Ecology Second Edition
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Ecology Second Edition N.S. Subrahmanyam A.V.S.S. Sambamurty Alpha Science International Ltd. Oxford, U.K. Contents Preface to the Second Edition vii Preface to the First Edition " ix 1. Ecology and Environment 1.1-1.20 1.1 Introduction 1.1 Branches of Ecology 1.2 Importancy of Ecology 1.2 1.2 History 1.2 1.3 Ecological Principles 1.3 Indian Ecology 1.4 1.4 Structural Concepts (Descriptive Ecology) 1.4 1.5 Functional Concepts 1.5' 1.6 Evolutionary Concepts 1.5 ° 1.7 Environmental Biotechnology 1.6 1.8 Atmosphere 1.12 2. Ecological Factors: Light and Temperature 2.1-2.28 2.1 Light 2.1 Quality of Light 2.1- Ultraviolet Light 2.2 Light in Aquatic Habitat 2.2 Effects of Light on the Plants 2.3 2.2 Transpiration 2.5 Light Absorption by Chlorophyll 2.5 Stomatal Movement 2.6 Photomorphogenesis 2.6 Phytochrome System 2.6 2.3 Phototropism 2.7 2.4 Photoperiodism 2.8 2.5 Vernalization 2.9 2.6 Bunning Hypothesis 2.10 Germination of Seeds 2.10 2.7 Measurement of Light Intensity 2.10 2.8 Light and Animals 2.11 Photoperiodism in Insects 2.11 Diapause 2.12 Seasonal Development 2.12 xii Contents 2.9 Circadian Rhythms 2.13 Behaviour Responses in Animals 2.13 2.10 Diurnation 2.14 2.11 Bioluminiscence 2.15 Types of Bioluminiscence 2.16 2.12 Temperature and Plants 2.17 2.13 Effects of Altitude 2.20 Temperature and Seed Germination 2.21 Temperature and Plant Reproduction 2.23 2.14 Thermal Constant 2.23 Thermal Stratification in Aquatic Ecosystems 2.24 2.15 Chemical Stratification 2.24 Temperature and Animals 2.25 Effect of Higher and Lower Temperatures 2.25 2.16 Temperature and Metabolism 2.25 2.17 Temperature and Reproduction 2.26 2.18 Temperature and Animal Behaviour 2.27 3. Water: Factor of Ecology 3.1-3.8 3.1 Introduction 3.1 Water and Animals 3.2 3.2 Hydrological Cycle 3.3 3.3 Measurement of Rainfalls 3.4 3.4 Humidity in India 3.5 Cloud, Mist and Fog 3.6 3.5 Frost 3.6 Thermal Insulation 3.7 3.6 Dew 3.7 4. Wind and Topographical Factors of Ecology 4.1-4.9 4.1 Wind 4.1 Flag Trees 4.1. Leaf Damage 4.2 Abrasion 4.2 Salt Spray 4.2 Lodging 4.2 Wind Erosion 4.2 Wind Speed 4.3 Windbreak or Shelter-belt 4.3 4.2 Topographic Factors 4.4 Behaviour of the Earth's Surface 4.4 4.3 Timber line or Tree Line 4.7 Exposure of the Slope to Light and Wind 4.9 Contents xiii 5. Fire Factor of Ecology 5.1-5.5 5.1 Introduction 5.1 5.2 Ecological Uses of Fire 5.4 5.3 Volcanoes 5.5 6. Soil (EDAPHIC) Factor of Ecology 6.1-6.31 6.1 Introduction 6.1 6.2 Residual and Transported Soils 6.2 6.3 Weathering 6.2 6.4 Soil Profile 6.4 6.5 Soil Texture 6.6 6.6 Soil Classification 6.7 Soil Moisture 6.10 6.7 Soil Water 6.11 Water Vapour 6.12 6.8 Physically and Physiologically Dry Soils 6.b3 Temporary and Permanent Wilting 6.13 Soil Density and Porosity 6.13 Soil Porocity 6.14 6.9 Chalkland Ecology 6.16 6.10 Soil Organic Matter 6.16 6.11 Soil Atmosphere 6.17 - 6.12 Soil Temperature 6.18 6.13 Soil Microbes 6.20 6.14 Soil Animals 6.20 Fauna and Flora of the Soil 6.20 6.15 Soil Erosion 6.21 Normal Erosion 6.21 Accelerated Soil Erosion 6.21 Agencies causing Soil Erosion 6.21 6.16 Soil Conservation 6.24 7. Biotic Factors of Ecology 7.1-7.21 7.1 Introduction 7.1 7.2 Mutualism 7.1 Mutualism between Plants and Plants 7.1 7.3 Syntrophy in Photosynthetic Bacteria 7.2 7.4 Mycorrhizae 7.2 7.5 Commensalism 7.7 Protocooperation (non-obligatory Mutualism) 7.9 7.6 Competition 7.9 7.7 Amensalism (Antibiosis) 7.10 Amensalism in Microorganisms 7.12 xiv Contents 7.8 Allelopathy 7.12 7.9 Predation 7.13 7.10 Slobodkin's Work 7.15 7.11 Insectivorous Plants 7.16 The Pitcher Plants 7.17 7.12 Myremecophily 7.18 7.13 Saprophytes 7.18 7.14 Ecology of Herbivores 7.19 Plant Defenses 7.20 Grazing and Browsing 7.21 8. Animal Inter Relationships 8.1-8.7 8.1 Introduction 8.1 8.2 Intraspecific Relationships 8.1 8.3 Interspecific Relationships 8.1 The Symbiosis of Dinoflagellates 8.3 8.4 Insect Symbiosis with Microorganisms 8.3 8.5 Mutualism without