, ♂, FW 45 mm, mm, SMFL. 5. aquamarina. Lectotype, ♂, FW 37 mm, SMFL. ET. 8. boholensis (Lexias). Holotype, ♂, FW 34.5 mm, FASU. 9. boholensis (Euthalia). Holotype, ♀, FW 38 mm, FASU. 14 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 15

Fig. 2. albusequus. Holotype, ♀, FW 54 mm, SK. 3. amlana. Holotype, ♂, FW 57 mm, USC. 4. antiquea. Holotype, ♂, FW 40.5 Fig. 6. balabacana (Bassarona). Holotype, ♂, FW 34 mm, ET. 7. balabacana (Dophla), balabacensis. Holotype, ♂, FW 45 mm, mm, SMFL. 5. aquamarina. Lectotype, ♂, FW 37 mm, SMFL. ET. 8. boholensis (Lexias). Holotype, ♂, FW 34.5 mm, FASU. 9. boholensis (Euthalia). Holotype, ♀, FW 38 mm, FASU. 16 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 17

Fig. 10. bongaoensis. Holotype, ♀, FW 37 mm, SMFL. 11. borromeoi. Holotype, ♂, FW 35.5 mm, SMFL. 12. cabigoni. Holotype, Fig. 14. candida. Holotype, ♂, FW 36 mm, SMFL. 15. cavarna. Lectotype, ♀, FW 49.5 mm, ZMHU. 16. chloe. Holotype, ♂, FW ♂, FW 36.5 mm, SMFL. 13. calliphorus. Syntype, ♂, FW 38.5 mm, BMNH, © The Natural History Museum, London. 46 mm, SMFL. 17. circe. Holotype, ♂, FW 40 mm, SMFL. 16 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 17

Fig. 10. bongaoensis. Holotype, ♀, FW 37 mm, SMFL. 11. borromeoi. Holotype, ♂, FW 35.5 mm, SMFL. 12. cabigoni. Holotype, Fig. 14. candida. Holotype, ♂, FW 36 mm, SMFL. 15. cavarna. Lectotype, ♀, FW 49.5 mm, ZMHU. 16. chloe. Holotype, ♂, FW ♂, FW 36.5 mm, SMFL. 13. calliphorus. Syntype, ♂, FW 38.5 mm, BMNH, © The Natural History Museum, London. 46 mm, SMFL. 17. circe. Holotype, ♂, FW 40 mm, SMFL. 18 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 19

Fig. 18. cusama. Syntype, ♀, FW 43 mm, SMFL. 19. dacasini. Holotype, ♂, FW 30.5 mm, HH. 20. darani. Holotype, ♀, FW 36.5 Fig. 22. dinagatensis. Holotype, ♂, FW 39 mm, ET. 23. dinorah. Syntype, ♀, FW 41 mm, MNHN. 24. dodong. Holotype, ♂, FW mm, BMNH. 21. dhayma. Holotype, ♀, FW 38 mm, BMNH. Figs. 20, 21, © The Natural History Museum, London. 39.5 mm, SMFL. 25. dohertyi. Syntype, ♂, FW 30.5 mm, BMNH. © The Natural History Museum, London. 18 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 19

Fig. 18. cusama. Syntype, ♀, FW 43 mm, SMFL. 19. dacasini. Holotype, ♂, FW 30.5 mm, HH. 20. darani. Holotype, ♀, FW 36.5 Fig. 22. dinagatensis. Holotype, ♂, FW 39 mm, ET. 23. dinorah. Syntype, ♀, FW 41 mm, MNHN. 24. dodong. Holotype, ♂, FW mm, BMNH. 21. dhayma. Holotype, ♀, FW 38 mm, BMNH. Figs. 20, 21, © The Natural History Museum, London. 39.5 mm, SMFL. 25. dohertyi. Syntype, ♂, FW 30.5 mm, BMNH. © The Natural History Museum, London. 20 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 21

Fig. 26. dumaranensis. Holotype, ♂, FW 35 mm, HH. 27. dunya. Syntype, ♂, FW 43 mm, BMNH, © The Natural History Fig. 30. eva. Syntype, ♂, FW 43.5 mm, BMNH, © The Natural History Museum, London. 31. exul. Holotype, ♂, FW 37 mm, ET. Museum, London. 28. ellora. Syntype, ♂, FW 42 mm, MNHN. 29. eson. Lectotype, ♂, FW 36.5 mm, SMFL. 32. felicitacionae. Holotype, ♂, FW 33.5 mm, JF. 33. gahiti. Holotype, ♀, FW 45 mm, FASU. 20 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 21

Fig. 26. dumaranensis. Holotype, ♂, FW 35 mm, HH. 27. dunya. Syntype, ♂, FW 43 mm, BMNH, © The Natural History Fig. 30. eva. Syntype, ♂, FW 43.5 mm, BMNH, © The Natural History Museum, London. 31. exul. Holotype, ♂, FW 37 mm, ET. Museum, London. 28. ellora. Syntype, ♂, FW 42 mm, MNHN. 29. eson. Lectotype, ♂, FW 36.5 mm, SMFL. 32. felicitacionae. Holotype, ♂, FW 33.5 mm, JF. 33. gahiti. Holotype, ♀, FW 45 mm, FASU. 22 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 23

