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Despite rhe lasting impact of Hotta's colrections and subsequenr :_:* dark green longitudinai r r-:r:. lications on the raxonomy of Bornean Araceae, many of his nex. sF---_:: inflation of the latter ciur:r- :l were long known from only the type coliection. The dearth of addii:c:_r in the lower 2r5 (corrrsp r .--- collections was in part a result of the woody foci of fieldwork in Bor:.: narrowly acuminate in r-i. :=: over the past 40 years, and also in no small measure a reflection of th. ,:- male anthesis, externall-, -, , ten resrricred distribution of many of the mesophytic herbaceous planrs -:. nal venation, inrernallr- ::-: Borneo. - appearance. Spadit mri -':. , - :
hou rglass-shaped : r'. - since 2003 the author ./i has been focussing fieldwork on the Arac-,- lower halF. 1.5 cm lon-': -. and as a resuit has located new popr,rlations of most of Hotta's aroid specir. 0.5 mm diam. Dn. br:--..: Two -:- of the most interesting scbisxtatagror.rls ciescribeci by Hotta, et.,.:_, s, diameter of the fresh cr.:, : and.l. garnoandra, are detailed here. Borh were included in the mosr recer: along the spathe/spadir, ,::.,:, taxonomic revision of scbismarag/attis (Hay & yuzammi,2000), but at rh.: flar-ropped, ,a, I mrr. -.,-. . time were known only from the type collections. mm long. composcc (: -: . -: dull cream agrng ro i::< :: -, scbismatoglottis erecta M. Horta, Mem. colr. Sci. univ. Kyoto, Se:. distally 1 cm diam.. iFi,:--- : 8,32 (1965) 233, fig.5, yuzammi, A-F; Hay & Telopea 9(1): 87 _ E! connecrive rhick and :-.- .' - (2000). Type: Malaysia, Bintulr-r ('4't"; Division, Binturu, Tatau, vaile,v o: across rhe rop ber*..- :-.: .. : Sungai Keyan, Ulu Sungai Kakus, 9 Nov 1953, h[. Hirano {z M. Hotta Lt-, boid. dull cream: ,', (KYO, holo). Piate VII. l a '.:. - ims; pollen reieased i:: .::,r,:. - reddish grecn. ;a. ).' - - - Erect to climbing herb to c.z. 30 cm tall. Stetn erect to decumbe nt, ca, -> mm diam., often rooting at the nodes, with internodes to 1.5 cm long, epi- Disrribrrtion: Encrr.- - )- geal, pleionanthic, glossy reddish brown, less often glossy medium gr..rr. through the Kanou:: .-..- : - I eaue' many rogerher, each module with ca.6 leaves; peilole to 10-i4 cm Belaga. long, slender, sub-rerete and very shaliowly canalicurate, green to reddish- brown, sheathing only at the exrreme base, the wings of tf,e rheurh extend- Habirar: Terresrriai r: :' : ed into a persistent, linear-lanceolate iigular portion to 4 cm long; /amina in perhumid densr ri'::=: erect, oblong-lanceolate to narrowly elliptic, margins somewhat crispulate, 10-15 cm long x 2-3.5 cm wide, rather thinly leathery, adaxially glossy Nores: The erecr Jn.- -:. - - bright medium green, -: abaxially paler green with mid-rib and all veins immediately disrin.rr.. : darker green, the base acute and somewhat = decurrent, the apex acute and and by having pair.------, rather abruptiy acuminate for ca. 2 cm; midrib abaxially proo,ir,r..rr, adaxi- mosr closely aliied r, ally grooved, with ca, ' B primary \aterarveins on each side^, alternating with sharing che somcu :r*: - = - -- lesser interprimaries and divergi ng at crt.60o; secondary venation ablxially sparhe and clavar. ,--:..: - t rather inconspicuous, - adaxially more or less obscure, arising from the mid- in the smaller inflorts--.:--=. :. rib; tertiary venarion obscure. Inflorescence soritary; pedunci ca. 5 cm long, interstice, the conn.,:..: :. . : slender, glossy reddish brown, apically curved bringing the inflorescence to theca, and in lacki:. :,-,-: subhorizontal to noddin g. Spatbe ca. 5 cm rong; loir lparbe subcylindric, ca' 2 cm iong, glossy medium green ro reddish brown, with conspicuous