Continuous Contact 8.4 Commensalism with Continuous Contact 8.5 Commensalism without Continuous Contact 8.5 8.6 Antagonism 8.5 8.7 Parasitic Adaptations 8.6 9. Animal Adaptations 9.1-9.10 9.1 Introduction 9.1 9.2 Pread'aptations 9.1 * . Desert Adaptations 9.1 9.3 Cursorial Adaptations 9.3 Change in Foot Posture and Loss of Digits 9.3 Fossorial Adaptations 9.4 9.4 Arboreal Adaptations 9.5 Volant Adaptations 9.5 9.5 Aquatic Adaptations 9.6 9.6 Adaptive Radiation 9.8 10. Ecological Concepts of the Species 10.1-10.20 10.1 Taxonomic Species 10.1 Ecological Species 10.1 10.2 Ecocline 10.2 10.3 Gregor's Work 10.3 10.4 Types of Ecotypes 10.4 10.5 Acclimation 10.5 Ecological Amplitude 10.6 10.6 The law of Tolerance 10.8 Contents XV 10.7 Ecological Equivalents 10.8 10.8 Dame Concept 10.9 10.9 Niche 10.10 Multidimensional or Hypervolume 10.13 10.10 Flora of Delhi 10.15 10.11 Flora of Vishakhapatnam 10.17 10.12 Flora of Osmanabad (Andhra Pradesh) 10.19 11. Population Ecology ' 11.1-11.17 11.1 Introduction 11.1 11.2 Influences on Populations 11.2 Population Growth and Density 11.2 Natality (Birth Rate) 11.2 11.3 Population Density 11.2 » 11.4 Mortality (Death Rate) 11.3 Population Growth Form 11.7 Age Distribution 11.7 11.5 Population Fluctuations 11.8 11.6 Population Dispersal 11.10 Home Range and Territoriality 11.11 11.7 Home Range and Territoriality in Animals 1*1.12 11.8 Autecology 11.14 ' Environmental Complex 11.14 11.9 Seed Germination and-Reproductive Capacity 11.14 11.10 Seed Dormancy 11.16 12. Nature and Structure of Plant Communities 12.1-12.10 12.1 Introduction 12.1 12.2 Organismic Concept 12.2 12.3 Individualistic Concept 12.3 12.4 Characteristics of Communities 12.3 12.5 Fidelity 12.8 12.6 Growth Form and Structure 12.8 12.7 Stratification 12.9 12.8 Ecotone 12.10 13. Analysis of Plant Communities: Community Classification 13.1-13.19 13.1 Introduction 13.1 13.2 Structure of Plant Community 13.1 13.3 Qualitative Characteristics 13.3 13.4 Physiognomy and Pattern 13.3 13.5 Biological Spectrum 13.6 Raunkiaer's Leaf Size Classification 13.8 xvi Contents 13.6 Quantitative Characteristics 13.8 Density 13.11 Basal Area 13.12 13.7 Phytosociological Methods 13.14 13.8 Line Intercept, Strip Transect and Bisect Methods 13.15 13.9 The Point Method 13.17 Distance (Plotless) Methods 13.17 Nearest Individual Method 13.17 14. Methods of Studying Animal Communities 14.1-14.9 14.1 Density 14.1 14.2 Estimation of Phytoplankton and Zooplankton in water bodies 14.2 14.3 Determination of Frequency of Phytoplankton 14.3 14.4 Zooplankton 14.4 14.5 Benthic Macro-invertebrates 14.6 14.6 Estimation of Secondary Production in Animals 14.6 Nearest Neighbour Techniques 14.6 Spatial Pattern 14.7 15. Plant Succession 15.1-15.13 15.1 Introduction 15.1 15.2 Trends in Succession 15.1 15.3 Process of Succession 15.4 . ' 15.4 Chronosequence Versus Toposequence 15.6 15.5 Cyclical Changes in vegetation 15.6 15.6 Xerarch 15.8 15.7 Succession on Sand Dunes (Psammosere) 15.9 15.8 Heterotrophic Succesion 15.10 15.9 Succession on Dung 15.11 15.10 Climax Community 15.12 16. Ecosystem Ecology 16.1-16.27 16.1 Introduction 16.1 16.2 Types of Ecosystems 16.2 16.3 Structure of Ecosystem 16.2 Producer Organisms (Autotrophic Component) 16.3 16.4 Decomposers and Transformers 16.3 Flow of Energy 16.4 16.5 The Ecosystem as a Cybernetic System 16.5 Homeostasis 16.5 16.6 Aquatic Ecosystems 16.6 Fresh Water Environments 16.6 Streams and Rivers 16.7 Lakes and Ponds 16.7 Contents xvii 16.7 The Lake Ecosystem 16.7 The Pond as an Ecosystem 16.9 16.8 Categories of Pond Flora and Fauna 16.10 Food Chains and Energy Flow 16.10 16.9 Types of Food Chains 16.13 Food Web 16.14 Detritus Food Web 16.16 16.10 Ecological Pyramids 16.16 16.11 Energy Flow in the Ecosystem 16.18 Lindeman's Work 16.21 Golley's Work 16.22 16.12 Ecological Efficiency 16.22 16.13 Biogeochemical Cycles 16.23 Hydrologic Cycle 16.24 Carbon Cycle 16.24 Nitrogen Cycle 16.24 ° Oxygen Cycle 16.25 Phosphorus Cycle 16.26 Sulphur Cycle 16.27 17.