Fig. 34. galoa. Syntype, ♂, FW 36.5 mm, BMNH, © The Natural History Museum, London. 35. goertzi. Holotype, ♀, FW 42 mm, Fig. 38. hiwaga. Paratype, ♀, FW 57 mm, ET. 39. howarthi. Holotype, ♂, FW 42.5 mm, USC. 40. imperator. Holotype, ♀, FW 56 ET. 36. hikarugenzi. Holotype, ♂, FW 46.5 mm, MKN. 37. hinunanganensis. Holotype, ♂, FW 39.5 mm, FASU. mm, BMNH, © The Natural History Museum, London. 41. ingae (Lexias). Holotype, ♂, FW 42.5 mm, SMFL. 22 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 23

Fig. 34. galoa. Syntype, ♂, FW 36.5 mm, BMNH, © The Natural History Museum, London. 35. goertzi. Holotype, ♀, FW 42 mm, Fig. 38. hiwaga. Paratype, ♀, FW 57 mm, ET. 39. howarthi. Holotype, ♂, FW 42.5 mm, USC. 40. imperator. Holotype, ♀, FW 56 ET. 36. hikarugenzi. Holotype, ♂, FW 46.5 mm, MKN. 37. hinunanganensis. Holotype, ♂, FW 39.5 mm, FASU. mm, BMNH, © The Natural History Museum, London. 41. ingae (Lexias). Holotype, ♂, FW 42.5 mm, SMFL. 24 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 25

Fig. 42. ingae (Euthalia). Holotype, ♀, FW 39 mm, SMFL. 43. irauana. Holotype, ♀, FW 36 mm, JF. 44. joloana. Lectotype, ♂, Fig. 46. kulaya. Holotype, ♂, FW 34.5 mm, SMFL. 47. laetitiae. Holotype, ♂, FW 30 mm, SMFL. 48. leopardina. Holotype, ♀, FW 31 mm, ZMHU. 45. kayumanggia. Holotype, ♂, FW 27.5 mm, SMFL. FW 40 mm, BMNH, © The Natural History Museum, London. 49. leucotaenia. Lectotype, ♂, FW 37 mm, SMFL. 24 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 25

Fig. 42. ingae (Euthalia). Holotype, ♀, FW 39 mm, SMFL. 43. irauana. Holotype, ♀, FW 36 mm, JF. 44. joloana. Lectotype, ♂, Fig. 46. kulaya. Holotype, ♂, FW 34.5 mm, SMFL. 47. laetitiae. Holotype, ♂, FW 30 mm, SMFL. 48. leopardina. Holotype, ♀, FW 31 mm, ZMHU. 45. kayumanggia. Holotype, ♂, FW 27.5 mm, SMFL. FW 40 mm, BMNH, © The Natural History Museum, London. 49. leucotaenia. Lectotype, ♂, FW 37 mm, SMFL. 26 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 27

Fig. 50. leytana. Holotype, ♂, FW 31 mm, SMFL. 51. leytensis. Holotype, ♀, FW 40 mm, USC. 52. liaoi. Holotype, ♂, FW 35.5 Fig. 54. lupina. Holotype, ♀, FW 46 mm, BMNH, © The Natural History Museum, London. 55. lusiada. Syntype, ♂, FW 31.5 mm, SMFL. 53. ludonia. Lectotype, ♂, FW 29.5 mm, ZMHU. mm, SMFL. 56. macer. Holotype, ♂, FW 36.5 mm, ET. 57. malissia. Syntype, ♂, FW 35 mm, MNHN. 26 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 27

Fig. 50. leytana. Holotype, ♂, FW 31 mm, SMFL. 51. leytensis. Holotype, ♀, FW 40 mm, USC. 52. liaoi. Holotype, ♂, FW 35.5 Fig. 54. lupina. Holotype, ♀, FW 46 mm, BMNH, © The Natural History Museum, London. 55. lusiada. Syntype, ♂, FW 31.5 mm, SMFL. 53. ludonia. Lectotype, ♂, FW 29.5 mm, ZMHU. mm, SMFL. 56. macer. Holotype, ♂, FW 36.5 mm, ET. 57. malissia. Syntype, ♂, FW 35 mm, MNHN. 28 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 29

Fig. 58. marinduque. Holotype, ♀, FW 57 mm, KMNH. 59. medaga. Syntype, ♂, FW 35.5 mm, MNHN. 60. mindanaensis. Holotype, Fig. 62. mindorana (Euthalia). Syntype, ♂, FW 33.5 mm, MNHN. 63. mindorensis. Holotype, ♂, FW 30.5 mm, SMFL. 64. miscus. ♂, FW 35.5 mm, SMFL. 61. mindorana (Tanaecia). Syntype, ♀, FW 45 mm, BMNH, © The Natural History Museum, London. Syntype, ♀, FW 42.5 mm, MNHN. 65. monara. Syntype, ♂, FW 41 mm, MNHN. 28 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 29

Fig. 58. marinduque. Holotype, ♀, FW 57 mm, KMNH. 59. medaga. Syntype, ♂, FW 35.5 mm, MNHN. 60. mindanaensis. Holotype, Fig. 62. mindorana (Euthalia). Syntype, ♂, FW 33.5 mm, MNHN. 63. mindorensis. Holotype, ♂, FW 30.5 mm, SMFL. 64. miscus. ♂, FW 35.5 mm, SMFL. 61. mindorana (Tanaecia). Syntype, ♀, FW 45 mm, BMNH, © The Natural History Museum, London. Syntype, ♀, FW 42.5 mm, MNHN. 65. monara. Syntype, ♂, FW 41 mm, MNHN. 30 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 31