THE SARAWAK MUSEUM TOURNAL 1VNUNOI )VMVUVS AHJ IMNMIIIIM '\INESN'^I
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EI^O8 f Ugrgd (/O OF HOTTA S BORNEAN .'CH IS N AT AG LOTT I \
orher specimens seen: sarawak, sri Aman Drvision: Lubok Antu, Batanr ca, 2 cm long, green. n:i::-rx _ Ai, Nanga Sumpa, 0I' 12' 07 .6"; Il2" 02' 5I.2" ,27 July 2004, p.C, 80.,.. rhe lrmb by an abrupr - ::: ak Kisai Jeland & N. Lernbaag AR-551(SAR+spirit); Lubok Anru, Barar: male zone and appendir i:. Ar, \i7ong -: Nanga Sumpa, Enseiuai, 01o 11'00.9"; II2" 04'20.g',,5 Apr,- long. caducous br d.lic-=:- - 2005, P.C. Boyce et al. AR-li4J (SAR+spirit); Lubok Antu, Batang A:" during male anchesis. i -:-;; Nanga Sumpa, Sungai Pedali,0lo j8.9"; 1t' LI2" 03'27.A',,25 IIai fetna/e //ower zone i//. I .i-- - :: 2008, P.C, Boyce, Vong Sin Yeng OJipont ak Tisai AR-2396 (SAR+sprrit : '24, narrowly spindle-sh.:"- j; Sarikei Division: Meradong, Sungai Matob, 01" 52' 06.1";111. j0 --. 55' upwards-direcred:,,r.;,' : --- B Dec. -. 2005, P.c. Boyce et al. AR-j610 (sAR+spirit) Kapit Division: Be- like. papillate. id. rt.. -:_ laga, km 10 Bakun - Bintulu-Miri road junction, 02" 50' 5L7,'; II4. 0I among the pistrls. cori:,=. : .6" 1 1 57 , Oct 2005, P.C. Boyce, Jeland ak Kisai & Jiporu ak Tisai AR- j +i, and concentrated in rhe sl=:--= (SAR+spirit); AR-2046 (SAR+spirit); Beiaga, Belaga rcad, 43' 4>.5' : 02" drying scale-Lke, ca. 7.: ::.::, - Il3" 45' .L" 37 , 12 Oct. 2005, P.C. Boyce, Jeland ak Kisai & Jipont ak Tii_t: with sraminodes, rhrs. it:-:. j- AR-1452 (SAR+spirit). drying somewhac lu, ..-- .- I = rnle f/ower zune obconi.. j-:r __ scbisneatoglottis gamoandra M. Hotta, Mem. coll. sci. Univ. Kyoro. across. truncate, connJ.:i. :.. .: _-. ser. B, (1965) yuzammi, 32 237, frg.7, A-G; Hay & Telopea 9G): I23-L2+ above the connecrivr ::: (2000) Type: Malaysia, Sarawak, Bintulu ('.i,h') Division, ":.- - Bintulu, Tarau. pores rz. 0.6 mm diam.: , --, - along sungaiLatai, small branch of ulu Sungai Kakus, T Nov 1963,,\1. yellowl apprndix tt.u,-i': -i: -rr: Hirana Hona (KyO, plate & M. B1Z holo). VIII. tops, ca. 0.8 mm dian" F'. ::, erect to weakly reile::ei" Small herb to ca. 15 cm tall. stem + hypogeal, creeping, condensed near soil level, more elongate below, pleionanthic, cut tissues plum-purple. Leatt; Distribution: Endemr-' i- ]ir: ca. together a rosette, 5 in tightiy appressed ro the ground; petio/e to ca. Division border in an;r.:.:= : 6 cm long, D-shaped in cross-section, sheathing in th. lo*ei v4, sheath (SSf) and Belaga (NE,. open, narrowly alate above the sheath, the wing mostly crisped or not; wing of sheath fully attached, broad, apically truncare; /arnina narrowly ovaro*- Habi rat: Gallery io rir.r.:-= :-_ sagittate, 8-15 cm long x 3.5-6.5 cm wide, stiffly chartaceous, somewhar overlying shales; 40-1lr'r - :i' brittle, metallic medium to dark green, pewter-grey adaxially, sometimes with a conspicuous paier grey or green median band, much paler abaxiallr-. Nores: Srbiswatugl,,/; j, -..' . .. - apex acute, base narrowly and rather shallowly cordate with posterior lobes chartaceous leaves in a rc,s-::= l-1.5 cm long; in nature often with adventitious plantlets arising from Iate tertiary venation. In :j:-: damaged edges; midrib adaxially somewhat prominent (fresh e< dry) less with very robust, apical'r :-: so abaxially, with B primarylateralveins 