Fig. 70. ornata. Holotype, ♂, FW 49 mm, SMFL. 71. palawana (Lexias). Lectotype, ♂, FW 35.5 mm, BMNH, © The Natural Fig. 66. negrosiana. Holotype, ♂, FW 39.5 mm, SMFL. 67. nirodha. Syntype, ♂, FW 31.5 mm, MNHN. 68. obatrata. Holotype, History Museum, London. 72. palawana (Tanaecia). Lectotype, ♂, FW 33.5 mm, ZMHU. 73. palawana (Euthalia). Lectotype, ♀, FW 36.5 mm, HH. 69. ormocana. Holotype, ♂, FW 50 mm, USC. ♂, FW 32.5 mm, ZMHU. 30 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 31

Fig. 70. ornata. Holotype, ♂, FW 49 mm, SMFL. 71. palawana (Lexias). Lectotype, ♂, FW 35.5 mm, BMNH, © The Natural Fig. 66. negrosiana. Holotype, ♂, FW 39.5 mm, SMFL. 67. nirodha. Syntype, ♂, FW 31.5 mm, MNHN. 68. obatrata. Holotype, History Museum, London. 72. palawana (Tanaecia). Lectotype, ♂, FW 33.5 mm, ZMHU. 73. palawana (Euthalia). Lectotype, ♀, FW 36.5 mm, HH. 69. ormocana. Holotype, ♂, FW 50 mm, USC. ♂, FW 32.5 mm, ZMHU. 32 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 33

Fig. 74. pallida. Holotype, ♂, FW 35 mm, SMFL. 75. panayana. Holotype, ♂, FW 43 mm, SMFL. 76. panopus. Syntype, ♂, FW Fig. 78. philippensis. Lectotype, ♂, FW 31 mm, MNHN. 79. phlegethon. Lectotype, ♀, FW 39.5 mm, SMFL. 80. pinamalayana, 37 mm, BMNH, © The Natural History Museum, London. 77. phelada. Lectotype, ♀, FW 40 mm, SMFL. tsukada. Holotype, ♂, FW 33 mm, ET. 81. piratica. Lectotype, ♂, FW 35 mm, SMFL. 32 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 33

Fig. 74. pallida. Holotype, ♂, FW 35 mm, SMFL. 75. panayana. Holotype, ♂, FW 43 mm, SMFL. 76. panopus. Syntype, ♂, FW Fig. 78. philippensis. Lectotype, ♂, FW 31 mm, MNHN. 79. phlegethon. Lectotype, ♀, FW 39.5 mm, SMFL. 80. pinamalayana, 37 mm, BMNH, © The Natural History Museum, London. 77. phelada. Lectotype, ♀, FW 40 mm, SMFL. tsukada. Holotype, ♂, FW 33 mm, ET. 81. piratica. Lectotype, ♂, FW 35 mm, SMFL. 34 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 35

Fig. 82. princesa. Holotype, ♀, FW 36.5 mm, MNHN. 83. proditrix. Syntype, ♂, FW 44.5 mm, MNHN. 84. producta. Syntype, ♀, Fig. 86. rodriguezi. Holotype, ♂, FW 30.5 mm, SMFL. 87. romeo. Holotype, ♀, FW 40.5 mm, SMFL. 88. rudraca. Lectotype, ♂, FW 37 mm, BMNH, © The Natural History Museum, London. 85. rhamases. Lectotype, ♂, FW 31 mm, ZMHU. FW 35.5 mm, ZMHU. 89. samarensis. Holotype, ♂, FW 46 mm, ET. 34 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 35

Fig. 82. princesa. Holotype, ♀, FW 36.5 mm, MNHN. 83. proditrix. Syntype, ♂, FW 44.5 mm, MNHN. 84. producta. Syntype, ♀, Fig. 86. rodriguezi. Holotype, ♂, FW 30.5 mm, SMFL. 87. romeo. Holotype, ♀, FW 40.5 mm, SMFL. 88. rudraca. Lectotype, ♂, FW 37 mm, BMNH, © The Natural History Museum, London. 85. rhamases. Lectotype, ♂, FW 31 mm, ZMHU. FW 35.5 mm, ZMHU. 89. samarensis. Holotype, ♂, FW 46 mm, ET. 36 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 37

Fig. 90. sarmana. Syntype, ♂, FW 34 mm, MNHN. 91. satrapes. Syntype, ♀, FW 55 mm, BMNH, © The Natural History Fig. 94. senilis. Holotype, ♂, FW 33.5 mm, HH. 95. sibutana. Holotype, ♂, FW 30 mm, SMFL. 96. sibuyana. Holotype, ♂, FW 34 Museum, London. 92. schoenigi. Holotype, ♀, FW 44 mm, SMFL. 93. semperi. Lectotype, ♂, FW 31.5 mm, ZMHU. mm, SMFL. 97. smaragdifera. Syntype, ♂, FW 41.5 mm, MNHN. 36 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 37

Fig. 90. sarmana. Syntype, ♂, FW 34 mm, MNHN. 91. satrapes. Syntype, ♀, FW 55 mm, BMNH, © The Natural History Fig. 94. senilis. Holotype, ♂, FW 33.5 mm, HH. 95. sibutana. Holotype, ♂, FW 30 mm, SMFL. 96. sibuyana. Holotype, ♂, FW 34 Museum, London. 92. schoenigi. Holotype, ♀, FW 44 mm, SMFL. 93. semperi. Lectotype, ♂, FW 31.5 mm, ZMHU. mm, SMFL. 97. smaragdifera. Syntype, ♂, FW 41.5 mm, MNHN. 38 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 39

Fig. 98. soloni. Holotype, ♂, FW 34 mm, FASU. 99. soregina. Syntype, ♀, FW 38 mm, BMNH. 100. subpiratica. Holotype, ♂, Fig. 102. susoni. Holotype, ♂, FW 33.5 mm, USC. 103. tanagra. Lectotype, ♂, FW 27 mm, ZMHU. 104. tawitawia. Holotype, ♂, FW 34.5 mm, SMFL. 101. suluana. Holotype, ♀, FW 34 mm, BMNH. Figs. 99, 101, © The Natural History Museum, London. FW 31.5 mm, SMFL. 105. tethys. Holotype, ♂, FW 42 mm, ET. 38 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 39

Fig. 98. soloni. Holotype, ♂, FW 34 mm, FASU. 99. soregina. Syntype, ♀, FW 38 mm, BMNH. 100. subpiratica. Holotype, ♂, Fig. 102. susoni. Holotype, ♂, FW 33.5 mm, USC. 103. tanagra. Lectotype, ♂, FW 27 mm, ZMHU. 104. tawitawia. Holotype, ♂, FW 34.5 mm, SMFL. 101. suluana. Holotype, ♀, FW 34 mm, BMNH. Figs. 99, 101, © The Natural History Museum, London. FW 31.5 mm, SMFL. 105. tethys. Holotype, ♂, FW 42 mm, ET. 40 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 41

Fig. 110. visayana (Lexias). Holotype, ♂, FW 42 mm, SMFL. 111. visayana (Cynitia). Holotype, ♂, FW 35.5 mm, SMFL. 112. Fig. 106. trapesa. Lectotype, ♂, FW 50.5 mm, SMFL. 107. treadawayi. Holotype, ♂, FW 36.5 mm, SMFL. 108. tyawena. vistrica. Syntype, ♂, FW 34.5 mm, BMNH, © The Natural History Museum, London. 113. volupia. Holotype, ♂, FW 40.5 Lectotype, ♀, FW 60 mm, ZMHU. 109. uposatha. Lectotype, ♂, FW 30 mm, ZMHU. mm, ET. 40 Takashi Yokochi and Fidel Bendanillo Adoliadini of the Philippine Is. 41

Fig. 110. visayana (Lexias). Holotype, ♂, FW 42 mm, SMFL. 111. visayana (Cynitia). Holotype, ♂, FW 35.5 mm, SMFL. 112. Fig. 106. trapesa. Lectotype, ♂, FW 50.5 mm, SMFL. 107. treadawayi. Holotype, ♂, FW 36.5 mm, SMFL. 108. tyawena. vistrica. Syntype, ♂, FW 34.5 mm, BMNH, © The Natural History Museum, London. 113. volupia. Holotype, ♂, FW 40.5 Lectotype, ♀, FW 60 mm, ZMHU. 109. uposatha. Lectotype, ♂, FW 30 mm, ZMHU. mm, ET. 42 Takashi Yokochi and Fidel Bendanillo Bull. Kitakyushu Mus. Nat. Hist. Hum. Hist., Ser. A, 18: 43–48, March 31, 2020 43