5- on each side, these alternating confluent pores. The sram.:-s . with lesser interprimaries and diverging at ca. secondary 6oa; uenaliou their union. and rhe rs( F --: rather sparse, arising mostly from the midrib; lertiary aenation forming confluent with each orher. a very conspicuous tesseilate reticulum abaxrally and adaxially (fresh t drv). Inflorescences l-3 together subtended by shorr, broad cataphylls, erecr Hay and Yuzammi placec -f or more often usually somewhat recumbent by reflexing of the peduncle: S. ca/yptr"ata Group, bur .:-s :: pedtncle to ca, 4 cm long, medium green. spathe ca. 5 cm long; /otuer spatlse S. gatnoandra is there cle=:-, ::
THE SARAWAK MUSEUM JOURNAL IHE : a,:,:-, .IVNUNO{ )VM\ilrIVS EHJ tY\-ll'to '\INESNIN
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r(t llf og')'d:Irsd sIlJoT90rwt )'2 T\7O OF HOTTA'S BORNEAN SCHISMATOGLOTTIS
clear to what s. gamoandra is most closely related. The stiffly chartaceous new Schismatoglutit leaf laminae with the petiolar sheath fully attached is reminiscent of these 5B(1): 7-18. states in species of the scbismatog/ottis neruosa complex, although S. gamo- andra lacks the aromatic vegetative tissues that define s. neruosa and its Boyce, P.C. and \Wong, S-Y kin. Scbismatog/ottis gamoant/ra also approaches the s. neraosa complex in the 2001 Studies on Schiqrn: intersticial staminodes being markedly compressed and in these drying liminary obserrri somewhat scale-like. Scbismatoglonis ZdL Aroidcana. 3O:.5G(i Other specimens seen: Sarawak: Kapit Division, Pelagus, Pelagus, Jeram 2008a Studies on Homel 02o 1l' 59"; 1I3" 04' 01", \Toodpecker Trail, 1 Dec. 2004, P.C. Boyce fz species and sperule Jeland ak Kisai AR-802 (SAR+spirit); Jeram Pelagus, Rapids Trail to wa- Gardens'Ball. terfall, 02" 17'35.1";113" 03'30.08", 15 March 2005,P.C. Boyce,Jeland Siq- ak Kisai & Tisai (SAR+spirit); J. AR-I062 Belaga, Belaga road, Sungai 2008b Studies on Schisn lJnan, 02" 55' 40.0"; 44' L5.4", Boyce, II3" 12 Oct 2005, P.C. Jeland ak Scbottarum and fuh Kisai &Jipom ak Tisai AR-l427 (SAR+spirit); Bintulu, Bukit Meraki,02" Borneo. Botaniul St 45' 26.9"; 113" 39' 19.8", p. Leong pL-4)(sub AR- j287) (SAR, SING) & P. Leong PL-251(sub AR-l274), Hay. A. and Yuzammi 2000 Schismatoglomidea Telopea 9 (1): 1-lTT
ACKNOSTLEDGEMENTS Hotta, M. 7965a Itinerary of the Bur Geobot.2L: L11-l& The collaboration and support of the Sarawak Forest Department, the English. Forest Research Centre (Kuching), and the Sarawak Biodiversity Centre are much appreciated. This study is under Sarawak Forest Department l95tb Notes on Schisrran Research Permit No. NP\7.9O7.4.2(I)-101 & Park Permit No. 58/2007. Kyoto Imp. Univ-, i
I955a Notes on Schism REFERENCES Kyoto Imp. Univ-, i
I956b Araceae. Pp. 1{.Ir Bogner, J. Geobot.22: I-lO. 1978 A critical list of the aroid genera. Aroicleana l(T: 63-73 (publ. 1979). 1967 Araceae.Pp. 1)i-1, Acta Phltotax Gwha Bogner, J. and Boyce, P.C. 2009 Studies on the Schismatoglottideae (Araceae) of Borneo VI: A 1975 Notes on Bornean new Scbismatlg/lttis species from Sarawak, Malaysian Borneo. two new gener,a of t Gardens' Bu//. S ing. 60(2): l-9. 1982 On the differend \Wong, Boyce, P.C. and S.Y Schismatoglortidin 2006 Studies on Schismatoglottideae (Araceae) of Borneo I: A trio of
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33: I2l-139 - inJapanese. \\'ong, S.Y and Boyce, P.C. 2007a studies on schismatogiotideae (Araceae) of Borneo v: prelimi- nary Ecological Observations of Scbismatoglottis on the Matan: Massif. Araideana. 30: 7 1-8 l.