塗り絵を利用した子供向けアンケート手法の有効性 エピソード記述やワークショップをとおして視覚化し関係 このような場合，保護者といえども子供の真意がくみ取れな 少数の簡単な設問（表 1）を提示したことが考えられる． 紙面は A4 などの用紙に比べて限られる（図 1D）．しかし 要約 による「放課後子ども教室」評価の支援－． 河村学園大学 者で共有する方法も検討されている（坂倉， 2015, 2017）． い可能性があることが指摘されている（吉田， 2012）． 放課後や週末に子供達が安全で安心した時間を過ごすこと 子供を主な対象とするためには設問の単純性やわかりやす 研究紀要， 19 (1): 85–104． 一方，日本では，エバリュエーターを専門職として配置し そこで筆者らは，多数の子供の生の声を，少ない労力で ができるよう公立小学校が実施している「放課後子ども教 さと，一定以上のフォントサイズが求められる．これにより， 特別展内における来館者参加型の「塗り絵」を利用した 倉田公裕・矢島國雄． 1993． 博物館展示評価の基礎的研究． 蓑島悠介1・真鍋 徹1・下村通誉2・上田恭一郎1 ている博物館がきわめて少ないため，体系的な展示評価や専 現出できる方法を検討した．この方法の立案に際し，北九州 室」の現状を，子供自身による評価を通して把握しようと 文字数や設問数に制限が生まれる．また，紙面が限られる 自由回答のアンケート手法について，回答率と回答者の面 明治大学人文科学研究所紀要， 33: 269–290． 門的な知識と経験が必要な来館者調査を実施することは難し 市立自然史・歴史博物館の特別展内で子供向けに行ってきた した研究でも，極力シンプルで複数項目からの選択式回答 ことで，インフォームドコンセントの文言を組み込むのが から有用性を検討した．試行は 2017 年の夏に北九州市立自 蓑島悠介・下村通誉・真鍋 徹・上田恭一郎． 2018． 実物資 1 805-0071 福岡県北九州市八幡東区東田2-4-1 北九州市立自然史・歴史博物館 い（倉田・矢島， 1993; 里見， 2014）．そのため，博物館を訪 塗り絵に着目した．この塗り絵は，はがきサイズの紙に学芸 のアンケートを用いたことが，子供の真意の現出につながっ 難しい．また，アンケート回答面は裏面であり，100 kg 前 然史・歴史博物館で行われた自然史の特別展において，10 料に内包された知覚効果を引き出す展示手法の開発 ～ E-mail: [email protected], [email protected], [email protected] れるさまざまなバックグラウンドを持つ多様な来館者の声を 員が描いた線画を印刷したものである．台紙を着色するため た要因の一つであると考察されている（北原ら， 2008）． こ 後の厚めの上質紙を使えば塗り絵を貼った後に表面から回 日間程度で行った．アンケート付の塗り絵は合計 1,064 枚 昆虫の多様性の知覚化に向けて． 全科協ニュース， 48 2 649-2211 和歌山県白浜町459 京都大学フィールド科学教育研究センター 瀬戸臨海実験所 拾い集め，展示評価に活かすため，国内では展示に対する対 の作業机を配置した「塗り絵コーナー」を設置し，さらに着 のように，子供の声を汲み取るためのアンケートの設計に 答が見えることはないものの，貼ってもらうという回答方 回収され，有効回答票が 340 枚（回収したアンケートに対 (4): 4–5． 面での聞き取り（インタビュー）あるいは用紙を設置して回 色した塗り絵を貼ることのできる壁を設置している（図 は，適切で簡単な設問を設定することは，非常に重要であ 法は箱に入れるアンケート回収とは同等ではない．例えば， する回答率として 32.0%）だった．各設問の回答率は 1 問 村田麻里子． 2003． 来館者研究の系譜とその課題―日本にお E-mail: [email protected] 答してもらうといったアンケートによっている場合が多い（例 1A–C）．塗り絵は，壁に貼る来館者もいる一方，思い出とし ると思われる．一方，複雑なものなど設問の内容によって 貼り付けてあった塗り絵は容易に剥がすことができるため， （82.4%）を除き，すべて 90% 以上の回答率となり，アンケー ける博物館コミュニケーションの展開のための一考察 （2019年6月1日受領，2019年10月19日受理） えば倉田・矢島， 1993; 伊丹市昆虫館， 2003; 鵜沢， 2016; 蓑島 て持ち帰る来館者もいる．また，時には大人も壁に塗り絵を は無回答が増えることが想定される．このような設問を設 回答面に記された情報を他人が見ることができる可能性も トとして十分に機能することが示唆された．回答率には配 ―． 日本ミュージアム・マネージメント学会研究紀要， ら， 2018）．しかし，これらのアンケートは大人を対象とした 貼っている．筆者らは，この塗り絵の裏に簡単な設問を書く ける必要があるアンケートの場合は，設問数を減らすなど 存在すると思われる．これらのことから，個人情報の収集 布方法や設問が関わっていると考えられる．有効回答中の 7: 95–104． ものがほとんどであり，子供の率直な生の声を広く集めるた ことで，来館者側も調査者側も手間を必要とせず，かつ自然 の方策が必要かもしれない． やセンシティブな質問事項にならないように，設問の設計 男女比は，男性が 55.3%，女性が 44.7% となった．回答者 Punch, K. F. 1998. Introduction to Social Research: Quantitative Effectiveness of questionnaire survey for children using picture coloring sheet めにはさまざまな工夫が求められる．例えば，小学校の低学 体でアンケートに回答してもらえるのではないかと着想し 今回の調査では，塗り絵のスペースおよび貼り付け用の には十分な考慮が必要であろう． の年齢平均は 8.8 歳で，中央値と最頻値ともに 7 歳となった． and Qualitative Approaches. Sage Publications Ltd., 年生と高学年生では発達段階がかなり異なるため，幅広い発 た．これは，展示会の感想や展示内容に対する質問を記した スペースという制限要因が想定され，塗り絵を持ち帰る来 これらの結果は本アンケート手法が博物館特別展において Thousand Oaks. [ 川合隆男監訳． 2005. 社会調査入門－ Yûsuke N. Minoshima1, Tohru Manabe1, Michitaka Shimomura2 and Kyoichiro Ueda1 達段階にある子供にも理解できる質問を設計する必要がある． カードや人気投票を来館者が自ら壁に貼って共有するという 館者が予想されることから，回収率（配布枚数に対する貼 7 ～ 8 歳前後の子供を対象に有用に機能する可能性を示唆 量的調査と質的調査の活用． 慶應義塾大学出版会 ] さらに，低学年生を対象とする場合，各児童に質問項目を読 参加型手法（例えば坂本・野本， 2016）を，アンケート用に 方法 にルビを振った．記述内容は表 1 のとおりで，名前などの り付け率）を求めず，本アンケート手法で十分な数の回答 まとめ している．一方でアンケート用紙のサイズがはがきサイズ 坂倉真衣． 2015． 来館者の「博物館体験」をどのように理解 み上げたり説明を加えたりしながら，回答を代筆することが 改変したものである．そこで，夏の特別展「大昆虫博」にお 個人を識別できる情報を記す欄は設けなかった． が見込まれるか、どのような回答者が期待されるのか，の である制約による面積の小ささや，アンケートが裏面で貼 し，関わることができるか－「学習の支援文脈モデル 1 Kitakyushu Museum of Natural History and Human History, 2-4-1 Higashida, Yahatahigashi-ku, Kitakyushu-shi, 必要な場合もある．このため，当方式でのアンケートの場合， いて，本アンケート手法により，どの程度の回答があるのか， アンケート調査は，北九州市立自然史・歴史博物館（福 塗り絵を壁に貼ることで回収する手法を採用したため， 分析のみを目的とした．しかし，回収率はアンケートの誤 本手法は，7 ～ 8 歳前後の子供を対象としたアンケート り出した際には見えないとはいえ，貼り出すことが前提で （Falk & Dierking, 2000）」の再解釈と展示物との「出会 Fukuoka, 805-0071 Japan. E-mail: [email protected], [email protected], [email protected] 岡県北九州市八幡東区東田）で開催された夏の特別展「大 配布数に対する回収数（回収率）は来館者数と混雑具合， 2Seto Marine Biological Laboratory, Field Science Education and Research Center, Kyoto University, 459 Shirahama-chô, 対応できる子供の数が限られてしまうといった課題も存在す またどのような回答者が見込まれるのかを確かめるため，塗 差を把握する上でも重要な指標となるため，今後の調査で 調査として十分に機能することが示唆された．本手法には あるため，設問に配慮が必要なことが本アンケート手法の い」という捉え方から－． 博物館学雑誌， 41 (1): 1–19． 昆虫博」（ 年 月 日（土）～ 年 月 日（日） すなわち塗り絵コーナーおよび休憩用に設けた机と椅子の Wakayama, 649-2211 Japan. E-mail: [email protected] る（北原ら， 2008）．また，保護者や教育者を通して間接的に り絵を利用した子供向けアンケートを試行した．本稿では本 2017 7 15 2017 9 3 は，回収率を把握し，より高めることのできる手法も検討 これまでに用いられてきたアンケートにはない利点がある 制限事項となる． 坂倉真衣． 2017． 来館者の博物館体験を視覚化，共有化する 子供の意見を収集する方法もある（伊丹市昆虫館， 2003 など）． アンケート試行の結果を報告する． の計 51 日間，特別展総入場者数は 122,294 名）の会期中， 数，加えて貼り出す壁の面積が制限要因となる．そのため， したい． が，トレードオフとなる欠点も存在する．すなわち，（1） 試み：ワークショップ「ミュージアムでの思い出を絵本 10 日間程度で以下のとおり行った． 本研究では第一段階として，配布数の計数および日時を正 回答者の性別については，男性が女性よりも 10% 多かっ 設問の単純さ，（2）設問数の少なさ，（3）倫理上の制限が にしよう」の事例から． 博物館学雑誌， 43 (1): 19–26． ABSTRACT − We evaluated the effectiveness of a survey questionnaire for children using a “picture アンケートは，特別展の期間中を通じて行っている参加 確に記載しての回収は行わずに，回収したアンケートに対 た．過去に行われた他の自然史系博物館におけるアンケー 大きいというデメリットと引き換えに，（1）面接アンケー 引用文献 坂本 昇． 2013． 博物館の調査収集活動を通じて地域の自然 coloring sheet” based on the response rate and the answers. The questionnaire was printed on the back of the 体験イベントの一つとして配布した塗り絵台紙の裏面を用 する回答率と属性のばらつきの検討のみに絞った．アンケー 回収された塗り絵は，回答があるものとないもの（白紙） その他）は回答なし，両方に丸がついているものが含まれる． トでは，博物館や展示内容により男女比が異なるものの， ト（インタビュー）のように調査者にとってプラス意見に を伝える展示 企画展「おきなわ～ちょうちょのふるさと picture coloring sheets (ca. 148 mm × 100 mm), which were distributed ca. 10 days at the special exhibition of いた．塗り絵の台紙は葉書サイズ（148 mm × 100 mm）で， トに用いた塗り絵の図柄（図 1E）はカブトムシ・チョウ・ に分けた．回答があるものは，無効回答（イタズラなど） 年代は，下は 1 歳，上は 61 歳で，年齢層に幅はあったも おおよそ男女ともに 4 割から 6 割程度の範囲に収まってい バイアスする（里見， 2014）可能性が低い，（2）設置型ア 萩原 剛・太田裕之・藤井 聡． 2006． アンケート調査回収 ～」開催報告． 伊丹市昆虫館研究報告， 1: 33–43． insects at the Kitakyushu Museum of Natural History and Human History in the summer of 2017. The completed 厚めの白上質紙（90 kg）を用いて，壁に貼り出した際に裏 サソリの 3 種である．