2001b Studies on Schismatoglottideae (Araceae) of Borneo II: Ar:- daruw crdssl.lnt, a new species from Sarawak, Malaysian Borneo. Gardens' Bu//, Sing. 5S(2): 279-286. 2008 Studies on Schismatoglotrideae (Araceae) of Borneo III: Schistnatoglottis confinis, a Putative Sister 'Iaxon to Schismatoglotti. baaensis from Sarawak, Malaysian Borneo. Gardens' Bu//. Sin,:. 60(1): 155-t63.
.,-:.' Plate VII: 5.'r.,-. .. -- elongated stem; :: -: :,- -. inflorescenrr.r[ L:.::: . - . olar shelr hs. C. l:.- :. - . . slighr l1 inHarec j:.-' - -. : rhe sparhe limb -'.-.:.:. .- - E. Inllorescencr i i- -- - lemale anthesis. ... :-t ,. -- - oping infructescei:-: -{-B G: AR-2046.
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Graeme Barker is Disnel P . bridge, Director of the ]IcD, :-' fessorial Fellow of St. John. C-. University. hc held positi,:.. -: Rome, and the Universirr' - L, His rescarch focuses on trc -: ..: :ion of modern humans. ;r..- .. -
Huw Barton is Lcctur.: :- -' Archaeology, Universirr ' : -. .. gatherer behaviour ani r--:. , usewear, and organic res:j*.' andLibya. Email: hjo-i.' = -
Besar Ketol is a iabor;: - :-" ,.,. nology, Universiti IIa,'-"'.:-. :*: versity surveys and co:1.-:- :
Efrosyni Boutsikas is L= .--= Schocrl of ELrropc.rn C- .-:. ,: sef,rches on astronor:-tr. *- -'. archaeoastronomical er -J::. :: publicatitrn as a monu-:rr.. :
Peter C. Boyce \\, 1;.- -, - higher plants of Sourh...: --.' -- from 198- ro 1001. tr -':: . b,orany. veg.etation *n* .. - Also worked on aroie . .-- . - - Other rcsearch are in ,.;-.,-- - of selected plants.
Daniel Britton rs a Fi..: , ,, Plate VIII: Scbismatoglottis garnoanclra M. Hotra. A, B. Plants in habi- by the Cambridge Ar.:.'= - - tat to show leaf variation. C. Detail of ressellate rerriary venarion. D. brittondan@hotmail. co::r Cut stem to show red internal tissue. E. Inflorescence at female an- Yvonne Michelle Campbei' thesis. F. Inflorescence near end of female anrhesis. G. Inflorescence UNIMAS. She obtainec :.: l at male anthesis; nore rhe spathe limb deliquescing at the limb/lower Nortingham. UK. H.: :..- spathe junction. H. Spadix at female anthesis, with spathe artificially gr-rage and culture of tl.rt B::," removed. A, B: AR- 1 062; C-H: AR-802.
THE SARAWAK MUSEUM JOURNAL l-: l-