これら図柄の生物は全て特別展に展 を除き，1 項目でも回答があれば有効と判断し，集計した． のの，平均年齢は 8.8 歳（標準偏差 7.5）で，中央値は 7 歳 る（橋本ら， 2007; 八木， 2010; 坂本， 2013; 鵜沢， 2016）． 今 ンケートのように自らアンケートを書きに行く手間がかか 率に関する：MM 参加率の効果的向上方策についてい 坂本 昇・野本康太． 2016． 企画展「ごきぶり」開催報告． 伊 coloring sheets were put on a wall at the special exhibition by children and/or parents; they were later collected 面の回答が衆目にふれることはないように配慮した．この 示したものであり，図柄の作成には，展示標本のほか，提 項目ごとの回答は Microsoft Excel ワークシートに入力し， である．年齢構成は 6～10 歳がおよそ 6 割を占め，次に 1 回みられた 10% の差が塗り絵参加者の男女比を反映してい らない，（3）大きな読みやすい文字を提示することができる， の基礎的検討． 土木計画学研究・論文集， 23 (1): 丹市昆虫館研究報告， 4: 23–33. and analyzed. The total valid responses in ca. 10 days, which is the number of the sheets with at least one answer 塗り絵は，予め学術的に正確な生物の形態を下絵として描 供者の許可を得た写真を利用した．アンケート付きの塗り 本ソフトウェア上で解析した．回収したアンケートに対す ～5 歳が多かった（表 3）．特に回答者が多い年代（1 歳か るのか，昆虫展入場者の比を反映しているのかは不明であ （4）アンケート回答用の場所を新たに用意する必要がない， 117–123． 里見 親幸． 2014. 博物館展示の理論と実践． v+229 p. 同成 excluding prank responses, were 340; response rates of the respective questions within the valid responses were いた上記台紙に，自らの好みの色を塗る作業を通し，生き 絵は 8 月に準備し，お盆休みのピークが明け来館者数が落 る回答率はアンケートが有効回答か否かの他に，それぞれ ら 15 歳）に注目すると，3 歳から回答者が増えだし，7 歳 るが，他館の結果と比べて大きな差はなかったことは興味 そして（5）塗り絵を塗らなければ壁に貼れない（貼れない 橋本佳延・坂田 宏 志・高 野 温子・三橋 弘宗・布施 静香． 2007． 社， 千代田区． 97.9% to 82.4%. The ratio of males and females within the valid responses of the question is 55.3% and 44.7% 物の体の作りについて知覚してもらうことを目的としたも ち着いた頃に，アンケートがない通常配布の塗り絵に補充 の設問に対しても計算した．割合については本文・図表と と 8 歳を頂点とするピークを経て，9 歳を境に回答者が減 深い． 訳ではないが，目立つ）点により一人による大量の回答を 過去と現在の里山における身近な動植物の利用～人と 鵜沢美穂子． 2016． ミュージアムパーク茨城県自然博物館第 respectively. Although more males than females answered the questions, this ratio is not very different from that のである．塗り絵は持ち帰ることも，特別展内で着色する する形で，会場に設置した塗り絵用の机に配置し，来館者 もに百分率で示し，小数点第二位を四捨五入した． 少するという傾向が見られた． 回答者は，10 歳以下が圧倒的に多く（表 3），さらに 7 歳 防ぐことができるというメリットを内在している．このよ 自然の博物館利用者へのアンケート調査から～． 人と自 57 回企画展「こけティッシュ苔ワールド！―ミクロの of questionnaires from other museums. The average age of the answerers was 8.8 years old (standard deviation: こともできる．今回のアンケートでは，塗り絵コーナー付 が自由に利用できるようにした．用意したアンケート付き なお，筆跡から保護者による代筆であろうと思われるも と 8 歳を頂点として 10 歳を超えると回答数が大きく減った うに，踏み込んだ精緻な情報ではなく，シンプルな子供の 然， 18: 151–162． 森に魅せられて―」のアンケートからみる来館者の意識 7.5) and the median value was 7. These results suggest that this questionnaire survey will work for children, 近に設置した壁に来館者が貼った塗り絵を回収する方法を の塗り絵がなくなり次第、アンケートのない塗り絵を補充 のは子供の意見を聞いて保護者が代筆しているものと判断 （表 4）ことから，小学校中・高学年生がアンケートの主な 生の声をランダムにかつ大量に収集するには有益な方法で Hein，G. E. 1998. Learning in the Museum. Routledge, 変化． 茨城県自然博物館研究報告， 19: 105–116． mainly 7 to 8 years old. This questionnaire survey, however, has some limitations. Due to the small size of the とった．塗り絵コーナーは記念撮影コーナー以外のおおよ した．そのため、アンケート付き塗り絵の配りはじめと配 し，アンケートに記された年齢の人の回答として扱った． 考察 回答者となっていたことが分かる．なお，先述したとおり， あると考えられる． Oxfordshire. [ 鷹野光行監訳 2010「博物館で学ぶ」同成 八木 剛． 2010. 昆虫リテラシー向上のための基礎資料－展 coloring sheet, we had to limit the number of questions included and it was difficult to incorporate sentences そすべての展示物を見たところに設置した（図 1A）． りおわりの時には，アンケートのある塗り絵とない塗り絵 筆跡から特に年少の子供で保護者による代筆だろうと思わ 社 ] 覧会「神戸元町・夏の昆虫館」における展示標本への人 concerning informed consent into the limited space. Another limitation is that the sheets were displayed on the アンケートは簡単な文字の読み書きができる子供が一人 が混在していたことになる．貼られた塗り絵は会場監視員 アンケートの回答数を増やすための手段として，アンケー れるものが散見された．このように，保護者と一緒に答え 伊丹市昆虫館． 2003． 平成 14 年度文部科学省科学系博物館 気投票から見た，男女別，年齢層別の昆虫の好み－． きべ wall. We concluded that we should carefully select questions in order to maximize the effectiveness of the でも回答できるよう，少数の簡単な設問とし，フォントサ （交流員）が適宜回収した．ただし塗り絵回収時に一部を壁 結果 ト用紙の郵送や調査員の雇用による対面調査量の増加など ることによるバイアスも今後の課題かもしれない． 謝辞 教育機能活用推進事業【ちょうちょのふしぎ いっぱ りはむし， 32 (2): 26–37． questionnaire survey for children as a research method. イズと解答欄を大きめに設定（図 1D）し，ほぼ全ての漢字 に残したため，正確な回答日時は不明であった． が行われている．さらに，事前報酬の提供や訪問留置など 他方，今回の結果では 16 ～ 20 歳の回答者が 3 名，26 ～ い】実施報告書． 54 p． 伊丹市昆虫館, 伊丹市． 吉田友英． 2012. 体平衡機能の発達と学童期のめまい． （1）アンケート回答数 アンケートの回収率を高める方策なども検討されている（萩 30 歳は 4 名，36 ～ 40 歳は 6 名と，小学生以上の回答数は 本研究に用いた特別展の開催・運営に関わった全ての方々 北原靖子・蓮見元子・川嶋健太郎・浅井義弘． 2008． 新規放 Equilibrium Research, 71 (4): 276–282． アンケートは合計 1,064 枚回収され，有効回答票が 340 枚， 原ら， 2006）．一方，質の高いあるいは均質なデータを得る きわめて少なかった．これまでの経験から，アンケートの の有形無形のご尽力にお礼申し上げる．特に特別展会場で 課後活動に対する小学児童の満足度調査－子ども自身 は じ め に 魅力的で満足度の高い博物館展示を行い，かつ展示の目 無効回答票が 10 枚，白紙が 714 枚だった（表 2）．回収し ためには，データ収集に係る手続きを可能な限り調査者以 有無にかかわらず，成人の塗り絵が張り出されることもあ 魅力的な展示を行っていただいた北九州・魚部の皆様，また， 的や理念を果たすためには，展示が来館者のニーズや期待 たアンケート数に対する有効回答票率は 32.0% であった． 外に委ねるべきではないとの指摘（Punch, 1998）には留意 るものの，成人の意見の収集に本手法を援用することは難 特別展期間中に貼り付けられた塗り絵を，アンケートの有 博物館の中核をなす役割の一つ，展示は大小さまざまな目 に応えているか，そして展示者の意図が的確に伝わってい 有効回答票内での回答率は表 2 の通りで，最も回答率が すべきであろう． しいものと思われる． 無にかかわらず一枚一枚丁寧に回収していただいた交流員 的や理念を持つ．例えば常設展示ではそれぞれの博物館がも るかなどを評価する必要がある（村田， 2003）．そのため， 高かった設問は設問 3「１．すきな虫をおしえてください」 今回行ったアンケートでは，保護者以外の手を煩わせる 今回の塗り絵を用いたアンケートに際して，展示会場で の皆様には深くお礼申し上げたい．最後に，原稿を注意深 つ目的や理念，また特別展示においては個別の目的や理念に これまでにさまざまな展示評価法が提示され，それらの効 で 97.9% であった．選択式の設問 6「４．すきなてんじを ことなく，また調査者側が多大な労力を投入することなく， 見た限りでは，男女どちらかが圧倒的に塗り絵を好んでい くお読みいただき，建設的なご意見をいただいた二名の匿 沿って行われる（里見， 2014）．これらの展示は，来館者に対 果も議論されてきた（倉田・矢島， 1993; Hein, 1998; 村田， ○でかこんでください」が 82.4％となったことを除きすべ 有効回答票を得ることができた．従って，アンケート実施 るという印象はなかった．アンケートへの回答の有無に絵 名査読者にお礼申し上げる．本研究の一部は JSPS 科研費 し豊かな教養を提供することによって，生活の質の向上を図 2003など）．例えば，展示に内包されたメッセージが来館者 て 90% 以上の回答率だった． 日を増やすことで，より多くの有効回答票を収集し，情報 柄が与える影響については，今後，同様のアンケートを複 JP17K15187 の助成によるものである． ることに繋がるほか，資料収集を通じて得られた研究成果が に伝わっているかどうかを評価するには，企画段階評価， の分析精度を高めることも可能であると考えられる．また， 数の展示会で実施し，総合的に評価していく必要がある． もたらす学問の面白さを伝える機能も併せ持っている．日々 形成的評価，総括的評価および修正的評価の実施といった （2）アンケート回答者の性別と年齢 調査者側からの回答促進策は行わなかったにも関わらず， 一方で，はがきサイズの塗り絵を用いることによる制限 Fig. 114. waltraudae. Holotype, ♀, FW 36 mm, SMFL. 115. yoshiroui. Holotype, ♂, FW 31.5 mm, KMNH. 116. yui. Holotype, ♂, の勉強の先にある世界を楽しいものと認識してもらえるなら 体系的な評価の実施が必要であるとされている（倉田・矢島， 性別は，男性が 172 枚（55.3%），女性が 139 枚（44.7%） 個別の設問に対する回答率は，一つの設問を除き 90% 以上 事項も存在するものと考えられる．例えば，アンケートの FW 31.5 mm, TY. 117. damalis, reproduced from the original description (Erichson, 1834). ば，学ぶことに対するモチベーションにもつながるだろう． 1993; 村田， 2003）．また，来館者の多様な博物館体験を， であった（表 3，有効回答 91.5%）．無効回答（表 3：無回答・ と高かった（表 2）．この高い回答率が得られた要因として，